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A rich fossil assemblage discovered from the classic Pennsylvanian locality of Joggins, Nova Scotia, is here described for the first time. The 2 m-thick Hebert sandstone, within the lower Joggins Formation of early Langsettian age, is the most productive stratum at Joggins of terrestrial tetrapods and the pulmonate gastropod Dendropupa vetusta exclusive of the historic fossil forest locality of Dawson and Lyell, and the most productive Pennsylvanian locality in the world of the large unionoid bivalve genus Archanodon. Although the land snail–archanodontid bivalve–tetrapod assemblage of the Hebert Sandstone comprises a unique late Paleozoic terrestrial dryland biota, individual taxa were not endemic to drylands alone. [ABSTRACT FROM AUTHOR]