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Anomala hualienensis Zhao, new species [ẄÄ异fflŵoi/ḢM條ŵff] (Figs. 1A–C, 2A–C, 3A–C) Type material. Holotype: ♂ (NMNS), Taiwan Island, Hualien, Kuangfu logging road [= 光復IJdz], 820~ 890m, 2020-V-30, Y.-T. Chien leg., by mercury lamp trap. Paratypes: 17♂♂, 25♀♀ (1♂, 2♀♀ in NMNS, 10♂♂, 17♀♀ in CCPC, 2♂♂, 2♀♀ in CZPC, 2♂♂, 2♀♀ in IZAS, 2♂♂, 2♀♀ in TARI), same data as holotype; 116♂♂, 125♀♀ (66♂♂, 75♀♀ in CCPC, 50♂♂, 50♀♀ in ZMPC), ditto, but “ 2021-V-13 ”; 47♂♂, 103♀♀ (CCPC), ditto, but “ 2021- V-21 ”; 4♂♂, 12♀♀ (CCPC), Kuangfu logging road, 1000m, 24-V-2009, Wen-I Chou leg. Description. Holotype male (Fig. 1A–C). Body length: 15.8 mm, greatest width: 8.1 mm. General appearance. Body shape elongated ovoid, convex. Entirely metallic green, elytra and abdomen predominantly red, antenna dark brown. Head. Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin nearly straight and weakly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons rugopunctate, gradually changing into small punctures, other portions of frons and vertex with sparse minute punctures. Antennal club slightly longer than antennomeres 1–6 combined. Inner margin of eye with several moderately long setae. Pronotum. Sides nearly subparallel in posterior two fifth, then gently curved, strongly convergent anteriad in anterior three fifth. Anterior angle acute and weakly protruding, posterior angle blunt. All marginal lines complete. With sparse small punctures. Lateral margin with several long setae. Scutellum. Subtriangular, lateral margin arched. With moderately dense minute punctures somewhat irregularly distributed, impunctate at central area and the curved margins. Elytra. Interstices flat, primary costae slightly convex; strial punctures moderately dense, annulated and large. Interstices I to IV each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria irregularly doubled; all these secondary striae obsolete before the level of apical umbone, except for that on interstice III, which has several scattered punctures before humeral umbone. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina distinct in basal half. Epipleura with a row of sparse short setae. Marginal membrane complete. Propygidium. With dense small and transverse punctures sometimes coalescent, punctures coarse near sides. Glabrous. Pygidium. Weakly bulging, posterior margin broadly arched apically. With dense transverse small punctures, punctures being denser at marginal portions. With several short setae distributed near posterior margin, posterior margin with a row of moderately long setae near apex. Ventral thoracic surface. Hypomeron irregularly striolate and sparse short setae. Collar of mesosternite with irregularly distributed small punctures and dense short setae at each side. Other portions of mesosternite with very dense coarse punctures and dense long setae. Ventral metathoracic surface with dense to very dense, semi-annulated large punctures which usually coalescent, and dense long setae at each side; with small dense punctures and sparse short setae medially. Abdominal ventrites. With moderately dense, transverse large punctures. Ventrites 2–5 each with a transverse row of sparse moderately long setae, the rows interrupted medially, setae sometimes becoming spiniform in ventrites 4–5; ventrite 6 with a complete row of setae along posterior margin. Legs. Protibia bidentate, both teeth distinct and sharp, apical tooth longer, extending to level of halfway of protarsomere 3. Inner spur situated at the level of proximal tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches distinctly longer and thicker, the lower branch of inner protarsal claw strongly concave internobasally. Protarsus slightly thickened. Each tarsomere 5 with a strong internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with three transverse rows of setae, first row at anterior margin with long setae, a second row at middle and a third row at posterior margin with short to moderately long, robuster setae; metafemoral surface with three transverse rows of moderately long setae, anterior row at anterior margin, a second row slightly behind middle with long setae, and a third row at posterior margin with short setae. Male genitalia. See Fig. 3A–C. Paratypes. Males. Body length: 14.6–16.4 mm, greatest width: 7.6–8.3 mm. External features and shape of parameres consistent. Sometimes the metallic green part with distinct blue sheen. Females (Fig. 2A–C). Body length: 16.2–17.8 mm, greatest width: 8.8–9.3 mm. Generally similar to male, but stouter and more convex. Antennal club almost equally as long as antennomeres 1–6 combined. Lateral carina more distinct in basal third of elytra. Pygidium more triangular in shape. Protibia and protarsus distinctly thinner than in male; both protibial teeth blunt at apex, apical tooth extending to level of base of protarsomere 3; inner spur situated at halfway of protibia; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length, but the lower branches are still thicker; metatibia more fusiform. Differential diagnosis. Anomala hualienensis Zhao, new species can be assigned to the Anomala semicastanea species-group (sensu Lin 2000) according to the morphological similarities. The shape of aedeagus suggests that its closest relatives are A. semicastanea Fairmaire, 1888 (Figs. 1D–F, 2D–F, 3D–F) from southeastern part of mainland China and A. quelparta Okamoto, 1924, endemic to Jeju Island off the coast of Korean peninsula. The new species can be distinguished by the wider protibia and protarsus, as well as the more divided parameres with distinctly sinuate inner margins and shorter apical denticles. The new species is externally similar to A. insulicola Lin, 2000 from Hainan and A. shimenensis Lin, 2000 from Hunan, but both species have bidentate and adjacent parameres (vs. unidentate and divided parameres in A. hualienensis). Remarks. The new species was misidentified as Anomala semicastanea by Kobayashi & Chou (2008). Etymology. This species is named after Hualien County, where the type material was collected. Distribution. Taiwan Island (Hualien)., Published as part of Zhao, Ming-Zhi & Zorn, Carsten, 2022, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan, pp. 175-195 in Zootaxa 5168 (2) on pages 176-179, DOI: 10.11646/zootaxa.5168.2.5, http://zenodo.org/record/6877267, {"references":["Lin, P. (2000) Anomala semicastanea species group (Coleoptera: Rutelidae) of China. Entomotaxonomia, 22 (1), 37 - 41.","Kobayashi, H. & Chou, W. I. (2008) Description of a new genus of Anomalini and two new species of Hopliini and Melolonthini from Taiwan, with two new records of scarabaeid beetles. Kogane, 9, 69 - 76."]}

Anomala kanshireiensis Zorn & Zhao, new species [关T岭异fflŵoi/ṞTĂ條ŵff] (Figs. 6, 8E–F, 9H–J, 9N–O) Type material. Holotype: ♂ (MFNB), Formosa, Sauter, Kanshirei [= Guanziling in Tainan City, ṞTĂ], 908. V. 19-27. Paratypes: 1♂, 3♀♀ (MFNB), same data as holotype; 3♂♂ (MFNB), Formosa, Sauter, Kanshirei, 908. V. 17-24.; 1♂ (MFNB), Formosa, Sauter, Kanshirei, 908. V. 1-9; 1♂ (MFNB), Formosa, Sauter, Kanshirei, 908. V. 19-26.; 3♂♂, 1♀ (SMNS), TAIWAN, Kanshirei, 2. - 14. 6. 1908, H. Sauter leg.; 1♂ (HNHM), Formosa, Sauter, Kanshirei, 908. V-1-2-14. Description. Holotype male (Figs. 6A–C, 8E, 9K–M). Body length: 11.8 mm, greatest width: 6.7 mm. General appearance. Body shape elongated ovoid, weakly convex. Entirely dark reddish brown, with weak green metallic sheen. Head (Fig. 8E). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons densely rugopunctate, other portions of frons and vertex with irregularly distributed dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several moderately long setae (mostly worn in the holotype). Pronotum. Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle obtuse, rounded off. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense small punctures, punctures being partly coalescent at marginal portions; with a very weakly indicated longitudinal medial furrow not reaching the base. Lateral margin with several long setae, two moderately long to long semierect setae on disc of pronotum. Scutellum. Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed moderately large punctures. Disc with two semierect long setae. Elytra. Intervals strongly convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae unequally spaced; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria shortly doubled in basal fourth; the other secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete between level of scutellar apex to midway of elytron. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete. Propygidium. With dense transverse punctures, usually coalescent into transverse striolation; glabrous. Pygidium. Distinctly bulging, posterior margin protruding. Disc with dense ovoid large punctures, becoming striate near all margins; punctures encircling greatest tumidity. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex. Ventral thoracic surface. Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with small punctures and somewhat rugopunctate, with dense short setae. Ventral metathoracic surface with dense annulated moderately large punctures and dense, rather long setae; with minute punctures and almost glabrous medially. Abdominal ventrites. Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae; ventrite 6 with a complete row of long setae along posterior margin. Legs. Protibia tridentate, apical and second tooth acute at apex, proximal tooth small; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted at the level of the indention between proximal and second tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and wider, the lower branch of inner protarsal claw concave internobasally. Each tarsomere 5 with an internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin usually with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae. Male genitalia. See Fig. 9H–J. Parameres asymmetric, right paramere longer. Ventral plate flat, apex acute. Paratypes. Males. Body length: 11.5–13.0 mm, greatest width: 6.5–7.5 mm. External features and shape of parameres consistent. Semierect longer setae on scutellum and disc of pronotum are only present in few males but might be worn off in other specimens. Females (Figs. 6D–F, 8F). Body length: 12.3–13.0 mm, greatest width: 7.0– 7.9 mm. Generally similar to male. Antennal club almost equally as long as antennomeres 2–6 combined. Sides of pronotum more strongly curved. Pygidium more bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, almost extends to level of apex of protarsomere 3; the other two teeth blunt at apex, proximal tooth indistinct; inner spur situated at the level of the proximal tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length. Ventral metathoracic surface less setose than in male. Semierect longer setae on scutellum and disc of pronotum are only present in the minority of females but might be worn off in some specimens. Differential diagnosis. Anomala kanshireiensis Zorn & Zhao, new species is most similar to A. inclinata Zhao & Zorn, new species. Above all, the general body color is reddish brown in A. kanshireiensis but blackish brown in A. inclinata. The interrupted secondary stria of interstice II is more complete than in A. inclinata. The aedeagi of the two species are similar in shape. The left paramere is shorter and has a less curved apex compared to A. inclinata. The left paramere is placed slightly above the right paramere in lateral view (parameres are overlapping in A. inclinata). The apex of the ventral plate is acute in A. kanshireiensis but arched in A. inclinata. The endophallites of the two species show considerable differences: the two dorsal endophallites are spaced with one being relatively long in A. kanshireiensis (Fig. 9N–O), they are adjacent and both short in A. inclinata (Fig. 9P–Q); the ventral endophallite is a curved spine in A. kanshireiensis, while it is a serrated sclerite in A. inclinata. Etymology. The specific epithet is alluding to the historical name of the type locality, Kanshirei. Distribution. Taiwan Island (Tainan)., Published as part of Zhao, Ming-Zhi & Zorn, Carsten, 2022, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan, pp. 175-195 in Zootaxa 5168 (2) on pages 184-187, DOI: 10.11646/zootaxa.5168.2.5, http://zenodo.org/record/6877267

Anomala linwenhsini Zhao & Zorn, new species [Ž信异fflŵoi/Ž信條ŵff] (Figs. 4, 8A–B, 9A–C) Type material. Holotype: ♂ (NMNS), Taiwan Island, Pingtung, Shuangliu [= Ḇ流], Yintun [= 尹屯], 2008-VI-29, Wen-Hsin Lin leg. Paratypes: 1♂ (HKPC), Nantou, Sun Moon Lake, 4-VI-1981, Chin-Kin Yu leg.; 2♂♂ (CCPC, ZMPC), Pingtung, Fengkang [= RAE], 2008-VI-11, Wen-Hsin Lin leg.; 4♂♂, 1♀ (CZPC), Taitung [actually Pingtung], Schouchia env. [= Ŕ+], 315m, 18.5.2012, N 22.20670°, E 120.86034°, Walter Grosser lgt. Description. Holotype male (Figs. 4A–C, 8A, 9A–C). Body length: 15.3 mm, greatest width: 7.9 mm. General appearance. Body shape elongated ovoid, weakly convex. Entirely reddish brown; sutural line of elytra black; joints of tibiae and femora, apices of tarsomeres dark brown. Head (Fig. 8A). Clypeus subrectangular, anterior corner broadly rounded; anterior margin straight and strongly reflexed; surface with dense coarse punctures, somewhat coalescent. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons rugopunctate, other portions of frons and vertex with dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several short setae. Pronotum. Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle round. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense annulated small punctures, punctures being coalescent near lateral margins, smooth before scutellum; with a longitudinal medial furrow not reaching the base. Lateral margin with several long setae. Scutellum. Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed small punctures. Disc with one semierect setae (others probably worn out). Elytra. Intervals convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae irregular in spaces; the whole surface with scattered minute punctures. Interstices I to IV each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria irregularly doubled in basal two third; all secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete from basal fourth to halfway of elytron. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete. Propygidium. With dense transverse large punctures, usually coalescent. Glabrous. Pygidium. Distinctly bulging, posterior margin protruding. With very dense transverse large punctures, punctures more ovoid near each side; punctures encircling greatest tumidity. Posterior margin with a row of long setae near apex. Ventral thoracic surface. Hypomeron with irregular striolation and sparse long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with shallow small punctures which usually coalescent into irregular striolation, with dense short setae. Ventral metathoracic surface with very dense, shallow and annulated large punctures and dense, rather long setae at each side; with minute punctures and glabrous medially. Abdominal ventrites. Ventrites 1–2, and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with transverse, small or large punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse, recumbent and moderately long setae, the rows broadly interrupted medially; ventrite 6 with a complete row of recumbent long setae along posterior margin. Legs. Protibia tridentate, apical and second teeth blunt at apex, proximal tooth indistinct; apical tooth almost extending to level of apex of protarsomere 2. Inner spur situated at the level of the second tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and distinctly wider, the lower branch of inner protarsal claw strongly concave internobasally. Protarsus slightly thickened. Each tarsomere 5 with an indistinct internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin regular with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin and somewhat doubled, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae. Male genitalia. See Fig. 9A–C. Parameres weakly asymmetric: the left paramere acute at apex, the right paramere round at apex. Ventral plate elongated and curved downwards at apex. Paratypes. Males. Body length: 14.4–15.9 mm, greatest width: 7.7–8.5 mm. External features and shape of parameres consistent. One male with a long erect seta on disc of pronotum. Female (Figs. 4D–F, 8B). Body length: 14.7 mm, greatest width: 8.2 mm. Generally similar to male. Anterior margin of clypeus weakly reflexed (Fig. 8B); antennal club almost equally as long as antennomeres 2–6 combined. Pygidium less bulging. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, extending to level of base of protarsomere 3; inner spur situated slightly before the level of the proximal tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length, but the lower branches are still thicker. Differential diagnosis. In the Anomala linwenhsini -species group, A. linwenhsini Zhao & Zorn, new species is easily characterized by the subrectangular clypeus in both sexes (subtrapezoidal in other three species). The parameres are almost symmetric with the apices slightly unequal. The ventral plate of aedeagus is elongated and curved downwards at apex (apex not elongated in other three species). Moreover, this is the largest species of this group. Remarks. The specimen identified as Anomala inconcinna in Yu et al. (1998) was examined and proved to be A. linwenhsini Zhao & Zorn, new species. This new species is active in March to April according to Yu et al. (1998), but the type series suggests that it is also active in May and June. Etymology. This species is named after the late Mr. Wen-Hsin Lin, who collected some of the type specimens. Distribution. Taiwan Island (Pingtung, Nantou)., Published as part of Zhao, Ming-Zhi & Zorn, Carsten, 2022, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan, pp. 175-195 in Zootaxa 5168 (2) on pages 180-182, DOI: 10.11646/zootaxa.5168.2.5, http://zenodo.org/record/6877267, {"references":["Yu, C. K., Kobayashi, H. & Chu, Y. I. (1998) The Scarabaeidae of Taiwan. Mu Sheng Company, Taipei, 263 pp."]}

Anomala inclinata Zhao & Zorn, new species [DZṜ异fflŵoi/Øae條ŵff] (Figs. 7, 8G–H, 9K–M, 9P–Q) Type material. Holotype: ♂ (NMNS), Taiwan Island, Taitung, Taimali [= kDzae], 2008-IV-12. Paratype: 1♀ (CCPC), same data as holotype. Description. Holotype male (Figs. 7A–C, 8G, 9H–J, 9P–Q). Body length: 13.0 mm, greatest width: 7.3 mm. General appearance. Body shape elongated ovoid, weakly convex. Entirely blackish brown, with weak green metallic sheen. Head (Fig. 8G). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons densely rugopunctate, other portions of frons and vertex with irregularly distributed, dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several moderately long setae. Pronotum. Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle round. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense small punctures, punctures being coalescent at marginal portions, impunctate before scutellum; with a longitudinal medial furrow not reaching the base. Lateral margin with several long setae. Scutellum. Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed coarse punctures. Disc with some semierect short setae. Elytra. Intervals strongly convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae unequal in spaces; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria shortly doubled in basal third; all secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete from the level of scutellar apex to apical two fifth. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete. Propygidium. With dense transverse punctures, usually coalescent into transverse striolation; glabrous. Pygidium. Distinctly bulging, posterior margin protruding. Disc with dense ovoid large punctures, punctures more transverse near all margins. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex. Ventral thoracic surface. Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with small punctures and somewhat rugopunctate, with dense short setae. Ventral metathoracic surface with dense annulated small punctures and dense, rather long setae at each side; with minute punctures and glabrous medially. Abdominal ventrites. Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae, interrupted medially in ventrites 3–4; ventrite 6 with a complete row of long setae along posterior margin. Legs. Protibia tridentate, apical and second teeth acute at apices, proximal tooth small; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted at the level indention between second and proximal tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and wider, the lower branch of inner protarsal claw concave internobasally. Each tarsomere 5 with an internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin usually with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae. Male genitalia. See Fig. 9K–M. Parameres asymmetric, right paramere longer. Ventral plate flat, apex arched. Paratype. Female (Figs. 7D–F, 8H). Body length: 13.5 mm, greatest width: 6.9 mm. Generally similar to male. Antennal club almost equally as long as antennomeres 2–6 combined. Pygidium more bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, almost extends to level of apex of protarsomere 2; other two teeth blunt at apex, the third one indistinct; inner spur situated at the level of the third tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length. Ventral metathoracic surface less setose than in male. Differential diagnosis. See differential diagnosis of Anomala kanshireiensis Zorn & Zhao, new species. Etymology. The specific epithet derives from the Latin adjective “ inclinatus, -a, -um ”, alluding to the inclined parameres of this species. Distribution. Taiwan Island (Taitung).

Anomala wutaiensis Zhao & Zorn, new species [Ẩ台异fflŵoi / ÃAE條ŵff] (Figs. 5, 8C–D, 9D–G) Type material. Holotype: ♂ (NMNS), Taiwan Island, Pingtung, Wutai Township [= ÃAEď], Ali [= Kae], 1300 m, 2.VI.2014, Yu-Feng Hsu leg. Paratypes: 1♂, 1♀ (ZMPC), Taiwan Island, [Pingtung], Wutai, 120.74E 22.74N 1040 m, light trap, 2018-V-5, Hai-Tian Song leg.; 4♂♂, 2♀♀ (ZMPC), Wutai, light trap, 2018-V-12, Hai-Tian Song leg; 1♀ (CCPC), Wutai, Shenshan [= Ñ山], 1998-IV-30, Chang-Chin Chen leg. Description. Holotype male (Figs. 5A–C, 8C, 9D–G). Body length: 14.0 mm, greatest width: 7.3 mm. General appearance. Body shape elongated ovoid, weakly convex. Entirely reddish brown, color darkened at head and pronotum, as well as sides of scutellum; sutural line of elytra black; joints of tibiae and femora, apices of tarsomeres dark brown. Head (Fig. 8C). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface with dense small punctures. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons rugopunctate, other portions of frons and vertex with dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with two moderately long setae. Pronotum. Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle round. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense annulated small punctures, punctures somewhat coalescent near lateral margins, a narrow longitudinal line at posterior third impunctate. Lateral margin with several long setae. Scutellum. Nearly triangular. Margins with polished appearance, disc with moderately dense large punctures. Disc with a few semierect moderately long setae. Elytra. Intervals convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae irregular in spaces; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria doubled in basal third; all secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete from basal fourth to apical third. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete. Propygidium. With dense transverse punctures coalescent into transverse striolation. Glabrous Pygidium. Distinctly bulging, posterior margin protruding. With very dense transverse large punctures, punctures more ovoid near each side; punctures encircling greatest tumidity. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex. Ventral thoracic surface. Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with shallow large punctures which usually coalescent into irregular striolation, with dense short setae. Ventral metathoracic surface with dense to very dense, shallow and annulated large punctures and dense, rather long setae at each side; with minute punctures and glabrous medially. Abdominal ventrites. Ventrites 1–2, and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small or large punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae, the setae sometimes recumbent, the rows broadly interrupted medially; ventrite 6 with a complete row of recumbent long setae along posterior margin. Legs. Protibia tridentate, the apical and the second teeth blunt at apex, proximal tooth small; apical tooth almost extending to level of apex of protarsomere 2. Inner spur situated at the level of the second tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and distinctly wider, the lower branch of inner protarsal claw strongly concave internobasally. Protarsus slightly thickened. Each tarsomere 5 with an indistinct internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin regular with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae. Male genitalia. See Fig. 9D–G. Parameres strongly asymmetric. The right paramere distinctly shorter than the left one. Ventral plate deflected at apex. Paratypes. Males. Body length: 12.0– 13.7 mm, greatest width: 6.7–7.6 mm. External features and shape of parameres consistent. A male has lighter body color and two short semierect setae on disc of pronotum. Another male with a long erect seta near posterior margin of pronotum. Females (Figs. 5D–F, 8D). Body length: 12.3–13.9 mm, greatest width: 6.8–8.1 mm. Generally similar to male, but less convex. Body color dark brown to blackish brown. Anterior margin of clypeus weakly reflexed (Fig. 8D); antennal club almost equal to antennomeres 2–6 combined. Scutellum has straighter lateral margin. Pygidium less bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial teeth tongue-shaped, extends to level of base of protarsomere 3; inner spur situated at the level of the third tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length, but the lower branches are still thicker. The setae of the setose rows situated behind the middle of meso- and metafemoral surfaces are longer than in male. Differential diagnosis. Anomala wutaiensis Zhao & Zorn, new species is similar to A. kanshireiensis Zorn & Zhao, new species and A. inclinata Zhao & Zorn, new species. However, interstice II of A. wutaiensis is slightly wider than in the other two species. The secondary stria of interstice II is obsolete approximately between basal fourth and apical third of elytron, while it is obsolete from level of scutellar apex to midway of elytron in A. kanshireiensis and to apical two fifth of elytron in A. inclinata. Moreover, A. wutaiensis has the thickest aedeagus among the A. linwenhsini -species group. The parameres are strongly asymmetric. The ventral plate of aedeagus is deflected at apex but flat in A. kanshireiensis and A. inclinata. Etymology. The specific epithet is derived from Wutai, where the type series was collected. Distribution. Taiwan Island (Pingtung)., Published as part of Zhao, Ming-Zhi & Zorn, Carsten, 2022, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan, pp. 175-195 in Zootaxa 5168 (2) on pages 182-184, DOI: 10.11646/zootaxa.5168.2.5, http://zenodo.org/record/6877267

The Anomala linwenhsini -species group Four of the new species from Taiwan described in the present paper, i.e. A. linwenhsini Zhao & Zorn, new species, A. wutaiensis Zhao & Zorn, new species, A. kanshireiensis Zorn & Zhao, new species and A. inclinata Zhao & Zorn, new species comprise a homologous group. They are characterized by the combination of the following characters: body moderately convex, anterior margin of clypeus distinctly reflexed, elytral intervals strongly convex, strial punctures dense and large, lateral carina indistinct, epipleura with a row of long setae, protibia distinctly tridentate in both sexes, metatibia rather slender and not fusiform, parameres fully separated and more or less asymmetric, scutellar surface and disc of pronotum before scutellum with few short to long erect setae. The latter setae have so far not be found in all examined specimens, and the authors are uncertain at this point whether they are worn off in these cases or not present in all specimens. Anomala inconcinna Bates, 1888, also only known from Taiwan, is superficially very similar to the species of the linwenhsini -group. However, in A. inconcinna the protibia is clearly bidentate, it does not have singular long setae on pronotum or scutellum, the epipleural setae are slightly shorter, and the aedeagus is more or less symmetric. Therefore, the authors currently do not include A. inconcinna in the linwenhsini -group.

Integrative taxonomy was employed to exploit the differences between the known Metaphire anomala (Michaelsen, 1907) and other specimens collected in Vietnam. The results brought to light two new species, namely Metaphire iranomala sp. nov. and Metaphire decemtheca sp. nov. The former is easily recognised by having male pores on xix and four pairs of spermathecal pores on 5/6/7/8/9 while the latter is distinguished by having five pairs of spermathecal pores on 4/5/6/7/8/9. The K2P distances of the fragment of the cytochrome c oxidase subunit I (COI) gene are 13.1% between M. iranomala sp. nov. and M. anomala (Michaelsen, 1907) and 18% between M. decemtheca sp. nov. and Metaphire grandiverticulata Nguyen & Lam, 2017. The intraspecific divergences are 1.5%–10.6% for M. iranomala sp. nov. and 2.1%–11.4% for M. decemtheca sp. nov.

The classification of Microhoriini Bonadona, 1974 is revised. Five genera are recognized: Aulacoderus LaFerté-Sénectère, 1849, Falsophilus Kejval, 2015, Liparoderus LaFerté-Sénectère, 1849, Microhoria Chevrolat, 1877, and Neocrohoria Telnov, 2019. (i) New species: Microhoria almukalla Kejval, sp. nov. (Yemen), M. anahita Kejval, sp. nov. (Iran), M. antalya Kejval, sp. nov. (Turkey), M. bacillisternum Kejval, sp. nov. (Iran), M. cervi Kejval, sp. nov. (Oman), M. fergana Kejval, sp. nov. (Kyrgyzstan), M. garavuti Kejval, sp. nov. (Tajikistan), M. gibbipennis Kejval, sp. nov. (Turkey), M. halophila Kejval, sp. nov. (Turkey), M. hazara Kejval, sp. nov. (Afghanistan), M. heracleana Kejval, sp. nov. (Greece), M. impavida Kejval, sp. nov. (Turkey), M. kabulensis Kejval, sp. nov. (Afghanistan), M. kermanica Kejval, sp. nov. (Iran), M. pahlavi Kejval, sp. nov. (Iran), M. persica Kejval, sp. nov. (Iran), M. strejceki Kejval, sp. nov. (Tajikistan), M. sawda Kejval, sp. nov. (Saudi Arabia), and M. sulaimanica Kejval, sp. nov. (Pakistan, Uzbekistan). (ii) New synonymies: Microhoria Chevrolat, 1877 = Clavicomus Pic, 1894 syn. nov. = Tenuicomus Pic, 1894 syn. nov.; Microhoria depressa (LaFerté-Sénectère, 1849) = Anthicus mollis Desbrochers des Loges, 1875 syn. nov.; Microhoria edmondi (Pic, 1893) = Anthicus spinosus Pic, 1912 syn. nov.; Microhoria globipennis (Pic, 1897) = Anthicus globipennis quercicola Sahlberg, 1913 syn. nov.; Microhoria luristanica (Pic, 1911) = Anthicus pietschmi Pic, 1938 syn. nov.; Microhoria ottomana (LaFerté-Sénectère, 1849) = Anthicus merkli Pic, 1897 syn. nov.; Microhoria pinicola (Reitter, 1889) = Microhoria feroni Bonadona, 1960 syn. nov.; Microhoria posthuma (Krekich-Strassoldo, 1931) = Anthicus fumeoalatus Krekich-Strassoldo, 1931 syn. nov.; Microhoria truncatipennis (Pic, 1897) = Anthicus mouzafferi Pic, 1910 syn. nov. (iii) Status changes. Anthicus tauricus var. inobscura Pic, 1908 is raised to species level as Microhoria inobscura (Pic, 1908) stat. nov.; Anthicus truncatus var. decoloratus Pic, 1897 is removed from synonymy with Anthicus truncatus Pic, 1895 and raised to species level as Microhoria decolorata (Pic, 1897) stat. restit. (iv) New combinations: Microhoria disconotata (Pic, 1907) comb. nov., M. fossicollis (LaFerté-Sénectère, 1849) comb. nov., M. gestroi (Pic, 1895) comb. nov., M. irregularis (Pic, 1932) comb. nov., M. lividipes (Desbrochers des Loges, 1875) comb. nov., M. marginicollis (Pic, 1951) comb. nov., M. nystii (LaFerté-Sénectère, 1849) comb. nov., M. schimperi (Pic, 1898) comb. nov., M. semiviridis (Pic, 1951) comb. nov., M. strandi (Krekich-Strassoldo, 1931) comb. nov., and M. yemenita (Nardi, 2004) comb. nov., all from Anthicus Paykull, 1798. Microhoria abscondita (Telnov, 2000) comb. nov., M. adusta (Krekich-Strassoldo, 1931) comb. nov., M. afghana (Telnov, 2010) comb. nov., M. almorae (Krekich-Strassoldo, 1931) comb. nov., M. ambusta (Krekich-Strassoldo, 1931) comb. nov., M. angulifer (Pic, 1893) comb. nov., M. anomala (Telnov, 1998) comb. nov., M. antinorii (Pic, 1894) comb. nov., M. apicordiger (Bonadona, 1954) comb. nov., M. aquatilis (Krekich-Strassoldo, 1931) comb. nov., M. assamensis (Pic, 1907) comb. nov., M. assequens (Krekich-Strassoldo, 1931) comb. nov., M. atrata (Krekich-Strassoldo, 1931) comb. nov., M. austriaca (Pic, 1901) comb. nov., M. bicarinifrons (Pic, 1892) comb. nov., M. biguttata (Bonadona, 1964) comb. nov., M. brevipilis (Pic, 1893) comb. nov., M. bruckii (Kiesenwetter, 1870) comb. nov., M. brunneipes (Krekich-Strassoldo, 1931) comb. nov., M. caeruleicolor (Pic, 1906) comb. nov., M. callima (Baudi di Selve, 1877) comb. nov., M. comes (Krekich-Strassoldo, 1931) comb. nov., M. cordata (Krekich-Strassoldo, 1931) comb. nov., M. curticeps (Pic, 1923) comb. nov., M. dichrous (LaFerté-Sénectère, 1849) comb. nov., M. doderoi (Pic, 1902) comb. nov., M. erythraea (Pic, 1899) comb. nov., M. erythrodera (Marseul, 1878) comb. nov., M. feai (Pic, 1907) comb. nov., M. fugax (LaFerté-Sénectère, 1849) comb. nov., M. fugiens (Marseul, 1876) comb. nov., M. garze (Telnov, 2018) comb. nov., M. gigas (Pic, 1899) comb. nov., M. gravida (Krekich-Strassoldo, 1931) comb. nov., M. harmandi (Pic, 1899) comb. nov., M. hauseri (Pic, 1906) comb. nov., M. henoni (Pic, 1892) comb. nov., M. heydeni (Marseul, 1879) comb. nov., M. himalayana (Pic, 1909) comb. nov., M. hummeli (Pic, 1933) comb. nov., M. immaculipennis (Krekich-Strassoldo, 1931) comb. nov., M. inabsoluta (Telnov, 2003) comb. nov., M. indeprensa (Telnov, 2000) comb. nov., M. kabyliana (Pic, 1896) comb. nov., M. kejvali (Telnov, 1999) comb. nov., M. kham (Telnov, 2018) comb. nov., M. kocheri (Pic, 1951) comb. nov., M. kuluensis (Pic, 1914) comb. nov., M. lepidula (Marseul, 1876) comb. nov., M. longiceps (LaFerté-Sénectère, 1849) comb. nov., M. longicornis (Uhmann, 1983) comb. nov., M. manifesta (Pic, 1907) comb. nov., M. martinezi (Pic, 1932) comb. nov., M. muguensis (Telnov, 2000) comb. nov., M. nigrocyanella (Marseul, 1877) comb. nov., M. nigrofusca (Telnov, 2000) comb. nov., M. nigroterminata (Pic, 1909) comb. nov., M. notatipennis (Pic, 1909) comb. nov., M. olivierii (Desbrochers des Loges, 1868) comb. nov., M. optabilis LaFerté-Sénectère, 1849) comb. nov., M. paganettii (Pic, 1909) comb. nov., M. phungi (Pic, 1926) comb. nov., M. picea (LaFerté-Sénectère, 1849) comb. nov., M. plagiostola (Bonadona, 1958) comb. nov., M. plicatipennis (Pic, 1936) comb. nov., M. posthuma (Krekich-Strassoldo, 1931) comb. nov., M. postimpressa (Pic, 1938) comb. nov., M. postluteofasciata (Pic, 1938) comb. nov., M. prolatithorax (Pic, 1899) comb. nov., M. proterva (Krekich-Strassoldo, 1931) comb. nov., M. ragusae (Pic, 1898) comb. nov., M. semidepressa (Pic, 1893) comb. nov., M. separatithorax (Pic, 1914) comb. nov., M. shibatai (Nomura, 1962) comb. nov., M. schrammi Pic, 1913) comb. nov., M. sikkimensis (Pic, 1907) comb. nov., M. sinensis (Pic, 1907) comb. nov., M. spinipennis (Pic, 1898) comb. nov., M. sporadica (Krekich-Strassoldo, 1931) comb. nov., M. striaticollis (Krekich-Strassoldo, 1931) comb. nov., M. subpicea (Pic, 1914) comb. nov., M. tersa (Krekich-Strassoldo, 1931) comb. nov., M. tonkinensis (Krekich-Strassoldo, 1928) comb. nov., M. truncatella (LaFerté-Sénectère, 1849) comb. nov., M. turgida (Krekich-Strassoldo, 1928) comb. nov., M. uhagoni (Pic, 1904) comb. nov., M. uniformis (Krekich-Strassoldo, 1931) comb. nov., M. variabilis (Telnov, 2003) comb. nov., M. weigeli (Telnov, 2000) comb. nov., M. versicolor (Kiesenwetter, 1866) comb. nov., M. wuyishanensis (Nardi, 2004) comb. nov., and Nitorus niger (Uhmann, 1996) comb. nov., all from Clavicomus Pic, 1894. Microhoria agriliformis (Pic, 1893) comb. nov., M. alfierii (Pic, 1923) comb. nov., M. angelinii (Degiovanni, 2012) comb. nov., M. babaulti (Pic, 1921) comb. nov., M. barnevillei (Pic, 1892) comb. nov., M. armeniaca (Pic, 1899) comb. nov., M. bonnairii (Fairmaire, 1883) comb. nov., M. cyanipennis (Grilat, 1886) comb. nov., M. depressa (LaFerté-Sénectère, 1849) comb. nov., M. dolichocephala (Baudi di Selve, 1877) comb. nov., M. duplex (Nardi, 2004) comb. nov., M. edmondi (Pic, 1893) comb. nov., M. escalerai (Pic, 1904) comb. nov., M. finalis (Telnov, 2003) comb. nov., M. fuscomaculata (Pic, 1893) comb. nov., M. insignita (Pic, 1906) comb. nov., M. luristanica (Pic, 1911) comb. nov., M. meloiformis (Reitter, 1890) comb. nov., M. mesopotamica (Pic, 1912) comb. nov., M. ocreata (LaFerté-Sénectère, 1847) comb. nov., M. olivacea (LaFerté-Sénectère, 1849) comb. nov., M. ottomana (LaFerté-Sénectère, 1849) comb. nov., M. pallicra (Dufour, 1849) comb. nov., M. paralleliceps (Reitter, 1890) comb. nov., M. paupercula (LaFerté-Sénectère, 1847) comb. nov., M. platiai (Degiovanni, 2000) comb. nov., M. siccensis (Normand, 1950) comb. nov., M. subaerea (Reitter, 1890) comb. nov., M. subcaerulea (Pic, 1906) comb. nov., M. subsericea (Pic, 1898) comb. nov., M. tarifana (Pic, 1904) comb. nov., M. tibialis (Waltl, 1835) comb. nov., M. velox (LaFerté-Sénectère, 1849) comb. nov., M. viridipennis (Pic, 1899) comb. nov., and M. viturati (Pic, 1893) comb. nov., all from Tenuicomus Pic, 1894. Microhoria decolorata (Pic, 1897) comb. nov. and M. truncata (Pic, 1895) comb. nov. from Stricticomus Pic, 1894. Microhoria truncatipennis (Pic, 1897) comb. nov. from Anthelephila Hope, 1833. (v) Lectotype designations. Lectotypes are designated for the following species: Anthicus depressus LaFerté-Sénectère, 1849, A. edmondi Pic, 1893, A. luristanicus Pic, 1911, A. merkli Pic, 1897, A. mouzafferi Pic, 1910, A. pietschmi Pic, 1938, A. pinicola Reitter, 1889, A. posthumus Krekich-Strassoldo, 1931, and A. spinosus Pic, 1912.

As a result of a field study, carried out in August 2019, 25 species of myxomycetes from 17 genera, 7 families, 6 orders and 2 subclasses of the class Myxomycetes were collected in the area of the Seymskyi Regional Landscape Park (Sumy region, Ukraine). According to the bias-corrected Chao1 estimator, the collection represents 77.6% of the myxomycetes species composition of the park. The biodiversity indexes of the studied biota are as follows: Margalef’s diversity index – 13.7, Shannon’s index – 1.32, Simpson’s index – 0.06, Pielou’s index – 0.28. All identified species are new to the Ukrainian part of the Central Russian Upland forests. One of them, Comatricha anomala, found on the dead wood of Betula pendula, is new to Ukraine. Among the substrate groups of myxomycetes, xylophilic (18 species, 65.4%) and corticolous (8; 31%) species dominate in the park. Arcyria cinerea and Stemonitis pallida demonstrate a mixed substrate strategy. Among the substrate-forming plants, the greatest diversity of myxomycetes was observed on substrates, formed by relatively unabundant plant species: Populus tremula (13 myxomycete species, 50.0%), Quercus robur (11; 42.3%) and Bbetula pendula (8 species; 30.8%), while only 6 species of myxomycetes (23.1%) were found on the dominant plant species of P. sylvestris. Among the orders of myxomycetes, by number of species, Stemonitidales (11 species; 42.3%) and Trichiales (8 species; 30.8%) prevail in the park, among the families the most abundant were Amaurochaetaceae (6; 23.1%) and Trichiaceae (8; 30.8%), among the genera they are Arcyria (4; 15.4%), Lycogala, Stemonitis, Stemonitopsis and Comatricha (2 species each; 7.7%). The taxonomic structure of myxomycete biota differs significantly on the substrates, formed by different plant species. According to the ratio between species Trichiales and Stemonitidales, the substrate-forming plants form the following row: P. sylvestris, B.pendula, P.tremula, R. pseudoacacia and Q. robur (3.0 → 1.5 → 1.5 → 0.5 → 0.25 correspondingly)

Anomala xiongi Wang & Zorn spec. nov. is described from Yunnan, China. This new species can be distinguished from all other known Anomala species by conspicuous long erect setae rising from the primary costae of elytra. Additionally, ten new distributional records of Anomala species from China and neighboring countries are provided. Nomenclatural act Anomala xiongi Wang & Zorn spec. nov. – urn:lsid:zoobank.org:act:F10AD66D-BEF8-41E2-B22D-42D0492D9E95, Contributions to Entomology = Beiträge zur Entomologie, Bd. 71 Nr. 1 (2021)

Anomala xanthoptera BLANCHARD, 1851 (Figs 21, 50���52) Distribution: India; Nepal; Pakistan; China (new country record): Yunnan. Type material: Syntype of Anomala prasinicollis BATES, 1891 [junior subjective synonym of Anomala xanthoptera]. 1 ��� (MFNB), PUNJAB Kulu Distr. / Euchlora prasinicollis Cotype Bts. [Ohaus���s handwriting] / Anomala prasinicollis Bates [Bates���s handwriting]. Other specimens examined: CHINA: Yunnan: 1 ���, 2 ������ (FLWC), Nabang, Yingjiang County, Dehong, 17.II.2016, Hao Xu & Jian-Yue Qiu leg. INDIA: 34 ������, 8 ������ (CZGG, ZMH), Indien, Bhimtal, F. Smetacek leg., Museum Hamburg, Eing. Nr. 3/1977; 1 ��� (SMTD), Sikkim India. NEPAL: 1 ��� (CZGG), Nepal, Narayani Prov. Sauraha Ufer Rapti River 180 m, 27��34'80"N, 84��29'49"E, 18.IV.2000, leg. A. Skale, LF [collected at light]; 1 ��� (CZGG), Nepal-Himalaya, Myagdi distr., Babichor-Beni, 16.III.1994, 900- 1000 m, leg. D. Ahrens. PAKISTAN: 1 ��� (HNHM), Pakistan, Kashmir, Himalaya, Mts. 20 km S Muzaffarabad, Nara village, 750 m, 73��29'E, 34��01'N, 20.V.1998, leg. M. L��szl�� & G. Ronkay. Remarks: This species was previously known from India, Nepal and Pakistan (BATES 1891; ARROw 1917; ZORN & BEZDĚK 2016), and is here newly recorded in China based on three specimens collected from Yunnan., Published as part of Wang, Fa-Lei & Zorn, Carsten, 2021, Description of a new species and new faunistic records of the genus Anomala SAMOUELLE (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions, pp. 147-159 in Beitr��ge Zur Entomologie = Contributions to Entomology 71 (1) on page 153, DOI: 10.21248/contrib.entomol.71.1.147-159, http://zenodo.org/record/5743084, {"references":["BATES, H. W. 1891: Coleoptera from Kulu in N. W. India. - The Entomologist 24 (Supplement): 7 - 23.","ARROw, G. J. 1917: The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). - Taylor & Francis, London: 387 pp + v pl.","ZORN, C. & BEZDEK, A. 2016: Rutelinae.: 317 - 358. - In: LOBL, I. & LOBL, D. (eds.): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. - Brill, Leiden, Boston: 317 - 358."]}

Anomala ahrensi ZORN, 2011 Distribution: Vietnam, China (new country record): Yunnan. Specimens examined: CHINA: Yunnan: 1 ���, 1 ��� (FLWC), Jinping County, Honghe Pref., V-VI.2018, Yun-Chuan Xu leg.; 1 ���(CCPC), Biaoshuiyan, Ma���andi, Jinping County, Honghe Pref., 2010.V.13 X. D. Yang leg.; 1 ��� (CCPC), Luobodi Village, Ma���andi, Jinping County, Honghe Pref., 2010.V.17, X. D. Yang leg. Remarks: Anomala ahrensi was previously known only from northern Vietnam (ZORN 2011, ZORN et al. 2017). Herein, we newly report this species from Yunnan, China., Published as part of Wang, Fa-Lei & Zorn, Carsten, 2021, Description of a new species and new faunistic records of the genus Anomala SAMOUELLE (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions, pp. 147-159 in Beitr��ge Zur Entomologie = Contributions to Entomology 71 (1) on page 149, DOI: 10.21248/contrib.entomol.71.1.147-159, http://zenodo.org/record/5743084, {"references":["ZORN, C. 2011: New species of the genus Anomala SAMOUELLE from mainland South East Asia and South China. - Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 4: 297 - 312. - http: // www. naturkundemuseum-bw. de / sites / default / files / publikationen / serie-a / ns 04 - 15 zorn. pdf [accessed 2017 / 06 / 26].","ZORN, C.; KOBAYASHI, H. & WADA, K. 2017: Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records. - Contributions to Entomology 67 (2): 325 - 352. - DOI: https: // doi. org / 10.21248 / contrib. entomol. 67.2.325 - 352."]}

Anomala siamensis (NONFRIED, 1891) (Figs 20, 46���49) Distribution: Thailand; China (new country record): Yunnan; Laos (new country record). Type material: Syntypes [?]. 1 ��� (MFNB), Coll. Nonfried. Siam / Aprosterna siamensis Type Nonfr. [Ohaus���s handwriting] / Anomala siamensis Bang-Pain Type A. F. Nonfried [possibly Nonfried���s handwriting]; 1 ��� (MFNB), Siam Nonfried / Heteroplia n. sp. / Aprosterna siamensis Cotype Nonfr. [Ohaus���s handwriting]. Note: It is not entirely certain that these two specimens from Ohaus���s collection are real syntypes of Aprosterna siamensis NONFRIED, 1891 because they do not bear Nonfried���s type label (compare with the label shown in HORN et al. 1990). Other specimens examined: CHINA: Yunnan: 1 ��� (FLWC), Menglun, Xishuangbanna, 15.IV.2019, Zi-Chun Xiong leg.; LAOS: 1 ��� (CZGG), Laos NW, Nam Tha NPA, 65 km nw Luang Nam Tha, 1050 m, 8-15.IV.2010, S. Murzin leg.; 1 ��� (PPPC), LAOS centr 27.IV.-1.V. 1997. 70 km NE Vientiane, Ban Phabat env., 150 m N 18��16.1'E 103��10.9'E. Jendek & O. Sausa leg.; 1 ��� (PPPC), S Laos Attapu, Nong Lom (lake), 18-30.IV. 1999, 800 m, 15��02'N, 106��35'E, M. Strba leg. THAILAND: 1 ��� (PLPC), THAILAND: Phayao Prov. Naresuan University; lights on campus; 19. April 2003; L-467 Vitheepradit, Prommi, Setaphan; 1 ��� (AHPC), Thailand, 9.11.1992 12 km SW Pak-Chong 130 km NO Bangkok Mischobstplantage, 400 m Lichtfang, Thielen leg.; 1 ��� (PLPC), Thailand, Ubon Ratchathani Prov., Phu Chong Na Yoi National Park, Kaeng Ka Loa, 11. April 2004, 182 m, 14��26.178'N 105��16.631E, Sites & Vitheepradit leg.; 3 ������, 1 ��� (CZGG), NW Thailand, Mae Hong Son, Pan Huai Po, 1600 m, 9-16.V.1991, leg. P. Pachl��tko; 1 ��� (CZGG), NW Thailand, Chom Thong, 24-27.IV.1991, leg. Pachol��tko. Remarks: PAULIAN (1959) misidentified Anomala anguliceps as A. siamensis, but later ZORN et al. (2017) corrected this identification based on the male genitalia illustrated in PAULIAN (1959). Also, YANG (1989) misidentified A. anguliceps as A. siamensis when reporting this species from Guizhou, China. Both species mentioned above were also confused by LIN (1987, 1992, 2002b), who consequently described A. corneola LIN, 2002, which was subsequently synonymized with A. anguliceps by PROKOFIEV (2014). In fact, Anomala siamensis is similar to A. bilobata ARROw, 1912 and A. fissilabris ARROw, 1912 rather than A. anguliceps. Both A. bilobata and A. fissilabris have a bilobed clypeus in males, which is the same in males of A. siamensis, while the clypeal front margin is straight in A. anguliceps. Anomala siamensis was previously known only from Thailand, here this species is newly recorded from China and Laos., Published as part of Wang, Fa-Lei & Zorn, Carsten, 2021, Description of a new species and new faunistic records of the genus Anomala SAMOUELLE (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions, pp. 147-159 in Beitr��ge Zur Entomologie = Contributions to Entomology 71 (1) on page 152, DOI: 10.21248/contrib.entomol.71.1.147-159, http://zenodo.org/record/5743084, {"references":["NONFRIED, A. F. 1891: Weitere Beitrage zur Kaferfauna von Sud-Asien und Neu-Guinea. - Berliner entomologische Zeitschrift 36: 359 - 380. - https: // www. biodiversitylibrary. org / item / 101306 page / 411 / mode / 1 up [accessed 2019 / 10 / 3].","HORN, W.; KAHLE, I.; FRIESE, G. & GAEDIKE, R. 1990: Collectiones entomologicae. Ein Kompendium uber denVerbleibentomologischerSammlungenderWeltbis 1960. - Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin: 473 pp.","PAULIAN, R. 1959: Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines). - Annales de la Societe entomologique de France 128: 35 - 136. - http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6142156 w / f 14 [accessed 2017 / 06 / 26].","ZORN, C.; KOBAYASHI, H. & WADA, K. 2017: Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records. - Contributions to Entomology 67 (2): 325 - 352. - DOI: https: // doi. org / 10.21248 / contrib. entomol. 67.2.325 - 352.","YANG, C. J. 1989: Scarabaeoidea. - In: GUO, Z. Z. (ed.): The Agricultural and Forestry Insect Fauna of Guizhou, Volume 2. - China: Guizhou People's Publishing House, Guiyang: 88 pp. (in Chinese).","PROKOFIEV, A. M. 2014: New and noteworthy scarab beetles from Asia and America (Coleoptera Lamellicornia). - Calodema 330: 1 - 25. - https: // www. zin. ru / Animalia / Coleoptera / pdf / prokofiev _ 2014 _ new _ and _ noteworthy _ scarab _ beetles. pdf [accessed 2017 / 10 / 12]."]}

Anomala bilobata ARROW, 1912 (Figs 15, 29���32) Distribution: India; Myanmar; Nepal; China (new country record): Yunnan. Specimens examined: CHINA: Yunnan: 7 ������, 11 ������ (FLWC), Hulukou, Xima, Yingjiang County, Dehong, VI-VII.2018, Wei-Zong Yang leg.; 1 ��� (FLWC), Yingjiang township, Yingjiang County, Dehong, VI.2018, Yang Yu leg. NEPAL: 1 ��� (NMEG), Nepal, Prov: Narayani vic. Chandranigahaui 27��07'25"N, 85��21'11"E, 130 m, 02.V.2003, leg. A. Weigel HF/LF; 1 ��� (NMEG), Nepal, Mechi / Taplejung, 9 km S Taplejung, Angbung Kabeli Khola (bridge), 450 m, 27��17'11"N, 87��43'21"E, 03.V.2003, leg. A. Weigel. Remarks: Anomala bilobata was originally described from Bengal, India, and several localities in Myanmar (ARROw 1912b) and subsequently reported from Nepal (ZORN & BEZDĚK 2016). It is here reported from Yunnan for the first time, representing a new country record for China., Published as part of Wang, Fa-Lei & Zorn, Carsten, 2021, Description of a new species and new faunistic records of the genus Anomala SAMOUELLE (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions, pp. 147-159 in Beitr��ge Zur Entomologie = Contributions to Entomology 71 (1) on page 150, DOI: 10.21248/contrib.entomol.71.1.147-159, http://zenodo.org/record/5743084, {"references":["ARROw, G. J. 1912 b: Descriptions of some new Burmese Species of Ruteline Coleoptera belonging to the genus Anomala. - The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, series 8, 10: 327 - 340.","ZORN, C. & BEZDEK, A. 2016: Rutelinae.: 317 - 358. - In: LOBL, I. & LOBL, D. (eds.): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. - Brill, Leiden, Boston: 317 - 358."]}

Anomala kuatuna (MACHATSCHKE, 1955) (Figs 19, 43���45) Distribution: China: Fujian, Guangdong, Guangxi, Zhejiang, Guizhou (new Chinese provincial record), Hunan (new Chinese provincial record). Specimens examined: CHINA: Fujian: 1 ��� (FLWC), Sanming, VII.2014, Bi-Huang Zhan leg.; 1 ��� (FLWC), Mangdang Mount, Nanping, 5.VII.2009, no leg.; 1 ���, 2 ������ (FLWC), Damo Mount, Sanming, 15.VI.2018, Xin Liu leg. Guangdong: 3 ������ (FLWC), Houzidong Mount, Wengyuan County, Shaoguan, 18-19.V.2018, Jian-Wei Chen leg. Guangxi: 2 ������, 6 ������ (FLWC), Dayao Mount, Jinxiu County, Laibin, Guangxi, China, VII.2015, Bao-Sheng Su leg.; 1 ��� (FLWC), Ziyuan County, Guilin, Guangxi, China, VII-VIII.2019, Wen Jin leg. Guizhou: 1 ��� (FLWC), Shunchangxiang, Shuicheng County, Shuicheng, Liupanshui, China, alt. 1100 m, 22.V.2019, Gui-Qiang Huang leg. Hunan: 1 ���, 1 ��� (FLWC), Mount Yangming, Yongzhou, Hunan, China, alt. 900 m, 1.VII.2019, Yu-Tang Wang & Yi-Ting Zhong leg. Zhejiang: 1 ���, 1 ��� (FLWC), Kuocang Mount, Taizhou, Zhejiang, China, VII.2015, Zhao-Yang Tang leg. Remarks: This species is currently only known from China. Previously it was recorded from Fujian, Guangdong and Guangxi (MACHATSCHKE 1955; LIN 1993; LIN 2002b; ZORN 2011; ZORN & BEZDĚK 2016). Here we newly report this species from Guizhou and Hunan., Published as part of Wang, Fa-Lei & Zorn, Carsten, 2021, Description of a new species and new faunistic records of the genus Anomala SAMOUELLE (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions, pp. 147-159 in Beitr��ge Zur Entomologie = Contributions to Entomology 71 (1) on page 152, DOI: 10.21248/contrib.entomol.71.1.147-159, http://zenodo.org/record/5743084, {"references":["LIN, P. 1993: A systematic revision of the China Mimela: (Coleoptera: Rutelidae). - The Publishing Company of Zhong Shan University, Guangzhou: 106 pp + xxii pl. (in Chinese and English).","LIN, P. 2002 b: Rutelidae.: 387 - 427. - In: HUANG, B. (ed.): Fauna of Insects of Fujian Province of China. Volume 6. - Fujian Science & Technology Press, Fuzhou: 894 pp. (in Chinese and English).","ZORN, C. 2011: New species of the genus Anomala SAMOUELLE from mainland South East Asia and South China. - Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 4: 297 - 312. - http: // www. naturkundemuseum-bw. de / sites / default / files / publikationen / serie-a / ns 04 - 15 zorn. pdf [accessed 2017 / 06 / 26].","ZORN, C. & BEZDEK, A. 2016: Rutelinae.: 317 - 358. - In: LOBL, I. & LOBL, D. (eds.): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. - Brill, Leiden, Boston: 317 - 358."]}

Based on observations on herbarium specimens (including type material) and living plants in the wild, we demonstrate that Artemisia anomala (Asteraceae, Anthemideae), a distinctive Chinese species yet morphologically imperfectly understood since its description, is variable with respect to its leaf indumentum and shape within and between populations. The leaves are ovate, ovate-lanceolate, elliptic-lanceolate, or lanceolate, and adaxially sparsely pubescent or glabrous, abaxially tomentose to sparsely pubescent, or rarely glabrescent. Morphologically, the two currently recognized varieties of A. anomala, var. tomentella and var. acuminatissima, fall within the variation range of the species in leaf indumentum and shape, and thus are synonymized herein. Lectotypification is proposed for A. anomala.

Anomalatibialis Lansberge, 1886 Anomalatibialis Lansberge, 1886: 101 [original combination] Anomala denuda Arrow, 1899: 258 [junior subjective synonym by Limbourg 2016: 10] Bibliography. Anomalatibialis: Lansberge 1886: 101 [description]; Ohaus 1918: 93 [catalogue, distribution]; Machatschke 1957: 84 [catalogue, distribution]; Machatschke 1972: 159 [catalogue, distribution]; Krajcik 2008: 50 [checklist]; Limbourg 2016: 10 [distribution]. Anomala denuda: Arrow 1899: 258 [description]; Ohaus 1918: 90 [catalogue, distribution]; Benderitter 1928: 412 [distribution]; Machatschke 1957: 82 [catalogue, distribution]; Frey 1968: 154 [distribution]; Machatschke 1972: 157 [catalogue, distribution]; Krajcik 2008: 31 [checklist]; Limbourg 2016: 10 [lectotype designation, synonymy]. Type locality. Democratic Republic of the Congo. Current status. Valid species. Literature records. BENIN (Limbourg 2016); BURKINA FASO (Limbourg 2016); CAMEROON (Limbourg 2016); DEMOCRATIC REPUBLIC OF THE CONGO (Lansberge 1886; Benderitter 1928; Ohaus 1918; Machatschke 1957, 1974; Limbourg 2016); EQUATORIAL GUINEA (Limbourg 2016); GABON (Limbourg 2016); GHANA (Limbourg 2016); IVORY COAST (Limbourg 2016); GHANA (Limbourg 2016); LIBERIA (Limbourg 2016); MALI (Limbourg 2016); NIGERIA (Arrow 1899; Machatschke 1957, 1974; Limbourg 2016); REPUBLIC OF CONGO (Frey 1968; Limbourg 2016); SENEGAL (Limbourg 2016); SIERRA LEONE (Limbourg 2016); TOGO (Limbourg 2016); ZAMBIA (Limbourg 2016). New records. ANGOLA (new country record): 1 ♂, 2 ♀ (ZMH): Cabinda, 6.xi.1955, F. Zielinski, ZMH842062 – ZMH842064 [WSD00347530–WSD00347532]; CAMEROON: 1 ♂ (MSPC): Northwest Region, x.2011, local collector [WSD00346920]; 1 ♀ (ZMH): Douala, 22.xi.1954, F. Zielinski, ZMH842061 [WSD00347529]; 1 ♂, 1 ♀ (ZMH): Buea, 1911 ZMH842068 – ZMH842069 [WSD00347536–WSD00347537]; 1 ♂ (ZMH): Doula, 16.v.1956, F. Zilinski, ZMH842084 [WSD00347552]; DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ (MSPC): North Kivu, Kasuo, i.2013, local collector [WSD00346899]; GABON: 1 ♀ (ZMH): Owendo, Libreville, 24.xi–4.xii.1954. F. Zielinski, ZMH842066 [WSD00347534]; 2 ♂, 4 ♀ (ZMH): Gabon, Libreville and environs, ZMH842078 – ZMH842083 [WSD00347546–WSD00347551]; GHANA: 1 ♂, 2 ♀ (MSPC): Volta Region, Hohoe environs, 3–6.vi.2006, L. Veselý, [WSD00346917–WSD00346919]; IVORY COAST: 1 ♀ (ZMH): Abidjan, Elfenbeinküste, 14.xi.1954, F. Zielinski, ZMH842067 [WSD00347535]; 1 ♂, 1 ♀ (ZMH): Elfenbeinküste, Abifjan, 30.v.1956, F. Zielinski, ZMH842085 – ZMH842086 [WSD00347553–WSD00347554]: MALI: 5 ♂, 8♀ (MSPC): Bamako, vi.1968, Verner [WSD00346900– WSD00346912]; 1 ♂, 3 ♀ (MSPC): Mali, Sikasso, v.1968, Verner [WSD00346913–WSD00346916]; NIGERIA: 1 ♂ (MSPC): Taraba State, 30 km SE Serti environs, Gashaka-Gumti, 380–560m, 24.iv–8.v.2011, J. Halada [WSD00347026]; 1 ♂, 1 ♀ (ZMH): Nord Nigeria, Lokoja 1911, ZMH842059 – ZMH842060 [WSD00347527–WSD00347528]; 2 ♀ (ZMH): Lagos, Nigeria, v.1956, F. Zielinski, ZMH842070 – ZMH842071 [WSD00347538–WSD00347539]; 1 ♂ (ZMH): Burutu, 22.ii.1962, ZMH842072 [WSD00347540]; REPUBLIC OF THE CONGO: 1 ♂, 1 ♀ (ZMH): Makouna, ZMH842057 – ZMH842058 [WSD00347525–WSD00347526]; 1 ♀ (ZMH): Pointe Noire, 1.xi.1955, F. Zielinski, ZMH842065 [WSD00347533]; 5 ♀ (ZMH): Brazzaville, ZMH842073 – ZMH842077 [WSD00347541–WSD00347545]; SIERRA LEONE: 7 ♂, 5♀ (MSPC): Eastern Province, 2 km W Potolu, Mobai district, SW Kailahun, 219 m, 4–5.v.2013, P. & B. Malec [WSD00346887–WSD00346898]., Published as part of Seidel, Matthias, 2021, New records of Anomala tibialis Lansberge, 1886 and a new replacement name for Anomala tibialis Schaeffer, 1906 (Coleoptera: Scarabaeidae: Rutelinae Anomalini), pp. 494-496 in Zootaxa 4966 (4) on page 494, DOI: 10.11646/zootaxa.4966.4.9, http://zenodo.org/record/4738895, {"references":["Lansberge, J. W. (1886) Scarabaeides, Buprestides et cerambycides de l'Afrique occidentale. Notes from the Leyden Museum, 8, 69 - 120.","Arrow, G. J. (1899) On sexual dimorphism in beetles of the family Rutelidae. The Transactions of the Entomological Society of London, 1899, 255 - 269. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00667. x","Limbourg, P. (2016) Contribution to the knowledge of Afrotropical Rutelinae IV (Coleoptera: Scarabaeoidea: Melolonthidae). Belgian Journal of Entomology, 38, 1 - 42.","Ohaus, F. (1918) Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus, 66, 1 - 241. [1915]","Machatschke, J. W. (1957) Coleoptera Lamellicornia. Fam. Scarabaeidae. Subfam. Rutelinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicule 199 b. Desmet-Verteneuil, Brussels, pp. 1 - 219.","Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. Coleopterorum Catalogus, Supplementa, 66 (1), 1 - 361.","Krajcik, M. (2008) Checklist of the Scarabaeoidea of the world. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. X, Supplement 4, 1 - 142.","Benderitter, E. (1928) Rutelides nouveaux du Musee du Congo belge. Revue Zoologique Africaine, 15, 410 - 414.","Frey, G. (1968) Melolonthini und Rutelini der Expedition des Ungarischen Nationalmuseums nach Kongo / Brazzaville (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 19, 151 - 163."]}

Anomala schaefferi new name Anomalatibialis Schaeffer, 1906: 5 [original combination] Anomala (Anomalopus) tibialis Schaeffer, 1906 [subgeneric placement by Casey 1915: 40] Bibliography. Schaeffer 1906: 5 [original description]; Schaeffer 1907: 69 [identification key]; Casey 1915: 40 [identification key]; Ohaus 1918: 101 [catalogue, distribution]; Machatschke 1957: 81 [catalogue, distribution]; Machatschke 1972: 156 [catalogue, distribution]; Potts 1977a: 40 [checklist, distribution]; Potts 1977b: 133 [identification key]; Riley & Wolfe 2003: 10, 25 [checklist, new record]; Krajcik 2008: 50 [checklist]. Type locality. Texas, United States of America. Current status. Valid species. Etymology. Anomala schaefferi is a replacement name for the primary junior homonym Anomala tibialis Schaeffer, 1906. The species is dedicated to Charles Schaeffer, the author who first recognized this taxon as distinct. Literature records. UNITED STATES OF AMERICA: Texas (Schaeffer 1906; Casey 1915; Machatschke 1957, 1972; Potts 1977a, 1977b; Riley & Wolfe 2003). Remarks. The species was only known from the female holotype with unknown locality in Texas, until Riley & Wolfe (2003) reported one specimen from Padre Island, Kleberg County, Texas, which is deposited in the Texas A & M University Collection (College Station, Texas, United States of America)., Published as part of Seidel, Matthias, 2021, New records of Anomala tibialis Lansberge, 1886 and a new replacement name for Anomala tibialis Schaeffer, 1906 (Coleoptera: Scarabaeidae: Rutelinae Anomalini), pp. 494-496 in Zootaxa 4966 (4) on page 495, DOI: 10.11646/zootaxa.4966.4.9, http://zenodo.org/record/4738895, {"references":["Schaeffer, C. (1906) Notes on some species of the genus Anomala with descriptions of new species. Journal of the New York Entomological Society, 14, 1 - 5.","Casey, T. L. (1915) A review of the American species of Rutelinae, Dynastinae, and Cetoniinae. Memoires on the Coleoptera, 6, 1 - 394.","Schaeffer, C. (1907) New Scarabaeidae. Journal of the New York Entomological Society, 15, 60 - 75.","Ohaus, F. (1918) Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus, 66, 1 - 241. [1915]","Machatschke, J. W. (1957) Coleoptera Lamellicornia. Fam. Scarabaeidae. Subfam. Rutelinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicule 199 b. Desmet-Verteneuil, Brussels, pp. 1 - 219.","Machatschke, J. W. (1972) Scarabaeoidea: Melolonthidae Rutelinae. Coleopterorum Catalogus, Supplementa, 66 (1), 1 - 361.","Potts, R. W. L. (1977 a) Revision of the Scarabaeidae: Anomalinae, 2: an annotated checklist of Anomala for the United States and Canada (Coleoptera). The Pan-Pacific Entomologist, 53, 34 - 42.","Potts, R. W. L. (1977 b) Revision of the Scarabaeidae: Anomalinae. 3. A key to the species of Anomala of America north of Mexico (Coleoptera). The Pan-Pacific Entomologist, 53, 129 - 134.","Riley, E. G. & Wolfe, C. S. (2003) An annotated checklist of the Scarabaeoidea of Texas (Coleoptera). Southwest Entomologist, Supplement 26, 1 - 37.","Krajcik, M. (2008) Checklist of the Scarabaeoidea of the world. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Animma. X, Supplement 4, 1 - 142."]}

Anomala blaisei Ohaus, 1914 (Figs. 7���10, 13���21, 31) Anomala blaisei Ohaus, 1914: 196 [type localities: Annam, Keng-Trap and Tonkin, Luc-Nam], fig. 2; Paulian 1959: 43, 47, figs. 99���100; Machatschke 1972: 111; Li et al. 2012: 275 [recorded from Bokeo, Laos]; Prokofiev 2013a: 109, figs. 56���57 [photos for specimen from Tonkin, Luc-Nam]. Type material examined. One male syntype (MFNB) was examined through photos, which were provided by Carsten Zorn (Gnoien, Germany). It is labeled ��� ANNAM / Keng. Trap 8.13 / R. V. d. Salvaza // Typus! // aprosterna / blaisei / Ohs.���. Additional material examined. CHINA: 2♂♂, 1♀ (1♂, 1♀ CCPC, 1♂ ZMPC), China: Yunnan, Xishuangbanna Pref., Mengla County, Lvshilin 2010.iv.20 Xiao-Dong Yang leg.; 1♂ (CCPC), CHINA, Yunnan, Mengla, Menglun, 538m 2018, V-12, C.-C. Chen Leg. mercury light trap; 1♂ (CCPC), CHINA, Yunnan, Mengla, Menglun, 1200m 2018, V-13, C.-C. Chen Leg. daytime on flowers; 1♀ (ZMPC), Yunnan, Xishuangbanna, Yexianggu 2015-VII-11 Yi-Fan Wang leg.; 2♀♀ (SCAU), CHINA �� YUNNAN Damangjie, Nuozhadu, Lancang 100��30���35.12���E 22��33���57.02���N 620m Zhen Zhou, Xiaolong Luo, Chengcong Yu, Jianfeng Liang 2017.VII.06 Light trap LT0001; 1♂ (ZMPC), CHINA, Yunnan, Luchen, Sanmeng 1069m 2018, V-01, Y.-Q. Lu leg.; THAILAND: 1♂ (ZMPC), N. THAILAND, Chiang Mai Reg., Chiang Dao Distr. N 19��33���28��� E 99��04���34��� 8.06.2017 Vishnyakov Alexey; 1♀ (ZMPC), THAILAND N., Chiang Mai prov., 5 km W of Chiang Dao 20.- 24.4.2017, 480m N 19��24���25.9��� E 098��55���30.7��� lgt. T. Staněk & T. Staněk jun. Remarks. Anomala blaisei was described based on specimens from Annam (Keng-Trap) and Tonkin (LucNam), which might not be conspecific. Hence, a lectotype designation is probably necessary (Carsten Zorn, personal communication, 2020). The examined specimens from China and Thailand are conspecific with the syntype from Annam. Distribution. China: Yunnan (new country record); Laos; Thailand (new country record); Vietnam., Published as part of Zhao, Ming-Zhi, 2021, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part I: descriptions of two new species and remarks on four species, pp. 578-590 in Zootaxa 4903 (4) on pages 587-588, DOI: 10.11646/zootaxa.4903.4.6, http://zenodo.org/record/4431451, {"references":["Ohaus, F. (1914) XV. Beitrag zur Kenntnis der Ruteliden. Stettiner Entomologische Zeitung, 75, 193 - 217.","Paulian, R. (1959) Coleopteres scarabeides de l'Indochine (rutelines et cetonines). Annales de la Societe Entomologique de France, 128, 35 - 136.","Machatschke, J. W. (1972) Coleopterorum catalogus supplementa. Pars 66. Fascicle 1. Superfamilie Scarabaeoidea, Familie Melolonthidae, Subfamilie Rutelinae. Editio Secunda. Dr. W. Junk, Gravenhage, 361 pp.","Li, J. K., Keith, D., Gao, M. X. & Lin, L. (2012) Coleopteres Scarabaeoidea de Pha Ngam (province de Bokeo, Laos). L'Entomologiste, 68 (5), 273 - 276.","Prokofiev, A. M. (2013 a) New Anomala species from Vietnam. Russian Entomological Journal, 22, 97 - 109."]}

Anomala bella Arrow, 1917 (Figs. 49���53) Anomala bella Arrow, 1917: 208 [type designated but type locality unspecified], Plate I, fig. 9; Machatschke 1972: 124; Zorn 2006: 257; Prokofiev 2015a: 39, figs. 10���13 [recorded from North Thailand, without detailed data]; Zorn & Bezděk 2016: 330. Type material examined. The type material was not examined by the author. Labels of the male holotype (BMNH) were provided by Carsten Zorn (Gnoien, Germany) and transcribed here. It is labeled ��� ♂ // Type H. T. Mungphu // Atkinson Coll. 92-3. // Anomala bella, Arrow Type���. Additional material examined. CHINA: 3♂♂, 1♀ (2♂♂ CCPC, 1♂, 1♀ ZMPC), CHINA: Yunnan, Dehong Pref., Yingjiang County, Nabang Town, Nabang Eletric Station, 2013.v.9 Xiao-Dong Yang leg.; 1♂ (CCPC), ditto, but 2013.v.10. Comments on type locality. Even Arrow (1917) designated type and cotypes for this species originally, the type locality was not specified by him. According to the above labels, Mungphu (currently Mungpoo in northeastern India) should be considered as type locality of this taxon. Remarks. This species is widely distributed from northern India to northern Thailand. The male genitalia of Chinese specimens well fits those figured by Prokofiev (2015a). Distribution. China: Yunnan (new country record); India; Myanmar; Thailand., Published as part of Zhao, Ming-Zhi, 2021, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part I: descriptions of two new species and remarks on four species, pp. 578-590 in Zootaxa 4903 (4) on page 587, DOI: 10.11646/zootaxa.4903.4.6, http://zenodo.org/record/4431451, {"references":["Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Des - monycinae, and Euchirinae). Taylor & Francis, London, 387 pp.","Machatschke, J. W. (1972) Coleopterorum catalogus supplementa. Pars 66. Fascicle 1. Superfamilie Scarabaeoidea, Familie Melolonthidae, Subfamilie Rutelinae. Editio Secunda. Dr. W. Junk, Gravenhage, 361 pp.","Zorn, C. (2006) Subfamily Rutelinae, tribe Anomalini C. E. Blanchard, 1851. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3, Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 276 - 277.","Prokofiev, A. M. (2015 a) New Anomala Samouelle, 1819 from South-East Asia. Russian Entomological Journal, 24, 37 - 59. https: // doi. org / 10.15298 / rusentj. 24.1.04","Zorn, C. & Bezdek, A. (2016) Subfamily Rutelinae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, pp. 317 - 358."]}

Anomala granuliformis Lin, 1996 Anomala granuliformis Lin, 1996: 310 [type locality: Yunnan, Lancang], fig. 8; Zorn 2006: 260; Zorn & Bezděk 2016: 334; Zorn et al. 2017: 342 [recorded from North Vietnam]. Type material examined. None. Additional material examined. CHINA: 1♂ (ZMPC), Yunnan, Honghe, Pingbian, Mt. Daweishan 2014-VII-15 Jun Xue leg.; 1♂ (ZMPC), Yunnan, Xishuangbanna, Mengyang, Manjingkan Village 2020. V.20 light trap Xun Li leg.; 1♀ (CCPC), Yunnan, Dehong, Yingjiang, Nabang Eletric Station 2013. V.9 Xiao-Dong Yang leg.; LAOS: 3♂♂, 1♀ (ZMPC), LAOS: Xieng Khouang prov., Muang Khoun 2019. V.5���15 local collector leg.; THAILAND : 1♂ (ZMPC), Thailand: Tambon Na Chom Thian, Amphoe Sattahip, Chang Wat Chon Buri. 12��51���04.1���N 100��54���13.7���E. 12-24.III.2013 S A. A. Potanina. D. V. Potanin leg.; VIETNAM: 2♂♂ (ZMPC), VIETNAM: Quang Nam prov., Tay Giang dist., Mt. Axan 1300m 2018.IV local collector leg. Remarks. This species was originally reported from southern China (Yunnan) and northern Thailand and subsequently from northern Vietnam (Zorn et al. 2017). It is reported from northern Laos (Xieng Khouang), eastern Thailand (Chon Buri) and central Vietnam (Quang Nam) herein. The examined material extends its known distribution range along the southern boundary of Yunnan. The female from Yingjiang County suggests possible occurrence of this species in northern Myanmar. Distribution. China; Laos (new country record); Thailand; Vietnam., Published as part of Zhao, Ming-Zhi, 2021, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part I: descriptions of two new species and remarks on four species, pp. 578-590 in Zootaxa 4903 (4) on pages 588-589, DOI: 10.11646/zootaxa.4903.4.6, http://zenodo.org/record/4431451, {"references":["Lin, P. (1996) Anomala cupripes species group of China and a discussion on its taxonomy (Coleoptera: Rutelidae). Entomologia Sinica, 3 (4), 300 - 313. https: // doi. org / 10.1111 / j. 1744 - 7917.1996. tb 00279. x","Zorn, C. (2006) Subfamily Rutelinae, tribe Anomalini C. E. Blanchard, 1851. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3, Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 276 - 277.","Zorn, C. & Bezdek, A. (2016) Subfamily Rutelinae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, pp. 317 - 358.","Zorn, C., Kobayashi, H. & Wada, K. (2017) Notes on the genus Anomala Samouelle, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records. Beitrage zur Entomologie, 67 (2), 325 - 352. https: // doi. org / 10.21248 / contrib. entomol. 67.2.325 - 352"]}

Two new species of the genus Anomala Samouelle, 1819 are described: A. dianopicta Zhao, new species from Yunnan and A. huangyuzhoui Zhao, new species from Hunan. New distributional records for Anomala bella Arrow, 1917, A. blaisei Ohaus, 1914, A. cyanipennis Lin, 1999 and A. granuliformis Lin, 1996 are presented.

Anomala cyanipennis Lin, 1999 Anomala cyanipennis Lin, 1999: 162, 169 [type locality: China, Yunnan, Menghai, Mt. Nannuoshan], fig. 10; Zorn 2006: 258; Zorn & Bezděk 2016: 331; Zorn et al. 2017: 340 [recorded from Laos and Vietnam]. Type material examined. None. Additional material examined. MYANMAR: 1♂, 1♀ (ZMPC), MYANMAR N (Burma) Putao, H-500M. 27.04.1998 leg. S. Murzin & V. Siniaev ex. coll. V. Sinaev. Remarks. This species was previously recorded from China, Laos and Vietnam (Lin 1999; Zorn et al. 2017). The two specimens from northern Myanmar greatly expand its known northern- and westernmost distribution border. The examined female belongs to an unicolor variation, which is entirely reddish brown. Females of the unicolor form coexist with females of the normal form (with generally yellowish to reddish body and blackish blue elytra, as in the original description) in the populations from northern Laos and northern Vietnam but are underrepresented (Carsten Zorn, personal communication, 2020). Distribution. China; Laos; Myanmar (new country record); Vietnam., Published as part of Zhao, Ming-Zhi, 2021, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part I: descriptions of two new species and remarks on four species, pp. 578-590 in Zootaxa 4903 (4) on page 588, DOI: 10.11646/zootaxa.4903.4.6, http://zenodo.org/record/4431451, {"references":["Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxo - nomia, 21, 157 - 176.","Zorn, C. (2006) Subfamily Rutelinae, tribe Anomalini C. E. Blanchard, 1851. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3, Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Apollo Books, Stenstrup, pp. 276 - 277.","Zorn, C. & Bezdek, A. (2016) Subfamily Rutelinae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, pp. 317 - 358.","Zorn, C., Kobayashi, H. & Wada, K. (2017) Notes on the genus Anomala Samouelle, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records. Beitrage zur Entomologie, 67 (2), 325 - 352. https: // doi. org / 10.21248 / contrib. entomol. 67.2.325 - 352"]}

Anomala ludoviciana Schaeffer, 1906 Potts 1977; Hardy 1991; ISIC; IWUC; UMRC; USNM; DAVC; ELFC. Collection dates: April, June–July. Collected sweeping prairie and in flight over prairie. Specimen label data: Cedar Hills Sand Prairie State Preserve, Black Hawk Co., Iowa, 5 July 2003, sweeping, ELF (ELFC); Marietta Sand Prairie State Preserve, Marshall Co., Iowa, 13 June 2003, vervain flower (Verbena stricta), ELF (ELFC). County records (Figure 166): Black Hawk, Linn, Marshall, Monona, Pottawattamie, Story, Woodbury., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 45, DOI: 10.5281/zenodo.5353792, {"references":["Potts, R. W. L. 1977. Revision of the Scarabaeidae: Anomalinae 2. An annotated checklist of the Anomala of the United States and Canada. The Pan-Pacific Entomologist 53 (1): 34 - 42.","Hardy, A. 1991. A catalog of Coleoptera of America north of Mexico. Family: Scarabaeidae. Subfamilies Rutelinae and Dynastinae. Volume 34, part 2. United States Department of Agriculture Handbook 529: 1 - 57."]}

Anomala binotata (Gyllenhal, 1817) Wickham 1911; Potts 1977; Hardy 1991; ISIC; IWUC; DAVC; ELFC. Collection dates: March–June. Collected at light and in flight during day. Specimen label data: Shimek State Forest, Van Buren Co., Iowa, 30 May 2003, light, ELF (ELFC). County records (Figure 163): Des Moines, Fremont, Henry, Linn, Louisa, Johnson, Monona, Page, Pottawattamie, Scott, Story, Van Buren, Webster., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 44, DOI: 10.5281/zenodo.5353792, {"references":["Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40.","Potts, R. W. L. 1977. Revision of the Scarabaeidae: Anomalinae 2. An annotated checklist of the Anomala of the United States and Canada. The Pan-Pacific Entomologist 53 (1): 34 - 42.","Hardy, A. 1991. A catalog of Coleoptera of America north of Mexico. Family: Scarabaeidae. Subfamilies Rutelinae and Dynastinae. Volume 34, part 2. United States Department of Agriculture Handbook 529: 1 - 57."]}

Anomala flavipennis Burmeister, 1844 DAVC; ELFC. Collection dates: June–July. Collected at light. Specimen label data: First known Iowa specimens and NEW STATE RECORD: Waubonsie State Park, Fremont Co., Iowa, 3 July 2003, UV light, DAV (DAVC); Shimek State Forest, Van Buren Co., Iowa, 4 July 2004, UV light, ELF (ELFC). County records (Figure 164): Fremont, Pottawattamie, Van Buren., Published as part of Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, pp. 1-83 in Insecta Mundi 2020 (787) on page 44, DOI: 10.5281/zenodo.5353792

This extinct cicadomorphan family Hylicellidae was a significant element of the Late Triassic (Norian) hemipteran fauna of south-eastern Queensland, represented by the tegmina of five monotypic genera from the fossil insect localities at Mount Crosby, Denmark Hill and Dinmore. Of the five, Mesothymbris Evans, 1956, from Mount Crosby, has previously been re-examined. The present work revises the remaining four genera: Hylicella Evans, 1956, with Hylicella colorata Evans, 1956, Mount Crosby; Crosbella Evans, 1956 (= Mesocicadella Evans, 1956, syn. nov.), with Crosbella elongata Evans, 1956 (= Crosbella alata Evans, 1956, syn. nov. = Mesocicadella venosa Evans, 1956, syn. nov. = Mesocicadella punctata Evans, 1961, syn. nov.), Mount Crosby; Triassoscelis Evans, 1956, with Triassoscelis anomala Evans, 1956, Mount Crosby; and Mesocixiodes Tillyard, 1922, with Mesocixiodes termioneura Tillyard, 1922 (= Mesocixiodes brachyclada Tillyard, 1922, syn. nov.), Denmark Hill, Dinmore. The Hylicellidae remains poorly defined as is its generic composition, however, the four Queensland Triassic genera are quite distinct. Hylicella is distinguished by the partial fusion of CuA1 with M3+4, and the very large intra-medial cell, and Crosbella and Triassoscelis by the peculiar parallelogram-like CuA1 cell and the widened peripheral membrane at CuA. Triassoscelis differs from Crosbella in its broader, more squat wing shape, CuA fork in line rather than proximal to the M fork, and much less proliferated apical venation. Mesocixiodes is distinguished by the alignment of the crossvein field at the claval apex to form a fairly even series of long, sub-equal apical cells. On the basis of the proliferated and variable apical venation, the presence of weakly developed costal veinlets, and the fusion of M and CuA at a single point to close the basal cell, Crosbella and Triassoscelis, are ascribed to the subfamily Vietocyclinae. The relationships of the three other Queensland genera, Hylicella, Mesocixiodes, and Mesothymbris remain obscure, and they are retained in the undefined catch-all subfamily Hylicellinae. With the exception of the hindwing, Mesojassula marginata Evans, 1956, the revision of the cicadomorphan fauna of the Mount Crosby Formation is now complete, resulting in an inventory of five species of Dysmorphoptilidae, four of Hylicellidae and two of Archijassidae, with the dysmorphoptilid Dysmorphoptiloides elongata Evans, 1956, as the dominant species.

We recognize 30 species of the pantropical genus Xylopia L. from Madagascar and an additional three species from the Mascarene Islands. Of the six sections of the genus, three are represented in the region: sect. Xylopia, sect. Verdcourtia, and sect. Stenoxylopia. All species are endemic, and many are microendemics of conservation concern. The greatest species richness occurs in humid forests below 1000 m. Taxonomic review supports the hypothesis that the genus dispersed to Madagascar from continental Africa five times. Ten new species are described: Xylopia anomala D.M.Johnson & N.A.Murray, sp. nov., X. australis D.M.Johnson & N.A.Murray, sp. nov., X. carinata D.M.Johnson & N.A.Murray, sp. nov., X. galokothamna D.M.Johnson & N.A.Murray, sp. nov., X. lokobensis D.M.Johnson & N.A.Murray, sp. nov., X. longirostra D.M.Johnson & N.A.Murray, sp. nov., X. marojejyana D.M.Johnson & N.A.Murray, sp. nov., X. ravelonarivoi D.M.Johnson & N.A.Murray, sp. nov., X. retusa D.M.Johnson & N.A.Murray, sp. nov., and X. sclerophylla D.M.Johnson & N.A.Murray, sp. nov. An identification key, technical descriptions, illustrations, and distribution maps are provided.

Anomala fuscicauda Lin, 1999 (Figs. 5, 11, 16, 21, 44) Material examined. CHINA: Yunnan. 4♂♂ and 18♀♀ (CFLW): Xingguo, Pingyuan, Yingjiang County, Dehong, 97��99���23.82��� E, 24��68���40.28��� N, 30.V.2018, Bao-Ying Jin leg.; 2♂♂ and 3♀♀ (CFLW): Xingguo, Pingyuan, Yingjiang County, Dehong, 15���30. VI. 2019, Chang-Gui Liu leg. Distribution. China: Yunnan; Myanmar (new country record). Remarks. This species was also recorded from the Oriental region (Zorn & Bezděk 2016), based on specimens from Putao, northern Myanmar (Carsten Zorn, personal communication)., Published as part of Huang, Gui-Qiang & Wang, Fa-Lei, 2019, Two new and one newly recorded species of Anomala Samouelle, 1819 (Coleoptera Scarabaeidae: Rutelinae) from Yunnan, China, pp. 366-374 in Zootaxa 4706 (2) on page 370, DOI: 10.11646/zootaxa.4706.2.9, http://zenodo.org/record/3567473, {"references":["Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxonomia, 21, 157 - 176.","Zorn, C. & Bezdek, A. (2016) Rutelinae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Revised and Updated Edition. Brill, Leiden & Boston, pp. 317 - 358."]}

Anomala qiului Huang & Wang, new species (Figs. 1���4, 10, 15, 20, 28, 44) Type material. Holotype: ♂ (MYNU): Nabang, Yingjiang County, Dehong, Yunnan, China, 97��56���26.28��� E, 24��75���04.56��� N, alt. 640 m, 2.IV.2018, Lu Qiu leg. Paratypes: 1♂ and 2♀♀ (CFLW): same data as the holotype; 1♂ (CMZZ): Nabang, Yingjiang County, Dehong, Yunnan, China, 9.V.2013, Xiao-Dong Yang leg.; 1♀ (CCCC): Nabang, Yingjiang County, Dehong, Yunnan, China, 8.V.2018, Xiao-Dong Yang leg. Description. Male, holotype (Figs. 1���2, 10, 15, 20). General. Length 13.6 mm, greatest width 9.4 mm; body elongate ovoid, moderately convex in profile, sub-lustrous. Color. Whole body black, antennae brown. Head. Clypeus sub-trapezoidal, rounded apically, with dense and fine punctures, anterior margin slightly reflexed towards dorsum; clypeofrontal suture extremely weak; frons with dense punctures (sparse at base), the punctures large on disc, small near eyes; interocular distance 0.63 times as wide as the maximum transverse width of head; antenna sparsely covered with yellow hairs, antennal club shorter than antennomeres 2���6. Pronotum. 1.5 times as broad as long, sides expanded in middle; basal part of the sides with short marginal line, basal marginal line not interrupted in the middle, gradually disappearing towards the sides; anterior angles obtuse, posterior angles rounded apically; disc densely covered with punctures but densest at sides; punctures on sides moderately large, gradually smaller towards anterior angles; with a distinct longitudinal smooth area from basal margin to the center of disc; with several long yellow setae along lateral margins. Scutellum. Tongue-shaped, 1.2 times as wide as long; with sparse punctures, punctures only dense basally; posterior margin smooth. Elytra. Approximately 1.17 times as long wide, humeral umbone and apical protuberance prominent and smooth, strial punctures dense, large and deep, forming several moderately deep longitudinal striae, intervals between striae smooth; elytra with large, dense and irregular punctures near base of suture, with small, sparse and rounded punctures close to sides of scutellum; epipleuron starting apical of level of humeral umbone, flat and gradually expanded at basal 1/2, ending at apical 1/4. Pygidium. Subtriangular and moderately convex in profile, with transverse punctures on disc, punctures gradually sparser towards sides; posterior marginal line complete, with brown setae at posterior margin. Abdominal ventrites. Abdominal ventrites 1���3 with fine punctures, the punctures denser in center and at sides; abdominal ventrite 4 with moderately dense transverse punctures, gradually finer from center to sides, denser in center and at sides; abdominal ventrite 5 with dense, fine punctures; abdominal ventrites 1���3 with a transverse rows of sparse, short yellow-brown setae near middle, the setae slightly denser at sides; abdominal ventrite 4 with a row of short yellowish brown setae near apical margin; abdominal ventrite 5 with a row of yellowish brown setae near apex. Legs. All tarsi short, metatarsus rather strong; protarsal inner claw and mesotarsal outer claw cleft, mesotarsal inner and metatarsal inner and outer claws simple and long; protibia bidentate, apical tooth produced and rounded apically, slightly extending beyond apex of first segment of tarsus, basal tooth short and obtuse, inner spur short; mesotibia slightly fusiform, metatibia very strong and fusiform; both mesotibia and metatibia densely covered with yellow hairs at inner sides, with two terminal spurs, the upper one 1.65 times as long as the lower one. Genitalia. As in Figs. 10, 15, 20. See differential diagnosis. Variability. Body length 12.5���13.6 mm, greatest width 9.1���9.4 mm in males. The holotype and male paratypes show no differences in the shape of the aedeagus. Females (Figs. 3, 4, 28). Similar to male, but body larger, body length 19.7���20.0 mm, greatest width 9.8���9.9 mm; antennal club shorter, apical margin of pronotum wider (compare with basal margin); apical tooth of protibia more rounded apically, distinctly extending beyond apex of second segment of tarsus, basal tooth right-angled. Differential diagnosis. Around Himalaya region (China, India, Bhutan and Myanmar), four morphologically similar species, Anomala fuscicauda Lin, 1999, A. merula Arrow, 1917, A. trochanterica Arrow, 1917 and A. validipes Arrow, 1917 share following characters with the new species: pronotum attenuated anteriorly, causing the anterior somewhat tapered body shape, sides of pronotum expanded in middle, posterior angles rounded apically; all tarsi short, hind legs extremely strong (compare with fore and mid legs). However, A. qiului new species can be distinguished from the four similar species by extremely asymmetric parameres of male genitalia (Figs. 10���26). Except for male genitalia, A. qiului new species differs from A. merula and A. trochanterica by the disc of pronotum with deeper punctures, elytra with large and irregular dense punctures near base of suture, with deeper striae and more convex intervals. Besides, females of A. qiului new species can be distinguished from A. trochanterica by two simple vaginal palps, whereas A. trochanterica with two additional sclerites in the membrane connecting the vaginal palps. Anomala qiului new species differs from A. fuscicauda by body black (brown in A. fuscicauda), and elytra with several moderately deep and regular longitudinal striae and smooth intervals. Anomala qiului new species differs from A. validipes by body black (reddish brown in A. validipes). Etymology. The new species is named after Dr. Lu Qiu (DZ���) from Southwest University, Chongqing, China, who collected most of type material. Distribution. The new species is only known from Nabang county, Yunnan province, China., Published as part of Huang, Gui-Qiang & Wang, Fa-Lei, 2019, Two new and one newly recorded species of Anomala Samouelle, 1819 (Coleoptera Scarabaeidae: Rutelinae) from Yunnan, China, pp. 366-374 in Zootaxa 4706 (2) on pages 367-370, DOI: 10.11646/zootaxa.4706.2.9, http://zenodo.org/record/3567473, {"references":["Lin, P. (1999) New species of the genus Anomala Samouelle (Coleoptera: Rutelidae) collected in Yunnan, China. Entomotaxonomia, 21, 157 - 176.","Arrow, G. J. (1917) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia part II (Rutelinae, Desmonycinae, and Euchirinae). Taylor & Francis, London, 387 pp."]}

Two new species, Anomala qiului Huang & Wang, new species and Anomala xuhaoi Huang & Wang, new species, are described and illustrated from Yunnan, China. Anomala trochanterica Arrow, 1917 is recorded from China for the first time, with the report of its intraspecific variation also discussed. Color plates (habitus and genitalia) of A. flavipunctuata Lin, 1999, A. flavoguttata Miyake, 2000, A. fuscicauda Lin, 1999 and A. trochanterica are provided for the first time.

Sponges of the Class Homoscleromorpha, despite their high evolutionary significance, are still poorly known due to their small size and cryptic habit. In this study, we describe five species of the homoscleromorph genus Plakina from submarine caves in the Aegean Sea (Lesvos Island, Greece): Plakina bowerbanki, P. trilopha, P. anisoactina sp. nov., P. anomala sp. nov., and P. hellenica sp. nov. Plakina anomala sp. nov. is the first species of the genus with papillae and the first Homoscleromorpha with a microcavity-filling habit. Plakina bowerbanki, P. anomala sp. nov. and P. anisoactina sp. nov. present exceptionally stout, annulated lophose spicules with highly variable ramification patterns, including some original types. The high diversity of Plakina in the two caves of Lesvos Island is similar to that of northwestern Mediterranean caves. Despite being much less studied, the Aegean Sea now equals the Western Mediterranean as the two ecoregions of the Mediterranean with the highest diversity of Plakina, with eight species each. The Aegean Sea encompasses a huge number of submarine caves, and therefore it has great potential for the discovery of new species of Homoscleromorpha.

Anomala undulata Melsheimer, 1846 Anomala undulata Melsheimer 1846: 140 Note. Seven presently recognized subspecies of Anomala undulata have been described, one of which occurs in the research area. The other six are A. undulata aeneipennis Blanchard, 1851 (recorded from Brazil), A. undulata brasiliensis Arrow, 1899 (recorded from Brazil, Venezuela, Colombia), A. undulata collaris Burmeister, 1844 (recorded from Colombia), A. undulata espiritosantensis Ohaus, 1902 (recorded from Brazil, Colombia), A. undulata peruviana (Gu��rin-M��neville, 1838) (recorded from Ecuador, Peru) and A. undulata undulata Melsheimer, 1846 (recorded from Belize, Guatemala, Jamaica, Mexico, U.S.A.). Subspecies Anomala undulata varians (Fabricius, 1801) Melolontha varians Fabricius 1801b: 173 (Guyana) Distribution. Guyana: Ohaus 1897: 401 (TYPE Melolontha varians Fabricius); Bodkin 1919: 217 (as A. undulata [s. l.]). French Guiana: Br��l�� and Dalens 2012b: 37 (as A. undulata [s. l.]); Br��l�� et al. 2014: 182 (as A. undulata [s. l.]; as Paranomala undulata [s. l.]). ���Guianas���: Blackwelder 1944: 245; Machatschke 1972: 146. Other: Fabricius 1801b: 173 (South America - as Melolontha varians). Note. The validity of the subspecific epithet of Anomala undulata varians (Fabricius, 1801) needs to be evaluated because it is a junior homonym of Anomala varians (Olivier, 1789) (p. 78) (as noted in Ohaus (1897: 401)). Both taxa were originally described in the genus Melolontha., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 120, DOI: 10.5281/zenodo.3678492, {"references":["Melsheimer, F. E. 1846. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2 (6): 134 - 160.","Blanchard, C. E. 1851. Ordre des Coleoptera. p. 129 - 240. In: H. Milne-Edwards. Catalogue de la collection entomologique du Museum d'Histoire Naturelle de Paris. Tome 1. Gide & Baudry; Paris. 240 p.","Arrow, G. J. 1899 a. Notes on the rutelid genera Anomala, Mimela, Popillia, and Strigoderma. Transactions of the Entomological Society of London 1899 (2): 271 - 276.","Burmeister, H. C. C. 1844. Handbuch der Entomologie. Vierter Band. Erste Abtheilung. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Anthobia et Phyllophaga Systellochela. Theod. Chr. Fr. Enslin; Berlin. 587 p.","Ohaus, F. 1902. Beitrage zur Kenntniss der Ruteliden. Entomologische Zeitung (Stettin) 63 (1): 3 - 57.","Guerin-Meneville, F. E. 1838 (1830). Crustaces, arachnides et insectes. p. 1 - 319. In: L. I. Duperrey. Voyage autour du monde, execute par ordre du roi, sur la corvette de la majeste, la Coquille, pendant les annees 1822, 1823, 1824 et 1825, sous le ministere et conformement aux instructions de s. e. m. le marquis de Clermont-Tonnerre, ministre de la marine; et publie sous les auspices de son excellence Mgr le Cte de Chabrol, ministre de la marine et des colonies. Zoologie. Tome second. 2 e Partie. Premiere division. Arthus Bertrand; Paris. 319 p. [Note. The date of publication is discussed in Sherborn and Woodward (1906: 336), Cowan (1970: 360), Jameson and Smith (2002: 325) and Bousquet (2016: 236)].","Fabricius, J. C. 1801 a. Systema eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Tomus I. Impensis Bibliopolii Academici Novi; Kiel, Germany. 506 p.","Fabricius, J. C. 1801 b. Systema eleutheratorum secundum ordines, genera, species: adjectis synonymis, locis, observationibus, descriptionibus. Tomus II. Impensis Bibliopolii Academici Novi; Kiel, Germany. 687 p.","Ohaus, F. 1897. Beitrage zur Kenntniss der Ruteliden. II. Anomaliden von Mittel- und Sud-Amerika. Entomologische Zeitung (Stettin) 58 (7 - 12): 383 - 440.","Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. The Entomologist's Monthly Magazine 55: 210 - 219.","Brule, S., and P. - H. Dalens. 2012 b. Resultats de la mission entomologique de AYA, Reserve de la Trinite (Guyane), Mai 2012. Rapport de la Societe entomologique Antilles-Guyane, SEAG, ONF. 31 p. + annexes.","Brule, S., J. Touroult, E. Poirier, and P. - H. Dalens. 2014. Resultats de l'etude-inventaire entomologique du site de la Montagne Pelee, Saul (Guyane), 2010 - 2012. Rapport de la Societe entomologique Antilles-Guyane. SEAG, PAG. 144 p. + annexes.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p.","Olivier, G. A. 1789. Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Tome premier. Baudouin; Paris. [Note. Each genus is separately paginated]."]}

Anomala brunnipennis (Gyllenhal, 1817) Melolontha brunnipennis Gyllenhal 1817: 108 (America) = Euchlora limbata Laporte 1840: 136 (French Guiana) Distribution. Suriname: Blackwelder 1944: 243; Machatschke 1972: 144; MHCN; NZCS. French Guiana: Laporte 1840: 136 (as Euchlora limbata); Blackwelder 1944: 244 (as A. limbata); Machatschke 1972: 144; Soula 2010a: 53 (as A. brunipennis [sic]). Other: Gyllenhal 1817: 108 (America - as Melolontha brunnipennis)., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 119, DOI: 10.5281/zenodo.3678492, {"references":["Gyllenhal, L. 1817. Melolontha brunnipennis. p. 108 - 109. In: Appendix ad C. J. Schonherr. Synonymiam insectorum. Tom. 1. Part 3. Sistens descriptiones novarum specierum. Lewerentziana; Skara, Sweden. 266 p.","Laporte de Castelnau, F. L. 1840. Histoire naturelle des insectes coleopteres. Tome deuxieme. p. 1 - 563. In: C. E. Blanchard, F. L. Laporte de Castelnau, and P. H. Lucas. Histoire naturelle des animaux articules, annalides, crustaces, arachnides, myriapodes et insectes. P. Dumenil; Paris. 564 p.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p.","Soula, M. 2010 a. Les Rutelinae: presentation des tribus et genres de Guyane (Coleoptera, Scarabaeidae). Contribution a l'etude des Coleopteres de Guyane 2: 50 - 61. [Supplement au Bulletin de liaison d'ACOREP-France \" Le Coleopteriste \"]."]}

Anomala vayana Ohaus, 1930 Anomala vayana Ohaus 1930: 147 (Guyana) Distribution. Guyana: Ohaus 1930: 147; Blackwelder 1944: 245; Machatschke 1972: 151., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 120, DOI: 10.5281/zenodo.3678492, {"references":["Ohaus, F. 1930. XXVI. Beitrag zur Kenntnis der Rutelinen (Col. Lamell.). Deutsche Entomologische Zeitschrift 1930 (2 - 3): 138 - 158.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p."]}

Anomala inconstans Burmeister, 1844 Anomala (Anomala) inconstans Burmeister 1844: 252 (Brazil���Mexico) Distribution. Suriname: MHCN - NEW COUNTRY RECORD (Suriname, Para.). Brazil: Burmeister 1844: 252; Ohaus 1897: 406; Blackwelder 1944: 244. Venezuela: Ohaus 1897: 406. Other: Burmeister 1844: 252 (Brazil���Mexico); Ohaus 1897: 406 (Paraguay); Blackwelder 1944: 244 (Colombia, Costa Rica, Guatemala, Mexico, Nicaragua, Panama, Saint Lucia, Saint Vincent); Machatschke 1972: 149 (Central America, South America)., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 119, DOI: 10.5281/zenodo.3678492, {"references":["Burmeister, H. C. C. 1844. Handbuch der Entomologie. Vierter Band. Erste Abtheilung. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Anthobia et Phyllophaga Systellochela. Theod. Chr. Fr. Enslin; Berlin. 587 p.","Ohaus, F. 1897. Beitrage zur Kenntniss der Ruteliden. II. Anomaliden von Mittel- und Sud-Amerika. Entomologische Zeitung (Stettin) 58 (7 - 12): 383 - 440.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p."]}

Anomala arrawaka Ohaus, 1902 Anomala arrawaka Ohaus 1902: 44 (Suriname) Distribution. Suriname: Ohaus 1902: 44; Blackwelder 1944: 243; Machatschke 1972: 147 (as A. arawaka [sic])., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 119, DOI: 10.5281/zenodo.3678492, {"references":["Ohaus, F. 1902. Beitrage zur Kenntniss der Ruteliden. Entomologische Zeitung (Stettin) 63 (1): 3 - 57.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p."]}

Anomala jordani Ohaus, 1902 Anomala jordani Ohaus 1902: 42 (Guyana) Distribution. Guyana: Ohaus 1902: 42; Machatschke 1972: 148. ���Guianas���: Blackwelder 1944: 244., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 120, DOI: 10.5281/zenodo.3678492, {"references":["Ohaus, F. 1902. Beitrage zur Kenntniss der Ruteliden. Entomologische Zeitung (Stettin) 63 (1): 3 - 57.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341."]}

Anomala limbaticollis Blanchard, 1851 Anomala limbaticollis Blanchard 1851: 188 (Nouv.- Grenade [= Colombia?]) Distribution. Guyana: Bodkin 1919: 217. Other: Blanchard 1851: 188 (Nouv.-Grenade [= Colombia?]); Blackwelder 1944: 244 (Colombia); Machatschke 1972: 151 (Neu Grenada [= Colombia?]). Note. Bodkin (1919: 217) records Anomala limbaticollis for Guyana. Because of the distance between this country and the assumed type locality, as well as the absence of any recent records, we consider the presence of this species in the research area in need of confirmation., Published as part of Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, pp. 1-306 in Insecta Mundi 732 (732) on page 120, DOI: 10.5281/zenodo.3678492, {"references":["Blanchard, C. E. 1851. Ordre des Coleoptera. p. 129 - 240. In: H. Milne-Edwards. Catalogue de la collection entomologique du Museum d'Histoire Naturelle de Paris. Tome 1. Gide & Baudry; Paris. 240 p.","Bodkin, G. E. 1919. Notes on the Coleoptera of British Guiana. The Entomologist's Monthly Magazine 55: 210 - 219.","Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185 (2): 189 - 341.","Machatschke, J. W. 1972. Scarabaeoidea, Melolonthidae, Rutelinae. p. 1 - 361. In: S. Schenkling. Coleopterorum Catalogus Supplementa Pars 66 (1). W. Junk; The Hague, Netherlands. 361 p."]}

Anomala esmeralda Prokofiev, 2013 (Figs 2A���B, 3D, 4D, 5D���F) Anomala esmeralda Prokofiev, 2013: 101 (original description); ZORN et al. (2017): 341 (collecting record). Type locality. ��� Vietnam, Khanh Hoa prov., Khanh Vinh distr., DT 652, 12��13.55���N, 108��46.02���E, alt. 877��� 879m.��� Material examined (3 spec.). 1 J 2 ♀♀ (ZMGC), Mt. Axan, Tay Giang, Quang Nam Province, Vietnam, ca. N 15��49���04��� E 107��18���12���, alt. 1200m, viii.2017, local collector leg. Notes. The examined material presents a new record for Vietnam (Qu���ng Nam Province), which documents the northernmost known distribution of this species. Distribution. Vietnam (Kh��nh Hoa, L��m Đ���ng, Gia Lai, Kon Tum, Qu���ng Nam)., Published as part of Zhao, Ming Zhi, 2019, A new species of the genusAnomala from southern China (Coleoptera: Scarabaeidae: Rutelinae), pp. 481-489 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 59 (2) on page 487, DOI: 10.2478/aemnp-2019-0037, http://zenodo.org/record/5454583, {"references":["PROKOFIEV A. M. 2013: New Anomala species from Vietnam. Russian Entomological Journal 22: 97 - 109.","ZORN C., KOBAYASHI H. & WADA K. 2017: Notes on the genus Anomala Samouelle, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records. Beitrage zur Entomologie 67: 325 - 352."]}

Some polysphinctine wasps of the genus Zatypota complete their life cycles upon theridiid host spiders. The host range of these wasps is usually species-specific, although in some less common associations more than one wasp species interacts with the same host spider. Here we describe and illustrate the polysphinctine wasps Zatypota baezae sp. n. and Zatypota mulunguensis sp. n. (Hymenoptera: Ichneumonidae), both koinobiont ectoparasitoids of the spider Anelosimus baeza (Theridiidae). The two parasitoid wasps show the same development time (12 days) which was longer when compared with other parasitoid wasps Z. anomala Holmgren and Z. riverai Gauld (nine days). As described for other species of Zatypota and Hymenoepimecis , the second larval instar remains attached to the spider by the remains of the chorion and also by a rigid brownish-semitransparent membrane called a saddle.

In this article, I describe a new endemic genus of New Caledonian Eumolpinae, Kumatoeides gen. nov. (tribe Eumolpini), and eight new species currently conforming the genus: K. anomala sp. nov., K. aulacia sp. nov. (generic type), K. leptalei sp. nov., K. megale sp. nov., K. metallica sp. nov., K. millei sp. nov., K. tarsalis sp. nov. and K. wanati sp. nov. Some easily recognizable defining traits of this new genus of fairly small species (2.23–3.20 mm) are the glabrous dorsum, aligned punctures on elytra in eight regular striae plus short scutellar and a subhumeral striae, convex elytral intervals, and the enlarged and elongate first pro- and mesotarsomeres of males. One species originally described in the genus Montrouzierella Jolivet, Verma et Mille is transferred to the new genus as Kumatoeides costata (Jolivet, Verma et Mille) nov. comb. A preliminary phylogenetic analysis using partial cox1 and rrnS mtDNA sequences of five species of Kumatoeides gen. nov. together with a representative sample of species of Eumolpinae from New Caledonia supports the monophyly of the genus and the existence of two main groups as recognized based on morphological traits. However, data are not conclusive about the relationship of the new genus with other New Caledonian lineages, although suggest a closest relationship with some unidentified species most similar to some representatives of the genus Samuelsonia Jolivet, Verma et Mille but with morphological similarities with Kumatoeides gen. nov., including dilated male protarsomeres, similar antennomere proportions and tendency to aligned rows of punctures on elytra, at least apically.

New information of the genus Barsine Walker, 1854 is provided. A new species, Barsine paraprominens Huang & Wang sp. nov. is described from Pailong, SE Xizang. Four species: B. pseudomactans Volynkin & Černý, 2016, B. anomala (Elwes, 1890), B. pseudocardinalis Volynkin & Černý, 2017 and B. ustrina Černý, 2009 are recorded as new to China. The female genitalia of B. ustrina are described for the first time. Adults and genitalia of all species mentioned are illustrated.

Anomala flavoguttata MIYAKE, 2000 Distribution: China (new country record), Vietnam. Specimens examined: 1 ���, N. VIETNAM ��� Lao Cai prov. Hoang Lien NP, surr. Sa Pa pass, at light (~ 1900 m) 27.V.-2.VI.2011, L. Bartolozzi, S. Bambi, F. Fabiano, E. Orbach leg. (CZPC); 2 ������, 1 ���, VIETNAM, Lao Cai Prov., Hoang Lien NP, Tram Ton, 1915 m, 22.3493723��N, 103.7704565��E, forest edge, at light, 8-11.IV.2010, leg. L. Papp, L. Peregovits & Z. Solt��sz VN 2010PL_4 (CZPC, HNHM); 1 ���, Yunnan, Dawei Mount, hotel at hilltop, Pingbian, 2014/5/25-30, Mao Ye leg. (WFPC) (examined by photographs); 1 ���, Yunnan, Hong Qi Reservoir, Pingbian, Dawei Mount, foothills, 2016/5/20, Lu Qiu leg. (WFPC) (examined by photographs). Remarks: Only the holotype of Anomala flavoguttata was known previously. The recently discovered specimens reported above were collected near the type locality, and they provide more precise locality data. Moreover, this apparently rare species is also known from the Dawei Mountain Nature Reserve, Yunnan, China., Published as part of Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, pp. 325-352 in Beitr��ge Zur Entomologie = Contributions to Entomology 67 (2) on page 341, DOI: 10.21248/contrib.entomol.67.2.325-352, http://zenodo.org/record/5742448, {"references":["MIYAKE, Y. 2000: New or little known scarabaeid beetles from Southeast Asia IV. - Science Report of the Research Institute of Evolutionary Biology 9: 105 - 120."]}

Anomala cyanipennis LIN, 1999 Distribution: China, Laos (new country record), Vietnam (new country record). Specimens examined: 1 ���, 1 ���, N. VIET NAM (Tonkin) pr. Hoang Lien Son SA PA 11.-15.V.1990 P. Pachol��tko leg. (CZPC, PPPC); 6 ������, 10 ������, NE-LAOS Hua Phan prov.: Ban Saleui, Phou Pan (Mt.) ~ 20��12'N 104��01'E; 1300-1900 m 01.-31.V.2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 (CZPC, ZFMK). Remarks: This species was previously only known from the type locality (Xishuangbanna, China). Our data confirm its occurrence in northwestern Vietnam, as well as in northern Laos., Published as part of Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, pp. 325-352 in Beitr��ge Zur Entomologie = Contributions to Entomology 67 (2) on page 340, DOI: 10.21248/contrib.entomol.67.2.325-352, http://zenodo.org/record/5742448, {"references":["LIN, P. 1999: New Species of the Genus Anomala SAMOUELLE (Coleoptera: Rutelidae) Collected in Yunnan, China. - Entomotaxonomia 21: 157 - 176."]}

Anomala jeanvoinei BENDERITTER, 1929 Distribution: Vietnam. Specimens examined: Holotype: ���, Chapa, Tonkin (ex. Jeanvoine) Coll. Clermont | Anomala Jeanvoinei Bend [Benderitter���s handwriting] E. Benderitter det. | Anomala jeanvoinei Cotype | MUS��UM PARIS 1944 Coll. E. BENDERITTER (MNHN). Additional material: 3 ������, VIETNAM, Lao Cai Prov., Hoang Lien NP, Tram Ton, 1915 m, 22.3493723��N, 103.7704565��E, forest edge, at light, 8-11.IV.2010, leg. L. Papp, L. Peregovits & Z. Solt��sz VN 2010PL_4 (CZPC, HNHM). Remarks: This species has not been reported since its description almost one hundred years ago. It was rediscovered in unidentified material within the HNHM, collected in 2010, not far from the type locality., Published as part of Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, pp. 325-352 in Beitr��ge Zur Entomologie = Contributions to Entomology 67 (2) on page 343, DOI: 10.21248/contrib.entomol.67.2.325-352, http://zenodo.org/record/5742448, {"references":["BENDERITTER, E. 1929: Contribution a l'etude des Rutelides du Tonkin. - Annales de la Societe Entomologique de France 98: 101 - 109."]}

Anomala bedeli OHAUS, 1914 Distribution: China, Laos, Thailand (new country record), Vietnam. Specimens examined: 1 ���, Vietnam - N (Na Hang) 160 km NNW Ha Noi, NE env. Na Hang, 1- 14.06.1996, h= 150-200 m, Lg. A. Napolov & I. Roma (ANPC); 3 ������, 1 ���, Vietnam-N, Lao Cai Prov., Hoang Lien Son Nat. Res., Sa Pa vill. env., 27.5.- 2.6.1998, Lg. A. Napolov | 22�� 20'N 103�� 50'E h= 1250 m (ANPC); 1 ���, 1 ���, N. Vietnam SA PA env. LAO CAI prov. 22��19'52"N 103��50'35"E 23.-27.v.1999 1630-1680 m leg. Fabrici, Jaeger, Ahrens (CZPC); 1 ���, LAOS 21��09'N 101��19'E Louangnamtha pr. Namtha-MuangSing 5-31.v.1997. 900 Vit Kuban leg. 1200 m (PPPC); 1 ���, LAOS Naka 1.V.1998 leg. Strba (CZPC); 6 ������, NE-LAOS Hua Phan prov.: Ban Saleui, Phou Pan (Mt.) ~ 20��12'N 104��01'E; 1300-1900 m 01.-31.V.2011; leg. C. Holzschuh Ankauf ZFMK Bonn 2011 (CZPC, ZFMK); 1 ���, LAOS-NE; HUA PHAN prov. BAN SALUEI; Phu Phan Mt.: 20��15'N 104��02'E; 1500-2000 m; 26.iv.-11.v.2001 J. Bezděk & D. Hauck leg. (CZPC); 1 ���, LAOS-N; Xieng Khuang prov. Nong Haet 19��30'N 104��03'E; 29.-31.v.2001 J. Bezděk leg. (CZPC); 1 ���, NW THAILAND, 1.-23.5. Mae Hong Son, 1991 Ban Huai Po, 1600-2000 m J. Horak leg. (CZPC); 2 ������, 1 ���, THAI, Loei prov. Phu Kradung N.P., 1000 m 16��52'N 101��49'E 16-1.v.1999, D. Hauck leg. (CZPC); 1 ���, THAI, 26.-28.v.1991 Palong 750 m 19��55'N 99��06'E Vit Kub��n leg. | Thailand ���91 Thanon Thong Chai D. Kr��l & V. Kub��n (CZPC); 8 ������, 4 ������, THAILAND: Nan Province Amphur Bo Kluea; Tumbon Phufa; Doi Phuka Park ranger station 6 21. April 2003; L-473 | 19��01'N 101��11'E 538 m UV pan trap colls: Vitheepradit, Prommi & Setaphan (CZPC, PLPC, UMO); 1 ���, Guizhou, Ceheng, Weinan 800-950 m, 1979.V.23-27 leg. Du Shaokun (GEI). Remarks: Anomala bedeli was incorrectly synonymized with A. holcoptera FAIRMAIRE, 1889 by PAULIAN (1959) (ZORN, 2005) and subsequently reported under this name from the southern Chinese provinces Guizhou and Yunnan in LIN (1992). The record ���Anhui��� in ZORN (2006) and ZORN & BEZDĚK (2016) is incorrect because it was based on incorrectly labeled material. Additionally, the species occurs in northern Thailand, Laos and Vietnam., Published as part of Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, pp. 325-352 in Beitr��ge Zur Entomologie = Contributions to Entomology 67 (2) on page 339, DOI: 10.21248/contrib.entomol.67.2.325-352, http://zenodo.org/record/5742448, {"references":["OHAUS, F. 1914: XV. Beitrag zur Kenntnis der Ruteliden. - Entomologische Zeitung 75: 193 - 217 - https: // www. biodiversitylibrary. org / page / 8825521 page / 205 / mode / 1 up [accessed 2017 / 10 / 12].","PAULIAN, R. 1959: Coleopteres Scarabeides de l'Indochine (Rutelines et Cetonines) (suite). - Annales de la Societe Entomologique de France 128: 35 - 136 - http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6142156 w / f 14 [accessed 2017 / 06 / 26].","ZORN, C. 2005: Taxonomical acts in the Anomalini initiated during the preparation of the \" Catalogue of Palaearctic Coleoptera \". - Acta Societatis Zoologicae Bohemicae 68: 310 - 328.","LIN, P. 1992: Coleoptera: Rutelidae: 503 - 524. - In: CHEN, S. (ed.): Insects of the Hengduan Mountains region. Volume 1., Beijing: Science Press, i-xii, 865 pp. (in Chinese and English).","ZORN, C. 2006: Scarabaeidae: Rutelinae: Anomalini. - In: LOBL, I. & SMETANA, A. (eds): Catalogue of Palaearctic Coleoptera. Vol. 3. - Apollo Books, Stenstrup, pp. 251 - 276."]}

Anomala iwasei MIYAKE, 1994 Distribution: China, Laos, Vietnam (new country record). Specimens examined: Holotype: ���, Samneua Laos 20.VI.91 | Holotype: Anomala iwasei Y. MIYAKE, 1994 (RIEB). Additional material: 1 ���, 3 ������, N. VIETNAM ��� Lao Cai prov., Van Ban dist. Van Ban Nature Reserve (at light) (~ 1000 m) 23.-26.V.2011, L. Bartolozzi, S. Bambi, F. Fabiano, E. Orbach leg. (MZUF); 1 ���, LAOS north, 5-11.V.1997 20 km NW Louang Namtha N21��09.2.E 101��18.7 alt. 900+- 100 m. M. Strba & R. Hergovits leg. (VMPC). Remarks: Anomala iwasei was originally described from Laos (Sam Neua) and occurs in Yunnan, China (LIN 1996a). Herein, we provide the first record of this species from Vietnam, where it interestingly occurs sympatrically with its very close relative A. sapa MIYAKE, 1994 in L��o Cai Province., Published as part of Zorn, Carsten, Kobayashi, Hirokazu & Wada, Kaoru, 2017, Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records, pp. 325-352 in Beitr��ge Zur Entomologie = Contributions to Entomology 67 (2) on page 343, DOI: 10.21248/contrib.entomol.67.2.325-352, http://zenodo.org/record/5742448, {"references":["MIYAKE, Y. 1994: New or little known Scarabaeid Beetles from Southeast-Asia II. - Special Bulletin Essa entomological Society 2: 139 - 156.","LIN, P. 1996 a: New species of Anomala hirsutula species group from China and discussion on their taxonomic problems (Coleoptera: Rutelidae). - Entomotaxonomia 18: 157 - 169."]}