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Genus Rhabdotis Burmeister, 1842 A diagnosis of the genus, a key to the nine species hitherto known, their types and distribution patterns were given by Holm (1991a). Since this date more sixteen species were described, as well as four subspecies from different African countries (see updates in Beinhundner 2017a). Within this assemblage, only two species were recorded for Angola with certitude (Paulino de Oliveira 1882; Rigout & Allard 1992; Beinhundner 2017a; Schoolmeesters 2018)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 39, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Holm, E. (1991 a) Synonymic notes on African Cetoniinae VI: Genus Rhabdotis Burmeister (Coleoptera: Scarabaeidae). Annals of the Transvaal Museum, 35 (22), 313 - 323.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Rigout, J. & Allard, V. (1992) Cetoniini 2: Les Coleopteres du monde. Sciences Naturelles Venette, 12, 1 - 135.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)"]}

Oplostomus fuligineus (Olivier, 1789) (Fig. 14c) Distribution: ANG, DOC, KEN, MOZ, NAB, NIG, RSA, SEN, ZAM, ZIM. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Ferreira 1965). Material examined: Angola, 1 ♀, N�� 1907 (blue label), old collection, box n�� XIV-83, MZUC. Remarks. The species seems to present a wide distribution in Africa (e.g. Schenkling 1921, Gomes Alves 1959, see Oldroyd & Allsopp 2017 also) and is known as the ���Large Hive Beetle���. The adults are associated with honey bees and paper wasps, while larvae and pupae live in dung (P��ringuey 1907; Scholtz & Holm 1985; Oldroyd & Allsopp 2017). The larvae description and life cycle were provided by Donaldson (1989) and Oldroyd & Allsopp (2017). A review on several aspects of its biology was also provided by these authors. The species seems to have potential to become invasive and highly damaging to beekeeping worldwide (Oldroyd & Allsopp 2017)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 65, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Schenkling, S. (1921) Scarabaeidae: Cetoninae. In: Junk, W. (Ed.), Coleopterorum Catalogus, 72, W. Junk, Berlin, 431 pp.","Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8.","Oldroyd, B. P. & Allsopp, M. H. (2017) Risk assessment for large African hive beetles (Oplostomus spp.). A review. Apidologie, 48, 495 - 503. https: // doi. org / 10.1007 / s 13592 - 017 - 0493 - 7","Peringuey, L. (1907) Descriptive catalogue of the Coleoptera of South Africa. Fam. Lucanidae. Fam. Scarabaeidae. Sub-families: Sericinae, Melolonthinae, Trichiinae, Valginae, Cetoniinae. Transactions of the South African Philosophical Society, 13, 289 - 546, pl. 47.","Scholtz, C. H. & Holm, E. (1985) Insects of Southern Africa. University of Pretoria, Pretoria, 502 pp.","Donaldson, M. I. J. (1989) Oplostomus fuligineus (Coleoptera Scarabaeidae): life cycle and biology under laboratory conditions, and its occurrence in bee hives. The Coleopterists Bulletin, 43 (2), 177 - 182."]}

Genus Hegemus J. Thomson, 1881 There are three species in the genus treated and illustrated by Lachaume (1983, but see also Wiebes 1968). An identification key for the species was given by Wiebes (1968). One species is known from Angola (Ferreira 1965; Beihnundner 2017a)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 110, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Lachaume, G. (1983) Goliathini 1. Les Coleopteres du Monde. Vol. 3. Goliathus-Argyrophegges-Fornasinius-Hegemus. Sciences Nat, Compiegne, 67 pp.","Wiebes, J. T. (1968) Catalogue of the Coleoptera Cetoniidae in the Leiden Museum. 1. Goliathus Lamarck, sensu lato. Zoologische Mededelingen, 43 (3), 19 - 40, 2 pls.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Phonotaenia scalaris scalaris (Gory & Percheron, 1833) (Fig. 7f) Distribution: ANG, BEN, CAM, CON, DOC, GAB, GHA, KEN, RCA, RRW, TOG, UGA. Distribution in Angola (Provinces): 1) CUS, LNO. Historic records: ��� Angola (without locality) (Paulino de Oliveira 1882 sub Gametis scalaris Gory & Perch.); ��� Luanda: rio Muka (not LUA, but LNO) (Gomes Alves 1959; Ferreira 1965; see Remarks); ��� Kwanza Sul (without locality) (CUS), Dundo (LNO) (Antoine 2005a). Material examined: Muita River (Luembe) (07�� 48��S, 21�� 26��E, 1260 m alt., 69) (LNO), -. VI.1948, 1 ex., N�� 2137, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC. Remarks. The adult specimen of Luembe was sampled in a forest (registered information). Gomes Alves (1959) recorded this specimen for River Muka (Luanda Province), but it was an erroneous interpretation of the register book of IICT which gives the following information: ���Rio Muka (afluente do Luembe���Lunda NE)���= [Muka River (tributary of Luembe���Lunda NE]. The true tributary of Luembe (River) is Muita River, not Muka River. Moreover, the reference to Lunda NE reinforces this interpretation. Some ecological remarks were provided for the subspecies Phonotaenia scalaris latefasciata Moser, 1910 in Sierra Leone by Malec (2015)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 38, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Gory, H. & Percheron, A. (1833) Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris, 406 pp. https: // doi. org / 10.5962 / bhl. title. 8957","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Antoine, Ph. (2005 a) Revision du genre Phonotaenia Kraatz, 1880, sensu lato (Coleoptera, Cetoniidae). Coleopteres, 11 (21), 227 - 265.","Malec, P. (2015) The Cetoniinae fauna of Sierra Leone and the basic guidelines to captive breeding of these beetles (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 7, 35 - 76."]}

Eudicella (Cyprolais) quadrimaculata (Fabricius, 1781) Distribution: ANG (?), CAM, GUI, RCA, RCI. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Paulino de Oliveira 1882 sub Cerathorrhina 4-maculata F.). ��� Angola (without locality) (Ferreira 1965 sub Coelorrhina pythia Kolbe). Material examined: none. Remarks. The species was well characterized (head and parameres) by Antoine (1996b). Ferreira (1965) reported the species for Angola under the name of Coelorrhina pythia Kolbe, 1899. Probably this species was misinterpreted by Paulino de Oliveira (1882) and Ferreira (1965) with C. loricata and its occurrence in Angola must be considered with some caution., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 91, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Antoine, Ph. (1996 b) Identification de Cetonia quadrimaculata Fabricius (Coleoptera, Cetoniidae). Revue Francaise d'Entomologie, Nouvelle Serie, 18 (3), 111 - 114."]}

Genus Spelaiorrhina Lansberge, 1886 The genus was revised by Allard (1991) who validated two species (S. lateralis Kolbe, 1892 and S. similis Moser, 1902) that were not recognized by the subsequent authors (e.g. Holm & Marais 1992; Krajcik 2008a). The genus is considered monotypic and the single species was recorded for Angola (Lansberge 1886; Ferreira 1965)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 104, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Lansberge, J. W. (1886) Scarabaeides, buprestides et cerambycides de l`Afrique Occidentale, envoyes au Musee de Leyden par MM. Veth et Van der Kellen. Notes from the Leyden Museum, 8, 69 - 120.","Allard, V. (1991) Goliathini 4: Les Coleopteres du monde. Sciences Naturelles de Venette, 11, 1 - 142.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Pseudinca incoides (J. Thomson, 1860) Distribution: ANG, BEN, CAM, CON, DOC, EQG, GAB, GHA, RCA, RCI, NRE, TOG. Distribution in Angola (Provinces): 1) CAB, ZAI. Historic records: ��� Cabinda (CAB) (Paulino de Oliveira 1882 sub Diplognatha incoides Thoms.); ���S. Salvador (ZAI) (Valck Lucassen 1933). Material examined: none. Remarks. The species in Angola seems restricted to its northwesternmost territory. Tchibozo et al. (2008) and Joly (2018) provided biodiversity information (abundance, habitat preference) of this species for Benin and Ghana, respectively., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 76, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Valck Lucassen, F. T. (1933) Revision du genre Pseudinca Kraatz, (Scarabaeidae: Cetoniinae). Tijdschrift voor Entomologie, 86, 277 - 308.","Tchibozo, S., Aberlenc, H. - P., Ryckewaert, Ph. & Le Gall, Ph. (2008) Premiere evaluation de la biodiversite des Odonates, des Cetoines et des Rhopaloceres de la foret marecageuse de Lokoli (Sud Benin). Bulletin de la Societe Entomologique de France, 113 (4), 497 - 509.","Joly, C. (2018) Deuxieme contribution a la connaissance des Cetoniidae du Ghana, sous-familles des Cetoniinae et des Trichinae (Coleoptera, Scarabaeidae). Lambillionea, 118 (1), 28 - 89."]}

Genus Caelorrhina Hope, 1841 The genus is close to Anisorrhina and Tmesorrhina, sharing some characters with them (see Holm & Marais 1992). According to Holm & Marais (1992) the genus was never revised since Bourgoin (1921), thus modern revision is urgently needed. Most of the species were treated under the genus Eccoptocnemis (e.g. see Allard 1991). Beinhundner (2017a) recognized eleven valid species, three of them recorded for Angola, one being endemic (Ferreira 1965). We give two new species records for this country., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 79, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Bourgoin, A. (1921) Resultats scientifiques. Insectes Coleopteres. Cetoniini. In: Babault, G. Voyage de Guy Babault en Afrique Orientale (1912 - 1913), Paris, 2, 1 - 46.","Allard, V. (1991) Goliathini 4: Les Coleopteres du monde. Sciences Naturelles de Venette, 11, 1 - 142.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

* Leucocelis (Leucocelis) niveoguttata (Blanchard, 1850) (Figs. 10e, f) Distribution: ANG, BOW, MOZ, NAB, RSA, ZIM. Distribution in Angola (Provinces): 2) BIE, HUI. Historic records: none. New localities reported: Chiapepe (coord. not available, 1280 m alt., 378) (HUI), 10.XII.2012, 1 ♂, P. Sch��le leg., PMC (P. Malec, personal communication). Material examined: Angola, 1 ex., N�� 1907 (blue label), n�� 201 (white label), old collection, box n�� IX-78, MZUC; Satchijamba (13�� 44��54.69���� S, 17�� 10��35.03���� E, 1579 m alt., 303) (BIe), 3.XI.2014, 1♂, 1♀, BT, 29.X.- 3.XI.2014, 1♀, CT, A. Serrano leg. & R. Capela, ASC. Remarks. The species was restablished by Antoine (2006b). The adult female specimen of Satchijamba fits very well with the description given by Antoine (2006b) and the parameres of the adult male specimen conforms with the same structures of L. vitticollis (Boheman, 1857) given by Holm & Marais (1992), a species which was synonymized with L. niveoguttata by Antoine (2006b). The Satchijamba adult specimens were sampled by BT and CT (Figs. 1a, b) in an open secondary forest. See also remarks under Pachnoda discolour kolbei. Ecological data were provided for the species (under the name of L. vitticollis) by Holm & Marais (1992). It is a new species record for Angola and Bi�� and Hu��la Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 49-50, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. (2006 b) Contribution a la connaissance des Cetoniidae africains II (Coleoptera, Scarabaeoidea). Coleopteres, 12 (30), 399 - 414.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

Gnathocera cruda lurida (Janson, 1877) Distribution: ANG, CON, DOC, TAN. Distribution in Angola (Provinces): 1) LNO. Historic records: ��� Angola (without locality) (Janson 1877b sub G. lurida); ��� Angola (without locality) (Basilewsky 1949 sub G. lurida); ���Dundo (LNO) (Gomes Alves 1959 and Ferreira 1965 sub G. lurida Janson). Material examined: none. Remarks. Antoine (2002a) treated this taxon as a subspecies of G. cruda while Krajcik (2008a), Beinhundner (2017a) and Schoolmeesters (2018) elevated it to the species level, recognising three subspecies. Gnathocera lurida was described originally from Angola based in specimens of ���Dr. Welwitsch who found it in considerable numbers��� Janson (1877b). Gomes Alves (1959) recorded the subspecies for Angola (Dundo) based on specimens of G. cruda gracilis (see Historic records under this subspecies). Biological or ecological data are not available for this species., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 95-96, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Janson, O. E. (1877 b) Notices of new or little known Cetoniidae No. 3. Cistula Entomologica, 2, 247 - 264.","Basilewsky, P. (1949) Les Gnathocera du Congo belge (Col. Cetoniinae). Revue de Zoologie et de Botanique Africaines, 28 (3 - 4), 259 - 273.","Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)"]}

Genus Eudicella White, 1839 The genus in the strict sense was revised by De Palma (2009), who provided an identification key. One year after, De Palma (2010) provides a new dichotomic key to all Eudicella sensu stricto. According to this work fourteen species and two subspecies remain valid. However, Beinhundner (2017a) recognized nineteen species as valid. The genus Cyprolais J. Thomson, 1880 was revised by Holm (1993b) who relegated it to the level of subgenus of Eudicella, a status followed by De Palma (2009) and Beinhundner (2017a). Oram (2015) says ��� Cyprolais J. Thomson, 1880 is usually treated as a subgenus of Eudicella (e.g. Krajcik, 2008���)���. However, Krajcik (2008a) raised the taxon to the genus level. Beinhundner (2017a) recognized fifteen species in Cyprolais subgenus. Two species of Eudicella (Eudicella) were recorded for Angola (Ferreira 1965; Beinhundner 2017a) and other two species of Eudicella (Cy- prolais) (Paulino de Oliveira 1882; Ferreira 1965), but one must be considered with caution., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 90, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["De Palma, M. (2009) Taxonomic revision of Eudicella White (Coleoptera: Cetoniinae) and Iconographic Catalogue. Natura Edizioni Scientifiche, Bologna, 48 pp.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Thomson, J. (1880) Mecynorhinus haroldii (sp. n.), Col. Bulletin de la Societe Entomologique de France, 1880, 110 - 111.","Holm, E. (1993 b) On the genera of African Cetoniinae I: Eudicella White, and the related genera with horned males (Coleoptera: Scarabaeidae). Journal of African Zoology, 107 (1), 65 - 81.","Oram, D. A (2015) A new species of Eudicella White subgenus Cyprolais Thomson from the Republic of Zambia, Africa (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 8, 77 - 84.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52."]}

Pedinorrhina viridicollis (Burgeon, 1932) Distribution: ANG, CAM, DOC, RCA. Distribution in Angola (Provinces): 1) LNO. Historic records: ���Dundo (LNO) (Antoine 2008). Material examined: none. Remarks. A species recorded for Dundo region in Angola where it seems overlap in some localities with the precedent species (Antoine 2008). Ecological data are not available for the species., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 103, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Burgeon, L. (1932) Catalogues raisonnes de la faune entomologique du Congo Belge. Coleopteres-Scarabeides- Cetoniines. Annales du Musee Royal du Congo Belge, Sciences Zoologiques, Series 3, 2 (2), 47 - 128.","Antoine, Ph. (2008) Remarques taxonomique sur le genre Pedinorrhina Kraatz, 1880 (Coleoptera, Cetoniidae). Coleopteres, 14 (3), 27 - 42."]}

Genus Cheirolasia Westwood, 1843 The genus is monotypic and was revised by Holm (1993a). It is represented in Angola by two subspecies (e.g. Paulino de Oliveira 1882; Bourgoin 1925)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 82, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Holm, E. (1993 a) On the genera of African Cetoniinae: Anisorrhina Westwood 1842, Melinesthes Kraatz 1880 and lnhambane Peringuey 1907. Tropical Zoology, 6, 165 - 177. https: // doi. org / 10.1080 / 03946975.1993.10539217","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Bourgoin, A. (1925) Coleopteres: Scarabaeidae. Tribu: Cetoniini. In: Mission Rohan-Chabot Angola et Rhodesia. 4 (3). Impremerie Nationale, Paris, pp. 141 - 146."]}

Genus Lissogenius Schaum, 1844 The genus was not revised and has seven recognized species (Krajcik 2008b). The species of this genus are myrmecophylous. There is only one species recorded for Angola (Moser 1911)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 61, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Krajcik, M. (2008 b) Cetoniidae of the World. Catalogue - Part II. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 72 + I-XXIII.","Moser, J. (1911) Beitrag zur kenntnis der Cetoniden IX. Annales de la Societe Entomologique de Belgique, 55, 119 - 129. https: // doi. org / 10.1002 / mmnd. 48019110505"]}

Genus Dicronorhina Hope, 1837 The genus was revised by Lekkerkerk & Krikken (1986). Beinhundner (2017a) recognized five valid species, two of them recorded for Angola (Ferreira 1965; Lekkerkerk & Krikken 1986)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 89, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Lekkerkerk, R. W. & Krikken, J. (1986) Taxonomic review of the Afrotropical genus Dicronorrhina Hope, with notes on its relatives (coleopteran: Cetoniidae). Zoologische Verhandelingen, 233, 3 - 46.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Genus Dischista Burmeister, 1842 The genus was revised by Rigout (1989), but some species/subspecies were described after this work (e.g. Antoine 2004a, 2010; Beinhundner 1998, 2006a). Eleven valid species are recognized (Beinhundner 2017a) of which three were recorded for Angola (Paulino de Oliveira 1882; Ferreira 1965)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 22, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Rigout, J. (1989) Cetoniini 1: Les Coleopteres du monde. Sciences Naturelles Venette, 9, 1 - 135.","Antoine, Ph. (2004 a) Contribution a la connaissance des Cetoniidae du Kenya (Coleoptera, Scarabaeoidea). Coleopteres, 10 (2), 13 - 28.","Antoine, Ph. (2010) Contribution a la connaissance des Cetoniidae de Zambie et regions limitrophes (Coleoptera, Scarabaeoidea). Coleopteres, 16 (9), 101 - 115.","Beinhundner, G. (1998) Beschreibung zwei neuer arten der gattung Dischista aus Ost Afrika. Lambillionea, 98 (3), 377 - 384.","Beinhundner, G. (2006 a) Description of a new species of Dischista Burmeister, 1842 and of a new species of Pseudinca Kraatz, 1880, from R. D. Congo. Lambillionea, 106 (1), 19 - 25.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Pseudoclinteria infuscata (Gory & Percheron, 1833) (Figs. 27c, d) Distribution: ANG, DOC, ETI, KEN, MAW, MOZ, RSA, SEN, TAN, ZIM, ZAM. Distribution in Angola (Provinces): 1) HUI; 2) BIE. Historic records: ���Angola (without locality) (Paulino de Oliveira 1882 sub Clinteria infuscata Gory & Percheron); ���Huilla (without locality) (HUI) (Schoch 1896 and Holm 1991b sub Clinteria maculicollis Schoch); ������Port. O. Afr.��� (Angola?, without locality) (Kuntzen 1930 and Holm 1991b sub Clinteria infuscata goetzei v. peringueyi Kuntzen); ��� Hu��la (without locality) (HUI) [Ferreira, 1965 sub Clinteria infuscata (Gory & Percheron) ab. macullicolis Schoch]. New localities reported: Negola (3,5 km SW) (14�� 08��53���� S, 14�� 28��16���� E, 1613 m alt., 317) (HUI), 11- 12.XI.2013, 1♂, P. Sch��le leg., PMC (P. Malec, personal communication). Material examined: Angola, 3 exs, N�� 1907 (blue labels), n��s 154, 187 and 200 (white labels), old collection, box n�� IX-78, MZUC; Satchijamba (13�� 44��54.69���� S, 17�� 10��35.03���� E, 1579 m alt., 303) (BIE), 3.XI.2014, 3♂, 3♀, BT, 29.X.-3.XI.2014, 1♂, 3♀, CT, A. Serrano leg. & R. Capela, ASC; Mumbu�� (Kwanza River spring) (13�� 49��41���� S, 17�� 19��26���� E, 1511 m alt., 303) (BIE), 8-17.XI.2015, 1♂, CT, A. Serrano leg. & R. Capela, ASC. Remarks. The species is very polychromatic. The adult specimens in Satchijamba were collected by BT and CT (Figs. 1a, b) in an open secondary forest (Fig. 1e). See also the remarks under Pachnoda discolour kolbei. Some ecological data were provided by Holm & Marais (1992). It is a new species record for the Bi�� Province., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 112-113, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Gory, H. & Percheron, A. (1833) Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris, 406 pp. https: // doi. org / 10.5962 / bhl. title. 8957","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Schoch, G. (1896) Einige neue Cetoniden. Entomologische Nachrichten, 22, 328 - 332.","Holm, E. (1991 b) Synonymic notes on the African Cetoniinae (Coleoptera: Scarabaeidae) VIII: tribe Gymnetini-genera Stethodesma Bainbridge, Amazula Kraatz and Pseudoclinteria. Cimbebasia, 13, 117 - 125.","Kuntzen, H. (1930) Analytische tabelle der unterarten und formen der Pseudoclinteria infuscata Gory & Perch. Mitteilungen des Zoologischen Museum Berlin, 15, 573 - 579.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

Amaurina spoliata (Harold, 1879) (Figs. 8i, 9a) Distribution: ANG, DOC. Distribution in Angola (Provinces): 1) HUI, MAL; 2) BGU, BIE, HUA. Historic records: ���Pungo Andongo (MAL) (Harold 1879 sub Leucocelis spoliata Har.); ��� Angola (without locality) (Ferreira 1965); ���S�� da Bandeira (=Lubango) (HUI) (Gomes Alves 1973 sub Leucocelis spoliata Har.). New localities reported: Lubango (10 km W) (14�� 50��32.5���� S, 13�� 24��06.2���� E, 2103 m alt., 335) (HUI), 1.XII.2017, 1 ♂, P. Sch��le leg., PMC (P. Malec, personal communication). Material examined: Angola, 10 exs, N�� 1907 (blue labels), n��s 155 (2 exs), 161, 162, 163, 210, 215, 216, 217 and 218 (white labels), old collection, box n�� IX-78, MZUC; S�� da Bandeira (= Lubango) (14�� 55��S, 13�� 30��E, 1750 m alt., 336) (HUI), 25.XI.1957, 1 ex., N�� 2970, Leucocelis spoliata Harold, Estudos Ap��colas do Ultramar n�� 122, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Chianga (12�� 44��S, 15�� 50��E, 1740 m alt., 256) (HUA), 26.IX.1964, 1 ex., N�� E.5695, n�� 606, 22.X.1964, 1 ex., N�� E.5729, n�� 605, E. Fonseca leg., JPCC, MUH- NAC; Chioa (coords not available) (BGU), 22.X.1964, 1 ex., N�� E.5728, n�� 607, C. Matos leg., JPCC, MUHNAC; Capingana���Satchijamba (13�� 42��47.18���� S, 17�� 05��23.81���� E, 1507m alt., 303) (BIE), 8.IV.2014, 1♂, DO, A. Serra- no leg., ASC; Carima (Cacuso���Pungandongo) (09�� 35��00���� S, 15�� 42��01���� E, 1117 m alt., 130) (MAL), 17.XI.2017, 9♂, 5♀, 20.XI.2017, 16♂, 4♀, 21.XI.2017, 2♂, 1♀, DO, A. Serrano & R. Capela leg., ASC. Remarks. The species was originally described on the basis of specimens from Angola (Harold 1879; Antoine 2000), but is also known from DOC (Beinhundner 2017a). The adult specimens from Chianga were found on Pandiaka welwitschii (Amaranthaceae) and Monotes calomerus (label information).The adult specimen of Capingana���Satchijamba was captured on a yellow flower and the Carima adult specimens were sampled feeding on the basis of tomentose leafs of spontaneous herbaceous plants (Fig. 9a), all from secondary open forests. In the latter locality (Fig. 10a), the species was abundant and syntopic with Leucocelis (Leucocelis) haemorrhoidalis (Fabricius, 1775), Plagiochilus angustatus (Westwood, 1873) and Cymophorus sp. It is a new species record for the Benguela, Bi�� and Huambo Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 43-45, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Harold, E. (1879) Bericht uber die von Herren A. v. Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Coleopteren. Coleopterologische Hefte, 16, 1 - 224.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Gomes Alves, M. G. (1973) Coleopteros da Africa Portuguesa (Lucanidae, Scarabaeidae: Cetoniinae; Chironinae, Phaenomerinae, Cicindelidae e Chrysomelidae: Hispinae). In: Livro de homenagem ao Prof. Fernando Frade por ocasiao do seu 70 º aniversario. Junta de Investigacoes do Ultramar, Lisboa, pp. 27 - 58.","Antoine, Ph. (2000) Nouveaux Cetonides de l`Angola (Coleoptera, Cetoniidae). Entomologia Africana, 5 (2), 20 - 29.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp."]}

Caelorrhina angolensis (Sch��rhoff, 1942) Distribution: ANG. Distribution in Angola (Provinces): 1) HUA; 2) HUI. Historic records: ���Caconda (HUA) (Sch��rhoff 1942 sub Eccoptocnemis angolensis); ��� Angola (without locality) (Ferreira 1965 sub Eccoptocnemis angolensis Sch��rhoff); ��� Angola (without locality) (Allard 1991 sub Eccoptocnemis barthi angolensis Sch��rhoff). New localities reported: (HUI) (S. Rojkoff, personal communication sub Caelorrhina barthi angolensis). Material examined: none. Remarks. An Angolan endemic species described based on one male and one female specimens from Caconda (Sch��rhoff 1942). Allard (1991) considered this taxon as a subspecies of Caelorrhina barthi, a criterion not followed by Krajcik (2008a) (but see New localities reported). No ecological data are available for this species. It is a new record for the Hu��la Province., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 79-80, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Schurhoff, P. N. (1942) Beitrage zur kenntnis der Cetoniden (Col.) IX. Mitteilungen der Munchner Entomologischen Gesellschaft, 32, 279 - 293.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Allard, V. (1991) Goliathini 4: Les Coleopteres du monde. Sciences Naturelles de Venette, 11, 1 - 142.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI."]}

Eudicella (Cyprolais) loricata loricata (Janson, 1877) (Figs. 20f, g) Distribution: ANG, DOC, MAW, RCA, TAN, ZAM. Distribution in Angola (Provinces): 1) HUI, 2) BIE. Historic records: ��� Angola (HUI) [Oram 2015, syntype]. Material examined: Angola, 1 ♂, N�� 1907 (blue label), n�� 60 (white label), Welwitch leg., 1 ♂, N�� 1907 (blue label), n�� 64 (white label), old collection, box n�� IX-78, MZUC; Angola, 1 ♀, N�� 1907 (blue label), old collection, box n�� IX-78, MZUC; Mumbu�� (Cuanza riverhead) (13�� 49��39.97���� S, 17�� 19��20.32���� E, 1524 m alt., 303) (BIE), 3���9.IV.2014, 1♂, CT, A. Serrano & R. Capela leg., ASC. Remarks. The species was originally described from Angola based on a specimen collected by Welwitsch among a dozen taken in a cluster on a flower (Janson 1877a). Eight subspecies are recognized in this species (Beinhundner 2017a). Antoine (1996b) and Oram (2015) gave some morphological details of head and parameres of this species. Single male of Mumbu�� (Fig. 20g) was collected by means of CT (e.g. Fig. 1b) in an open secondary forest together with A. picturata. Some ecological data are provided by Holm & Marais (1992). It is a new species record for the Bi�� Province., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 90, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Oram, D. A (2015) A new species of Eudicella White subgenus Cyprolais Thomson from the Republic of Zambia, Africa (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 8, 77 - 84.","Janson, O. E. (1877 a) Notices of new or little known Cetoniidae No. 2. Cistula Entomologica, 2, 141 - 147.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Antoine, Ph. (1996 b) Identification de Cetonia quadrimaculata Fabricius (Coleoptera, Cetoniidae). Revue Francaise d'Entomologie, Nouvelle Serie, 18 (3), 111 - 114.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

Pachnoda cordata cordata (Drury, 1773) Distribution: ANG, GHA, GUI, MOZ, RCI, RSA, SLN, TOG. Distribution in Angola (Provinces): 1) LNO. Historic records: ���Quango (=Cuango) (LNO) (Quedenfeldt 1884); ��� Angola (without locality) (Ferreira 1965). Material examined: none. Remarks. The species is polytypic with six to eight known subspecies in Africa (Krajcik 2008a; Beinhundner 2017a; Schoolmeesters 2018). Some biological and ecological data were provided by Holm & Marais (1992)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 28, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Quedenfeldt, G. (1884) Verzeich niss der von Herrn Major a. D. von Mechow in Angola und am Quango-Strom 1878 - 1881 gesammelten Pectinicornen und Lamellicornen. Berliner Entomologiche Zeitschrift, 28 (2), 265 - 340. https: // doi. org / 10.1002 / mmnd. 47918840107","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

* Chthonobius conspersus Burmeister, 1847 (Fig. 13g) Distribution: ANG, DOC, RSA, TAN, ZAM. Distribution in Angola (Provinces): 2)? Historic records: none. Material examined: Angola, 1 ex., N�� 1907 (blue label), old collection, box n�� IX-78, MZUC. Remarks. The adult specimen of MZUC despite be very old and damaged (Fig. 13g), fits well this species description. The adults are generally associated with ants or termits, but the larvae can be found on decaying organic matter. Malec & ����pek (2016) refer that it is possible to rear adults from wild larvae. It is a new genus/species record for Angola., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 61, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Malec, P. & Sipek, P. (2016) On the cetoniine fauna of Eastern Cape (EC) and KwaZulu-Natal (KZN) and the basic guidelines to captive breeding of these beetles (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 9, 52 - 78."]}

Genus Pseudalleucosma Antoine, 1989 The genus was established by Antoine (1989b) under the revision of the genus Alleucosma Schenkling, 1921. There are four recognized species (Beinhundner 2017a), one of them recorded for Angola (Ruter 1960)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 53, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. (1989 b) Revision des genres Alleucosma Schenkling et Molynoptera Kraatz (Coleoptera: Cetoniidae). Revue de Zoologie Africaine, 103, 243 - 252 + 260 - 261.","Schenkling, S. (1921) Scarabaeidae: Cetoninae. In: Junk, W. (Ed.), Coleopterorum Catalogus, 72, W. Junk, Berlin, 431 pp.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Ruter, G. (1960) Description d`un Cetonide nouveau de l`Angola et remarques synonymiques sur quelques Cetonides (Coleoptera). Beitrage zur Entomologie, 10, 434 - 437."]}

Phonotaenia hassoni (Antoine & Legrand, 2005) Distribution: ANG, CAM, DOC, GAB, RCA. Distribution in Angola (Provinces): 1) CUS. Historic records: ��� Kwanza Sul (without locality) (CUS) (Antoine & Legrand 2005, Antoine 2005a). Material examined: none. Remarks. The species mimics Phonotaenia scalaris scalaris Gory & Percheron, 1833 with which can be mistaken (Antoine & Legrand 2005). Only the holotype specimen of the typical series is known from Angola. Biological and ecological data are not available for this species., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 38, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. & Legrand, J. - Ph. (2005) Rehabilitation du genre Phonotaenia Kraatz, 1880, et description d'especes nouvelles (Coleoptera, Cetoniidae). Coleopteres, 11 (3), 13 - 20.","Antoine, Ph. (2005 a) Revision du genre Phonotaenia Kraatz, 1880, sensu lato (Coleoptera, Cetoniidae). Coleopteres, 11 (21), 227 - 265.","Gory, H. & Percheron, A. (1833) Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris, 406 pp. https: // doi. org / 10.5962 / bhl. title. 8957"]}

Trogodes (Trogodes) angolensis Rojkoff, 2011 Distribution: ANG. Distribution in Angola (Provinces): 1) BIE. Historic records: ���Chitengue (BIE) (Rojkoff 2011b). Material examined: none. Remarks. An endemic species of Angola (see Figs 9���11 and Photo 2 in Rojkoff 2011b). Biological and ecologi- cal data are not available., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 67, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Rojkoff, S. (2011 b) Revision du genre Trogodes Westwood, 1857 (Coleoptera, Cetoniidae, Cremastocheilini). Cetoniimania, Nouvelle Serie, 2, 37 - 59."]}

Genus Campsiura Hope, 1831 The genus is distributed throughout Afrotropical, Oriental and Palaearctic Regions (Krajcik 2008a). In the former Region there are fourteen or fifteen recognized species, depending of the authors (cf. Marais & Holm 1992 and Krajcik 2008a). However, two additional species [Campsiura (Macroma) pavlae Legrand & Malec, 2006 and Campsiura (Macroma) kerleyi Antoine, 2006], are not listed in these catalogues. Antoine (2006a) recognized four subgenera and provided an identification key for them, as well as a key to the species of the Campsiura (Macroma) scutellaris group. There are four species recorded for Angola (Paulino de Oliveira 1882, Ferreira 1965, Antoine 2006a, Krajcik 2008a), all in the subgenus Macroma., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 62, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Legrand, J. Ph. & Malec, P. (2006) Une nouvelle espece de Campsiura Hope de Zambie (Coleoptera, Cetoniidae). Les Cahiers Magellanes, Hors-Serie, 20, 1 - 6.","Antoine, Ph. (2006 a) Le genre Campsiura Hope, 1831: Remarques taxonomiques et description d'une espece nouvelle (Coleoptera, Cetoniidae). Coleopteres, 12 (26), 399 - 414.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Genus Tephraea Burmeister, 1842 The genus was revised by Holm (1992) and includes sixteen species (Beinhundner 2017a). Within this assemblage, one species was previuously recorded for Angola (Beinhundner 2017a, Schoolmeesters 2018). A new species record for this country is given in this work., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 41, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)"]}

Porphyronota tottenhami (Holm, 1990) Distribution: ANG. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Holm 1990b sub P. carnifex tottenhami Holm); ��� Angola (without locality) (Perissinotto 2012). Material examined: none. Remarks. An endemic species of Angola firstly described as a subspecies of Porphyronota carnifex (Fabricius, 1781) by Holm (1990b) and later elevated to species level (Perissonoto 2012). Only the type series is currently known for this species. Some ecological data were only provided for P. carnifex by Holm & Marais (1992)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 75, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Holm, E. (1990 b) Revision of African cetoniinae III. Genus Porphyronota Burmeister including Poecilophila Kolbe (coleoptera scarabaeidae). Cimbebasia, 12, 111 - 126.","Perissinotto, R. (2012) Review of Porphyronota carnifex (Fabricius, 1781) (Coleoptera: Scarabaeidae: Cetoniinae). African Entomology, 20 (1), 150 - 160. https: // doi. org / 10.4001 / 003.020.0119","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

Genus Clastocnemis Burmeister & Schaum, 1840 The genus was not revised. There are five recognized species (Antoine 2005b; Krajcik 2008a; Schoolmeesters 2018), one of them recorded for Angola (Harold 1879)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 115, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. (2005 b) A propos de Clastocnemis quadrimaculatus (Afzelius, 1817) (Coleoptera, Cetoniidae, Trechiinae). Ento- mologia Africana, 10 (1), 2 - 7.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Harold, E. (1879) Bericht uber die von Herren A. v. Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Coleopteren. Coleopterologische Hefte, 16, 1 - 224."]}

Pachnoda sinuata flaviventris (Gory & Percheron, 1833) (Figs. 6b, c, d) Distribution: ANG, RSA, ZIM. Distribution in Angola (Provinces): 1) HUI, LNO; 2) LUA, MOX. Historic records: ��� Hu��la (HUI) (Paulino de Oliveira 1882 sub Cetonia flaviventris Gory & Perch.); ���Dundo (LNO) (Gomes Alves 1959 sub P. sinuata Kraatz). Material examined: Angola, 5 ♀, N�� 1965 (blue labels), n�� 189 (white labels), RQC/CSFC, box n�� II-71, MZUC; Angola, 2 ♀, N�� 1907 (blue labels), n��s 85 and 87 (white labels), old collection, box n�� XIV-83, MZUC; Dundo (7�� 22��S, 20�� 50��E, 1470 m alt., 51) (LNO), -. XI.1948, 1 ♀, N�� 2140, -. XII.1948, 1♂, N�� 2146, -. I.1949, 1 ♀, N�� 2131, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Calombe (riverhead) (11�� 50��S, 19�� 56��E, 1360 m alt., 218) (MOX), 17.X.1959, 1 ♀, N�� 3249, Estudos Ap��colas do Ultramar, n�� 46, Junta de Investiga- ����es Coloniais (Entomologia), IICT, MUHNAC; Luanda (8�� 50��S, 13�� 15��E, 70 m alt., 89) (LUA), 15.VIII.1970, 4 ex., n��s 4342, 4343, 4344, 4345, A. Bivar de Sousa leg., ABSC, MUHNAC; Luanda (8�� 48��28���� S, 13�� 14��19���� E, 7 m alt., 89) (LUA), 26.XI. 2015, 1♀, DO, A. Serrano leg., ASC. Remarks. Some adult specimens collected in Luanda like the ones deposited in MZUC fit very well the habitus (e.g. Figs. 6b, c) of the nominal species (see p. 221 in Beinhundner 2017a) and others (e.g. Fig. 6d) the habitus of P. sinuata machadoi Ruter (in Rigout 1982) (see also Beinhundner 2017a, figs 1���5, p. 224). However, all present the underside yellow, a character associated to P. sinuata flaviventris. The larva has been described by Donaldson (1987). Biological and ecological data were provided by Holm & Marais (1992). A basic guideline to captive breeding was also provided by Malec & ����pek (2016) for the nominal species. It is a new species record for the Luanda and Moxico Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 34, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Gory, H. & Percheron, A. (1833) Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris, 406 pp. https: // doi. org / 10.5962 / bhl. title. 8957","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Rigout, J. (1982) La determination des Pachnoda (2 º article). Bulletin des Sciences Naturelles, 36, 8 - 11.","Donaldson, M. I. J. (1987) Descriptions of, and a key to larvae of some South African Cetoniinae (Coleoptera: Scarabaeidae). Entomologist's Monthly Magazine, 123, 1 - 13.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Malec, P. & Sipek, P. (2016) On the cetoniine fauna of Eastern Cape (EC) and KwaZulu-Natal (KZN) and the basic guidelines to captive breeding of these beetles (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 9, 52 - 78."]}

Genus Lophorrhina Westwood, 1842 Most of the representatives of this genus were described under genera Daedycorrhina Bates, 1880 and Chordodera Burmeister, 1842 (e.g. see Allard 1985; Marais & Holm 1992; Krajcik 2008a), nowadays synonyms of Lophorrhina together with Aphanesthes Kolbe, 1892, Aphanochroa Kolbe, 1893 and Pseudoheteromorpha Allard, 1990 (Krikken 1984; Marais & Holm 1992; Krajcik 2008a). Beinhundner (2017a) recognized thirteen valid species. It is represented by two species in Angola (Paulino de Oliveira 1882) one of them a new record for this country., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 99, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Allard, V. (1985) Goliathini 2: Les Coleopteres du monde. Sciences Naturelles de Venette, 6, 1 - 89.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Krikken, J. (1984) A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated lists of the known genera. Zoologische Verhandelingen, 210, 1 - 75.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52."]}

Pachnoda picturata Boheman, 1860 (Fig. 5b) Distribution: ANG, BOW, NAB, RSA, TAN. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Paulino de Oliveira 1882 sub C etonia picturata Boh.). Material examined: Angola, 1 ♀, N�� 1907 (blue label) and 139 (white label), old collection, box XIV-83, MZUC. Remarks. The distribution of this species is not consensual. While Paulino de Oliveira (1882) recorded it to Angola and Holm & Marais (1992) to Botswana and Republic of South Africa, Schoolmeesters (2018) cites it to Tanzania also, but Beinhundner (2017a) only refers Namibia. There is one adult female specimen of Angola (Fig. 5b) deposited in the ���old collection��� of MZUC which confirms its presence in this country. Some ecological data were provided by Holm & Marais (1992)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 31, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp."]}

Plagiochilus pilosus Moser, 1913 (Fig. 13e) Distribution: ANG, DOC. Distribution in Angola (Provinces): 1) Huambo. Historic records: ���Bailundo (HUAMBO) (Moser 1913a); ��� Angola (without locality) (Ferreira 1965). Material examined: Alto Hama���Bailundo (9 km) (12�� 11��� 27.62���S, 15�� 35��� 58.56���E, 1450 m alt., 232) (HUAMBO), 4.XII.2017, 1♀, DO, A. Serrano leg., ASC. Remarks. The species was described from Angola (Moser 1913a), but later recorded for DOC also (Basilewsky 1956). Basilewsky (1956) remarked that it is a true termitophilic species. The Alto Hama���Bailundo adult specimen was collected in a yellow flower of a spontaneous herbaceous plant in a secondary open forest. Biological and other ecological data are not available for the species, but see general comments on the genus Coenochilus., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 59, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Moser, J. (1913 a) Beitrag zur kenntnis der Cetoniden. XII. Annales de la Societe Entomologique de Belgique, 57, 155 - 169. https: // doi. org / 10.5962 / bhl. part. 4592","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Basilewsky, P. (1956) Exploration du Parc National de l`Upemba, Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. Van Meel et R. Verheyen (1946 - 1949). 41: Cetoniinae, Trichiinae, Valginae (Coleoptera, Polyphaga). Institut des Parcs nationaux du Congo Belge, Bruxelles, 53 pp."]}

Grammopyga cincticollis (Hope, 1842) Distribution: ANG, CAM, CON, DOC, GHA, GUI, LIR, RCI, TAN, UGA. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Beinhundner 2017a, 2017b). Material examined: none. Remarks. Beinhundner (2017a, 2017b) recorded the species for Angola without any precise locality. Jerath & Unny (1965) described the larvae of this species. Joly (2018) provided some biodiversity information (abundance, habitat preference) of this species for Ghana., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 46, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Beinhundner, G. (2017 b) Descriptions from West Africa of a new species of Grammopyga Kolbe, 1895 and a new species of Leucocelis Burmeister, 1842 (Coleoptera, Cetoniidae). Entomologia Africana, 22 (2), 49 - 54.","Jerath, M. L. & Unny, K. L. (1965) Larvae of six genera of Cetoniinae from eastern Nigeria (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 19 (2), 59 - 64.","Joly, C. (2018) Deuxieme contribution a la connaissance des Cetoniidae du Ghana, sous-familles des Cetoniinae et des Trichinae (Coleoptera, Scarabaeidae). Lambillionea, 118 (1), 28 - 89."]}

Dicronorhina johnstoni (Waterhouse, 1902) (Figs. 20d, e) Distribution: ANG, BUR, CON, DOC, KEN, NRE, RCA, RRW, SUD, TAN, UGA, ZAM. Distribution in Angola (Provinces): 1) CAB, BGU, CNN, LNO, MAL. Historic records: ���Pungo Andongo (MAL) (Harold 1879 sub Dicranorrhina micans Drury); ���Quilo river (CAB), Humbe (CNN) (Paulino de Oliveira 1882 sub Ceratorrhina micans Drury); ���Quango (=Cuango) (LNO) (Quedenfeldt 1884 sub Dicranorrhina micans Drury); ��� Angola (without locality) (Ferreira 1965 and Ruter 1967a sub D. micans Drury); ���Chinchoxo, Luango (CAB); Kimbundo, Pungandongo (MALANJE); Benguela (BGU) (Lekkerkerk & Krikken 1986 sub D. micans (Drury); ���Northwestern Angola (without localities) (De Palma & Franz 2011. Material examined: Angola, 1 ♂, N�� 1907 (blue label), n�� 32 (white label), Welwitsch leg., old collection, box n�� VI-75, MZUC; Angola, 1 ♀, N�� 1907 (blue label), n�� 33 (white label), old collection, box n�� IX-78, MZUC. Remarks. For long time, this species was considered a synonym of D. micans (Drury, 1773) (e.g. Lekkerkerk & Krikken 1986; Marais &Holm 1992; Krajcik 2008a), but recently De Palm & Frantz (2011) clarified the true status of D. johnstoni, elevating it to a valid species. The localities recorded for Angola suggested that it seems to be widespread in the territory. The larva of the third instar was described by Carl & K��hbander (1993) under the name D. micans. A synthesis of ecological data were provided by De Palm & Frantz (2011)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 89-90, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Harold, E. (1879) Bericht uber die von Herren A. v. Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Coleopteren. Coleopterologische Hefte, 16, 1 - 224.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Quedenfeldt, G. (1884) Verzeich niss der von Herrn Major a. D. von Mechow in Angola und am Quango-Strom 1878 - 1881 gesammelten Pectinicornen und Lamellicornen. Berliner Entomologiche Zeitschrift, 28 (2), 265 - 340. https: // doi. org / 10.1002 / mmnd. 47918840107","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Ruter, G. (1967 a) Contribution a l'etude du genre Leucocelis Burmeister. Revue de Zoologie et de Botanique Africaine, 76, 37 - 42.","Lekkerkerk, R. W. & Krikken, J. (1986) Taxonomic review of the Afrotropical genus Dicronorrhina Hope, with notes on its relatives (coleopteran: Cetoniidae). Zoologische Verhandelingen, 233, 3 - 46.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Carl, M. & Kuhbander, M. (1993) Die Praimaginalstadien der Rosenkafer. 1. Beschreibung der Larven von Dicronorhina micans (Drury, 1773) und Dicronorhina derbyana subsp. oberthuri Deyrolle, 1876 (Coleoptera, Scarabaeidae, Cetoniinae). Entomofauna, 14, 489 - 500."]}

Diplognatha (Diplognatha) silicea (MacLeay, 1838) (Fig. 15f) Distribution: ANG, BOW, MAW, MOZ, NAB, RSA, TAN, ZAM, ZIM. Distribution in Angola (Provinces): 1) CCU, HUI, LNO, MAL; 2) BIE, CUS, HUA, LSU, MOX. Historic records: ���Pungo Andongo (MAL) (Harold 1879); ���Malange (= Malanje) (MAL), Quango (=Cuango) (LNO) (Quedenfeldt 1884); ���Kapelongo (=Capelongo), Dongo (HUI), Luiana (CCU) (Bourgoin 1925); Material examined: Angola, 5 exs, N�� 1907 (blue labels), old collection, box n�� XIV-83, MZUC; Angola, 1 ex., N�� 1907 (blue label), n�� 22 (white label), old collection, box VI-75, MZUC; Angola, 2 exs, N�� 1975 (blue label), n�� 208 (white label), RQC / CSFC, box n�� II-71, MZUC; Angola, 1 ex., N�� 2110 (blue label), J. Pimentel leg., box n�� I-70, MZUC; Huambo (12�� 46��S, 15�� 44��E, 1650 m alt., 256) (HUA), VIII.1934, 2 exs, N��s 1984 and 1989 (blue labels), n�� 67a (white label, red number), J. Pimentel leg., X.1934, 1 ex., N�� 1987 (blue label), n�� 72 (white label, red number), J. Pimentel leg., X.1934, 3 exs, N�� 1989 (blue labels), n�� 77 (white labels, red numbers), J. Pimentel leg., XI.1934, 16 exs, N�� 1989 (blue labels), n�� 77c (white labels, red numbers), J. Pimentel leg., XII.1934, 1 ex., N�� 1991 (blue label), n�� 88 (white label, red number), J. Pimentel leg., box n�� I-70, MZUC; Huambo (12�� 46��S, 15�� 44��E, 1650 m alt., 256) (HUA), 20 exs, N�� 1989 (white labels), J. Pimentel leg., box n�� 97, MZUC; Luma���Cassai (Dala) (coord. not available, 197) (LSU), 9.X.1957, 1 ♂, N�� 2927, Estudos Ap��colas do Ultramar n�� 79, Junta de Investiga- ����es do Ultramar (Entomologia), IICT, MUHNAC; Dala (=Lunda) (11�� 02��S, 20�� 12��E, 1180 m alt., 197) (LSU), 10.X.1957, 1 ♂, N�� 2929, Estudos Ap��colas do Ultramar n�� 82, Junta de Investiga����es do Ultramar (Entomologia), IICT, MUHNAC; Silva Porto (=Kuito=Cuito) (12�� 23��S, 16�� 57��E, 1450 m alt., 234) (BIE), 19.X.1957, 1 ♂, 1 ♀, N�� 2946, Estudos Ap��colas do Ultramar n�� 89, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Vouga (Qt�� Vista Alegre) (12�� 14��S, 16�� 47��E, 1766 m alt., 234) (HUA), 25.X.1957, 1 ♂, N�� 2948, Estudos Ap��colas do Ultramar n�� 2948, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Mutumbo (13�� 10��S, 17�� 23��E, 1457 m alt., 283) (BIE), 30.X.1957, 1 ♀, N�� 3087, Estudos Ap��colas do Ultramar, Junta de Investiga����es Co- loniais (Entomologia), IICT, MUHNAC; Nova Lisboa (= Huambo) (12�� 46��S, 15�� 44��E, 1650 m alt., 256) (HUA), 6.XI.1957, 1 ♂, N�� 2956, Estudos Ap��colas do Ultramar n�� 106, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Calombe (=Calomba) (11�� 50��S, 19�� 55��E, 1360 m alt., 218) (MOX), 1.XII.1959, 1 ♀, N�� 3277, Estudos Ap��colas do Ultramar n�� 74, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Chianga (12�� 44��S, 15�� 50��E, 1740 m alt., 256) (HUA), 25.X.1963, 1 ex., N�� E.3472, n�� 611, E. Fonseca leg., 12.XI.1964, 1 ex., N�� E.5818, n�� 610, Jos�� 5�� leg., JPCC, MUHNAC; Lumeje (P.N. Cameia) (11�� 33��S, 20�� 47��E, 1150 m alt., 220) (MOX), 15.IV.1965, n�� 4353, A. Bivar de Sousa leg., ABSC, MUHNAC; Ciclotroco (Nova Lisboa= Huambo) (coords not available, 256) (HUA), 15.XII.1974, 3 ♀, DO, A. Serrano leg., ASC; Dundo���Chitato (7�� 22��41.93���� S, 20�� 50��04.81���� E, 733 m alt., 51) (LNO), 26.X.���7.XI.1995, 1 ♀, L. Mendes leg., IICT, MUHNAC; Mussende (Calulo) (CUS) (9�� 57��00���� S, 14�� 47��36���� E, 905m alt., 128), 5.XII.2015, 1♀, BT, A. Serrano & R. Capela leg., ASC. Remarks. Despite the species be known for Angola since a long time ago (e.g. Bourgoin 1925), both Beinhundner (2017a) and Schoolmeesters (2018) do not consider its presence in that country. All the examined specimens fit the mesotibial character (only one tooth on the external margin) proper of this species (Rigout & Allard 1992). The adult specimens from Chianga were collected on bean plants (label information). The Mussende adult specimen was sampled by BT in a secondary moist forest together with Lophorrhina quinquelineata (Fabricius, 1791), Tmesorrhina laeta burgeoni Allard, 1991 and Myodermum alutaceum (Afzelius, 1817). The larva of this species was described by Oberholzer (1959). Ecological data were provided by Holm & Marais (1992), but see Malec & ����pek (2016) for breeding habits also. It is a new record for the Bi��, Huambo, Kwanza Sul, Lunda Sul and Moxico Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 71-72, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Harold, E. (1879) Bericht uber die von Herren A. v. Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Coleopteren. Coleopterologische Hefte, 16, 1 - 224.","Quedenfeldt, G. (1884) Verzeich niss der von Herrn Major a. D. von Mechow in Angola und am Quango-Strom 1878 - 1881 gesammelten Pectinicornen und Lamellicornen. Berliner Entomologiche Zeitschrift, 28 (2), 265 - 340. https: // doi. org / 10.1002 / mmnd. 47918840107","Bourgoin, A. (1925) Coleopteres: Scarabaeidae. Tribu: Cetoniini. In: Mission Rohan-Chabot Angola et Rhodesia. 4 (3). Impremerie Nationale, Paris, pp. 141 - 146.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Rigout, J. & Allard, V. (1992) Cetoniini 2: Les Coleopteres du monde. Sciences Naturelles Venette, 12, 1 - 135.","Allard, V. (1991) Goliathini 4: Les Coleopteres du monde. Sciences Naturelles de Venette, 11, 1 - 142.","Oberholzer, J. J. (1959) A morphological study of some South African Lamellicorn larvae. I-descriptions of the third instar larvae. II-Comparative morphology. South African Journal of Agricultural Science, 2, 41 - 88.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Malec, P. & Sipek, P. (2016) On the cetoniine fauna of Eastern Cape (EC) and KwaZulu-Natal (KZN) and the basic guidelines to captive breeding of these beetles (Coleoptera, Scarabaeidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 9, 52 - 78."]}

Mecynorhina (Mecynorhinella) torquata torquata (Drury, 1782) Distribution: ANG, CAM, COM, DOC, GHA, RCA, RCI, SLN. Distribution in Angola (Provinces): 1) CAB. Historic records: ��� Cabinda (CAB) (Paulino de Oliveira 1882 sub Ceratorrhina torquata Drury); ��� Angola (without locality) (Ruter 1967a). Material examined: none. Remarks. The species is polytypic with two to four subspecies recognized, depending of the authors (see Marais & Holm 1992; Krajcik 2008a; Beinhundner 2017a; Schoolmeesters 2018). It was recorded for CAB, the northernmost Province of Angola (Paulino de Oliveira 1882). This species has been used in insect flight studies like remote radio control or the role of the stretched forelegs in flight (Sato et al. 2009; Li et al. (2017)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 101, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Ruter, G. (1967 a) Contribution a l'etude du genre Leucocelis Burmeister. Revue de Zoologie et de Botanique Africaine, 76, 37 - 42.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Sato, H., Berry, C. W., Peeri, Y., Baghoomian, E., Casey, B. E., Lavella, G., VandenBrooks, J. M., Harrison, J. F. & Maharbiz, M. M. (2009) Remote radio control of insect flight. Frontiers in Integrative Neuroscience, 3 (24), 1 - 11. https: // doi. org / 10.3389 / neuro. 07.024.2009","Li, Y., Cao, F., Vo Doan, T. T. & Sato, H. (2017) Role of outstretched forelegs of flying beetles revealed and demonstrated by remote leg stimulation in free flight. Journal of Experimental Biology, 220, 3499 - 3507. https: // doi. org / 10.1242 / jeb. 159376"]}

Diploa tridens Arrow, 1906 (Figs. 28a, b, c, d) Distribution: ANG, DOC. Distribution in Angola (Provinces): 1) BIE. Historic records: ���Bihe (BIE) (Arrow 1906). Material examined: Chitembo (10 km N) (13�� 22��40.78���� S, 16�� 41��58.07���� E, 1666 m alt., 282) (BIE), 1.XI.2014, 1♂, 1♀, LT, A. Serrano & R. Capela leg., ASC. Remarks. The species was originally described from Angola and D.R. of the Congo (Katanga: Garenganze) (Arrow 1906). The adult specimens from Chitembo were collected by LT in an open secondary forest. Biological and other ecological data are not available., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 115, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Arrow, G. J. (1906) On Lamellicorn Coleoptera from Portuguese West Africa, with descriptions of new species. The Annals and Magazine of Natural History, Series 7, 18 (104), 127 - 136. https: // doi. org / 10.1080 / 00222930608562586"]}

Genus Platygenia MacLeay, 1819 The genus was revised by Antoine & Camiade (1995), who recognized four valid species. There is one species recorded for Angola (Paulino de Oliveira 1882)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 113, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. & Camiade, D. (1995) Revision du genre Platygenia MacLeay et description d'une espece nouvelle (Coleoptera, Cetoniidae). Coleopteres, 1 (1), 1 - 8.","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52."]}

Tephraea leucomelona (Gory & Percheron, 1833) Distribution: ANG, BOW, MOZ, NAB, RSA, TAN, ZIM. Distribution in Angola (Provinces): 1) BIE, CNN. Historic records: ���Bihe (= Bi��) (without locality) (BIE) (Gory & Percheron 1833); ���Merero (CNN) (Paulino de Oliveira 1882 sub Cetonia leucomelana Gory & Perch.). Material examined: none. Remarks. The species was originally described from Angola (Gory & Percheron 1833). The larvae instars of T. leucomelona were described by Donaldson (1987) and oviposition by Donaldson & Todner (1992). Biological and ecological data were provided by Holm (1992) and Holm & Marais (1992)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 41, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Gory, H. & Percheron, A. (1833) Monographie des Cetoines et genres voisins, formant, dans les familles naturelles de Latreille, la division des Scarabees melitophiles. J. - B. Bailliere, Paris, 406 pp. https: // doi. org / 10.5962 / bhl. title. 8957","Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Donaldson, M. I. J. (1987) Descriptions of, and a key to larvae of some South African Cetoniinae (Coleoptera: Scarabaeidae). Entomologist's Monthly Magazine, 123, 1 - 13.","Donaldson, M. I. J. & Todner H. J. (1992) Brief observation on the chorion patterns of Cetoniine eggs (Coleptera: Scarabaeidae). Journal of the Entomological Society of South Africa, 55, 9 - 19.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

* Mecynorhina (Mecynorhina) polyphemus confluens (Kraatz, 1890) (Fig. 23i) Distribution: ANG, CON, RRW, UGA. Distribution in Angola (Provinces): 2)? Historic records: none Material examined: Angola, 1 ♀, N�� 1907 (blue label), n��36 (white label), Welwitsch leg., old collection, n�� VI-75, MZUC. Remarks. The adult female specimen deposited in MZUC fits well the habitus of this subspecies (cf. Fig. 3 in Beinhundner 2017a, p. 1002, with Fig. 23i). See also the remarks under the precedent species. It is a new subspecies record for Angola., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 101, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp."]}

Coenochilus parrianus Westwood, 1883 Distribution: ANG. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Westwood 1883). Material examined: none Remarks. The species was described from Angola (Westwood 1883). It is considered a controversial species and an incertae sedis (Schein 1954a). Schein (1954a) in the ���general comments��� (���I. Allgemeines���) referred to Coenochilus parrianus based in Schenkling (1921) catalogue (��� parrianus Westwood Tijdschr. voor Ent. 26 Seite 65 Angola ���). Taking into account this reference, it seems that the species considered by Schein (1954a) is indeed that which was described by Westwood (1883), i.e. Coenochilus parrianus from Angola. The same author stated also that the type of C. parrianus was unfortunately lost, concluding that ���The Parrianus not mentioned in the Junk Schenkling catalog should therefore remain omitted��� (���Der im Junk-Schenkling-Katalog ausgelassene Parrianus muss also weiter im Dunklen bleiben���). Biological and ecological data are not available, but see general comments in Schein (1954a) and the remarks under the genus Coenochilus., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on pages 55-56, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Westwood, J. O. (1883) Descriptions of some new exotic coleoptera. Tijdschrift voor Entomologie, 26, 61 - 78, 3 pls.","Schein, H. (1954 a) Uber Coenochilus (Coleoptera, Cetoniidae). Eine Revision der afrikanischen Arten mit einer Bestimmungstabelle und ein Nachtrag zu meiner Revision der asiatischen Arten. Entomologische Arbeiten aus dem Museum G. Frey, 5, 271 - 325.","Schenkling, S. (1921) Scarabaeidae: Cetoninae. In: Junk, W. (Ed.), Coleopterorum Catalogus, 72, W. Junk, Berlin, 431 pp."]}

Lobovalgus murinus Moser, 1913 Distribution: ANG, DOC. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Antoine 2002c). Material examined: none. Remarks. As the precedent, the species is only known for ANG and DOC (Antoine 2002c). Biological and ecological data are not available, but like the other representatives of Valgini, is probably a flower visitor., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 118, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Antoine, Ph. (2002 c) Revision du genre Lobovalgus Kolbe (Coleoptera, Cetoniidae, Valginae). Coleopteres, 8 (12), 191 - 209."]}

Spelaiorrhina rufopyga (Lansberge, 1886) Distribution: ANG, CAM, DOC, TAN, ZIM. Distribution in Angola (Provinces): 1) HUI. Historic records: ���Humpata (HUI) (Lansberge 1886, Ferreira 1965, Krajcik 2008a). Material examined: none. Remarks. Brief ecological data were provided by Holm & Marais (1992)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 104, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Lansberge, J. W. (1886) Scarabaeides, buprestides et cerambycides de l`Afrique Occidentale, envoyes au Musee de Leyden par MM. Veth et Van der Kellen. Notes from the Leyden Museum, 8, 69 - 120.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125."]}

Leucocelis (Leucocelis) cincticollis Moser, 1908 Distribution: ANG, CON, DOC, TAN. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Beinhundner 2017a). Material examined: none. Remarks. The species was originally described from ���Deutsch Ost-Afrika��� (Moser 1908), but curiously Schoolmeesters (2018) refers only East Africa for its distribution. Beinhundner (2017a) indicates Angola in its distribution without reference to any locality. Biological and ecological data are not available for this species., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 47, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Moser, J. (1908) Beitrag zur kenntnis der Cetoniden IV. Annales de la Societe Entomologique de Belgique, 52, 85 - 96. https: // doi. org / 10.5962 / bhl. part. 5012","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)"]}

Genus Coenochilus Schaum, 1841 Some Cremastocheilini to which this and other genera belong (e.g. Anatonochilus P��ringuey, 1907, Cymophorus Kirby, 1827, Plagiochilus Wasmann, 1900, Trogodes Westwood in Boheman, 1857, Spilophorus Westwood in Schaum, 1848) are a group of Cetoniinae that are reported to be myrmecophilous or termitophilous (e.g. Schein 1954a) and, as adults, with some exceptions, are mainly active at night (Holm & Perissinotto 2010). Some species are also predators of ant larvae (e.g. see Ghorpade 1975; Mynhardt & Wenzel 2010). The genus Coenochilus is the most speciose with about 100 species distributed throughout the Afrotropical, Oriental and Palaearctic Regions (Krajcik 2008b). Schein (1954a) revised the 41 African species known by that time and defined seven groups within the genus. There are about 48 species recognized for the former Region (Schoolmeesters 2018), eight of them previously recorded for Angola (Schein 1954a; Ferreira 1965). We give two new species records for this country., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 53, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Schaum, H. R. (1841) Analecta entomologica. Vol. 4. Voss, Halle, 49 pp.","Peringuey, L. (1907) Descriptive catalogue of the Coleoptera of South Africa. Fam. Lucanidae. Fam. Scarabaeidae. Sub-families: Sericinae, Melolonthinae, Trichiinae, Valginae, Cetoniinae. Transactions of the South African Philosophical Society, 13, 289 - 546, pl. 47.","Schein, H. (1954 a) Uber Coenochilus (Coleoptera, Cetoniidae). Eine Revision der afrikanischen Arten mit einer Bestimmungstabelle und ein Nachtrag zu meiner Revision der asiatischen Arten. Entomologische Arbeiten aus dem Museum G. Frey, 5, 271 - 325.","Holm, E. & Perissinotto, R. (2010) Revision of the Afrotropical Species of the Genus Spilophorus Westwood (in Schaum), 1848 (Coleoptera: Scarabaeidae: Cetoniinae: Cremastocheilini). African Entomology, 18 (1), 47 - 65. https: // doi. org / 10.4001 / 003.018.0107","Ghorpade, K. D. (1975) A remarkable predacious cetoniid, Spilophorus maculatus (Gory & Percheron), from southern India (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 29 (4), 226 - 230.","Mynhardt, G. & Wenzel, J. W. (2010) Phylogenetic analysis of the myrmecophilous Cremastocheilus Knoch (Coleoptera, Scarabaeidae, Cetoniinae), based on external adult morphology. ZooKeys, 34, 129 - 140. https: // doi. org / 10.3897 / zookeys. 34.262","Krajcik, M. (2008 b) Cetoniidae of the World. Catalogue - Part II. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 72 + I-XXIII.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317."]}

Lophorrhina penthacordia (Klug, 1835) Distribution: ANG, CAM, DOC, GHA, RCI, SEN. Distribution in Angola (Provinces): 1)? Historic records: ��� Angola (without locality) (Paulino de Oliveira 1882 sub Chordodera penthacordia Klug). Material examined: none. Remarks. A species apparently very common (Allard 1985; Le Gall 2015; Schoolmeesters 2018) and recorded to Angola by Paulino de Oliveira (1882). Eventually, it was misinterpreted by Paulino de Oliveira (1882) with C. quinquelineata (Fabricius, 1781) and its occurrence in Angola must be considered with some caution (see next species remarks)., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 100, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Paulino de Oliveira, M. (1882) Etudes sur les insectes d'Angola qui se trouvent au Museum National de Lisbonne. Jornal de Sciencias Mathematicas Physicas e Naturaes, 33, 40 - 52.","Allard, V. (1985) Goliathini 2: Les Coleopteres du monde. Sciences Naturelles de Venette, 6, 1 - 89.","Le Gall, Ph. (2015) La faune entomologique du Benin: Les Cetoniinae (Coleoptera, Cetoniidae, Cetoniinae). Cetoniimania, Nouvelle Serie, 8, 25 - 53.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)"]}

* Trogodes (Trogodes) rotundicollis Westwood, 1857 (Figs. 14g, h, i) Distribution: ANG, RSA, DOC, TAN, ZAM, ZIM. Distribution in Angola (Provinces): 2) BIE, HUI. Historic records: none. Material examined: Huilla (= Hu��la) (15�� 05��S, 13�� 33��E, 1720 m alt., 356) (HUI), 1 ex., Lobo d`Avila leg., old collection, box n�� VI-75, MZUC; Trumba (12�� 11��50.47���� S, 17�� 00��7.22���� E, 1674 m alt., 235) (BIE), 28.XI.2017, 1♀, DO, A. Serrano leg., ASC. Remarks. The adult specimem of MZUC and the adult female specimen of Trumba fit well the description of the species, presenting in each side of the pronotal disk one small smooth plate and the absence of posterior angles. However, the schematic outline of pronotum with the anterior angles obtuse (dorsal view) given by Rojkoff (2011b, Fig. 6) does not conform with the same structure of our specimens (anterior angles acute, Fig. 14i).The adult specimen of Trumba was captured on sandy whitish soil in a secondary open forest. Biological and ecological data are not available for the species, but see general comments on the genus Coenochilus. It is a new species record for Angola and the Bi�� and Hu��la Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 68, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Rojkoff, S. (2011 b) Revision du genre Trogodes Westwood, 1857 (Coleoptera, Cetoniidae, Cremastocheilini). Cetoniimania, Nouvelle Serie, 2, 37 - 59."]}

Chlorocala smaragdina angolensis (Schauer, 1941) Distribution: ANG. Distribution in Angola (Provinces): 1) BGO. Historic records: ���Dembos (BGO) (Schauer 1941 and Ferreira 1965 sub Smaragdesthes smaragdina angolensis Schauer). Material examined: none. Remarks. This is an enigmatic taxon. We did not find any record in Marais & Holm (1992), Sakai & Nagai (1998), Krajcik (2008a) or Beinhundner (2017a). The subspecies was briefly described by Schauer (1941) in a key of the representatives of the genus Smaragdesthes. While Beinhundner (2017a) considers C. smaragdina as a subspecies of C. africana, Schoolmeesters (2018) lists it as a valid species. Chlorocala smaragdina angolensis seems to be an endemic subspecies of Angola (Schauer 1941), but the validity of the taxon is dubious. The larva of the third instar of C. smaragdina was described by Jerath & Unny (1965). Ecological data are not available for this subspecies., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 84, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Schauer, G. (1941) Revision der Gattung Smaragdesthes Kraatz (Coleoptera: Cetonidae). Arbeiten uber morphologische und taxonomische Entomologie aus Berlin-Dahlem, 8 (3), 161 - 178.","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Marais, E. & Holm, E. (1992) Type catalogue and bibliography of the Cetoniinae of Sub-Saharan Africa. Cimbebasia Memoir, 8, 1 - 125.","Sakai, K. & Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Tokyo, Japan, 3, 1 - 421.","Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI.","Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp.","Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L. (Eds.), Species 2000 & ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)","Jerath, M. L. & Unny, K. L. (1965) Larvae of six genera of Cetoniinae from eastern Nigeria (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 19 (2), 59 - 64."]}

Leucocelis (Leucocelis) triliturata triliturata Quedenfeldt, 1884 (Fig. 10h) Distribution: ANG, DOC. Distribution in Angola (Provinces): 1) MAL, MOX; 2) BIE, HUA, HUI. Historic records: ���Malauge (= Malanje) (MAL) (Quedenfeldt 1884); ��� Angola (without locality) (Ferreira 1965). ���Vila Luso (=Luena) (MOX) (Gomes Alves 1973). New localities reported: Leba pass top (15�� 04��19.48���� S, 13�� 14��39.43���� E, 1690 m alt., 355) (HUI), 4- 5.XII.2017, 1 ♂, P. Sch��le leg., PMC (P. Malec, personal communication). Material examined: Vila Luso (= Luena) (Luena River edge) (coord. not available, 218) (MOX), 20.IX.1957, 1 ex., N�� 2896, Junta de Investiga����es Coloniais (Entomologia), IICT, MUHNAC; Nova Lisboa (= Huambo) (12�� 46��S, 15�� 44��E, 1650 m alt., 256) (HUA), 22.V.1969, 1 ex., N�� 6536, Lonete leg., JPCC, MUHNAC; Capingana- Satchijamba (13�� 42��47.18���� S, 17�� 05��23.81���� E, 1507m alt., 303) (BIE), 8.IV.2014, 1♂, 3♀, DO, A. Serrano leg., ASC; Lumbanje (Morro Lubiri) (12�� 07��55���� S, 15�� 30��59���� E, 1424 m alt., 232) (HUA), 31.X.2015, 3♂, 1♀, DO, A. Serrano & R. Capela leg., ASC. Remarks. The species is polytypic with four recognized subspecies. The adult specimens of Capingana-Satchijamba and Lumbanje were sampled on yellow flowers of unidentified plants from meadows (Fig. 10b). Biological and ecological data are not available for this species. It is a new species record for the Huambo, Hu��la and Bi�� Provinces., Published as part of Serrano, Artur R. M., Capela, R��ben A., Nunes, Telmo & Santos, Carmen Van-D��- Nem Neto, 2020, The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes, pp. 1-130 in Zootaxa 4776 (1) on page 50, DOI: 10.11646/zootaxa.4776.1.1, http://zenodo.org/record/3821288, {"references":["Quedenfeldt, G. (1884) Verzeich niss der von Herrn Major a. D. von Mechow in Angola und am Quango-Strom 1878 - 1881 gesammelten Pectinicornen und Lamellicornen. Berliner Entomologiche Zeitschrift, 28 (2), 265 - 340. https: // doi. org / 10.1002 / mmnd. 47918840107","Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317.","Gomes Alves, M. G. (1973) Coleopteros da Africa Portuguesa (Lucanidae, Scarabaeidae: Cetoniinae; Chironinae, Phaenomerinae, Cicindelidae e Chrysomelidae: Hispinae). In: Livro de homenagem ao Prof. Fernando Frade por ocasiao do seu 70 º aniversario. Junta de Investigacoes do Ultramar, Lisboa, pp. 27 - 58."]}