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Ivmaka longicaudatus Xu & Dietrich, sp. nov. (Figs. 1–14) Type species: Holotype. J (UCD), Papua New Guinea Gulf: Ivimka Res. Station Lakekamu Basin, 120m 7’44’S 146’30’E, 7–8, March, 2000, T. Sears, MT. Paratypes. 2J (INHS), same data as holotype. Description. Body length: male 3.1–3.3 mm. General color of body orange brown (Figs. 1–3). Vertex orange yellow; coronal suture brown, with pair of irregular dark brown spots on each side (Figs. 1–3). Eyes silvery gray to black (Figs. 1–3). Ocelli surrounded by yellowish creamy patches (Fig. 3). Face orange to pale yellow from vertex to anteclypeus; lateral frontal suture brown (Fig. 3). Pronotum with median yellow transverse band bordered by thin dark brown lines, anterior margin white, posterior part tan (Fig. 1). Mesonotum with basolateral triangles black, otherwise dark brown (Fig. 1). Forewing dark brown, subhyaline distally with veins brown and contrasting with pale membrane; hind wing transparent, veins brown (Figs. 1, 2). Legs orange except the pretarsus dark brown (Figs. 1–3). Male 2S abdominal apodemes parallel, moderately broad, reaching middle of segment V (Fig. 4). Male pygofer bearing 2–3 long and 10–11 small rigid setae on dorsal margin, group of 17–20 feeble microsetae near middle of lobe; distal process of pygofer lobe nearly straight and evenly tapered; dorsal bridge extended almost four-fifths length of lobe (Figs. 7, 8). Anal tube process slender, sinuate, apex sharply narrowed to point hooked anterad (Fig. 9). Style sinuate, dentifer bearing about 13 tiny teeth preceded by 2 fine setae and few sensory pits (Fig. 10). Connective 1amellate, with medial sclerotization; anterior margin straight or weakly concave; lateral margin distinctly constricted (Fig. 11). Subgenital plate in lateral view with apex round and bent dorsad along ventral margin of pygofer lobe; basal setae (7–8) uniseriate, distinctly longer than marginal setae, marginal setae (29–30) arranged in 1–2 rows occupying nearly half length of dorsal margin, macrosetae (21–22) arising near based of plate, terminated at marginal setae, feeble microsetae (54–60) irregularly arranged in 3–6 irregular rows (Figs. 7, 12). Aedeagus slender, V-shaped in lateral view, shaft with few tiny teeth dorsally, wide basally, then abruptly narrowed, apex bifid in posterior view giving rise to pair of short distally directed processes (Figs. 13, 14). Etymology. The specific epithet is a compound Latin adjective “ longicaudatus ”, which refers to the elongate male pygofer of the new species. Distribution. Papua New Guinea., Published as part of Xu, Ye, Zhang, Peng-Wen, Dietrich, Christopher H. & Qin, Dao-Zheng, 2023, A new genus of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae) from Papua New Guinea, pp. 175-179 in Zootaxa 5271 (1) on pages 176-177, DOI: 10.11646/zootaxa.5271.1.9, http://zenodo.org/record/7864498

Ivmaka longicaudatus Xu & Dietrich, sp. nov. (Figs. 1–14) Type species: Holotype. J (UCD), Papua New Guinea Gulf: Ivimka Res. Station Lakekamu Basin, 120m 7’44’S 146’30’E, 7–8, March, 2000, T. Sears, MT. Paratypes. 2J (INHS), same data as holotype. Description. Body length: male 3.1–3.3 mm. General color of body orange brown (Figs. 1–3). Vertex orange yellow; coronal suture brown, with pair of irregular dark brown spots on each side (Figs. 1–3). Eyes silvery gray to black (Figs. 1–3). Ocelli surrounded by yellowish creamy patches (Fig. 3). Face orange to pale yellow from vertex to anteclypeus; lateral frontal suture brown (Fig. 3). Pronotum with median yellow transverse band bordered by thin dark brown lines, anterior margin white, posterior part tan (Fig. 1). Mesonotum with basolateral triangles black, otherwise dark brown (Fig. 1). Forewing dark brown, subhyaline distally with veins brown and contrasting with pale membrane; hind wing transparent, veins brown (Figs. 1, 2). Legs orange except the pretarsus dark brown (Figs. 1–3). Male 2S abdominal apodemes parallel, moderately broad, reaching middle of segment V (Fig. 4). Male pygofer bearing 2–3 long and 10–11 small rigid setae on dorsal margin, group of 17–20 feeble microsetae near middle of lobe; distal process of pygofer lobe nearly straight and evenly tapered; dorsal bridge extended almost four-fifths length of lobe (Figs. 7, 8). Anal tube process slender, sinuate, apex sharply narrowed to point hooked anterad (Fig. 9). Style sinuate, dentifer bearing about 13 tiny teeth preceded by 2 fine setae and few sensory pits (Fig. 10). Connective 1amellate, with medial sclerotization; anterior margin straight or weakly concave; lateral margin distinctly constricted (Fig. 11). Subgenital plate in lateral view with apex round and bent dorsad along ventral margin of pygofer lobe; basal setae (7–8) uniseriate, distinctly longer than marginal setae, marginal setae (29–30) arranged in 1–2 rows occupying nearly half length of dorsal margin, macrosetae (21–22) arising near based of plate, terminated at marginal setae, feeble microsetae (54–60) irregularly arranged in 3–6 irregular rows (Figs. 7, 12). Aedeagus slender, V-shaped in lateral view, shaft with few tiny teeth dorsally, wide basally, then abruptly narrowed, apex bifid in posterior view giving rise to pair of short distally directed processes (Figs. 13, 14). Etymology. The specific epithet is a compound Latin adjective “ longicaudatus ”, which refers to the elongate male pygofer of the new species. Distribution. Papua New Guinea.

Ivmaka Xu & Dietrich, gen. nov. Type species: Ivmaka longicaudatus Xu & Dietrich sp. n., here designated. Description. Body robust. Head including eyes broader than pronotum in dorsal aspect, crown short and broad, round anteriorly, length less than half width between eyes, coronal suture long, restricted by vertex and postfrons, dorsal part of the frontoclypeus (above antennal pits) with median longitudinal suture continuous with coronal suture (Figs. 1, 3). Ocelli distinct, located on anterior margin of crown midway between eyes and coronal suture (Fig. 3). Face moderately broad; lateral frontal suture well delimited, curved mesad above antennal pit and meeting epistomal suture at midline ventromesad of ocelli; anteclypeus slightly convex, not expanded; rostrum distinctly overlapping base of hind coxae (Fig. 3). Pronotum large, almost twice as long as crown, with sinuate transverse depression (Fig. 1). Forewing narrow, round apically, apical cells occupying nearly one-third of total length, c and r cells nearly equal in width, both narrower than m and cua cells; vein R2 and RM arising from r cell, connected by cross-vein, vein MCu arising from m cell, veins RM and MCu evenly curved and almost parallel throughout their length (Fig. 5). Hind wing with CuA unbranched (Fig. 6). Front femur seta AM1 stout, situated near ventral margin; intercalary row with one large basal seta and five smaller setae more distad. Hind femur with macrosetal formula 2+1+1. Hind-tibia row AV with six preapical macrosetae (Figs. 1–3). Male 2S abdominal apodemes well developed (Fig. 4). Male pygofer elongate, posterior margin extended into sharp point, with conspicuous basolateral setal group and macrosetae, distal setal group scattered on each side of lobe, dorsal bridge well delimited, ventral appendage absent (Figs. 7, 8). Anal tube process well developed, extended to ventral margin of pygofer lobe (Fig. 9). Style broadened at base, sinuate in ventral and lateral views, apex tapered, dentifer adorned with tiny teeth and preapical setae (Fig. 10). Connective much longer than broad, caudal margin notched medially (Fig. 11). Subgenital plate extended beyond main part of pygofer lobe but not surpassing distal process, broadest near base and slightly trapered and upturned apically, basolateral lobe rounded; four categories of setae present: basal setae slim, arranged in row along basolateral lobe; marginal setae well defined but somewhat irregularly arranged, limited to distal part of plate margin; macrosetae uniseriate, reaching apex of plate; feeble microsetae numerous, scattered, longer than macrosetae (Figs. 7, 12). Aedeagus shaft tubular, preatrium short, dorsoatrium well developed, gonopore apical on ventral surface (Figs. 13, 14). Notes. This new genus will run to Mindanaoa Mahmood, 1967 and Smyga Dworakowska, 1995 in the generic key of Xu et al. (2021) but differs from Mindanaoa Mahmood and Smyga Dworakowska in presence of a well-developed dorsoatrium on the aedeagus (Fig. 13) (dorsoatrium absent in Mindanaoa Mahmood and Smyga Dworakowska). Among other previously described genera in Empoasca -group, this new genus is superficially similar to Dapitana Mahmood, 1967 and Serratulus Mahmood, 1967 in having the coronal suture well developed (Fig. 3), and vein R2 and RM connected by a cross-vein in the forewing (Fig. 5). However, this new genus differs from Dapitana Mahmood in having the pygofer with a conspicuous basolateral setal group (Fig. 7) and vein ScP+RA present on the forewing (Fig. 5) (absent in Dapitana Mahmood). It differs from Serratulus Mahmood, 1967 in having the subgenital plate with macrosetae uniseriate (Fig. 12) (vs. arranged in two rows). Furthermore, this new genus is unique in having the pygofer lobe extended into a distal spine-like process surpassing the subgenital plate (Figs. 7, 8). Distribution. Papua New Guinea. Etymology. The generic name alludes to the locality of the type material (Ivimka Res. Station). Gender: masculine., Published as part of Xu, Ye, Zhang, Peng-Wen, Dietrich, Christopher H. & Qin, Dao-Zheng, 2023, A new genus of Empoascini (Hemiptera, Cicadellidae, Typhlocybinae) from Papua New Guinea, pp. 175-179 in Zootaxa 5271 (1) on page 176, DOI: 10.11646/zootaxa.5271.1.9, http://zenodo.org/record/7864498, {"references":["Mahmood, S. H. (1967) A study of the typhlocybine genera of the Oriental region (Thailand, the Philippines and adjoining areas). Pacific Insects Monographs, 12, 1 - 52.","Dworakowska, I. (1995) Szara gen. nov. and some other Oriental Empoascini (Insecta: Auchenorrhyncha: Cicadellidae: Typhlocybinae). Entomologische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 56, 129 - 160.","Xu, Y., Dietrich, C. H., Zhang, Y. L. Dmitriev, D. A., Zhang, L., Wang, Y. M., Lu, S. H. & Qin, D. Z. (2021) Phylogeny of the tribe Empoascini (Hemiptera: Cicadellidae: Typhlocybinae) based on morphological characteristics, with reclassification of the Empoasca generic group. Systematic Entomology, 46, 266 - 286. https: // doi. org / 10.1111 / syen. 12461"]}