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27. Scieroptera sanaoensis Schmidt, 1924 Scieroptera sanaoensis: Lee 2010: 26; Lee 2015: 554. Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Schmidt E. 1924. Neue Zikaden-Gattungen und Arten. Entomologische Mitteilungen. 13: 285 - 297.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Genus Oncotympana Stål, 1870 Remarks Currently, Oncotympana includes nine species, all from the Philippines: O. pallidiventris Stål, 1870 (Samar and Mindanao), O. grandis Lee, 2010 (Luzon), O. viridicincta Stål, 1870 (Philippines) [= O. consobrina (Distant, 1906)], O. nigristigma (Walker, 1850) (Philippines), O. capitulum Lee and Mohagan, 2021 (Negros), O. simonae Lee, 2010 (Luzon), O. brevis Lee, 2010 (Luzon), O. undata Lee, 2010 (Luzon) and O. averta Lee, 2011 (Mindanao). Lee and Mohagan (2021) provided a key to the nine species. With the new species described below, Oncotympana now has 10 species., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 211, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ, Mohagan A. 2021. A new species of Oncotympana Stal, 1870 (Hemiptera: Cicadidae: Oncotympanini) from Negros, Philippines. J Asia Pac Biodivers. 14 (4): 651 - 653. doi: 10.1016 / j. japb. 2021.09.002.","Lee YJ. 2011. New genus and two new species of Oncotympanina stat. nov. (Hemiptera: Cicadidae: Cicadini) and the erection of Sonatini new tribe. J Asia Pac Entomol. 14 (2): 167 - 171. doi: 10.1016 / j. aspen. 2010.12.001."]}

24. Mogannia tenebrosa Lee and Marshall sp. nov. (Figures 15, 16) Type material Holotype. Male, specimen code 12.PH. MN.CAB.01, PHILIPPINES, Mindanao, Cateel / Compostela Rd., nr bdry, 646 m, 07°40.39 ʹ N, 126°12.77 ʹ E, 6 May 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (NMPM). Paratypes. 1 male, same location as holotype, genitalia dissection G103 (UCONN); 1 male, specimen code 12. PH. MN.HWM.01, PHILIPPINES, Mindanao, W side Mt. Hamiguitan, Ca. 1.5 h up trail from Tumalite, 282 m, 06°43.95 ʹ N, 126°08.13 ʹ E, 26 April 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (UCONN). Etymology The specific name is a Latin feminine adjective meaning ̍dark̾, in reference to the darkcoloured body and the base of the fore wings of this species. Measurements of types (2 males, holotype (larger) and paratype from Mt. Hamiguitan) Length of body: 12.5 (11.2–13.7). Length of head and thorax together: 6.3 (5.4–7.1). Length of abdomen: 6.2 (5.8–6.6). Width of head including compound eyes: 3.2 (3.1–3.2). Width of pronotum: 4.6 (4.1–5.0). Width of mesonotum: 4.0 (3.6–4.4). Width of abdominal tergite 3: 5.3 (4.6–5.9). Length of fore wing: 15.4 (14.0–16.8). Width of fore wing: 5.2 (4.8–5.5). Wing span: 33.4 (30.8–36.0). Description of male Head black to fuscous; narrow, much narrower than mesonotum. Distance between lateral ocelli and compound eyes slightly shorter than distance between lateral ocelli. Antenna mostly dull ochraceous but with black pedicel and scape. Postclypeus greatly protruding anteriad. Rostrum with small dull ochraceous patch proximally, with apex slightly passing beyond posterior margin of mid coxae. Thorax including cruciform elevation black to fuscous. Anterolateral pronotal collar not dentate. Fore-, mid- and hind coxae each with light green distal membrane. Wings: Fore wing partly hyaline with venation ochraceous except for basal half of costal vein pinkish brown and with broad dark brown patch on basal half of fore wing. Basal cell tinged with pinkish brown. Basal membrane pinkish brown. Hind wing hyaline. Hind wing costal cell, radial cell and medial cell with small pinkish brown patch basally. Vannus with pinkish brown patch basally on about basal one-fourth. Hind wing jugum pinkish brown. Operculum black to fuscous; hook- or meniscus-shaped, with length of 2.5–3 times the width, with apex heading towards centre of body; not reaching posterior margin of abdominal sternite II. Opercula nearly touching each other medially. Abdomen wide and short, about as long as head and thorax together; mostly black with small reddish brown patches, especially on median and sublateral parts. Tergite 3 much wider than pronotum. Timbal cover black but tinged with reddish brown, very small, triangular with anterior angle rounded, wider than long. Timbal mostly exposed. Abdominal sternites dark brown to reddish brown with black caudal margin on each of sternites II–VII. Sternite VIII and epipleurite VIII nearly red. Genitalia. Pygofer circular in ventral view. Dorsal beak narrow and long. Uncus short with widely concave apex. Median clasper process extending posteriad in ventral view, but curved inward in lateral view. Lateral clasper lobe short. Aedeagus with spine-like processes apically. Distal shoulder with short triangular tooth in lateral view. Upper lobe uncurved, with rounded apex in lateral view. Basal lobe indistinct. Remarks This species resembles Mogannia conica (Germar, 1830) (TL: Java) because its fore wing also has a broad dark-coloured patch halfway from the base, but M. conica has the reddish postclypeus (vs black in the new species),narrower patch on the fore wing, smoothly rounded distal shoulder of the male pygofer (vs with a short triangular tooth in the new species), and a lobe-like process under the lateral clasper lobe (vs absent in the new species). Also similar is Mogannia funettae Boulard, 2008 from northern Thailand, but like M. conica, M. funettae has the fore wing basal cell clear or tinged with bright colour rather than infuscated with dark brown to match the remainder of the basal fore wing (see Boulard 2013). As discussed below, the song of Mogannia tenebrosa Lee and Marshall sp. nov. also suggests a relationship to M. funettae. This species is similar also to Mogannia formosana Matsumura,1907 because of the blackish body colour and the brownish patch on the fore wing, but M. formosana has a much more brightly coloured patch on the fore wing and light colouration on the ventral side of the body. Two other Mogannia previously recorded from the Philippines are Mogannia indigotea Distant, 1917 (TL: Tonkin) and Mogannia viridis (Signoret, 1847) (TL: Java) although we suspect that they are not distributed in the Philippines. Both differ from M. tenebrosa in the main body colour (M. indigotea is dark indigo blue, and M. viridis green to ochraceous). While the dark body colour of M. indigotea could be confused in poorly preserved specimens, the fore wing colour pattern is different, with some sections of the fore wing hyaline or tinted in indigotea but fully infuscated in tenebrosa (see Chen et al. 2012). Song (Figure 16) The song of the holotype male was recorded and is illustrated here. The sound is a highpitched staccato buzz that continues indefinitely (at least 62s in our recording). Syllables containing approximately six pulses each are closely grouped or coalesced into echemes comprising four syllables, with the beginning and ending pulses of each syllable sometimes overlapping. The syllables are produced at about 200/s, and the echemes at about 20/s. There is minor amplitude modulation that helps to define the syllables,but otherwise sound intensity and frequency are not regularly modulated.The frequency spectrum is broad,reaching from at least 8 kHz to above 20 kHz, but most energy is concentrated within the range 8–12 kHz. The dominant frequency is about 10 kHz.Males were observed to sing while perching on the upper sides of leaves, as noted by Boulard (2013) for other Mogannia in Thailand. No songs have been published previously for the genus Mogannia in the Philippines. The song of M. funettae, as shown in Boulard (2013), is very similar to that of M. tenebrosa, differing mainly in having a somewhat faster echeme repetition rate of about 28/s and inter-echeme intervals that are about the same duration as the echemes themselves., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 228-232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Boulard M. 2013. The cicadas of Thailand, volume 2: taxonomy and sonic ethology. Manchester (United Kingdom): Siri Scientific Press; p. 436.","Signoret MV. 1847. Description de deux Hemipteres Homopteres, tribu des Octicelles groupe des cicadides. Annales de la Societe entomologique de France. 1847: 293 - 296.","Chen X, Yang M, Wei C 2012. Review of the cicada genus Mogannia Amyot & Serville from China, with descriptions of three new species (Hemiptera: Cicadidae). Zootaxa. 3568 (1): 1 - 35."]}

Genus Neopurana Lee and Marshall gen. nov. Type species Neopurana bouptera Lee and Marshall sp. nov., monotypic. Etymology The generic name is feminine, a combination of the Greek prefix neo- meaning ̍new̾ and the name of an allied genus, Purana. Description Body small-sized (24.4 mm long in holotype of the type species). Head including compound eyes (6.8 mm wide in holotype) about as wide as mesonotum (6.7 mm wide in holotype). Postclypeus moderately swollen. Rostrum slightly passing beyond hind coxae. Anterolateral pronotal collar not dentate. Wings hyaline. Fore wing with infuscations on crossveins and along its hind submargin. Fore wing with very long apical cells, with ulnar cell 2 about as long as apical cell 4. Male operculum small, about as long as wide, extending to anterior margin of abdominal sternite III. Opercula widely separated from each other. Male abdomen long obconical, distinctly longer than head and thorax together. Male abdominal tergite 3 wider than mesonotum. Timbal cover about as long as wide, concealing timbal in dorsal view. Male abdominal sternite III with tubercle-like projection on each posterolateral surface, long protruding posteriorly. Male pygofer oval in ventral view. Uncus simple, short, not bifurcate. Distal shoulder not pointed. Basal lobe of pygofer absent. Remarks Neopurana Lee and Marshall gen. nov. resembles various species and species groups of the genus Purana Distant, 1905 in the following characters, and especially in the similarlooking marks and patterns on the mesonotum: body small-sized; wings hyaline; male operculum small; and male abdominal sternite III with tubercle-like projection on each lateral surface. However, this genus can be distinguished from Purana by the following characteristics: anterolateral pronotal collar not dentate; fore wing with very long apical cells; male abdominal sternite IV without tubercle-like projections; and aedeagus thick apically. This genus can be distinguished from all other genera of Puranina Lee, 2013 by the fore wing with very long apical cells and the aedeagus very thick apically (as illustrated in Figure 11 (C))., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 217-218, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ, Emery D. 2013. New genera and species of Leptopsaltriini (Hemiptera: Cicadidae: Cicadinae) from India and Tibet, with the descriptions of five new subtribes. Zoosystema. 35 (4): 525 - 535. doi: 10.5252 / z 2013 n 4 a 6."]}

Subtribe EUTERPNOSIINA Lee, 2013 Euterpnosiina Lee, 2013 in Lee and Emery, 2013: 527–528. Remarks The position of the new genus described immediately below is unique and independent from any other known species of Leptopsaltriini in the phylogenetic tree constructed by Hill et al. (2021). Therefore, this genus could form a separate new subtribe within Leptopsaltriini. This genus has very long fore wing apical cells, which is one of its peculiar characteristics within Leptopsaltriini, and has tubercle-like projections only on the sternite III, which is seen only in some members of Tanna Distant, 1905 and Vietanna Lee and Pham, 2021. This genus is different from the genera of Puranina Lee, 2013 because of the anterolateral pronotal collar which is not dentate, the male abdominal sternite IV which does not have tubercle-like projections, and the aedeagus which is thick apically. This genus is tentatively placed in Euterpnosiina because the type specimen of the genus was largely grouped with genera of Euterpnosiina in the phylogenetic tree constructed by Hill et al. (2021) in spite of many morphological differences, including the following: timbal cover well developed (vs minute in Euterpnosiina); male abdominal sternite III with tubercle-like projections (vs mostly without them in Euterpnosiina); aedeagus thick (vs thin in Euterpnosiina); and basal lobe of pygofer absent (vs present in Euterpnosiina). Further taxonomic study is required to find its proper subtribe placement.

Tribe LEPTOPSALTRIINI Moulton, 1923 Remarks Hill et al. (2021) noted molecular phylogenetic evidence of confusion between the tribes Gaeanini Distant, 1905 and Leptopsaltriini Moulton, 1923 and suggested that Gaeanini could perhaps be redefined as a subtribe within Leptopsaltriini. However, in such a case tribe Gaeanini would take priority since it is the older family group. In this paper, we refrain from synonymising Leptopsaltriini with Gaeanini, awaiting a further comprehensive morphological review., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 216, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079."]}

17. Neoncotympana leeseungmoi Lee 2011 Neoncotympana leeseungmoi Lee 2011: 169–171, fig. 2 [TL: Mt. Kitanlad Intavas, Mindanao]; Lee 2015: 551. Additional material examined 2 males, specimen code 12. PH. MN.KGS.05 and 12. PH. MN.KGS. 06, PHILIPPINES, Mindanao, Mt. Kitanglad, 1773 m, 08°09.91 ʹ N, 124°55.96 ʹ E, 4 May 2012, K.B. R. Hill, D.C. Marshall, D. Mohagan and T. Catanach (UCONN); 1 female, PHILIPPINES, Mindanao, Cotabato, Mt. Apo, Agro-ecosystem (Sudsuwayan), 25 May 2010, D. Mohagan (UCONN). Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 216, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2011. New genus and two new species of Oncotympanina stat. nov. (Hemiptera: Cicadidae: Cicadini) and the erection of Sonatini new tribe. J Asia Pac Entomol. 14 (2): 167 - 171. doi: 10.1016 / j. aspen. 2010.12.001.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

6. Chremistica minor Bregman, 1985 Chremistica minor: Lee 2010: 16; Lee 2015: 546. Distribution Malaysia (peninsular Malaysia and Sarawak), Indonesia (Kalimantan) and Philippines? (Mindanao?). Remarks Bregman (1985, p. 48) stated that ̍ Two female specimens from Mindanao, Philippines showed great resemblance to the male types of C . minor. It is not quite certain whether these females indeed belong to C . minor ̾. Chremistica minor has not been found in Mindanao or any other part of the Philippines by the present authors or by any other authors after Bregman (1985). We suspect that C . minor (TL: Kalimantan, Indonesia) is not distributed in Mindanao. As the body colouration of C . minor somewhat resembles that of Chremistica kyoungheeae, it is possible that the two Mindanao females noted by Bregman (1985) are specimens of C. kyoungheeae., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 206, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Bregman R. 1985. Taxonomy, phylogeny and biogeography of the tridentigera group of the genus Chremistica Stal, 1870 (Homoptera, Cicadidae). Beaufortia. 35: 37 - 60.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

29. Philipsalta exilis Lee and Marshall sp. nov. (Figures 17, 18) Type material Holotype. Male, specimen code 12.PH. MN. SSM.02, PHILIPPINES, Mindanao, Davao City, Marilog District, Datu Salumay, Mt. Malambo, 7°29.28 ʹ N, 125°15.18 ʹ E, 18 April 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (NMPM). Paratype. 1 male, specimen code 12. PH. MN. SSM.01, DNA voucher, same collection data. Etymology The specific name is a Latin adjective meaning ̍slender̾, ̍thin̾, ̍feeble̾ or ̍small̾, in reference to the small and slender body of this species. Measurements of holotype (1 male) Length of body: 14.2. Length of head and thorax together: 5.5. Length of abdomen:8.7. Width of head including compound eyes: 3.6. Width of pronotum: 4.2. Width of mesonotum: 3.5. Width of abdominal tergite 3: 3.9. Length of fore wing: 16.8. Width of fore wing: 6.3. Wing span: 37.0. Description of male Body tiny and slender. Head: Vertex ochraceous with the following fuscous marks: inverted triangular mark surrounding median ocellus; a pair of rather indistinct marks surrounding lateral ocelli; a pair of spots on inner corners of supra-antennal plates; and a pair of spots on posterolateral corner of vertex (touching compound eyes). Distance between lateral ocelli and compound eyes about as long as distance between lateral ocelli. Antennae black except for reddish-brown scape and base of pedicel. Postclypeus not swollen anteriad and rather flat ventrally; reddish brown to ochraceous with V-shaped black median mark, of which posterior part merged into a large patch. Anteclypeus reddish ochraceous with median fuscous spot and fuscous posterior margin. Rostrum brown with black apex; with apex extending slightly beyond posterior margin of mid trochanter. Lorum ochraceous with large fuscous spot posteriorly. Gena ochraceous without distinct marks. Thorax Pronotum ochraceous. Inner area of pronotum with the following fuscous marks: tiny spot on posteromedian margin of inner area;a pair of short longitudinal fasciae between about middle of paramedian fissures and posterior ends of lateral fissures; a pair of fasciae along lateral fissures; irregular indistinct marks between paramedian fissures and lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar very narrow; without marks. Anterolateral pronotal collar not dentate. Mesonotum dull ochraceous with a pair of fuscous patches on entire submedian sigilla and a pair of longitudinal larger patches on almost entire lateral sigilla. Cruciform elevation light ochraceous or yellow without distinct marks. Thoracic sternites greenish light ochraceous. Legs ochraceous to reddish brown with some fuscous spots. Fore femur with primary, secondary and subapical spines. Wings hyaline without infuscation; with extremely narrow marginal areas. Fore wing venation reddish brown basally but fuscous distally. Basal membrane yellowish grey. Hind wing with five or six apical cells (possibly with six apical cells normally, considering other Philipsalta species have six apical cells). Operculum small, semicircular, not reaching posterior margin of sternite II; greenish light ochraceous. Opercula separated from each other. Abdomen long triangular in dorsal view, much longer than head and thorax together, with ridge formed on tergites 4–7 dorsomedially; slightly greenish light ochraceous with grey-tinged lateral margins. Tergite 1 very short, less than one-fourth as long as tergite 2 in median length. Tergites 3–8 black to fuscous on about anterior half and brown to ochraceous on about posterior half dorsomedially; upper lateral surfaces slightly translucent. A pair of fuscous globose bumps present on lateral margins of tergite 2. Timbal cover absent. Timbals not extending below wing bases ventrally. Abdominal sternites translucent, ochraceous indistinctly with pinkish caudal margin on each of sternites 3–6. Genitalia Pygofer ovate with triangular dorsal beak in ventral view. Upper lobe moderately pointed. Dorsal beak triangular. Median lobe of uncus moderately protruding. A pair of clasper-like processes protruding straight downward. Remarks This new species can be distinguished from Philipsalta nigrina by body colouration and by the comparative size and shape of each part of the body and wings as follows: body colouration not monotonous(vs entirely black in P.nigrina);head comparatively short,about half as long as pronotum (vs about two-thirds as long as pronotum in P. nigrina); fore wing radial cell comparatively narrow, narrower than ulnar cell 3 (vs spacious, broader than ulnar cell 3 in P. nigrina); and male abdomen much longer than head and thorax together (vs slightly longer in P. nigrina). Song (Figure 18) About 190 s of song was recorded from this species at the type locality, including that of the paratype male, which is illustrated here. Males produce song phrases of about 0.3– 0.85 s duration at a regular pace of about 0.5 phrases/s. Each phrase begins with about 8–20 doublet pulses produced at 40 doublets/s, with the first pulse of each doublet louder. The doublets increase in intensity towards the end of this long syllable. After a gap equivalent to the time needed for one additional doublet, the phrase concludes with a syllable of 6– 13 doublets produced at a faster rate (60/s) and with a greater intensity. In our recording, it is difficult to discern the quieter pulse of each doublet in the second syllable. There is a tendency, across phrases, for pulse doublets to drop out occasionally within both syllables. The recording was taken with the recordist close to the cicada and attempting to attract it with finger-snaps, so disturbance could have occurred. The frequency spectrum is broad, reaching from at least 8 kHz to above 20 kHz, but most energy is concentrated within the range 8–12 kHz. The dominant frequency is about 10 kHz. Males sang from perches on vertical surfaces (e.g. ferns, tree trunks, small branches) around 2–4 m aboveground in the forest interior. The song of Philipsalta exilis Lee and Marshall sp. nov. can be distinguished from that of the next new species of Philipsalta (below) by the absence of a short single-doublet element ending many of the phrases and by the uniform rate of production of pulse doublets within the first syllable/echeme. Philipsalta exilis and P. nigrina have a similar song structure, but in the latter species the pulses in the first section of each phrase form widely separated four-pulse syllables, the intervening gaps being much longer than the syllables., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 233-236, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268

Tribe CICADATRINI Distant, 1905 Cicadatraria Distant 1905a: 304–305., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 228, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Distant WL. 1905 a. Rhynchotal notes. - XXX. Ann Mag Nat Hist. (7) 15 (): 304 - 319. doi: 10.1080 / 03745480509443047."]}

21. Purana guttularis (Walker, 1858) Purana guttularis: Lee 2010: 22; Lee 2015: 553. Distribution S. China, Philippines (including Mindanao) (not previously reported from specific islands), Indonesia (Borneo, Java, Sumatra and Nias), Malaysia, Thailand, Myanmar and India. Remarks As Lee (2010, 2015) indicated, the identity of this species in the Philippines should be reconsidered. The female type of P. guttularis is hardly distinguishable from the female paratype of P. nana as shown in Lee (2009a). We suspect that the record of P. guttularis from Mindanao is due to misidentification. Purana guttularis may have to be removed from the cicada list of Mindanao., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 223, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5.","Lee YJ. 2009 a. Descriptions of two new species of the Purana abdominalis species group (Hemiptera: Cicadidae: Cicadini) from the Philippines, with a key to the species groups of Purana. J Nat Hist. 43 (25 - 26): 1487 - 1504. doi: 10.1080 / 00222930902941848."]}

8. Cryptotympana consanguinea Distant 1916 Cryptotympana consanguinea: Lee 2010: 19; Lee 2015: 546. Distribution Philippines (Dinagat, Mindanao and Basilan)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 210, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Distant WL. 1916. On the Cicadidae found in the Philippine Islands. Entomologist. 49: 203 - 204.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Tribe NELCYNDANINI Moulds and Marshall, 2018 Nelcyndanini Moulds and Marshall, 2018 in Marshall et al., 2018b: 26., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Marshall DC, Moulds M, Hill KBR, Price BW, Wade E, Owen CL, Goemans G, Marathe K, Sarkar V, Cooley JR, et al. 2018 b. A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa. 4424 (1): 1 - 64. doi: 10.11646 / zootaxa. 4424.1. 1."]}

19. Purana nana Lee, 2009 Purana nana: Lee 2010: 22; Lee 2015: 551. Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 222, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Genus Mogannia Amyot and Audinet-Serville,1843 Mogannia Amyot and Audinet-Serville 1843: 467. Type species Mogannia illustrata Amyot and Audinet-Serville 1843 (Java) [junior synonym of Mogannia conica (Germar, 1830)]. Cephaloxys Signoret, 1847: 294. Type species Cicada conica Germar, 1830 (Java)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 228, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Amyot CJB, Audinet-Serville JG. 1843. Homopteres. Homoptera Latr. In: Histoire naturelle des insectes, hemipteres. Paris (France): Librairie Encyclopedique de Roret. pp. 455 - 588; p. 676. Banks CS. 1910. Rhynchota Palawanica, Part II. Homoptera. Philipp J Sci. 5: 33 - 55.","Signoret MV. 1847. Description de deux Hemipteres Homopteres, tribu des Octicelles groupe des cicadides. Annales de la Societe entomologique de France. 1847: 293 - 296."]}

7. Chremistica flavialata Lee and Marshall sp. nov. (Figures 7, 8) Type material Holotype. Male, specimen code 12.PH. MN. MUS.15, PHILIPPINES, Mindanao, Bukidnon, Maramag, Mt. Musuan, 7°52.62 ʹ N, 125°4.19 ʹ E, 5 May 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (NMPM). Paratypes. 1 male, PHILIPPINES, Mindanao, Bukidnon, Maramag, Bayabason, P 4B, 8 May 2012, R. Estorco and A.B. Mohagan (UCONN); 3 males, PHILIPPINES, Mindanao, Malaybalay City, San Martin, 24 August 2010, A.B. Mohagan (UCONN). Etymology The specific name is the Latin prefix flavi -, meaning ̍yellowish̾, plus the feminine form of the Latin adjective alata, meaning ̍winged̾, in reference to the wings of this species, which are deeply tinged with yellowish brown. Measurements of types (5 males) Length of body: 33.8 (31.9–37.1). Length of head and thorax together: 15.9 (15.0–16.9). Length of abdomen: 17.9 (16.4–20.2). Width of head including compound eyes: 12.8 (12.4–13.5). Width of pronotum: 13.3 (12.5–14.1). Width of mesonotum: 11.2 (10.7–11.7). Width of abdominal tergite 3: 13.2 (12.7–13.9). Length of fore wing: 42.0 (39.3–45.6). Width of fore wing: 13.7 (12.9–14.7). Wing span: 94.3 (88.2–102.3). Description of male Head. Eyes slightly narrower than pronotum (including pronotal collar).Vertex black with the following reddish-brown marks, sometimes interrupted or incomplete: a pair of T-shaped marks between ocelli and compound eyes; and a pair of inverted L-shaped marks between postclypeus and compound eyes. Distance between lateral ocelli and compound eyes distinctly longer than distance between lateral ocelli. Antenna black to fuscous. Postclypeus moderately swollen.Postclypeus black in about anterior one-fourth but reddish brown (or dull ochraceous in discoloured specimens) in remaining part with black transverse fasciae along transverse grooves. Anteclypeus reddish brown with a pair of large elliptical black marks occupying most of the anteclypeus except for median longitudinal part and anterior and posterior margins. Rostrum reddish brown to ochraceous but black apically and with longitudinal median fascia on labium; nearly reaching posterior margin of hind coxae. Lorum dull brown to ochraceous with a large black patch on inner corner. Gena reddish brown to ochraceous with broad black patch on anterior margin. Thorax. Inner area of pronotum black to fuscous with a pair of narrow transverse ochraceous fascia on anterior margin and median longitudinal thick ochraceous fascia, and irregularly spread dull purplish patches. Pronotal collar dull light green (or ochraceous in discoloured specimens) with dull ochraceous antero-lateral part and a pair of branch-like black marks derived from lateral corner of inner area. Anterolateral pronotal collar slightly developed and roundly angulated. Mesonotum black with the following dull greenish ochraceous (or reddish ochraceous in discoloured specimens) marks: median W-shaped mark along outside of submedian sigilla, extending to anterior arms of cruciform elevation; and a pair of longitudinal marks along lateral margin of mesonotum, connected to the median W-shaped mark posteriorly. Cruciform elevation reddish brown (dull brown in discoloured specimens) medially but dull light green (ochraceous in discoloured specimens) laterally with the following black marks: antero-median spot; narrow fascia along posterior margin; and a pair of spots on middle of anterior arms. Thoracic sternites dull brown to dull ochraceous with some black trivial marks. Legs reddish brown to ochraceous. Fore femur with a small subapical spine as well as primary and secondary spines; with black to fuscous fascia along midline of ventral side. Fore tibia black to fuscous. Fore- and mid tarsi mostly black to fuscous. Fore-, mid- and hind pretarsal claws fuscous apically. Wings hyaline, tinged slightly with greenish yellow, especially basally. Fore wing venation dull greenish ochraceous (ochraceous in discoloured specimens) basally and brownish to fuscous apically, without infuscation. Basal cell more tinged with green. Basal membrane grey (tinged with green in fresh specimens). Hind wing jugum grey. Operculum reddish brown with black antero-lateral margin; much longer than wide, with slightly oblique and convex lateral margin and rounded apical margin; nearly reaching or slightly passing beyond posterior margin of abdominal sternite II. Opercula meeting medially. Abdomen slightly longer than head and thorax together; black with a pair of narrow white pubescent fasciae along anterior margin of tergite 3. Abdominal tergite 3 wider than head including compound eyes. Timbal cover tinged with purple, wider than long. Abdominal sternites mostly reddish brown, but sternite II ochraceous to reddish ochraceous or greenish ochraceous with fuscous mark. Genitalia. Pygofer diamond-shaped in ventral view with angles rounded. Uncus shaped like flower shovel, with roundly pointed apex and without lateral protrusions. A pair of claspers originating from lateroventral uncus hidden behind uncus and lateral lobes. Aedeagus without subapical or ventral processes. Basal lobe of pygofer long, with roundish apex and with slender finger- or thumb-like protrusion branching out at subapical inner part of the basal lobe. Lateral lobe present, much longer than basal lobe and obliquely covering basal lobe. Dorsal beak narrowly triangular. Remarks This new species is similar to Chremistica tagalica (Stål, 1870) (TL: Philippine Islands) in marks and colours on the body but can be distinguished from C. tagalica mainly by the following characteristics: head comparatively narrow (vs wider than width of pronotal collar in C. tagalica); pronotal collar with short and wide triangular black mark on medioposterior margin (vs without such a mark in C. tagalica); male abdomen comparatively wide or thick, with tergite 3 wider than head including compound eyes (vs comparatively narrow or slender, with tergite 3 narrower than head including compound eyes in C. tagalica); transverse white marks on abdominal tergite 3 narrow (vs broader, covering whole tergite 3 laterally, in C. tagalica); uncus without bumps on underside (vs with bumps in C. tagalica); and, finally and most importantly, basal lobe of pygofer with slender finger- or thumb-like protrusion branching out at subapical inner part of the basal lobe (vs with very short protrusion at subapical inner part of the basal lobe in C. tagalica). Song (Figure 8) The holotype male was recorded for about 102 s at 2.58pm on 5 May 2012.A further recording of 103 s of anonymous males from the same location on 7 July 2010 was also examined. Song is produced in phrases of 9.5– 11.5 s duration, separated by slightly shorter gaps. A song phrase contains a complex range of 12–13 major frequency bands appearing at intervals of about 0.65 kHz from 1.8–8.9 kHz,with the middle frequencies being the loudest (much weaker bands extend beyond this range). The acoustic character of the song is dominated by a synchronised sharp frequency and amplitude modulation of about 8–9 cycles/s at the beginning of the phrase, accelerating to 12/s during the middle section, which is also louder, and then slowing again. The spectrum of frequencies also shifts slightly downward in concert by about 100–200 Hz during the central section of the phrase, and then rebounds. Sound pulses produced at about 588–606/s form the most basic structure of the song., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 206-209, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268

23. Maua philippinensis Schmidt, 1924 Maua philippinensis: Lee 2010: 22, fig. 5; Lee 2015: 553. Distribution Philippines (Mindanao). Additional material examined 1 male, specimen code 12. PH.MN.HWF. 01, PHILIPPINES, Mindanao, W side of Mt. Hamiguitan, 648 m, 6°43.358 ʹ N, 126°9.211 ʹ E, 25 April 2012, K.B.R. Hill, D.C. Marshall and A.B. Mohagan (UCONN)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 228, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Schmidt E. 1924. Neue Zikaden-Gattungen und Arten. Entomologische Mitteilungen. 13: 285 - 297.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

1. Platypleura bella Lee and A. Mohagan sp. nov. (Figures 2, 3, 6 (B)) Type material Holotype. Male, PHILIPPINES, Mindanao, Bukidnon, Mt. Kiamo, 7 May 2012, A.B. Mohagan (NMPM). Paratypes. 1 male, 1 female, same data as holotype, A.B. Mohagan (UCONN); 2 males, PHILIPPINES, Mindanao, Mt. Kitanglad trailhead, 1689 m, 08°10.22 ʹ N, 124°56.14 ʹ E, 4 May 2012, K.B. R. Hill, D.C. Marshall and D. Mohagan (UCONN); 1 female, same location, 1 May 2012, K.B. R. Hill, D.C. Marshall and D. Mohagan (UCONN). Etymology The specific name is the Latin feminine adjective bella meaning ̍pretty̾ or ̍handsome̾. Measurements of types (4 males, 2 females) Length of body: males 20.6 (19.6–21.2), females 19.8 (18.7–20.8). Length of head and thorax together: males 10.5 (9.9–10.7), females 10.6 (10.3–10.8). Length of abdomen: males 10.1 (9.7–10.6), females 9.2 (8.4–10.0). Width of head including compound eyes: males 8.5 (8.4–8.6), females 8.8 (8.5–9.1). Width of pronotum: males 11.5 (10.8–12.2), females 11.7 (11.2–12.1). Width of mesonotum: males 8.8 (8.7–9.1), females 8.9 (8.4–9.4). Width of abdominal tergite 3: males 9.3 (9.0–9.7), females 9.4 (9.3–9.5). Length of fore wing: males 28.4 (27.9–28.9), females 29.6 (29.2–30.0). Width of fore wing: males 10.2 (9.8–10.4), females 10.7 (10.6–10.8). Wing span: males 65.8 (65.4–66.6), females 67.5 (65.6–69.4). Diagnosis This new species is very similar to Platypleura elizabethae Lee, 2009 (Figure 6 (C)) from Palawan but has much less infuscation on the inner half of the fore wing and the hind wing and has the following differences: fore wing apical cell 6 seldom with infuscation or with tiny spot on its proximal end (vs with quite large infuscation on about its proximal sixth in P. elizabethae); fore wing medial cell with a few tiny grey spots (vs with cloudy patterns covering about half of the cell in P. elizabethae); and fore wing apical cells 5 and 6 each with multiple tiny sub-distal spots (vs with no such spots in P. elizabethae). This species can be distinguished from Platypleura transitiva Lee, 2021 (from Mindanao) (Figure 6 (D)) by the following characteristics: fore wing basal cell with black patch or infuscation on about half of its area (vs basal cell entirely black in P. transitiva); and hind wing apical cells 1–2 fuscous on about or less than half of their area (vs fuscous on far more than half of their area in P. transitiva). Platypleura bella sp. nov. is also similar to P. minima sp. nov., as explained in the Diagnosis for that taxon below. Description of male Head. Vertex ochraceous to castaneous with the following black marks: median large spot enclosing ocelli, with anterior and posterior ends reaching anterior and posterior margins of vertex, and with short transverse branch at each lateral end; a pair of small spots between lateral ocelli and compound eyes; a pair of small spots on posterolateral corners of vertex; a pair of short obliquely longitudinal fasciae between postclypeus and compound eyes; and a pair of narrow fasciae along compound eyes. Distance between lateral ocelli and compound eyes about twice the distance between lateral ocelli. Antenna mostly castaneous but pedicel and scape black to fuscous. Postclypeus not swollen anteriad; ochraceous with a medial longitudinal black fascia, wider anteriorly, a pair of large spots antero-laterally through anterior five transverse grooves, which are connected to the medial longitudinal fascia, and short transverse fasciae along lateral parts of the remaining transverse grooves. Anteclypeus black except ochraceous median ridge. Rostrum ochraceous but darkened towards apex to become black apically; with apex reaching medial part of posterior margin of abdominal sternite II. Lorum black except ochraceous anterior and lateral margins. Gena black to dark brown with dull brown spot on posterior margin. Thorax. Pronotum ochraceous to greenish ochraceous. Inner area of pronotum with the following black to fuscous marks: posteromedial inverted T mark; indistinct spot on anteromedial part of inner area; irregularly shaped marks along paramedian fissures; a pair of irregularly shaped marks along lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar darkened anteriorly and laterally on both sides. Anterolateral pronotal collar broadly developed and roundly angulated with an angle of 100–110°, expanding to a width much greater than the widest width of posterior pronotum, mesonotum and abdomen. Mesonotum reddish ochraceous with the following black marks: median longitudinal fascia, with its posterior end widened and reaching anterior margin of cruciform elevation, but its anterior end not reaching anterior margin of mesonotum; a pair of small roundish spots enclosing scutal depressions; a pair of large semicircular spots on submedian sigilla, with their anterior ends reaching anterior margin of mesonotum; and a pair of long inverted triangular marks on lateral sigilla, with their anterior ends reaching anterior margin of mesonotum. Cruciform elevation ochraceous to greenish ochraceous. Fore-, mid- and hind trochanters and femora brown to dark brown. Wings. Fore wing outer margin (not anterior margin) slightly curved outwardly (not straight or linear). Fore wing with venation ochraceous to reddish ochraceous, darker apically; furnished with white patch on about basal two-fifths to one-half (hyaline on remaining part: about apical half) and with the following infuscations: three spots on radial cell; mark on both ends of both ulnar cells 1 and 2; rather indistinct marks on basal half and tiny spot on apical end of ulnar cell 3; indistinct small marks on medial cell; mark on both apical and basal ends of apical cell 1; irregularly shaped small marks on subapical parts of apical cells 2–7, sometimes indistinct; one or two tiny dots on apical ends of apical cells 2–7; two spots on apical end of apical cell 8. Basal membrane dark grey with light green or green margin. Hind wing fuscous except outer marginal area and apical portion of apical cells 1–6. Anal cell 2 fuscous on about basal one-third to two-thirds. Anal cell 4 fuscous on about basal half to two-thirds. Anal cell 5 fuscous on about basal half to twothirds. Hind wing jugum light grey with light green margin. Operculum mostly fuscous except greenish ochraceous margin; semicircular, transversely wide; short, nearly reaching posterior margin of abdominal sternite II. Opercula overlapping medially. Abdomen obconical in dorsal view, short, about as long as or slightly shorter than head and thorax together; mostly black with narrow greenish ochraceous caudal margin on each of tergites 3–8. Timbal cover black to fuscous with greenish ochraceous inner anterior margin; semicircular, wider than long. Timbal concealed by timbal cover in dorsal and lateral views. Abdominal sternites mostly black with greenish ochraceous caudal margin on each of sternites III–VI. Genitalia. Pygofer long ellipsoidal with acuminate apex (wide dorsal beak) in ventral view. Uncus simple, not bifurcate, with apical margin rounded. Aedeagus not tapering down to apex, with short subapical process. Description of female Head, thorax and wings similar to male. Abdomen obconical in dorsal view, short, shorter than head and thorax together; mostly black with narrow greenish ochraceous to dull brown (probably due to discoloration) caudal margin on each of tergites 3–7. Abdominal sternites mostly black with greenish ochraceous caudal margin on each of sternites III–VI. Abdominal sternite VII with a pair of large paramedian patches, extending to posterior margin. Gonocoxite IX greenish ochraceous to dull brown (probably due to discoloration). Ovipositor not extending beyond dorsal beak. Song (Figure 3) About 500 s of song was recorded from uncollected males on 1–2 May 2012, at three locations on Mt. Kitanglad, including the paratype location. The sound is a continuous whining buzz with a rapid frequency modulation. Pulses are regularly produced at a rate of 434–450/s. Pulse amplitude more than doubles across oscillations that occur at a rate of 35–40/s, with the peak frequency subtly shifting within the range 8–9 kHz across these cycles. Most sound energy lies within the range 7.5–10 kHz. The distinguishing features of the song are discussed along with those of the next species below., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 196-199, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268

Subtribe EUTERPNOSIINA Lee, 2013 Euterpnosiina Lee, 2013 in Lee and Emery, 2013: 527–528. Remarks The position of the new genus described immediately below is unique and independent from any other known species of Leptopsaltriini in the phylogenetic tree constructed by Hill et al. (2021). Therefore, this genus could form a separate new subtribe within Leptopsaltriini. This genus has very long fore wing apical cells, which is one of its peculiar characteristics within Leptopsaltriini, and has tubercle-like projections only on the sternite III, which is seen only in some members of Tanna Distant, 1905 and Vietanna Lee and Pham, 2021. This genus is different from the genera of Puranina Lee, 2013 because of the anterolateral pronotal collar which is not dentate, the male abdominal sternite IV which does not have tubercle-like projections, and the aedeagus which is thick apically. This genus is tentatively placed in Euterpnosiina because the type specimen of the genus was largely grouped with genera of Euterpnosiina in the phylogenetic tree constructed by Hill et al. (2021) in spite of many morphological differences, including the following: timbal cover well developed (vs minute in Euterpnosiina); male abdominal sternite III with tubercle-like projections (vs mostly without them in Euterpnosiina); aedeagus thick (vs thin in Euterpnosiina); and basal lobe of pygofer absent (vs present in Euterpnosiina). Further taxonomic study is required to find its proper subtribe placement., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 217, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ, Emery D. 2013. New genera and species of Leptopsaltriini (Hemiptera: Cicadidae: Cicadinae) from India and Tibet, with the descriptions of five new subtribes. Zoosystema. 35 (4): 525 - 535. doi: 10.5252 / z 2013 n 4 a 6.","Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079."]}

22. Purana mindanaoensis Lee and Marshall sp. nov. (Figures 13, 14) Purana cf. barbosae: Hill et al. 2021: table 1, figs 5, 13, 14. Type material Holotype. Male, specimen code 12.PH. MN. MUS.01, PHILIPPINES, Mindanao, Bukidnon, Maramag, Mt. Musuan, 7°52.62 ʹ N, 125°4.19 ʹ E, 19 April 2012, K.B. R. Hill, D.C. Marshall and A. B. Mohagan (NMPM). Paratypes. 1 male, specimen code 12. PH. MN. MUS. 05, same collection data (UCONN); 1 male, specimen code 12. PH. MN. HWB.01, PHILIPPINES, Mindanao, W side Mt. Hamiguitan, 1 h up trail from Tumalite Village, 6°43.849 ʹ N, 126°7.856 ʹ E, 24 April 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (UCONN). Additional location audio recorded PHILIPPINES, Mindanao, Cateel / Compostela Rd., nr bdry, 646 m, 07°40.39 ʹ N, 126°12.77 ʹ E, PH. MN.CAB, 6 May 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan. Etymology The new species is named after the type locality, Mindanao. Measurements of types (3 males) Length of body: 20.8 (20.2–21.4). Length of head and thorax together: 10.1 (9.8–10.3). Length of abdomen: 10.7 (10.4–11.1). Width of head including compound eyes: 6.5 (6.2–6.6). Width of pronotum: 7.2 (7.1–7.4). Width of mesonotum: 6.3 (6.3–6.4). Width of abdominal tergite 3: 6.9 (6.8–6.9). Length of fore wing: 27.2 (24.7–28.9). Width of fore wing: 8.9 (8.5–9.3). Wing span: 60.2 (56.1–63.8). Description of male Head. Vertex dark ochraceous tinged with green and with the following black marks: inverted triangular large median mark surrounding ocelli, with its anterior tip not reaching frontoclypeal suture; a pair of longitudinal F-shaped marks on both sides of the median mark, extending to supra-antennal plates; a pair of narrow fasciae along compound eyes; and a pair of tiny spots posteromedially. Distance between lateral ocelli and compound eyes longer than twice the distance between lateral ocelli. Antennae dark brown to fuscous. Postclypeus moderately swollen anteriad; light ochraceous, tinged with green anteriorly, with fasciae or spots along transverse grooves, all connected at inner ends, of which first five long but remainder short. Anteclypeus black except ochraceous median ridge. Rostrum ochraceous but black to fuscous apically; not reaching abdominal sternite III. Lorum ochraceous on anterior one-third but black on posterior two-thirds. Gena ochraceous with obliquely transverse black fascia between postclypeus and compound eye and fascia along compound eye. Thorax. Pronotum mostly ochraceous but partly light green. Inner area of pronotum ochraceous but light green posteromedially with the following black to fuscous marks: a pair of median longitudinal fasciae slightly broadened at both anterior and posterior ends; a pair of short, oblique indistinct branches along paramedian fissures arising from around middle of the median longitudinal fasciae; a pair of indistinct short longitudinal fasciae between about middle of paramedian fissures and posterior ends of lateral fissures; a pair of fasciae along lateral fissures, interrupted in the middle; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar greenish ochraceous with narrow transverse black fascia along posterior margin and a pair of fuscous spots on sublateral parts. Anterolateral pronotal collar dentate. Mesonotum brown (medially) to dark greenish ochraceous (laterally) with the following black marks: median longitudinal fascia slightly broadened posteriorly to reach anterior margin of cruciform elevation; a pair of small round spots enclosing scutal depressions; a pair of inwardly curved fasciae along inner side of parapsidal sutures; a pair of longitudinally arranged two spots on lateral sigilla: anterior ones longitudinally long touching anterior margin of mesonotum but posterior ones roundish. Cruciform elevation greenish ochraceous but tinged with brown centrally with black posterior margin. Thoracic sternites light greenish ochraceous with black to fuscous spot each on basisternum 2, episternum 2, meron 2, basisternum 3, episternum 3 and meron 3. Legs ochraceous to greenish ochraceous. Fore femur with black primary, secondary and subapical spines and with black fascia along midline of ventral side of fore femur. Fore-, mid- and hind femora with large brown to light brown patches. Fore tibia mostly fuscous except small proximal part. Mid tibia with small distal fuscous part. Fore- and mid pretarsi black. Wings hyaline and without infuscations but slightly tinged with brown distally. Costal vein greenish ochraceous basally but darkening to become castaneous to fuscous distally. Basal cell slightly tinged with greenish ochraceous. Basal membrane and hind wing jugum dark grey. Ulnar cell 3 very broad, with border adjoining radial cell more than twice as long as that between radial cell and ulnar cell 2. Operculum inverted triangular with posterior angle rounded, and extending far beyond middle of abdominal sternite III but not reaching posterior margin of sternite III; dull light green laterally and widely black to fuscous along inner and posterior margins, and covered with short silvery hairs and white pollinosity. Anterior one-fourth of inner margin deeply concave. Opercula separated from each other, with gap about one-fourth as wide as operculum. Abdomen castaneous to light brown; long obconical, longer than head and thorax together. Posterior margins of each tergite narrowly black to fuscous, sometimes indistinct. Tergite 3 wider than mesonotum. Timbal cover greenish ochraceous with a large fuscous patch sublaterally; nearly quarter circle with rounded anterolateral corner, about as long as wide. Timbal concealed by timbal cover in dorsal view. Abdominal sternites dull ochraceous to light brown. Sternites III and IV each with a pair of black tubercle-like projections on posterolateral surfaces, protruding ventrolaterally but slightly obliquely downward. Genitalia. Pygofer barrel-shaped in ventral view. Uncus short, consisting of median lobe with blunt bicuspidate apex and a pair of lateral lobes. Distal shoulder not pointed. Basal lobe of pygofer triangularly prominent. Remarks The existing 16 species of the Purana carmente species group are very similar to each other and often hard to distinguish. This new species has two longitudinally arranged fasciae or spots on the lateral sigillum, which are always separate from each other. This characteristic is seen only in Purana mindanaoensis Lee and Marshall sp. nov. and Purana natae Boulard, 2007. Other species of the Purana carmente species group have only one long, continuous longitudinal fascia on the lateral sigillum, except for Purana barbosae (Distant, 1889), which has no such distinct fasciae or spots at all on the lateral sigillum. The male operculum of this new species has a broadly black patch along the inner margin, about half as wide as the operculum. The following eight species have a much thinner or nearly absent black patch on the male operculum: Purana barbosae, Purana obducta Schouten and Duffels, 2002, Purana sagittata, 2002, Purana infuscata, 2002, Purana mickhuanae Boulard, 2009, Purana natae, Purana phetchabuna Boulard, 2008 and Purana tanae Boulard, 2007. The following eight species have their own peculiar uncus shapes, respectively: Purana carmente (Walker, 1850), Purana hermes Schouten and Duffels, 2002, Purana infuscata, Purana sagittata, Purana chueatae Boulard, 2007, Purana mickhuanae, Purana natae and Purana tanae (see Schouten and Duffels 2002; Boulard 2013). The remaining species of the Purana carmente species group, including Purana mindanaoensis, have somewhat similar uncus shapes, but Purana mindanaoensis has largely expanded lateral uncal lobes, which are larger or more developed than those of any other species of the Purana carmente species group. Purana dimidia Chou and Lei, 1997 has a comparatively short male operculum. Purana trui Pham et al., 2012 has very thin longitudinal fascia on the timbal cover, a very broad black patch on the male operculum, and a blunt apical angle of the basal lobe of pygofer. Purana opaca Lee, 2009 has darker body pigmentation and an almost entirely black timbal cover and male operculum. Song (Figure 14) Recordings of the song of this species, totalling several minutes, were obtained at the type locality. The sound is a resonant, stuttering buzz with a bell-like quality, and it contains amplitude and frequency modulation as well as a complex pattern of fundamental pulses, syllables and echemes. The song is arranged into phrases that change in structure as a male continues to sing, progressing from shorter, usually isolated phrases, to longer ones up to ca.2 s often concatenated into sets lasting up to 49s in our limited sample. In its most developed form, a single phrase usually consists of 4–6 repeated echemes, each comprising (1) an initial syllable containing coalesced pulses lasting about 40–80 ms, (2) a shorter syllable containing isolated pulses repeated at about 180/s, and (3) ca. 1–3 higher amplitude syllables lasting 15–50 ms and containing pulses produced at about 360/s. The phrase begins and ends with less well-defined strings of pulses. Complex frequency and amplitude modulation patterns occur, with the loudest syllables having a lower dominant pitch (ca. 6.8 kHz) than the syllables containing coalesced pulses (14.2 kHz), and with the transition occurring during the syllables containing isolated pulses. Frequency modulation is also apparent at a shorter time scale, within individual pulses., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 223-227, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079.","Schouten MA, Duffels JP. 2002. A revision of the cicadas of the Purana carmente group (Homoptera, Cicadidae) from the Oriental region. Tijdschrift voor Entomologie. 145 (1): 29 - 46. doi: 10.1163 / 22119434 - 900000097.","Boulard M. 2013. The cicadas of Thailand, volume 2: taxonomy and sonic ethology. Manchester (United Kingdom): Siri Scientific Press; p. 436."]}

12. Champaka solivenae Lee, 2015 Champaka solivenae Lee 2015: 548–550, fig. 1. Additional material examined 1 male, specimen code 12. PH. MN.KGS. 10, PHILIPPINES, Mindanao, Mt. Kitanglad, 1773 m, 08°09.91 ʹ N, 124°55.96 ʹ E, 2 May 2012, K.B. R. Hill, D.C. Marshall and D. Mohagan (UCONN); 1 male, PHILIPPINES, Mindanao, Cotabato, Mt. Apo, small volcano, 6374 feet, 07°00.49 ʹ N, 125°15.28 ʹ E, 20 May 2010, D. Mohagan (UCONN). Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 211, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Genus Chremistica Stål, 1870 Remarks With the presence of the claspers and the long lateral lobes in the male pygofer, the following three species seem to belong to the Chremistica tridentigera species group, as suggested by Bregman (1985) and Yaakop et al. (2005), together with other Philippine species such as Chremistica semperi Stål, 1870 and Chremistica tagalica (Stål, 1870)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 204, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Bregman R. 1985. Taxonomy, phylogeny and biogeography of the tridentigera group of the genus Chremistica Stal, 1870 (Homoptera, Cicadidae). Beaufortia. 35: 37 - 60.","Yaakop S, Duffels JP, Visser H. 2005. The cicada genus Chremistica Stal (Hemiptera: Cicadidae) in Sundaland. Tijdschrift voor Entomologie. 148 (2): 247 - 306. doi: 10.1163 / 22119434 - 900000172."]}

15. Oncotympana averta Lee, 2011 Oncotympana sp.: Lee 2010: 20. Oncotympana averta Lee, 2011: 168–169 [TL: Philippines]; Lee 2015: 550–551. Distribution Philippines (Mindanao). Additional material examined 1 male, PHILIPPINES, Mindanao, Cotabato, Mt. Apo, small volcano, 6374 feet, 07°00.49 ʹ N, 125°15.28 ʹ E, 20 May 2010, D. Mohagan (UCONN); 2 males, same data except for 20–23 May 2010 (UCONN); 3 males, same data except for 20–24 May 2010 (UCONN)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 212, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2011. New genus and two new species of Oncotympanina stat. nov. (Hemiptera: Cicadidae: Cicadini) and the erection of Sonatini new tribe. J Asia Pac Entomol. 14 (2): 167 - 171. doi: 10.1016 / j. aspen. 2010.12.001.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

9. Dundubia vaginata (Fabricius, 1787) Dundubia vaginata: Lee 2010: 24; Lee 2015: 546. Distribution S. China, Philippines (Luzon, Sibuyan, Panay, Palawan, Dinagat and Mindanao), Malay Archipelago, S. Myanmar and India., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 210, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

14. Orientopsaltria fuliginosa (Walker, 1850) Orientopsaltria fuliginosa: Lee 2010: 24; Lee 2015: 550. Distribution Philippines (Luzon, Mindoro, Homonhon, Samar, Leyte, Panay, Negros, Mindanao and Basilan)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 211, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

28. Ggomapsalta fragilis Lee, 2015 Ggomapsalta fragilis Lee 2015: 554–555, fig. 3. Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

18. Neopurana bouptera Lee and Marshall sp. nov. (Figures 11, 12) Cicadinae sp. C: Hill et al. 2021: table 1, figs 5, 9, 10, 13, 14. Type material Holotype. Male, specimen code 12.PH. MN.KGS.01, PHILIPPINES, Mindanao, Mt. Kitanglad, 1773 m, 08°09.91 ʹ N, 124°55.96 ʹ E, 3 May 2012, K.B. R. Hill, D.C. Marshall and D. Mohagan (NMPM). Paratype. 1 male, specimen code 12. PH. MN.KGS. 02, same collection data (UCONN). Etymology The specific name is the feminine noun, a combination of the Greek prefix bou - meaning ̍large̾ and the Greek ptera meaning ̍wing̾. Measurements of types (2 males, holotype is larger) Length of body: 23.6 (22.7–24.4). Length of head and thorax together: 10.2 (9.8–10.6). Length of abdomen: 13.4 (12.9–13.8). Width of head including compound eyes: 6.6 (6.4– 6.8). Width of pronotum: 7.4 (7.0–7.8). Width of mesonotum: 6.5 (6.2–6.7). Width of abdominal tergite 3: 7.5 (7.3–7.7). Length of fore wing: 31.4 (29.4–33.3). Width of fore wing: 9.4 (9.1–9.7). Wing span: 68.1 (64.7–71.5). Description of male Head. Vertex light green with the following black marks: median mark surrounding ocelli, with its anterior tip broadened and nearly touching frontoclypeal suture; a pair of small longitudinal spots on both sides of the median mark; a pair of small transverse spots on antero-sublateral corners of vertex, with their inner tips connected to anterior tips of the small longitudinal spots; a pair of indistinct transverse spots on supra-antennal plates; and a pair of tiny spots posteromedially. Distance between lateral ocelli and compound eyes slightly longer than twice the distance between lateral ocelli. Postclypeus moderately swollen. Antennae black but lighter towards tip. Postclypeus light green with the following black marks: short fasciae or spots along inner parts of anterior half of transverse grooves; and large median spot on posterior one-third of postclypeus. Anteclypeus light green to greenish yellow with W-shaped mark in the middle. Rostrum greenish ochraceous to ochraceous but black apically; slightly passing beyond hind coxae. Lorum ochraceous with large black patch on inner corner. Gena greenish ochraceous with transverse spot between postclypeus and compound eye, touching postclypeus. Thorax. Pronotum green, greenish ochraceous, and ochraceous (medially). Inner area of pronotum with the following marks: a pair of median longitudinal black fasciae slightly broadened at both anterior and posterior ends; tiny spot medio-posteriorly; irregularly shaped brown spots and marks along or near paramedian and lateral fissures; transverse narrow black to fuscous fascia along anterior margin of pronotum; and a pair of curved broad fasciae along lateral margins of inner area. Pronotal collar greenish ochraceous with narrow transverse fascia along posterior margin. Anterolateral pronotal collar not dentate. Mesonotum green, greenish ochraceous, and ochraceous (medially) with the following black to fuscous marks: median longitudinal fascia broadened anteriorly and posteriorly, not reaching anterior margin of cruciform elevation; a pair of small round spots enclosing scutal depressions; a pair of inwardly curved fasciae along inner side of parapsidal sutures; a pair of two longitudinally arranged large spots on lateral sigilla; and a pair of tiny spots on lateral margin of mesonotum. Cruciform elevation light green to greenish ochraceous with a pair of black patches on apices of arms. Thoracic sternites greenish ochraceous to ochraceous with a pair of black spots on episterna 3. Legs light green to greenish ochraceous with black to fuscous marks. Fore femur with black primary, secondary and subapical spines and with black patch posteriorly on inner side. Fore- and mid tibiae fuscous apically. Fore- and mid pretarsi black. Wings hyaline. Fore wing with infuscations on radial, radiomedial, medial and mediocubital crossveins. Small, roundish infuscation present on each hind margin of radius posterior, median 1–4, and cubitus anterior 1 veins but not distinct on radius posterior and median 1 veins among them. Basal cell slightly tinged with ochraceous. Basal membrane dark grey. Hind wing jugum dark grey (proximally) to light grey (distally). Operculum slightly greenish ochraceous; obliquely trapezoidal with posterior angles rounded, extending to anterior margin of abdominal sternite III. Anterior inner margin concave. Opercula widely separated from each other, with gap about two-thirds as wide as operculum. Abdomen obconical, distinctly longer than head and thorax together. Tergites 2–6 light green (sub-medially) to ochraceous with a pair of brown sublateral spots. Posterior margins of tergite 2 narrowly fuscous. Tergite 7 castaneous with a pair of fuscous sublateral spots. Tergite 8 fuscous with brown median patch. Tergite 3 wider than mesonotum. Timbal cover slightly greenish ochraceous; nearly quarter circle with rounded anterolateral corner, about as long as wide. Timbal concealed by timbal cover in dorsal view. Abdominal sternites III–VI slightly greenish ochraceous. Sternite VII castaneous with black posterior margin. Sternite VIII castaneous with lighter postero-lateral parts. Sternites III with light ochraceous tubercle-like projection on each posterolateral surface, protruding posteriorly to partly cross anterior margin of sternite IV. Epipleurites 3–6 greenish ochraceous. Epipleurites 7–8 castaneous. Epipleurites 6–8 with black posterior margins. Genitalia. Pygofer oval in ventral view. Uncus simple, short, not bifurcate and with a widely truncate apex in ventral view; without median groove. Uncus slightly curved inward in lateral view. Aedeagus protruding from uncus; thick and bifurcate at apex. Basal lobe of pygofer absent. Song (Figure 12) About six minutes of song was recorded for this species at Mt. Kitanglad, including a sample from the holotype male, which is illustrated. The sound is a complex, resonant, amplitude-modulated buzz comprised mainly of pulse pairs (doublets) produced at approximately 100–150/s, although in a few samples the doublets are paired and in one segment the pulses appear briefly in threes. The song is produced in phrases containing, usually, one short echeme of ca. 0.6–1 s followed by a much longer echeme of ca. 8.5–11 s, sometimes with the phrases paired or strung together into longer series. One male produced a series of phrases lasting at least 125 s. Both short and long echemes begin with a string of single pulses and continue with (usually) doublets. Phrases are separated by short silent gaps of ca. 40–110 ms, as are (commonly) the short and long echemes contained within a phrase, and occasionally a gap of a few tens of milliseconds in duration occurs within what otherwise appears to be one of the long syllables. Amplitude modulates considerably, with intensity increasing to nearly double on the waveform and then back down in one cycle for each short echeme, and once or sometimes twice for long echemes. In the latter case, the initial amplitude modulation resembles a second short echeme appended to the long echeme. This gives the song a slow ̍revving̾ quality. Frequency content is not modulated but is complex, including two major peaks around 3.5–5 (usually dominant) and 11–13 kHz and a least one weaker peak around 8.5 kHz. The holotype specimen was collected by attracting it down from its perch on a vertical tree trunk, by movement of the collector around the base of the tree and gentle scratching and tapping of the trunk, followed by climbing and hand-capture. The male continued to face upward while backing down the tree trunk., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 218-222, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079."]}

20. Purana crassinotata Lee, 2015 Purana crassinotata Lee 2015: 551–553, fig. 2. Additional material examined 1 male, specimen code 12. PH. MN.IMP. 04, PHILIPPINES, Mindanao, Bukidnon Province, Impasugong Municipality, Impalutao Barangay, Impalutao Forest, 08°15.072 ʹ N, 125° 01.891 ʹ E, 21 April 2012, K. Hill, D. Marshall, T. Catanach and A Mohagan (UCONN). Distribution Philippines (Camiguin and Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 222-223, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

11. Champaka virescens (Distant, 1905) Champaka virescens: Lee 2010: 25; Lee 2015: 547. Distribution Philippines (Luzon, Camiguin Island north of Luzon, Negros, Samar, Mindanao and Basilan) and Borneo., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 210-211, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

13. Orientopsaltria inermis (Stål, 1870) Orientopsaltria inermis: Lee 2010: 22–24, fig. 6A, B; Lee 2015: 550. Distribution Philippines (Samar, Camiguin and Mindanao). Additional material examined 2 males, PHILIPPINES, Mindanao, Cotabato, Mt. Apo, Agro-ecosystem (Sudsuwayan), 25 May 2010, D. Mohagan (UCONN)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 211, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Genus Purana Distant, 1905 Remarks The Purana carmente species group was proposed by Schouten and Duffels (2002), of which 16 member species have been found so far in southern China, Vietnam, Thailand, Malaysia, Indonesia and the Philippines. The independence of the Purana carmente species group was also brought up by Lee (2009b). The new Purana species described below belongs to this species group. Two of the other Purana species of Mindanao, Purana nana Lee, 2009 and Purana crassinotata Lee, 2015, are included in the Purana abdominalis species group proposed by Lee (2009a). Lee (2009a) suggested Purana guttularis (Walker, 1858) be placed in the Purana ubina species group, but no male specimens have been confirmed. Its type specimen, which is a female, seems close to Purana kpaworensis Boulard, 2007, which was placed in the Purana tigrina species group by Lee (2009a), considering the similarities in fore wing venation. However, the species group placements of P. guttularis and P. kpaworensis need further study., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 222, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Schouten MA, Duffels JP. 2002. A revision of the cicadas of the Purana carmente group (Homoptera, Cicadidae) from the Oriental region. Tijdschrift voor Entomologie. 145 (1): 29 - 46. doi: 10.1163 / 22119434 - 900000097.","Lee YJ. 2009 b. A checklist of Cicadidae (Insecta: Hemiptera) in Palawan with five new species. J Nat Hist. 43 (41 - 42): 2617 - 2639. doi: 10.1080 / 00222930903244002.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5.","Lee YJ. 2009 a. Descriptions of two new species of the Purana abdominalis species group (Hemiptera: Cicadidae: Cicadini) from the Philippines, with a key to the species groups of Purana. J Nat Hist. 43 (25 - 26): 1487 - 1504. doi: 10.1080 / 00222930902941848."]}

Subtribe PURANINA Lee, 2013 Remarks We continue to use subtribe Puranina here. Unfortunately, no species of the Purana tigrina species group (see Duffels et al. 2007; Lee 2009a) or the true Purana Distant, 1905 (type species: Dundubia tigrina Walker, 1850) was included in the study by Hill et al. (2021).

16. Oncotympana obesa Lee and Marshall sp. nov. (Figures 9, 10) Oncotympana cf. averta: Hill et al. 2021: table 1, figs 4, 12, 14. Type material Holotype. Male, specimen code 12.PH. MN.KGC.01, PHILIPPINES, Mindanao, ca. 1/2 h up trail to Mt. Kitanglad, 1776 m, 08°09.81 ʹ N, 124°55.95 ʹ E, 1 May 2012, K.B. R. Hill, D.C. Marshall and D. Mohagan (NMPM). Paratypes. 2 males, PHILIPPINES, Mindanao, Bukidnon, Mt. Kiamo, 7 May 2012, A.B. Mohagan (UCONN); 2 males, PHILIPPINES, Mindanao, Mt. Kitanglad, 1773 m, 08°09.91 ʹ N, 124°55.96 ʹ E, PH. MN. KGS, 3 May 2012 (UCONN). Etymology The specific name is the Latin feminine adjective obesa, meaning ̍fat̾, ̍obese̾, ̍plump̾ or ̍swollen̾. Measurements of types (5 males) Length of body: 22.3 (21.3–23.3). Length of head and thorax together: 11.3 (11.0–11.6). Length of abdomen: 11.0 (10.2–11.7). Width of head including compound eyes: 6.3 (6.1– 6.4). Width of pronotum: 8.9 (8.6–9.4). Width of mesonotum: 8.0 (7.7–8.4). Width of abdominal tergite 3: 9.8 (9.7–10.1). Length of fore wing: 31.2 (30.6–32.3). Width of fore wing: 10.0 (9.5–10.2). Wing span: 68.6 (67.1–71.7). Diagnosis This new species resembles Oncotympana averta but has a distinctively larger body, more distinct infuscations on the fore wing, a proportionally much larger male abdomen, and differently shaped uncal lobes of the male genitalia, as detailed in the Remarks below. Description of male Head. Vertex yellowish green with the following black to dark brown marks: median mark enclosing ocelli, reaching posterior margin of vertex; a pair of spots on supraantennal plates; a pair of obliquely longitudinal small spots on posterolateral corners of vertex; a pair of indistinct tiny spots on anterolateral parts of vertex; and irregularly shaped transverse fascia along anterior margin of vertex. Supra-antennal plates produced anteriad, covering scape. Compound eyes brown in life. Distance between lateral ocelli and compound eyes about twice the distance between lateral ocelli. Antenna mostly black but with dark brown to fuscous pedicel. Postclypeus moderately swollen; yellowish green with a few black to dark brown fasciae along anterior transverse grooves. Anteclypeus ochraceous with a pair of central fuscous spots. Rostrum ochraceous with black apex, passing posterior margin of hind trochanter. Thorax. Pronotum greenish ochraceous. Inner area of pronotum with the following fuscous marks: a pair of medial longitudinal fasciae not reaching anterior margin of pronotum, broadened near posterior ends and meeting each other at posterior ends; a pair of marks along paramedian fissures; a pair of short longitudinal fasciae between median parts of paramedian fissures and posterior ends of lateral fissures; a pair of marks along lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar with a pair of indistinct marks near posterolateral corners; posterior marginal area very narrow. Anterolateral pronotal collar not dentate but sinuate. Mesonotum greenish ochraceous with the following fuscous to black marks: medial longitudinal fascia reaching anterior margin of cruciform elevation but not reaching anterior margin of mesonotum; a pair of small roundish spots enclosing scutal depressions; a pair of large obconical paramedian marks on submedian sigilla, indistinct in their inner parts; a pair of long obconical sublateral marks on lateral sigilla, discontinued in the middle; and a pair of small spots on posterolateral corners of mesonotum. Cruciform elevation greenish ochraceous without distinct marks. Thoracic sternites greenish ochraceous. Legs greenish ochraceous to green with some brown parts and black to fuscous marks. Fore femur with primary and secondary spines and minute subapical spines. Wings hyaline, slightly tinged with brown. Fore wing costal vein mostly ochraceous but fuscous near node. Basal vein of apical cell 1 about one-third to one-fourth as long as longitudinal vein of the apical cell 1. Infuscation present at bases of apical cells 1–5 and 7– 8 and on cubitus anterior 2 vein. A distinct spot appearing on each hind margin of radius anterior 2, radius posterior, median 1–4 and cubitus anterior 1 veins, forming a series of spots on subapical margin of fore wing. Basal membrane and base of hind wing jugum greenish (bright blue in life). Operculum light greenish ochraceous without distinct marks; much wider than long, with rounded posterior apex passing beyond posterior margin of abdominal sternite II and with posterolateral corner almost straight but slightly convex. Opercula separated from each other. Abdomen obconical in dorsal view, short, shorter than or about as long as head and thorax together. Tergites 2–8 mostly castaneous, with light green to green posterior margins (blue in life). Tergites 2–5 with a pair of fuscous paramedian patches. Tergites 4–8 with a pair of lateral fuscous patches. Posterior margin of tergite 3 much wider than pronotum or mesonotum. Timbal cover dull greenish ochraceous with black to fuscous patch along posterior margin; globose, wider than long, with anterior and posterior inner corners smoothly rounded. Timbal completely concealed by timbal cover in dorsal view. Abdominal sternites II–VIII fuscous, with green posterior margins of sternites III–VI. Genitalia. Pygofer barrel-shaped in ventral view. Uncus bifurcate. Uncal lobes each divided into two parts: a long external part, and a shorter internal part, which is less than half as long as external part: in ventral view, external parts curved and extended obliquely laterad, and internal parts convexly curved with tips directed inward; in lateral view, external parts convexly curved with tips directed inward, and internal parts curved inward. Aedeagus slender. Dorsal beak long, narrow, about as long as anal styles. Remarks This species is very similar to Oncotympana averta but can be distinguished from O. averta by the following characteristics: body distinctly larger; infuscation present at base of apical cells 4 and 8 (vs absent there in O. averta); basal membrane and base of hind wing jugum greenish (vs greyish in O. averta); operculum with posterolateral corner almost straight but slightly convex (vs slightly concave in O. averta); posterior margin of tergite 3 much wider than pronotum or mesonotum (vs about as wide as or slightly narrower than mesonotum in O. averta); abdominal sternites II–VIII fuscous (vs sternites II– V mostly ochraceous but sternites VI–VIII fuscous in O. averta); timbal cover globose but more or less flat (vs much more prominently globose in O. averta); dark-coloured patch absent on inner corner of timbal cover (vs present in O. averta); external parts of uncal lobes curved and extended obliquely laterad in ventral view (vs nearly parallel to each other but with curved tips extended obliquely laterad in O. averta); and internal parts of uncal lobes convexly curved with tips directed inward in ventral view (vs concavely curved with hooked tips extended laterad in O. averta). Song (Figure 10) Recordings of male calling song totalling 320 s were taken at the type locality (the holotype male is illustrated). Males were recorded producing song in phrases of 4–11 s duration (4–6 most common) comprised of two-syllable echemes repeated at about 5.2– 6.5 echemes/s. Note that such values are commonly dependent on temperature. The syllables within each echeme contain strongly contrasting sound frequencies. The first syllable contains most of its sound energy within the range 7–15 kHz, with a peak at about 10 kHz, while the second syllable, which is nearly an order of magnitude quieter than the first, contains sound energy with a peak of 4.5 kHz and a broad shoulder of quieter frequencies ranging from 2–16 kHz. The fundamental sound pulses are produced at about 225/s in the first syllable, which lasts 0.04 s, and slightly faster at about 235/s in the second syllable which is twice as long. This species sang from low stations on tree trunks in the forest understory in low light conditions, as early as 7.00am. Males could be attracted by mouth-clicks or finger-snaps produced a fraction of a second after the end of the complete song phrase. Immediately after hearing the response, the male would flick his wings with a synchronised timbal click, before turning or moving in the direction of the sound. Several specimens from the paratype series were collected with this technique., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 212-216, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079."]}

30. Philipsalta lata Lee and Marshall sp. nov. (Figures 19, 20) Type material Holotype. Male, specimen code 12.PH. MN.CAC.01, PHILIPPINES, Mindanao, Cateel / Compostela Rd., nr bdry, 726 m, 07°39.93 ʹ N, 126°12.82 ʹ E, 6 May 2012, K.B. R. Hill, D.C. Marshall and A.B. Mohagan (NMPM). Etymology The specific name is a Latin feminine adjective meaning ̍broad̾ or ̍wide̾, in reference to the broader body of this species compared with the previous species. Measurements of type (1 male) Length of body: 15.4. Length of head and thorax together: 6.4. Length of abdomen: 9.0. Width of head including compound eyes: 4.1. Width of pronotum: 4.6. Width of mesonotum: 4.0. Width of abdominal tergite 3: 4.3. Length of fore wing: 17.7. Width of fore wing: 6.6. Wing span: 39.1. Description of male Body tiny. Head: Vertex ochraceous with the following black marks, connected to each other: inverted triangular mark surrounding median ocellus; a pair of marks surrounding lateral ocelli, expanding to anterior and lateral margins of vertex; a pair of spots on inner corners of supra-antennal plates; and a pair of spots on posterolateral corner of vertex (touching compound eyes). Compound eyes light brown in life. Distance between lateral ocelli and compound eyes about as long as distance between lateral ocelli. Antennae black to fuscous. Postclypeus not swollen anteriad and rather flat ventrally; dull ochraceous with broad black median fascia. Anteclypeus dull ochraceous with median fuscous fascia and fuscous lateral margins. Rostrum ochraceous with black apex; with apex extending slightly beyond posterior margin of mid coxa. Lorum dull ochraceous with black longitudinal spot on inner corner of lorum. Gena dull ochraceous with black spot on inner corner of gena. Thorax. Pronotum dull ochraceous. Inner area of pronotum with the following black to fuscous marks: tiny indistinct spot on posteromedian margin of inner area; a pair of short longitudinal fasciae between about middle of paramedian fissures and posterior ends of lateral fissures; a pair of fasciae along paramedian fissures, with their posterior parts indistinct; a pair of fasciae along lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar very narrow; without distinct marks. Anterolateral pronotal collar not dentate. Mesonotum dull ochraceous with a pair of black to fuscous patches on entire submedian sigilla and a pair of larger longitudinal patches on almost entire lateral sigilla but partly indistinct. Cruciform elevation light ochraceous without marks. Thoracic sternites ochraceous. Legs ochraceous to dull brown with some fuscous marks. Fore femur with primary, secondary and subapical spines. Wings hyaline without infuscation; with extremely narrow marginal areas. Fore wing venation dull brown basally but fuscous distally. Basal membrane grey. Hind wing with six apical cells. Operculum small, semicircular, with lateral margin slightly concave, not reaching posterior margin of sternite II; light ochraceous. Opercula separated from each other. Abdomen triangular in dorsal view, much longer than head and thorax together; ochraceous. Tergite 1 about half as long as tergite 2 in median length. Tergites 3–7 each with fuscous spot anteromedially. A pair of fuscous globose bumps present on lateral margins of tergite 2. Timbal cover absent. Timbals not extending below wing bases ventrally. Abdominal sternites ochraceous with greyish patches irregularly. Genitalia. Pygofer ovate with triangular dorsal beak in ventral view. Upper lobe moderately pointed. Dorsal beak triangular. Median lobe of uncus very short. A pair of clasperlike processes protruding downward but slightly curved, with apices heading outward. Remarks This new species can be distinguished from Philipsalta exilis Lee and Marshall sp. nov. mainly by the following characteristics, besides the differences in colours and marks: fore wing ulnar cell 2 more spacious than ulnar cell 1 (vs smaller than ulnar cell 1 in P. exilis); male operculum with lateral margin slightly concave (vs not concave in P. exilis); male abdomen with convex lateral margins in dorsal view (vs with almost linear marginal lines in P. exilis); median lobe of uncus comparatively short (vs moderately protruding, longer in P. exilis); and clasper-like processes slightly curved outward in ventral view (vs straight in P. exilis). Song (Figure 20) A 70s song sample was recorded from the holotype male and is illustrated here. Males produce phrases of 0.1– 0.16 s duration at 1.8–2.6 phrases/s. Most phrases contain a single echeme followed by an isolated doublet after a silent gap of about 11–13 ms; some phrases lack the final doublet. The quality of the one recording is weak, in part due to environmental noise and echo, but for the clearest phrases the main echeme exhibits a complex pulse pattern, with sets of three stronger pulses or doublets produced at 350 pulses/s separated by single weaker pulses, and with the gaps preceding and following the weaker pulses being shorter than those separating the main pulses. The frequency spectrum is broad, reaching from at least 8 kHz to well above 20 kHz, and has a peak around 13 kHz. Both the sound frequency and amplitude increase somewhat during the first half of each song phrase. The song of Philipsalta lata Lee and Marshall sp. nov. is distinguishable from those of P. exilis and P. nigrina by the short single-doublet element ending most phrases, as well as by the complex 3 + 1 pulse pattern within the main echeme (P. exilis and P. nigrina have pulses or doublets produced at a uniform rate)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 236-239, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268

5. Chremistica kyoungheeae Lee 2010 Chremistica kyoungheeae Lee 2010: 16–18, fig. 2; Lee 2015: 546. Distribution Philippines (Dinagat and Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 205, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

4. Hamza ciliaris (Linnaeus, 1758) Hamza ciliaris: Lee 2010: 15, fig. 1C, D; Lee 2015: 546. Distribution Philippines (Mindanao), Palau and Indonesia (Maluku, Timor and Banggai Archipelago)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 204, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

2. Platypleura minima Lee and Marshall sp. nov. (Figures 4, 5, 6 (A)) Type material Holotype. Male, specimen code 12.PH. MN. MUS.06, PHILIPPINES, Mindanao, Bukidnon, Maramag, Mt. Musuan, 7°52.62 ʹ N, 125°4.19 ʹ E, 21 April 2012, K. B. R. Hill, D. C. Marshall and A. B. Mohagan (NMPM). Etymology The specific name is the feminine form of the Latin adjective minimus meaning ̍smallest̾ or ̍least̾, in reference to the black and grey patterns on the wings, which is the lightest or least infuscated among the Philippine species of Platypleura. Measurements of type (1 male) Length of body: 18.7. Length of head and thorax together: 9.6. Length of abdomen: 9.1. Width of head including compound eyes: 8.3. Width of pronotum: 10.3. Width of mesonotum: 7.9. Width of abdominal tergite 3: 8.4. Length of fore wing: 25.5. Width of fore wing: 9.2. Wing span: 59.1. Diagnosis This new species is very similar to Platypleura bella Lee and A. Mohagan sp. nov. but smaller in body size (outside of the range of all the body part sizes of the available male specimens of P. bella) and has less infuscation on the fore wing and the hind wing with some distinguishing points: anal cell 2 fuscous on about basal one-fourth (vs fuscous on about basal one-third to two-thirds in P. bella); anal cell 4 fuscous on about basal one-third (vs fuscous on about basal half to two-thirds in P. bella); anal cell 5 fuscous on about basal one-third (vs fuscous on about basal half to two-thirds in P. bella); longitudinal length of light greenish ochraceous portion of tergite 8 long, about four times as long as black portion (vs very short, shorter than one-third as long as black portion in P. bella); and timbal cover mostly greenish ochraceous (vs mostly black to fuscous in P. bella). The calling songs are readily distinguishable from each other, as discussed below. Description of male Head. Vertex slightly greenish ochraceous with the following black marks: median large spot enclosing ocelli, with its anterior end reaching anterior margin of vertex; a pair of small longitudinal spots between lateral ocelli and compound eyes; a pair of small spots on posterolateral corners of vertex; a pair of short obliquely longitudinal fasciae between postclypeus and compound eyes; and a pair of narrow fasciae along compound eyes. Distance between lateral ocelli and compound eyes distinctly longer than twice the distance between lateral ocelli. Antenna mostly castaneous but pedicel fuscous. Postclypeus not swollen anteriad; greenish ochraceous with a medial longitudinal black fascia, wider anteriorly, and transverse fasciae along transverse grooves, becoming indistinct going posteriad. Anteclypeus dull brown except ochraceous median ridge. Rostrum slightly greenish ochraceous but darkened towards apex to become black apically; with apex not reaching medial part of posterior margin of abdominal sternite II. Lorum black except ochraceous anterior and lateral margins. Gena black to fuscous with greenish ochraceous spot on posterior margin. Thorax. Pronotum ochraceous to greenish ochraceous. Inner area of pronotum with the following black to fuscous marks: posteromedial inverted T mark; indistinct spot on anteromedial part of inner area; rather indistinct irregularly shaped marks along paramedian fissures; a pair of irregularly shaped marks along lateral fissures; and a pair of curved fasciae along lateral margins of inner area. Pronotal collar darkened anteriorly and laterally on both sides. Anterolateral pronotal collar broadly developed and roundly angulated with an angle of 100–110°, expanding to a width much greater than the widest width of posterior pronotum, mesonotum and abdomen. Mesonotum reddish ochraceous with the following black marks: median longitudinal fascia, with its posterior end widened and reaching anterior margin of cruciform elevation, but its anterior end not reaching anterior margin of mesonotum; a pair of small transverse spots enclosing scutal depressions; a pair of large semicircular spots on submedian sigilla, with their anterior ends reaching anterior margin of mesonotum; and a pair of long inverted triangular marks on lateral sigilla, with their anterior ends reaching anterior margin of mesonotum. Cruciform elevation greenish ochraceous. Fore-, mid- and hind trochanters and femora brown to dark brown. Wings. Fore wing outer margin slightly curved outwardly (not straight or linear). Fore wing with venation ochraceous to reddish ochraceous, darker apically; furnished with white patch on about basal two-fifths to one-half (hyaline on remaining part: about apical half) and with the following infuscations: three spots on radial cell; mark on both ends of both ulnar cells 1 and 2; rather indistinct marks on basal half and tiny spot on apical end of ulnar cell 3; indistinct small marks on medial cell; mark on both apical and basal ends of apical cell 1; irregularly shaped small marks on subapical parts of apical cells 2–7, sometimes indistinct; one or two tiny dots on apical ends of apical cells 2–7; and two spots on apical end of apical cell 8. Basal membrane dark grey with light green margin. Hind wing fuscous except outer marginal area and apical portion of apical cells 1–6. Anal cell 2 fuscous on about basal one-fourth. Anal cell 4 fuscous on about basal one-third. Anal cell 5 fuscous on about basal one-third. Hind wing jugum grey with light green margin. Operculum mostly fuscous except greenish ochraceous margin; semicircular, transversely wide; short, nearly reaching posterior margin of abdominal sternite II. Opercula overlapping medially. Abdomen obconical in dorsal view, short, slightly shorter than head and thorax together; mostly black with light greenish ochraceous caudal margin on each of tergites 3–7. Tergite 8 mostly light greenish ochraceous except for narrow anterior part, with longitudinal length of light greenish ochraceous portion about four times as long as black portion. Timbal cover mostly greenish ochraceous with black to fuscous patch posteriorly;semicircular, wider than long. Timbal concealed by timbal cover in dorsal and lateral views. Abdominal sternites black with greenish ochraceous to dull brown caudal margin on each of sternites II–VII. Genitalia. Pygofer long ellipsoidal with acuminate apex (wide dorsal beak) in ventral view. Uncus simple, not bifurcate, with apical margin rounded. Aedeagus not tapering down to apex. Song (Figure 5) About 103 s of song was recorded from uncollected males at Mt. Musuan, Bukidnon, on 21 April 2012, at 11:05 AM. An additional 95s of song was recorded on 7 July 2010, at the same location, although no specimens were collected. The sound is a continuous frequency-modulated whining buzz, similar in quality to a car alarm. Pulses of approximately uniform amplitude are regularly produced at about 435–455 pulses/s in the 2012 recordings, and at 380 pulses/s in the 2010 recording. The sound sweeps upward and downward (the latter more slowly) in pitch at about 3.7–3.8 cycles/s in all recordings, with amplitude subtly shifting concordantly. Most sound energy lies within the range 6.5–12.5 kHz, with the peak shifting from about 8.5 to about 9.5 kHz across the cycles. Males sang from the upper surface of lateral branches of an Acacia mangium tree with the head elevated and the abdomen and wings angled downward. This together with their lichenlike colour pattern contributed to making them difficult to locate. The song of P. minima differs from that of P. bella in the much slower rate of concordant amplitude and frequency oscillation,ca.3.7–3.8 cycles/s (vs 35–40 cycles/s in P. bella). This rate difference, nearly an order of magnitude, is not likely to be caused by environmental temperature differences during recording because the underlying pulse rate is similar for both species. Furthermore, some platypleurine species have been shown to thermoregulate (e.g. Sanborn et al. 2004), which reduces the significance of environmental temperature. Remarks Endo and Hayashi (1979) reported Platypleura sp. aff. nobilis (Germar, 1830) (TL: Java) based on a male specimen collected from Mt. Apo. Lee (2010) identified the specimen as P. elizabethae. However, this record is more likely to be Platypleura minima because the cloudy patterns on the wings of Platypleura nobilis are more similar to those of P. minima than to P. bella, P. elizabethae or P. transitiva, which was misidentified as P. elizabethae by Lee (2010, 2015). The dorsal habiti of male specimens of several Philippine Platypleura species are shown in Figure 6 for comparison., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on pages 199-204, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Sanborn AF, Villet MH, Phillips P. 2004. Endothermy in African platypleurine cicadas: the influence of body size and habitat (Hemiptera: Cicadidae). Physiol Biochem Zool. 77 (5): 816 - 823. doi: 10.1086 / 422226.","Endo T, Hayashi M. 1979. A note on Cicadidae collected and observed by T. Endo in the Philippines. Cicada Tokyo. 1: 35 - 38. (In Japanese with English summary).","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

3. Platypleura transitiva Lee, 2021 (Figure 6 (D)) Platypleura elizabethae: Lee 2010: 15, fig. 1A, B; Lee 2015: 546; Lee et al. 2016: 190 (nec Lee 2009b). Platypleura transitiva Lee 2021: 261–263, figs 1, 2(B) [TL: Surigao del Sur, Mindanao, Philippines]. Distribution Philippines (Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 204, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5.","Lee YJ, Marshall DC, Mohagan A, Hill KBR. 2016. Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species. Zootaxa. 4097 (2): 187 - 202. doi: 10.11646 / zootaxa. 4097.2.2.","Lee YJ. 2009 b. A checklist of Cicadidae (Insecta: Hemiptera) in Palawan with five new species. J Nat Hist. 43 (41 - 42): 2617 - 2639. doi: 10.1080 / 00222930903244002."]}

25. Nelcyndana tener (Stål, 1870) Nelcyndana tener: Lee 2010: 26, fig. 7; Lee 2015: 553. Distribution China, Philippines (including Mindanao) (not previously reported from specific islands) and Borneo., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Genus Philipsalta Lee, Marshall and Hill, 2016 Philipsalta Lee, Marshall and Hill, 2016 in Lee et al., 2016: 197. Type species Philipsalta nigrina Lee, Marshall and Hill, 2016 (Dinagat, Philippines). Remarks The two new species described below under Philipsalta are placed in this monotypic genus because their morphological characteristics fit many of the characteristics listed in the description of the genus given by Lee et al. (2016), which are as follows. Body tiny. Postclypeus rather flat ventrally. Pronotal collar very narrow, not dentate. Wings hyaline without infuscation. Fore wing cubital cell spacious, about as broad as medial cell. Hind wing with six apical cells. Male operculum small, not reaching posterior margin of sternite II. Male abdomen triangular in dorsal view, longer than head and thorax together. Timbal cover absent. Timbals not extending below wing bases ventrally. Pygofer ovate with triangular dorsal beak in ventral view. Upper lobe of pygofer present. Dorsal beak triangular. A pair of clasper-like processes protruding downward., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 233, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ, Marshall DC, Mohagan A, Hill KBR. 2016. Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species. Zootaxa. 4097 (2): 187 - 202. doi: 10.11646 / zootaxa. 4097.2.2."]}

25. Nelcyndana tener (Stål, 1870) Nelcyndana tener: Lee 2010: 26, fig. 7; Lee 2015: 553. Distribution China, Philippines (including Mindanao) (not previously reported from specific islands) and Borneo.

Subtribe PURANINA Lee, 2013 Remarks We continue to use subtribe Puranina here. Unfortunately, no species of the Purana tigrina species group (see Duffels et al. 2007; Lee 2009a) or the true Purana Distant, 1905 (type species: Dundubia tigrina Walker, 1850) was included in the study by Hill et al. (2021)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 222, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ, Emery D. 2013. New genera and species of Leptopsaltriini (Hemiptera: Cicadidae: Cicadinae) from India and Tibet, with the descriptions of five new subtribes. Zoosystema. 35 (4): 525 - 535. doi: 10.5252 / z 2013 n 4 a 6.","Duffels JP, Schouten MA, Lammertink M. 2007. A revision of the cicadas of the Purana tigrina group (Hemiptera, Cicadidae) in Sundaland. Tijdschrift voor Entomologie. 150 (2): 367 - 387. doi: 10.1163 / 22119434 - 900000237.","Lee YJ. 2009 a. Descriptions of two new species of the Purana abdominalis species group (Hemiptera: Cicadidae: Cicadini) from the Philippines, with a key to the species groups of Purana. J Nat Hist. 43 (25 - 26): 1487 - 1504. doi: 10.1080 / 00222930902941848.","Hill KBR, Marshall DC, Marathe K, Moulds M, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price BW, Duffels JP, et al. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebr Syst. 35 (5): 570 - 601. doi: 10.1071 / IS 20079."]}

26. Huechys parvula Haupt, 1924 Huechys parvula: Lee 2010: 25; Lee 2015: 553–554. Distribution Philippines (Samar, Camiguin and Mindanao)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 232, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Haupt H. 1924. Die Gattung Huechys Am. et Serv. (Homopt.) Beitrage zu einer Monographie. Deutsche entomologische Zeitschrift. 1924: 201 - 225.","Lee YJ. 2010. Cicadas (Insecta: Hemiptera: Cicadidae) of Mindanao, Philippines, with the description of a new genus and a new species. Zootaxa. 2351 (1): 14 - 28. doi: 10.11646 / zootaxa. 2351.1.2.","Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

31. Lembeja fatiloqua (Stål, 1870) Lembeja fatiloqua: Lee 2015: 555. Distribution Philippines (Mindanao and Basilan) and Malaysia (Sabah). Additional material examined 2 males, PHILIPPINES, Mindanao, Bukidnon, Maramag, Mt. Musuan, 7 May 2012, K.B.R. Hill, D.C. Marshall and A.B. Mohagan (UCONN); 1 male, PHILIPPINES, Mindanao, Davao City, Bunawan Hilltop, 22 April 2011, A. Baguhin (UCONN)., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 240, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Lee YJ. 2015. Additional records of cicadas (Hemiptera: Cicadidae) to the fauna of Mindanao, Philippines, with the description of three new species and a key to the species of Champaka Distant. Zootaxa. 4007 (4): 545 - 556. doi: 10.11646 / zootaxa. 4007.4.5."]}

Tribe PLATYPLEURINI Schmidt 1918 Hamzaria Distant 1905b: 382. Hamzini: Lee 2014: 61 (Platypleurini synonymised with Hamzini). Platypleurini Schmidt 1918: 378; Marshall et al. 2018a: 50–52; ICZN, 2020: 147–149 (Platypleurini conserved over Hamzaria); ICZN, 2021: 60 (date correction); Lee 2021: 261., Published as part of Lee, Young June, Marshall, David C., Mohagan, Alma B., Hill, Kathy B. R. & Mohagan, Dave P., 2023, Revised checklist of Cicadidae (Insecta: Hemiptera) of Mindanao, Philippines, with descriptions of a new genus and nine new species, pp. 193-242 in Journal of Natural History 57 (1 - 4) on page 196, DOI: 10.1080/00222933.2023.2171820, http://zenodo.org/record/7737268, {"references":["Schmidt E. 1918. Platypleura frontalis, eine neue Singzikade von Sumatra. (Rhynchota Homoptera). Stettiner Entomologische Zeitung. 79: 378.","Distant WL. 1905 b. Rhynchotal notes. - XXXI. Ann Mag Nat Hist. (7) 15 (): 379 - 387. doi: 10.1080 / 03745480509443064.","Lee YJ. 2014. Cicadas (Hemiptera: Cicadidae) of Laos, with the description of four new genera and two new species. Annales de la Societe entomologique de France (N S). 50 (1): 59 - 81. doi: 10.1080 / 00379271.2014. 934038.","Marshall DC, Moulds M, Boulard M, Sanborn AF, Ewart A, Wei C, Marathe K, Popple LW, Price BW, Simon C. 2018 a. Case 3761 - Platypleurini Schmidt, 1918: proposed precedence over Hamzaria Distant, 1905 to conserve nomenclatural stability in the Cicadidae (Insecta: Hemiptera: Auchenorrhyncha: Cicadoidea). Bull Zool Nomencl. 75 (1): 49 - 54. doi: 10.21805 / bzn. v 75. a 012."]}

Semia spiritus sp. nov. (Figs 2 B, 3C–D, 5) Type material. Holotype: ♂, Kon Tum Prov., Ngoc Linh Nature Reserves, 1900 m, 15°00’39”N, 107°57’53”E, III 2016; (typed ink label) (VNMN; E-00112). Paratypes: 2♂♂, Lam Dong Prov., Bidoup Nui Ba National Park, 2000 m, 12°06’43”N, 108°57’53”E, 10 V 2013; 6♂♂, 1♀, same location as holotype, 20 IV 2016 (VNMN; E-113-121), 2♂♂, same data as holotype (AM, K493593-94); 1♀, Lai Chau Prov., 1600 m, 22°18' 42" N 102°09'22" E, 18 V 2015; 2♂♂, Thua Thien Hue, Bach Ma National Park, 1400 m, 16°11’51”N, 107°51’36”E, 25 IV 2016 (DE). Other Material. 5♂♂, Kon Tum Prov., Ngoc Linh Nature Reserves, 1900 m, 15°00’39”N, 107°57’53”E, IV 2014 (DE); 7♂♂, same location, 15 V 2015 (DE); 2♂♂, Lam Dong, Bidoup Nui Ba National Park, 2000 m, 12°06’43”N, 108°57’53”E, 10 V 2013 (DE); 2♂♂, Thua Thien Hue, Bach Ma National Park, 1400 m, 16°11’51”N, 107°51’36”E, 25 IV 2016 (DE). Description of male ( Figs 2 B, 3C–D, 5A–B). Head mostly brown, brown anteriorly, with black markings around ocelli; ocelli pink; postclypeus brown, with black “V” marking dorsally, black horizontal striations along transverse grooves mid-anteriorly, pale brown patch on either side of midline over central third, and prominently shiny black marking over posterior third; anteclypeus mainly brown; gena and lorum dark brown; rostrum brown, black at tip, extending to mid-sternite II; compound eyes dark brown; antennae light brown; supra-antennal plates brown. Thorax: Pronotum mostly pale brown-ochraceous, with black markings outlining anterior brown stripe and posterior triangle marking on midline, darker brown patches laterally along ridges between parapsidal sutures, and black strip along lateral border; pronotal collar evenly brown-red to ochraceous but greenish on lateral margins, with median black spot on posterior margin, expanded laterally. Mesonotum ranging from light to olive-brown, with black median stripe, black patterned motif on submedian sigella on most specimens, but central area olive brown on majority of specimens from Bach Ma Mountain, with dull black pattern radiating over lateral sigilla and reddish brown patch along lateral margin; scutal depressions black; cruciform elevation light brown, with lateral depressions dark and golden hairs sparsely spread. Metanotum dark brown. Legs predominantly light brown; coxa brown with anterior black stripes; trochanters brown; femora with small black markings at ends, with primary spine erect, brown; tibiae similarly patterned to femora, but black around tarsi; tarsi brown anteriorly, blackish at tips; claws dark brown. Meracanthus narrow, pale brown, sharply pointed, overlapping anterior half of corresponding operculum. Wings hyaline, veins cream to brown, cream at node; fore wing with infuscations along crossveins r, m, r-m, and m-cu, vein CuA2, bases and apices of veins RA2, RP, M, M1-4, and CuA1, nodal line intersection, and outer margins of apical cells 1–7; basal membrane uniformly grey; basal cell translucent; hind wing with 6 apical cells; with continuous infuscations along proximal veins of apical cells 1–3 and light infuscations along outer margins of apical cells 1–3 in most specimens; plaga grey, covering jejum, with translucent area near distal margin. Opercula pale yellowish brown, with black lateral margins; spatulate, not extending beyond posterior margin of sternite II, rounded medially, almost meeting on midline. Timbals (Fig. 2 B) with 5 long ribs; ribs 1–4 dark brown, full length fused ventrally, thinner through dorsal half to prominent central expansion, thicker ventrally, pale brown to yellow, interspersed by three intercalary ribs; rib 5 discontinuous centrally. Timbal plate prominent; timbal dome pale brown, extending over half width of timbal, inverted distally. Apodeme pit small, slit like, conspicuous. Abdomen with tergites mainly brown, darker medially, lighter laterally, with black longitudinal stripe near lateral margins; tergite 1 brown; timbal covers pale brown, triangulate and convex, completely covering timbals; tergites 2–7 brown; tergite 8 dark brown; sternites pale brown, translucent. Genitalia (Fig. 3 C–D): Pygofer trapezoid in ventral view; dorsal beak short, rounded, protruding upwards in lateral view, not extending to elongated anal styles; distal shoulder acute, directed and curved laterally; uncal lobe long, with apposed inner margins, with lateral margins angled anteriorly towards rounded apex, spatulate and gently curved in lateral view; basal lobe stout and curved inward in ventral view. Aedeagus short, concealed within uncus. Description of female (Fig. 5 C–D). Head and thorax similar to male, with generally darker and greenish coloration; pronotal collar olive green, ampliate laterally; mesonotum distinctly olive green with similar black patterns to male. Wings similar to male, darker infuscations along posterior margins of apical cells 1–6. Abdomen dark brown; tergite 1 brown; tergites 2–3 with black anterior margins, brown posteriorly; tergites 3– 8 black-brown medially, browner laterally with distinct golden hairs; sternites I–VII dark brown to black, black mottling along anterior margins; abdominal segment 9 black, dark brown laterally, green along dorsal midline. Ovipositor dark brown, not extending beyond abdomen. Measurements (12 males, 2 females). Body length: male 35.2–39.1 (36.8); female 27.8–31.0 (29.9). Length of head + thorax: male 13.8–16.0 (14.7); female 13.4–14.9 (14.2). Length of abdomen: male 19.6–23.8 (22.0); female 14.4–16.1 (15.2). Fore wing length: male 39.6–44.8 (42.0); female 41.5–44.5 (42.9). Head width: male 8.3–9.3 (8.9); female 8.8–9.2 (9.0). Pronotum width: male 9.9–11.1 (10.5); female 9.0–9.4 (9.2). Abdomen width: male 12.0–13.9 (12.8); female 11.8–11.8 (11.8). Etymology. The name “ spiritus ” (Latin masculine noun for spirit) derives from the Vietnamese “ Linh ” (spirit, soul), reflecting the predominant location of this species on Ngoc Linh Nature Reserves. Distinguishing features. Males of S. spiritus sp. nov. lack the continuous infuscation along the posterior margin of the hind wing apical cells 1–6, so are readily distinguished from S. klapperichi, S. majuscula, S. magna sp. nov., and S. tibetensis. Semia spiritus sp. nov. (body length> 35.2 mm) is larger than S. spinosa (body length S. gialaiensis (body length S. albusequi sp. nov., (body length S. brevidilata (body length 34 mm). Semia spiritus sp. nov. lacks the distinct white pollinosity present on the tergite 8 of S. watanabei. The large timbal covers on S. spiritus sp. nov. contrast with rudimentary covers which leave the majority of timbals exposed in S. lachna, S. brevidilata, and S. watanabei. The dark brown coloration of S. spiritus sp. nov. contrasts with the pallor of S. pallida sp. nov. and the olive-brown markings on S. hainanensis . Semia spiritus sp. nov. also lacks the black markings seen laterally on the pronotal collar and tibiae of S. hainanensis . Distribution, habitat, and seasonality. This species has been found in Central Vietnam at Bach Ma National Park and Ngoc Linh Nature Reserves and 500 km southwards at Bidoup Nui Ba National Park (Fig. 4). Specimens have been collected between March and May.

Semia magna sp. nov. (Figs 1, 2 A, 3A���B) Type material. Holotype: 1♂, Kon Tum Prov., Ngoc Linh Nature Reserves, 1900 m, 15��00���39���N, 107��57���53���E, 20 IV 2016 (typed ink label) (VNMN; E-00105). Paratypes: 4♂♂, 2♀♀, same data as holotype (VNMN;E- 00106- 111); 2♂♂, 2♀♀, same data as holotype (AM; K.493586-89); 5♂♂, 4♀♀, same data as holotype (DE). Other material. 7♂♂, same location, 15 III 2016 (DE); 1♂, Thua Thien Hue Prov., Bach Ma National Park, 1400 m, 16��11���51���N, 107��51���36���E, 20 III 2016 (DE). Description of male (Figs 1 A���B, 2A, 3A���B). Head mostly brown, brown anteriorly with irregular black markings around ocelli and margins of eyes; ocelli pink, distance between lateral ocellus and corresponding eye much longer than that between lateral ocelli; postclypeus with brown marking along anterior third of midline, black horizontal striations along transverse grooves mid-anteriorly, brown patch either side of midline over central third, and shiny black patch posteriorly; anteclypeus mainly brown with black anterior and posterior markings; gena and lorum mostly black with lateral margins brown; rostrum brown, black at tip, extending to anterior margin of abdominal sternite II; compound eyes dark red; antennae black; supra-antennal plates black. Thorax: Pronotum mostly pale brown-ochraceous with black markings outlining two midline brown triangles and black patch along parapsidal sutures and lateral margins; pronotal collar evenly brown-red to ochraceous, expanded laterally. Mesonotum olive-brown with black median stripe, black patterned motif on submedian sigella, impressive black patterning stippled over lateral sigella, black strip on distal half of lateral margin, and black scutal depressions. Cruciform elevation dark brown with black anterior borders, lateral depressions brown, and golden hairs sparsely spread. Metanotum dark brown. Legs variously patterned brown and black; coxa brown with anterior black stripes; trochanters brown; fore femur with equivalent black-brown-black annular markings and with primary spine prominent, erect, black; mid and hind femora with increasingly brown central segments and black extremities; tibia similarly patterned to femora; tarsi brown anteriorly, black at tips; claws black. Meracanthus narrow, striped brown and black, sharply pointed, overlapping anterior half of corresponding operculum. Wings hyaline, forewing with faint milky glaze proximal to nodal line, veins brown to black, cream at node; fore wing with infuscations along crossveins r, m, r-m, and m-cu, vein CuA2, bases and apices of veins RA2, RP, M1-4, and CuA1, nodal line intersection, and outer margins of apical cells 1���7; basal membrane pale anteriorly, blackening posteriorly; basal cell orange-brown; hind wing with 6 apical cells; with continuous infuscations along proximal veins of apical cells 1���3 and outer margins of apical cells 1���6; plaga grey, covering jejum, with ovoid translucent area near distal margin. Opercula pale yellowish brown, triangulate, ridged laterally, not extending beyond posterior margin of sternite II, rounded and meeting on midline. Timbals (Fig. 2 A) with 5 long ribs; ribs 1���4 dark brown, full length and fused ventrally, thinner through dorsal half to prominent central expansion, thicker ventrally, pale brown to yellow, interspersed by three intercalary ribs; rib 5 discontinuous centrally. Timbal plate prominent when timbal plate removed; timbal dome pale brown, extending over half width of timbal, inverted distally. Apodeme pit small, slit like, conspicuous. Abdomen with tergites mainly brown, darker medially, lighter laterally, darker brown along posterior and lateral margins; tergite 1 dark brown medially with black anterior margin; timbal covers pale brown, ovoid and convex, completely covering timbals; tergite 2 with black anterior margin, brown over remainder; tergites 3���7 brown with lateral golden hairs; tergite 8 black; sternites I���VII dark brown, translucent, blackish laterally; tergite VIII black, brown along lateral third. Genitalia (Fig. 3 A���B): Pygofer trapezoid in ventral view; dorsal beak short, rounded, not extending to anal styles; distal shoulder acute, curved laterally; uncal lobe long, with inner margins separating towards apex, apex bevelled inwards, rounded at tip, gently curved in lateral view, with lateral projection short, rounded; basal lobe stout and curved inward in ventral view; aedeagus slender, short, entirely within uncus. Description of female (Fig. 1 C, D). Head and thorax similar to male, with generally darker coloration; pronotal collar olive green; mesonotum distinctly olive green with similar black patterns to male. Abdomen dark brown; tergite 1 brown; tergites 2���3 with black anterior margins, brown posteriorly; tergites 3��� 8 black-brown medially, browner laterally with distinct golden hairs; sternites I���VII dark brown to black, black mottling along anterior margins; abdominal segment 9 black, dark brown laterally, with dark green strip along dorsal midline. Ovipositor black, dark brown at tip, not extending beyond abdomen. Measurements (12 males, 8 females). Body length: male 43.4���46.8 (45.2); female 34.9���37.4 (36.0). Length of head + thorax: male 17.0���18.2 (17.4); female 15.6���16.4 (16.0). Length of abdomen: male 26.4���28.6 (27.8); female 18.4���21.0 (19.9). Fore wing length: male 50.1���52.9 (51.0); female 50.6���52.8 (51.6). Head width: male 11.3���12.2 (11.7); female 10.7���12.0 (11.6). Pronotum width: male 12.8���14.6 (13.4); female 13.2���14.5 (13.8). Abdomen width: male 14.1���16.6 (15.4); female 13.2���15.0 (14.1). Etymology. The name ��� magna ��� (Latin feminine adjective for large) is appropriate for the size of this species, the largest of the Semia species described to date. Distinguishing features. Semia magna sp. nov. is readily distinguished from all other Semia species except S. klapperichi by its larger size (body length> 43mm); the fore wing radial crossvein which is nearly right-angled to the outer margin of the fore wing; and the well developed timbal cover which is distinctly longer than the tergite 2. With these very distinguishing characteristics of the two species, S. magna and S. klapperichi can be grouped separately from other congeners under the name of 'the S. klapperichi species group'. This species group is thought to be very closely related to the Pomponia linearis species group of the genus Pomponia St��l, 1866, which has these characters and also the well-developed anterolateral pronotal collar as the common characters. Semia magna sp. nov. can be distinguished from S. klapperichi by the following combination of characters: less pronounced black infuscations on fore and hind wings compared with very thick infuscations in S. klapperichi; inner area of pronotum largely light coloured compared with mostly black inner area in S. klapperichi; uncal lobes with distinctly narrowed apices, with rounded apices in S. klapperichi; majority of mesonotum pale brown, black in S. klapperichi; ventral postclypeus and anteclypeus with conspicuous pale brown markings, which are mostly black in S. klapperichi; opercula light brown, dark brown to black on S. klapperichi. Distribution, habitat, and seasonality. This species has been found on Ngoc Linh Nature Reserves in Kon Tum Province and 350 km north at Bach Ma National Park, Thua Thien Hue Province. At both locations, the species co-exists with S. spiritus sp. nov. and S. pallida sp. nov. on Bach Ma National Park (Fig. 4). Specimens have been collected from March to May., Published as part of Emery, David L., Lee, Young June & Pham, Hong-Thai, 2017, Descriptions of four new species of Semia Matsumura (Hemiptera: Cicadidae: Psithyristriini) from Vietnam, with a key to the species of Semia, pp. 153-166 in Zootaxa 4216 (2) on pages 154-159, DOI: 10.5281/zenodo.230748

Key to the males of the species of Semia 1. Fore wing radial crossvein not parallel but about right-angled to outer margin of fore wing; timbal cover well developed, dis- tinctly longer than tergite 2.................................................... 2 (S. klapperichi species group) - Fore wing radial crossvein nearly parallel to outer margin of fore wing; timbal cover about as long as or shorter than tergite 2....................................................................................................3 2. Inner area of pronotum largely light coloured; uncal lobes with rounded apices........................... S. klapperichi - Inner area of pronotum mostly black; uncal lobes with distinctly narrowed apices..................... S. magna sp. nov. 3. Body without median fascia continued from head to abdomen; uncal lobes without multiple spines.................... 4 - Body with median black to dark brown fascia continued from head to abdomen; uncal lobes with multiple spines on each lobe............................................................................................. S. spinosa 4. Uncal lobe without long extended projection protruding laterally from underside of uncus............................ 5 - Uncal lobe with long extended projection protruding laterally from underside of uncus, being laterally parallel to uncal lobe........................................................................................... S. gialaiensis 5. Hind wing with continuous infuscation along hind margins of apical cells 1–6.................................... 6 - Hind wing without continuous infuscation along hind margins of apical cells..................................... 7 6. Uncal lobes comparatively slender, with length about four times the width, tapering to subapex, with apex roundly expanded and slightly twisted longitudinally, in ventral view................................................. S. majuscula – Uncal lobes comparatively broad, with length about three times the width, and parallel to each other from bases to truncated apices, in ventral view......................................................................... S. tibetensis 7. Uncal lobes very long, with length more than four times the central width........................................ 8 - Uncal lobes short, with length less than four times the central width............................................. 10 8. Abdominal tergite 8 covered with white pollinosity; uncal lobe in similar width from base to apex.....................9 - Abdominal tergite 8 without white pollinosity; uncal lobe with much widened apex.......................... S. lachna 9. Fore wing ulnar cell 2 comparatively long, with length more than four times the width; uncal lobes widened apically in lateral view; abdominal tergites 6 and 7 seldom covered with white pollinosity................................. S. watanabei - Fore wing ulnar cell 2 comparatively short, with length less than four times the width; uncal lobes narrowed apically in lateral view; abdominal tergites 6 and 7 thickly covered with white pollinosity............................... S. hainanensis 10. Timbal cover moderately or well developed, with its anterior margin at similar level of or more anteriorly positioned than antero-median margin of tergite 2........................................................................ 11 - Timbal cover rudimentary, very short, with its anterior margin far behind (far more posteriorly positioned than) antero-median margin of tergite 2.................................................................................. 12 11. Abdomen shorter than 1.6 times the length of head and thorax together; uncal lobes triangulate with narrowed apices, in ventral view.............................................................................. S. spiritus sp. nov. - Abdomen longer than 1.8 times the length of head and thorax together; uncal lobes wide, wider in apices than in bases, in ventral view............................................................................... S. pallida sp. nov. 12. Uncal lobes comparatively long, with length more than twice the width, nearly parallel to each other........................................................................................................ S. albusequi sp. nov. - Uncal lobes very short, with length much less than twice the width, widely separated from each other........ S. brevidilata

Semia pallida sp. nov. (Figs 2 C, 3E���F, 6) Type material. Holotype: ♂, Lam Dong Prov., Bidoup Nui Ba National Park, 2000 m, 12��06���43���N, 108��57���53���E, 10 V 2013 (at light) (VNMN; E-00122). Paratypes: 2♂♂, 1♀, same data as holotype (AM, K493590-92); 3♂♂, 1♀, same location, 20 III 2016; 3♂♂, Khanh Hoa Prov., Hon Ba Nature Reserves, 1500 m, IV 2013 (VNMN; E- 00123-129); 2♂♂, 4♀♀, Thua Thien Hue Prov., Bach Ma National Park, 1400 m, 16��11���51���N, 107��51���36���E, 20 III 2016 (at light); 1♂, Nghe An Prov., Pu Mat National Park, 1200 m, 19��02���21���N, 104��45���50���E, 20 IV 2014 (DE). Other Material. 4♂♂, Lam Dong Prov., Lac Duong, Bidoup Nui Ba National Park, 2000 m, 12��06���43���N, 108��57���53���E, 20 IV 2016, (at light) (DE). Description of male ( Figs 6 A���B). Entire body light grey-brown and covered with whitish pollinosity (Fig. 6 A). Head mostly light grey with black markings between compound eyes and ocelli, black median line, and black spot posterior to eye; ocelli light orange; postclypeus brown dorsally, black medially; ventral postclypeus pale orange with dark brown horizontal striations along transverse grooves mid-anteriorly, brown midline, thin black surrounds, pale brown patch over remainder, and prominent white pollinosity spread; anteclypeus mainly brown; gena and lorum light brown, with medial margin black; rostrum pale brown, black at tip, extending to mid-sternite II; compound eyes reddish brown; antennae light brown; supra-antennal plates light brown. Thorax: Pronotum mostly pale brown-ochraceous with prominent white pillinosity, midline brown markings with black borders; parapsidal sutures black; pronotal collar evenly orange-brown, expanded laterally. Mesonotum olive-brown; submedian sigella margins black, dull black pattern radiating over lateral sigella; scutal depressions black; cruciform elevation light brown, with lateral depressions darker brown and white pollinosity. Metanotum light brown. Legs predominantly light brown; coxa brown; trochanters brown; femora light brown, black at extremities, with primary spine small, obtuse; tibiae similarly patterned to femora, black at centre and extremities, with spines black; tarsi brown anteriorly, blackish at tips; claws black. Meracanthus narrow, pale brown, sharply pointed, overlapping anterior half of corresponding operculum. Wings hyaline; costal vein green-brown proximal to node, olive green distally; veins cream to brown, with white pollinosity proximal to intermodal line; fore wing with infuscations along crossveins r, m, r-m, and m-cu, vein CuA2, junction of veins CuP+1A and 2A+3A, bases and apices of veins RA2, RP, M, M1-4, and CuA1, nodal line intersection, and outer margins of apical cells 1���7; basal membrane uniformly light grey; basal cell translucent; hind wing with 6 apical cells; with infuscations along proximal veins of apical cells 2���3, absent from margins of apical cells 1���6; plaga white, covering jejum, with translucent area near distal margin. Opercula light brown under white pillosity, with black lateral margins; spatulate, not extending beyond posterior margin of sternite II, rounded medially, almost meeting on midline. Timbals (Fig. 2 C) with 5 long ribs; ribs 1���4 dark brown, full length, fused ventrally, thinner through dorsal half to prominent central expansion, thicker ventrally, pale brown to yellow, interspersed by four intercalary ribs; rib 5 discontinuous centrally. Timbal plate prominent; timbal dome pale brown, extending over half width of timbal, inverted distally. Apodeme pit slit-like, conspicuous. Abdomen with tergites mainly brown, darker medially, lighter laterally, with black longitudinal stripe along lateral margins; tergite 1 brown;, timbal covers pale brown, triangulate and convex, covering around two-thirds of timbals; tergites 2���7 brown; tergite 8 dark brown; sternites I���VI pale brown, translucent, white pollinosity over lateral half; sternite VII dark brown to black. Genitalia (Fig. 3 E���F). Pygofer rectangular in ventral view; dorsal beak short, pilose, rounded, not extending posteriorly to anal styles; distal shoulder inconspicuous, rounded, pilose; uncal lobe dark brown posteriorly, becoming pale brown towards apex, short, with inner margins becoming separated from base to apex and lateral margins rounded at apex, bevelled posteriorly towards midline to form a ���V���, strongly curved in lateral view; basal lobe stout, solidly curved inward in ventral view. Aedeagus short, entirely within uncus. Description of female (Fig. 6 C���D). Head and thorax similar to male, with generally darker and greenish coloration; pronotal collar olive green, ampliate laterally; mesonotum olive green with similar black patterns to male. Abdomen dark brown; tergites brown, black on lateral margins; tergite 1 ochraceous; tergites 2���8 ochraceous, darker medially, lighter laterally; sternite I pale brown, sternites II ���VII light brown becoming darker posteriorly; abdominal segment 9 ochraceous, with black stripe adjacent to midline and brown dorsal midline. Ovipositor dark brown, black at tip, not extending beyond abdomen. Measurements (12 males, 6 females). Body length: male 33.0���36.6 (35.3); female 26.2���31.0 (28.3). Length of head + thorax: male 12.4���13.8 (13.4); female 12.2���14.5 (13.4). Length of abdomen: male 20.7���23.0 (21.9); female 14.0���16.51 (14.9). Fore wing length: male 34.4���41.2 (38.9); female 38.2���44.5 (40.4). Head width: male 8.0���9.4 (8.6); female 8.3���9.2 (8.9). Pronotum width: male 9.3���10.9 (10.0); female 9.0���10.4 (9.7). Abdomen width: male 11.5���12.7 (12.0); female 10.1���11.8 (11.0). Etymology. The name ��� pallida ��� (Latin feminine adjective) refers to the pale grey coloration of this species, emanating in part from the extensive white pollinosity over the entire body, giving it a ���ghostly��� appearance. Distinguishing features. The grey coloration of S. pallida sp. nov. distinguishes it from all other Semia species which are pale, olive, or dark brown. Compared to those Semia species lacking continuous infuscations along hind margins of the apical cells 1���6 (S. brevidilata, S. hainaensis, S. spinosa, S. spiritus sp. nov., S. albusequi sp. nov., and S. watanabei), S. pallida sp. nov. (body length 33���37 mm) is smaller than S. hainanensis (body length> 37 mm) and larger than S. spinosa and S. albusequi sp. nov. (body lengths S. pallida sp. nov. is slightly larger than S. gailaiensis (body length 32.8���33.6 mm), the opercula of S. gailaiensis meet in the midline while those of S. pallida sp. nov. are clearly separated. The rounded ventral apices on the uncal lobes of S. brevidilata, the lateral ���spine-like��� projection from the uncus of S. spinosa, the ornate lateral projections of the uncal apices in S. lachna, the darker lateral uncal projections on S. gialaiensis, and the rectangular appearance of the apical lobes of S. watanabei contrast with the ���V-shaped��� angulation produced at the apices of the uncal lobes on S. pallida sp. nov. Semia pallida sp. nov. is distinguished from the single male S. brevidilata by its narrow head width (S. brevidilata) and presence of lateral black stripes on the tergites, which are lacking in S. brevidilata. Semia pallida sp. nov. also lacks the white pollinosity displayed on the terminal tergites of S. watanabei. Distribution, habitat, and seasonality. This species has a wide distribution along the east coast of Vietnam from Bidoup Nui Ba National Park in the south to Pu Mat National Park in northern Vietnam (Fig. 4). Specimens have been collected between March and May., Published as part of Emery, David L., Lee, Young June & Pham, Hong-Thai, 2017, Descriptions of four new species of Semia Matsumura (Hemiptera: Cicadidae: Psithyristriini) from Vietnam, with a key to the species of Semia, pp. 153-166 in Zootaxa 4216 (2) on pages 161-163, DOI: 10.5281/zenodo.230748

Semia spiritus sp. nov. (Figs 2 B, 3C���D, 5) Type material. Holotype: ♂, Kon Tum Prov., Ngoc Linh Nature Reserves, 1900 m, 15��00���39���N, 107��57���53���E, III 2016; (typed ink label) (VNMN; E-00112). Paratypes: 2♂♂, Lam Dong Prov., Bidoup Nui Ba National Park, 2000 m, 12��06���43���N, 108��57���53���E, 10 V 2013; 6♂♂, 1♀, same location as holotype, 20 IV 2016 (VNMN; E-113-121), 2♂♂, same data as holotype (AM, K493593-94); 1♀, Lai Chau Prov., 1600 m, 22��18' 42" N 102��09'22" E, 18 V 2015; 2♂♂, Thua Thien Hue, Bach Ma National Park, 1400 m, 16��11���51���N, 107��51���36���E, 25 IV 2016 (DE). Other Material. 5♂♂, Kon Tum Prov., Ngoc Linh Nature Reserves, 1900 m, 15��00���39���N, 107��57���53���E, IV 2014 (DE); 7♂♂, same location, 15 V 2015 (DE); 2♂♂, Lam Dong, Bidoup Nui Ba National Park, 2000 m, 12��06���43���N, 108��57���53���E, 10 V 2013 (DE); 2♂♂, Thua Thien Hue, Bach Ma National Park, 1400 m, 16��11���51���N, 107��51���36���E, 25 IV 2016 (DE). Description of male ( Figs 2 B, 3C���D, 5A���B). Head mostly brown, brown anteriorly, with black markings around ocelli; ocelli pink; postclypeus brown, with black ���V��� marking dorsally, black horizontal striations along transverse grooves mid-anteriorly, pale brown patch on either side of midline over central third, and prominently shiny black marking over posterior third; anteclypeus mainly brown; gena and lorum dark brown; rostrum brown, black at tip, extending to mid-sternite II; compound eyes dark brown; antennae light brown; supra-antennal plates brown. Thorax: Pronotum mostly pale brown-ochraceous, with black markings outlining anterior brown stripe and posterior triangle marking on midline, darker brown patches laterally along ridges between parapsidal sutures, and black strip along lateral border; pronotal collar evenly brown-red to ochraceous but greenish on lateral margins, with median black spot on posterior margin, expanded laterally. Mesonotum ranging from light to olive-brown, with black median stripe, black patterned motif on submedian sigella on most specimens, but central area olive brown on majority of specimens from Bach Ma Mountain, with dull black pattern radiating over lateral sigilla and reddish brown patch along lateral margin; scutal depressions black; cruciform elevation light brown, with lateral depressions dark and golden hairs sparsely spread. Metanotum dark brown. Legs predominantly light brown; coxa brown with anterior black stripes; trochanters brown; femora with small black markings at ends, with primary spine erect, brown; tibiae similarly patterned to femora, but black around tarsi; tarsi brown anteriorly, blackish at tips; claws dark brown. Meracanthus narrow, pale brown, sharply pointed, overlapping anterior half of corresponding operculum. Wings hyaline, veins cream to brown, cream at node; fore wing with infuscations along crossveins r, m, r-m, and m-cu, vein CuA2, bases and apices of veins RA2, RP, M, M1-4, and CuA1, nodal line intersection, and outer margins of apical cells 1���7; basal membrane uniformly grey; basal cell translucent; hind wing with 6 apical cells; with continuous infuscations along proximal veins of apical cells 1���3 and light infuscations along outer margins of apical cells 1���3 in most specimens; plaga grey, covering jejum, with translucent area near distal margin. Opercula pale yellowish brown, with black lateral margins; spatulate, not extending beyond posterior margin of sternite II, rounded medially, almost meeting on midline. Timbals (Fig. 2 B) with 5 long ribs; ribs 1���4 dark brown, full length fused ventrally, thinner through dorsal half to prominent central expansion, thicker ventrally, pale brown to yellow, interspersed by three intercalary ribs; rib 5 discontinuous centrally. Timbal plate prominent; timbal dome pale brown, extending over half width of timbal, inverted distally. Apodeme pit small, slit like, conspicuous. Abdomen with tergites mainly brown, darker medially, lighter laterally, with black longitudinal stripe near lateral margins; tergite 1 brown; timbal covers pale brown, triangulate and convex, completely covering timbals; tergites 2���7 brown; tergite 8 dark brown; sternites pale brown, translucent. Genitalia (Fig. 3 C���D): Pygofer trapezoid in ventral view; dorsal beak short, rounded, protruding upwards in lateral view, not extending to elongated anal styles; distal shoulder acute, directed and curved laterally; uncal lobe long, with apposed inner margins, with lateral margins angled anteriorly towards rounded apex, spatulate and gently curved in lateral view; basal lobe stout and curved inward in ventral view. Aedeagus short, concealed within uncus. Description of female (Fig. 5 C���D). Head and thorax similar to male, with generally darker and greenish coloration; pronotal collar olive green, ampliate laterally; mesonotum distinctly olive green with similar black patterns to male. Wings similar to male, darker infuscations along posterior margins of apical cells 1���6. Abdomen dark brown; tergite 1 brown; tergites 2���3 with black anterior margins, brown posteriorly; tergites 3��� 8 black-brown medially, browner laterally with distinct golden hairs; sternites I���VII dark brown to black, black mottling along anterior margins; abdominal segment 9 black, dark brown laterally, green along dorsal midline. Ovipositor dark brown, not extending beyond abdomen. Measurements (12 males, 2 females). Body length: male 35.2���39.1 (36.8); female 27.8���31.0 (29.9). Length of head + thorax: male 13.8���16.0 (14.7); female 13.4���14.9 (14.2). Length of abdomen: male 19.6���23.8 (22.0); female 14.4���16.1 (15.2). Fore wing length: male 39.6���44.8 (42.0); female 41.5���44.5 (42.9). Head width: male 8.3���9.3 (8.9); female 8.8���9.2 (9.0). Pronotum width: male 9.9���11.1 (10.5); female 9.0���9.4 (9.2). Abdomen width: male 12.0���13.9 (12.8); female 11.8���11.8 (11.8). Etymology. The name ��� spiritus ��� (Latin masculine noun for spirit) derives from the Vietnamese ��� Linh ��� (spirit, soul), reflecting the predominant location of this species on Ngoc Linh Nature Reserves. Distinguishing features. Males of S. spiritus sp. nov. lack the continuous infuscation along the posterior margin of the hind wing apical cells 1���6, so are readily distinguished from S. klapperichi, S. majuscula, S. magna sp. nov., and S. tibetensis. Semia spiritus sp. nov. (body length> 35.2 mm) is larger than S. spinosa (body length S. gialaiensis (body length S. albusequi sp. nov., (body length S. brevidilata (body length 34 mm). Semia spiritus sp. nov. lacks the distinct white pollinosity present on the tergite 8 of S. watanabei. The large timbal covers on S. spiritus sp. nov. contrast with rudimentary covers which leave the majority of timbals exposed in S. lachna, S. brevidilata, and S. watanabei. The dark brown coloration of S. spiritus sp. nov. contrasts with the pallor of S. pallida sp. nov. and the olive-brown markings on S. hainanensis . Semia spiritus sp. nov. also lacks the black markings seen laterally on the pronotal collar and tibiae of S. hainanensis . Distribution, habitat, and seasonality. This species has been found in Central Vietnam at Bach Ma National Park and Ngoc Linh Nature Reserves and 500 km southwards at Bidoup Nui Ba National Park (Fig. 4). Specimens have been collected between March and May., Published as part of Emery, David L., Lee, Young June & Pham, Hong-Thai, 2017, Descriptions of four new species of Semia Matsumura (Hemiptera: Cicadidae: Psithyristriini) from Vietnam, with a key to the species of Semia, pp. 153-166 in Zootaxa 4216 (2) on pages 159-161, DOI: 10.5281/zenodo.230748