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We unfold the bifurcation involving the loss of evolutionary stability of an equilibrium of the canonical equation of Adaptive Dynamics (AD). The equation deterministically describes the expected long-term evolution of inheritable traits - phenotypes or strategies - of coevolving populations, in the limit of rare and small mutations. In the vicinity of a stable equilibrium of the AD canonical equation, a mutant type can invade and coexist with the present - resident - types, whereas the fittest always win far from equilibrium. After coexistence, residents and mutants effectively diversify, according to the enlarged canonical equation, only if natural selection favors outer rather than intermediate traits - the equilibrium being evolutionarily unstable, rather than stable. Though the conditions for evolutionary branching - the joint effect of resident-mutant coexistence and evolutionary instability - have been known for long, the unfolding of the bifurcation has remained a missing tile of AD, the reason being related to the nonsmoothness of the mutant invasion fitness after branching. In this paper, we develop a methodology that allows the approximation of the invasion fitness after branching in terms of the expansion of the (smooth) fitness before branching. We then derive a canonical model for the branching bifurcation and perform its unfolding around the loss of evolutionary stability. We cast our analysis in the simplest (but classical) setting of asexual, unstructured populations living in an isolated, homogeneous, and constant abiotic environment; individual traits are one-dimensional; intra- as well as inter-specific ecological interactions are described in the vicinity of a stationary regime. [ABSTRACT FROM AUTHOR]

Abstract: We analyze a codimension-two bifurcation in generic -dimensional Filippov (autonomous, piecewise smooth, discontinuous) systems, when a hyperbolic equilibrium collides with the discontinuity boundary. At generic (codim 1) boundary equilibria (BE), two different scenarios can occur: the ‘persistence’ of the stationary solution, that changes from a standard equilibrium on one side of the discontinuity boundary into a pseudo-equilibrium, and the ‘nonsmooth-fold’ between the standard equilibrium and a coexisting pseudo-equilibrium. Generalized (codim 2) boundary equilibria (GBE) separate persistence and nonsmooth-fold scenarios and are here shown to occur together with a fold bifurcation between pseudo-equilibria (PLP). In a two-parameter plane, the PLP bifurcation curve emanates from the GBE point tangentially to the BE curve. The asymptotic of the PLP curve locally to the GBE point is also derived and tested on a 4-dimensional ecological example. [Copyright &y& Elsevier]

We contribute to the analysis of codimension-two bifurcations in discontinuous systems by studying all equilibrium bifurcations of 2D Filippov systems that involve a sliding limit cycle. There are only two such local bifurcations: (1) a degenerate boundary focus, which we call the homoclinic boundary focus (HBF), and (2) the boundary Hopf (BH).We prove that-besides local bifurcations of equilibria and pseudoequilibria-the universal unfolding of the HBF singularity includes a codimension-one global bifurcation at which a sliding homoclinic orbit to a pseudosaddle exists, while that of the BH singularity has a codimension-one bifurcation curve along which a cycle grazing occurs. We define two canonical forms, one for each singularity, to which a generic 2D Filippov system can be locally reduced by smooth changes of variables and parameters and time reparametrization. Explicit genericity conditions are also provided, as well as the asymptotics of the bifurcation curves in the two-parameter space. We show that both studied codimension-two bifurcations occur in a known 2D Filippov system modeling an ecosystem subject to on-off harvesting control, and we provide two Mathematica scripts that automatize all computations. [ABSTRACT FROM AUTHOR]

Through the analysis of a mathematical model we construct in this paper the full catalog of possible love stories among standard (i.e. secure and non-synergic) individuals. For this, we first distinguish between robust and fragile couples, i.e. couples which have one or two stable equilibria. We also distinguish between high and low quality relationships by looking at the sign of the feelings among the partners. Finally, we split fragile couples into two distinct classes, namely those that start from the state of indifference to evolve toward their most desirable equilibrium, as opposed to those that have an unfavorable evolution. The boundaries separating the various classes in any two-dimensional parameter space are either bifurcation curves or special curves characterized by the annihilation of a suitably defined indicator associated to the equilibria or to their basins of attraction. The analysis is performed in part analytically and in part numerically (through continuation methods), and is mainly focused on the role of the appeals of the individuals. The results are in agreement with common wisdom and observations and show, in particular, that love stories can evolve positively even if one of the two individuals has negative feelings in the initial phase of the relationship. [ABSTRACT FROM AUTHOR]

We analyze three codimension-two bifurcations occurring in nonsmooth systems, when a nonhyperbolic cycle (fold, flip, and Neimark-Sacker cases, in both continuous and discrete time) interacts with one of the discontinuity boundaries characterizing the system's dynamics. Rather than aiming at a complete unfolding of the three cases, which would require specific assumptions on both the class of nonsmooth system and the geometry of the involved boundary, we concentrate on the geometric features that are common to all scenarios. We show that, at a generic intersection between the smooth and discontinuity induced bifurcation curves, a third curve generically emanates tangentially to the former. This is the discontinuity induced bifurcation curve of the secondary invariant set (the other cycle, the double-period cycle, or the torus, respectively) involved in the smooth bifurcation. The result can be explained intuitively, but its validity is proved here rigorously under very general conditions. Three examples from different fields of science and engineering are also reported. [ABSTRACT FROM AUTHOR]

The paper describes the slow evolution of two adaptive traits that regulate the interactions between two mutualistic populations (e.g. a flowering plant and its insect pollinator). For frozen values of the traits, the two populations can either coexist or go extinct. The values of the traits for which populations extinction is guaranteed are therefore of no interest from an evolutionary point of view. In other words, the evolutionary dynamics must be studied only in a viable subset of trait space, which is bounded due to the physiological cost of extreme trait values. Thus, evolutionary dynamics experience so-called border collision bifurcations, when a system invariant in trait space hits the border of the viable subset. The unfolding of standard and border collision bifurcations with respect to two parameters of biological interest is presented. The algebraic and boundary-value problems characterizing the border collision bifurcations are described together with some details concerning their computation. [ABSTRACT FROM AUTHOR]

We show in this paper how numerical bifurcation analysis can be used to study the evolution of genetically transmitted phenotypic traits. For this, we consider the standard Rosenzweig-MacArthur prey-predator model [The American Naturalist, 97 (1963), pp. 209-223] and, following the so-called adaptive dynamics approach, we derive from it a second-order evolutionary model composed of two ODEs, one for the prey trait and one for the predator trait. Then, we perform a detailed bifurcation analysis of the evolutionary model with respect to various environmental and demographic parameters. Surprisingly, the evolutionary dynamics turn out to be much richer than the population dynamics. Up to three evolutionary attractors can be present, and the bifurcation diagrams contain numerous global bifurcations and codimension-2 bifurcation points. Interesting biological properties can be extracted from these bifurcation diagrams. In particular, one can conclude that evolution of the traits can be cyclic and easily promote prey species diversity. [ABSTRACT FROM AUTHOR]