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Sympatric host-associated genetic differentiation is a prominent pattern that could lead to speciation. In insects, there are numerous examples of host-associated differentiation among herbivores that prefer different plants, and parasitoids that prefer different hosts, but few examples for specialist predators. We developed new microsatellite loci for two species of silver fly, Leucotaraxis argenticollis (Zetterstedt) and L. piniperda (Malloch) (Diptera: Chamaemyiidae), being evaluated as biological control agents for the hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), in eastern North America where it is a nonnative pest. We obtained DNA from specimens of both fly species feeding on native A. tsugae in western North America, as well as on other western and eastern adelgid species. We performed population genetic analyses using the new loci and DNA barcode sequences. Our results confirmed east–west allopatric divergence and uncovered nested genetic differentiation associated with different adelgid prey species and their host plants in western North America for both species of silver flies. For both species, there is also evidence for a longer history of diversification in the west, with ancestral specialization of feeding on pine adelgids, which was retained after range expansion to the east. More recently, divergence to feeding on new adelgid prey species occurred in the west. Our findings are consistent with the hypothesis that host-alternating life cycles in Adelgidae may provide temporary escape from specialist predators. We discuss the implications for biological control efficacy and potential for lineage hybridization as western flies are released in the east to control A. tsugae.

The Neotropical genus Lopesiodinia Prado is represented by two species, L. diversa Prado (Brazil: Rio de Janeiro) and L. alvarengai Prado (Brazil: Pará). The genus is reviewed here, and three new species are described and illustrated: Lopesiodinia marcusi sp. nov. (Brazil: Amazonas, Maranhão), L. argentata sp. nov. (Brazil: Amazonas) and L. pontarolloi sp. nov. (Brazil: Amazonas, Maranhão). A key to identify the subfamilies and extant genera and the species of Traginopinae from the Neotropical Region is presented.

Korneyev, Severyn V., Hancock, David L., Hauser, Martin, Korneyev, Valery A., Gaimari, Stephen D. (2021): A new species of the genus Themarictera Hendel (Diptera: Tephritidae: Phytalmiinae: Acanthonevrini) from Madagascar. Zootaxa 4996 (2): 383-391, DOI: https://doi.org/10.11646/zootaxa.4996.2.12

We present a summary and analysis of the Diptera-related information published in Zootaxa from 2001 to 2020, with a focus on taxonomic papers. Altogether, 2,527 papers on Diptera were published, including 2,032 taxonomic papers and 1,931 papers containing new nomenclatural acts, equivalent to 22% of all publications with new nomenclatural acts for Diptera. The new nomenclatural acts include 7,431 new species, 277 new genera, 2,003 new synonymies, and 1,617 new combinations. A breakdown by family of new taxa and new replacement names proposed in the journal during the last two decades is provided, together with a comparison of Zootaxa’s output to that of all other taxonomic publications on Diptera. Our results show that the journal has contributed to 20% of all biodiversity discovery in this megadiverse insect order over the last 20 years, and to about 31% in the last decade.

Two new genera of Chamaemyiidae (Diptera: Lauxanioidea) are described and illustrated, including: Chamaethrix gen. nov. (type species Chamaethrix necopina sp. nov.), possibly a predator of Cinara ponderosae (Williams) on Pinus ponderosa in the southwestern United States; and Vitaleucopis gen. nov. (type species Vitaleucopis nidolkah sp. nov.; other included species Vitaleucopis astonea (McAlpine), comb. nov., and Vitaleucopis scopulus sp. nov.), predators of Cinara aphids and possibly adelgids on Pinaceae in western North America. Immature stages are discussed or described and illustrated for some taxa, including the eggs of Chamaethrix necopina and Vitaleucopis nidolkah; and the third instars and puparia of Vitaleucopis nidolkah.

Umbodinia bella gen. nov. et sp. nov. is described and illustrated from specimens collected in the canopy of an ombrophilous Amazonian forest, Manaus, Brazil. The genus is characterized by a unique combination of diagnostic features: body predominantly yellow; frons with two shiny dark brown to black lobules in lower corners, between the lunule and eye margins; lunule high-arched with two subtriangular spots covered with black pubescence; parafacial with tufts of long and strong setae, almost as long and stout as the fronto-orbital setae; and anepisternum with setulae dorsally and posteriorly, in addition to posterior setae.

A new genus of odiniids (Odiniidae: Traginopinae) is described from Brazil, Pauximyia gen. nov., with two species, P. oliveirai sp. nov. (type species) and P. vidali sp. nov. A modified key to the Neotropical genera of the family and for the two Pauximyia gen. nov. species is provided.

Study of all flies (Diptera) collected for one year from a four-hectare (150 x 266 meter) patch of cloud forest at 1,600 meters above sea level at Zurquí de Moravia, San José Province, Costa Rica (hereafter referred to as Zurquí), revealed an astounding 4,332 species. This amounts to more than half the number of named species of flies for all of Central America. Specimens were collected with two Malaise traps running continuously and with a wide array of supplementary collecting methods for three days of each month. All morphospecies from all 73 families recorded were fully curated by technicians before submission to an international team of 59 taxonomic experts for identification. Overall, a Malaise trap on the forest edge captured 1,988 species or 51% of all collected dipteran taxa (other than of Phoridae, subsampled only from this and one other Malaise trap). A Malaise trap in the forest sampled 906 species. Of other sampling methods, the combination of four other Malaise traps and an intercept trap, aerial/hand collecting, 10 emergence traps, and four CDC light traps added the greatest number of species to our inventory. This complement of sampling methods was an effective combination for retrieving substantial numbers of species of Diptera. Comparison of select sampling methods (considering 3,487 species of non-phorid Diptera) provided further details regarding how many species were sampled by various methods. Comparison of species numbers from each of two permanent Malaise traps from Zurquí with those of single Malaise traps at each of Tapantí and Las Alturas, 40 and 180 km distant from Zurquí respectively, suggested significant species turnover. Comparison of the greater number of species collected in all traps from Zurquí did not markedly change the degree of similarity between the three sites, although the actual number of species shared did increase. Comparisons of the total number of named and unnamed species of Diptera from four hectares at Zurquí is equivalent to 51% of all flies named from Central America, greater than all the named fly fauna of Colombia, equivalent to 14% of named Neotropical species and equal to about 2.7% of all named Diptera worldwide. Clearly the number of species of Diptera in tropical regions has been severely underestimated and the actual number may surpass the number of species of Coleoptera. Various published extrapolations from limited data to estimate total numbers of species of larger taxonomic categories (e.g., Hexapoda, Arthropoda, Eukaryota, etc.) are highly questionable, and certainly will remain uncertain until we have more exhaustive surveys of all and diverse taxa (like Diptera) from multiple tropical sites. Morphological characterization of species in inventories provides identifications placed in the context of taxonomy, phylogeny, form, and ecology. DNA barcoding species is a valuable tool to estimate species numbers but used alone fails to provide a broader context for the species identified.

Made available in DSpace on 2019-10-06T15:56:52Z (GMT). No. of bitstreams: 0 Previous issue date: 2018-12-01 Estimations of tropical insect diversity generally suffer from lack of known groups or faunas against which extrapolations can be made, and have seriously underestimated the diversity of some taxa. Here we report the intensive inventory of a four-hectare tropical cloud forest in Costa Rica for one year, which yielded 4332 species of Diptera, providing the first verifiable basis for diversity of a major group of insects at a single site in the tropics. In total 73 families were present, all of which were studied to the species level, providing potentially complete coverage of all families of the order likely to be present at the site. Even so, extrapolations based on our data indicate that with further sampling, the actual total for the site could be closer to 8000 species. Efforts to completely sample a site, although resource-intensive and time-consuming, are needed to better ground estimations of world biodiversity based on limited sampling. Entomology Section Natural History Museum of Los Angeles County, 900 Exposition Boulevard Royal British Columbia Museum and the American Museum of Natural History, 691-8th Ave. SE Department of Plant and Environmental Sciences Poole Agricultural Center Clemson University, 130 McGinty Court, E-143 Depto. de Biologia FFCLRP Universidade de São Paulo, Av. Bandeirantes 3900 Great Lakes Forestry Centre Canadian Forest Service Natural Resources Canada, 1219 Queen St. E. Australian Museum, 1 William Street Department of Natural Resource Sciences McGill University Macdonald Campus Canadian National Collection of Insects Invertebrate Biodiversity Agriculture and Agri-Food Canada, K.W. Neatby Building, 960 Carling Avenue Department of Biological Sciences Spaulding Hall University of New Hampshire Department of Biological Sciences Wright State University, 3640 Colonel Glenn Hwy Instituto Federal do Triângulo Mineiro – Campus Uberaba. Rua João Batista Ribeiro 4000 Distrito Industrial II Departamento de Zoologia Universidade Federal do Paraná Jardim das Américas Mississippi Entomological Museum Mississippi State University, 100 Old Highway 12, P.O. Drawer 9775 Department of Biology Western Kentucky University Integrative Research Center Field Museum of Natural History Independent Investigator California State Collection of Arthropods, 2683 Tam O’ Shanter Dr., El Dorado Hills California Department of Food and Agriculture California State Collection of Arthropods, 3294 Meadowview Rd. The Academy of Natural Sciences of Drexel University, 1900 Ben Franklin Parkway American Museum of Natural History, Central Park West at 79th St. Department of Entomology University of California Riverside, 900 University Ave. Zoological Museum Biodiversity Unit FI-20014 University of Turku Instituto de Ecología A.C. (INECOL) Red Ambiente y Sustentabilidad, Carretera Antigua a Coatepec 351, Col El Haya, Xalapa Station Linné, Ölands Skogsby 161 I. I. Schmalhausen Institute of Zoology of the National Academy of Sciences of Ukraine, Bogdan Chmielnicki St. 15 Universidad de Panama Department of Natural History University Museum of Bergen University of Bergen, P.O. Box 7800 Agriculture and Agri-Food Canada, 960 Carling Avenue School of Environmental Sciences University of Guelph Department of Entomology Smithsonian Institution, PO Box 37012, MRC 169 Natural History Museum of Denmark, Universitetsparken 15 Institute of Evolutionary Biology and Environmental Studies University of Zurich, Winterthurerstrasse 190 Systematic Entomology Laboratory USDA ARS c/o National Museum of Natural History, MRC-168, P.O. Box 37012 Departamento de Sistemática e Ecologia CCEN Universidade Federal da Paraíba, Castelo Branco, s/n Research Institute for Nature and Forest (INBO), Kliniekstraat 25 Research Group Terrestrial Ecology (TEREC) Ghent University, K.L.Ledeganckstraat 35 Entomology Unit Royal Belgian Institute for Natural Sciences (RBINS), Vautierstraat 29 USDA Forest Service Rocky Mountain Research Station Forestry Sciences Laboratory, 1648 S. 7th Avenue Department of Biological Sciences Bishop’s University, 2600 College Street UNESP - Univ Estadual Paulista Faculdade de Ciências Agrárias e Veterinárias Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane s/n Canadian National Collection of Insects & Canadian Food Inspection Agency OPL-Entomology, K.W. Neatby Bldg., C.E.F., 960 Carling Ave. Department of Biological Sciences University of Calgary, 2500 University Drive NW Finnish Museum of Natural History Zoology Unit University of Helsinki Washington State University, 2533 Inter Avenue Bohart Museum of Entomology University of California, One Shields Avenue Instituto Nacional de Biodiversidad (INBio) UNESP - Univ Estadual Paulista Faculdade de Ciências Agrárias e Veterinárias Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane s/n

Five species of the genus Tetroxyrhina Hendel from China are described as new to science: Tetr. dashahensis sp. nov., Tetr. dentata sp. nov., Tetr. jinpingensis sp. nov., Tetr. menglunensis sp. nov. and Tetr. tengchongensis sp. nov., and the species Tetr. sauteri (Hendel), also from China, is redescribed. The genitalia of these species are illustrated and described. Trigonometopus alboapicalis Shatalkin, is synonymized under Tetr. sauteri , syn. nov. A key to separate all species of the genus is provided, along with nomenclatural details for all species, and photographs of all primary type specimens.

In the course of two field trips to Northern Vietnam during March 2012 and June 2014 the dragonflies and damselflies of three National Parks (Cuc Phuong, Tam Dao, and Ba Be) and one Biodiversity Station (Melinh) were sampled. A total of 90 species of odonates in 60 genera and 15 families was recorded, including two new records for Vietnam: Macromia katae (Macromiidae) and Indothemis carnatica (Libellulidae). Diagnostic illustrations for these two species are provided, as well as the listing of the species recorded from the surveyed areas.

The new Afrotropical genus and type species Hamecamyia stuckenbergi gen. et sp. n. (Diptera: Chamaemyiidae), from KwaZulu-Natal Province, South Africa, are described and figured. Characters are provided to differentiate this new genus from other Afrotropical chamaemyiids. Discussion is provided comparing this new genus to Parochthiphila Czerny, 1904, and an apparently unique (in Chamaemyiidae) set of characteristics of the female terminalia is described, figured and discussed.

Made available in DSpace on 2018-12-11T17:19:22Z (GMT). No. of bitstreams: 0 Previous issue date: 2018-03-14 Museu de Zoologia da Universidade de São Paulo Chelonian Research Institute Bernice Pauahi Bishop Museum International Commission on Zoological Nomenclature Department of Marine Invertebratesn Museum School of Biological Earth & Environmental Sciences University of New South Wales Departamento de Biodiversidad y Biología Evolutiva Museo Nacional de Ciencias Naturales (CSIC) Department of Biology University of Washington Universidad de Atacama Copiapó and Programa de Doctorado en Sistemática y Biodiversidad Facultad de Ciencias Naturales y Oceanográficas Universidad de Concepción Department of Biological Sciences National University of Singapore Biodiversity and Biocomplexity Unit Okinawa Institute of Science and Technology, Kunigami District Laboratório de Ictiologia Departamento de Zoologia IBB-UNESP Campus de Botucatu FishBase Information and Research Group IRRI Campus Comparative Plant and Fungal Biology Department Royal Botanic Gardens Kew Zoologische Staatssammlung München Staatliche Naturwissenschaftliche Sammlungen Bayerns Department of Life Sciences Natural History Museum Australian National Herbarium Centre for Australian National Biodiversity Research CSIRO National Research Collections Australian Capital Territory College of Medicine Biology and Environment Research School of Biologyn National Universityn Capital Territory Hawkesbury Institute for the Environment Western Sydney University Canadian National Collection of Insects Arachnids and Nematodes Agriculture and Agri-Food Canada Institut de Systématique Evolution Biodiversité Muséum National d'Histoire Naturelle Sorbonne Universités Division of Invertebrate Zoology American Museum of Natural History Department of Biology Hofstra University Naturalis Biodiversity Center Royal Belgian Institute of Natural Sciences School of Life Sciences University of KwaZulu-Natal Plant Gateway Ltd. Department of Natural Sciences Gardner-Webb University Department of Biology Villanova University Potato Research Center Georgikon Faculty University of Pannonia Dipartimento di Biologia e Biotecnologia 'Charles Darwin' Sapienza Università di Roma Department of Earth and Planetary Sciences Johns Hopkins University Department of Environmental Science and Technology University of Maryland Department of Plant Medicine Chungbuk National University Hawai‘i Institute of Marine Biology University of Hawai‘i Institute of Marine Science University of Auckland Centre for Agricultural Research Hungarian Academy of Sciences Department of Zoology Eszterházy Károly University Herbarium School of Biological Sciences University of Reading Instituto de Ciencias Ambientales y Evolutivas Facultad de Ciencias Universidad Austral de Chile Department of Biosciences University of Helsinki Ornithological Nomenclature Expert Green Plant Herbarium Department of Natural History Royal Ontario Museum Nomenclature Specialist CITES Animals Committee; Turtle Conservancy Global Wildlife Conservation Department of Plant Anatomy Institute of Biology Eötvös Loránd University Department of Biosciences (Plant Biology) Viikki Plant Science Centre Illinois Natural History Survey Institute of Biodiversity Science and Sustainability California Academy of Sciences Royal Botanic Gardens Kew NTNU University Museum Norwegian University of Science and Technology Institut de Biologia Evolutiva Universitat Pompeu Fabra Bioveyda Biodiversity Inventories and Research Department of Plant Biology Ecology & Evolution and Herbarium Oklahoma State University Research School of Earth Sciencesn National Universityn Capital Territory Departamento de Biologia Animal Instituto de Biologia Universidade Estadual de Campinas Senckenberg Naturhistorische Sammlungen Department of Biology University of Copenhagen Department of Botany Smithsonian Institution National Museum of Natural History Plant Pest Diagnostics Branch California Department of Food and Agriculture PPG Zoologia Departamento de Zoologia Instituto de Ciências Biológicas Universidade Federal de Minas Gerais Macleay Museum University of Sydney Centre for Ecology and Hydrology Edinburgh Research Station Laboratório de Coleções Zoológicas Instituto Butantan Environmental Management School of Applied and Biomedical Science Federation University PPG Biodiversidade Animal Universidade Federal de Santa Maria National Museum of Natural History Smithsonian Institution Department of Terrestrial Zoology Western Australian Museum Museu Nacional da História Natural e da Ciência Universidade de Lisboa The Kyoto University Museum Kyoto University Finnish Museum of Natural History (Botany) University of Helsinki Royal Botanic Garden Edinburgh School of Biological Sciences and Environment Institute University of Adelaide Nees Institut für Biodiversität der Pflanzen Universität Bonn Department of Plant Science and Biotechnology University of Pannonia Lee Kong Chian Natural History Museum National University of Singapore Denver Botanic Gardens Department of Biology Earlham College Montana Entomology Collection Montana State University School of Environmental Sciences University of Guelph Instituto de Ambiente de Montaña y Regiones Áridas Universidad Nacional de Chilecito Martha N. and John C. Moser Chair in Arthropod Systematics and Biological Diversity The Ohio State University Department of Biology Victor Valley College Identification and Naming Department Royal Botanic Gardens Kew Natural History Laboratory Faculty of Science Ibaraki University Natural History Museum and Botanical Garden University of Tartu Plant Protection Institute Centre for Agricultural Research Hungarian Academy of Sciences Department of Zoology Denver Museum of Nature & Science Department of Systematic and Evolutionary Botany Universität Wien FishBase Department of Zoology Swedish Museum of Natural History Department of Zoology Museo Civico di Storia Naturale of Verona Departamento de Zoologia Universidade Federal do Paraná Sezione di Zoologia degli Invertebrati e Idrobiologia MUSE-Museo delle Scienze di Trento Centro de Ciências Agrárias Ambientais e Biológicas Universidade Federal do Recôncavo da Bahia Department of Biological Sciences University of Toronto Department of Environmental Science and Policy George Mason University Natural History Museum of Geneva Department of Genetics and Evolution University of Geneva Departamento de Zoologia Universidade de Brasília Department of Integrative Biology Oregon State Arthropod Collection Oregon State University Henares 16 Velilla de San Antonio Department of Vertebrate Zoology National Museum of Natural History Smithsonian Institution International Committee on Bionomenclature Universidade Estadual Paulista (Unesp) Faculdade de Ciências Agrárias e Veterinárias Jaboticabal Departamento de Biologia Aplicada à Agropecuária Faculty of Biology Alexandru Ioan Cuza University of Iași Departamento de Biología Universidad de Los Andes Laboratorio de Morfología Animal Universidad Autónoma del Estado de Hidalgo H.W. 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The Chinese fauna of the genus Cestrotus Loew (Diptera: Lauxaniidae) is revised, including the five new species Cestrotus acuticurvus sp. nov., Cestrotus heteropterus sp. nov., Cestrotus liui sp. nov., Cestrotus longinudus sp. nov. and Cestrotus obtusus sp.nov., and the two previously described species Cestrotus apicalis (Hendel) and Cestrotus flavoscutellatus de Meijere. Extralimital records are also given for Cestrotus flavoscutellatus and Cestrotus heteropterus, including paratypes of the latter species from Thailand. A key to the species of Cestrotus from China is presented, and a list of all described species of Cestrotus is provided, with information on type locality, primary type specimen(s) and depository, synonymy, and distribution.

Three new genera of Odiniidae (Diptera) are described from Costa Rica, including Helgreelia gen. nov. (type species, albeto sp. nov.; additional species, parkeri sp. nov.) and the monotypic genera Neoschildomyia gen. nov. (type species, fusca sp. nov.) and Pradomyia gen. nov. (type species, hadromera sp. nov.). A key to the New World genera of the family is provided.

Fauna Europaea provides a public web-service with an index of scientific names (including important synonyms) of all extant multicellular European terrestrial and freshwater animals and their geographical distribution at the level of countries and major islands (east of the Urals and excluding the Caucasus region). The Fauna Europaea project comprises about 230,000 taxonomic names, including 130,000 accepted species and 14,000 accepted subspecies, which is much more than the originally projected number of 100,000 species. Fauna Europaea represents a huge effort by more than 400 contributing taxonomic specialists throughout Europe and is a unique (standard) reference suitable for many user communities in science, government, industry, nature conservation and education. The Diptera-Brachycera is one of the 58 Fauna Europaea major taxonomic groups, and data have been compiled by a network of 55 specialists.Within the two-winged insects (Diptera), the Brachycera constitute a monophyletic group, which is generally given rank of suborder. The Brachycera may be classified into the probably paraphyletic 'lower brachyceran grade' and the monophyletic Eremoneura. The latter contains the Empidoidea, the Apystomyioidea with a single Nearctic species, and the Cyclorrhapha, which in turn is divided into the paraphyletic 'aschizan grade' and the monophyletic Schizophora. The latter is traditionally divided into the paraphyletic 'acalyptrate grade' and the monophyletic Calyptratae. Our knowledge of the European fauna of Diptera-Brachycera varies tremendously among families, from the reasonably well known hoverflies (Syrphidae) to the extremely poorly known scuttle flies (Phoridae). There has been a steady growth in our knowledge of European Diptera for the last two centuries, with no apparent slow down, but there is a shift towards a larger fraction of the new species being found among the families of the nematoceran grade (lower Diptera), which due to a larger number of small-sized species may be considered as taxonomically more challenging.Most of Europe is highly industrialised and has a high human population density, and the more fertile habitats are extensively cultivated. This has undoubtedly increased the extinction risk for numerous species of brachyceran flies, yet with the recent re-discovery of Thyreophora cynophila (Panzer), there are no known cases of extinction at a European level. However, few national Red Lists have extensive information on Diptera.For the Diptera-Brachycera, data from 96 families containing 11,751 species are included in this paper.

We summarise the findings of arthropods collected by fogging the canopy of 24 pine trees in two sites in Eastern and Western Norway. From the samples, taken in 1998 and in 1999, almost 30,000 specimens were determined to 512 species,with Diptera being most species rich (210 species), followed by Coleoptera (76 species) and Araneae (49 species). Of the 96 new species records, nine were new to science (5 Diptera and 4 Oribatida), two were new to the European, three to the Scandinavian and 82 to the Norwegian faunas. The paper demonstrates the need for detailed faunistical inventories of European forests.

A new genus and species of Chamaemyiidae (Diptera: Lauxanioidea) is described and illustrated, namelyChamaeleucopis, gen. nov. (type species Chamaeleucopis trevas, sp. nov.), a predator of Ceroplastes wax scales on guavafrom Brazil. In addition, the previously unrecognized genus Ortalidina Blanchard 1852 (type species Ortalidina cellularisBlanchard 1852), mainly from the Neotropics, is recognized as a chamaemyiid, and is the senior synonym of ToropameciaCogan 1978, syn. nov. (type species Acrometopia punctata Coquillett 1902). The species Toropamecia grossa Cogan1978, syn. nov., is synonymized under Ortalidina cellularis. All species therein become new combinations withinOrtalidina, including (in their original combinations) Acrometopa australis Malloch 1933, comb. nov., Acrometopiamaculata Coquillett 1902, comb. nov., Acrometopia punctata Coquillett, comb. nov., Toropamecia apaxa Cogan 1978,comb. nov., Toropamecia caribbea Cogan 1978, comb. nov., Toropamecia hendeli Cogan 1978, comb. nov., Toropameciahyalipennis Cogan 1978, comb. nov., Toropamecia jujuyensis Cogan 1978, comb. nov., Toropamecia longipennis Cogan1978, comb. nov., Toropamecia macalpinei Cogan 1978, comb. nov., Toropamecia multipunctata Cogan 1978, comb. nov.,Toropamecia nigripalpis Cogan 1978, comb. nov., Toropamecia reducta Cogan 1978, comb. nov., Toropamecia smithiCogan 1978, comb. nov., Toropamecia veenota Cogan 1978, comb. nov., Trigonometopus reticulatus Johnson 1913, comb. nov. A habitus photo of the primary type for each species of Ortalidina is provided.

The proportion of the width of the lower margin of the face and the height of the gena is proposed to sort several speciesof Homoneura Wulp in the two subgenera Euhomoneura Malloch and Homoneura where the anteriormost dc is locatedat the transverse suture of the mesonotum. One species Homoneura (Homoneura) litorea sp. nov. is described as new toscience, and two species, Homoneura (Homoneura) stackelbergiana Papp and Homoneura (Minettioides) orientis (Hen-del) are recorded from China for the first time. The former species is elevated out of synonymy from Homoneura(Homoneura) stackelbergi Czerny. The species Homoneura alini Hering is synonymized under Homoneura (Homoneura)brevicornis Kertész. A key is presented to separate the five subgenera of Homoneura, and another key is presented to separate the Chinese species of the subgenera Euhomoneura and Minettioides Malloch.

The following three new species of the genus Hybos Meigen, from Guangdong Province, China, are described as new to science: Hybos mangshanensis, H. nankunshanensis, H. xiaohuangshanensis. Their relationships with related species are discussed.

The new genus Evertomyia (Diptera: Lauxaniidae) from New Caledonia is described and illustrated, along with six new species, albeto, frankadelia, irwini, helenae, matilei and webbi. Phylogenetic relationships among the six species are hypothesized, character states are discussed, and a key to the genus and its species is provided.