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Nerita cf. funata Dujardin, 1837 Plate 1 E 1 -E 3 cf. * Nerita funata Dujardin 1837: 281, pl. 19, fig. 14. ? Nerita Plutonis Bast. —Bronn in Reiss 1862: 33. ? Nerita Plutonis Bast. — Mayer 1864: 63, 93. cf. Nerita (Theliostyla) funata Dujardin, 1837 — Glibert 1949: 69, pl. 4, fig. 8. Santa Maria material examined. Maximum height 4.0 mm, width 6.0 mm. DBUA-F 917-1 (1), Ponta do Castelo, Santa Maria Island, Azores, Touril Complex, Lower Pliocene. Description. Apical fragment with completely flattened spire. Teleoconch bearing about 22 subequal rounded ribs, slightly narrower than their interspaces, shoulder cord slightly strengthened. Discussion. Mayer (1864: 63) recorded two specimens from Bocca do Cré (= Cré), Santa Maria Island, without indicating their size. The Azorean material is very incomplete, but clearly represents a Nerita species of the Theliostyla M̂rch, 1852 group. Two species occurred along the French Atlantic Frontage during the Miocene. Nerita plutonis de Basterot, 1825, from the Lower Miocene, has relatively coarse cords of irregular strength and N. funata Dujardin, 1837, from the Middle Miocene, has finer, subequal cords. Lozouet et al. (2001: 20) considered the Lower Miocene records of N. funata in Cossmann & Peyrot (1917: pl. 7, figs. 77-82) to represent N. plutonis. The fragment at hand from Santa Maria Island, has numerous subequal ribs more closely similar to those seen in N. funata than N. plutonis. However, we hesitate to make a definitive identification based on this very incomplete specimen. The historical Azorean specimens identified by Bronn in Reiss (1862: 33) and Mayer (1864: 63, 93) as N. plutonis are likely to represent the same species. This group of Nerita species with spiral cords is not represented in the Pliocene Mediterranean. Today, the group is represented in the eastern Atlantic by N. senegalensis Gmelin, 1791, distributed from the Cabo Verde Islands to Angola, whereas in the western Atlantic a total of five species are reported, four of them from Bermuda, the Gulf of Mexico and the Caribbean (Nerita fulgurans Gmelin, 1791; Nerita peloronta Linnaeus, 1758; Nerita tessellata Gmelin, 1791; and Nerita versicolor Gmelin, 1791), and one species endemic to Brazil (Nerita chlorostoma Lamarck, 1816). Nerita senegalensis differs in having flattened cords of irregular width, separated by narrow grooves. Distribution. Lower Pliocene: Atlantic, Santa Maria Island, Azores (?Bronn in Reiss 1862;? Mayer 1864; this paper)., Published as part of Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, pp. 1-150 in Zootaxa 5295 (1) on pages 27-28, DOI: 10.11646/zootaxa.5295.1.1, http://zenodo.org/record/7965273, {"references":["Dujardin, F. (1837) Memoire sur les couches du sol en Touraine et description des coquilles de la craie et des faluns. Meimoire de la Socieitei Geiologique de la France, 2, 211 - 311.","Reiss, W. (1862) Mittheilungen u ¨ ber die tertia ¨ ren Schichten von Santa Maria, der su ¨ dlichsten der Azoren, und ihre organischen Einschlu ¨ sse. Nebst Beschreibung dieser letzten und Abbildung der neuen Arten, von H. G. Bronn. Neues Jahrbuch fu ¨ r Mineralogie, Geognosie, Geologie und Petrefactunkunde, 1862, 1 - 48.","Mayer, K. (1864). Die Tertiar-fauna der Azoren und Madeiren. Zurich, published by the author. Z ¸ rich: vi + 107 pp. + 7 pls.","Glibert, M. (1949) Gastropodes du Miocene moyen du Bassin de la Loire. 1. Memoires de l'Institut Royal des Sciences Naturelles de Belgique, 2 (30), 1 - 240.","Basterot, B. de (1825) Meimoire geiologique sur les environs de Bordeaux. PremieIre partie, comprenant les observations geineirales sur les mollusques fossiles, et la description particulieIre de ceux qu'on rencontre dans ce bassin. Tastu, J., Paris, 100 pp. [reprinted from Meimoires de la Socieitei d'Histoire Naturelle de Paris, 2, 1 - 100]","Lozouet, P., Lesport, J. F. & Renard, P. (2001) Revision des Gastropoda (Mollusca) du stratotype de l'Aquitanien (Miocene inf.): site de Saucats ' Lariey', Gironde, France. Cossmanniana, 3, 1 - 189.","Gmelin, J. F. (1791) Caroli a Linnei systema natura per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, disserentis, synonymis, locis etc. Editio decima tertia, aucta, reformata, cura J. F. Gmelin. 1 (6). Vermes testacea. G. E. Beer, Lipsiae, pp. 3021 - 4120.","Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1. Editio decima, reformata. Laurentii Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542","Lamarck, J. B. P. A. de M. (1816) TTableau encyclope´dique et me´thodique des trois re`gnes de la nature ..., 23. Mollusques et polypes divers. Liste des objets repre´sente´s dans les planches de cette livraison. Veuve Agasse, Paris, 16 pp."]}

Genus Nerita Linnaeus, 1758 Type species. Nerita peloronta Linnaeus, 1758 (Linnaeus 1758: 776), by subsequent designation (Montfort 1810). Present-day, Caribbean., Published as part of Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, pp. 1-150 in Zootaxa 5295 (1) on page 27, DOI: 10.11646/zootaxa.5295.1.1, http://zenodo.org/record/7965273, {"references":["Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. 1. Editio decima, reformata. Laurentii Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542","Montfort, D. de (1810) Conchyliologie syste´matique, ou classification me´thodique des coquilles; offrant leurs figures, leur arrangement ge´ne´rique, leurs descriptions caracte´ristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Ouvrage destine´a`faciliter l'e´tude dew coquilles, ainsi que leur disposition dans les cabinets d'histoire naturelle. Coquilles univalves, non cloissonne´es. Coquilles univalves, non cloisonne´es. Schoell, F., Paris, 676 pp."]}

Nabq as “A Managed Resource Protected Area” or as “Multiple Use Management Area” is subjected to human activities that influence its biodiversity. Therefore, many field trips from January 2016 to January 2017 were carried out to detect the main activities in Nabq protectorate and their impacts on faunal macro-benthos biodiversity. Four stations representing two major habitats (mangrove and rocky shore) were chosen depending on anthropogenic activities. A total of 112 macro-benthos taxa were recorded in Nabq dominated with Planaxis salcatus, Nerita spp., Barbatia trapezina and Ophiocoma scolopendrina. The anthropogenic activities don't only affect the presence and absence of species, but also influence on the dominance status of species in the investigated stations. El-Rowayseaa and El-Dagal were unaffected stations showing more abundance and diversification than El-Gharkana and El-Monkateaa which were affected mangrove and rocky shore stations, respectively. In conclusion, anthropogenic activities are the main cause of recent changes to the marine biodiversity in Nabq Protected Area and their continuation may cause habitat damage. Keywords: Macro-benthos, Biodiversity, Human activities, Ecotourism, Fishing, Protected areas

Genus Nerita Linnaeus, 1758 Type species: Nerita peloronta Linnaeus, 1758, by subsequent designation (Montfort 1810). Recent, Florida., Published as part of DeVries, Thomas J., 2019, Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru), pp. 1533-1584 in Journal of Natural History 53 (25) on page 1552, DOI: 10.1080/00222933.2018.1524032, http://zenodo.org/record/3670229, {"references":["Montfort P (D). 1810. Conchyliologie systematique et classification methodique des coquilles. Volume II. Paris: Schoell."]}

Nerita undata Linnaeus, 1758 Figs 3 G–H Nerita undata Linnaeus, 1758: 779. Nerita (Nerita) undata: Okutani 2000: 102 –103, pl. 51, fig. 5; Thach 2005: 36, pl. 6, figs 33–34. Nerita undata: Poppe 2008a: 274 –275, pl. 82, fig. 10; Tan & Clements 2008: 487, figs 3-21, 3-22; Zvonareva et al. 2015: 8. Material examined. Dam Bay, natural mangrove, about 10 specimens. Abundance. Uncommon. Measurements. Up to 33 mm (Tan & Clements 2008). Ecological notes. Development with free swimming veliger (Fukumori & Kano 2014). Found in crevices and under rocks of breakwaters and rocky shores of middle to upper intertidal zones. Nocturnal, emerging from their hiding places at dusk (Tan & Clements 2008). In studied sites it was found on rocks in the lower intertidal zone. Relation to mangroves. Eurybiotic. Distribution in Vietnam. From northern-east coast to southern delta (Thach 2005). General distribution. IWP: northeastern South Africa to Kenya, ranging through central region, including Indonesia and the Philippines, to Melanesia and Micronesia, southern Japan to northern Australia (Frey 2010)., Published as part of Zvonareva, Sofya & Kantor, Yuri, 2016, Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam, pp. 401-437 in Zootaxa 4162 (3) on page 408, DOI: 10.11646/zootaxa.4162.3.1, http://zenodo.org/record/254973, {"references":["Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae. Vol. 1. 10 th Edition. Salvius, Stockholm, 823 pp.","Okutani, T. (2000) Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pp.","Thach, N. N. (2005) Shells of Vietnam. Conchbooks, Hackenheim, 338 pp.","Poppe, G. T. (2008 a) Volume I: Philippine marine mollusks. ConchBooks, Hackenheim, 759 pp.","Tan, S. & Clements, R. (2008) Taxonomy and distribution of the Neritidae (Mollusca: Gastropoda) in Singapore. Zoological Studies, 47, 481 - 494.","Zvonareva, S., Kantor Yu., Xinzheng, L. & Britayev, T. (2015) Long-term monitoring of Gastropoda (Mollusca) fauna in planted mangroves in central Vietnam. Zoological studies, 54, 1 - 39.","Fukumori, H. & Kano, Y. (2014) Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology, 161, 213 - 227. http: // dx. doi. org / 10.1007 / s 00227 - 013 - 2330 - 5","Frey, M. A. (2010) The relative importance of geography and ecology in species diversification: evidence from a tropical marine intertidal snail (Nerita). Journal of Biogeography, 37, 1515 - 1528. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2010.02283. x"]}

Nerita histrio Linnaeus, 1758 Figs 3 C–D Nerita histrio Linnaeus, 1758: ii, 824 pp. Nerita (Theliostyla) squamulata Le Guillou, 1841: Okutani 2000: 102 –103, pl. 51, fig. 8; Thach 2005: 38, pl. 6, figs 1, 6, 21. Nerita histrio: Lozouet & Plaziat 2008: 106 –107, pl. 17, figs 5–8; Tan & Clements 2008: 486, figs 2-11, 2-12. Nerita squamulata: Zvonareva et al. 2015: 8. Material examined. Nha Phu, Dam Bay, in natural and planted mangroves, about 20 specimens. Abundance. Common. Measurements. Up to 20 mm (Okutani 2000). Ecological notes. Development with free swimming veliger (Fukumori & Kano 2014). Found on mangrove tree trunks and stilt roots, on muddy rocks at coastal fringes of mangroves, on boulders in sheltered waters or estuary (Okutani 2000; Tan & Clements 2008). In studied sites it was found on sediment surface under mangrove trees and occasionally on pneumatophores of Rhizophora trees. Relation to mangroves. Eurybiotic. Distribution in Vietnam. Northern - east coast, central coast, southern delta (Thach 2005). General distribution. IWP: India to Samoa, and southern Japan to northern Australia (Frey 2010)., Published as part of Zvonareva, Sofya & Kantor, Yuri, 2016, Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam, pp. 401-437 in Zootaxa 4162 (3) on pages 407-408, DOI: 10.11646/zootaxa.4162.3.1, http://zenodo.org/record/254973, {"references":["Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae. Vol. 1. 10 th Edition. Salvius, Stockholm, 823 pp.","Okutani, T. (2000) Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pp.","Thach, N. N. (2005) Shells of Vietnam. Conchbooks, Hackenheim, 338 pp.","Lozouet, P. & Plaziat, J. - C. (2008) Mangrove environments and mollusks. Abatan River, Bohol and Panglao Islands, Central Philippines. ConchBooks, Hackenheim, 166 pp.","Tan, S. & Clements, R. (2008) Taxonomy and distribution of the Neritidae (Mollusca: Gastropoda) in Singapore. Zoological Studies, 47, 481 - 494.","Zvonareva, S., Kantor Yu., Xinzheng, L. & Britayev, T. (2015) Long-term monitoring of Gastropoda (Mollusca) fauna in planted mangroves in central Vietnam. Zoological studies, 54, 1 - 39.","Fukumori, H. & Kano, Y. (2014) Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology, 161, 213 - 227. http: // dx. doi. org / 10.1007 / s 00227 - 013 - 2330 - 5","Frey, M. A. (2010) The relative importance of geography and ecology in species diversification: evidence from a tropical marine intertidal snail (Nerita). Journal of Biogeography, 37, 1515 - 1528. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2010.02283. x"]}

Nerita albicilla Linnaeus, 1758 Figs 3A–B Nerita albicilla Linnaeus, 1758: 778. Nerita (Theliostyla) albicilla: Okutani 2000: 104 –105, pl. 52, fig. 13; Thach 2005: 37, pl. 6, figs 24–25. Nerita albicilla: Tan & Clements 2008: 483, figs 2-1, 2-2; Poppe 2008a: 268 –268, pl. 79, fig. 6; Zvonareva et al. 2015: 8. Material examined. Dam Bay, natural mangrove, about 10 specimens. Abundance. Uncommon. Measurements. Up to 32 mm (Tan & Clements 2008). Ecological notes. Development with free swimming veliger (Fukumori & Kano 2014). Found on breakwaters and seawalls in lower to middle intertidal zone (Tan & Clements 2008). In studied sites it was found on rocks in the lower intertidal zone. Relation to mangroves. Eurybiotic. Distribution in Vietnam. Northern-east coast, northern delta, central coast (Thach 2005). General distribution. IWP: Southeastern South Africa to Hawaiian Is (Frey 2010)., Published as part of Zvonareva, Sofya & Kantor, Yuri, 2016, Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam, pp. 401-437 in Zootaxa 4162 (3) on page 407, DOI: 10.11646/zootaxa.4162.3.1, http://zenodo.org/record/254973, {"references":["Linnaeus, C. (1758) Systema Naturae per Regna tria Naturae. Vol. 1. 10 th Edition. Salvius, Stockholm, 823 pp.","Okutani, T. (2000) Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pp.","Thach, N. N. (2005) Shells of Vietnam. Conchbooks, Hackenheim, 338 pp.","Tan, S. & Clements, R. (2008) Taxonomy and distribution of the Neritidae (Mollusca: Gastropoda) in Singapore. Zoological Studies, 47, 481 - 494.","Poppe, G. T. (2008 a) Volume I: Philippine marine mollusks. ConchBooks, Hackenheim, 759 pp.","Zvonareva, S., Kantor Yu., Xinzheng, L. & Britayev, T. (2015) Long-term monitoring of Gastropoda (Mollusca) fauna in planted mangroves in central Vietnam. Zoological studies, 54, 1 - 39.","Fukumori, H. & Kano, Y. (2014) Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology, 161, 213 - 227. http: // dx. doi. org / 10.1007 / s 00227 - 013 - 2330 - 5","Frey, M. A. (2010) The relative importance of geography and ecology in species diversification: evidence from a tropical marine intertidal snail (Nerita). Journal of Biogeography, 37, 1515 - 1528. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2010.02283. x"]}

Nerita planospira Anton, 1838 Figs 3 E–F Nerita planospira Anton, 1838: 30. Nerita (Theliostyla) planospira: Okutani 2000: 104 –105, pl. 52, fig. 14. Nerita planospira: Poppe 2008a: 272 –273, pl. 81, fig. 3; Tan & Clements 2008: 486, figs 2-13, 2-14; Lozouet & Plaziat 2008: 106 –107, pl. 17, figs 11–12. Material examined. Dam Bay, natural mangrove, 5 specimens. Abundance. Uncommon. Measurements. Up to 29 mm (Tan & Clements 2008). Ecological notes. Development with free swimming veliger (Fukumori & Kano 2014). Found on trunks and roots of mangrove trees, on rocks and dead wood from sea edge to the upper estuary, mainly in peripheral belt but also within internal mangroves (Lozouet & Plaziat 2008; Tan & Clements 2008; Frey 2010). In studied sites it was found on rocks and Rhizophora pneumatophores. Relation to mangroves. Predominantly mangrove-associated. Distribution in Vietnam. Central coast. General distribution. From Indo-Malayan core to the Ryu Kyu Is (Lozouet & Plaziat 2008)., Published as part of Zvonareva, Sofya & Kantor, Yuri, 2016, Cheсklist of gastropod molluscs in mangroves of Khanh Hoa province, Vietnam, pp. 401-437 in Zootaxa 4162 (3) on page 408, DOI: 10.11646/zootaxa.4162.3.1, http://zenodo.org/record/254973, {"references":["Anton, H. E. (1838) [\" 1839 \"] Verzeichnis der Conchylien welche sich in der Sammlung von Herrmann Eduard Anton befinden. Herausgegeben von dem Besitzer. Anton, Halle, xvi + 110 pp. [date of publication after W. O. Cernohorsky, 1978. The Veliger, 20 (3), 299.] http: // dx. doi. org / 10.5962 / bhl. title. 11509","Okutani, T. (2000) Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pp.","Poppe, G. T. (2008 a) Volume I: Philippine marine mollusks. ConchBooks, Hackenheim, 759 pp.","Tan, S. & Clements, R. (2008) Taxonomy and distribution of the Neritidae (Mollusca: Gastropoda) in Singapore. Zoological Studies, 47, 481 - 494.","Lozouet, P. & Plaziat, J. - C. (2008) Mangrove environments and mollusks. Abatan River, Bohol and Panglao Islands, Central Philippines. ConchBooks, Hackenheim, 166 pp.","Fukumori, H. & Kano, Y. (2014) Evolutionary ecology of settlement size in planktotrophic neritimorph gastropods. Marine Biology, 161, 213 - 227. http: // dx. doi. org / 10.1007 / s 00227 - 013 - 2330 - 5","Frey, M. A. (2010) The relative importance of geography and ecology in species diversification: evidence from a tropical marine intertidal snail (Nerita). Journal of Biogeography, 37, 1515 - 1528. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2010.02283. x"]}

Marine biogeographers have long recognized broad east‐west differentiation of marine communities across southern Australia, but few studies have explicitly assessed the site of disjunction or the factors potentially underlying this biodiversity. A recent, detailed phylogeographical and distributional study of the dispersive gastropod genus Nerita revealed an abrupt shift in species abundance across mainland Australia, apparently correlated with the site of an historical vicariant barrier: the Bassian Isthmus. In the current study I provide an independent assessment of this vicariant hypothesis by morphologically analysing over 3000 intertidal Nerita specimens from eight coastal sites around Tasmania. Consistent with the Bassian Isthmus hypothesis, the study reveals a dramatic east‐west disjunction across north-east Tasmania. A survey of marine biogeographical literature reveals several additional examples supporting the east‐west species turnover detected in Nerita . I discuss the role of contemporary oceanographic factors in preserving the biogeographical signature of marine vicariance, even in highly dispersive taxa. Based on recent marine connectivity data, the east‐west disjunction in Nerita taxa can be interpreted as an historic vicariant pattern perpetuated by contemporary oceanographic conditions. The results of this study emphasize the potential importance of considering relative abundance data ‐ rather than just species range data ‐ in marine biogeographical analyses. As the observed disjunction is likely to have broad implications for Australia’s marine biodiversity, it is imperative that conservation biologists incorporate such data in the design of marine protected areas.

urceus Nerita urceus M��ller, 1774. Additional detail of type material ��� syntype: ZMUC GAS 242 (ex Moltke collection); possible syntypes: ZMUC GAS 243 (1 spm.), ZMUC GAS 244 (1 spm.) (both probably ex Moltke collection) [not found in ZMUC in 2002 (Cowie & Thiengo, 2003: 76)]., Published as part of Cowie, Robert H., 2015, The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda, pp. 1-92 in Zootaxa 3940 (1) on page 65, DOI: 10.11646/zootaxa.3940.1.1, http://zenodo.org/record/288204, {"references":["Muller, O. F. (1774) Vermium terrestrum et fluviatilium, seu animalium infusoriorum, helminthicorum, et testaceorum, non marinorum, succincta historia. Volumen alterum. Heineck & Faber, Havniae [= Copenhagen] & Lipsiae [= Leipzig], xxxv + [1] + 214 + [10] pp. http: // dx. doi. org / 10.5962 / bhl. title. 46299","Cowie, R. H. & Thiengo, S. C. (2003) The apple snails of the Americas (Mollusca: Gastropoda: Ampullariidae: Asolene, Felipponea, Marisa, Pomacea, Pomella): a nomenclatural and type catalog. Malacologia, 45 (1), 41 - 100. [29 August; Malacologia 46 (1); un-numbered page at end of issue]"]}

Nerita vaitapihana Curtiss, 1938, a synonym of Nerita argus R��cluz, 1841 Original description (pp. 192, 193). ���The black Tahiti nerita, which shares with the last the name of hihi, is fiveeighths of an inch long, dull black above and whitish underneath. Its shell is smooth, without the furrows of the last-described sort. Its spire is very indistinct, much more so than that of the whitish nerita. The opening is semicircular, with very small teeth. The edge of the outer lip is sharp and black, further down, the lip becomes thicker and whiter. The animal resembles a snail, and closes its entrance with the brown membrane on the bottom of its foot. It is found on rocks along the seaside. (Nerita vaitapihana. (Seaside, near Tautira.))���. Identity. The description of this ���black Tahiti nerita ��� agrees well with Nerita argus R��cluz, 1841, which is known from the French Polynesia (Tr��ndl�� & Boutet 2009). Although the similarly relatively smooth and black N. picea was not listed by Tr��ndl�� & Boutet (2009), it is apparently also found in Tahiti (Spencer et al. 2007). However, N. picea has incised spiral grooves on its dorsal surface (see Kay 1979) and its conspecificity with N. vaitapihana is unlikely., Published as part of Low, Martyn E. Y. & Tan, Siong Kiat, 2014, On the identities of the molluscan names described in A Short Zoology of Ta h i t i in the Society Islands by Anthony Curtiss in 1938 (Mollusca: Cephalopoda, Gastropoda), pp. 394-400 in Zootaxa 3764 (3) on page 399, DOI: 10.11646/zootaxa.3764.3.9, http://zenodo.org/record/226594, {"references":["Curtiss, A. (1938) A Short Zoology of Tahiti in the Society Islands. [Published by the Author]. Printed by Guide Printing, Brooklyn, New York, xvi + 193 pp.","Trondle, J. & Boutet, M. (2009) Inventory of marine molluscs of French Polynesia. Atoll Research Bulletin, 570, i - ii, 1 - 87. http: // dx. doi. org / 10.5479 / si. 00775630.570.1","Spencer, H. G., Waters, J. M. & Eichhorst, T. E. (2007) Taxonomy and nomenclature of black nerites (Gastropoda: Neritimorpha: Nerita) from the South Pacific. Invertebrate Systematics, 21 (3), 229 - 237. http: // dx. doi. org / 10.1071 / is 06038","Kay, E. A. (1979) Hawaiian marine shells. Reef and shore fauna of Hawaii, section 4: Mollusca. Bernice P. Bishop Museum Special Publication, 64 (4), i - xviii, 1 - 653."]}

International audience; The Southwest Indian Ocean (SWIO) is a striking marine biodiversity hotspot. Coral reefs in this region host a high proportion of endemics compared to total species richness and they are particularly threatened by human activities. The island archipelagos with their diverse marine habitats constitute a natural laboratory for studying diversification processes. Rocky shores in the SWIO region have remained understudied. This habitat presents a high diversity of molluscs, in particular gastropods. To explore the role of climatic and geological factors in lineage diversification within the genus Nerita, we constructed a new phylogeny with an associated chronogram from two mitochondrial genes [cytochrome oxidase sub-unit 1 and 16S rRNA], combining previously published and new data from eight species sampled throughout the region. All species from the SWIO originated less than 20 Ma ago, their closest extant relatives living in the Indo-Australian Archipelago (IAA). Furthermore, the SWIO clades within species with Indo-Pacific distribution ranges are quite recent, less than 5 Ma. These results suggest that the regional diversification of Nerita is closely linked to tectonic events in the SWIO region. The Reunion mantle plume head reached Earth's surface 67 Ma and has been stable and active since then, generating island archipelagos, some of which are partly below sea level today. Since the Miocene, sea-level fluctuations have intermittently created new rocky shore habitats. These represent ephemeral stepping-stones, which have likely facilitated repeated colonization by intertidal gastropods, like Nerita populations from the IAA, leading to allopatric speciation. This highlights the importance of taking into account past climatic and geological factors when studying diversification of highly dispersive tropical marine species. It also underlines the unique history of the marine biodiversity of the SWIO region.

Five gastropod taxa of the family Neritidae in the Ypresian and Bartonian (Eocene) of the Paris and Basse-Loire Basins, are introduced: Cuisenerita gen. nov., Tomostoma angusta sp. nov., Nerita gouetensis sp. nov., Clithon barbei sp. nov., Neritodryas guillioui sp. nov. and Cuisenerita tuberosa sp. nov. New combinations introduced are: Neritodryas dutemplei (Deshayes, 1864) and Neritodryas globosa (J. de C. Sowerby, 1823). This is the earliest record of Neritodryas and the first record for this genus from Europe.

Genus Nerita Linn��, 1758 Type species. By subsequent designation, Montfort (1810: 347): Nerita peloronta Linn��, 1758. Recent, marine, Caribbean. Diagnosis. Sturdy shells, smooth to spirally ribbed; inner lip septum well developed, its surface commonly pustulose or irregularly ribbed (Keen 1960)., Published as part of Symonds, Malcolm Francis & Pacaud, Jean-Michel, 2010, New species of Neritidae (Neritimorpha) from the Ypresian and Bartonian of the Paris and Basse-Loire Basins, France, pp. 55-68 in Zootaxa 2606 on page 58, DOI: 10.5281/zenodo.197721, {"references":["Linne, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio decima, reformata. Vol. 1. Regnum Animale. Salvius, Stockholm, 824 pp.","Montfort, D. de. (1810) Conchyliologie Systematique et Classification Methodique des Coquilles. Vol 2. F. Schoell, Paris, 676 pp.","Keen, M. (1960) In: Moore, R. (Ed.), Treatise on Invertebrate Palaeontology. Part 1. Mollusca 1. The Geological Society of America & The University of Kansas, U. S. A., 351 pp., 216 figs."]}

Nerita (Amphinerita) gouetensis sp. nov. (Fig. 2 a���c) Type material. Holotype MNHN A 31489 (Pacaud leg., Fig. 2 a���c). Stratum typicum. Late Eocene, Bartonian, ���Sands of Bois-Gou��t���. Locus typicus. Bois-Gou��t, Saffr��, Loire-Atlantique, France. 47 ˚ 29 �� 9.25 ˝ N 1 ˚ 36 �� 45.55 ˝ E. Derivatio nominis. Named after Bois-Gou��t, the type locality and the only locality at which this species has, as yet, been found. Diagnosis. A small Nerita with prominent, rounded, spiral ribs and a smooth, edentate septum. Description. Protoconch badly worn in only known specimen. Teleoconch of just over two whorls increasing rapidly in size, earlier rather worn in holotype. Last whorl bears nine prominent, smooth, rounded, spiral ribs, mostly evenly spaced; 6 th and 7 th below suture closer together. Apart from spiral ribs, surface of teleoconch smooth, lacking any collabral ornamentation other than some fine, rather faint growth lines. No colour pattern discernable but brown colour remains between ribs where shell relatively unworn. Slightly worn septum smooth, edge slightly concave, edentate. Outer lip thickened within, smooth apart from small protuberance about 1 mm from adapical end of septum. Gutter at junction of adapical end of septum and outer lip. Apertural tooth prominent, below abapical end of septum, in form of apostrophe with thicker end inward, higher, sloping to tail of apostrophe. Size. The holotype is approximately 4 mm high, and 4 mm wide. Remarks. Nerita (Amphinerita) gouetensis does not fit precisely within any of the generally accepted subgenera of Nerita. Those that share diagnostic characteristics with Na. (A.) gouetensis are: Lisanerita Krinjen, 2002 (Type species by original designation: Nerita lirellata Rehder, 1980); Heminerita Martens, 1887 (type species by monotypy: Nerita pica Gould, 1859 = Nerita japonica Dunker, 1861); Amphinerita Martens, 1887 (type species by subsequent designation Crosse, 1892: 98: Nerita umlaasiana Krauss, 1848); Cymostyla Martens, 1887 (type species by subsequent designation Crosse, 1892: 98: Nerita undata Linn��, 1758); Theliostyla M��rch, 1852 (type species by subsequent designation Kobelt, 1879: 147: Nerita albicilla Linn��, 1758); Subgenus Septum Septum edge Teleoconch Lisanerita Smooth Edentate or with very small teeth Smooth or with finely incised lines Heminerita Smooth Edentate or with 1 or 2 obscure Smooth or with low spiral lirae low broad projections Amphinerita Smooth Edentate or with small, obscure Smooth or with spiral ribs or well defined teeth In his subgenus Amphinerita, Martens (1887) included Na. umlaasiana and Na. georgina R��cluz, 1841. The former has an almost smooth shell but the latter has numerous well developed spiral ribs. Amphinerita is recorded in Europe from as early as the Thanetian [Na. (A.) semilugubris Deshayes, 1864, Le Renard & Pacaud 1995]. Although Na. (A.) gouetensis has fewer and broader ribs than Na. (A.) georgina, we consider that it can nevertheless be placed within Amphinerita. Nerita (A.) gouetensis is superficially similar to some juvenile specimens of Nerita (?) plutonis Basterot, 1825 from the Miocene of the south of France but its septum is normally neither smooth nor edentate but has granules on the surface and several well defined teeth on the edge. Range and distribution. Only known from type locality., Published as part of Symonds, Malcolm Francis & Pacaud, Jean-Michel, 2010, New species of Neritidae (Neritimorpha) from the Ypresian and Bartonian of the Paris and Basse-Loire Basins, France, pp. 55-68 in Zootaxa 2606 on pages 58-59, DOI: 10.5281/zenodo.197721, {"references":["Krinjen, C. (2002) The subgenera of the genus Nerita Linnaeus, 1758. Gloria Maris, 41, 38 - 69.","Rehder, H. A. (1980) The marine mollusks of Easter Island (Isla de Pascua) and Sala y Gomez. Smithsonian Contributions to Zoology, 289, 1 - 167.","Martens, E. von. (1887 - 1889) Die Gattungen Nerita und Neritopsis. In: Kuster, H. C. and Kobelt, W. (Eds.), Systematisches Conchylien-Cabinet von Martini und Chemnitz, 2 (11). Bauer & Raspe, Nurnberg, pp. 1 - 147, pls 1 - 15.","Gould, A. A., (1859) Descriptions of new species of shells. Proceedings of the Boston Society of Natural History, 7, 40 - 45.","Dunker, W. B. R. H. (1861) Mollusca Japonica Descripta et Tabulis Tribus Iconum. E. Schweizerbart, Stuttgart, 36 pp., 3 pls.","Crosse, H. (1892) Faune malacologique terrestre et fluviatile de l'ile de Portorico. Journal de Conchyliologie, 32, 5 - 71.","Krauss, F. (1848) Die Sudafrikanischen Mollusken. Ebner & Seubert, Stuttgart, 140 pp., 6 pls.","Linne, C. (1758) Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio decima, reformata. Vol. 1. Regnum Animale. Salvius, Stockholm, 824 pp.","Morch, O. A. L. (1852) Catalogus Conchyliorum quae Reliquit d'Alphonso d'Aguirra & Gadea Comes de Yoldi. 1. Hafni, Copenhagen, 1 - 170.","Kobelt, W. (1879) Illustrirtes Conchylienbuch. Bauer & Raspe, Nurnberg, 391 pp., 112 pls.","Recluz, C. A. (1841) Description de quelques nouvelles especes de nerites vivantes. Revue Zoologique par la Societe Cuvierienne, 4, 177 - 182.","Deshayes, G. P. (1864) Description des Animaux sans Vertebres Decouverts dans le Bassin de Paris. J. B. Bailliere et fils, Paris, 3, 1 - 200, pls 63 - 85.","Le Renard, J. & Pacaud, J-M. (1995) Revision des mollusques Paleogenes du Bassin de Paris II - Liste des references primaires des especes. Cossmanniana, 3, 65 - 132.","Basterot, B. de, (1825) Description geologique du Bassin Tertiaire du sud-ouest de la France. Memoires de la Societe d'Histoire Naturelle de Paris, 2, 1 - 100."]}

To determine how historical processes, namely speciation, extinction, and dispersal, have contributed to regional species diversity patterns across the marine tropics, we examined the biogeographical history of a circumtropical genus of intertidal gastropods. A species-level phylogeny of Nerita, representing approximately 87% of extant species, was developed from 1608 bp of mitochondrial (COI and 16S) and nuclear (ATPSα) markers. Phylogenetic relationships generally corresponded to prior classifications; however, comprehensive sampling revealed a number of previously undetected ESUs. Using the resulting tree as a framework, we combined geographical distributions and fossil evidence to reconstruct ancestral ranges, produce a time-calibrated chronogram, and estimate diversification rates. Analyses revealed two monophyletic eastern Pacific + Atlantic (EPA) clades, each of which likely split from an Indo-West Pacific (IWP) sister clade prior to an early Miocene Tethys Seaway closure. More recent diversification throughout the IWP appears to have been driven by both vicariance and dispersal events; EPA diversity has been further shaped by speciation across the Central American Seaway prior to its closure and dispersal across the Atlantic. Despite the latter, inter-regional dispersal has been rare, and likely contributes little to regional diversity patterns. Similarly, infrequent transitions into temperate regions combined with reduced diversification rates may explain low diversity in West and South Pacific clades. Since origination, Nerita diversification appears remarkably constant, with the exception of a lag in the late Eocene–early Oligocene and elevated rates in the late Oligocene–early Miocene. However, a comparison among regions suggested that IWP clades have experienced, on average, higher rates of speciation. Fossil evidence indicates that the EPA likely witnessed greater extinction relative to the IWP. We propose that regional differences in species diversity in Nerita have been largely shaped by differential rates of speciation and extinction.

Nerita albicilla [spec. nov.] N. testa striata, labiis subdentatis: interiore tuberculato. Rumph. mus. t. 22. f. 8. Habitat ad Hitoe., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 778, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita albume [spec. nov.] N. testa umbilicata convexa, umbilico subcordato: lobo explanato. List. conch. 4. s. 5. c. 3. f. 1, 2. Rumph. mus. t. 22. f. B. Vitellus compressus. Gvalt. test. t. 67. f. A, B. Argenv. conch. t. 10. f. T. Kratzenst. Regenf. 20. t. 5. f. 54. Habitat in O. Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 776, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita vitellus [spec. nov.] N. testa umbilicata subglobosa, umbilico perforato aequali. Rumph. mus. t. 22. f. D. Valvata 5. Habitat in O. Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 776, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita lacustris [spec. nov.] N. testa laeviuscula cornea, apice exquisito, labiis edentulis. Habitat in Europae lacubus. Testa similis N. littorali, sed cornea, nec vertice carioso. * * * Imperforatae labio dentato., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 778, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita plicata [spec. nov.] N. testa sulcata, labiis dentatis: interiore rotundato; exteriore utrinque dentibus acutis conicis. M. L. U. Habitat.., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 779, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita polita [spec. nov.] N. testa laevi, labiis utrisque dentatis. Rumph. mus. t. 22. f. I, 2. Valvata alpina. Gvalt. test. t. 66. f. C. Argenv. conch. t. 10. f. K. Kratzenst. Regenf. 16. t. 4. f. 43. Habitat in O. Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 778, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita fluviatilis [spec. nov.] N. testa rugosa, labiis edentulis. Fn. svec. 1318. List. angl. 136. t. 2. f. 20. conch. 2. p. 1. t. 38. Pet. mus. 67. n. 718. Habitat in Europae cataractis., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 777, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita histrio [spec. nov.] N. testa sulcata transverseque striata, labio interiore dentato. Rumph. mus. t. 22. f. 6. Habitat.., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 778, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita grossa [spec. nov.] N. testa sulcata, labiis dentatis: interiore convexo rugoso. Rumph. mus. t. 22. f. N. Gvalt. test. t. 66. f. V. Habitat in O. Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 779, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita bidens [spec. nov.] N. testa laevi, labio interiore bidentato. M. L. U. Habitat.., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 778, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita radula [spec. nov.] N. testa sulcata tuberculis aequalibus, labiis edentulis. M. L. U. Habitat.., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 777, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita mammilla [spec. nov.] N. testa umbilicata ovata glabra, umbilico obtecto, apertura ovata. Column. aqv. t. 52. f. ult. Cochlea nivea. List. conch. 4. s. 5. c. 3. t. 1. f. 3, 4. Rumph. mus. t. 22. f. E. Valvata 7: Albula. Gvalt. test. t. 67. f. C, D. Argenv. conch. t. 10. f. X. Uber tenue. Habitat ad Barbados. * * Imperforatae labio edentulo., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on pages 776-777, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita nodosa [spec. nov.] N. testa nodosa pluribus ordinibus solida, labro angulato. Bonan. recr. 3. t. 173, 174. List. conch. t. 804. f. 13. Gvalt. test. t. 66. f. BB. Klein. ostr. 17. t. 1. f. 30. Habitat.. Affinis Murici Nodo tota structura, sed postice lacuna destituta, qua neglecta ruerent genera., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 777, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita exuvia [spec. nov.] N. testa sulcata, labiis dentatis: interiore tuberculato. Bonan. recr. 3. t. 220. List. conch. 4. s. 1. c. 1. t. 2. f. 5. Rumph. mus. t. 22. f. M. Pet. gaz t. 100. f. 6. Gvalt. test. t. 66. f. CC. Argenv. conch. t. 10. f. I. Habitat in O. Americae & Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 779, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita glaucina [spec. nov.] N. testa umbilicata laevi, spira obtusiuscula, umbilico semiclauso: lobo gibbo dicolore. List. conch. 4. s. 5. c. 1. t. 5. f. 1. Rumph. mus. t. 22. f. A. Valvata laevis. Gvalt. test. t. 67. f. M, P, T. Habitat in O. Europaeo., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 776, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita canrena [spec. nov.] N. testa umbilicata laevi, spira submucronata, umbilico gibbo bifido. Rumph. mus. t. 22. f. C. Valvata 4. Gvalt. test. t. 67. f. E, Q, R, S, V, X. Argenv. conch t. 10. f. C. Kratzenst. Regenf. 10. t. 3. f. 34. Habitat in O. Asiae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 776, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita [gen. nov.] Animal Limax. Testa univalvis, spiralis, gibba, subtus planiuscula. Apertura semiorbicularis: labio columellae transverso, truncato, planiusculo. * Umbilicatae., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 776, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206

Nerita undata [spec. nov.] N. testa sulcata sulcis triginta, labiis dentatis: interiore rugoso tuberculato. Rumph. mus. t. 22. f. 4. Gvalt. test. t. 66. f. X. Habitat in O. Asiae ad Amboinam. Differt a praecedenti etiam apice magis prominente, & fasciis latiusculis, nec tenuibus striis., Published as part of Linnaeus, Carolus, 1758, Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, Stockholm :Laurentius Salvius on page 779, DOI: 10.5962/bhl.title.542, http://zenodo.org/record/3922206