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We use DNA sequence data to generate the first phylogenetic hypothesis for the weevil tribe Lymantini. These are leaf litter inhabiting beetles generally regarded as restricted to the New World and taxonomically arranged in two subtribes, 11 genera and some 150 named species. An additional genus of questionable affinities to the tribe, Devernodes Grebennikov, 2018, has five described species in Southeastern Asia. All these beetles are flightless and some have eyes reduced in size or absent, traits normally associated with limited dispersal capacity. We performed a phylogenetic analysis of 153 terminals (50 of them belong to Lymantini representing Devernodes and all but three named genera) based on 4,174 bp alignment of one mitochondrial (cox1) and two nuclear fragments (ITS2 and 28S). We find that both Lymantini subtribes Lymantina and Caecossonina are monophyletic, the latter sister to the amphi-Atlantic tribe Anchonini. The Asian genus Devernodes is deeply nested among American Lymantina. The clade of Anchonini plus Lymantini is consistently recovered outside of the CCCMS clade of “higher” weevils (Curculioninae, Conoderinae, Cossoninae, Molytinae and Scolytinae). We hypothesize that the polished head capsule of adult beetles is an apomorphy of Anchonini and Lymantini, the 8-segmented antennal funicle is an apomorphy of Anchonini plus Caecossonina. We attribute the origin of the currently observed amphi-Pacific distribution of Lymantina to normal ecological dispersal facilitated by the warmer periods of the Cenozoic such as the Eocene, and by presently submerged Arctic land bridges. Using parsimony we hypothesize a North American origin for the Anchonini plus Lymantini crown group, as well as that of Lymantina. We argue that Bronchotibia adunatus Poinar & Legalov, 2021, a Dominican amber adult weevil fossil, is not a member of Lymantini and re-classify it as Curculionidae incertae sedis. We present an image gallery of 28 Lymantini specimens to document the morphological diversity of the tribe. We hypothesize the existence of unnamed American genera of Lymantina and make public the DNA-barcode dataset of 89 Lymantini specimens.

Melanophilharmostes tuber Grebennikov, new species Figs 1���5. Type locality. Cameroon, Mt. Cameroon, 4.0681��N, 9.0717��E, 233m. Type material. Holotype (deposited in the Canadian National Collection of Insects, Arachnids, and Nematodes, Ottawa, Canada), female, ��� CAMEROON, Mt. Cameroon, 4.0681, 9.0717, 233m, 28.xii.2015, sift. CM09, V. Grebennikov.���, ���CNCCOLVG00009477���. Paratype (deposited in the Canadian National Collection of Insects, Arachnids, and Nematodes), not sexed, ��� CAMEROON, Mt. Kupe, 4.8193, 9.7075, 1525m, 29.xii.2015, sift. CM13, V. Grebennikov.���, ���CNCCOLVG00009488���. Description. Holotype maximal body length (in rolled position, Fig. 1A): 2.5 mm (paratype: 2.5 mm), maximal body width (Fig. 1): 2.2 mm (paratype: 2.3 mm). Head distinctly convex, clypeal disc gibbose. Interocular bridge completely subdividing each eye into smaller dorsal and larger ventral parts (Fig. 2E); antenna with seemingly 10 antennomeres (Fig. 2F, not dissected to make exact count); head, pronotum, and elytra with deep and dense microsculpture (horseshoe-shaped and comma-shaped), as well as with sparse, short, and distinct pilosity; surface of pronotum uneven, with bumps and depressions (Fig. 2A); each elytron with notable lateral longitudinal sharp carina interrupted in posterior part (Figs 2B, D); hind wings fully developed; distal fork of vein CuA present; distal expansion of vein MP 1+2 cannot be verified using single available wing (Fig. 2D). Male genitalia unknown. DNA data of holotype: MW899169 (COI) and MW845067 (28S); DNA data of paratype: MW899176 (COI), MW845087 (ITS2), and MW845074 (28S). Diagnosis. Within the Synarmostes group of four Afrotropical genera (including Madagascar and Comoros), this species can be immediately recognised by any of its four diagnostic characters: (1) head distinctly convex with clypeal disc gibbose (Fig. 2F), (2) bumpy pronotum, (3) notably developed longitudinal carina on each elytron, and (4) deep and dense microsculpture on the head, pronotum, and elytra. Distribution. This species is known from two Cameroonian localities about 100 km apart: Mt. Cameroon (233 m) and Mt. Kupe (1523 m). Natural History. Both specimens were collected by sifting forest leaf litter in rainforest. Derivation of name. The species epithet is the Latin noun meaning ���a hump, bump, swelling, protuberance���, with reference to the peculiarly bumpy pronotum of the new species distinguishing it from all its congeners; it is used as a noun in apposition., Published as part of Grebennikov, Vasily V., 2022, Phylogenetic placement of a new Melanophilharmostes Paulian, 1968 pill scarab (Coleoptera: Hybosoridae: Ceratocanthinae) from Cameroon: molecular results decipher misleading morphology, pp. 441-450 in Zootaxa 5087 (3) on page 446, DOI: 10.11646/zootaxa.5087.3.3, http://zenodo.org/record/5826878

Melanophilharmostes Paulian, 1968 Type species: Philharmostes (Melanophilharmostes) zicsii Paulian, 1968, by original designation. Composition and distribution: 17 named species (including the one described below) from sub-Saharan Africa and the Gulf of Guinea island of Annob��n. Monophyly, sister group, and morphological synapomorphies: the genus formed a moderately supported clade in both the morphology-based (Ballerio 2016) and DNA-based (Fig. 5) analyses. Its three morphological synapomorphies (Ballerio 2016) are (1) dorsal body surface with setae, (2) larger elytral punctures are ���large horseshoe-shaped���, and (3) elytra with a lateral carina. In both analysis the genus formed either moderately (Ballerio 2016) or weakly-supported (Fig. 5) sister group to the genus Pseudopterorthochaetes; the only morphological synapomorphy of both genera is that the dorsal puncture on the head is shallow (not deep; Ballerio 2016). Two hind wing characters seem to additionally support the Melanophilharmostes and Pseudopterorthochaetes clade (presence of distal fork in vein CuA and presence of the distal expansion of vein MP 1+2; characters 15/1 and 16/1, respectively, in Ballerio 2016; see also Fig. 2D); however, ambiguity remains due to the widespread loss of hind wings within this clade. The genus Pseudopterorthochaetes was recovered either as a moderately-supported (Ballerio 2016) or strongly-supported clade (Fig. 5) with a single morphological autapomorphy: the inner mesotibial apical spur in males is parallel to the tibial axis (not bent inwards at a right angle, as in Melanophilharmostes and most other pill scarabs)., Published as part of Grebennikov, Vasily V., 2022, Phylogenetic placement of a new Melanophilharmostes Paulian, 1968 pill scarab (Coleoptera: Hybosoridae: Ceratocanthinae) from Cameroon: molecular results decipher misleading morphology, pp. 441-450 in Zootaxa 5087 (3) on page 445, DOI: 10.11646/zootaxa.5087.3.3, http://zenodo.org/record/5826878

Aside: Cleonini are not universally flightless Regardless of whether the tribe is monophyletic or not, some Cleonini species do actively fly. This is, however, a poorly documented and rarely observed phenomenon, which misled Arzanov & Grebennikov (2017) into the erroneous conclusion that all Cleonini are flightless. Active flight has been reported for at least three economically important species: Asproparthenis punctiventris (Germar, 1823) (e.g., Orlachiova 1956; Auersch 1961; Tóth et al. 2006), Bothynoderes affinis (Schrank, 1781) (Kokovikhin 1966), and Temnorhinus brevirostris (Gyllenhal, 1834) (Brunner 1959). Meregalli (2005) reported flight in Eurycleonus talamellii Meregalli, 2005. One of us (BI) observed in Mongolia active flight of, likely, Conorhynchus conirostris Gebler, 1829. Robert Stejskal (personal communication) observed in central Europe active flight of Bothynoderes affinis, Asproparthenis punctiventris, Cleonis pigra, and Cyphocleonus dealbatus Gmelin, 1790., Published as part of Volovnik, Semyon V., Boldgiv, Bazartseren, Iderzorig, Badamnyambuu, Khobrakova, Larisa Ts., Kolov, Sergey V., Rudykh, Sergey G., Zabaluev, Ilya A. & Grebennikov, Vasily V., 2021, The first molecular phylogeny of the weevil subfamily Lixinae (Coleoptera Curculionidae) casts doubts on the monophyly of its tribes, pp. 201-220 in Zootaxa 5026 (2) on page 216, DOI: 10.11646/zootaxa.5026.2.3, {"references":["Arzanov, Yu. G. & Grebennikov, V. V. (2017) Cleonini (Coleoptera: Curculionidae: Lixinae) are monophyletic and flightless: tribe overview, rampant adult homoplasy and illustrated global diversity. Zootaxa, 4329 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4329.1.1","Orlachiova, K. A. (1956) Faktory, opredelyayushchie perelioty zhukov sveklovichnogo dolgonosika. In: Vasiliev, V. P. (Ed.), Sveklovichnyi Dolgonosik i Borba s Nim. Sbornik Nauchnykh Trudov Instituta Entomologii i Fitopatologii AN USSR. Vol. 7. Akademia Nauk USSR, Kiev, pp. 158 - 194 [in Russian].","Auersch, O. (1961) Zur Kenntnis des Rubenderbrusslers (Bothynoderes punctiventris Germ.). Teil 2. Zeitschrift fur Angewandte Entomologie, 49, 50 - 77. https: // doi. org / 10.1111 / j. 1439 - 0418.1961. tb 02855. x","Toth, M., Ujvary, I., Sivcev, I., Imrei, Z., Szarukan, I., Farkas, O., Gomory, A., Gacs-Baitz, E. & Francke, W. (2006) An aggregation attractant for the sugar-beet weevil, Bothynoderes punctiventris. Entomologia Experimentalis et Applicata, 122, 125 - 132 https: // doi. org / 10.1111 / j. 1570 - 7458.2006.00502. x","Kokovikhin, A. I. (1966) K biologii Chromoderus fasciatus Mull. (Coleoptera, Curculionidae) v stepnoy zone Omskoi oblasti. In: Cherepanov, A. I. (Ed.), Fauna i Ekologija Chlenistonogikh Sibiri. Nauka - Sibirskoe otdelenie, Novosibirsk, рp. 29 - 32. [in Russian]","Brunner, Yu. N. (1959) Samarkandskyi sveklovichnyi dolgonosik. In: Savchenko, E. N. (Ed.), Sveklovodstvo. Vol. 3. Gosselkhozizdat USSR, Kiev, pp. 135 - 136. [in Russian]","Meregalli, M. (2005) Eurycleonus talamellii n. sp. of Cleonine weevil from the Moroccan desert (Coleoptera: Curculionidae: Lixinae: Cleonini). Zootaxa, 1053 (1), 23 - 34. https: // doi. org / 10.11646 / zootaxa. 1053.1.2"]}

Sampling of Lixinae Most herein analysed specimens of Lixinae were sampled during our fieldwork in Kazakhstan, Mongolia, Russia and Ukraine; the remaining specimens were accumulated through a network of correspondents (see Acknowledgements). All Cleonini specimens sampled by us were either hand-picked during the daytime from under items lying on the ground (mainly dry dung of domesticated ungulates, Fig. 2L)), or caught in unbaited pitfall traps; a few specimens were picked when walking on the ground or reared from root galls (Fig. 2J). Conversely, all Lixini and Rhinocyllini specimens were swept or hand-picked from vegetation. All specimens sequenced herein (including the outgroup) are deposited in the Canadian National Collection of Insects, Arachnids, and Nematodes (Ottawa). Images and geographic data of each specimen reported in this work, either of Lixinae or non-Lixinae, are available online in the Barcode of Life Data System (BOLD, Ratnasingham & Hebert 2007; dx.doi.org/10.5883/DS-VGDS21)., Published as part of Volovnik, Semyon V., Boldgiv, Bazartseren, Iderzorig, Badamnyambuu, Khobrakova, Larisa Ts., Kolov, Sergey V., Rudykh, Sergey G., Zabaluev, Ilya A. & Grebennikov, Vasily V., 2021, The first molecular phylogeny of the weevil subfamily Lixinae (Coleoptera Curculionidae) casts doubts on the monophyly of its tribes, pp. 201-220 in Zootaxa 5026 (2) on page 207, DOI: 10.11646/zootaxa.5026.2.3, {"references":["Ratnasingham, S. & Hebert, P. D. N. (2007) BOLD: The Barcode of Life Data System (www. barcodinglife. org). Molecular Ecology Notes, 7, 355 - 364. https: // doi. org / 10.1111 / j. 1471 - 8286.2007.01678. x"]}

Tazarcus gen. n. http://zoobank.org/ urn:lsid:zoobank.org:act: 09E4CE1E-BFE6-47E9-89BD-2A529F652DA4 Type species. Tazarcus aeaea sp. nov., here designated. Diagnosis. Known representatives of the new genus can be recognized among weevils by the unique combination of the following easily observed characters: body small (combined dorsal length of pronotum and elytra 2.0��� 3.4 mm); ratio of combined pronotal and elytral length in dorsal view to the maximal body width 1.6 x or less; elytra laterally rounded, without shoulders; rostrum relatively short and straight; eyes present, small and flat; antennal funicle with seven segments; anterior margin of prothorax laterally without postocular lobes; prosternum with canal laterally delimited by ridges tapering anteriad and linking anterior edge of prosternum with anterior edge of procoxae (thus canal only present in front of procoxae, not behind); procoxae separate; metanepisternal sutures absent, their place occupied by longitudinally oriented metapleural ridge bearing on its crest a line of short and robust small setae (sclerolepidia?); femora without teeth-like projections; tarsal claws small, separate and not appendiculate; elytra with nine longitudinal punctate striae; legs and rostrum dorsally covered with dense matt whitish pilosity; hind wings absent. Adults of the new genus (particularly those of the type species) habitually resemble comparably rounded, flightless and globular members of the distantly related (McKenna et al. 2009) genus Ocladius Sch��nherr, 1825 (Meregalli & Colonnelli 2006). At least three Ocladius species have been reported from the highlands of northern Tanzania (Aurivillius 1910), one of them (O. seriatus Faust, 1896) from ���Usambara���. Adults of Tazarcus are externally distinguishable from those of Ocladius by the straight, shorter and thicker rostrum lacking longitudinal keels and furrows.Adults of all other genera of flightless East African Molytinae have a more slender body (a ratio of dorsal combined length of pronotum and elytra to the maximal body width of> 1.6x). Four additional diagnostic characters aid in the identification of Tazarcus: (1.) small body size, combined length of pronotum and elytra in dorsal view 2.0��� 3.4 mm; (2.) a short and straight rostrum; (3.) small flat eyes; and (4.) a short prosternal canal, tapering anteriad. Species composition and distribution. This genus consists of two herein described allopatric species restricted to the EAM Chain in Tanzania. The two species can be easily separated by the form of the elytral punctation (Figs. 1A, B vs Fig. 2A, B). Biology. Adult specimens of the new genus were found by sifting forest floor litter in wet Afromontane forests at elevations between 1576 m and 2318 m. Host plants, immature life stages, parasites or any other biological aspects are unknown. Etymology. Toponymic, formed by abbreviating ���Tanzanian Arc���, with reference to the EAM occurring predominantly in Tanzania; gender masculine., Published as part of Grebennikov, Vasily V., 2020, Tazarcus, a new phylogenetically unplaced genus of two flightless weevils with metapleural ridge from the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae: Molytinae), pp. 421-434 in Zootaxa 4766 (3) on pages 425-426, DOI: 10.11646/zootaxa.4766.3.2, http://zenodo.org/record/3765549, {"references":["McKenna, D. D., Sequeira, A. S., Marvaldi, A. E. & Farrell, B. D. (2009) Temporal lags and overlap in the diversification of weevils and flowering plants. Proceedings of the National Academy of Sciences USA, 106, 7083 - 7088. https: // doi. org / 10.1073 / pnas. 0810618106","Meregalli, M. & Colonnelli, E. (2006) The genus Ocladius Schonherr, 1825 (Coleoptera: Curculionoidea: Curculionidae) in the Arabian subregion, with description of six new species. Fauna of Arabia, 21, 251 - 306.","Aurivillius, C. (1910) 7. Coleoptera. 21 Curculionidae. In: Sjostedt, Y. (Ed.), Wissenschaftliche Ergebnisse der Schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906, unter Leitung von Prof. Dr. Yngve Sjostedt. Volume 1, Abteilung 1 - 7. P. Palmquists Aktiebolag, Stockholm, pp. 403 - 435."]}

Tazarcus aeaea sp. n. Figs 1, 4, 5 http://zoobank.org/ urn:lsid:zoobank.org:act: 3EA886A2-BA5C-4F0A-AA07-49558CD9A936 Type locality. Tanzania, West Usambara Mts., Mkusu Forest, S4.76585�� E38.36022��, 1576 m. Description. Length of pronotum and elytra in dorsal view 2.0��� 2.5 mm (n=7); lateral body contour in dorsal view not constricted between pronotum and elytra; lateral sides of posterior half of pronotum and elytra at middle evenly curved; disc of pronotum completely covered by dense whitish pilosity; procoxae separated by distance greater than half their length; elytral punctures on striae small, their diameter not greater than 0.2 of width of interstriae; elytral interstriae with row of prominent appressed scales as long as width of interstriae; male genitalia: Figs 5 E���F; DNA: Table 2. Geographic variability. Adults of the new species from South Pare and West Usambara populations appear morphologically indistinguishable, even though genetically distinct (Fig. 4). Material examined. Holotype (CNC), male, ��� TANZANIA, W. Usambara Mts., Mkusu For., S4.76585�� E38.36022��, 1576m, 14.i.2013, sift47, V.Grebennikov���, ���CNCCOLVG00005480���. Paratypes (CNC), 3: same data as holotype, specimens 5479, 9337, 9338. Additional material: 4 (CNC): in ethanol, same data as holotype; 1 (in collection of D. McKenna): same data as holotype; 1 (CNC): ��� TANZANIA, South Pare Mts., Chome For., S4.26865�� E37.92944��, 2318m, 3.i.2013, sift36, V.Grebennikov���, 2 (CNC): ��� TANZANIA, South Pare Mts., Chome For., S4.26945�� E37.92814��, 2239m, 3.i.2013, sift37, V.Grebennikov���. Distribution. This species is known from two nearby EAM localities in Tanzania, South Pare and West Usambara. Etymology. The species epithet is a Latinized Greek mythical name of Aeaea, an island of the goddess Circe, who is known for transforming her offenders into animals, and it is a noun in apposition., Published as part of Grebennikov, Vasily V., 2020, Tazarcus, a new phylogenetically unplaced genus of two flightless weevils with metapleural ridge from the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae: Molytinae), pp. 421-434 in Zootaxa 4766 (3) on page 426, DOI: 10.11646/zootaxa.4766.3.2, http://zenodo.org/record/3765549

Grebennikov, Vasily V. (2020): Tazarcus, a new phylogenetically unplaced genus of two flightless weevils with metapleural ridge from the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae: Molytinae). Zootaxa 4766 (3): 421-434, DOI: 10.11646/zootaxa.4766.3.2

Agrilus draco sp. nov. (Fig. 3A) Description of holotype. Body length: 17.2 mm. BODY. Shape: cuneiform, Build: robust, Color (dorsally): unicolored. HEAD. Medial impression: deep. Vertex. Sculpture elements: rugae, Sculpture aspect: arcuate. Eyes. Size: subequal to half width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: converging ventrally. Antennae. Length: reach to anterior pronotal angles, Serration: from antennomere 4. PRONOTUM. Shape: visually transverse, Sides: subangulate, Maximal width: at posterior margin, Anterior margin: narrower than posterior. Anterior lobe. Development: absent or vague. Posterior angles. Shape: obtuse, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: entire, Lateral impressions (width): narrow. Prehumerus. Development: obsolete, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: distant from lateral pronotal carina, Posterior end: distant from posterior pronotal angle or margin, Arc: moderate. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior angles. Scutellum. Carina: obsolete or absent. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate, Modifications: widely expanded. Pubescence. Extent: distal only, Distal (shape): preapical. STERNUM. Prosternal lobe. Distal margin: arcuate, Emargination (width): wide. Prosternal process. Shape: subparallel, Sides: arcuate, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (length): protruding distinctly beyond angles. ABDOMEN. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate, Sinuosity (depth): very shallow. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Ovipositor. Shape: square (uritiform). Variability. The species is known only from the female holotype. Diagnosis. A. draco sp. nov. has no similar species in the Oriental fauna, but resembles Neotropical A. pyropygus Thomson, 1878. Among Oriental species, the new species distantly resembles A. bicornis Th��ry, 1930 by having expanded elytral apices, but can be easily recognized by the much greater size. Type material. Type locality: Malaysia, Sabah, Crocker Range, Mt. Trus Madi, N 05��33'00���, E 116��31'00���E, altitude 1000 m. Type specimens. Holotype ♀ (EJCB): ��� Malaysia: Sabah, Crocker Range, Mt. Trus Madi, 1000m \ N05��33'00���, E 116��31'00���E, 20.iii.���28.iv.2006, S. Chew coll.���. Distribution. MALAYSIA: Sabah. Etymology. The specific name draco (dragon) is the Latin noun in apposition. It refers to the remarkable species appearance more typical for Neotropical Agrilus., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on pages 450-451, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus xia sp. nov. (Figs. 1A, 5A) Description of holotype. Body length: 7.3 mm. BODY. Shape: subparallel, Color (dorsally): unicolored. HEAD. Size: very large, Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate, Sculpture density: dense. Eyes. Size: subequal or larger than width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: converging ventrally. Antennae. Serration: from antennomere 4, Antennomeres 7–10: with obvious petiole. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: obtuse, Apex: sharp. Disk. Convexity: flat, Impressions: medial and lateral, Medial impression: entire, Lateral impressions (depth): shallow, Lateral impressions (width): narrow. Prehumerus. Development: obsolete, Extent: adjoining posterior angle, Posterior end: joined with posterior pronotal angle or margin. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5–1/4 of marginal carina. Scutellum. Size: robust. ELYTRA. Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate. Pubescence. Extent: distal only, Distal (shape): perisutural stripe. STERNUM. Pubescence. Strip of erect pubescence: absent. Prosternal lobe. Size: large, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): wide. Prosternal process. Width: wide, Shape: markedly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, with cluster of setae, Projection (length): protruding distinctly beyond angles. Metasternum. Modification: with cluster of setae above metacoxal plates. ABDOMEN. Tomentum: absent. Basal ventrite. Modifications: with tubercle(s). Pygidium. Apical margin: arcuate. Sternal groove. Extent: on all ventrites, Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to mesotarsus: obviously longer, Length to metatibia: somewhat shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4. GENITALIA. Aedeagus. Symmetry: symmetric, Shape: widest in apical part, Modifications: parameres with obvious membranous lobes. Variability. The species is known only from the holotype. Diagnosis. Agrilus xia sp. nov. is similar to the Palaearctic A. graminis Kiesenwetter, 1857 by the habitus, by the pattern of the elytral pubescence, and by the clusters of setae on the prosternum. The new species can be distinguished by the unmodified antennae in the males, by the obsolete prehumerus, by the large prosternal lobe, by two clusters of setae on the prosternum, with the larger cluster at the base and the smaller cluster on the apex of prosternal process (one cluster in A. graminis), and by the presence of a single hook-like tubercle in the middle of the basal ventrite. Type material. Type locality: North Laos, Phongsaly province, Phongsaly env., 21°41'–2'N, 102°06'–08'E, altitude ~ 1500m. Type specimens. Holotype ♂ (EJCB): “Lao-N, Phongsaly prov., 21°41'–2'N, 102°06'–08'E, 28.v.–20.vi.2003 Phongsaly env., ~ 1500m, Vít Kubáň leg.”. Distribution. LAOS: Phôngsali.

Agrilus jum sp. nov. (Figs. 2D, 5J) Description of holotype. Body length: 4.4 mm. BODY. Shape: cuneiform, Posterior tapering part: long with narrow apex, Color (dorsally): unicolored. HEAD. Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate, Sculpture density: dense. Eyes. Size: larger than half width but smaller than width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually transverse, Sides: markedly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: obtuse, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (depth): shallow, Lateral impressions (width): broad. Prehumerus. Development: filamentary, Shape: arcuate, Extent: beyond 1/2 of pronotal length, Modifications: with rudiment at anterior part, Anterior end: adjoining to lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: moderate. Lateral carinae. Interspace: narrow, Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5–1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate. Pubescence. Extent: proximal and distal, Proximal (shape): quadrangular, Distal (shape): preapical. STERNUM. Pubescence. Strip of erect pubescence: absent. Prosternal lobe. Position to sternum: markedly declivous, Distal margin: arcuately emarginate, Emargination (width): wide. Prosternal process. Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: impressed, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate, Sinuosity (depth): markedly deep. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4. GENITALIA. Aedeagus. Symmetry: feebly asymmetric, Profile: flat, Modifications: apex of medial lobe markedly obtuse. Variability. Body length: 4.3–4.4 mm. The species is known only from males. Sexual dimorphism. The male has a medial strip of erect pubescence from the prosternal lobe to the metasternal projection. Diagnosis. Agrilus jum sp. nov. is similar to A. friebi Obenberger, 1922 by the size, by the color, and by the habitus, but it can be distinguished by having a subparallel prosternal process and a feebly asymmetric aedeagus (Fig. 5J). Type material. Type locality: Northeastern Laos, Houa Phan province, 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei + Phou Pane Mt., altitude 1340–1870 m. Type specimens. Holotype ♂, 2 ♂ paratypes (EJCB): “Laos- NE, Houa Phan prov., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei + Phou Pane Mt., 1340–1870m, 15.iv.–15.v.2008, Lao collectors leg.”. Distribution. LAOS: Houaphan.

Agrilus cicadelloides sp. nov. (Fig. 3C) Description of holotype. Body length: 6.8 mm. BODY. Shape: fusiform, Profile: convex, Color (dorsally): unicolored. HEAD. Eyes. Size: smaller than half width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Length: reach to anterior pronotal angles, Width: antennomeres 5–10 obviously short and broad, Serration: from antennomere 5. PRONOTUM. Shape: visually transverse, Sides: slightly arcuate, Maximal width: at posterior margin, Anterior margin: narrower than posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: projecting beyond anterior angles. Posterior angles. Shape: acute, Apex: blunt. Disk. Convexity: strongly convex, Impressions: absent. Prehumerus. Development: obsolete, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: distant from lateral pronotal carina, Posterior end: distant from posterior pronotal angle or margin, Arc: weak. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5–1/4 of marginal carina. Scutellum. Size: robust. ELYTRA. Humeral carina: absent. Apices. Arrangement: conjoint, Shape: arcuate. Pubescence. Extent: entire ornamental. STERNUM. Prosternal lobe. Distal margin: subtruncate. Prosternal process. Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disc: flat. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: angulately sinuate. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4. GENITALIA. Ovipositor. Shape: square (uritiform). Variability. The species is known only from the female holotype. Diagnosis. Agrilus cicadelloides sp. nov. is similar to A. scutellaris Deyrolle, 1864 by the small eyes, by the trapezoid pronotum, by the sculpture of the pronotal disc, by the absence of a prehumerus, and by the robust scutellum. The new species can be distinguished by having a cluster of reddish setae on head, by an obvious pronotal lobe, and by conjointly arcuate elytral apices. Type material. Type locality: Malaysia, Sarawak, Kampung Kuap [01°24'28”N, 110°21'38”E]. Type specimens. Holotype ♀ (EJCB): “ Malaysia, Sarawak, Kampung Kuap, iii. 1914 ”. Distribution. MALAYSIA: Sarawak. Etymology. The specific name cicadelloides is derived from the family name Cicadellidae (Hemiptera), and the suffix- oides, denoting form or resemblance to stress similarity with a leafhopper.

Agrilus kon sp. nov. (Figs. 2F, 5L) Description of holotype. Body length: 4.1 mm. BODY. Shape: subparallel, Profile: convex, Color (dorsally): unicolored. HEAD. Size: very large, Modification: strongly convex. Vertex. Shape: markedly convex, Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate, Sculpture density: dense. Eyes. Size: subequal or larger than width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Maximal width: at anterior margin, Anterior margin: wider than posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: obtuse, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: entire, Lateral impressions (depth): shallow. Prehumerus. Development: carinal, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: adjoining to lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: weak. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5���1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Shape: arcuate. Pubescence. Extent: proximal and distal, Proximal (shape): triangular, Distal (shape): preapical. STERNUM. Pubescence. Strip of erect pubescence: absent. Prosternal lobe. Size: narrow, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): wide. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: impressed, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: with tubercle(s). Last three ventrites. Modifications: with patches of erect pubescence. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate, Sinuosity (depth): markedly deep. LEGS. Metafemora. Modifications: with comb of setae on inner margin. Metatibiae. Modification: with comb of setae on inner side. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Aedeagus. Symmetry: symmetric, Shape: widest in basal part. Variability. The species is known only from the male holotype. Diagnosis. Agrilus kon sp. nov. is similar to A. relegatus Curletti, 1990 by the size, by the habitus, and by the color. The new species can be distinguished by the shape of the aedeagus, which is strongly enlarged apically (Fig. 5L), and by very specific male sexual characters in the form of a pair of tubercles on the basal ventrite, by clusters of fine, erect pubescence in the middle of the last three ventrites, and by a row of setae on the inner margin of the metatibiae and metafemora. Type material. Type locality: South West Cambodia, 20 km SE Koh Kong, Tatai river, 11��34'N, 103��07'E, altitude 50���300 m. Type specimens. Holotype ♂ (EJCB): ���SW Cambodia, 20km SE Koh Kong, Tatai river 11��34'N, 103��07'E, 3���19.v.2005, 50��� 300m, E. Jendek & O. ��au��a leg.���. Distribution. CAMBODIA: Kaol Kong., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on pages 454-455, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus ika sp. nov. (Figs. 2E, 5K) Description of holotype. Body length: 6.6 mm. BODY. Shape: cuneiform, Profile: convex, Color (dorsally): unicolored. HEAD. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate, Sculpture density: dense. Eyes. Size: larger than half width but smaller than width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: converging ventrally. Antennae. Length: reach to anterior pronotal angles, Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Sides (modification): sinuate before posterior angles, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: acute, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: entire. Prehumerus. Development: carinal, Shape: arcuate, Extent: to 1/2 of pronotal length, Modifications: with rudiment at anterior part, Anterior end: distant from lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: moderate. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5–1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: conjoint, Shape: arcuate. Pubescence. Extent: entire except epipleura. STERNUM. Pubescence. Strip of erect pubescence: extending from prosternal lobe to metasternal projection. Prosternal lobe. Size: large, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): narrow. Prosternal process. Width: wide, Shape: markedly dilated, Sides: straight, Angles: acute, Angles (tips): sharp, Disc: flat, Projection (length): adjoining to angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate, Sinuosity (depth): very shallow. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4. GENITALIA. Aedeagus. Symmetry: symmetric, Modifications: medial lobe distinctly wider than parameres. Variability. Body length: 5.9–6.6 mm. The species is known only from males. Sexual dimorphism. The male has a medial strip of erect white pubescence from the prosternal lobe to the metasternal projection. Diagnosis. Agrilus ika sp. nov. is similar to A. dahoi Baudon, 1965 by habitus, by the color, and by the obviously dilated sternal process. The new species can be distinguished by the pronotum with sinuate sides at the posterior angles; by the elytra having a wide strip of adsutural pubescence; by conjointly arcuate elytral apices; and by the apically enlarged aedeagus(Fig. 5K). Type material. Type locality: Solomon Islands, Quadalcanal Island, ca 3.5 km SE Barana village, 09°29.8'S, 159°59.5'E. Type specimens. Holotype ♂, 1 ♂ paratype (EJCB): “ Solomon isl., Quadalcanal, ca 3.5km SE Barana vill., 09°29.8'S, 159°59.5'E, 23.xi.–15.xii.2013, J. Horak leg.”. Distribution. SOLOMON ISLANDS.

Agrilus wos sp. nov. (Figs. 1D, 5D) Description of holotype. Body length: 5.7 mm. BODY. Shape: subparallel, Profile: convex, Color (dorsally): unicolored. HEAD. Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate. Eyes. Size: subequal or larger than width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4, Antennomeres 7–10: with obvious petiole. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Sides (modification): sinuate before posterior angles, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: projecting beyond anterior angles. Posterior angles. Shape: acute, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: entire, Lateral impressions (depth): shallow. Prehumerus. Development: obsolete, Shape: arcuate, Extent: adjoining posterior angle, Anterior end: distant from lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5–1/4 of marginal carina. Scutellum. Size: robust. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: conjoint, Shape: subtruncate. Pubescence. Extent: proximal and distal, Proximal (shape): quadrangular, Distal (shape): apical. STERNUM. Pubescence. Strip of erect pubescence: extending from prosternal lobe to metasternal projection. Prosternal lobe. Size: large, Position to sternum: markedly declivous, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): wide. Prosternal process. Width: wide, Shape: markedly dilated, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to metatibia: somewhat shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4, Modification: with comb of setae on lower side. GENITALIA. Aedeagus. Symmetry: symmetric, Modifications: apex of medial lobe markedly obtuse, medial lobe distinctly wider than parameres. Variability. Body length: 5.1–5.7 mm. The paratype has broadly, separately arcuate elytral apices. Sexual dimorphism. The male has a strip of erect pubescence on the sternum and a comb of erect setae on the first metatarsomere. The female is unknown. Diagnosis. Agrilus wos sp. nov. is similar to A. volkovitshi Jendek, 2007 by the color, by the habitus, by the size of the eyes, by the shape of the pronotum, and by the pattern of the elytral pubescence. The new species can be distinguished by the obliterate prehumerus, by the shape of aedeagus (Fig. 5D), by the lack of tubercles on the basal ventrite, and by the presence of long setae on the first metatarsomere in the males. Type material. Type locality: Northeastern Laos, Houa Phan province, 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei + Phou Pane Mt., altitude 1340–1870 m. Type specimens. HT, 1 ♂ paratype (EJCB): “Laos-NE, Houa Phan prov., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei + Phou Pane Mt., 1340–1870m, 15.iv.–15.v.2008, Lao collectors leg.”. Distribution. LAOS: Houaphan.

Agrilus titi sp. nov. (Fig. 4C) Description of holotype. Body length: 4.3 mm. BODY. Shape: subparallel, Build: robust, Posterior tapering part: short with broad apex, Color (dorsally): unicolored. HEAD. Size: very large, Modification: obviously flat. Vertex. Outline: markedly protruding head outline, Medial impression: present, Sculpture elements: rugae, Sculpture density: sparse, Sculpture intensity: superficial. Eyes. Size: smaller than half width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually transverse, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: absent or vague. Posterior angles. Shape: rectangular, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: entire, Lateral impressions (depth): deep. Prehumerus. Development: carinal, Shape: bisinuate, Extent: to anterior angle, Anterior end: joined with anterior pronotal angle, Posterior end: joined with posterior pronotal angle or margin, Arc: weak. Lateral carinae. Interspace: narrow, Convergence: moderate, Junction: present, Narrowest point: at posterior 1/3 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate. Pubescence. Extent: entire ornamental with indication of stripes. STERNUM. Prosternal lobe. Size: large, Distal margin: arcuate. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Ovipositor. Shape: markedly elongate. Variability. The species is known only from the female holotype. Diagnosis. Agrilus titi sp. nov. is similar to A. evolutus Descarpentries & Villiers, 1963 by the size, by the color, by the habitus, by the very small eyes, by the markedly transverse pronotum, and by the similar elytral pubescence. The new species can be distinguished by the sinuate carinal prehumerus extending to the anterior pronotal angles. Type material. Type locality: Malaysia, West Perak, 25 km Northeast of Ipoh, Banjaran Titi Wangsa mts., Korbu Mt., altitude 1200 m. Type specimens. Holotype (EJCB): ��� Malaysia, W Perak, 25 km NE of Ipoh, 1200 m, Banjaran Titi Wangsa mts., Korbu Mt., 6���12.v.2001, P. Čechovsk�� leg.���. Distribution. MALAYSIA: Perak. Etymology. The specific name is derived from a part of the name of the Banjaran Titi Wangsa Mountains, the type locality., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on pages 458-459, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus uxo sp. nov. (Figs. 1E, 5E) Description of holotype. Body length: 5.7 mm. BODY. Shape: subparallel, Profile: convex. HEAD. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: subparallel, Sculpture density: dense. Eyes. Size: larger than half width, but smaller than width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4, Antennomeres 7���10: with obvious petiole. PRONOTUM. Shape: visually transverse, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: narrower than posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: rectangular, Apex: blunt. Disk. Impressions: medial and lateral, Medial impression: entire, Lateral impressions (depth): shallow, Lateral impressions (width): broad. Prehumerus. Development: carinal, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: distant from lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: weak. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5���1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate. Pubescence. Presence: without discernible pubescence. STERNUM. Pubescence. Strip of erect pubescence: absent. Prosternal lobe. Size: large, Position to sternum: markedly declivous, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): wide. Prosternal process. Shape: markedly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: flat with cluster of setae, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: impressed. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate, Sinuosity (depth): markedly deep, Sinuosity (width): very narrow. LEGS. Metacoxal plates. Modifications: with tubercle on inner margin. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: longer than 2���3 but shorter than 2���4. GENITALIA. Aedeagus. Symmetry: feebly asymmetric, Profile: flat. Variability. The species is known only from the male holotype. Diagnosis. Agrilus uxo sp. nov. is similar to the Palaearctic A. graminis Kiesenwetter, 1857 by the habitus and by the presence of a cluster of setae on the base of the prosternal process. The new species can be distinguished by unmodified antennae in males, by the lack of white elytral pubescence, by a small cluster of setae on the inner margin of the metacoxae, and by the shape of the aedeagus (Fig. 5E). Type material. Type locality: North Vietnam, Ninh Binh province, Cuc Phuong National Park, N 20��21'10���, E105��35'00���, altitude 440 m. Type specimens. Holotype ♂ (EJCB): ���N Vietnam, Ninh Binh prov., Cuc Phuong N. Park, N20��21'10���, E105��35'00���, 24���28.iv.2012, alt 440m, E. Jendek leg.���. Distribution. VIETNAM: Ninh Binh., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on page 459, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus qom sp. nov. (Figs. 2C, 5I) Description of holotype. Body length: 4.3 mm. BODY. Shape: cuneiform, Color (dorsally): unicolored. HEAD. Size: very large, Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: subparallel, Sculpture density: sparse, Sculpture intensity: superficial. Eyes. Size: subequal or larger than width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Maximal width: at anterior margin, Anterior margin: wider than posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: obtuse, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (depth): shallow. Prehumerus. Development: filamentary, Shape: arcuate, Extent: to 1/2 of pronotal length, Modifications: with rudiment at anterior part, Anterior end: adjoining to lateral pronotal carina, Posterior end: distant from posterior pronotal angle or margin, Arc: moderate. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5���1/4 of marginal carina. ELYTRA, Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate. Pubescence. Extent: entire. STERNUM. Pubescence. Strip of erect pubescence: absent. Prosternal lobe. Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): narrow. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: impressed, with row of setae on sides, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: impressed. ABDOMEN. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Aedeagus. Symmetry: symmetric, Shape: widest in apical part, Modifications: apex of medial lobe sharply pointed, parameres with obvious membraneous lobes. Variability. The species is known only from the male holotype. Sexual dimorphism. In the male, sides of prosternal process with a row of setae. Diagnosis. Agrilus qom sp. nov. is similar to Agrilus pusillesculptus Obenberger, 1940 by the size, by the color, by the habitus, by the large eyes protruding outside the outline of the head in dorsal view, and by the obvious pronotal lobe. The new species can be distinguished by the maximum width located at the anterior margin of the pronotum; by the shorter prehumerus, extending to half the pronotal length; by the separately arcuate elytral apices; and by the shape of the aedeagus (Fig. 5I). Type material. Type locality: Northeastern Laos, Houa Phan province, 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei + Phou Pane Mt., altitude 1340���1870 m. Type specimens. Holotype ♂ (EJCB): ���Laos-NE, Houa Phan prov., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei + Phou Pane Mt., 1340���1870m, 15.iv.���15.v.2008, Lao collectors leg.���. Distribution. LAOS: Houaphan., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on page 458, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus yuk sp. nov. (Figs. 2A, 5G) Description of holotype. Body length: 7.2 mm. BODY. Shape: subparallel, Posterior tapering part: short with broad apex, Profile: dorsally flattened, Color (dorsally): unicolored. HEAD. Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture density: dense, Sculpture intensity: rough. Eyes. Size: subequal or larger than width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: converging ventrally. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: projecting beyond anterior angles. Posterior angles. Shape: obtuse, Apex: blunt. Disk. Convexity: flat, Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (depth): shallow, Lateral impressions (width): broad. Prehumerus. Development: carinal, Shape: straight, Extent: adjoining posterior angle, Anterior end: distant from lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5���1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate. Pubescence. Extent: distal only, Distal (shape): preapical. STERNUM. Pubescence. Strip of erect pubescence: extending from prosternal lobe to metasternal projection. Prosternal lobe. Size: narrow, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): wide. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Angles (tips): sharp, Disc: impressed, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: angulately sinuate, Sinuosity (depth): markedly deep. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Aedeagus. Symmetry: symmetric, Shape: widest in apical part. Variability. Body length: 5.9���7.4 mm. Some paratypes have additional pubescenceon the anterior half of the elytra. Sexual dimorphism. The males have longer antennae, broader elytral apices, and a medial strip of erect pubescence on the sternum. The females have a markedly elongate ovipositor. Diagnosis. Agrilus yuk sp. nov. is similar to the Palaearctic A. sulcicollis Lacordaire, 1835 by the size, by the general habitus, by the shape of the pronotum, by the widely arcuate elytral apices, and by the narrow sternal lobe. The new species can be distinguished by the presence of the elytral pubescence, by the short prehumerus, by the lack of tubercles on basal ventrite in males, and by the shape of the aedeagus (Fig. 5G). Type material. Type locality: Northeastern Laos, Houa Phan province, 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei + Phou Pane Mt., altitude 1340���1870 m. Type specimens. Holotype ♂, 1 ♂ paratype, 4 ♀ paratypes (EJCB): ���Laos-NE, Houa Phan prov., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei + Phou Pane Mt., 1340���1870m, 15.iv.���15.v.2008, Lao collectors leg.���. Other types. LAOS: Houaphan: ���Laos-NE, Hua Phan Province, Ban Saleui, Phou Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. Holzschuh ��� 1 ♀ paratype (USNM); ���Laos-NE, Houa Phan prov., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy + Phou Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & Lao coll. leg.��� 1 ♂ paratype (EJCB). Distribution. LAOS: Houaphan., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on pages 464-466, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus mimicus sp. nov. (Fig. 4A) Description of holotype. Body length: 5.1 mm. BODY. Shape: cuneiform, Color (dorsally): unicolored. HEAD. Vertex. Outline: markedly protruding head outline, Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate. Eyes. Size: subequal to half width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Sides (modification): sinuate before posterior angles, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Shape: arcuate, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: acute, Apex: sharp. Disk. Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (width): narrow. Prehumerus. Development: carinal, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: adjoining to lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: obvious. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior 1/5���1/4 of marginal carina. ELYTRA. Color: unicolored, Humeral carina: absent. Apices. Arrangement: separate, Shape: cuspidate, Number of cusps or spines: 2, Position of dominant projection: sutural margin. Pubescence. Color: bicolored, Character: with patches or spots of denser pubescence, Extent: entire ornamental with indication of stripes. STERNUM. Prosternal lobe. Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): narrow. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: impressed, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2���4. GENITALIA. Ovipositor. Shape: markedly elongate. Variability. This species is known only from the female holotype. Diagnosis. By having bicuspidate elytral apices, A. mimicus resembles species related to A. albogaster Deyrolle, 1864. It differs by the markedly longer pronotum when compared to the elytra, by the rough sculpture of the head and pronotum, and by the camouflaging pattern of the elytral pubescence. Type material. Type locality: Laos, Louang Namtha province, Namtha���Muang Sing, 21��09'N, 101��19'E, altitude 900���1200 m. Type specimens. Holotype ♀ (EJCB): ��� Laos, Louang Namtha pr., 21��09'N, 101��19'E, Namtha���Muang Sing, 5���31.v.1997, 900��� 1200 m, Vit Kub��ň leg.���. Distribution. LAOS: Louang Namtha. Etymology. The specific name mimicus (mimic) is the Latin adjective referring to the camouflage-like appearance of this species., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on pages 455-457, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus hergovitsi sp. nov. (Figs. 1B, 5B) Description of holotype. Body length: 5.4 mm. BODY. Shape: subparallel, Profile: convex, Color (dorsally): unicolored. HEAD. Size: very large, Modification: strongly convex. Vertex. Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate, Sculpture density: dense, Sculpture intensity: superficial. Eyes. Size: subequal or larger than width of vertex (dorsal view), Lower margin: in line or below antennal socket, Median orbit: converging ventrally. Antennae. Serration: from antennomere 4, Antennomeres 7���10: markedly longer than wide. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Maximal width: at anterior margin, Anterior margin: wider than posterior. Anterior lobe. Development: obvious, Shape: arcuate, Position: at level with anterior angles. Posterior angles. Shape: rectangular, Apex: sharp. Disk. Convexity: flat, Impressions: medial and lateral, Medial impression: anteromedial and posteromedial, Lateral impressions (depth): shallow, Lateral impressions (width): broad. Prehumerus. Development: filamentary, Shape: arcuate, Extent: beyond 1/2 of pronotal length, Anterior end: adjoining to lateral pronotal carina, Posterior end: distant from posterior pronotal angle or margin, Arc: weak. Lateral carinae. Convergence: moderate, Junction: present, Narrowest point: at posterior angles. ELYTRA. Color: unicolored. Apices. Arrangement: separate, Shape: subangulate. Pubescence. Extent: distal only, Distal (shape): perisutural stripe. STERNUM. Prosternal lobe. Size: large, Position to sternum: markedly declivous, Distal margin: arcuately emarginate, Emargination (depth): shallow, Emargination (width): narrow. Prosternal process. Shape: slightly dilated, Sides: straight, Angles: obtuse, Disc: impressed with sides exserted, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: impressed. ABDOMEN. Basal ventrite. Modifications: with impression(s). Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuate. LEGS. Metafemora. Modifications: with comb of setae on inner margin. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: longer than 2���3 but shorter than 2���4. GENITALIA. Aedeagus. Symmetry: symmetric, Shape: widest in basal part, Modifications: apex of medial lobe markedly obtuse. Variability. The species is known only from the holotype. For male sexual dimorphism see Diagnosis. Diagnosis. The species is similar to A. syrphus Descarpentries & Villiers, 1963 in color and general appearance, but can be distinguished by the longer pronotum, by the gracile antennae, by the obliterated elytral pubescence, and by the shape of the aedeagus (Fig. 5B). Type material. Type locality: Malaysia, Johor, Endau Rompin National Park, 2��37'20���N 103��20'E, altitude 120���300 m. Type specimens. Holotype ♂ (EJCB): ��� Malaysia, Johor, Endau Rompin NP, 2��37'20���N 103��20'E, 23.ix.���3.x.2012, 120��� 300m, R. Hergovits leg.���. Distribution. MALAYSIA: Johor. Etymology. Patronymic; the species is named in honor of entomologist Roman Hergovits, Slovakia, who collected the holotype., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2019, Twenty more new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental region, pp. 449-469 in Zootaxa 4564 (2) on page 451, DOI: 10.11646/zootaxa.4564.2.7, http://zenodo.org/record/3713725

Agrilus zim sp. nov. (Fig. 4B) Description of holotype. Body length: 5.2 mm. BODY. Shape: cuneiform, Color (dorsally): unicolored. HEAD. Modification: strongly convex. Vertex. Outline: markedly protruding head outline, Medial impression: present, Sculpture elements: rugae, Sculpture aspect: arcuate. Eyes. Size: subequal or larger than width of vertex (dorsal view), Shape: markedly protruding head outline, Lower margin: in line or below antennal socket, Median orbit: subparallel. Antennae. Serration: from antennomere 4. PRONOTUM. Shape: visually elongate or square, Sides: slightly arcuate, Sides (modification): sinuate before posterior angles, Maximal width: at middle, Anterior margin: subequal to posterior. Anterior lobe. Development: obvious, Width: broad, Position: at level with anterior angles. Posterior angles. Shape: acute, Apex: sharp. Disk. Convexity: flat, Impressions: medial and lateral, Medial impression: entire, Lateral impressions (width): broad. Prehumerus. Development: carinal, Shape: arcuate, Extent: to 1/2 of pronotal length, Anterior end: joined with lateral pronotal carina, Posterior end: joined with posterior pronotal angle or margin, Arc: moderate. Lateral carinae. Interspace: narrow, Convergence: moderate, Junction: present, Narrowest point: at posterior angles. ELYTRA. Humeral carina: absent. Apices. Arrangement: separate, Shape: arcuate, Modifications: margin obviously denticulate. Pubescence. Color: bicolored, Character: with patches or spots of denser pubescence, Extent: entire ornamental with indication of stripes. STERNUM. Prosternal lobe. Distal margin: arcuate. Prosternal process. Shape: subparallel, Sides: straight, Angles: obtuse, Angles (tips): blunt, Disc: impressed, Projection (length): protruding distinctly beyond angles. Metasternum. Metasternal projection: flat. ABDOMEN. Basal ventrite. Modifications: absent. Pygidium. Apical margin: arcuate. Sternal groove. Shape on apex of last ventrite: arcuately sinuate. LEGS. Metatarsus. Length to metatibia: distinctly shorter. Metatarsomere 1. Length to following tarsomeres: subequal or longer than 2–4. GENITALIA. Ovipositor. Shape: markedly elongate. Variability. The species is known only from the female holotype. Sexual dimorphism. Females with an elongate ovipositor. Diagnosis. Agrilus zim sp. nov. is similar to A. mimicus sp. nov. (Fig. 4A) by the elongated pronotum, by the strongly arcuate carinal prehumerus, and by the bicolored elytral pubescence. It differs by having the anterior end of the prehumerus joined with the lateral pronotal carina, by the separately subarcuate elytral apices, by the arcuate anterior margin of the sternal lobe, and by the subparallel sternal process. Type material. Type locality: Malaysia, [Pahang], Benom Mts, 15 km East of Kampong Dong, 3°53' N, 102°01'E, altitude 700 m. Type specimens. Holotype ♀ (EJCB): “ Malaysia, Benom Mts, 15 km E Kampong Dong, 1.iv.1998, alt. 700 m, 3.53° N, 102.01°E, D. Hauck leg.”. Distribution. MALAYSIA: Pahang. Remarks. Coordinates cited on the locality label are stated incorrectly. The proper format is 3°53'N 102°01'E.

Oreoscotus Aurivillius, 1910 Oreoscotus Aurivillius, 1910: 421 (species included: serratus, minor, laevis). Type species: not yet designated. Remarks. Judging by the results of the morphological comparison of the type specimens, both of Hustache���s taxa are not congeneric with M. carinatus and have to, therefore, be transferred to another genus. Choosing such a genus requires a phylogenetic hypothesis of African Molytinae, and such a hypothesis is currently absent. To the best of available knowledge, both Hustache���s species likely belong to a possible clade distinguished by the adult prosternal canal delimited laterally by carinae and uniting Afrotropical species currently assigned to the genera Niphades Pascoe, 1871, Oreoscotus Aurivillius, 1910 and Niphadonothus Voss, 1965. This agrees with an earlier thought captured on one of the labels of the O. puncticollis holotype, where the specimen is related to Niphades (written perhaps by Guy A.K. Marshall, Figs 2C, D). These three genera, however, have themselves never been assessed from a phylogenetic viewpoint and their taxonomic validity is based on the pre-Henniginan typological concept. Alonso-Zarazaga & Lyal (1999) placed Oreoscotus as Molytinae incertae sedis, while Alonso-Zarazaga & Lyal (2002) transferred it to Aminyopini. Lyal (2014) included Niphades and other Aminyopini in the tribe Euderini; in this paper characters for the redefined Euderini were provided but not, however, cladistically analysed. Moreover, preliminary data suggest that the genus Niphades is widely distributed in the Afrotropical, Palaearctic and Oriental regions, has its type species in Sarawak and is not monophyletic. Considering all this uncertainty and pending further studies, both of Hustache���s nominal species originally described as Molytophilus (affinis and puncticollis), are herein transferred to the so far exclusively Tanzanian genus Oreoscotus to become O. affinis (Hustache, 1936) comb. n. and O. puncticollis (Hustache, 1936) comb. n., Published as part of Grebennikov, Vasily V., 2018, Both non-type species of Molytophilus (Coleoptera: Curculionidae: Molytinae) are transferred to Oreoscotus, pp. 393-396 in Zootaxa 4418 (4) on page 395, DOI: 10.11646/zootaxa.4418.4.6, http://zenodo.org/record/1244950, {"references":["Aurivillius, C. (1910) 7. Coleoptera. 21 Curculionidae. In: Sjostedt, Y. (Ed.), Wissenschaftliche Ergebnisse der Schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906, unter leitung von Prof. Dr. Yngve Sjostedt. Vol. 1. Abteilung 1 - 7. P. Palmquists Aktiebolag, Stockholm, pp. 403 - 435.","Pascoe, F. P. (1871) Contributions towards a knowledge of the Curculionidae. Part II. Journal of the Linnaean Society of London, Zoology, 11 (51), 154 - 218, pls. vi - ix.","Voss, E. (1965) Mission zoologieque de l'I. R. S. A. C. en ' Afrique orientale (P. Basilewsky et N. Leleup, 1957). XCVIII. - Coleoptera Curculionidae. II. Annales du Musee Royal de l'Afrique Centrale, Serie in 8 °, 138, 293 - 375.","Alonso-Zarazaga, M. A. & Lyal, C. H. C. (1999) A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (excepting Scolytidae and Platypodidae). Entomopraxis, Barcelona, 315 pp.","Alonso-Zarazaga, M. A. & Lyal, C. H. C. (2002) Addenda and corrigenda to ' A World Catalogue of Families and Genera of Curculionoidea (Insecta: Coleoptera) '. Zootaxa, 63 (1), 1 - 37. https: // doi. org / 10.11646 / zoootaxa. 63.1.1","Lyal, C. H. (2014) 3.7. 7 Molytinae Schoenherr, 1823. In: Leschen, R. A. B. & Beutel, R. G. (Eds.), Handbook of Zoology, Arthropoda: Insecta: Coleoptera. Volume 3: Morphology and Systematics (Phytophaga). Walter de Gruyter, Berlin, pp. 529 - 570.","Hustache, A. (1936) XLVII. - Entomological Expedition to Abyssinia, 1926 - 7: Coleoptera, Curculionidae. Annals and Magazine of Natural History, Series 10, 18, 353 - 373, 419 - 446, 483 - 505, 563 - 587."]}

Molytophilus Hartmann, 1904 Hartmann, 1904: 388 (species included: carinatus). Type species: Molytophilus carinatus Hartmann, 1904 fixed by monotypy. Remarks. The herein re-defined genus Molytophilus is monotypic and its only species M. carinatus is known from two east African localities separated by about one thousand kilometres: the type locality in the Usambara mountains in northeastern Tanzania and the Genale region in vicinity of Mogadishu in south-eastern Somalia. Its current taxonomic assignment to Molytini: Plinthina (Alonso-Zarazaga & Lyal 1999) or Plinthini (Lyal 2014) suggests a close relationship with at least some of the approximately 20 genera traditionally attributed to this group (Grebennikov 2014). This hypothesis was first implicitly suggested by Hartmann (1904: 389), who noted an external resemblance of M. carinatus and ��� Plinthus caliginosus ��� (= Mitoplinthus caliginosus (F.), figs 15A���B in Grebennikov 2014). The subtribe Plinthina (sensu Grebennikov 2014), like the great majority of other weevil tribes and subtribes, has never been subjected to a phylogenetic analysis and, therefore, its monophyly and the relationships of its members to Molytophilus have never been tested. Biological preferences of the genus and its immature stages are unknown., Published as part of Grebennikov, Vasily V., 2018, Both non-type species of Molytophilus (Coleoptera: Curculionidae: Molytinae) are transferred to Oreoscotus, pp. 393-396 in Zootaxa 4418 (4) on page 393, DOI: 10.11646/zootaxa.4418.4.6, http://zenodo.org/record/1244950, {"references":["Hartmann, F. (1904) Neue Russelkafer aus Ostafrika. Deutsche Entomologische Zeitschtift, 1904, 369 - 419. https: // doi. org / 10.1002 / mmnd. 48019040206","Alonso-Zarazaga, M. A. & Lyal, C. H. C. (1999) A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (excepting Scolytidae and Platypodidae). Entomopraxis, Barcelona, 315 pp.","Lyal, C. H. (2014) 3.7. 7 Molytinae Schoenherr, 1823. In: Leschen, R. A. B. & Beutel, R. G. (Eds.), Handbook of Zoology, Arthropoda: Insecta: Coleoptera. Volume 3: Morphology and Systematics (Phytophaga). Walter de Gruyter, Berlin, pp. 529 - 570.","Grebennikov, V. V. (2014) Morimotodes, a new genus for two minute wingless litter species from southwest China and Taiwan with an illustrated overview of Molytina and Plinthina genera (Coleoptera: Curculionidae: Molytini). Bonn Zoological Bulletin, 63, 123 - 147."]}

Molytophilus carinatus Hartmann, 1904 (Figs. 1A���G) Molytophilus carinatus Hartmann, 1904: 388 ���389 (description) Type locality: Ostafrika, Usambara. Type specimens: Primary: Lectotype (Figs. 1A���E), male (MTD), herein designated to maintain taxonomic stability: ���Usambara Speyer [handwritten, =h] \ [male sign] [h] \ Typus [h] [red label] \ Molytophilus carinatus Hrtm. [h] \ Samml. K.F. Hartmann Ankauf 1941.1 [printed, =p] [blue label] \ Staatl. Museum f��r Tierkunde. Dresden [p] \ LECTOTYPE Molytophilus carinatus Hartmann, 1904 V.Grebennikov des., 2013 [p] [red label]���. Secondary: 2 Paralectotypes (Fig. 1F) (MTD), each with male and female handwritten sign, respectively, same labels as lectotype, except the locality label ���Usambara��� is written in another hand and without ���Speyer���, and with labels ��� PARALECTOTYPE Molytophilus carinatus Hartmann, 1904 V.Grebennikov des., 2013��� [p; yellow label]; handwritten identification label is absent. Described from unknown number of syntypes, but since ���Hartmann��� appears on labels of these specimens I consider them part of the type series. Non-type specimens: (Fig. 1G) 2 (MNHN): ��� SOMALIA Genale, F.Bili 5-935 [p] \ Molytophilus carinatus Hartm. [h] MUSEUM PARIS 1968 Coll. A.HOFFMANN [p]���. Remarks. For diagnostic purposes this species is easily recognized among likely all Molytinae by its characteristic habitus, along with the following three characters: 1, all tibiae with large spines on inner apical part (arrows, Fig. 1F); 2, lateral side of rostrum with deep groove adjacent to anterior edge of eye (arrow, Fig. 1B); and 3, robust antennae with narrow club (Fig. 1B). It remains uncertain in which of the two currently recognized Usambara regions, Western versus Eastern Usambara (Grebennikov 2017), the type series has been collected. Phylogenetic affinities of this species and, consequently, of the herein re-defined genus Molytophilus are entirely unknown. It appears probable that, unlike many East African Molytinae, this species does not inhabit the forest leaf litter since not a single specimen has been re-sampled in Western or Eastern Usambara during my recent litter sifting campaign (Grebennikov 2017)., Published as part of Grebennikov, Vasily V., 2018, Both non-type species of Molytophilus (Coleoptera: Curculionidae: Molytinae) are transferred to Oreoscotus, pp. 393-396 in Zootaxa 4418 (4) on pages 393-394, DOI: 10.11646/zootaxa.4418.4.6, http://zenodo.org/record/1244950, {"references":["Hartmann, F. (1904) Neue Russelkafer aus Ostafrika. Deutsche Entomologische Zeitschtift, 1904, 369 - 419. https: // doi. org / 10.1002 / mmnd. 48019040206","Grebennikov, V. V. (2017) Phylogeography and sister group of Lupangus, a new genus for three new flightless allopatric forest litter weevils endemic to the Eastern Arc Mountains, Tanzania (Coleoptera: Curculionidae, Molytinae). Fragmenta entomologica, 49, 37 - 55. https: // doi. org / 10.4081 / fe. 2017.229"]}

Oreoscotus affinis (Hustache, 1936) comb. n. (Figs. 2A, B) Molytophilus affinis Hustache, 1936: 492 (description) Type locality: Abyssinie: entre Djem-Djem et Wouramboulchi, vers 9000 pieds. Type specimens: Primary: Holotype (by monotypy, Figs. 2A, B), unsexed (NHMUK): ��� Type [p] \ TYPE [h] \ Abyssinia: Between Djem-Djem and Wouramboulchi. 8,000���9,000ft. 30.ix���1.x.1926. J.Omer Cooper. [p] \ Specimen figured [h] \ Molytophilus affinis [h] Hustache det. [p]���. Remarks. The only known specimen of this nominal species was collected on the watershed separating the Blue Nile basin and that of Awash River (spelt as ���Hawash��� in Omer-Cooper [1930], map 2) some 75 km W of Addis-Ababa and at relatively high altitude of about 2438���2743 meters., Published as part of Grebennikov, Vasily V., 2018, Both non-type species of Molytophilus (Coleoptera: Curculionidae: Molytinae) are transferred to Oreoscotus, pp. 393-396 in Zootaxa 4418 (4) on page 395, DOI: 10.11646/zootaxa.4418.4.6, http://zenodo.org/record/1244950, {"references":["Hustache, A. (1936) XLVII. - Entomological Expedition to Abyssinia, 1926 - 7: Coleoptera, Curculionidae. Annals and Magazine of Natural History, Series 10, 18, 353 - 373, 419 - 446, 483 - 505, 563 - 587.","Omer-Cooper, J. (1930) 13. Dr. Hugh Scott's expedition to Abyssinia. - A preliminary investigation of the freshwater fauna of Abyssinia. Proceedings of the Zoological Society of London, 100 (1), 195 - 207, 3 pl."]}

Oreoscotus puncticollis (Hustache, 1936) comb. n. (Figs. 2C, D) Molytophilus puncticollis Hustache, 1936: 492 (description), 493 (habitus illustration) Type locality: Abyssinie. Type specimens: Primary: Holotype (by original designation, Fig. 2C, D), female (MNHN): ���Abyss. R. [h] \ Gen. nov. nr. Niphades. Another sp. in BM. [h] \ TYPE [p]���. Secondary: Paratype (MNHN): not studied. Described from two specimens (holotype, paratype). Remark: Hustache noted that the holotype has been deposited in MNHN, while the paratype has remained in his collection. Later the Hustache collection was incorporated into MNHN. The only specimen located in MNHN in 2011 has the label ��� Type ��� and is, therefore, considered to be the holotype., Published as part of Grebennikov, Vasily V., 2018, Both non-type species of Molytophilus (Coleoptera: Curculionidae: Molytinae) are transferred to Oreoscotus, pp. 393-396 in Zootaxa 4418 (4) on page 396, DOI: 10.11646/zootaxa.4418.4.6, http://zenodo.org/record/1244950, {"references":["Hustache, A. (1936) XLVII. - Entomological Expedition to Abyssinia, 1926 - 7: Coleoptera, Curculionidae. Annals and Magazine of Natural History, Series 10, 18, 353 - 373, 419 - 446, 483 - 505, 563 - 587."]}

Devernodes gen. n. Type species: Devernodes chthonia sp. n., here designated. Diagnosis. Adults of the genus Devernodes can be most efficiently distinguished from those of other Curculionidae distributed in the Palaeaerctic or Oriental Regions by a unique combination of two easy-to-see head characters: antennae with 6-segmented funicle and apparently unsegmented club, as well as a strong constriction separating the eye-bearing rostrum from the head capsule. Life stages of Devernodes other than adults are unknown and cannot be identified using data other than the herein released DNA sequences. Description. Black or dark-brown, with compact and strongly sclerotized body, about 3���5 mm in body length, in dorsal view elongate, parallel-sided or slightly rounded. Rostrum variously curved, about 4 times as long as wide, with longitudinal lines of punctures sometimes merging in grooves. Prementum distinctly visible in ventral view. Antennae with apparently unsegmented club and funicle consisting of six antennomeres. Eyes fully developed and rounded or partly reduced and crescentic, each consisting of about 7���30 ommatidia. Head capsule spherical, smooth and shiny, without punctures or pilosity, separated from eye-bearing rostrum by notable constriction. Pronotum densely and evenly punctate, prothorax without ocular lobes, procoxae contiguous or narrowly separated, all femora without spurs or spines. Elytral shoulders effaced, each elytron with 7 or 9 punctate striae and with short, sparse setae on intrestriae. Pygidium in both sexes formed by tergite VII. Male sternite IX Ishaped (=not Y-shaped), male genitalia of pedal type, without large internal sclerotization. Each of two female hemisternites IX entire (not subdivided into apical ���stylus��� and basal ���coxite���). Diversity, distribution and biology. Biological information on all five Devernodes species is restricted to the collecting circumstances suggesting that the adult beetles inhabit forest floor litter in Southeast Asia. It should be noted that despite intense sampling in China and Taiwan (Grebennikov 2018b), Mount Emei Shan is the only locality where specimens of Devernodes were recorded. Etymology. Devernodes, patronymic, derived by adding the Latin suffix -odes to the family name of Linda DeVerno (retired, formerly Director of the Ottawa Plant Laboratory, Canadian Food Inspection Agency, Ottawa, Canada), in recognition of her role in development of the sampling program in the Asia-Pacific Region, which lead to the discovery of this new genus. Gender masculine., Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 365, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691, {"references":["Grebennikov, V. V. (2018 b) Dryophthorinae weevils (Coleoptera: Curculionidae) of the forest floor in Southeast Asia: three marker analysis reveals monophyly of Asian Stromboscerini and new identity of rediscovered Tasactes. European Journal of Entomology, 115, 437 - 444."]}

Devernodes asteria sp. n. Figs 3A���G, 7C,I, 8C,I, 9C, 10, 11. Diagnostic description. Holotype male (Figs 3A���G, 7C,I, 8C,I, 9C), body length 3.2 mm, DNA barcode GenBank accession number in Fig. 10; unique combination of morphological characters in Table 3. Material examined. Holotype (CNC): ��� VIETNAM, Tam Dao, N 2127'54" E10538 '44", 12.v.2012, 1218m, sift02, V.Grebennikov���, ���CNCCOLVG00004340���. Paratypes (CNC): 4 specimens, as in Fig. 10, same locality as holotype. Distribution. Known only from the type locality, the Tam Dao Mountains in northern Vietnam (Fig. 12). Elevation: 1,219 ���1,240 m. Etymology. One of the seven daughters of Alcyoneus; noun in apposition., Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 368, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691

Key to Devernodes adult beetles 1 Each elytron with seven striae (Figs 8G,H.J); body form more elongate in dorsal view, ratio of length to maximal width 2.1���2.5 (Figs. 1A, 2A, 4A).................................................................................... 2 1* Each elytron with nine striae (Figs 8I,K,L); body form more rounded in dorsal view, ratio of length to maximal width 1.9���2.2 (Figs 3A, 5A, 6A)..................................................................................... 3 2 Eyes larger, each with at least 15 ommatidia, eyes oval in shape, their posterior edge rounded (Fig. 7G) or straight (Fig. 7H); rostrum dorsally with central longitudinal groove (Figs 7A,B).................... D. alkippe sp. n. (Mount Emei, China) 2* Eyes smaller, each with less than 10 ommatidia, eyes crescentic in shape, their posterior edge concave (Fig 7J); rostrum dorsally with two central longitudinal lines of punctures (Fig. 7D)....... D. chthonia sp. n. (the Tam Dao Mountains, Vietnam) 3 Pronotum with central longitudinal groove containing two lines of punctures (Figs 3A,D); each eye with less than 15 ommatidia (Fig. 7I)................................................ D. asteria sp. n. (the Tam Dao Mountains, Vietnam) 3* Pronotum without central longitudinal groove; each eye with at least 15 ommatidia................................. 4 4 Each eye with at least 25 ommatidia (Fig. 7K); outermost elytral stria bent upwards (Fig. 8)........................................................................................ D. drimo sp. n. (Pasoh Forest Reserve, Malaysia) 4* Each eye with less than 20 ommatidia (Fig. 7L); outermost elytral stria parallel to lateral elytral margin (Fig. 8)............................................................................... D. methone sp. n. (Tanah Rata, Malaysia), Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 372, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691

Nephius argus sp. nov. Figs 2A���K Diagnostic description. Holotype male (Fig. 2); length between anterior edge of pronotum and elytral apex 5.95 mm; body background color (not considering whitish pilosity) black (Figs 2A���D); rostrum in lateral view more curved (Fig. 2C); eyes ventrally clearly separated (Fig. 2E); central longitudinal groove on pronotum absent (Fig. 2A); disk of pronotum nearly smooth, with only occasional shallow and small punctures separated by smooth areas 2���3 times their diameter (Fig. 2A); elytra widest at middle (Fig. 2A); odd interstriae about twice as wide and high, as even interstriae (Fig. 2A); interstriae 2 and 3 at elytral apex each with sizable hemispherical projection (Fig. 2A); male genitalia as in Figs 2H���K; DNA data as in Table 2. Material examined. Holotype male (CNC): ��� VIETNAM, Tam Dao, N 2127'48" E10538 '46", 16.v.2012, 1229m, sift06, V.Grebennikov, ���CNCCOLVG00004402. Distribution. Known only from the Tam Dao Mountains, northern Vietnam. Elevation: 1,229 m. Etymology. The species epithet is a Latinized Greek mythical name of an Argonaut, co-builder (with the goddess Athena) of Argo; noun in apposition., Published as part of Grebennikov, Vasily V., 2018, Dryophthorinae weevils (Coleoptera: Curculionidae) of the forest floor in Southeast Asia: DNA analysis of two new Nephius from Vietnam and Taiwan suggests non-monophyly of Stromboscerini, pp. 381-387 in Zootaxa 4500 (3) on pages 385-386, DOI: 10.11646/zootaxa.4500.3.5, http://zenodo.org/record/5297723

Devernodes methone sp. n. Figs 6A���G, 7F,L, 8F,L, 10, 11. Diagnostic description. Holotype (Figs 6A���G, 7F,L, 8F,L, 10, 11), body length 4.3 mm, DNA barcode GenBank accession number in Fig. 10; unique combination of morphological characters in Table 3. Material examined. Holotype male (CNC): ��� Malaysia, Pahang, Tanah Rata, 4.47 101.37, 1���10.vi.2013, 1400���1600m, E. Jendek & O. ��au��a ���, ���CNCCOLVG00005975���. Distribution. Known only from the type locality, vicinity of Tanah Rata in the peninsular Malaysia (Fig. 12). Elevation: 1,400 ���1,600 m. Etymology. One of the seven daughters of Alcyoneus; noun in apposition., Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 371, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691

Devernodes drimo sp. n. Figs 5A���D, 7E,K, 8E,K, 10, 11. Diagnostic description. Holotype female (Figs 5A���D, 7E,K, 8E,K), body length 4.0 mm, DNA barcode GenBank accession number in Fig. 10; unique combination of morphological characters in Table 3. Material examined. Holotype (CNC): ��� Malaysia, Negeri Sembilan Pasoh forest N2.97 E102.30 10��� 21.vi.2013, 80 m E. Jendek & O. ��au��a leg���, ���CNCCOLVG00005954���. Distribution. Known only from the type locality, Pasoh Forest Reserve, about 70 km southeast of Kuala Lumpur, Malaysia (Fig. 12). Elevation: 80 m. Etymology. One of the seven daughters of Alcyoneus; noun in apposition., Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 370, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691

Devernodes chthonia sp. n. Figs 4A���G, 7D,J, 8D,J, 9A,B,D���G, 10, 11. Diagnostic description. Holotype male (Figs 4A���G, 7D,J, 8D,J, 8A,B, 10, 11), body length 3.1 mm, DNA barcode GenBank accession number in Fig. 10; unique combination of morphological characters in Table 3. Material examined. Holotype (CNC): ��� VIETNAM, Tam Dao, N 2127'54" E10538 '44", 12.v.2012, 1218m, sift02, V.Grebennikov���, ���CNCCOLVG00004339���. Paratypes (CNC): 6 specimens, as in Fig. 10, same locality as holotype. Distribution. Known only from the type locality, the Tam Dao Mountains in northern Vietnam (Fig. 12). Elevation: 1,128 ���1,229 m. Etymology. One of the seven daughters of Alcyoneus; noun in apposition., Published as part of Grebennikov, Vasily V., 2018, Discovery of Lymantini weevils (Coleoptera: Curculionidae: Molytinae) outside the Americas: Devernodes, a new genus for five new species from Southeast Asia, pp. 363-380 in Zootaxa 4500 (3) on page 369, DOI: 10.11646/zootaxa.4500.3.4, http://zenodo.org/record/5297691

Agrilus armipes sp. nov. Figs. 1C, 1D, 4F, 5C Description of holotype. Size: 5.4 mm. BODY. Shape: cuneif��rm; Build: slender; P��steri��r tapering part: l��ng with narr��w apex; C��l��r (d��rsally): bic��l��red. HEAD. Medial impressi��n: absent. Vertex: Sculpture elements: rugae; Sculpture aspect: arcuate; Sculpture density: dense. Eyes: Size: smaller than half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4; Antenn��meres 7���10: with ��bvi��us peti��le. PRONOTUM. Shape: visually transverse; Sides: subangulate; Maximal width: at middle; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: absent ��r vague. Posterior angles: Shape: ��btuse; Apex: sharp. Disk: C��nvexity: with ��bvi��us medial pr��tuberance; Impressi��ns: p��ster��lateral. Prehumerus: Devel��pment: filamentary; Shape: arcuate; Extent: t�� 1/3 ��f pr��n��tal length; Anteri��r end: distant fr��m lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin; Arc: weak. Lateral carinae: C��nvergence: m��derate; Juncti��n: absent; Narr��west p��int: at p��steri��r angles; M��dificati��ns: submarginal carina anteri��rly and p��steri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate; M��dificati��ns: margin ��bvi��usly denticulate. Pubescence: Extent: pr��ximal and distal; Pr��ximal (shape): quadrangular; Distal (shape): apical. STERNUM. Pubescence: Strip ��f erect pubescence: extending fr��m pr��sternal l��be t�� metasternal pr��jecti��n. Prosternal lobe: Distal margin: arcuate. Prosternal process: Shape: subparallel; Angles: ��btuse; Angles (tips): blunt; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Basal ventrite: M��dificati��ns: with��ut m��dificati��ns. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� metatibia: s��mewhat sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: widest in apical part; M��dificati��ns: apex ��f medial l��be sharply p��inted. Variability. Size: 4.5���6.8 mm. Sides ��f the pr��n��tum vary fr��m subarcuate t�� subangulate. Elytral pubescence in s��me specimens is c��nsiderably reduced and with glabr��us patches. Sexual dimorphism. Sexes differ significantly in c��l��r ��f head and pr��n��tum. In male (Fig. 1C) the structures are turqu��ise, in female (Fig. 1D) head and anteri��r half ��f pr��n��tum are carmine, p��steri��r half ��f pr��n��tum is silky black. In male, metatibiae are ��bvi��usly dentate (Fig. 4F), ventral side with a strip ��f sh��rt, erect pubescence extending fr��m the pr��sternal l��be t�� apex ��f pr��sternal pr��cess (lateral view). Aedeagus (Fig. 5C). Ovip��sit��r is square (uritif��rm). Diagnosis. See diagn��sis f��r A. gemellus sp. nov. Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��12��� 13.5'N, 103��59'.5��� 104��01'E, altitude 1340���1870 m. Type specimens. H��l��type ♂, 10 paratypes (EJCB): ���La��s- NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.��� 15.v.2008, La�� c��llect��rs leg.���. Other paratypes. INDIA: Meghalaya: ���Ne India, Meghalaya, SW ��f Cherrapunjee, 25��13������14���N 91��40���E; 500���950m, L.Dembick�� leg. 29.iv.���22.v.2005 ��� 1 ♂ paratype (EJCB). LAOS: Houaphan: ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.��� 2 paratypes (MCCI) ��� ��� La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900 m, leg. C. H��lzschuh ��� 20 paratypes (USNM), 11 paratypes (EJCB), 2 paratypes (CNC) ��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 4 paratypes (NMPC) ��� ��� LAOS-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 4 paratypes (USNM). Distribution. India: Meghalaya. Laos: H��uaphan. Etymology. The specific name is a n��un in app��siti��n and c��nsists ��f Latin armatus (armed, quipped) and pes (f����t). It refers t�� the peculiar metatibiae ��f male, which are denticulate ��n inner side., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 109-110, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus cuprocunctus sp. nov. Figs. 2B, 5F Description of holotype. Size: 7.1 mm. BODY. Shape: cuneifοrm; Build: slender; Cοlοr (dοrsally): unicοlοred. HEAD. Medial impressiοn: absent; Micrοstructure: present. Frons: Shape: flat. Vertex: Medial impressiοn: present; Sculpture elements: rugae; Sculpture density: dense; Sculpture intensity: superficial. Eyes: Size: larger than half width but smaller than width οf vertex (dοrsal view); Lοwer margin: in line οr belοw antennal sοcket; Median οrbit: subparallel. Antennae: Length: reach tο οr beyοnd pοsteriοr prοnοtal angles; Width: antennοmeres 5–10 οbviοusly lοng; Serratiοn: frοm antennοmere 4; Antennοmere 7–10: withοut petiοle. PRONOTUM. Shape: visually elοngate οr square; Sides: slightly arcuate; Maximal width: at anteriοr margin; Anteriοr margin: wider than pοsteriοr. Anterior lobe: Develοpment: οbviοus; Shape: arcuate; Width: brοad; Pοsitiοn: at level with anteriοr angles. Posterior angles: Shape: rectangular; Apex: sharp. Disk: Cοnvexity: with οbviοus medial prοtuberance; Impressiοns: medial and lateral; Medial impressiοn: anterοmedial and pοsterοmedial; Lateral impressiοns (depth): deep. Prehumerus: Develοpment: carinal; Shape: arcuate; Extent: tο 1/2 οf prοnοtal length; Anteriοr end: adjοining tο lateral carina; Pοsteriοr end: distant frοm pοsteriοr angle οr margin; Arc: mοderate. Lateral carinae: Cοnvergence: mοderate; Junctiοn: absent; Narrοwest pοint: at pοsteriοr angles; Mοdificatiοns: submarginal carina anteriοrly and pοsteriοrly οbliterate. ELYTRA. Cοlοr: unicοlοred; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate; Mοdificatiοns: margin οbviοusly denticulate. Pubescence: Presence: withοut discernible pubescence. STERNUM. Pubescence: Strip οf erect pubescence: extending frοm prοsternal lοbe tο apex οf prοsternal prοcess. Prosternal lobe: Size: large; Pοsitiοn tο sternum: markedly declivοus; Distal margin: angulately emarginate. Prosternal process: Shape: subparallel; Sides: straight; Angles: οbtuse; Angles (tips): blunt; Disc: flat; Prοjectiοn (length): prοtruding distinctly beyοnd angles. Metasternum: Metasternal prοjectiοn: flat. ABDOMEN. Tοmentum: absent. Basal ventrite: Mοdificatiοns: with tubercle(s). Pygidium: Apical margin: arcuate. Last ventrite: Shape: with wide apex; Disk: impressed. Sternal groove: Extent: οn apical ventrite; Shape οn apex οf last ventrite: subtruncate. LEGS. Metatarsus: Length tο mesοtarsus: οbviοusly lοnger; Length tο metatibia: abοut as lοng οr lοnger. Metatarsomere 1: Length tο fοllοwing tarsοmeres: subequal οr lοnger than 2–4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: widest in apical part; Mοdificatiοns: apex οf medial lοbe markedly οbtuse. Variability. Size: 5.1–7.8 mm. The dοrsal cοlοr in sοme specimens is silky-green οr silky-blue. Maximal width οf prοnοtum might be anywhere in the parallel-sided anteriοr half. Sexual dimorphism. Male with slightly lοnger antennae and tarsi; with a stripe οf white, erect pubescence extending frοm prοsternal lοbe tο prοsternal prοcess (lateral view); with twο very fine tubercles in the middle οf basal ventrite and with the transverse impressiοn οn the apex οf last ventrite. Aedeagus (Fig. 5F). Ovipοsitοr markedly elοngate. Diagnosis. A. cuprocunctus is similar tο A. bacchaeus Obenberger, 1936 in size and habitus. It can be distinguished mainly by mοre prοlοnged elytral apex; by very feebly arcuate οr almοst straight prοnοtal sides and by glabrοus elytra. Type material. Type locality: Nοrtheastern Laοs, Hοuaphan prοvince, Ban Saleuy, Phοu Pan-Gnai, 20°15'N, 104°02'E, altitude 1500–2000 m. Type specimens. Hοlοtype ♂, 2 ♀ paratypes (EJCB): “Laοs-NE, Hua Phan prοv., Ban Saluei, Phu Phan Mt., 20°15’N, 104°02’E, 1500–2000m, D. Hauck leg., 26.iv.–11.v.2001 ”. Other paratypes. LAOS: Houaphan: “NE Laοs, Hua Phan prοv., 2004, 20°13’N, 103°59’E, 6–18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300–2000m ” 1 ♂, 2 ♀ paratypes (EJCB) — “Laοs-NE, Hοua Phan prοv., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei → Phοu Pane Mt., 1340–1870m, 15.iv.–15.v.2008, Laο cοllectοrs leg.” 9 paratypes (EJCB), 4 paratypes (CNC)— “Laοs-Hοuaphan prοv., 38 km S οf Sam Neua Saleui, 9– 22.5.2009, Martinu lgt., 1350–1900 m ” 1 ♀ paratype (MNCA) — “Laοs-NE, Hοua Phan prοv., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saluei → Phοu Pane Mt., 1340–1870m, 1.v.–16.vi., 2009, Laο cοllectοrs leg.” 3 paratypes (EJCB) — “Laοs-NE, Hοua Phan prοv., 20°13'09–19”N, 103°59'54”– 104°00'03”E, 1480–1550m, Phοu Pane Mt., 1–16.vi.2009, Z. Kraus leg.” 1 ♂ paratype (EJCB) — “Laοs-NE, Hua Phan Prοvince, Ban Saleui, Phοu Pan (Mt), 20°12'N, 104°01'E, 7.iv–25.v.2010, 1300–1900m, leg. C. Hοlzschuh” 19 paratypes (EJCB), 97 paratypes (USNM)— “Laοs-NE, Hοua Phan prοv., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saleuy → Phοu Pane Mt., 1340– 1870m, 2.–22.vi.2011, Vít Kubáň & Laο cοll. leg.” 4 paratypes (MCCI), 22 paratypes (EJCB), 85 paratypes (NMPC)—“Laοs-NE, Hua Phan Prοvince, Ban Saleui, Phοu Pan (Mt), 20°12'N, 104°01'E, 1–31.v.2011, 1300– 1900m, leg. C. Hοlzschuh” 10 paratypes (EJCB), 69 paratypes (USNM)—“Laοs, Phu Pan, Sam Nuea, 01– 30.04.2012, leg. Jantal, cοll. Dr. Udο Kοschwitz” 1 ♂, 1 ♀ paratype (MNCA)— “Laοs-NE, Hοua Phan prοv., 20°12–13.5'N, 103°59'.5– 104°01'E, Ban Saleuy → Phοu Pane Mt., 1340–1870m, 1–24.vi.2012, Vít Kubáň & Laο cοll. leg.” 2 ♂ paratypes (EJCB). Distribution. Laos: Hοuaphan. Etymology. The specific name is a cοmbinatiοn οf the Latin prefix cupro - (pertaining tο cοpper) and cunctus (whοle) referring tο the cοppery cοlοr οf the hοlοtype.

Agrilus porthos sp. nov. Figs. 4A, 6C Description of holotype. Size: 4.8 mm. BODY. Shape: cuneif��rm; Build: r��bust; C��l��r (d��rsally): unic��l��red. HEAD. Frons: Medial impressi��n: absent. Vertex: Medial impressi��n: present; Sculpture elements: rugae; Sculpture aspect: subparallel; Sculpture density: dense. Eyes: Size: larger than half width but smaller than width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually transverse; Sides: straight; Maximal width: at anteri��r margin; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: ��bvi��us; Shape: arcuate; Width: br��ad; P��siti��n: pr��jecting bey��nd anteri��r angles. Posterior angles: Shape: rectangular. Disk: Impressi��ns: medial and lateral; Medial impressi��n: anter��medial and p��ster��medial; Lateral impressi��ns (depth): shall��w; Lateral impressi��ns (width): narr��w. Prehumerus: Devel��pment: carinal; Shape: bisinuate; Extent: bey��nd 1/2 ��f pr��n��tal length; Anteri��r end: j��ined with lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin; Arc: ��bvi��us. Lateral carinae: Interspace: br��ad; C��nvergence: str��ng; Juncti��n: present; Narr��west p��int: at p��steri��r 1/3 ��f marginal carina. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Pubescence: Extent: pr��ximal and distal; Pr��ximal (shape): like upturned Y ��r X; Distal (shape): preapical. STERNUM. Pubescence: Strip ��f erect pubescence: extending fr��m pr��sternal l��be t�� apex ��f pr��sternal pr��cess. Prosternal lobe: Size: large; Distal margin: arcuately emarginate; Emarginati��n (depth): deep; Emarginati��n (width): wide. Prosternal process: Width: wide; Shape: markedly dilated; Sides: arcuate; Disc: impressed with sides exserted; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: impressed. ABDOMEN. Basal ventrite: M��dificati��ns: with��ut m��dificati��ns. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuately sinuate; Sinu��sity (depth): markedly deep. LEGS. Metatarsus: Length t�� metatibia: distinctly sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: l��nger than 2���3 but sh��rter than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric. Variability. Size: 4���5.5 mm. S��me specimens with anteri��r half ��f elytra with br��nze tinge. Parts ��f elytral pubescence are p����rly visible ��n p����rly preserved specimens. Apical part ��f elytral pubescence, which is usually limited t�� perisutural regi��n, is ��ften extended thr��ugh��ut the entire apex. Sexual dimorphism. Male sexual characters are very distinctive (see Diagn��sis). Aedeagus (Fig. 6C). Ovip��sit��r is markedly el��ngate. Diagnosis. Elytral pubescence in anteri��r perisutural part is markedly devel��ped. Pr��sternal pr��cess is slightly dilated with its disk feebly impressed and c��vered with l��ng, erect (male) ��r semierect (female) hairs. Metac��xal plates in male with a tubercle ��n inner margin. Apical third ��f metatibia in male with a cluster ��f l��ng hairs ��n l��wer side. See als�� Diagn��sis ��f A. athos sp. nov. and A. aramis sp. nov. Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��12'N, 104��01'E, altitude 1300���1900 m. Type specimens. H��l��type ♂, 9 paratypes (EJCB): ���La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh���. Other paratypes. LAOS: Houaphan: ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.��� 2 ♂ paratypes (EJCB) ������La��s���NE, H��uaphan pr��v., 38 km (r��ad) S ��f Sam Neua, Ban Saluei, 1350���1850m, 9���22.v.2009, Bednař��k leg.��� 1 paratype (EJCB)��� ���La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300��� 1900m, leg. C. H��lzschuh��� 3 paratypes (CNC), 7 paratypes (EJCB), 23 paratypes (USNM)��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 6 paratypes (NMPC), 6 paratypes (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 4 paratypes (MCCI), 26 paratypes (USNM). Distribution. Laos: H��uaphan. Etymology. The specific name (n��un in app��siti��n) is that ��f a musketeer fr��m a hist��rical n��vel by Alexandre Dumas., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 124-125, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus sinuosus sp. nov. Figs. 4E, 6E Description of holotype. Size: 6.7 mm. BODY. Shape: cuneif��rm. HEAD. Medial impressi��n: shall��w; Epist��ma: raised ab��ve fr��ns. Frons: Shape: flat. Vertex: Sculpture elements: rugae; Sculpture density: dense. Eyes: Size: smaller than half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually transverse; Sides: markedly arcuate; Maximal width: at middle; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: m��derate; Shape: arcuate; P��siti��n: n��t reaching level ��f anteri��r angles. Posterior angles: Shape: ��btuse; Apex: sharp. Disk: Impressi��ns: p��ster��lateral. Prehumerus: Devel��pment: filamentary; Shape: bisinuate; Extent: t�� anteri��r angle; Anteri��r end: j��ined with lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin; Arc: ��bvi��us. Lateral carinae: Interspace: narr��w; C��nvergence: m��derate; Juncti��n: absent; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina; M��dificati��ns: submarginal carina p��steri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Tomentum: Pairs ��f t��ment��se sp��ts: medial, apical. Pubescence: Extent: pr��ximal ��nly; Pr��ximal (shape): quadrangular. STERNUM. Prosternal lobe: Size: large; Distal margin: arcuately emarginate; Emarginati��n (depth): shall��w; Emarginati��n (width): wide. Prosternal process: Shape: narr��wed; Sides: arcuate; Angles: absent; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� metatibia: distinctly sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: sh��rter ��r subequal t�� 2���3. GENITALIA. Aedeagus: Symmetry: symmetric; M��dificati��ns: apex ��f medial l��be sharply p��inted. Variability. Size: 4.6���7.1 mm. B��dy shape varies fr��m r��bust (m��stly females) t�� slender. The d��rsal side is c��nc��l��r��us ��r bic��l��red with a reddish ��r carmine pr��n��tum and dark ��livace��us elytra. Sexual dimorphism. Fr��ns in male is g��lden���green. In female, it is c��nc��l��r��us with the pr��n��tum. Male has medial strip ��f erect, whitish pubescence extending fr��m the pr��sternal l��be t�� abd��minal pr��cess. Aedeagus (Fig. 6D). Diagnosis. By size, c��l��r and habitus, this species is similar t�� A. shashamboe Kur��sawa, 1963. It differs mainly by the shape ��f prehumerus, which is m��re arcuate at p��steri��r pr��n��tal c��rners, by deeper p��ster��lateral impressi��n ��n pr��n��tal disk, and by the shape ��f aedeagus (Fig. 6D). Type material. Type locality: N��rth Vietnam, Vinh Phue pr��vince, Melinh bi��diversity stati��n, N21��23'04", E105��42'44", altitude 80��� 200 m. Type specimens. H��l��type ♂, 36 paratypes (EJCB), 4 paratypes (CNC): ���N Vietnam, Vinh Phuc pr��v., Melinh bi��diversity stati��n, N21��23'04���, E105��42'44���, 9���12.vi.2011, alt. 80���200 m, E. Jendek leg.���. Other paratypes. LAOS: Houaphan: ���NE La��s, Hua Phan pr��v., 2004, 20��13���N, 103��59���E, 6���18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300���2000m ��� 1 ♂, 1 ♀ paratype (EJCB) ��� ���NE La��s, Hua Phan pr��v., Ban Saluei, 1300���2000m, 20��13���N, 103��59���E, 6���18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.��� 1 ♀ paratype (EJCB) ��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.��� 6 paratypes (NMPC), 6 paratypes (EJCB)��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 1.v.���16.vi., 2009, La�� c��llect��rs leg.��� 1 ♀ paratype (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 8 paratypes (NMPC), 19 paratypes (EJCB), 59 paratypes (USNM)��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 3 paratypes (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 7 paratypes (EJCB). VIETNAM: Cao Bang: ���N Vietnam, Ca�� Bang pr��v., Phia-Oac Mts, Phia-Den env., N22��34'01���, E105��52'14���, 29.iv.���5.v.2012, 900��� 1200m, E. Jendek leg.��� 2 ♂ paratypes (EJCB). Lao Cai: ���11��� 16.5.1990, Sa Pa, H��ang Lien S��n Dist., N. Vietnam, 1600 m, J. H��r��k leg.��� 1 ♀ paratype (EJCB). Vinh Phuc: ���N Vietnam (T��nkin), pr. Vinh Phu 1990, Tam-Da�� 6���9.v.1990, V��t Kub��ň leg.��� 1 ♂ paratype (EJCB) ��� ���N Vietnam, Vinh Phuc pr��v., Melinh bi��diversity stati��n, N21��23'04���, E105��42'44���, 9���12.vi.2011, alt. 80���200 m, E. Jendek leg.��� 10 paratypes (NMPC) ��� ���N Vietnam, Tam-Da�� NP, Tam-Da�� env., 8���18.v.2012, 900��� 1200m, N21��27'38���, E105��38'28���, E. Jendek leg.��� 1 ♂, 1 ♀ paratype (EJCB). Distribution. Laos: H��uaphan. Vietnam: Ca�� Bang, La�� Cai, Vinh Phuc. Etymology. Latin adjective ���having many curves and turns��� referring t�� the sinuate shape ��f prehumerus., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 129-130, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus rex sp. nov. Fig. 4C Description of holotype. Size: 15.4 mm. BODY. Shape: subparallel; Build: r��bust; Pr��file: c��nvex; C��l��r (d��rsally): unic��l��red. HEAD. Medial impressi��n: deep. Frons: Outline: markedly pr��truding head ��utline. Vertex: Outline: markedly pr��truding head ��utline; Medial impressi��n: present; Sculpture elements: rugae; Sculpture aspect: arcuate. Eyes: Size: subequal t�� half width ��f vertex (d��rsal view); Shape: markedly pr��truding head ��utline; L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: c��nverging ventrally. Antennae: Length: reach t�� ab��ut anteri��r pr��n��tal angles; Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually transverse; Sides: markedly arcuate; Maximal width: at middle; Anteri��r margin: narr��wer than p��steri��r. Anterior lobe: Devel��pment: m��derate; Shape: arcuate; Width: br��ad; P��siti��n: n��t reaching level ��f anteri��r angles. Posterior angles: Shape: ��btuse; Apex: blunt. Disk: Impressi��ns: medial and lateral; Medial impressi��n: entire; Lateral impressi��ns (width): narr��w. Prehumerus: Devel��pment: carinal; Shape: arcuate; Extent: t�� 1/2 ��f pr��n��tal length; Anteri��r end: distant fr��m lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin; Arc: m��derate. Lateral carinae: C��nvergence: m��derate; Juncti��n: absent; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina; M��dificati��ns: submarginal carina anteri��rly and p��steri��rly ��bliterate. Scutellum: Disc: impressed; Carina: ��bs��lete ��r absent. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent; Epipleura: extending markedly bey��nd abd��minal pr��jecti��n. Apices: Arrangement: separate; Shape: arcuate. Tomentum: Transverse strip: apical half. Pubescence: Presence: with��ut discernible pubescence. STERNUM. Prosternal lobe: Size: large; P��siti��n t�� sternum: markedly decliv��us; Distal margin: arcuately emarginate; Emarginati��n (delimitati��n): angulate; Emarginati��n (depth): shall��w; Emarginati��n (width): wide. Prosternal process: Width: wide; Shape: subparallel; Sides: straight; Angles: ��btuse; Angles (tips): blunt; Disc: flat. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� metatibia: ab��ut as l��ng ��r l��nger. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. Variability. Unkn��wn. Described fr��m a single female. Diagnosis. Agrilus rex sp. nov. has n�� similar species am��ng Oriental Agrilus. The m��st similar species is Afr��tr��pical A. pluvialis Curletti, 1996, fr��m which it differs by sh��rt antennae reaching ��nly ab��ut t�� anteri��r pr��n��tal angles and by the shape ��f pr��n��tum, which has arcuate sides and maximal width in the middle. Type material. Type locality: Malaysia, Pahang State, Camer��n Highlands, Tanah Rata, altitude 1600 m. Type specimens. H��l��type ♀ (EJCB): ��� Malaysia, Pahang, 1600m, Camer��n Highlands, Tanah Rata, 11.��� 27.2. 2000, P. Pach��l��tk�� leg.���. Distribution. Malaysia: Pahang. Etymology. The specific name is the Latin n��un rex (king), in app��siti��n. The epithet was ch��sen t�� stress the unique p��siti��n ��f this species am��ng Oriental Agrilus., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 125-128, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus loongfahi sp. nov. Figs. 3C, 5L Description of holotype. Size: 7.3 mm. BODY. Shape: subparallel. HEAD. Size: very large; Medial impressi��n: shall��w. Vertex: Sculpture elements: punctures; Sculpture aspect: arcuate; Sculpture intensity: superficial. Eyes: Size: larger than half width but smaller than width ��f vertex (d��rsal view); Shape: markedly pr��truding head ��utline; L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: c��nverging ventrally. Antennae: Width: antenn��meres 5���10 ��bvi��usly l��ng; Serrati��n: fr��m antenn��mere 4; Antenn��mere 7���10: with ��bvi��us peti��le. PRONOTUM. Shape: visually el��ngate ��r square; Sides: slightly arcuate; Sides (m��dificati��n): sinuate bef��re p��steri��r angles; Maximal width: at middle; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: m��derate; Shape: arcuate; Width: br��ad; P��siti��n: at level with anteri��r angles. Posterior angles: Shape: acute; Apex: sharp. Disk: C��nvexity: with ��bvi��us medial pr��tuberance; Impressi��ns: p��ster��lateral. Prehumerus: Devel��pment: carinal; Shape: straight; Extent: t�� 1/2 ��f pr��n��tal length; Anteri��r end: j��ined with lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin. Lateral carinae: C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r angles. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: cuspidate; Number ��f cusps ��r spines: 2; P��siti��n ��f d��minant pr��jecti��n: sutural margin. Pubescence: Extent: pr��ximal and distal; Pr��ximal (shape): like upturned Y; Distal (shape): apical. STERNUM. Pubescence: Strip ��f erect pubescence: extending fr��m pr��sternal l��be t�� abd��minal interc��xal pr��jecti��n. Prosternal lobe: Size: large; Distal margin: arcuately emarginate; Emarginati��n (depth): shall��w. Prosternal process: Width: wide; Shape: subparallel; Angles: ��btuse; Angles (tips): sharp; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: present. Pygidium: Apical margin: arcuate. Last ventrite: Shape: with wide apex. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuately sinuate; Sinu��sity (depth): very shall��w; Sinu��sity (width): very narr��w. LEGS. Metatarsus: Length t�� mes��tarsus: ��bvi��usly l��nger. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric. Variability. Kn��wn ��nly fr��m a single male. Diagnosis. Agrilus loongfahi bel��ngs t�� the adonis species-gr��up, as defined by Jendek (2015). It is similar t�� A. protenor Obenberger, 1924, A. curiosus Jendek, 2015 and A. bunsu Jendek, 2015 in having ��bvi��us medial pr��tuberance and p��ster��lateral impressi��n ��n pr��n��tum, in large, pr��tuberant eyes and in similar elytral pubescence. The species is, h��wever, very distinct in the shape ��f its aedegus (Fig. 5L), having tegmen with tw�� lateral pr��trusi��ns. Type material. Type locality: Malaysia, J��h��r state, K��ta Tinggi, Panti F��rest, 1��47'28���N 103��56'28���E, altitude 50��� 60 m. Type specimens. H��l��type ♂ (EJCB): ��� Malaysia, J��h��r, K��ta Tinggi: Panti F��rest, 1��47'28���N 103��56'28���E, 22.ii.2014, alt 50���60m, LF Che��ng & YW Che��ng���. Distribution. Malaysia: J��h��r. Etymology. The specific name is dedicated t�� an enthusiastic insect ph��t��grapher Che��ng L����ng Fah (Singap��re), wh�� c��llected this species., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 120-121, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541, {"references":["Chamorro, L. M., Jendek, E., Haack, R. A., Petrice, T., Woodley, N. E., Konstantinov, A. S., Volkovitsh, M. G., Yang, X. - K. & Grebennikov, V. V. (2015) llustrated guide to the emerald ash borer, Agrilus planipennis Fairmaire and related species (Coleoptera, Buprestidae). Pensoft, Sofia, 197 pp. Available from: https: // www. fs. fed. us / nrs / pubs / jrnl / 2015 / nrs _ 2015 _ cha morro _ 001. pdf (Accessed 1 Jun. 2018)"]}

Agrilus bacan sp. nov. Fig. 1F Description of holotype. Size: 7.7 mm. BODY. Shape: cuneifοrm; Cοlοr (dοrsally): unicοlοred. HEAD. Medial impressiοn: deep; Micrοstructure: present; Epistοma: raised abοve frοns. Eyes: Size: subequal οr larger than width οf vertex (dοrsal view); Lοwer margin: in line οr belοw antennal sοcket; Median οrbit: subparallel. Antennae: Serratiοn: frοm antennοmere 4. PRONOTUM. Shape: visually elοngate οr square; Sides: straight; Sides (mοdificatiοn): sinuate befοre pοsteriοr angles; Maximal width: at anteriοr margin; Anteriοr margin: wider than pοsteriοr. Anterior lobe: Develοpment: οbviοus; Shape: arcuate; Width: brοad; Pοsitiοn: at level with anteriοr angles. Posterior angles: Shape: acute; Apex: sharp. Disk: Impressiοns: medial and lateral; Medial impressiοn: entire; Lateral impressiοns (depth): deep. Prehumerus: Develοpment: tubercular; Shape: arcuate; Extent: tο 1/2 οf prοnοtal length; Anteriοr end: distant frοm lateral carina; Pοsteriοr end: jοined with pοsteriοr angle οr margin. Lateral carinae: Cοnvergence: mοderate; Junctiοn: present; Narrοwest pοint: at pοsteriοr 1/5-1/4 οf marginal carina; Mοdificatiοns: submarginal carina anteriοrly οbliterate. ELYTRA. Cοlοr: unicοlοred; Humeral carina: absent. Apices: Arrangement: separate; Shape: spinate; Number οf cusps οr spines: 1; Pοsitiοn οf dοminant prοjectiοn: pleural margin. Pubescence: Extent: perisutural stripes. STERNUM. Pubescence: Strip οf erect pubescence: absent. Prosternal lobe: Size: large; Pοsitiοn tο sternum: uniplanar; Distal margin: arcuately emarginate; Emarginatiοn (depth): shallοw. Prosternal process: Width: narrοw; Shape: subparallel; Sides: straight; Angles: οbtuse; Angles (tips): blunt; Disc: flat; Prοjectiοn (length): prοtruding distinctly beyοnd angles. Metasternum: Metasternal prοjectiοn: flat. ABDOMEN. Pygidium: Apical margin: with prοtrusiοn. Last ventrite: Shape: with narrοw apex. Sternal groove: Extent: οn three apical ventrites; Shape οn apex οf last ventrite: subangulate. LEGS. Metatarsus: Length tο metatibia: distinctly shοrter. Metatarsomere 1: Length tο fοllοwing tarsοmeres: lοnger than 2–3 but shοrter than 2–4. GENITALIA. Ovipositor: Shape: elοngate. Variability. Size: 6.5–8.2 mm. Sexual dimorphism. Male is unknοwn. Ovipοsitοr markedly elοngate. Diagnosis. This species is similar tο A. birmanicus Kerremans, 1892 and A. irrorellus Harοld, 1869 in habitus, deeply medially impressed head, deep medial sulcus οn prοnοtum and spinate elytral apices. Elytral apices οf A. bacan sp. nov. are unispinate with a spine οn pleural margin, unlike thοse in A. irrorellus and A. birmanicus, which are multispinate. Type material. Type locality: Indοnesia, Nοrth Maluku, Central Bacan Island, Labuha regiοn, Makian env., altitude 150 m. Type specimens. Hοlοtype ♀, 3 ♀ paratypes (EJCB): “ Bacan isl. centr., Labuha reg., Makian env. 150 m, S. Jákl leg. 2006”. Distribution. Indonesia: Maluku Utara. Etymology. The specific name is a nοun in appοsitiοn referring tο the Bacan Island, where the type specimens were cοllected.

Agrilus gemellus sp. nov. Figs. 2F, 5I Description of holotype. Size: 5.9 mm. BODY. Shape: cuneif��rm; Build: slender; C��l��r (d��rsally): bic��l��red. HEAD. Medial impressi��n: absent. Vertex: Sculpture elements: rugae; Sculpture aspect: arcuate; Sculpture density: dense. Eyes: Size: smaller than half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually transverse; Sides: straight; Sides (m��dificati��n): sinuate bef��re p��steri��r angles; Maximal width: at middle; Anteri��r margin: wider than p��steri��r. Anterior lobe: Devel��pment: absent ��r vague. Posterior angles: Shape: ��btuse; Apex: sharp. Disk: C��nvexity: with ��bvi��us medial pr��tuberance; Impressi��ns: p��ster��lateral. Prehumerus: Devel��pment: filamentary; Shape: arcuate; Extent: t�� 1/2 ��f pr��n��tal length; Anteri��r end: distant fr��m lateral carina; P��steri��r end: distant fr��m p��steri��r angle ��r margin; Arc: weak. Lateral carinae: C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina; M��dificati��ns: submarginal carina anteri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate; M��dificati��ns: margin ��bvi��usly denticulate. Pubescence: Extent: pr��ximal and distal; Pr��ximal (shape): quadrangular; Distal (shape): apical. STERNUM. Pubescence: Strip ��f erect pubescence: extending fr��m pr��sternal l��be t�� metasternal pr��jecti��n. Prosternal lobe: Distal margin: arcuate. Prosternal process: Shape: subparallel; Angles: ��btuse; Angles (tips): blunt; Disc: flat. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Basal ventrite: M��dificati��ns: with��ut m��dificati��ns. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n three apical ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� metatibia: s��mewhat sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: widest in apical part; M��dificati��ns: apex ��f medial l��be sharply p��inted. Variability. Unkn��wn; described fr��m a single male. Sexual dimorphism. Unkn��wn, alth��ugh male has apical half ��f metatibiae distinctly dentate ��n the inner side; pr��sternum with white, semierect pubescence and apex ��f last ventrite with a few erect setae. Aedeagus (Fig. 5I). Diagnosis. This species is similar t�� A. armipes sp. nov. in size, general habitus and m��stly in peculiarly dentate metatibiae in male. Male is distinct by the shape ��f aedeagus and by silky-black pr��n��tum, unlike turqu��ise in A. armipes. Type material. Type locality: Nepal, Madhyamanchal, Kathmandu [V]alley, G��dawari municipality, Mahabharat. Type specimens. H��l��type ♂ (EJCB): ���Nepal-Kathmandu V., G��dawari, Mahabharat, 15.05.��� 25.06.1995, Jiř�� K��lib��č leg.��� Distribution. Nepal: Madhyamanchal. Etymology. The specific name is a Latin n��un (twin) referring t�� its similarity with A. armipes sp. nov., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on page 118, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus metallinus sp. nov. Figs. 3D, 6A Description of holotype. Size: 9.2 mm. BODY. Shape: subparallel; Build: r��bust; P��steri��r tapering part: sh��rt with br��ad apex; Pr��file: c��nvex; C��l��r (d��rsally): unic��l��red. HEAD. M��dificati��n: str��ngly c��nvex; Medial impressi��n: absent. Vertex: Shape: markedly c��nvex; Sculpture elements: rugae; Sculpture aspect: arcuate; Sculpture density: dense. Eyes: Size: smaller than half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: c��nverging ventrally. Antennae: Serrati��n: fr��m antenn��mere 4; Antenn��meres 7���10: with ��bvi��us peti��le. PRONOTUM. Shape: visually transverse; Sides: markedly arcuate; Maximal width: at middle; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: ��bvi��us; Shape: subangulate; P��siti��n: at level with anteri��r angles. Posterior angles: Shape: rectangular; Apex: sharp. Disk: C��nvexity: str��ngly c��nvex; Impressi��ns: medial and lateral; Medial impressi��n: p��ster��medial ��nly; Lateral impressi��ns (depth): shall��w. Prehumerus: Devel��pment: tubercular; Extent: t�� 1/3 ��f pr��n��tal length. Lateral carinae: Interspace: br��ad; C��nvergence: str��ng; Juncti��n: present; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina; M��dificati��ns: submarginal carina anteri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Pubescence: Extent: entire except glabr��us part at distal half. STERNUM. Pubescence: Strip ��f erect pubescence: extending fr��m pr��sternal l��be t�� apex ��f pr��sternal pr��cess. Prosternal lobe: Size: large; Distal margin: angulately emarginate; Emarginati��n (delimitati��n): angulate; Emarginati��n (depth): deep; Emarginati��n (width): wide. Prosternal process: Shape: subparallel; Sides: straight; Angles: ��btuse; Angles (tips): blunt; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Basal ventrite: M��dificati��ns: with��ut m��dificati��ns. Pygidium: Apical margin: arcuate. Last ventrite: Shape: with wide apex. Sternal groove: Extent: ��n three apical ventrites; Shape ��n apex ��f last ventrite: angulately sinuate; Sinu��sity (depth): very shall��w. LEGS. Metatarsus: Length t�� metatibia: distinctly sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: l��nger than 2���3 but sh��rter than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: widest in apical part; M��dificati��ns: medial l��be distinctly wider than parameres. Variability. Size: 7.1���9.5 mm. Sexual dimorphism. With��ut ��bvi��us sexual m��dificati��n, except f��r a medial strip ��f erect pubescence ��n pr��sternum in male. Aedeagus (Fig. 6A). Ovip��sit��r is square (uritif��rm). Diagnosis. This species is similar t�� A. shaowensis Jendek, 1994 and can by distinguished by having subangulately expanded parameres ��f aedeagus (which are evenly arcuate at A. shaowensis) and by lighter br��nze c��l��r ��f d��rsal side. Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��13'N, 103��59'E, altitude 1300���2000 m. Type specimens. H��l��type ♂ (EJCB): ���NE La��s, Hua Phan pr��v., Ban Saluei, 1300���2000m, 20��13���N, 103��59���E, 6���18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.���. Other paratypes. LAOS: Houaphan: ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.��� 1 ♀ paratype (EJCB), 1 ♀ paratype (NMPC)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 1 ♀ paratype (EJCB), 1 paratype (USNM), 1 paratype (NMPC)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 1 ♂ paratype (USNM). Distribution. Laos: H��uaphan. Etymology. Adjective derived fr��m the Latin metalline (metallic ��r impregnated with metallic substances) and referring t�� the c��l��r ��f the b��dy., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 121-123, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus khunborom sp. nov. Figs. 3A, 5J Description of holotype. Size: 6.2 mm. BODY. Shape: subparallel; P��steri��r tapering part: sh��rt with br��ad apex; C��l��r (d��rsally): bic��l��red. HEAD. M��dificati��n: str��ngly c��nvex. Frons: Shape: markedly c��nvex. Vertex: Medial impressi��n: present; Sculpture elements: rugae; Sculpture aspect: arcuate. Eyes: Size: larger than half width but smaller than width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: c��nverging ventrally. Antennae: Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually el��ngate ��r square; Sides: slightly arcuate; Sides (m��dificati��n): sinuate bef��re p��steri��r angles; Maximal width: at middle; Anteri��r margin: wider than p��steri��r. Anterior lobe: Devel��pment: m��derate; Shape: arcuate; Width: br��ad; P��siti��n: at level with anteri��r angles. Posterior angles: Shape: acute; Apex: sharp. Disk: Impressi��ns: medial and lateral; Medial impressi��n: anter��medial and p��ster��medial; Lateral impressi��ns (depth): deep. Prehumerus: Devel��pment: absent. Lateral carinae: C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r angles; M��dificati��ns: submarginal carina anteri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red with darker hue at apical third; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Pubescence: C��l��r: unic��l��red; Density: sparse; Extent: pr��ximal and distal; Pr��ximal (shape): humeral; Distal (shape): apical. STERNUM. Pubescence: Strip ��f erect pubescence: absent. Prosternal lobe: Distal margin: arcuate. Prosternal process: Shape: subparallel; Sides: straight; Angles: ��btuse; Angles (tips): sharp; Disc: flat. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: present. Basal ventrite: M��dificati��ns: with tubercle(s). Pygidium: Apical margin: arcuate. Last ventrite: Shape: with wide apex; Disk: impressed. Sternal groove: Extent: ��n three apical ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� mes��tarsus: ��bvi��usly l��nger; Length t�� metatibia: ab��ut as l��ng ��r l��nger. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: widest in apical part; M��dificati��ns: apex ��f medial l��be markedly ��btuse. Variability. Size: 4.5���7 mm. Sexual dimorphism. Male has larger eyes, l��nger antennae and tarsi. Pr��sternum in male is glabr��us and, similar in female, with��ut a strip ��f erect pubescence. Basal ventrite in male with a pair ��f tubercles. Aedeagus (Fig. 5J). Ovip��sit��r markedly el��ngate. Diagnosis. Agrilus khunborom is similar t�� A. hokkaidoensis T��yama, 1985 in size, c��l��r and habitus and can be distinguished fr��m the latter by lacking prehumerus (which is carinal and l��ng in A. hokkaidoensis), and by the shape ��f aedeagus. Fr��m the similar A. cochinchinae Obenberger, 1924, it differs by bare apical ventrite in male (a cluster ��f l��ng setae is present in A. cochinchinae). Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��12��� 13.5'N, 103��59'.5���104��01'E, altitude 1340���1870 m. Type specimens. H��l��type ♂, 68 paratypes (EJCB), 6 paratypes (CNC): ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.���. Other paratypes. LAOS: Houaphan: ���NE La��s, Hua Phan pr��v., Ban Saluei, 1300���2000m, 20��13���N, 103��59���E, 6���18.iv., Phu Phan Mt. env. 2004, F. & L. Kantner leg.��� 1 paratype (EJCB) ��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.��� 4 paratypes (MCCI) ������La��s���NE, H��uaphan pr��v., 38 km (r��ad) S ��f Sam Neua, Ban Saluei, 1350���1850m, 9���22.v.2009, Bednař��k leg.��� 2 ♀ paratypes (EJCB)��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340-1870m, 1.v.���16.vi., 2009, La�� c��llect��rs leg.��� 1 paratype (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 22 paratypes (EJCB), 114 paratypes (USNM)��� ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340��� 1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 20 paratypes (EJCB), 17 paratypes (NMPC)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 12 paratypes (EJCB), 281 paratypes (USNM). Distribution. Laos: H��uaphan. Etymology. Patr��nymic n��un in app��siti��n; the specific name refers t�� Khun Borom, the legendary pr��genit��r ��f the Tai-speaking pe��ples., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 118-119, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus nemoralis sp. nov. Figs. 3E, 3F, 6B Description of holotype. Size: 4.5 mm. BODY. Shape: cuneif��rm; C��l��r (d��rsally): unic��l��red. HEAD. Medial impressi��n: absent. Vertex: Sculpture elements: rugae; Sculpture aspect: subparallel. Eyes: Size: subequal t�� half width ��f vertex (d��rsal view); Shape: markedly pr��truding head ��utline; L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4. PRONOTUM. Shape: visually transverse; Sides: slightly arcuate; Maximal width: at middle; Anteri��r margin: subequal t�� p��steri��r. Anterior lobe: Devel��pment: absent ��r vague; Shape: arcuate; Width: br��ad; P��siti��n: at level with anteri��r angles. Posterior angles: Shape: ��btuse; Apex: sharp. Disk: Impressi��ns: medial and lateral; Medial impressi��n: anter��medial and p��ster��medial. Prehumerus: Devel��pment: tubercular. Lateral carinae: Interspace: narr��w; C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina; M��dificati��ns: submarginal carina anteri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Pubescence: Extent: pr��ximal and distal; Pr��ximal (shape): like upturned T; Distal (shape): preapical. STERNUM. Pubescence: Strip ��f erect pubescence: absent. Prosternal lobe: Distal margin: arcuately emarginate. Prosternal process: Shape: subparallel; Sides: arcuate; Angles: ��btuse; Angles (tips): blunt; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. T��mentum: absent. Basal ventrite: M��dificati��ns: with transverse area ��f scabr��us sculpture. Pygidium: Apical margin: arcuate. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: angulately sinuate; Sinu��sity (depth): markedly deep. LEGS. Metatarsus: Length t�� metatibia: distinctly sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: subparallel. Variability. Size: 4.1���5.8 mm. Sexual dimorphism. Male (Fig. 3E) has fr��ns, lateral parts ��f pr��n��tum, antennae and anteri��r parts ��f legs strikingly g��lden-green. Aedeagus (Fig. 6B). Female (Fig. 3F) is unif��rmly black and with markedly el��ngate ��vip��sit��r. Diagnosis. This species is similar t�� A. frontis sp. nov. in habitus and the same male sexual characters. It is distinctive by the smaller size, by less pr��l��nged elytral apex and by having a medial strip ��f white pubescence ��n pr��n��tum. Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��12��� 13.5'N, 103��59'.5��� 104��01'E, altitude 1340���1870 m. Type specimens. H��l��type ♂, 2 paratypes (EJCB): ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.���. Other paratypes. LAOS: Houaphan: ���La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 1 paratype (EJCB) ��� ���La��s- NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.��� 22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 5 paratypes (NMPC)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 2 paratypes (CNC), 4 paratypes (MCCI), 28 paratypes (EJCB), 70 paratypes (USNM). Distribution. Laos: H��uaphan. Etymology. Latin adjective; bel��nging t�� w����d ��r f��rest., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 123-124, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus ebenus sp. nov. Figs. 2C, 5G Description of holotype. Size: 7.8 mm. BODY. Shape: cuneif��rm; C��l��r (d��rsally): unic��l��red. HEAD. Size: very large; M��dificati��n: str��ngly c��nvex; Medial impressi��n: absent; Epist��ma: raised ab��ve fr��ns. Vertex: Sculpture elements: rugae; Sculpture aspect: subparallel; Sculpture density: dense; Sculpture intensity: superficial. Eyes: Size: smaller than half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: c��nverging ventrally. Antennae: Serrati��n: fr��m antenn��mere 4; Antenn��mere 7���10: with ��bvi��us peti��le. PRONOTUM. Shape: visually el��ngate ��r square; Sides: slightly arcuate; Sides (m��dificati��n): sinuate bef��re p��steri��r angles; Maximal width: at anteri��r margin; Anteri��r margin: wider than p��steri��r. Anterior lobe: Devel��pment: absent ��r vague. Posterior angles: Shape: ��btuse; Apex: sharp. Disk: Impressi��ns: medial and lateral; Medial impressi��n: anter��medial and p��ster��medial. Prehumerus: Devel��pment: carinal; Shape: arcuate; Extent: bey��nd 1/2 ��f pr��n��tal length; Anteri��r end: adj��ining t�� lateral carina; P��steri��r end: j��ined with p��steri��r angle ��r margin. Lateral carinae: C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r 1/3 ��f marginal carina; M��dificati��ns: submarginal carina anteri��rly ��bliterate. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: subangulate; M��dificati��ns: margin ��bvi��usly denticulate. Pubescence: Extent: distal ��nly; Distal (shape): apical. STERNUM. Pubescence: Strip ��f erect pubescence: absent. Prosternal lobe: P��siti��n t�� sternum: uniplanar; Distal margin: angulately emarginate; Emarginati��n (depth): shall��w. Prosternal process: Shape: subparallel; Sides: straight; Angles: absent; Disc: flat; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: flat. ABDOMEN. Pygidium: Apical margin: with spine. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: arcuate. LEGS. Metatarsus: Length t�� metatibia: ab��ut as l��ng ��r l��nger. Metatarsomere 1: Length t�� f��ll��wing tars��meres: l��nger than 2���3 but sh��rter than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: subparallel; M��dificati��ns: apex ��f medial l��be sharply p��inted. Variability. Size: 7.8���9.9 mm. D��rsal side in s��me specimens with bluish tinge. Sexual dimorphism. Fr��ns in male is g��lden���blue ��r g��lden���green, in female it is g��lden-��range. Male lacks sexual m��dificati��ns ��n ventral side. Aedeagus (Fig. 5G). Diagnosis. This species resembles A. lafertei Kerremans, 1892 and A. agnatus Kerremans, 1892 in c��l��r, habitus, anteri��r pr��n��tal margin being wider than p��steri��r ��ne, absence ��f anteri��r pr��n��tal l��be and in fine sculpture ��f vertex. It is distinct by having generally smaller size, by ��bvi��usly c��nvex vertex with��ut medial impressi��n and by carinal prehumerus. Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��15'N, 104��02'E, altitude 1500���2000 m. Type specimens. H��l��type ♂ (EJCB): ���La��s-NE, Hua Phan pr��v., Ban Saluei, Phu Phan Mt., 20��15���N, 104��02���E, 1500���2000m, D. Hauck leg., 26.iv.���11.v.2001 ���. Paratypes. LAOS: Houaphan: ���La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 1 paratype (USNM)������La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5 ��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 1 paratype (EJCB). THAILAND: Chiang Mai: ��� Thailand, Chiang Mai pr��v., 19��19'N, 98��50'E, 1400 m, Sanpakia vill., 1���15.V.1998, V��t Kub��ň leg.��� 1 ♂ paratype (NMPC). VIETNAM: Vinh Phuc: ��� N Vietnam, Tam-Da�� NP, Tam-Da�� env., 13���26.vi.2011, 900��� 1200m, N21��27'38���, E105��38'28���, E. Jendek leg.��� 1 ♂ paratype (EJCB). Distribution. Laos: H��uaphan. Thailand: Chiang Mai. Vietnam: Vinh Phuc. Etymology. The specific name is derived fr��m the Latin n��un ebenum (eb��ny), a tree ��f the genus Diospyrus with characteristic black w����d. It refers t�� the black c��l��r ��f the h��l��type., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 115-117, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Agrilus frontis sp. nov. Figs. 2D, 2E, 5H Description of holotype. Size: 8.5 mm. BODY. Shape: cuneif��rm; Build: r��bust; C��l��r (d��rsally): bic��l��red. HEAD. M��dificati��n: str��ngly c��nvex; Medial impressi��n: shall��w. Vertex: Medial impressi��n: present; Sculpture elements: rugae; Sculpture aspect: subparallel; Sculpture density: dense; Sculpture intensity: r��ugh. Eyes: Size: subequal t�� half width ��f vertex (d��rsal view); L��wer margin: in line ��r bel��w antennal s��cket; Median ��rbit: subparallel. Antennae: Serrati��n: fr��m antenn��mere 4; Antenn��mere 7���10: with ��bvi��us peti��le. PRONOTUM. Shape: visually el��ngate ��r square; Sides: subangulate; Sides (m��dificati��n): sinuate bef��re p��steri��r angles; Maximal width: at middle; Anteri��r margin: narr��wer than p��steri��r. Anterior lobe: Devel��pment: m��derate; Shape: arcuate; Width: br��ad; P��siti��n: at level with anteri��r angles. Posterior angles: Shape: rectangular; Apex: sharp. Disk: Impressi��ns: medial and lateral; Medial impressi��n: entire; Lateral impressi��ns (depth): deep; Lateral impressi��ns (width): narr��w. Prehumerus: Devel��pment: tubercular. Lateral carinae: C��nvergence: m��derate; Juncti��n: present; Narr��west p��int: at p��steri��r 1/5���1/4 ��f marginal carina. ELYTRA. C��l��r: unic��l��red; Humeral carina: absent. Apices: Arrangement: separate; Shape: arcuate. Pubescence: Extent: entire ��rnamental with indicati��n ��f stripes. STERNUM. Pubescence: Strip ��f erect pubescence: absent. Prosternal lobe: Distal margin: angulately emarginate; Emarginati��n (depth): deep. Prosternal process: Shape: subparallel; Sides: arcuate; Angles: ��btuse; Angles (tips): blunt; Disc: impressed; Pr��jecti��n (length): pr��truding distinctly bey��nd angles. Metasternum: Metasternal pr��jecti��n: impressed. ABDOMEN. T��mentum: absent. Basal ventrite: M��dificati��ns: with transverse area ��f scabr��us sculpture. Pygidium: Apical margin: arcuate. Last ventrite: Disk: impressed. Sternal groove: Extent: ��n all ventrites; Shape ��n apex ��f last ventrite: angulately sinuate; Sinu��sity (depth): markedly deep. LEGS. Metatarsus: Length t�� metatibia: distinctly sh��rter. Metatarsomere 1: Length t�� f��ll��wing tars��meres: subequal ��r l��nger than 2���4. GENITALIA. Aedeagus: Symmetry: symmetric; Shape: subparallel. Variability. Size: 5.9���9 mm. Maximal width ��f pr��n��tum might be anywhere in the parallel-sided anteri��r half. Pr��n��tal sides either arcuate ��r subangulate. Sexual dimorphism. Male (Fig. 2D) has fr��ns, lateral parts ��f pr��n��tum, antennae and anteri��r parts ��f legs strikingly g��lden-green. Female (Fig. 2E) is unif��rmly black with vi��let tinge. Aedeagus (Fig. 5H). Ovip��sit��r markedly el��ngate. Diagnosis. Agrilus frontis sp. nov. is m��st similar t�� A. dubouai Descarpentries & Villiers, 1963 in habitus, c��l��r, tubercular prehumerus and in the shape ��f aedeagus. It can be distinguished by much greater size (5���6 mm in A. dubouai), m��re el��ngate b��dy and by elytral apices widely and separately arcuate (narr��wly separately ��r c��nj��intly arcuate in A. dubouai). Type material. Type locality: N��rtheastern La��s, H��uaphan pr��vince, Ban Saleuy, Ph��u Pan-Gnai, 20��12��� 13.5'N, 103��59'.5��� 104��01'E, altitude 1340���1870 m. Type specimens. H��l��type ♂ (EJCB): ���La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5 ��� 104��01'E, Ban Saluei ��� Ph��u Pane Mt., 1340���1870m, 15.iv.���15.v.2008, La�� c��llect��rs leg.���. Other paratypes. LAOS: Bolikhamxai: ���La��s centr., B��likhamsai pr��v., Ban Nape-Kaew Nua Pass, 18.4. ��� 1.5.1998, alt. 600�� 100 m, N 18��22.3, E 105��09.1 (GPS), E. Jendek & O. ��au��a leg.��� 2 paratypes (EJCB). Houaphan: ���NE La��s, Hua Phan pr��v., 2004, 20��13���N, 103��59���E, 6���18.iv., Ban Saluei, Phu Phan Mt. env., 2004, J. Bezděk leg., 1300���2000m ��� 1 paratype (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 7.iv���25.v.2010, 1300���1900m, leg. C. H��lzschuh��� 8 paratypes (EJCB)������La��s-NE, H��ua Phan pr��v., 20��12���13.5'N, 103��59'.5��� 104��01'E, Ban Saleuy ��� Ph��u Pane Mt., 1340���1870m, 2.���22.vi.2011, V��t Kub��ň & La�� c��ll. leg.��� 6 paratypes (NMPC), 5 paratypes (EJCB)������La��s-NE, Hua Phan Pr��vince, Ban Saleui, Ph��u Pan (Mt), 20��12'N, 104��01'E, 1���31.v.2011, 1300���1900m, leg. C. H��lzschuh��� 2 paratypes (MCCI), 25 paratypes (EJCB), 22 paratypes (USNM), 2 paratypes (CNC). Distribution. Laos: B��likhamxai, H��uaphan. Etymology. The specific name is the Latin n��un frontis (face, f��re part ��f anything, l����k) in app��siti��n. It refers t�� the striking g��lden-green fr��ns ��f male (see Sexual dim��rphism), which is in the sharp c��ntrast with the c��l��r ��f the d��rsal side., Published as part of Jendek, Eduard & Grebennikov, Vasily V., 2018, Twenty new species of Agrilus (Coleoptera: Buprestidae: Agrilinae) from the Oriental Region, pp. 107-131 in Zootaxa 4429 (1) on pages 117-118, DOI: 10.11646/zootaxa.4429.1.4, http://zenodo.org/record/1280541

Grebennikov, Vasily V. (2017): Cleonini (Coleoptera: Curculionidae: Lixinae) are monophyletic and flightless: tribe overview, rampant adult homoplasy and illustrated global diversity. Zootaxa 4329 (1): 1-63, DOI: https://doi.org/10.11646/zootaxa.4329.1.1

Cotasterosoma Konishi, 1962 Konishi 1962: 3 (species included: omogoensis). Type species: Cotasterosoma omogoensis Konishi, 1962, by original designation. Diagnosis. Species of the genus Cotasterosoma are recognizable among all weevils by the unique combination of the following nine non-unique adult characters: (1.) small-sized relatively glabrous weevils of characteristic ���edaphic Cossoninae ��� habitus (Figs 1 A���D, 2A���D); (2.) antennae consisting of scape, funicle with six (Fig. 2 F) or seven antennomeres (the type species, but see Discussion) and seemingly one-segmented club (Fig. 2 F); (3.) all tarsi fivesegmented (Figs 2 G���I); (4.) all tarsomeres subcylindrical in shape (Figs 2 G���I); (5.) fore-legs with unci arising from the middle of apical margin of tibia (Fig. 2 G), and not from the dorso-apical corner; (6.) tarsomere 3 not enlarged (Figs 2 G��� I); (7.) tarsomere 4 short and inserting at apical surface of tarsomere 3, with a few setae on ventral side (Figs 2 G���I); (8.) male genitalia with U- or Y-shaped tegmen broadly abbreviated dorsally (Figs 1 E���H); (9.) collum of spermatheca bent at apex and almost parallel to ramus (Fig. 2 K)., Published as part of Grebennikov, Vasily V. & Morimoto, Katsura, 2016, Flightless litter-dwelling Cotasterosoma (Coleoptera: Curculionidae: Cossoninae) found outside of Japan, with mtDNA phylogeography of a new species from Southwest China, pp. 133-138 in Zootaxa 4179 (1) on pages 134-135, DOI: 10.11646/zootaxa.4179.1.11, http://zenodo.org/record/267439, {"references":["Konishi, M. (1962) Taxonomic studies on the Cossoninae of Japan (Coleoptera: Curculionidae), Part 1. Insecta Matsumurana, 25, 1 - 17."]}

Catapionus mopsus sp.n. Figs 1���7. Diagnosis. Adult specimens of this nominal species can be likely distinguished from all other Catapionus (and, in fact, from all organisms), by having unique combination of external and genital morphological characters as illustrated in Figs 3���6, and unique DNA barcodes. Description. Adults with dense scales on body surface giving beetles bluish or greenish luster (unless scales are partly abrased and body appears more blackish, Fig. 4); antennomere 3 slightly longer than antennomere 2; temples behind eyes with longitudinal black strip bearing no scales (Figs 7A���D); pronotum along midline about 1.1���1.3x as long as its maximal width; elytral lateral contour more rounded in females (Figs 4 A, 6A) than in males (Figs 3 A, 5A). Holotype, male (Figs 3 A���H). GenBank accession: KU748534. Intraspecific variation. Male (Figs 5 A���H) and females (Figs 4 A���G, 6A���H); GenBank accessions as in Fig.7 E. Material examined. Holotype (ZCAS): ��� CHINA, Yunnan, Haba Shan, N27��20'51" E100��05'33", 27.vi. 2012, 4158m, under rock, V. Grebennikov ���, ���CNCCOLVG00005402���. Paratypes (CNC, ZCAS; specimen numbers as in Fig. 7 E): 5 exx in total: same locality data as holotype. Additional specimens (CNC): six specimens from Mt. Haba (collected either together with the holotype, or on June 28, 2012 at 4195 m) and seven specimens from Mt. Gongga with labels ���P.R. CHINA, Sichuan, NE slope Gongga Shan, N29��52'04" E102��01'53", 12.vi. 2011, 3533m, under stone, V.Grebennikov ��� or ���P.R. CHINA, Sichuan, NE slope Gongga Shan, N29��54'40" E102��00'37", 13.vi. 2011, 4143m, under stone, V.Grebennikov���, specimen numbers of six of them used for DNA barcodes are in Fig. 7 E. Distribution. This species consists of two populations (Fig. 1): one on Mt. Haba in Yunnan (the type locality) at altitude of 4,158���4,195 m and another on Mt. Gongga in Sichuan at altitude of 3,533���4,143 m. Etymology. The species epithet is a Latinized Greek mythical name of Mopsus, an Argonaut, who understood the language of birds and died from a snakebite in Libya; noun in apposition. Results of mtDNA analysis. Phylogeographic analysis grouped 12 sequences of the new Catapionus species in two clusters corresponding to both localities: Mt. Haba and Mt. Gongga (Fig. 7 E). The time of mtDNA divergence between both populations was estimated at about 3.65 Mya (Fig. 7 E)., Published as part of Grebennikov, Vasily V., 2016, Flightless Catapionus (Coleoptera: Curculionidae: Entiminae) in Southwest China survive the Holocene trapped on mountaintops: new species, unknown phylogeny and clogging taxonomy, pp. 243-254 in Zootaxa 4205 (3) on pages 247-248, DOI: 10.11646/zootaxa.4205.3.4, http://zenodo.org/record/194300