Search

Dundadela maculata Zahradník & Trýzna sp. nov. (Figs 1–16) Type material. Holotype (male): Madagascar, Isalo N.P., Oasïs, 879 m, S22°37′05″, E45°21′08″, 19.–20.i.2013, M. Trýzna leg. (FGMRI). Paratypes (7 males): 1 male: Madagascar, Isalo N.P., Piscine nat., 850 m, S22°33′, E45°23′, 21.i.2013, M. Trýzna leg; 1 male: Madagascar, Isalo N.P., Namaza camp, 776 m, S22°32′22.2′′, E45°22′46.0′′, 22.i.2013, M. Trýzna leg. [all specimens come from Fianarantsoa province] (FGMRI); 2 males: Madagascar, Tamatave prov. [= Toamasina province], Moramanga env., 21.–24.xii.1996, I. Jeniš leg. (FGMRI); 3 males: idem but 27.-30.xii.1996, I. Jeniš leg. (BMNH, FGMRI, MTDC). Description. Male (holotype). General appearance. Body long oval (Fig. 1), mostly parallel, body length 7.0 mm, the greatest width (on elytral base) 2.7 mm. Coloration. Body brown, antennae, palpi and legs light brown. Pubescence short, recumbent, white-silver, in nebulous pattern (Figs 1, 4, 7). Vestiture and structure. Head (Figs 2–3, 10) slightly transverse convex, matt shining (velvety), densely and finely punctuated, punctures almost contiguous. Pubescence very sparse, short, recumbent, inclined forward. Eyes large, globular, glabrous. Frons 2.3 times wider than width of eye in dorsal view. The last segment of maxillary palpi spindle shaped, 1.3 times longer than wide. Antennae (Figs 1, 8, 12, 14) with eleven antennomeres, last three antennomeres strongly enlarged, penultimate two strongly pectinate, others filiform. Antennomere I robust, 1.5 times as long as wide. Antennomere II 0.25 of length of I, width 0.7 of length. III twice as long as wide and 1.4 times as long as previous. IV the same width as III, 0.55 of length of III. V twice as long as wide, 1.1 times as long as previous. Antennomeres VI and VII of the same length and width, half as long as V. VIII transverse, twice as wide as long, one-fifth of length of VII. IX six times as long as wide, X seven times as long as wide, their length longer than IV to VIII combined. Ramus on IX 2.2 times as long as length of antennomere IX, ramus on X 1.8 times as long as length of antennomere X. Rami of antennomeres IX and X widened towards the apex. Antennomere XI filiform, long elliptical, the same width as I, 4.6 times as long as wide. Pronotum (Figs 1, 4, 7) transverse, without hump, ratio of its length to maximum width 0.8, widest in the middle, strongly oblongly narrowed backwards. Surface matt-shining, finely and densely punctuated, punctures almost contiguous. Pubescence short, recumbent, white-silver, nebulous pattern. Elytra (Fig. 1) parallel, without distinct humeral angles, with four very fine and indistinct longitudinal carinae (not visible on Fig. 1), equally distributed on elytra. Ratio of maximum length to maximum width of elytra 1.7. Surface and pubescence the same as on pronotum (Fig. 1). Scutellum small, triangular, 1.2 times as long as wide. Legs (Fig. 1) slim, femora 0.8 of length of the tibia. Tibia triangular in cross-section, with two sharp longitudinal edges. Tarsi two-thirds of length of tibiae. Tarsomere I as long as II and III together, II 1.2 times as long as III, III slightly transverse, 1.1 times as wide as previous. All these three tarsomeres widened at apex. Tarsomere IV shallowly emarginate from above, the same length as III, 1.2 times as wide as previous. The last tarsomere, V, robust, as long as II, the same width as IV at apex, widest at the apex, with two claws, widely spaced apart. Claws on interior part without teeth. Abdomen (Fig. 11) consists of five ventrites, ventrite I in the middle with short blunt wide promontory. Terminalia (Figs 13, 15–16). Aedeagus symmetrical (Figs 13, 16). Median lobe wide, parameres narrow, as long as median lobe; on apex with two more or less sharp small teeth inclined to median lobe. Each paramere with long, slim promontory. Lateral corner of tegmen [= Genitalbügel (Gottwald 1977), = genital stirrup (e. g. Zahradník 2006, 2013)] without promontory, rounded and broadened (Fig. 15). Female. Unknown. Variability. Differs in size of body—total length varies from 5.6 to 7.0 mm, width at base of elytra from 2.2 to 2.7 mm. Colour of body brown, to reddish-brown (possibly on teneral specimen). Differential diagnosis. The species has forward oriented head (probably semi-prognathous in natural position), antennae with 11 antennomeres, and only antennomeres IX and X strongly pectinate (see Differential diagnosis of the genus above). Etymology. From the nebulous (cloud like) arrangement of pubescence on the body, forming small rounded spots. Collecting circumstances. Three specimens of the type series (including holotype) were collected in Parc National de l´Isalo near Ranohira in Fianarantsoa province, S Madagascar (Figs 17–18). This area is dominated by a sub-arid climate. The territory consists of sandstones, shale and conglomerate (Fig. 17). The low, sparse forests are located mainly in the deeply carved canyons of the sandstone massif, and accompanied by sclerophyllous woodland with secondary grasslands and pastures in the vicinity (see also Goodman et al. 2018). These specimens from Isalo were caught at night at light near dry, dead trees (Fig. 18). Another five specimens of the type series, however, come from the vicinity of Moramanga in Toamasina province, C Madagascar. Climatic conditions are completely different here. A humid climate prevails with a much larger amount of precipitation compared to Isalo. There are remnants of degraded medium altitude moist evergreen forest. However, we do not know the collecting details of the specimens from Moramanga. Distribution. S and C Madagascar: Fianarantsoa (holotype) and Toamasina provinces (Fig. 19)., Published as part of Zahradník, Petr & Trýzna, Miloš, 2023, Description of a new genus and species, Dundadela maculata, from Madagascar (Coleoptera: Ptinidae), pp. 276-286 in Zootaxa 5244 (3) on pages 278-283, DOI: 10.11646/zootaxa.5244.3.5, http://zenodo.org/record/7656391, {"references":["Gottwald, J. (1977) Die palaarktischen Xyletinus - Arten (Coleoptera, Anobiidae). Acta entomologica Bohemoslovaca, 74 (3), 158 - 177.","Zahradnik, P. (2006) New species of the genus Metholcus Jacquelin du Val, 1860 (Coleoptera: Bostrichoidea: Anobiidae) from South Africa. Studies and reports of District Museum Prague-East, Taxonomical Series, 2 (1 - 2), 137 - 141.","Zahradnik, P. (2013) Brouci celedi cervotocoviti (Ptinidae) stredni Evropy - Beetles of the family Ptinidae of Central Europe. Academia, Praha, 351 pp. [in Czech and English]","Goodman, S. M., Raherilalao, M. J. & Wohlauser, S. (Eds.) (2018) s. n. In: Les aires protegees terrestres de Madagascar: Leur histoire, description et biote. Tome III. [The terrestrial protected areas of Madagascar: Their history, description, and biota. Vol. III]. Association Vahatra, Antananarivo, pp. 1233 - 1716."]}

A new genus and species, Dundadela maculata gen. nov. & sp. nov. (Ptinidae: Anobiinae: tribe incertae sedis) from Madagascar is described, male genitalia are illustrated. Comparison with similar genera is given. Ecological notes, including photographs of habitats of the new species, are provided.

Key to genera of Madagascan Anobiinae 1 Antennae with 10 antennomeres......................................................................... 2 - Antennae with 11 antennomeres......................................................................... 3 2 Elytra with distinct striae............................................... Hemigastrallus Español & Comas, 1991 - Elytra without distinct striae, only on lateral margin with 1–2 small striae and on the disc of elytra with traces of shallow striae formed by small sparse punctures ……………….................................. Gastrallus Jacquelin du Val, 1860 3 Antennae serrate................................................................. Mimotrypopitys Pic, 1931 - Only antennomeres IX and X strongly pectinate............................................ Dundadela gen. nov., Published as part of Zahradník, Petr & Trýzna, Miloš, 2023, Description of a new genus and species, Dundadela maculata, from Madagascar (Coleoptera: Ptinidae), pp. 276-286 in Zootaxa 5244 (3) on page 278, DOI: 10.11646/zootaxa.5244.3.5, http://zenodo.org/record/7656391, {"references":["Pic, M. (1931) Nouveautes diverses. Melanges Exotico-entomologiques, 57, 1 - 36."]}

Dundadela Zahradník & Trýzna gen. nov. (Figs 1–16) Type species. Dundadela maculata sp. nov., by original designation. Placement. Subfamily Anobiinae, tribe incertae sedis. Name derivation. Dundadela, gender feminine, dedicated to our colleague and friend Mr. Radek Dunda (*30 June, 1969–†30 March, 2020), specialist in the family Attelabidae (Coleoptera: Curculionoidea) (Figs 20–21). Description. Male. Body long oval, mostly parallel (Fig. 1). Head (Figs 2–3, 7, 10) forward oriented (probably semi-prognathous in natural position), slightly transverse convex, eyes large, globular, glabrous.Antennae (Figs 1, 8, 12, 14) with eleven antennomeres. Flagellum filiform, antennomere VIII strongly transverse, last three antennomeres strongly enlarged, IX–X strongly pectinate, XI simple and robust. Pronotum (Fig. 4) transverse, slightly concave dorsally, without central bump; pronotal lateral margin only partly visible, on posterior part invisible (in dorsal view). Elytra (Fig. 1) parallel, without distinct humeral angles, with four very fine and indistinct longitudinal carinae (not visible on Fig. 1), equally distributed on elytra. Legs (Fig. 1) short and slim. Anterior coxae (Fig. 10) very close, touching, mesocoxae (Fig. 9) very similar. Metacoxae (Fig. 9) slightly separated. Metathoracic ventrite (Fig. 9) with short longitudinal depression, posteriorly sharpened. All coxae conical. Claws (Fig. 6) widely spaced, not bifurcate at apex. Mesosternum (Fig. 9) posteriorly without hook, metasternum without tarsal grooves. Abdomen (Fig. 11) with five ventrites, I with short blunt wide promontory in middle. Aedeagus symmetrical (Figs 13, 16). Differential diagnosis. Dundadela gen. nov. differs from other genera of subfamily Anobiinae by antennal structure—no other genus of this subfamily has antennomeres IX and X pectinate. Only genus Megorama Fall, 1905 (subfamily Xyletininae) has similar structure of antennae, but it has a distinctly hypognathous (orthognathous) head. Megorama (unlike Dundadela) lacks elytral carinae. Distribution. So far known only from Madagascar., Published as part of Zahradník, Petr & Trýzna, Miloš, 2023, Description of a new genus and species, Dundadela maculata, from Madagascar (Coleoptera: Ptinidae), pp. 276-286 in Zootaxa 5244 (3) on pages 277-278, DOI: 10.11646/zootaxa.5244.3.5, http://zenodo.org/record/7656391

Six new species of spider beetle are described from southwestern Peru: Cordielytrum silvae, sp. n., C. inca, sp. n., C. fimbriatum, sp. n., C. obscurum, sp. n., C. adustum, sp. n., and C. globuloides, sp. n. The potential for undescribed diversity in this genus is discussed, as well as the geographic distribution and biology of species.

7.2. Ptinidae Symmetrischer Bau: Abb. 7: (7a) Gynopterus aubei ( BOIELDIEU, 1854); (7b) Dignomus frivaldskyi (REITTER, 1884); (7c) Hanumanus ludovici (PIC, 1919); (7d) Ptinus brevivittis (REITTER, 1881); (7e) Ptinus disconotatus PIC, 1907; (7f) Kedirinus bisbinotatus (PIC, 1910). Asymmetrischer Bau: Abb. 8: (8a) Ptinus bicinctus STURM, 1837; (8b) Ptinus japonicus REITTER, 1877; (8c) Ptinus montanus ESCALERA, 1914; (8d) Ptinus obesus LUCAS, 1849; (8e) Ptinus pilosus MÜLLER, 1821; (8f) Ptinus subpilosus STURM, 1837. Es ist mir ein starkes Anliegen, hier auf die ausgezeichnete Arbeit zur Gattung Sphaericus (ERBER & HINTERSEHER 2000) hinzuweisen, deren Genitalabbildungen vorbildlich die Determination dieser schwierigen Gattung erleichtern., Published as part of Eifler, Michael, 2022, Contribution to the preparation and determination of families Anobiidae und Ptinidae (Coleoptera, Teredilia), pp. 63-84 in Linzer biologische Beiträge 54 (1) on pages 82-83, DOI: 10.5281/zenodo.7507174

7.2. Ptinidae Symmetrical construction: Abb. 7: (7a) Gynopterus aubei ( BOIELDIEU, 1854); (7b) Dignomus frivaldskyi (REITTER, 1884); (7c) Hanumanus ludovici (PIC, 1919); (7d) Ptinus brevivittis (REITTER, 1881); (7e) Ptinus disconotatus PIC, 1907; (7f) Kedirinus bisbinotatus (PIC, 1910). Asymmetrical construction: Abb. 8: (8a) Ptinus bicinctus STURM, 1837; (8b) Ptinus japonicus REITTER, 1877; (8c) Ptinus montanus ESCALERA, 1914; (8d) Ptinus obesus LUCAS, 1849; (8e) Ptinus pilosus MÜLLER, 1821; (8f) Ptinus subpilosus STURM, 1837. It is very important to me to refer to the excellent work on the genus Sphaericus (ERBER & HINTERSEHER 2000), whose genital illustrations facilitate the determination of this difficult genus in an exemplary manner.

Eifler, Michael (2022): Contribution to the preparation and determination of families Anobiidae und Ptinidae (Coleoptera, Teredilia). Linzer biologische Beiträge 54 (1): 63-84, DOI: http://doi.org/10.5281/zenodo.7507174, {"references":["ESPANOL F. (1992): Coleoptera Anobiidae. En: Fauna Iberica, vol. 2., RAMOS M.A. et al. (eds). - Museo Nacional de Ciencias N aturales. CSIC. Madrid, 195 pp.","ERBER D. & W. HINTERSEHER (2000): Revision der Gattung Sphaericus (Col., Ptinidae) der Lauri-Makaronesischen Region (Azoren, Madeira, Selvagens, Kanaren) einschliesslich Nordafrika und des europaischen Festlandes. - Coleoptera 4 (2000): 153-282 ISSN 0945-1889","FREUDE H., HARDE K. & G. LOHSE (1969): Die Kafer Mitteleuropas. - Goecke & Evers Verlag, Bd. 8, 28-74 pp., Krefeld 1969.","LOMPE A. (1989): Ein bewahrtes Einbettungsmittel fur Insektenpraparate. - In: L OHSE & LUCHT (eds): Die Kafer Mitteleuropas, 1. Supplementband 12: 17-18 pp. Goecke & Evers Verlag, Krefeld."]}

7.2. Ptinidae Symmetrischer Bau: Abb. 7: (7a) Gynopterus aubei ( BOIELDIEU, 1854); (7b) Dignomus frivaldskyi (REITTER, 1884); (7c) Hanumanus ludovici (PIC, 1919); (7d) Ptinus brevivittis (REITTER, 1881); (7e) Ptinus disconotatus PIC, 1907; (7f) Kedirinus bisbinotatus (PIC, 1910). Asymmetrischer Bau: Abb. 8: (8a) Ptinus bicinctus STURM, 1837; (8b) Ptinus japonicus REITTER, 1877; (8c) Ptinus montanus ESCALERA, 1914; (8d) Ptinus obesus LUCAS, 1849; (8e) Ptinus pilosus MÜLLER, 1821; (8f) Ptinus subpilosus STURM, 1837. Es ist mir ein starkes Anliegen, hier auf die ausgezeichnete Arbeit zur Gattung Sphaericus (ERBER & HINTERSEHER 2000) hinzuweisen, deren Genitalabbildungen vorbildlich die Determination dieser schwierigen Gattung erleichtern.

7.2. Ptinidae Symmetrical construction: Abb. 7: (7a) Gynopterus aubei ( BOIELDIEU, 1854); (7b) Dignomus frivaldskyi (REITTER, 1884); (7c) Hanumanus ludovici (PIC, 1919); (7d) Ptinus brevivittis (REITTER, 1881); (7e) Ptinus disconotatus PIC, 1907; (7f) Kedirinus bisbinotatus (PIC, 1910). Asymmetrical construction: Abb. 8: (8a) Ptinus bicinctus STURM, 1837; (8b) Ptinus japonicus REITTER, 1877; (8c) Ptinus montanus ESCALERA, 1914; (8d) Ptinus obesus LUCAS, 1849; (8e) Ptinus pilosus MÜLLER, 1821; (8f) Ptinus subpilosus STURM, 1837. It is very important to me to refer to the excellent work on the genus Sphaericus (ERBER & HINTERSEHER 2000), whose genital illustrations facilitate the determination of this difficult genus in an exemplary manner., Published as part of Eifler, Michael, 2022, Contribution to the preparation and determination of families Anobiidae und Ptinidae (Coleoptera, Teredilia), pp. 63-84 in Linzer biologische Beiträge 54 (1) on pages 71-72, DOI: 10.5281/zenodo.7507174

Cordielytrum globuloides, sp. n. (Fig. 8) Holotype: PERU, AR, Yarabamba, Linga Alta 16°34’23” / 71°33’45” 2759 m 29.III.2017 C. Bravo. Holotype has been deposited in MUSM. Paratypes (6): PERU, AR, Yarabamba, San Jose 2622 m 16°33’32” / 71°37’51” 18.III.2017 C. Bravo (1); PERU, AR, Yarabamba, San Juan 2438 m 16°34’00” / 71°38’27” 20.III.2017 C. Bravo (2); PERU, AR, Yarabamba, Linga Alta, 2760 m 16°33’32” / 71°37’51” 28.III.2017 C. Bravo (1); PERU, AR, Tiabaya, Quebradita 2500 m 16°28’53” / 71°36’02” 22.III.2017 C. Bravo (2). Paratypes have been deposited in MUSM, KAWC, and TKPC. Diagnosis: This species is distinguished from congeners by its yellowish tan and speckled dark brown elytral coloration, the absence of carinae on the elytra, and the distinct rectangular or conical shaped elytral scales. Description. Body small, ovate, convex; mottled yellowish tan. Length (anterior of pronotum to apex of elytra) 2.19–2.88 mm. Head completely covered with depressed, light tan, ovoid scales; slight superantennal carinae; eyes surrounded by very narrow bare patch lacking scales dorsally, ventrally with scales slightly protruding; antennae stout, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 longer than wide, others subequal; labrum with scattered short erect orange setae apically; frons with elongate ligulate scales pointing mainly obliquely toward middle, especially prominent toward apex. Pronotum covered with densely matted setae, relatively few short, erect setae; medial cavity very deep, extending over half total length of pronotum, border well-defined except at middle anterior and posterior; two posterolateral cavities on each side deep and distinct, lateralmost cavities completely anterior to adjacent cavities. Elytra: Setation yellowish tan, speckled with dark brown in random somewhat clumped pattern, background color reddish brown; surface mostly covered in conical or flattened rectangular scales, some scales instead acutely pointed; basal 1/3 of anterolateral margin with row of long bristly setae; elytral carinae absent; ~4 rows of short erect setae on each elytron. Ventral Surface: Ventral part of pro-, meso-, and, metathorax, legs, and abdominal ventrites with appressed oval scales. Etymology: The name “globuloides” is for the similar color pattern of this species to Trigonogenius globulus Solier, for which it was initially mistaken among museum material., Published as part of Whorrall, Kyle & Philips, Keith, 2021, Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips, pp. 414-427 in Zootaxa 5040 (3) on pages 423-425, DOI: 10.11646/zootaxa.5040.3.6, http://zenodo.org/record/5531508

Cordielytrum silvae, sp. n. (Fig. 3) Holotype: Peru: Ica Region, Lomas de San Fernando. 11–12.VI.2019, S15.13°, W75.31° dung pitfall traps, 1425 m K.A.Whorrall & T. K.Philips. Holotype has been deposited in MUSM. Paratypes (16): Same data as holotype (11); PE: ICA, Prov. de Nazca Lomas de San Fernando 15°08’51.7” S 75°15’27.8” W 489m, 03.iv.2010, D. Silva y M. Vilchez (1); Prov. de Nazca Lomas de San Fernando 15°07’29.0” S 75°20’19.8” W 430m, 03.iv.2010, D. Silva y M. Vilchez (4). Paratypes have been deposited in MUSM, KAWC, TKPC, and JBHC. Diagnosis. This species is distinguished from its congeners by its relatively large size, the smooth appearance of its setation, its small bead-like scales, and its pale, brassy coloration. It can be distinguished from C. peruvianum by the larger depressions in elytral scales, more transverse pronotum, and less pronounced elytral carinae. Description. Body small, compact, subovate, convex; brass colored with a slight sheen. Length (anterior of pronotum to apex of elytra) 2.93–3.25 mm. Head completely covered with depressed, light tan, ovoid scales; pronounced superantennal carinae; antennae slender, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 slightly longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with short ligulate scales pointing toward middle. Pronotum covered with densely matted setae, short, erect setae protruding from deep cavities within matted setation; medial cavity deep, broad, when viewed from above, extending nearly ½ total length, border well-defined laterally but indistinct medially, extending as a shallow groove anteriorly; two posterolateral cavities very shallow with indistinct borders. Elytra: Setation pale brassy tan, darker tan pattern medially, background color dark brown; densely covered in small, broad, ovate scales; basal ¼ of anterolateral margin with row of long bristly setae; four low longitudinal carinae extending length of each elytron, including one at suture; 4–5 rows of deep depressions between carinae. Ventral Surface: Prosternum and mesoventrite with matted scales somewhat similar to pronotum; remainder of ventral mesothorax, metathorax, legs, and abdominal ventrites with scales similar to elytra. Etymology: The name “silvae” is for Diana Silva, who collected the first specimens of this species., Published as part of Whorrall, Kyle & Philips, Keith, 2021, Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips, pp. 414-427 in Zootaxa 5040 (3) on pages 417-418, DOI: 10.11646/zootaxa.5040.3.6, http://zenodo.org/record/5531508

Cordielytrum obscurum, sp. n. (Fig. 6) Holotype: Peru: Arequipa Reg, 16 km NW Matarani, 18–21.VI.2019 S16.8754° W72.1968°, 285 m dune veg, dung pitfalls K.A.Whorrall & T. K. Philips. Holotype has been deposited in MUSM. Paratypes (24): Same data as holotype. Paratypes have been deposited in MUSM, KAWC, TKPC, and JBHC. Diagnosis: This species is distinguished from most congeners by its relatively slender shape, the more rounded shape of its pronotum, the rough, disheveled appearance of its vestiture, and the tassel-like shape of some setae. It can be distinguished from C. fimbriatum (Fig. 5) by the longer and more filiform setae of the elytral humeral angles, narrower and more sparsely spaced scales of the elytra, and darker coloration. Description. Body small, ovate, convex; mottled dark brown. Length (anterior of pronotum to apex of elytra) 2.16–2.35 mm. Head completely covered with depressed, light tan, ovoid scales; pronounced superantennal carinae; antennae stout, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with elongate ligulate scales pointing toward middle, especially prominent toward apex. Pronotum covered with densely matted setae, relatively few short, erect setae protruding from deep cavities within matted setation; anteriorly depressed, especially laterally, with distinct frontal ridge; medial cavity very deep, extending over half total length of pronotum, border well-defined except at middle anterior and posterior; two posterolateral cavities on each side deep and distinct. Elytra: Setation mottled tan with darker brown patches, background color dark brown; surface sparsely covered in slender scoop-shaped setae, gaps between setae showing bare elytra; basal 1/3 of anterolateral margin with row of long setae, setae mostly simple, but some variably tassel-like; five low longitudinal carinae extending length of each elytron, including one at suture, each carina with one row of short tassel-like setae with highly branched apices. Ventral Surface: Ventral part of pro-, meso, and, metathorax, legs, and abdominal ventrites with scales similar to elytra. Etymology: The name “obscurum” is from the Latin word for dark, referring to the coloration of this species. Secondarily, this name alludes to the difficulty of separating this species from others closely related., Published as part of Whorrall, Kyle & Philips, Keith, 2021, Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips, pp. 414-427 in Zootaxa 5040 (3) on pages 421-422, DOI: 10.11646/zootaxa.5040.3.6, http://zenodo.org/record/5531508

Cordielytrum silvae, sp. n. (Fig. 3) Holotype: Peru: Ica Region, Lomas de San Fernando. 11–12.VI.2019, S15.13°, W75.31° dung pitfall traps, 1425 m K.A.Whorrall & T. K.Philips. Holotype has been deposited in MUSM. Paratypes (16): Same data as holotype (11); PE: ICA, Prov. de Nazca Lomas de San Fernando 15°08’51.7” S 75°15’27.8” W 489m, 03.iv.2010, D. Silva y M. Vilchez (1); Prov. de Nazca Lomas de San Fernando 15°07’29.0” S 75°20’19.8” W 430m, 03.iv.2010, D. Silva y M. Vilchez (4). Paratypes have been deposited in MUSM, KAWC, TKPC, and JBHC. Diagnosis. This species is distinguished from its congeners by its relatively large size, the smooth appearance of its setation, its small bead-like scales, and its pale, brassy coloration. It can be distinguished from C. peruvianum by the larger depressions in elytral scales, more transverse pronotum, and less pronounced elytral carinae. Description. Body small, compact, subovate, convex; brass colored with a slight sheen. Length (anterior of pronotum to apex of elytra) 2.93–3.25 mm. Head completely covered with depressed, light tan, ovoid scales; pronounced superantennal carinae; antennae slender, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 slightly longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with short ligulate scales pointing toward middle. Pronotum covered with densely matted setae, short, erect setae protruding from deep cavities within matted setation; medial cavity deep, broad, when viewed from above, extending nearly ½ total length, border well-defined laterally but indistinct medially, extending as a shallow groove anteriorly; two posterolateral cavities very shallow with indistinct borders. Elytra: Setation pale brassy tan, darker tan pattern medially, background color dark brown; densely covered in small, broad, ovate scales; basal ¼ of anterolateral margin with row of long bristly setae; four low longitudinal carinae extending length of each elytron, including one at suture; 4–5 rows of deep depressions between carinae. Ventral Surface: Prosternum and mesoventrite with matted scales somewhat similar to pronotum; remainder of ventral mesothorax, metathorax, legs, and abdominal ventrites with scales similar to elytra. Etymology: The name “silvae” is for Diana Silva, who collected the first specimens of this species.

Cordielytrum adustum, sp. n. (Fig. 7) Holotype: Arequipa Region, nr. Pescadores, 17–22.VI.2019 S16.3876° W73.2588° 19m riparian veg, dung pitfalls K.A.Whorrall & T. K.Philips. Holotype has been deposited in MUSM. Paratype (1): Same data as holotype. Paratype will be deposited in TKPC. Diagnosis: This species is distinguished from most congeners by the rough, disheveled appearance of its vestiture, and its unique scoop-shaped scales. The only two specimens are of notably small size, though this may not be representative of the true size variation within the species. It can be distinguished from C. fimbriatum (Fig. 5) and C. obscurum (Fig. 6) by the short erect filiform setae covering its body, its more convex shape, the dark bands on the carinae of its elytra, and the relatively large depressions in its elytral setation. The species can be distinguished from C. inca (Fig. 4) by the more rounded pronotum, larger scale size relative to the body, and much shorter setae at the humeral angles of the elytra. Description. Body very small, ovate, convex; mottled reddish brown with a slight sheen. Length (anterior of pronotum to apex of elytra) 1.81 mm. Head completely covered with depressed, light tan, ovoid scales; pronounced superantennal carinae; antennae stout, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with elongate ligulate scales pointing toward middle, especially prominent toward apex. Pronotum covered with densely matted setae, relatively few short, erect setae protruding from deep cavities within matted setation; anteriorly depressed, especially laterally; medial cavity shallow broad, extending over half total length of pronotum, border well-defined except at middle anterior and posterior; two posterolateral cavities on each side deep and distinct. Elytra: Setation mottled tan and dark brown, background color reddish brown; surface covered in triangular scoop-shaped setae; basal 1/3 of anterolateral margin with row of relatively short bristly setae; five low longitudinal carinae extending length of each elytron, including one at suture, each carina with one row of very short truncate setae; ~3 rows of deep, open depressions between carinae. Ventral Surface: Ventral part of pro-, meso, and metathorax, legs, and abdominal ventrites with scales similar to elytra. Etymology: The name “adustum” is from the Latin word for singed, referring to the darkened appearance of the elytral pattern., Published as part of Whorrall, Kyle & Philips, Keith, 2021, Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips, pp. 414-427 in Zootaxa 5040 (3) on page 422, DOI: 10.11646/zootaxa.5040.3.6, http://zenodo.org/record/5531508

Cordielytrum fimbriatum, sp. n. (Fig. 5) Holotype: Peru: Arequipa Reg, 22 km NW Matarani, 18–21.VI.2019 S16.8548° W72.2306°, 304 m ravine veg, dung pitfalls K.A.Whorrall & T. K. Philips. Holotype has been deposited in MUSM. Paratypes (36): Same data as holotype. Paratypes have been deposited in MUSM, KAWC, TKPC, and JBHC. Diagnosis: This species is distinguished from most congeners by its relatively slender shape, the more rounded shape of its pronotum, the rough, disheveled appearance of its vestiture, and the tassel-like shape of its setae. It can be distinguished from C. obscurum (Fig. 6) by the shorter and more truncate setae of the elytral humeral angles, broader and more densely spaced scoop-like setae on the elytra, and paler coloration. Description. Body small, ovate, convex; mottled tan and brown. Length (anterior of pronotum to apex of elytra) 1.90–2.81 mm. Head completely covered with depressed, light tan, ovoid scales; pronounced superantennal carinae; antennae stout, antennomeres densely covered in scales, setae increasingly fine toward apex, antennomeres 1–4 and 11 longer than wide, others subequal; clypeus distinct; labrum with short erect orange setae apically; frons with elongate ligulate scales pointing toward middle, especially prominent toward apex. Pronotum covered with densely matted setae, relatively few short, erect setae protruding from deep cavities within matted setation; anteriorly depressed, especially laterally, with distinct frontal ridge; medial cavity very deep, extending over half total length of pronotum, border well-defined except at middle anterior and posterior; two posterolateral cavities on each side deep and distinct. Elytra: Setation mottled tan with darker brown patches, background color dark brown; surface covered in triangular scoop-shaped setae; basal 1/3 of anterolateral margin with row of relatively short tassel-like setae; five low longitudinal carinae extending length of each elytron, including one at suture, each carina with one row of short tassel-like setae with highly branched apices. Ventral Surface: Ventral part of pro-, meso, and, metathorax, legs, and abdominal ventrites with scales similar to elytra. Etymology: The name “fimbriatum” is from the Latin word meaning fringed, referring to the rows of tassel-like setae on the elytral carinae., Published as part of Whorrall, Kyle & Philips, Keith, 2021, Six new species of spider beetles (Coleoptera: Ptinidae) in the genus Cordielytrum Philips, pp. 414-427 in Zootaxa 5040 (3) on pages 419-420, DOI: 10.11646/zootaxa.5040.3.6, http://zenodo.org/record/5531508

Gearner, Olivia M., Philips, T. Keith (2021): Notaferrum n. gen. (Coleoptera: Ptinidae): the first known spider beetle associated with weaver ants. Zoosystema 43 (2): 29-36, DOI: 10.5252/zoosystema2021v43a2

Notaferrum natalensis (Pic, 1906) n. comb. (Figs 1-4) Ptinus elegans P��ringuey, 1904: 225. Ptinus natalensis Pic, 1906: 227. MATERIAL EXAMINED. ��� Holotype. South Africa ��� 1 ♀; [Natal] Isipingo beach; I.1899; SAM type Acc. No.1261. ADDITIONAL LABEL DATA. ��� ���Type��� (red label); ���natalensis���; ��� Lectotypus, Ptinus elegans, P��ringuey, des J. Borowski ���98 (SAMC)���. Abdominal ventrites on a separate card. OTHER MATERIAL. ��� South Africa ��� 1 ♂; Kwazulu-Natal, iSimangaliso Wetland Park [Previously known as Greater St. Lucia Wetland Park], Sodwana Bay campgrounds; 27.54��S, 32.67��E; c. 40 m; 21- 24.X.2014; R. Stals leg.; Vegetation type: FOz Northern Coastal Forest; SANC ��� 1 ♀; Natal, Charters Creek, St. Lucia; 28.12��S, 32.25��E; 14-16.I.1981; R. Oberprieler leg.; from nest of Oecophylla longinoda; AcP. 8497; SANC. DIAGNOSIS. ��� The generic diagnosis together with the white scale setal pattern on the elytra (Fig. 1) should distinguish this taxon, if other species within this genus are discovered in the future. SPECIES REDESCRIPTION Body Color dark brown to black on head, pronotum, and elytra. Body small and elongate. Length 3.4 �� 0.33 mm (n = 3). Head (Fig. 2) Head covered in punctures, large shallow irregular and contiguous mediolaterally, medially finer, longitudinally elongate; scattered, white small appressed scales; antennae dark brown, antennomeres 1-8 with moderately dense white appressed scales, first antennomere largest, second small as long as wide, remaining antennomeres longer than wide. Pronotum Pronotum mostly smooth with very fine alutaceous sculpture, glabrous except anterior �� with round and elongate scattered punctures; pair of medial vertical ridges approximately broadly rounded from anterior to posterior dorsal edge, then declivous to base just past middle, short thin yellow setae scattered on top of ridges; three tubercles on each lateral edge, anterior tubercle rounded, middle concave triangular, somewhat ear-shaped, posterior tubercle small, pointed, recumbent clumped orange trichome setae on lateral edge in between and around tubercles; a broadly rounded U-shaped smooth impression; posterior margin of pronotum narrow anterior of scutellum. Elytra Elytral surface covered in rows of round to elongate punctures separately longitudinally by half or less their length, puncture rows separated by about 1.5-2.0 times their diameters, 1 st and 2nd rows slightly obliquely oriented, 3rd and 4th less so; humeral angles raised; scutellum triangular, lateral edges below elytral surface; interpuncture rows of small recumbent dull pale orange setae; white scales forming an irregular transverse band at anterior 1/3, extending medially in an irregular longitudinal band from anterior transverse band to posterior 1/3, lateral clump of setae on each elytron at about posterior 1/3, transverse band of setae at apex and sometimes extending anteriorly along suture. Ventrites Ventral surface reddish to dark brown, covered in small white appressed scales mostly covering surface; proventrite smooth, mesoventrite with irregularly shaped, glabrous indentation at middle; lateral edge of fourth ventrite and apical edge of fifth ventrite with scattered fine yellowish setae. Legs Legs reddish to dark brown, covered in white scales as on ventral surface; 1 st tarsomere longer than wide, others short in length and width, internal surface of tarsomeres covered in fine gold recumbent setae. Sexual dimorphism Antennomeres 6-11 in the maledistinctly more elongate compared to those seen in the female. Male genitalia with elongate narrow parameres with a thick median lobe that is about the same length as the parameres. DISCUSSION There is little doubt that Notaferrum n. gen. shares close common ancestry especially with Silisoptinus Pic, 1917 (see Bell��s 1988, 2009) and more distantly with Eutaphrimorphus Pic, 1898 (see Bell��s 1992). All three genera are hypothesized to compose a single monophyletic lineage based on the possession of a very distinct and similarly shaped large medial smooth and flattened U-shaped shallow depression with a broadly rounded posterior border located medially at the base of the pronotum. Further similarity in these three genera is the anteriorly expanded prosternum that covers in part the ventral portion of the head, a feature that may have evolved to help protect mouthparts, most likely from ants. There are additional taxa from the Americas and southeast Asia (e.g. Prosternoptinus Bell��s, 1985, and Sundaptinus Bell��s, 1991) that also share this feature but based on other characteristics are not closely related (Mynhardt et al. unpublished). Another similarity between Notaferrum n. gen. and Silisoptinus is the greatly reduced fourth abdominal ventrite (c. �� the length of the third). Further is the strong declivity on the third ventrite that slopes dramatically down to the fourth. In contrast, Eutaphrimorphus has a more typical spider beetle abdomen with the fourth reduced (c. half the length of the third) and the ventrites are all in roughly the same plane. Notable differences of Notaferrum n. gen. with the other two genera is the lack of two short knobs near the pronotal base positioned laterally. They are very pronounced in Eutaphrimorphus while in Silisoptinus they are smaller and of various shapes depending upon the species. Eutaphrimorphus has two additional knobs positioned more centrally as well. These additional projections are also seen in at least one species of Silisoptinus but are much less pronounced and relatively gently rounded. Both Notaferrum n. gen. and Silisoptinus also exhibit depression(s) on the ventral surface that are lacking in Eutaphrimorphus; in Notaferrum n. gen., it is located only on the metaventrite near the middle and is similar to at least two other species of Silisoptinus (Bell��s 1988) including one undescribed (Philips unpublished). But the two described species of Silisoptinus also have a single depression on the abdomen while two undescribed (Philips unpublished) have either only a single depression on the metaventrite or both very weakly developed depressions on both the metaventrite and abdomen. Silisoptinus species are known from the eastern side of the African continent (Zanzibar and Socotra Islands) as well as Ghana (Philips unpublished) while Eutaphrimorphus is known only from South Africa. Also possibly related to Notaferrum n. gen. is Dignomus Wollaston, 1862, and the flightless Pseudomezium Pic, 1897, with the latter representing a flightless Dignomus (Philips unpublished and see Smiley & Philips 2011). These genera together with Silisoptinus and Eutaphrimorphus may form a monophyletic clade or at least with some close common ancestry. Dignomus as currently defined has a diverse set of pronotal modifications (Bell��s 1996; Smith & Philips unpublished) and may not form a monophyletic clade. But the overall similarity of the pronotum having an anterior and posterior lobe laterally and without much lateral projection in both Notaferrum n. gen. and Dignomus (plus Pseudomezium) is notable. One other characteristic feature of the new genus is the presence of a transverse posterior pronotal groove of various degrees and shapes. Many other spider beetle taxa also possess a basal groove of various shapes including the African genera Eutaphrimorphus Pic, 1898, Silisoptinus Pic, 1917, Eutaphroptinus Borowski, 2009, Dignomus Wollaston, 1862, Pseudomezium Pic, 1897, Dignomorphus Borowksi, 2009, as well as some species of the non-African Ptinus (e.g. P. espanyoli Bell��s, 1997), Tropicoptinus Bell��s, 1998, and members of Trymolophus Bell��s, 1990. DISTRIBUTION Notaferrum natalensis n. gen., n. comb. is known only from KwaZulu-Natal province in South Africa (Fig. 4). The current range is a little over 300 km in length, and all records are within 30 km of the coast.This distribution is within the Maputuland-Pondoland-Albany hotspot, one of three biodiversity hotspots in South Africa. Maputuland-Pondoland-Albany is notable in particular for its high plant endemism and is the second richest floristic region in South Africa (Myers et al. 2000). As the distribution of the weaver ant Oecophylla longinoda extends from Southern Africa through Eastern and Western Africa (Bolton 1995), one might expect the distribution of Notaferrum natalensis n. gen., n. comb. to be much larger. BIOLOGY Notaferrum natalensis n. gen., n. comb. is hypothesized to be a myrmecophile, as one individual was collected in a weaver ant nest (Oecophylla longinoda, Fig. 3B). There are no other spider beetles known to be associated with Oecophylla ants anywhere in the world. Only one other southern African genus appears to be a true symphile; Diplocotidus P��ringuey, 1899 includes three species currently known from South Africa and Namibia (Bell & Philips 2008). Unlike this new genus, species are wingless and found in arid or semiarid regions and are associated with ground nesting ants. The food type of Notaferrum natalensis n. gen., n. comb. is unclear, but it may feed on nest detritus, as most species of spider beetles feed on accumulated organic matter, particularly dung and detritus., Published as part of Gearner, Olivia M. & Philips, T. Keith, 2021, Notaferrum n. gen. (Coleoptera: Ptinidae): the first known spider beetle associated with weaver ants, pp. 29-36 in Zoosystema 43 (2) on pages 31-34, DOI: 10.5252/zoosystema2021v43a2, http://zenodo.org/record/4464304, {"references":["PIC M. 1906. - Sur divers Ptinides et Anobiides [Col.]. Bulletin de la Societe entomologique de France 11: 227 - 228.","PERINGUEY L. 1904. - Sixth contribution to the South African coleopterous fauna. Annals of the South African Museum 3: 167 - 300.","BELLES X. 1988. - El genero Silisoptinus Pic, 1917 (Coleoptera, Ptinidae). Miscellania Zoologica 12: 371 - 374.","BELLES X. 2009. - Spider beetles (Coleoptera, Ptinidae) from the Socotra Archipelago. Fauna of Arabia 24: 145 - 154.","BELLES X. 1992. - El genere Eutaphrimorphus Pic (Coleoptera: Ptinidae) de l'Africa austral. Butlleti de la Institucio Catalana d'Historia Natural 60: 129 - 132.","SMILEY M. E. & PHILIPS T. K. 2011. - Review of the South African spider beetle genus, Pseudomezium Pic, 1897 (Coleoptera: Ptinidae: Ptininae). African Entomology 19: 572 - 596.","BELLES X. 1996. - El genero Dignomus Wollaston (Coleoptera, Ptinidae). Bolleti de la Societat d'Historia Natural de Balears 39: 209 - 228.","BOROWSKI J. 2009 a. - The spider beetles of the continental Africa (Coleoptera: Ptinidae). Part I-Genus Mezium Curtis. Studies and Reports of the District Museum Prague-East Taxonomical Series 5: 1 - 10.","MYERS N., MITTERMEIER R., MITTERMEIER C., DA FONSECA G. A. B & KENT J. 2000. - Biodiversity hotspots for conservation priorities. Nature 403: 853 - 858. https: // doi. org / 10.1038 / 35002501","BOLTON B. 1995. - A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, Mass., 504 p.","BELL K. L. & PHILIPS T. K. 2008. - A revision of the South African myrmecophile Diplocotidus (Coleoptera: Ptinidae). African Entomology 16: 33 - 40. https: // doi. org / 10.4001 / 1021 - 3589 - 16.1.33"]}

Genus Notaferrum n. gen. urn:lsid:zoobank.org:act: A9D794F8-7079-4544-A08D-FCE368A035B5 TYPE SPECIES. ��� Ptinus natalensis Pic, 1906 by original designation. ETYMOLOGY. ��� Notaferrum n. gen. is derived from the Greek nota for back and the Latin ferrum for blade in reference to the longitudinal blade-like ridges on the pronotum. DIAGNOSIS. ��� This genus can be recognized by the pair of longitudinal blade-like ridges located medially on the pronotum as well as a transverse, flattened, and U-shape depression with sides diverging anteriorly at the basal 2/5 of the pronotum. GENERIC DESCRIPTION Body Small, length approximately 3.4 mm, elongate. Head Eyes convex, ovoid, slightly projecting, visible dorsally; vertex flat at middle; antennal fossae deep and distinct medially and ventrally, antennae long, filiform, 11 antennomeres; interantennal space between antennal insertions narrow, with a longitudinal groove; clypeus triangular, concave; labrum appearing truncate, maxillary and labial palpomeres fusiform. Pronotum Two thin longitudinal ridges positioned on either side of midline; two lateral tubercles on either side of the pronotum, flanking a transverse groove, basally forming a large posterior flat U-shaped depression with sides diverging anteriorly; anterior margin with deep irregular punctures. Elytra With longitudinal rows of punctures; interpuncture rows of small setae; scattered scale-like setae. Thoracic ventrites Proventrite small, about as long as wide; anteriorly, ventrally expanded to partially cover mouthparts, proventral process extending beyond procoxae, expanded anteriorly, tip truncate; mesoventrite U-shaped, surrounding proventral process apex, much wider than long, metaventrite large, wider than mesoventrite. Abdominal ventrites First three ventrites large, each as long or longer than 4th and 5th combined, fourth very short, fifth smaller than first three but much longer than fourth. Sutures visible between all ventrites; fifth ventrite distinctly transversely concave. Legs Femora expanded apically, tibia similar in length to femora; first tarsomere largest; metacoxae with posterior extension lateral to the trochanter; pro-, mesocoxae conical shaped, moderately projecting, metacoxae transverse, extending laterally to elytral margin. Sexual dimorphism Male antennomeres 6-11 distinctly more elongate than those of the female. Eye size in both sexes approximately equal., Published as part of Gearner, Olivia M. & Philips, T. Keith, 2021, Notaferrum n. gen. (Coleoptera: Ptinidae): the first known spider beetle associated with weaver ants, pp. 29-36 in Zoosystema 43 (2) on pages 30-31, DOI: 10.5252/zoosystema2021v43a2, http://zenodo.org/record/4464304, {"references":["PIC M. 1906. - Sur divers Ptinides et Anobiides [Col.]. Bulletin de la Societe entomologique de France 11: 227 - 228."]}

Meurgey, François, Ramage, Thibault (2020): Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography. Insecta Mundi 2020 (786): 1-183, DOI: http://doi.org/10.5281/zenodo.5353608, {"references": ["Adamson, A. M. 1938. Notes on termites destructive to buildings in the Lesser Antilles. Tropical Agriculture (Trinidad) 15: 220-224.", "Aiguillon B., and B. Borthury. 2017. Les termites des iles de la Guadeloupe. Rapport d'etude. Parc National de la Guadeloupe; Saint-Claude, Guadeloupe. 162 p.", "Allemand, R., C. Lemaitre, F. Frey, M. Bouletreau, F. Vavre, G. Nordlander, J. van Alphen, and Y. Carton. 2002. Phylogeny of six African Leptoptilina species (Hymenoptera; Cynipoidea, Figitidae) parasitoids of Drosophila, with description of three new species. Annales de la Societe Entomologique de France 38: 319-332.", "Ashmead, W. H. 1900. Report upon the aculeate Hymenoptera of the islands of St Vincent and Grenada, with additions to the parasitic Hymenoptera and a list of the described Hymenoptera of the West Indies. Transactions of the Entomological Society of London 2: 207-367.", "Badonnel, A. 1981. Psocopteres geophiles des Petites Antilles. Revue d'Ecologie et de Biologie du sol 18(3): 425-434.", "Balke, M., J. Hajek, and L. Hendrich. 2017. Generic reclassification of species formerly included in Rhantus Dejean (Coleoptera, Dytiscidae, Colymbetinae). Zootaxa 4258(1): 91-100.", "Balmford, A., and K. J. Gaston. 1999. Why biodiversity surveys are good value. Nature 398: 204-205.", "Barbut, J., and B. Lalanne-Cassou. 2009. Contribution a la connaissance des Noctuidea des Antilles et descriptions de quatre nouvelles especes (Lepidoptera). Bulletin de la Societe Entomologique de France 114(4): 409-418.", "Barbut, J., and B. Lalanne-Cassou. 2010. Description d'une nouvelle especede Mythimna Ochsenheimer, 1816, sous-genre Pseudaletia Franclemont, 1951 (Lepidoptera Noctuidae Hadeninae). L'Entomologiste 66(3): 117-121.", "Barnes, M. J. C. 1993. Papillons de nuit (Lepidopteres) recoltes par M.J.C. Barnes dans le Parc National de la Guadeloupe. Rapport dactylographie. PRAG; Guadeloupe. 25 p. + pl.", "Barre, N. 1991. Arthropodes d'importance veterinaire pour les petits ruminants des Antilles et de Guyane. Revue d'Elevage et de Medecine Veterinaire des Pays Tropicaux, Numero Special: 133-138.", "Beard, J. S. 1949. The natural vegetation of the Windward and Leeward Islands. Oxford Forestry Memoirs 21: 1-192.", "Beccaloni, G. and P. Eggleton. 2013. Order Blattodea. p. 46-48. In: Z.-Q. Zhang, (ed.). Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703: 1-82.", "Becker, V. O. 2002. The Geometroidea (Lepidoptera) from Cuba described by Herrich-Schaffer in the Gundlach collection, Havana. Revista Brasileira de Zoologia 19(2): 393-417.", "Belkin, J. N. and S. J. Heinemann. 1976. Collection Records of the Project \"Mosquitoes of Middle America\". 5. French West Indies: Guadeloupe (FWI) and Martinique (FWIM, MAR). Mosquito Systematics 8(2): 163-193.", "Bennett, F. D., H. Glenn, and R. M. Baranowski. 1993. Goetheana shakespearei (Hymenoptera: Eulophidae) an immigrant parasitoid of thrips in Florida and Guadeloupe. The Florida Entomologist 76(2): 395-397.", "Bensted-Smith, R. 2002. A biodiversity vision for the Galapagos Islands. Charles Darwin Foundation and World Wildlife Fund; Galapagos, Ecuador. 141 p.", "Bequaert, J. C. 1948. The Genus Pachodynerus (Hymenoptera, Vespidae) in the Antilles. Psyche 55(3): 105-112.", "Bigot, L., and J. Etienne. 2009. Les Pterophoridae de l'ile de la Guadeloupe (Lepidoptera). Bulletin de la Societe entomologique de France 114(4): 463-467.", "Bland, R. G., and L. Desutter-Grandcolas. 2003. An annotated list of Orthoptera from St Eustatius and Saba, Dutch West Indies, with descriptions of new cricket species (Trigonidiidae, Mogoplistidae). Journal of Orthoptera Research 12(2): 115-126.", "Bonfils, J. 1966. Contribution a l'etude des Orthopteroides des Antilles. II. Sur deux especes nouvelles ou peu connues (Orth. Tettigoniidae Pseudoplyllinae). Bulletin de la Societe entomologique de France 71: 77-83.", "Bonfils J. 1969. Catalogue raisonne des insectes des Antilles francaises. 2. Dictyoptera: Blattaria et Mantida. Annales de zoologie, ecologie animale 1(2): 107-120.", "Bonfils, J. 1972. Catalogue raisonne des insectes des Antilles francaises. 5. Dermaptera. Annales de zoologie, ecologie animale 4(3): 271-280.", "Bonfils, J., and A. Delplanque. 1971. Distribution des principales cicadelles des prairies aux Antilles francaises. Annales de zoologie, ecologie animale 3(2): 135-150.", "Bonfils, J., S. Quilici, and B. Reynaud. 1994. Les Hemipteres Auchenorhynques de l'Ile de la Reunion. Bulletin de la Societe entomologique de France 99(3): 227-240.", "Borkent, A. 2008. The frog-biting midges of the world (Corethrellidae: Diptera). Zootax, 1804: 1-456.", "Borkent, A. 2014. The pupae of the biting midges of the world (Diptera: Ceratopogonidae), with a generic key and analysis of the phylogenetic relationships between genera. Zootaxa 3879(1): 1-327.", "Botosaneanu, L. 1988. Trichopteres de la Martinique. Annales de la Societe Entomologique de France (Nouvelle Serie) 24: 215-228.", "Botosaneanu, L. 1989. Seconde contribution a l'etude des Trichopteres de la Martinique. Annales de la Societe entomologique de France (Nouvelle Serie) 25(1): 95-104.", "Botosaneanu, L. 1990. Results of a trichopterological (Insecta: Trichoptera) travel to the Lesser Antilles in 1989. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 60: 39-48.", "Botosaneanu, L. 1994. Les Trichopteres de la Guadeloupe. Annales de la Societe entomologique de France (Nouvelle Serie) 30(1): 33-54.", "Botosaneanu, L. 2000. Etude d'une faunule madicole de Guadeloupe: complements a la connaissance des Trichopteres (Tricoptera) de l'ile. Annales de Limnologie 36(4): 249-259.", "Botosaneanu, L. 2002. An annotated checklist of caddisflies from the Caribbean islands, with distribu- tion and bibliography (Insecta, Trichoptera). Bulletin de la Societe entomologique de France 107(1): 79-108.", "Botosaneanu, L., and J. Sykora. 1973. Sur quelques Trichopteres (Insecta: Trichoptera) de Cuba. p. 379-424. In: T. Orghidan (ed.). Resultats des Expeditions Biospeologiques cubano-roumaines a Cuba, volume 1. Editura Academiei Republicii Socialiste Romania; Bucharest. 424 p.", "Boucher, S., A. M. Dutrillaux, and B. Dutrillaux. 2015. Parthenogenetic reproduction demonstrated in the diploid Spasalus puncticollis (Le Peletier & Serville 1825), n. stat., from the Antilles (Coleoptera, Scarabaeoidea, Passalidae). Comptes Rendus Biologies 338: 738-744.", "Boykin, L. M., P. De Barro, D. G. Hall, W. B. Hunter, C. L. McKenzie, C. A. Powell, and R. G. Shetters. 2012. Overview of worldwide diversity of Diaphorina citri Kuwayama mitochondrial cytochrome oxydase 1 haplotypes: two Old World lineages and a New World invasion. Bulletin of Entomological Research 102(5): 573-582.", "Brambila, J. 2000. A Review of Cligenes with the description of a new genus, Valeris (Hemiptera: Rhyparochromidae: Antillocorini). The Florida Entomologist 83(3): 303-315.", "Branger, G. 1972. Observations sur Uthetheisa ornatrix L. en Martinique et Guadeloupe. Alexanor 7(7): 290-296.", "Brevignon, C. 2000. Description d'un nouveau Lycaenidae provenant de la foret cotiere guadeloupeenne (Lepidoptera, Lycaenidae, Theclinae). Lambillionea 100(4): 625-629.", "Brevignon, C. 2001. Le genus Chlorostrymon Clench, 1961 en Guadeloupe (Lepidoptera, Lycaenidae, Theclinae). Lambillionea 101(4): 561-566.", "Brevignon, C. 2003. Les papillons diurnes de la Guadeloupe (Lepidoptera, Papilionidae, Hesperioidea). Lambillionea 103(2): 1-29.", "Brevignon, C. 2004. Description de deux nouvelles sous-especes guadeloupeennes du genus Junonia Hubner, 1819 (Lepidoptera, Nymphalidae, Nymphalinae). Lambillionea 104(1): 72-80.", "Brevignon, C., and L. Brevignon. 2003. Papillons de jour des Antilles francaises. PLB editions; Abymes, Guadeloupe. 64 p.", "Budesinsky, M. I., E. Valtarova, E. Semon, E. Cancello, and C. Bordereau. 2005. NMR structure determination of (11E)-trinervita-1(14),2,11 -triene, a new diterpene from sexual glands of termites. Tetrahedron 61: 10699-10704.", "Buenaventura, E., and T. Pape. 2013. Revision of the New World genus Peckia Robineau-Desvoidy (Diptera: Sarcophagidae). Zootaxa 3622 (1): 1-87.", "Capy, P., J. R. David, R. Allemand, Y. Carton, G. Febvay, and A. Kermarec. 1986. Genetic analysis of Drosophila melanogaster in the French West Indies and comparison with populations from other parts of the world. Genetica 69: 167-176.", "Cardoso, P., T. L. Erwin, P. A. V. Borges, and T. R. New. 2011. The seven impediments in invertebrate conservation and how to overcome them. Biological Conservation 144: 2647-2655.", "Carpintero, D. L. 2002. Catalogue of the Neotropical Anthocoridae (Heteroptera). Revista de la Sociedad Entomologica Argentina 61(1-2): 25-44.", "Carpintero, D. L., and P. M. Anddelape. 2012. Neotropical Scolopini (Hemiptera: Heteroptera: Anthocoridae): new taxa, diagnostic characters and a key to the genera of the tribe. Acta Entomologica Musei Naturalis Pragae 52: 49-66.", "Carvalho, J. C. M. 1985. Mirideos neotropicais, CCXLIX: Especies novas da tribo Hyaliodini Carvalho Drake (Hemiptera). Revista Brasileira de Biologia 44(3): 363-376.", "Carvalho, J. C. M. 1989. Mirideos Neotropicais CCCXV: novas especies da subfamilia Bryocorinae Douglas and Scott (Hemiptera). Anais da Academia Brasileira de Ciencias 61(1): 105-113.", "Carvalho, J. C. M., and L. A. A. Costa. 1996. The genus Fulvius from the Americas (Hemiptera: Miridae). Annales del Institudo de Biologia del Universidad Autonomia de Mexico 65(1): 63-135.", "Celini, L., V. Roy, J. H. C. Delabie, S. Caquineau, K. Questel, and P. Mora. 2013. Premiers signalements de Cyphomyrmex minutus Mayr, 1862 a Saint-Barthelemy, Petites Antilles, et au Mexique (Hymenoptera, Formicidae, Attini). Bulletin de la Societe entomologique de France 118(3): 393-398.", "Chalumeau, F. 1983. Les Coleopteres scarabaeides des Petites Antilles (Guadeloupe a Martinique). Encyclopedie Entomologique (serie 44). Editions Lechevalier; Paris. 295 p.", "Chalumeau, F., and E. Benito-Espinal. 1984. Deux Ctenuchidae (Lepidoptera) ravageurs du laurierrose aux Antilles francaises. Bulletin mensuel de la Societe linneenne de Lyon 5: 175-182.", "Chalumeau, F., and A. Delplanque. 1978. Catalogue commente des Ctenuchidae (Lepidoptera) des Antilles francaises. Bulletin de la Societe Linneenne de Lyon 47(4): 176-187.", "Chalumeau, F., and A. Delplanque. 1985. Les Insectes de la Guadeloupe et des iles avoisinantes. Centre Departemental de la Documentation Pedagogique; Pointe-a-Pitre, Guadeloupe. 45 p.", "Chalumeau, F., and J. Touroult. 2005. Les longicornes des Petites Antilles (Coleoptera, Cerambycidae): taxonomie, ethologie, biogeographie. Series Faunistica 51. Pensoft Publishers; Sofia-Moscow. 241 p.", "Chassain, J., and J. Touroult. 2014. Les Eucnemides des Antilles francaises (Coleoptera, Eucnemidae). p. 39-45. In: J. Touroult (coord.). Contribution a l'etude des Coleopteres des Petites Antilles - Tome II. Supplement au Bulletin de liaison d'ACOREP-France \" Le Coleopteriste \": 1-112.", "Claret, J., and Y. Carton. 1980. Diapause in a tropical species, Cothonaspis boulardi (Parasitic Hymenoptera). Oecologia 45: 32-34.", "Clastrier, J., and J. Legrand. 1984. Forcipomyia (Pterobosca) pinheyi nouvelle espece de l'ile Maurice parasite des ailes de libellules et nouvelles localisations du Subgenus (Diptera, Ceratopogonidae; Odonata). Revue francaise d'entomologie (N.S.) 6(4): 173-180.", "Cochereau, P. 1989. Les travaux de recherche francais en zone tropicale sur les noctuelles et pyrales du mais: synthese bibliographique. L'Agronomie tropicale 44(2): 137-142.", "Cochereau, P. 1991. Installation en Guadeloupe de Cotesia flavipes Cameron (Hymenoptera: Braconidae), un parasite larvaire foreur de la canne a sucre Diatrea saccharalis L. (Lepidoptera: Crambidae). p. 443-450. In: INRA (ed.). Rencontres Caraibes en lutte biologique, Colloques de l'INRA, 58. INRA; Paris.", "Cochereau, P., and A. Jean-Bart. 1990. Echantillonnages sur canne a sucre en Grande-Terre (Guadeloupe). Les pertes de tonnage sur pied et leurs causes. ORSTOM, INRA; Pointe a Pitre. 24 p.", "Constantin, R. 2017. Le genre Ablechrus Waterhouse en Martinique, avec la description d'une espece nouvelle (Coleoptera, Cleroidea, Malachiidae). p. 34-37. In: J. Touroult (coord.). Contribution a l'etude des Coleopteres des Petites Antilles - Tome III. Supplement au Bulletin de liaison d'ACOREP-France \" Le Coleopteriste \": 1-72.", "Corbet, P. S. 1999. Dragonflies: Behaviour and ecology of Odonata. Harley Books; Colchester. 829 p.", "Cordeiro, I. R., and F. F. Moreira. 2015. New distributional data on aquatic and semiaquatic bugs (Hemiptera: Heteroptera: Gerromorpha & Nepomorpha) from South America. Biodiversity Data Journal 3: 1-30.", "Courtney, G. W., and C. L. Hogue. 2018. A review of the net-winged midges (Diptera: Blephariceridae) of the West Indies, with description of a new species from Puerto Rico. Aquatic Insects 39(2-3): 117-154.", "D'Aguilar, J. 1966. Catalogue raisonne des Insectes des Antilles francaises 1. Lepidopteres, Sphingidae. Annales des Epiphyties 1950 17: 251-262.", "Daigle, J. J. 2007. Macrothemis meurgeyi spec. nov. from Guadeloupe (Anisoptera: Libellulidae). Odonatologica 36(2): 191-195.", "Darlington, J. P. E. C. 1992. Survey of termites in Guadeloupe, Lesser Antilles (Isoptera: Kalotermitidae, Rhinotermitidae, Termitidae). The Florida Entomologist 75(1): 104-109.", "David, J. 1973. Activite nycthemerale de quelques especes de Drosophila (Diptera, Drosophilidae) de la Guadeloupe: Importance de la methode de piegeage. Annales de zoologie, ecologie animale 5: 499-506.", "Delattre, P. 1978. Conditions d'etablissement et de dispersion en Guadeloupe d' Apanteles flavipes (Hym; Braconidae), parasite des Pyrales de la Canne a sucre du genus Diatraea (Lep; Pyralidae). Entomophaga 23(1): 43-50.", "Delecolle, J. C. and J. P. Rieb. 1994. Contribution a l'etude des Ceratopogonides de la Guadeloupe. Description de trois especes nouvelles, appartenant aux genres Dasyhelea et Atrichopogon (Diptera, Nematocera). Bulletin de la Societe Entomologique de France 99: 267-279.", "Delplanque, A. 1976. Les insectes ravageurs du sorgho en Guadeloupe. Nouvelles agronomiques des Antilles et de la Guyane 2(1): 15-20.", "Delplanque, A., and F., Chalumeau. 1985. Insectes de la Guadeloupe et iles avoisinantes. CDDP de la Guadeloupe, Pointe-a-Pitre, Guadeloupe. 45 p.", "Delvare, G. 1993. Sur les Megaphragma de Guadeloupe avec la description d'une espece nouvelle (Hymenoptera, Trichogrammatidae). Revue Francaise d'Entomologie (N.S.) 15(4): 149-152.", "Descoins, C., B. Lalanne-Cassou, C. Malosse, and M. L. Milat. 1986. Analyse de la pheromone sexuelle emise par la femelle vierge de Mocis latipes (Guenee) (Noctuidae, Catocalinae) de Guadeloupe, Antilles francaises. Comptes rendus de l'Academie des Sciences de Paris 302: 509-512.", "Descoins, C., J. F. Silvain, B. Lalanne-Cassou, and H. Cheron. 1988. Surveillance d'insectes ravageurs des cultures par piegeage sexuel des males dans les departements de la Guadeloupe et de la Guyane. Agriculture, ecosystems and environment 21(1-2): 53-65.", "Dommanget, J. L. 2000. Note preliminaire sur les collections d'Odonates exotiques mises a disposition de la SFO. Martinia 16(3): 133.", "Donnelly, T. W. 1970. The Odonata of Dominica, British West Indies. Smithsonian Contributions to Zoology 37: 1-20.", "Dumas, L. L., and J. L. Nessimian. 2012. Faunistic catalog of the caddisflies (Insecta: Trichoptera of Parque Nacional do Itatiaia and its surroundings in South Brazil. Journal of Insect Science 12(25): 1-40.", "Dumbardon-Martial, E. 2015. Les Asilidae des Petites Antilles (Diptera). Bulletin de la Societe entomologique de France 120(4): 465-472.", "Dumbardon-Martial, E. 2016. Pollen feeding in the larva of Toxomerus pulchellus (Diptera, Syrphidae). Bulletin de la Societe Entomologique de France 121(4): 413-420.", "Dumbardon-Martial, E. 2017. Decouverte en Guadeloupe de la Dexiine Beskia aelops (Walker, 1849) (Diptera, Tachinidae, Dexiinae). Bulletin de la Societe entomologique de France 122: 357-358.", "Dumbardon-Martial, E., and S. A. Marshall. 2015. New records and behavioral observations for Grallipeza Rondani from Guadeloupe and Martinique (Diptera, Micropezidae, Taeniapterinae). Bulletin de la Societe entomologique de France 120(1): 79-82.", "Dumbardon-Martial, E., and C. Pierre. 2016. Contribution a la faune entomologique de Guadeloupe: Diptera & Cicadomorpha. Parc National de Guadeloupe; Guadeloupe. 38 p.", "Esmenjaud, D. 1984. Les noctuelles (Lepidopteres Noctuidae) des Antilles francaises. Donnees biologiques pour la conception d'une protection integree du mais contre Spodoptera frugiperda (J.E. Smith) et Heliothis zea (Boddie) en Guadeloupe. Ecole Nationale Superieure Agronomique; Montpellier, France. 194 p.", "Etienne, J. 2005. Les pucerons de Guadeloupe, des Grandes et des Petites Antilles (Hemiptera, Aphididae). Bulletin de la Societe entomologique de France 110(4-5): 455-462.", "Etienne, J., D. Burckhardt, and C. Grapin. 1998. Diaphorina citri (Kuwayama) en Guadeloupe, premier signalement pour les Caraibes (Hem., Psyllidae). Bulletin de la Societe entomologique de France 103(1): 32.", "Etienne, J., and P. Champoiseau. 2011. Signalement de deux Pucerons nouveaux pour la Guadeloupe (Hemiptera, Aphididae). Bulletin de la Societe entomologique de France, 116(3): 327-328.", "Etienne, J., and E. Dumbardon-Martial. 2013. Quadrastichus erythrinae Kim: un redoutable ravageur pour les erythrines de Guadeloupe et de Martinique (Hymenoptera, Eulophidae, Tetrastichinae). Bulletin de la Societe entomologique de France 118(2): 155-158.", "Etienne, J., and R. Gagne. 2017. Les Cecidomyiidae de l'ile de la Guadeloupe et recapitulatif des especes connues des Caraibes (Diptera). Bulletin de la Societe entomologique de France 122(4): 455-466.", "Etienne, J., and M. Martinez. 2003. Les Agromyzidae de Guadeloupe: especes nouvelles et notes additionnelles (Diptera). Nouvelle revue d'Entomologie 19(3): 249-272.", "Etienne, J., M. Martinez, and G. Boecasse. 2004. Premiere signalisation averee du ravageur Melanagromyza obtusa (Malloch) dans la region neotropicale (Dipt. Agromyzidae). Bulletin de la Societe entomologique de France 109(1): 93-105.", "Etienne, J., and D. Matile-Ferrero. 2008. Crypticerya genistae (Hempel), nouveau danger en Guadeloupe (Hemiptera, Coccoidea, Monophlebidae). Bulletin de la Societe entomologique de France 113(4): 517-520.", "Etienne, J., D. Matile-Fererro, and T. Kondo. 2014. Phalacrococcus howertoni Hodges and Hodgson (Hemiptera: Coccoidea: Coccidae), a new soft scale record for the island of Guadeloupe. Revista Corpoica, Ciencia y Tecnologia Agropecuaria 15(1): 115-118.", "Etienne, J., D. Matile-Ferrero, F. Leblanc, and D. Marival. 1998. Premier signalement de la cochenille Maconellicoccus hirsutus (Green) en Guadeloupe; situation actuelle de ce ravageur des cultures dans les Antilles francaises (Hemiptera; Pseudococcidae). Bulletin de la Societe entomologique de France 103(2): 159-173.", "Etienne, J., B. Michel, and C. Grapin. 2018. Premier signalement du Puceron Sarucallis kahawaluokalani (Kirkaldy, 1907) et de ses ennemis naturels en Guadeloupe (Hemiptera, Aphididae). Bulletin de la Societe entomologique de France 123(4): 447-450", "Etienne, J., S. Quilici, D. Marival, and A. Franck. 2001. Biological control of Diaphorina citri (Hemiptera: Psyllidae) in Guadeloupe by imported Tamarixia radiata (Hymenoptera: Eulophidae). Fruits 56(5): 307-315.", "Etienne, J., P. Ryckewaert, and B. Michel. 2015. Thrips (Insecta: Thysanoptera) of Guadeloupe and Martinique: Updated check-list with new information on their ecology and natural enemies. Florida Entomologist 98(1): 298-304.", "Etienne, J., and J. C. Streito. 2008. Premier signalement en Guadeloupe et en Martinique de Pseudacysta perseae (Heidemann, 1908), u

Cavoptinus palawanicus nov.sp. (Figs 1 a-d) T y p e m a t e r i a l: Holotype (♁): Philippines, Palawan, San Vicente, October 2019, local collector. Holotype deposited in coll. J. Háva (Jiří Háva, Private Entomological Laboratory & Collection, Únětice u Prahy, Prague-West, Czech Republic). D e s c r i p t i o n: Male. Longly oval, slightly transversally convex, body length 3.2 mm, greatest width 1.2 mm (Fig. 1a). Piceous black, antennae, palpi and legs dark brown, brown setation erect and long, grey setation recumbent and short. Head slightly convex, shiny, finely and densely punctuate, punctures almost touched, with intermixed grey and brown setation, on vertex sparser. Eyes relatively large, globular, glabrous. On front between eyes with a few long erect setae. Palpi dark brown. Antennae consist of eleven antennomeres, filiform, without antennal club (Fig. 1c). All antennomeres with dense, long semi-erect, brown setae. Pronotum slightly longitudinal (Fig. 1b). Sides of pronotum parallel (from dorsal view), in the middle slightly narrowed. Disc of pronotum with short, very narrow, longitudinal depression. The posterior part of pronotum with depression with long grey setae, posteriorly sharpened. Surface of pronotum shiny, finely punctuate. Part of surface covered with short grey setae (especially on border of depression) and with long dense erect, brown setae, especially on border of pronotum. Scutellum triangular, covered densely by grey setae. Elytra parallel, obtusely ended, with distinct shoulders, shiny, with long erect, brown setae and short grey setae. Each elytron with tenth striae consisting of large, longitudinal punctures. Interstriae narrower than striae. Each elytron with one humeral spots of grey setae and the posterior third quarter of grey setation. Legs dark brown, slim, covered with intermixed yellow and white setation, femur more robust. Tibiae as long as tarsi. All tarsomeres short and wide, slightly emarginate. Claws large, without teeth. Metasternum covered with grey setation, with longitudinal suture, inclined anteriorly from base (Fig. 2a). Male genitalia see Fig. 1d. Female: unknown. D i f f e r e n t i a l d i a g n o s i s The new species is similar to Cavoptinus albonotatus PIC, 1931 and C. luzonicus ZAHRADNÍK & HÁVA, 2017 but differs from them by the form of pronotal carinae, elytral spots and the structure of male genitalia. E t y m o l o g y: Named according to the type locality, Palawan Island, the Philippines., Published as part of Háva, Jiří & Zahradník, Petr, 2020, Cavoptinus palawanicus nov. sp., a new species from the Philippines (Coleoptera, Bostrichoidea, Ptinidae), pp. 85-88 in Linzer biologische Beiträge 52 (1) on pages 85-86, DOI: 10.5281/zenodo.4526789

Háva, Jiří, Zahradník, Petr (2020): Cavoptinus palawanicus nov.sp., a new species from the Philippines (Coleoptera, Bostrichoidea, Ptinidae). Linzer biologische Beiträge 52 (1): 85-88, DOI: http://doi.org/10.5281/zenodo.4526789, {"references":["BELLES X. (1983): Revision des Maheoptinus PIC, 1903, et des genres voisins (Col., Ptinidae). - Annales de la Societe Entomologique de France (N.S.) 19: 7-16.","BELLES X. (1991): Los generos Kedirinus nov., Sundaptinus nov. y Hanumanus nov. enel Archipelago Indo-Australiano y sureste Asiatico, y nuevos datos sobre el genero Maheoptinus PIC (Coleoptera, Ptinidae). - Graellsia 47: 71-96.","ZAHRADNIK P. & J. HAVA (2014): Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia). - Zootaxa 3754: 301-352.","ZAHRADNIK P. & J. HAVA (2017): Cavoptinus luzonicus sp. nov., a new species from the Philippines (Coleoptera: Bostrichoidea: Ptinidae). - Studies and Reports, Taxonomical Series 13(2): 511-515."]}

A new genus of flightless spider beetle from Peru with two new species is described. It is characterized by unique heart-shaped fused elytra and a broad pronotum with five basal depressions. The characters of this new genus and species are illustrated and discussed and the possible phylogenetic placement of this taxon is also included.

Sanchez, Andreas, Chittaro, Yannick (2019): Dignomus irroratus (Ptinidae) et Isomira testacea (Tenebrionidae), deux Coléoptères nouveaux pour la faune de Suisse. Entomo Helvetica 12: 161-163, DOI: 10.5169/seals-985875

Dignomus irroratus (Kiesenwetter, 1852) (Ptinidae) (Fig. 1a) • 1 ex., Chamoson (VS), 16.7.– 11.8.2015, 630 m, leg. & coll. A. Sanchez, det. Y. Chittaro, rev. B. Calmont. Aucun spécimen valide de cette espèce n’existant à notre connaissance dans les collections muséales et privées de Suisse consultées, la capture de cet individu constitue sa première mention suisse. L’individu a été récolté au moyen d’un piège d’interception placé dans une chênaie pubescente, en bordure de pelouse steppique. Cette espèce, répandue en Europe centrale et méditerranéenne, remonte en France le Rhône jusqu’à Lyon seulement (Calmont 2016). Sa capture en Valais est dès lors extrêmement intéressante. L’écologie de l’espèce est encore partiellement méconnue. Néanmoins, il semblerait que l’adulte se rencontre sous les écorces, dans les cavités d’arbres et sur les lierres morts ou dépérissants, et parfois sous les pierres à proximité de bergeries (Calmont 2016). La larve se développerait à l’intérieur du crottin épars de brebis (Xambeu 1894)., Published as part of Sanchez, Andreas & Chittaro, Yannick, 2019, Dignomus irroratus (Ptinidae) et Isomira testacea (Tenebrionidae), deux Coléoptères nouveaux pour la faune de Suisse, pp. 161-163 in Entomo Helvetica 12 on page 161, DOI: 10.5169/seals-985875, http://zenodo.org/record/8086607, {"references":["Calmont B. 2016. Coleopteres Gibbinae et Ptininae de France. Arvensis, hors-serie 2, 202 pp.","Xambeu P. 1894. Moeurs et metamorphoses d'insectes. Annales de la societe entomologique de France 63: 455 - 503."]}

Data from the literature and from specimens preserved at the Colección Nacional de Insectos (CNIN) Instituto de Biología, UNAM were used to compile a checklist of the fauna of Coleoptera of the state of Morelos, México. A sum of 70 families, 167 subfamilies, 361 tribes, 1,022 genera, and 2,606 species are recorded; from this 24 species are new records for Morelos. The State of Morelos ranks fourth in Coleoptera species richness for Mexico, following Veracruz (3,176 spp.), Oaxaca (2,148 spp.) and Chiapas (1,734 spp.). The checklist presented here provides a summary that can serve as a basis for future progress in the knowledge of Mexican Coleoptera.

This paper treats 134 new records of Coleoptera for the province of New Brunswick, Canada from the following 41 families: Gyrinidae, Carabidae, Dytiscidae, Histeridae, Leiodidae, Scarabaeidae, Scirtidae, Buprestidae, Elmidae, Limnichidae, Heteroceridae, Ptilodactylidae, Eucnemidae, Throscidae, Elateridae, Lampyridae, Cantharidae, Dermestidae, Bostrichidae, Ptinidae, Cleridae, Melyridae, Monotomidae, Cryptophagidae, Silvanidae, Laemophloeidae, Nitidulidae, Endomychidae, Coccinellidae, Corylophidae, Latridiidae, Tetratomidae, Melandryidae, Mordellidae, Tenebrionidae, Mycteridae, Pyrochroidae, Aderidae, Scraptiidae, Megalopodidae, and Chrysomelidae. Among these, the following four species are newly recorded from Canada: Dirrhagofarsus ernae Otto, Muona & McClarin (Eucnemidae), Athous equestris (LeConte) (Elateridae), Ernobius opicus Fall (Ptinidae), and Stelidota coenosa Erichson (Nitidulidae). The Family Limnichidae is newly reported for New Brunswick, and one species is added to the fauna of Nova Scotia. Stephostethus productus Rosenhauer (Latridiidae), Tetratoma (Abstrulia) variegata Casey (Tetratomidae), and Chauliognathus marginatus (Fabricius) (Cantharidae) are removed from the faunal list of New Brunswick, and additional records of Lacconotus punctatus LeConte (Mycteridae) are presented and discussed. Lindgren funnel traps provided specimens for 104 (78%) of the species and were the sole source of specimens for 89 (66%) of the species reported here, suggesting they are a very useful tool for sampling Coleoptera fauna in the forests of New Brunswick.

Dignomus francescovitalii sp. nov. (Figs 1���6) Type material. Holotype: collection number 1045-1b [CCHH], adult, probably a male. A practically complete beetle (right protarsus lacking) with partially exposed genitalia, included in a small, transparent yellow amber piece embedded in block of GTS-polyester resin with overall dimensions 9��9�� 4 mm. Syninclusions: one stellate Fagaceae trichome. Type strata. Baltic amber, Ypresian to Priabonian, Eocene, 56��33.9 Ma. Type locality. Baltic Sea coast, the Samland Peninsula, the Kaliningrad Region, Russia. Etymology. Patronymic, this new species is dedicated to our colleague and leading researcher of amber Cerambycidae, Dr. Francesco Vitali (Luxembourg). Description. Body length 2.3 mm, maximum width about 1.4 mm; body shape ovoid; body color (as preserved) unicolorous black. Head hypognathous, turned downwards and not visible in dorsal view, evenly convex; frons with sparse erect setae, and with transverse row of erect setae above antennal pits; vertex glabrous, shiny and apparently impunctate; antennal pits contiguous, interantennal space with a sharp, narrow carina. Compound eyes relatively small, hemispherical, convex, entire, with well differentiated facets, apparently without ommatidial setae; distance between compound eyes nearly equal to 1.6�� diameter of one eye. Antennae filiform, with eleven antennomeres, exceeding in length half of the elytra; densely pubescent and with sparse, semierect setae, except antennomere 11 which has only short and recumbent pubescence; scape subcylindrical, robust, about 2�� as wide and 1.3�� as long as pedicel, antennomeres 2���10 subequal in shape and size, elongate, slightly dilated apically, antennomere 11 spindle-shaped; relative length ratios of antennomeres 1��� 11 equal to 10-6-6-6 -6-7- 7-7-7-8-12. Last maxillary palpomere cylindrical. Pronotum slightly transverse, about 1.15�� as wide as long; with distinct constriction in posterior one-fourth; covered with small punctures, each puncture bearing seta; with elevated discal protuberance widely divided medially by longitudinal groove, protuberance becoming concave postero-medially and narrowly reflexed at lateral sides, which show an acute, triangular, small projection; whole pronotum with small dense granules and semierect setae, distance between granules being less than the diameter of one granule; two lateral sides of discal protuberance with very dense, long, rufous hairs. Scutellum small, slightly transverse, semioval, densely pubescent. Elytra sub-ovoid, convex, about 1.2�� as long as wide, with visible humeral calli; each elytron with five small spots of rufous scale-like hairs: three round shaped spots at elytral base, and two oval shaped spots on disc. Elytral punctures small, arranged in 10 striae that extend along entire length of elytron, punctures round to oval showing medial constriction (punctures of striae 9���10); intervals flat, with sparse heterogeneous pubescence. Pro-, meso- and metathotax densely pubescent, with moderately large, deep, round punctures; prosternal intercoxal process wide, about 0.7�� as wide as diameter of procoxa, slightly convex, with lateral margins parallel; metathoracic ventrite almost flat, medially and slightly convex laterally, distance between punctures distinctly larger than diameter of one puncture; metepisternum about 3.9�� as long as wide, with row of punctures at interior lateral margin, slightly widened anteriorly and gradually narrowed posteriorly, anterior margin oblique, lateral margins nearly straight. Legs long, moderately robust, densely pubescent. Procoxae subglobose, separated by about 0.7�� procoxal diameter; mesocoxae subglobose, distinctly separated by almost one mesocoxal diameter; metacoxae narrow, transverse, with groove for reception of metafemora. Femora clavate, with few erect setae ventro-apically. Tibiae almost straight, slightly dilated apically, nearly as long as tibia. Tarsi poorly visible because of beetle location in amber piece; metatarsi about 0.6�� as long as metatibia. Claws thin, free and simple. Abdomen with five visible ventrites, densely pubescent, with sparse, fine semierect setae (distinctly visible on ventrite 5), sparsely punctured (punctures distinctly smaller than metathoratic punctures); abdominal sutures distinct in all their length; relative length (laterally) ratios of ventrites 1���5 equal to 10-12-10-5-8. Genitalia exposed but poorly defined, although apparently corresponding to male aedeagus with median lobe and parameres slender and symmetrical, as usual in extant species of the genus Dignomus (see Bell��s 1996). Diagnosis. Dignomus francescovitalii sp. nov. assigned to Dignomus irroratus species-group based on a combination of the following morphological characters (sensu Bell��s 1996): the structure of pronotum with discal protuberance showing subparallel sides, convex but not very elevated, widely divided medially by longitudinal groove, and with triangular lateral projections located behind the discal protuberance in lateral view. The characteristic sub-ovoid elytra clearly distinguishes this new species from others of the group. Only D. subelongatus (Pic, 1917) from East Africa (Somalia) has similar sub-ovoid elytra, but differs from new species by pronotum with discal protuberance more elevated and elytra more elongated. Dignomus francescovitalii sp. nov. distinctly differs from D. regiomontanus (described from a Baltic amber) in pronotal structure: the latter species has pronotum with discal longitudinal protrusion flanked by groove on each side, but in new species pronotum with elevated discal protuberance widely divided medially by longitudinal groove, protuberance becoming concave postero-medially and narrowly reflexed at lateral sides. Additionally, these species can be distinguished by body length (D. regiomontanus is 3.7 mm long; 2.3 mm in new species), eyes (large and with ommatidial setae in D. regiomontanus; small, without ommatidial setae in new species), and antennae (slender, exceeding in length apex of the elytra in D. regiomontanus; D. francescovitalii sp. nov. possesses comparatively shorter and thicker antennae)., Published as part of Bukejs, Andris, Bell��s, Xavier & Alekseev, Vitalii I., 2018, A new species of Dignomus Wollaston (Coleoptera: Ptinidae) from Eocene Baltic amber, pp. 195-200 in Zootaxa 4486 (2) on pages 196-198, DOI: 10.11646/zootaxa.4486.2.9, http://zenodo.org/record/1436761, {"references":["Belles, X. (1996) El genero Dignomus Wollaston (Coleoptera, Ptinidae). Bolleti de la Societat d'Historia Natural de Balears, 39, 209 - 228."]}

Genus Dignomus Wollaston, 1862 Remarks. The specimen under consideration belongs to the genus Dignomus within Ptinini on the basis of the following morphological characters (Bell��s 1996): (1) antennae 11-segmented, filiform; (2) prothorax constricted near base; (3) pronotum with a protuberance on the disc, divided in two parts by a longitudinal groove; and (4) elytra elongate., Published as part of Bukejs, Andris, Bell��s, Xavier & Alekseev, Vitalii I., 2018, A new species of Dignomus Wollaston (Coleoptera: Ptinidae) from Eocene Baltic amber, pp. 195-200 in Zootaxa 4486 (2) on page 196, DOI: 10.11646/zootaxa.4486.2.9, http://zenodo.org/record/1436761, {"references":["Belles, X. (1996) El genero Dignomus Wollaston (Coleoptera, Ptinidae). Bolleti de la Societat d'Historia Natural de Balears, 39, 209 - 228."]}

Dignomus francescovitalii sp. nov. (Figs 1–6) Type material. Holotype: collection number 1045-1b [CCHH], adult, probably a male. A practically complete beetle (right protarsus lacking) with partially exposed genitalia, included in a small, transparent yellow amber piece embedded in block of GTS-polyester resin with overall dimensions 9×9× 4 mm. Syninclusions: one stellate Fagaceae trichome. Type strata. Baltic amber, Ypresian to Priabonian, Eocene, 56¯33.9 Ma. Type locality. Baltic Sea coast, the Samland Peninsula, the Kaliningrad Region, Russia. Etymology. Patronymic, this new species is dedicated to our colleague and leading researcher of amber Cerambycidae, Dr. Francesco Vitali (Luxembourg). Description. Body length 2.3 mm, maximum width about 1.4 mm; body shape ovoid; body color (as preserved) unicolorous black. Head hypognathous, turned downwards and not visible in dorsal view, evenly convex; frons with sparse erect setae, and with transverse row of erect setae above antennal pits; vertex glabrous, shiny and apparently impunctate; antennal pits contiguous, interantennal space with a sharp, narrow carina. Compound eyes relatively small, hemispherical, convex, entire, with well differentiated facets, apparently without ommatidial setae; distance between compound eyes nearly equal to 1.6× diameter of one eye. Antennae filiform, with eleven antennomeres, exceeding in length half of the elytra; densely pubescent and with sparse, semierect setae, except antennomere 11 which has only short and recumbent pubescence; scape subcylindrical, robust, about 2× as wide and 1.3× as long as pedicel, antennomeres 2–10 subequal in shape and size, elongate, slightly dilated apically, antennomere 11 spindle-shaped; relative length ratios of antennomeres 1– 11 equal to 10-6-6-6 -6-7- 7-7-7-8-12. Last maxillary palpomere cylindrical. Pronotum slightly transverse, about 1.15× as wide as long; with distinct constriction in posterior one-fourth; covered with small punctures, each puncture bearing seta; with elevated discal protuberance widely divided medially by longitudinal groove, protuberance becoming concave postero-medially and narrowly reflexed at lateral sides, which show an acute, triangular, small projection; whole pronotum with small dense granules and semierect setae, distance between granules being less than the diameter of one granule; two lateral sides of discal protuberance with very dense, long, rufous hairs. Scutellum small, slightly transverse, semioval, densely pubescent. Elytra sub-ovoid, convex, about 1.2× as long as wide, with visible humeral calli; each elytron with five small spots of rufous scale-like hairs: three round shaped spots at elytral base, and two oval shaped spots on disc. Elytral punctures small, arranged in 10 striae that extend along entire length of elytron, punctures round to oval showing medial constriction (punctures of striae 9–10); intervals flat, with sparse heterogeneous pubescence. Pro-, meso- and metathotax densely pubescent, with moderately large, deep, round punctures; prosternal intercoxal process wide, about 0.7× as wide as diameter of procoxa, slightly convex, with lateral margins parallel; metathoracic ventrite almost flat, medially and slightly convex laterally, distance between punctures distinctly larger than diameter of one puncture; metepisternum about 3.9× as long as wide, with row of punctures at interior lateral margin, slightly widened anteriorly and gradually narrowed posteriorly, anterior margin oblique, lateral margins nearly straight. Legs long, moderately robust, densely pubescent. Procoxae subglobose, separated by about 0.7× procoxal diameter; mesocoxae subglobose, distinctly separated by almost one mesocoxal diameter; metacoxae narrow, transverse, with groove for reception of metafemora. Femora clavate, with few erect setae ventro-apically. Tibiae almost straight, slightly dilated apically, nearly as long as tibia. Tarsi poorly visible because of beetle location in amber piece; metatarsi about 0.6× as long as metatibia. Claws thin, free and simple. Abdomen with five visible ventrites, densely pubescent, with sparse, fine semierect setae (distinctly visible on ventrite 5), sparsely punctured (punctures distinctly smaller than metathoratic punctures); abdominal sutures distinct in all their length; relative length (laterally) ratios of ventrites 1–5 equal to 10-12-10-5-8. Genitalia exposed but poorly defined, although apparently corresponding to male aedeagus with median lobe and parameres slender and symmetrical, as usual in extant species of the genus Dignomus (see Bellés 1996). Diagnosis. Dignomus francescovitalii sp. nov. assigned to Dignomus irroratus species-group based on a combination of the following morphological characters (sensu Bellés 1996): the structure of pronotum with discal protuberance showing subparallel sides, convex but not very elevated, widely divided medially by longitudinal groove, and with triangular lateral projections located behind the discal protuberance in lateral view. The characteristic sub-ovoid elytra clearly distinguishes this new species from others of the group. Only D. subelongatus (Pic, 1917) from East Africa (Somalia) has similar sub-ovoid elytra, but differs from new species by pronotum with discal protuberance more elevated and elytra more elongated. Dignomus francescovitalii sp. nov. distinctly differs from D. regiomontanus (described from a Baltic amber) in pronotal structure: the latter species has pronotum with discal longitudinal protrusion flanked by groove on each side, but in new species pronotum with elevated discal protuberance widely divided medially by longitudinal groove, protuberance becoming concave postero-medially and narrowly reflexed at lateral sides. Additionally, these species can be distinguished by body length (D. regiomontanus is 3.7 mm long; 2.3 mm in new species), eyes (large and with ommatidial setae in D. regiomontanus; small, without ommatidial setae in new species), and antennae (slender, exceeding in length apex of the elytra in D. regiomontanus; D. francescovitalii sp. nov. possesses comparatively shorter and thicker antennae).

Dignomus francescovitalii sp. nov. is described and illustrated based on a well-preserved specimen from Eocene Baltic amber. This new extinct species is placed into D. irroratus species-group. The distribution of the genus Dignomus including fossil records is mapped.

Anitys rubens (J. J. Hoffmann, 1803) (Ptinidae, Dorcatominae) (Fig. 2b) – 1 ex. (mort) et nombreuses larves, Orvin (BE), 876 m, 5.4.2017, leg., det. & coll. Chittaro Y. – 20 ex. et quelques larves, Orvin (BE), 876 m, 1.6.2017, leg. & det. Chittaro Y., coll. Chittaro Y. & Sanchez A. Tous les exemplaires ont été découverts dans la carie sèche du tronc d’un vieux chêne (Fig 4a) en bordure d’un pâturage boisé. Ils étaient localisés sur une petite surface d’environ 10 x 10 cm de carie sèche pulvérulente rougeâtre, issue de l’activité du polypore Laetiporus sulphureus, située environ 1 cm sous la surface du tronc (Figs 4b, c). De nombreux Pentaphyllus testaceus (Hellwig, 1792) (Tenebrionidae) colonisaient ce même habitat, troué de galeries larvaires, alors que plusieurs Mycetophagus piceus (Fabricius, 1777) (Mycetophagidae) se trouvaient à quelques centimètres de distance. Les autres parties du tronc ne présentaient pas ce degré de décomposition très particulier, correspondant aux indications de la littérature (Möller & Schneider 1992), et aucun autre Anitys rubens n’a été trouvé lors des quelques pointages effectués., Published as part of Chittaro, Yannick & Sanchez, Andreas, 2018, Limoniscus violaceus (P. W. J. Müller, 1821), Anitys rubens (J. J. Hoffmann, 1803) et Philothermus evanescens (Reitter, 1876), trois Coléoptères « reliques de forêts primaires » nouveaux pour la Suisse, pp. 9-15 in Entomo Helvetica 11 on page 12, DOI: 10.5169/seals-986019, http://zenodo.org/record/8033757, {"references":["Moller G. & Schneider M. 1992. Koleopterologisch-entomologische Betrachtungen zu Alt- und Totholzbiotopen in der Umgebung Berlins - Teil 1. Entomologische Nachrichten und Berichte 36: 73 - 86."]}

Chittaro, Yannick, Sanchez, Andreas (2018): Limoniscus violaceus (P. W. J. Müller, 1821), Anitys rubens (J. J. Hoffmann, 1803) et Philothermus evanescens (Reitter, 1876), trois Coléoptères «reliques de forêts primaires» nouveaux pour la Suisse. Entomo Helvetica 11: 9-15, DOI: 10.5169/seals-986019

Nine new species of the genusClada(s. str.) Pascoe, 1887 (Bostrichoidea: Ptinidae: Eucradinae) are described from Madagascar: Clada (Clada) barclayisp. n., C. (C.) dimbyisp. n., C. (C.) fasciatasp. n., C. (C.) lalaesp. n., C. (C.) madagascarensissp. n., C. (C.) mamyisp. n., C. (C.) njakaisp. n., C. (C.) obesasp. n., and C. (C.) rindraisp. n.No species of this genus were previously known from Madagascar. Photographs of the dorsal habiti and drawings the male and female antennae and aedeagi of most of these species are given.

Mitter, Heinz (2017): Bemerkenswerte Käferfunde aus Oberösterreich XIV und Anhang weiterer bemerkenswerter Funde aus Österreich (Insecta: Coleoptera). Linzer biologische Beiträge 49 (2): 1489-1504, DOI: http://doi.org/10.5281/zenodo.5412239, {"references":["BENSE U. (1995): Bockkafer. Illustrierter Schlussel zu den Cerambyciden und Vespreiden Europas, Weikersheim.","DALLA TORRE K.W. v. (1879): Die Kaferfauna von Oberosterreich. Systematisches Verzeichnis der in Oberosterreich bisher beobachteten Kafer. - 10. Jber. Ver. Natkde. in Oesterr. ob der Enns zu Linz: 1-125.","ECKELT A. & M. KAHLEN (2012): Die holzbewohnende Kaferfauna des Nationalperk Kalkalpen in Oberosterreich (Coleoptera). - Beitr. Naturkunde Oberosterreichs 22: 3-57, Linz.","ERMISCH K. (1969): 79. Familie: Mordellidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","FRANZ H. (1970): Die Nordostalpen im Spiegel ihrer Landtierwelt 3, Innsbruck.","FRANZ H. (1974): Die Nordostalpen im Spiegel ihrer Landtierwelt 4, Innsbruck.","FREUDE H. (1971): 12. Familie: Silphidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 3, Krefeld.","FREUDE H. (1976): 1. Familie: Carabidae (Laufkafer). - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 2, Krefeld.","GEISER E. (2001): Die Kafer des Landes Salzburg. - Monographs on Coleoptera, Vol. 2.","HARDE K.W. (1966): 87. Familie: Cerambycidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 9, Krefeld.","HARDE K.W. (1979): 38. Familie: Buprestidae (Prachtkafer). - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","HORION A. (1949): Faunistik der mitteleuropaischen Kafer 2, Frankfurt a.M.","HORION A. (1956): Faunistik der mitteleuropaischen Kafer 5, Tutzing.","HORION A. (1958): Faunistik der mitteleuropaischen Kafer 6, Uberlingen.","HORION A. (1974): Faunistik der mitteleuropaischen Kafer 12, Uberlingen.","KASZAB Z. (1969): 71. Familie: Pythidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KASZAB Z. (1969): 73. Familie: Scraptiidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KASZAB Z. (1969): 75. Familie: Anthicidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KASZAB Z. (1969): 80. Familie: Serropalpidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KASZAB Z. (1969): 82. Familie: Alleculidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KASZAB Z. (1969): 83. Familie: Tenebrionidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","KLOIBER J. (1956): Das Vorkommen der Curculioniden im Linzer Raum und in einigen anderen oberosterreichischen Landesteilen. - Naturkundl. Jb. Stadt Linz 1956: 241-292.","KLOIBER J. (1957): Das Vorkommen der Curculioniden im Linzer Raum und in einigen anderen oberosterreichischen Landesteilen. - Naturkundl. Jb. Stadt Linz 1957: 91-161.","KOLLER F. (1961): Die Bockkafer (Cerambycidae) Oberosterreichs mit besonderer Berucksichtigung des Grossraumes von Linz. - Naturk. Jb. Stadt Linz 1961: 235-276.","LOHSE G.A. (1964): 23. Familie: Staphylinidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mittleuropas 4, Krefeld.","LOHSE G.A. (1969): 68. Familie: Anobiidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","LOHSE G.A. (1971): 9. Familie: Hydrophilidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 3, Krefeld.","LOHSE G.A. (1979): 32. Familie: Derodontidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE: Die Kafer Mitteleuropas 6, Krefeld.","LOHSE G.A. (1979): 34. Familie: Elateridae. - In: FREUDE H., HARDE K.W. & G.A.LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","LOHSE G.A. (1979): 36. Familie: Eucnemidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","LOHSE G.A. (1979): 40. Familie: Helodidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","LOHSE G.A. (1979): 45. Familie: Dermestidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","LOHSE G.A. (1983): 93. Familie: Curculionidae, 11. Unterfamilie: Cleoninae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 11, Krefeld.","LOHSE G.A. (1983): 93. Familie: Curculionidae, 18. Unterfamilie: Notarinae. - In: FREUDE H., HARDE K.W. & G.A, LOHSE, Die Kafer Mitteleuropas 11, Krefeld.","LOHSE G.A. (1983): 93. Familie: Curculionidae, 20. Unterfamilie: Pissodinae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 11, Krefeld.","LOHSE G.A. (1983): 93. Familie: Curculionidae, 28. Unterfamilie: Ceutorhynchinae. - In: FREUDE H. HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 11, Krefeld.","LOHSE G.A. & W.H. LUCHT (1989): 9. Familie: Hydrophilidae. - In: Lohse G.A. & W.H. Lucht, Die Kafer Mitteleuropas 12 (1. Supplementband), Krefeld.","LUCHT W. (1998): 32.a Familie: Trogossitidae. - In: LUCHT W. & B. KLAUSNITZER, Die Kafer Mitteleuropas 15 (4. Supplementband), Jena.","MACHATSCHKE J.W. (1969): 85. Familie: Scarabaeidae (Blatthornkafer). - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 8, Krefeld.","MITTER H. (1981): Zur Verbreitung einiger Arten der Familie Serropalpidae in Oberosterreich (Coleoptera, Serropalpidae). - NachrBl. bayer. Ent. 30: 79-80.","MITTER H. (2000): Die Kaferfauna Oberosterreichs (Coleoptera: Heteromera und Lamellicornia). - Beitr. Naturk. Oberosterreichs 8: 3-192.","MITTER H. (2000): Verbreitung und Biologie der Eucnemidae und Lissomidae (Coleoptera, Elateroidea) in Oberosterreich. - Beitr. Naturk. Oberosterreichs 9: 39-45.","MITTER H. (2001): Die Kaferfauna des Naturschutzgebietes \"Staninger Leiten\" (Oberosterreich, Unteres Ennstal). - Beitr. Naturk. Oberosterreichs 10: 415-437.","MITTER H. (2001): Bestandsanalyse und Okologie der nach FFH-Richtlinie geschutzten Kafer in Oberosterreich (Insecta, Coleoptera). - Beitr. Naturk. Oberosterreichs 10: 439-448.","MITTER H. (2002): Vergleichende Untersuchung der Kaferfauna an drei Halbtrockenrasen im Unteren Steyrtal (Keltenweg, Kreuzberg, Kuhschellenboschung Neuzeug, Oberosterreich). - Beitr. Naturk. Oberosterreichs 11: 5-34.","MITTER H. (2005): Bemerkenswerte Kaferfunde aus Oberosterreich VIII (Insecta: Coleoptera). - Beitr. Naturk. Oberosterreichs 14: 411-433.","MITTER H. (2007): Bemerkenswerte Kaferfunde aus Oberosterreich IX (Insecta: Coleoptera). - Beitr. Naturk. Oberosterreichs 17: 169-194.","MITTER H. (2008): Bemerkenswerte Kaferfunde aus Oberosterreich X (Insecta: Coleoptera). - Beitr. Naturk. Oberosterreichs 18: 297-303.","MITTER H. (2009): Bemerkenswerte Kaferfunde aus Oberosterreich XI (Insecta: Coleoptera). - Beitr. Naturk. Oberosterreichs 19: 339-346.","MITTER H. (2012): Bemerkenswerte Kaferfunde aus Oberosterreich XII (Insecta: Coleoptera). - Beitr. Naturk. Oberosterreichs 22: 69-82.","MITTER H. (2015): Bemerkenswerte Kaferfunde aus Oberosterreich XIII (Insecta: Coleoptera). - Linzer biol. Beitr. 47 (1): 691-706.","MULLER- MOTZFELD G. (Hrsg.) (2004): Bd. 2 Adephaga 1: Carabidae (Laufkafer). - In: FREUDE H., HARDE K.W., LOHSE G.A. & B. KLAUSNITZER, Die Kafer Mitteleuropas 2, 2. Aufl., Heidelberg/Berlin.","REDTENBACHER L. (1874): Fauna Austriaca, 3. Aufl., Wien.","SCHUH R, PLONSKI I.S. et al. (2011): Bemerkenswerte Kaferfunde aus Osterreich (XVIII) (Coleoptera). - Koleopt. Rdsch. 81: 327-332, Wien.","SPORNRAFT K. (1967): 50. Familie: Nitidulidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 7, Krefeld.","VOGT H. (1967): 59. Familie: Mycetophagidae. - In: FREUDE H, HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 7, Krefeld.","VOGT H. (1967): 60. Familie: Colydiidae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 7, Krefeld.","WITTMER W. (1979): 27. Familie: Cantharidae, pars Malthinini. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 6, Krefeld.","WITZGALL K. (1971): 10. Familie: Histeridae. - In: FREUDE H., HARDE K.W. & G.A. LOHSE, Die Kafer Mitteleuropas 3, Krefeld."]}

Ochina ptinoides (MARSHAM, 1802) M a t e r i a l: Haid bei Ansfelden, 1.6.2015 und 4.6.2017, leg. A. Link. An trockenem Efeu, im Süden und Westen und in wärmeren Gegenden durchaus nicht selten; in den meisten Gebieten aber fehlend. E r s t n a c h w e i s f ü r O b e r ö s t e r r e i c h!, Published as part of Mitter, Heinz, 2017, Bemerkenswerte Käferfunde aus Oberösterreich XIV und Anhang weiterer bemerkenswerter Funde aus Österreich (Insecta: Coleoptera), pp. 1489-1504 in Linzer biologische Beiträge 49 (2) on page 1495, DOI: 10.5281/zenodo.5412239

Hemicoelus favonii sp. nov. is described and illustrated from Eocene Baltic amber. This new fossil species differs from extant congeners in having 11-segmented antennae; a metathoracic ventrite with large impression in its anterior portion; a pronotum distinctly narrower than the elytral base region; the posterior suture of abdominal ventrite 1 weakly arcuate medially; sharp lateral pronotal margins that are incomplete and distinct in their basal half only; elytral striae that are not grouped in pairs; posterior pronotal angles that are rounded; elytral intestriae 3, 5, 7 and 9 that are distinctly convex; and a comparatively small total body size. The presence of Hemicoelus in Baltic amber suggests that moist, rotting wood was available as a microhabitat in the ancient forest. Beyond the new species description, the systematic placement of Anobium jacquelinae Hawkeswood, Makhan & Turner is discussed. A new fossil record for Microbregma waldwico Bukejs & Alekseev, and the first report of the genus Trichodesma LeConte from Eocene Baltic amber are also presented.

Tribe Anobiini Fleming, 1821 Genus Hemicoelus LeConte, 1861 Remarks. The specimen considered here was assigned to the genus Hemicoelus within the tribe Anobiini, on the basis of the following morphological characters: (1) distinctly clubbed antennae with antennomeres 9���11 longer than the preceding five antennomeres combined; (2) procoxae distinctly separated by a parallel-sided intercoxal process; (3) pubescence of body dorsal surface unicolourous and homogeneous, without long erect setae; (4) all abdominal ventrite sutures distinct throughout length, with ventrites 3 and 4 subequal in length; (5) metathoracic ventrite with a deep anterior impression not reaching its middle; (6) elytral apices indistinctly truncated, not tapered., Published as part of Bukejs, Andris, Alekseev, Vitalii I., Cooper, David M. L., King, Gavin A. & Mckellar, Ryan C., 2017, Contributions to the palaeofauna of Ptinidae (Coleptera) known from Baltic amber, pp. 181-188 in Zootaxa 4344 (1) on page 182, DOI: 10.11646/zootaxa.4344.1.12, http://zenodo.org/record/1042239

Peck, Stewart B. (2016): The beetles of the Lesser Antilles (Insecta, Coleoptera): diversity and distributions. Insecta Mundi 2016 (460): 1-360, DOI: http://doi.org/10.5281/zenodo.5182961, {"references":["Platerosfraternus Gorham 1898a: 317; Lengand Mutchler 1922: 430; Blackwelder 1944-1957: 349. Distribution. St. Vincent; singleisland endemic.","Platerospalliatus Gorham 1898a: 317; Lengand Mutchler 1922: 430; Blackwelder 1944-1957: 349. Distribution. Grenada*, St. Vincent; Lesser Antillesendemic. Plate 21."]}

Sucinoptinus brevipennis sp. n. M a t e r i a l. Holotype. Possibly male. SIZK D-2295, Dubrovitsa, Rovno amber, Late Eocene (fig. 1, 2). Syninclusions: Mycetophilidae, Acari (Parasitengona), numerose stellate hairs. D e s c r i p t i o n. Length: 1.6 mm. Slender, parallel-sided; color brownish-piceus (fig. 1, 2). Head finely punctuated and pubescent; interantennal space narrow and rather flat; eyes hemispherical, quite prominent and finely faceted. Antennae eleven segmented, slender and long, slightly longer than the elytral length, covered with semierect pubescence; antennomere proportions according to the formula: 7–3–4–4–4–4–4–4–4.5–5–8.5 (fig. 1, 2). Pronotum slightly longer than wide and wider than a single elytron; apical margin widely convex anteriorly, partially concealing the head; sides slightly rounded, only slightly constricted near the base, moderately convex at the disk and showing a quite apparent transversal depression in the posterior third; surface irregularly sculptured in the anterior third, with round, small tubercles in the disk, and with small elongated tubercles in the posterior third transversal depression; pubescence formed by short semi-recumbent hairs evenly distributed (fig. 1, 2). Legs short and robust covered with short recumbent pubescence; tarsi relatively long, nearly as long as tibiae; mesotarsi with tarsomere proportions according to the formula: 5–1.5–1.5–1–5. Scutellum triangular, as long as wide. Elytra subparallel and short, only slightly longer than twice the length of the pronotum; humeri prominent; elytral surface serially punctuated by elongated strial punctures, leaving an interstriae interval about twice wider as the width of the striae; pubescence formed by semirecumbent short setae inserted in the punctures (with a length similar to that of the puncture) and erect and moderately long (somewhat longer than those of the punctures) setae inserted in the intervals (fig. 1, 2). D i a g n o s t i c c h a r a c t e r s. S. brevipennis is similar to the previously known species, S. bukejsi and S. sucini in having the same general structure of the prothorax, but differing from these species in having much shorter elytra, with a proportion length of the pronotum (LP)/length of the elytra (LE) = 2.1. This clearly differs from the proportion found in S. sucini (LE/LP = 2.7), in S. bukejsi (LE/LP = 2.6, according to measurement made on the pictures sent by V. Alekseev) and in S. rovnoensis, described above, where LE/LP = 3.0. E t y m o l o g y. The specific name refers to the shortness of the elytra, which is the most typical feature of the new species., Published as part of Bellés, X. & Perkovsky, E. E., 2016, New Data On The Genus Sucinoptinus (Coleoptera, Ptinidae) From Rovno Amber, pp. 17-22 in Vestnik Zoologii 50 (1) on pages 18-20, DOI: 10.1515/vzoo-2016-0002, http://zenodo.org/record/6414445

Sucinoptinus ? sp.? M a t e r i a l. SIZK K-25483, Klesov, Rovno amber, Late Eocene. The inclusion contains a spider beetle, with only the ventral and right lateral parts preserved. Moreover, the remains are affected by a sort of a cloudy secretion (fig. 1, 5) that makes more difficult the observations. However, the lateral part of the prothorax showing a characteristic depression near the base in the dorsal part, suggests that it belongs to the genus Sucinoptinus., Published as part of Bellés, X. & Perkovsky, E. E., 2016, New Data On The Genus Sucinoptinus (Coleoptera, Ptinidae) From Rovno Amber, pp. 17-22 in Vestnik Zoologii 50 (1) on page 20, DOI: 10.1515/vzoo-2016-0002, http://zenodo.org/record/6414445

Bellés, X., Perkovsky, E. E. (2016): New Data On The Genus Sucinoptinus (Coleoptera, Ptinidae) From Rovno Amber. Vestnik Zoologii 50 (1): 17-22, DOI: 10.1515/vzoo-2016-0002, URL: http://content.sciendo.com/view/journals/vzoo/50/1/article-p17.xml

Sucinoptinus rovnoensis sp. n. M a t e r i a l. Holotype. Possibly female, only thorax and elytra present. SIZK UA-335, Rovno amber, Late Eocene. D e s c r i p t i o n. Length (pronotum plus elytra): 1.5 mm. Relatively robust, roundedsided; color brownish (fig. 1, 1). Pronotum slightly wider than long and neatly wider than a single elytron; apical margin widely convex anteriorly; sides only slightly rounded, practically not constricted near the base, moderately convex at the disk and showing a quite apparent semicircular depression in the posterior third; surface covered with dense tubercles, oval and large, specially within the semicircular depression, where they are almost as large as half the size of the scutellum; pubescence formed by short semi-recumbent hairs inserted between the tubercles (fig. 1, 1). Scutellum triangular, as long as wide. Elytra slightly roundsided and relatively short, 1.35 as long as wide, but clearly longer than twice the length of the pronotum; humeri prominent; elytral surface serially punctuated by elongated strial punctures, leaving an interstriae interval about twice wider as the width of the striae; pubescence formed by recumbent short setae inserted in the punctures (with a length somewhat longer that the puncture), semirecumbent and moderately long (somewhat longer than those of the punctures) setae inserted in the intervals and evenly distributed, and erect and still longer setae sparsely distributed in the interstriae of the apical part (fig. 1, 1). D i a g n o s t i c c h a r a c t e r s. The new species is similar to S. bukejsi, from amber inclusions found at Yantarny (formerly Palmnicken) in the Kaliningrad region, Russia (Alekseev, 2012) in having the same general structure of pronotum and elytra, but differing from this species by the morphology of the pronotum, with sides only slightly rounded and practically not constricted near the base, whereas that of S. bukejsi has the sides well rounded leaving a slight but clear constriction near the base. Moreover, the semicircular depression of the posterior third of the pronotal disk of S. rovnoensis is not present in S. bukejsi, which shows a regularly convex pronotal disk, with a depression in the posterior third only slightly marked. The large tubercles forming the surface of the semicircular depression of S. rovnoensis are neither present in S. bukejsi, which shows a type of tubercles smaller and evenly distributed. S. sucini Bellés et Vitali (2007) also has a semicircular depression in the posterior third of the pronotal disk, but it is narrower and the tubercles of the sculpture within it are much smaller and rounded in shape. E t y m o l o g y. The specific name refers to Rovno region, the geographical origin of the holotype., Published as part of Bellés, X. & Perkovsky, E. E., 2016, New Data On The Genus Sucinoptinus (Coleoptera, Ptinidae) From Rovno Amber, pp. 17-22 in Vestnik Zoologii 50 (1) on page 18, DOI: 10.1515/vzoo-2016-0002, http://zenodo.org/record/6414445, {"references":["Alekseev, V. I. 2012. Sucinoptinus bukejsi sp. nov. (Coleoptera: Ptinidae: Ptinini), the second species of the Tertiary genus from the Baltic amber. Baltic Journal of Coleopterology, 12 (2), 145 - 148.","Belles, X., Vitali, F. 2007. New fossil spider beetles from Baltic amber (Coleoptera Ptinidae). Entomapeiron (P. S.), 2 (2), 17 - 28."]}

Sucinoptinus sp.? M a t e r i a l. Two specimens, SIZK UA-284, Rovno amber, Late Eocene and SIZK K-6067, Klesov, Rovno amber, Late Eocene. The inclusion labeled UA-284 contains a well recognizable spider beetle belonging to the genus Sucinoptinus sp. The specimen is practically complete except the antennae and parts of the legs, but its orientation in the inclusion makes difficult a specific identification. However, it looks like a S. rovnoensis, but with the typical semicircular depression on the posterior third of the pronotum somewhat deeper, and with the strial punctures of the elytra somewhat broader, which leave the interstriae intervals only slightly wider than the punctures (fig. 1, 3). The specimen SIZK K-6067 is also practically complete, but again the orientation makes difficult a complete identification. It looks very similar to the specimen UA-284, but with the pronotum slightly more transversal and the elytra somewhat shorter (fig. 1, 4)., Published as part of Bellés, X. & Perkovsky, E. E., 2016, New Data On The Genus Sucinoptinus (Coleoptera, Ptinidae) From Rovno Amber, pp. 17-22 in Vestnik Zoologii 50 (1) on page 20, DOI: 10.1515/vzoo-2016-0002, http://zenodo.org/record/6414445

PTILININAE Shuckard, 1839 = Sclerasteidae Gistl, 1856., Published as part of Zahradn��k, Petr & H��va, Ji����, 2014, Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia), pp. 301-352 in Zootaxa 3754 (4) on page 329, DOI: 10.11646/zootaxa.3754.4.1, http://zenodo.org/record/224794, {"references":["Shuckard, W. E. (1839) [text and new taxa]. In: Spry, W. & Shuckard, W. E. (Eds.), The British Coleoptera delineated, consisting of figures of all genera of British beetles, drawn in outline by W. Spry, M. E. S. W. Crofts, London, viii + 83 pp. + 86 + 8 pls. (Pt)","Gistl, J. N. F. X. (1856) Die Mysterien der europaischen Insectenwelt. Ein geheimer Schlussel fur Sammler aller Insecten - Ordnungen und Stande, behufs des Fangs, des Aufenhalts - Orst, der Wohnung, Tag - und Jahreszeit u. s. w., oder autoptische Darstellung des Insectenstaats in seinem Zusammenhange zum Bestehen des Naturhaushaltes uberhaupt und insbesondere in seinem Einflusse auf die phanerogamische und cryptogamische Pflanzenberotzerrung Europa´s. Zum ersten Male nach 25 jahrigen einegen Erfahrungen zusammengestellt und herausgegeben. T. Dannheimer, Kempten, xii + 530 + [2] pp. (D, No)"]}

Lasiodermini Böving, 1927 Lasioderma Stephens, 1835: 417. Type species Ptilinus testaceum Duftschmid, 1825 (A, E, I, N, Nt, P, F). Hypora Mulsant et Rey, 1864 b: 294. Type species Ptinus serricorne Fabricius, 1792. Pseudochina Jacquelin du Val, 1860 b: 143. Type species Ptilinus haemorrhoidalis Illiger, 1807. Megorama Fall, 1905: 206. Type species Catorama simplex LeConte, 1865 (E, I, N, Nt, P). Catoramina Pic, 1951 b: 6. Type species Catoramina signata Pic, 1951. Falsoptilinus Pic, 1950 a: 10. Type species Falsoptilinus modestus Pic, 1950. Pseudolasioderma Logvinovskiy, 1978: 26. Type species Lasioderma brevis Reitter, 1891 (P).

Xyletinini Gistl, 1848 = Vrilettini B��ving, 1927. Euvrilletta Fall, 1905: 192, 196. Type species Euvrilletta xyletinoides Fall, 1905 (N, Nt). Oligomerodes Fall, 1905: 154, 161. Type species Oligomerodes catalinae Fall, 1905. Neoxyletinus Espa��ol, 1983: 83. Type species Xyletinus angustatus Pic, 1907 (I, P). Paraxyletinus Espa��ol, 1972 a: 60. Type species Paraxyletinus israelsoni Espa��ol, 1972 (E, P). Pseudoptilinus Leiler, 1969: 57. Type species Ptilinus fissicollis Reitter in Reitter, Saulcy et Weise, 1877 (P). Pseudoptilinus Leiler, 1963: 112. Ptilinus banatensis Pic, 1912. Nomen nudum Trachelobrachys Gemminger, 1870: 124. Type species Brachytrachelus kiesenwetteri Morawitz, 1861 (P). Replacement name. Brachytrachelus Morawitz, 1861: 290. Type species Brachytrachelus kiesenwetteri Morawitz, 1861. Homonym. Vrilletta LeConte, 1874: 64. Type species Vrilletta murrayi LeConte, 1874 (N, F). Pseudoxyletinus Pic, 1903 b: 182. Type species Pseudoxyletinus fulvolineatus Pic, 1903. Xyletinodes Espa��ol, 1983: 87. Type species Xyletinus hiekei Espa��ol, 1972 (E, I). Xyletinomorphus Pic, 1923 a: 6. Type species Xyletinomorphus (Xyletinomorphus) chilensis Pic, 1923. subg. Macroxyletinus Pic, 1923 a: 6. Type species Xyletinomorphus (Macroxyletinus) longipennis Pic, 1923 (Nt). subg. Xyletinomorphus Pic, 1923 a: 6. Type species Xyletinomorphus (Xyletinomorphus) chilensis Pic, 1923 (Nt). Xyletinus Latreille, 1809: 376. Type species Ptilinus ater Creutzer in Panzer, 1796 (E, I, N, Nt, P). subg. Calypterus Mulsant et Godart, 1859: 181. Type species Xyletinus sericans Mulsant et Godart, 1859 (E, P). Notiomimus Wollaston, 1861: 15. Type species Notiomimus fimicola Wollaston, 1861. subg. Evaniocerius Gottwald, 1983: 136. Type species Xyletinus semilimbatus Pic, 1899 (P). sbg. Pseudocalypterus Gottwald, 1977: 159. Type species Xyletinus pectiniferus Fairmaire, 1879 (P). subg. Xeronthobius Morawitz, 1863: 164. Type species Ptilinus pallens Germar, 1824 (P). subg. Xyletinus Latreille, 1809: 376. Type species Ptilinus ater Creutzer in Panzer, 1796 (E, N, P). Sternoplus Mulsant et Rey, 1864 b: 263. Type species Xyletinus ater Creutzer in Panzer, 1796. subg. Xyletomimus Reitter, 1901: 28. Type species Xyletinus sanguineocinctus Fairmaire, 1859 (P). Xyletomerus Fall, 1905: 192, 197. Type species Xyletomerus histricus Fall, 1905 (E, N, Nt). subg. Xyletomeridius Espa��ol, 1968 b: 9. Type species Xyletomerus (Xyletomeridius) longitarsis Espa��ol, 1968 (E). subg. Xyletomerus Fall, 1905: 192, 197. Type species Xyletomerus histricus Fall, 1905 (N, Nt)., Published as part of Zahradn��k, Petr & H��va, Ji����, 2014, Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia), pp. 301-352 in Zootaxa 3754 (4) on pages 333-334, DOI: 10.11646/zootaxa.3754.4.1, http://zenodo.org/record/224794, {"references":["Gistl, J. N. F. X. (1848) Faunula monacensis cantharologica. Isis von Oken, 1848, 13 unpp. [inserted in numbers 6 - 11]. (D, Pt)","Boving, A. G. (1927) The larva of Nevermannia dorcatomoides Fisher with comments on the classification of the Anobiidae according to their larvae (Coleoptera: Anobiidae). Proceedings of the Entomological Society of Washington, 29 (3), 51 - 62. (Pt)","Fall, H. C. (1905) Revision of the Ptinidae of Boreal America. Transaction of the American Entomological Society, 31, 97 - 296. (Pt)","Espanol, F. (1983) Nuevos Anobiidae (Col.) del sudeste Asiatico. Nota 99. Miscelanea Zoologica (Barcelona), 7, 75 - 90. (Pt)","Espanol, F. (1972 a) Notas sobre Anobidos (Col.). LVI. - LVIII. Publicaciones del Instituto de Biologia Aplicada, 52, 49 - 65. (Pt)","Leiler, T. - E. (1969) Ptilinus banatensis Pic = Pseudoptilinus fissicollis Rtt. Entomologisk Tidskrift, 90, 57 - 58. (Pt) Lepesme, P. (1947) Bostrychoidea. In: Fleutiaux, E. J. B., Legros, C., Lepesme, P. & Paulian, R. (Eds.), Coleopteres des Antilles. Vol. I. Faune de L'Empire Francais VII., pp. 194 - 233. (Pt)","Leiler, T. - E. (1963) Eine neue Gattung der Unterfamilie Ptilininae oder Xyletininae, Col., nebst Beschreibung der Puppe von Pseudoptilinus n. gen. (Ptilinus banatensis Pic. Entomologisk Tidskrift 84, 110 - 112. (Pt)","Pic, M. (1912 a) Coleopterorum catalogus. Pars 41 - Ptinidae. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. W. Junk, Berlin, 46 pp. (X)","Gemminger, M. (1870) [Abanderungen vergebener Namen]. Coleopterologische Hefte, 6, 119 - 124. (Pt)","Morawitz, F. F. (1861) Einige fur die russisch - europaische Fauna neue Kafer beschreiben. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 34, 284 - 294. (Pt)","LeConte, J. L. (1874) Descriptions of new Coleoptera chifly from the Pacific slope of North America. Transaction of the American Entomological Society, 5, 43 - 72. (Pt)","Pic, M. (1903 b) Diagnoses generiques et specifiques de divers Coleopteres exotiques. L'Echange, Revue Linneenne, 19, 182 - 183. (Pt)","Pic, M. (1903 a) Coleopteres exotiques nouveaux. L'Echange, Revue Linneenne, 19, 105 - 107. (Pt)","Pic, M. (1923 a) Nouveautes diverses. Melanges Exotico - entomologiques, 38, 1 - 32. (Pt)","Latreille, P. A. (1809) Genera Crustaceorum et Insectorum secundum ordinem naturalem in familias disposita, iconibus exemplisque plurimis explicata. Tomus quartus et ultimus. A. Koenig, Parisiis et Argentorati, 399 pp. (Pt) http: // d. doi. org / 10.5962 / bhl. title. 4819","Mulsant, E. & Godart, A. (1859) Description de trois especes nouvelles de Coleopteres. Opuscules Entomologiques de Lyon, 9, 177 - 183. (Pt)","Wollaston, T. V. (1861) On the Anobiidae of the Canary Islands. The Annals and Magazine of Natural History, ser. 3, 7, 11 - 19. (Pt)","Gottwald, J. (1983) Zur Taxonomie und Faunistik der palaarktischen Xyletinus Latreille, 1809, - Arten (Coleoptera, Anobiidae). Entomofauna, Zeitschrift fur Entomologie, 4, 133 - 137. (Pt)","Pic, M. (1899) Description d'un genre nouveau de Coleopteres d'Algerie. Bulletin de la Societe Entomologique de France, 1899, 28 - 29. (D)","Gottwald, J. (1977) Die palaarktischen Xyletinus - arten (Coleoptera, Anobiidae). Acta Entomologica Bohemoslovaca, 74, 158 - 177. (Pt)","Morawitz, F. F. (1863) Notiz uber die russischen Xyletininae. Horae Societatis Entomologicae Rossicae, 2, 161 - 37. (Pt)","Mulsant, E. & Rey, C. (1864 b) Histoire naturelle des Coleopteres de France. Terediles. F. Savy, Paris, 8 + 392 + 13 pp. + 10 pls. (Pt)","Reitter, E. (1901) Analytische Uebersicht der palaearctischen Gattungen und Arten der Coleopteren - Familien: Byrrhidae (Anobiidae) und Cioidae. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 3 - 64. (Pt)","Espanol, F. (1968 b) Notas sobre anobidos (Col.). XXXIV. Generos de Xyletininae de la fauna etiopica. Publicaciones del Instituto de Biologia Aplicada, 45, 5 - 25. (Pt)","Espanol, F. (1968 a) Notas sobre Anobidos (Coleoptera). XXXI. Generos de Dorcatominae de la fauna etiopica. Publicaciones del Instituto de Biologia Aplicada, 44, 103 - 136. (Pt)"]}

Eucradini LeConte, 1861 Eucrada LeConte, 1861: 202. Type species Hedobia humeralis Melsheimer, 1845 (N, F). Crichtonia Abdullah et Abdullah, 1967: 26. Type species Crichtonia macleani Abdullah et Abdullah, 1967.

Ptinini Latreille, 1802 Casapus Wollaston, 1862: 194. Type species Casapus bonvouloirii Wollaston, 1862. subg. Casapus Wollaston, 1862: 194. Type species Casapus bonvouloirii Wollaston, 1862 (P). Casopus Wollaston, 1864: 237. Type species Casapus bonvouloirii Wollaston, 1862. Pithodes Wollaston, 1862: 200. Type species Casapus (Pithodes) subcalvus Wollaston, 1862. Syn. nov. subg. Piotes Wollaston, 1862: 211. Type species Piotes inconstans Wollaston, 1862 (P). Cyphoniptus Bell��s, 1992 a: 257. Type species Ptinus sulcithorax Pic, 1899 (P). Dignomus Wollaston, 1862: 201. Type species Dignomus gracilipes Wollaston, 1862 (E, I, P). Eutaphrus Mulsant et Rey, 1868 a: 53, 55. Type species Ptinus irroratus Kiesenwetter, 1851. Pseudentaphrus Pic, 1896 b: 109. Type species Ptinus fairmairei Pic, 1895. Pseudeutaphrus Pic, 1930: 1. Type species Ptinus tewfiki Pic, 1930. Epauloecus Mulsant et Rey, 1868 a: 49, 198. Type species Ptinus crenatus Fabricius, 1801 (N, P). Tipnus Thomson, 1863: 141. Type species Tipnus unicolor Piller et Mitterpacher, 1783. Eurostodes Reitter, 1884: 299. Type species Niptus (Eurostodes) quisquiliarum Baudi di Selve, 1874 (P). Eurostoptinus Pic, 1895: 80. Type species Eurostus (Eurostoptinus) algericus Pic, 1895 (P). Eutaphrimorphus Pic, 1898 a: 54. Type species Ptinus (Eutaphrimorphus) raffrayi Pic, 1898 (E). Hanumanus Bell��s, 1991 a: 83. Type species Hanumanus sumateraensis Bell��s, 1991 (I, P). Kedirinus Bell��s, 1991 a: 72. Type species Kedirinus papuanus Bell��s, 1991 (A, I, P). Lapidoniptus Bell��s, 1981: 70. Type species Piarus hispanicus Pic, 1953 (P). Myrmecoptinus Wasmann, 1916: 206. Type species Myrmecoptinus butteli Wasmann, 1916 (A, I, P). Sundaptinus Bell��s, 1991 a: 76. Type species Sundaptinus planophthalmus Bell��s, 1991. Niptodes Reitter, 1884: 299. Type species Niptus (Niptodes) nobilis Reitter, 1884. subg. Niptodes Reitter, 1884: 299. Type species Niptus (Niptodes) nobilis Reitter, 1884 (E, P). Microptinus Kiesenwetter, 1877: 79. Type species Niptus elongatus Boiledieu, 1854. Homonym. subg. Paraniptus Kiesenwetter, 1877: 79. Type species Ptinus globulus Illiger, 1807 (P). subg. Pseudoniptus Reitter, 1884: 298. Type species Niptus (Pseudoptinus) globipennis Reitter, 1884 (P). Niptus Boieldieu, 1856: 662. Type species Ptinus hololeucus Faldermann, 1836 (A, E, I, N, Nt, P, F). Oviedinus Bell��s, 2011: 78. Type species Ptinus angustithorax Bell��s, 1986 (Nt). Piarus Wollaston, 1862: 209. Type species Piarus basalis Wollaston, 1862 (E, P). Pseuderostus Heyden in Heyden, Reitter et Weise, 1906: 424. Type species Ptinus frigidus Boieldieu, 1854 (N, P). Eurostus Mulsant et Rey, 1868 a: 49, 185. Type species Ptinus frigidus Boieldieu, 1854. Homonym. Niptinus Yang 1980: 26. Type species Niptus hilleri Reitter, 1877. Homonym. Niptomimus Pic, 1950 a: 10. Type species Niptus (Niptomimus) subelongatus Pic, 1950. Ptinus Linnaeus, 1767: 565. Type species Cerambyx fur Linnaeus, 1758 (A, E, I, N, Nt, P). subg. Bruchoptinus Reitter, 1884: 303. Type species Ptinus rufipes Olivier, 1790 (P). subg. Cyphoderes Mulsant et Rey, 1868 a: 54, 173. Type species Ptinus bidens Olivier, 1790 (N, P). subg. Gynopterus Mulsant et Rey, 1868 a: 53, 70. Type species Ptinus sexpunctatus Panzer, 1789 (E, N, Nt, P). Ptinobruchus Brenske et Reitter, 1884: 83. Type species Ptinus crassicornis Kiesenwetter, 1877. subg. Pseudoptinus Reitter, 1884: 303. Type species Ptinus coarcticollis Sturm, 1837 (P). subg. Ptinus Linnaeus, 1767: 565. Type species Cerambyx fur Linnaeus, 1758 (A, E, I, N, Nt, P, F). Bruchus M��ller, 1776: xxiii. Type species Cerambyx fur Linnaeus, 1758. Heteroplus Mulsant et Rey, 1868 a: 54, 70. Type species Ptinus (Heteroplus) pusillus Sturm, 1837. Heteroptinus Reitter, 1884: 303 type species Ptinus obesus Lucas, 1849. Pseudobruchus Pic, 1894: 203. Type species Ptinus (Pseudobruchus) pustuliferus Pic, 1894. subg. Tectoptinus Iablokoff-Khnzorian et Karapetyan, 1986: 513. Type species Ptinus tectus Boieldieu, 1856 (A, E, I, N, Nt, P). Scaleptinus Borowski, 2006 b: 17. Type species Ptinus sericans P��ringuey, 1888 (E)., Published as part of Zahradn��k, Petr & H��va, Ji����, 2014, Catalogue of the world genera and subgenera of the superfamilies Derodontoidea and Bostrichoidea (Coleoptera: Derodontiformia, Bostrichiformia), pp. 301-352 in Zootaxa 3754 (4) on pages 330-331, DOI: 10.11646/zootaxa.3754.4.1, http://zenodo.org/record/224794, {"references":["Latreille, P. A. (1802) Histoire Naturelle, Generale et Particuliere des Crustaces et des Insectes. III. F. Dufart, Paris, i - xii + 13 - 468 pp. (B, D, Pt)","Wollaston, T. V. (1862) On the Ptinidae of the Canary Islands. The Transaction of the Entomological Society of London, ser. 3, 1, 190 - 214 + 1 pl. (Pt)","Wollaston, T. V. (1864) Catalogue of the Coleopterous Insects of the Canaries in the collection of the British Museum. printed by order of the Trustees, London, xiii + 648 pp. (Pt)","Belles, X. (1992 a) Cyphoniptus (Coleoptera: Ptinidae) a new genus from the Tibet upland and its southeastern counterparts. Elytron, Suppl. 5, 257 - 260. (Pt)","Pic, M. (1899) Description d'un genre nouveau de Coleopteres d'Algerie. Bulletin de la Societe Entomologique de France, 1899, 28 - 29. (D)","Mulsant, E. & Rey, C. (1868 a) Histoire naturelle des Coleopteres de France. Gibbicolles. Deyrolle, Naturaliste, Paris, 224 + 16 pp. + 14 pls. (Pt)","Pic, M. (1896 b) Habitats de Ptinides du nord de l'Afrique. L'Echange, Revue Linneenne, 12, 107 - 110. (Pt)","Pic, M. (1895) Descriptions de Coleopteres d'Algerie. L'Echange, Revue Linneenne, 11, 78 - 82. (Pt)","Pic, M. (1930) Deux nouveaux Coleopteres d'Egypte et Sinad. Bulletin de la Societe Royale Entomologique d'Egypte, 1930, 1 - 2. (Pt)","Thomson, C. G. (1863) Skandinaviens Coleoptera, synoptisk bearbetade. Tom V. Lundbergska, Lund, 340 pp. (Pt)","Reitter, E. (1884 a) Bestimmungs - Tabellen der europaischen Coleopteren. XI. Bruchidae (Ptinidae). Verhandlungen des Naturforschenden Vereines in Brunn, 22, 295 - 323. (Pt)","Baudi di Selve, F. (1874) Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio: de Europaeis notis quibusdam additis. Pars quinta. Berliner Entomologische Zeitschrift, 17, 317 - 338. (Pt) http: // dx. doi. org / 10.1002 / mmnd. 47918710113","Pic, M. (1898 a) Diagnoses de deux Ptinus de l'Afrique australe et sous - genre Eutaphrimorphus. Miscellanea Entomologica, 6, 54 - 55. (Pt)","Belles, X. (1991 a) Los generos Kedirinus nov., Sundaptinus nov. y Hanumanus nov. en el Archipelago Indo - Australiano y sureste Asiatico, y nuevos datos sobre el genero Maheoptinus Pic (Coleoptera, Ptinidae). Graellsia, 47, 71 - 96. (Pt)","Belles, X. (1981) Lapidoniptus nov. gen. pour Piarus hispanicus Pic, 1953 (Coleoptera, Ptinidae). Spixiana, 62, 69 - 72. (Pt)","Wasmann, E. (1916) Wissenschaftliche Ergebnisse einer Forschungreise nach Ostindien, ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel - Reepen. V. Termitophile und myrmecophile Coleopteren. Zoologische Jahrbucher, Abteilung fur Systematik, Geografie und Biologie der Tiere, 39, 169 - 210 + 2 pl. (Pt)","Kiesenwetter, E. A. H. von (1877) Anobiidae und Cioidae. In: Kiesenwetter, E. A. H. von & Seidlitz, G. 1877 - 1898: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags - Buchhandlung R. Stricker, Berlin, pp. 1 - 200. (E, Pt)","Boieldieu, A. A. (1856) Monographie des Ptiniores. (Suite). Annales de la Societe Entomologique de France, ser. 3, 4, 662. (Pt)","Belles, X. (2011) Systematics of the genus Oviedinus nov. (Coleoptera: Ptinidae), including a fossil new species from Dominican amber, biogeographical remarks and an account on fossil ptinids. Elytron, 24, 77 - 88. (Pt)","Heyden, L. F. J. D. von, Reitter, E. & Weise, J. (1906) Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. Editio secunda. Berlin, Paskau, Caen, E. Reitter, 6 + 775 pp. (Pt)","Yang, C. K. (1980) Niptinus nom. nov. for Eurostus Brown 1940 (Coleoptera: Ptinidae) nec Dallas 1851 (Hemiptera: Pentatomidae). Entomotaxonomia, 2, 26. (Pt)","Pic, M. (1950 a) Coleopteres du globe. L'Echange, Revue Linneenne, 66, 9 - 12. (Pt)","Linnaeus, C. (1767) Systema naturae, per regna tria naturae, secundum classes, ordines, genera. Species cum characteribus, differentiis, synonimis, locis. Tomus I. Editio duodecima. Laurentii Salvii, Holmiae, 533 - 1327 pp. (Pt)","Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus I. Laurenti Salvii, Holmiae, iv + 824 + 1 pp. (D)","Brenske, E. & Reitter, E. (1884) Neuer Beitrag zur Kaferfauna Griechenlands. Unter Mitwirkung der Herren Dr. Eppelsheim und L. Ganglbauer. Deutsche Entomologische Zeitschrift, 28, 17 - 100 + 2 pl. (Pt)","Muller, O. F. (1776) Zoologiae Danicae prodromus, seu animalium daniae et norvegiae indigenarum characteres, nomina et synonyma imprimis popularium. Hallageriis, Hafniae, xxxii + 282 pp. (Pt) http: // dx. doi. org / 10.5962 / bhl. title. 13268","Pic, M. (1894) Descriptions de Coleopteres. Memoires de la Societe Zoologique de France, 7, 203 - 207. (Pt)","Iablokoff - Khnzorian, S. M. & Karapetyan, A. P. (1986) Issledovaniya po rodu Ptinus L. (Coleoptera, Ptinidae). Biologicheskiy Zhurnal Armenii, 39, 512 - 513. (Pt)","Borowski, J. (2006 b) Scaleptinus - a new genus for Ptinus sericans Peringuey, 1888 and Ptinus squamulosus Pic, 1908 and description of two new species benging to the new genus (Coleoptera, Ptinidae). Annals of Warszaw Agricultural University - SGGW, Forestry and Wood Technology, 60, 17 - 23. (Pt)"]}

Sphaericini Portevin, 1931 Neoptinus Gahan in Waterhouse, Gahan et Arrow, 1900: 102. Type species Neoptinus parvus Gahan in Waterhouse, Gahan et Arrow, 1900 (A, I). Ptinosphaerus Bellés et Lawrence, 1984: 35. Type species Ptinosphaerus marginicollis Bellés et Lawrence, 1984. Pitnus Gorham, 1883: 197. Type species Pitnus pygmaeus Gorham, 1883. subg. Octopitnus Bellés, 1992 b: 170, 184. Type species Pitnus australiae Lea, 1923 (A). subg. Pitnus Gorham, 1883: 197. Type species Pitnus pygmaeus Gorham, 1883 (N, Nt). Sphaericus Wollaston, 1854: 263. Type species Ptinus (Sphaericus) dawsoni Wollaston, 1854. subg. Doramasus Bellés, 1994: 74. Type species Sphaericus crotchianus Wollaston, 1854 (P). subg. Leasphaericus Bellés, 1998 b: 264. Type species Sphaericus (Leasphaericus) flavipennis Bellés, 1998 (A). subg. Nitpus Jacquelin du Val, 1860 a: 138. Type species Nitpus gonospermi Jacquelin du Val, 1860 (E, P). Microptinus Wollaston, 1865: 215. Type species Nitpus gonospermi Jacquelin du Val, 1860. subg. Sphaericus Wollaston, 1854: 263. Type species Ptinus (Sphaericus) dawsoni Wollaston, 1854 (A, E, I, N, Nt, P). Outipnus Gozis, 1886: 24. Type species Trigonogenius exiguus Boieldieu, 1854. Ptinodes Wollaston, 1865: 219. Type species Ptinus (Sphaericus) fragilis Wollaston, 1854. Homonym. Tipnus Jacquelin du Val, 1860: 137 type species Trigonogenius gibboides Boieldieu, 1854 Wollastonella R. Lucas, 1920: 559. Type species Ptinus (Sphaericus) fragilis Wollaston, 1854. Replacement name. Stereocaulophilus Bellés, 1995: 43. Type species Strereocaulophilus volcanicus Bellés, 1995 (P). Trigonogenius Solier, 1849: 464. Type species Ptinus (Trigonogenius) globulus Solier, 1849 (A, E, I, N, Nt, P).