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Loretoniscus Ocampo-Maceda, López-Orozco & Campos-Filho gen. nov. urn:lsid:zoobank.org:act: 293A9B09-3480-4B20-807B-64C27672ACD0 Type species Loretoniscus mashiriaensis gen. et sp. nov., by present designation and monotypy. Diagnosis Body convex. Animals with partial conglobation ability; exoantennal conglobation. Cephalon with welldeveloped antennary lobes, triangular frontal shield slightly developed, laterally interrupted by antennal grooves, frontal line delimiting frontal shield on upper portion. Pereonite 1 epimera without dorsolateral furrows and schisma. Pereonites 1–3 without ventral lobes. Pleon outline continuous with that of pereon. Telson triangular with distal portion covering uropod endopods. Antennula with three articles. Antenna with flagellum of three articles, second and third articles divided by thin suture. Mandible with molar penicil dichotomized. Maxillula outer endite with outer group bearing teeth cleft at apex. Uropod protopod subrectangular, exopod inserted distally. Pleopod exopods 1 and 2 with respiratory areas. Etymology The new genus is named after the department where the specimens were collected: Loreto. Remarks In the complex shape of the depressions in the frontal shield of the cephalon, and the absence of dorsolateral furrow on pereonite 1 epimera, Loretoniscus gen. nov. resembles Iansaoniscus; however, it differs in the broad shape of the telson with lateral margins straight (vs concave), uropod protopod subrectangular (vs subquadrangular), and pleopod 1 and 2 exopods with respiratory areas (vs absent). The Pudeoniscidae were recently revised by Campos-Filho et al. (2018b). The diagnostic characters of the family do not comprise the variation observed in Iansaoniscus and Loretoniscus gen. nov. Thus, the diagnosis of the family after Campos-Filho et al. (2018b) is emended as follows: animals with exoantennal conglobation; cephalon with triangular frontal shield and profons having complex depressions, and pereonite 1 epimera with dorsolateral furrow to fit antennae during conglobation (reduced or absent in cave-dwellers or endogean species); epimera of pereonites 1–3 with small ventral lobes or schisma on pereonite 1 (reduced or absent in cave-dwellers or endogean species); pleon outline continuous to that of pereonite 7, neopleurae 3–5 well developed and rectangular, telson triangular or trapezoidal with proximal portion wider than distal portion; antennal flagellum of three articles, second and third divided by suture; uropod protopod subquadrangular, exopod inserted distally, endopod not surpassing the distal margin of telson; and pleopod exopods with respiratory structures. Loretoniscus gen. nov. is included in the family Pudeoniscidae by sharing most of these mentioned characters. The absence of dorso-lateral furrow and schisma on pereonite 1 epimera, and absence of ventral lobes on pereonites 1–3 epimera are closely related with the conglobation ability, an ecomorphological strategy of oniscideans used against predation and to reduce water loss (Edney 1951; Schmalfuss 1984). The new genus includes one new species with endogean habits, and probably the absence of these structures is related with habitat selection.

Loretoniscus Ocampo-Maceda, López-Orozco & Campos-Filho gen. nov. urn:lsid:zoobank.org:act: 293A9B09-3480-4B20-807B-64C27672ACD0 Type species Loretoniscus mashiriaensis gen. et sp. nov., by present designation and monotypy. Diagnosis Body convex. Animals with partial conglobation ability; exoantennal conglobation. Cephalon with welldeveloped antennary lobes, triangular frontal shield slightly developed, laterally interrupted by antennal grooves, frontal line delimiting frontal shield on upper portion. Pereonite 1 epimera without dorsolateral furrows and schisma. Pereonites 1–3 without ventral lobes. Pleon outline continuous with that of pereon. Telson triangular with distal portion covering uropod endopods. Antennula with three articles. Antenna with flagellum of three articles, second and third articles divided by thin suture. Mandible with molar penicil dichotomized. Maxillula outer endite with outer group bearing teeth cleft at apex. Uropod protopod subrectangular, exopod inserted distally. Pleopod exopods 1 and 2 with respiratory areas. Etymology The new genus is named after the department where the specimens were collected: Loreto. Remarks In the complex shape of the depressions in the frontal shield of the cephalon, and the absence of dorsolateral furrow on pereonite 1 epimera, Loretoniscus gen. nov. resembles Iansaoniscus; however, it differs in the broad shape of the telson with lateral margins straight (vs concave), uropod protopod subrectangular (vs subquadrangular), and pleopod 1 and 2 exopods with respiratory areas (vs absent). The Pudeoniscidae were recently revised by Campos-Filho et al. (2018b). The diagnostic characters of the family do not comprise the variation observed in Iansaoniscus and Loretoniscus gen. nov. Thus, the diagnosis of the family after Campos-Filho et al. (2018b) is emended as follows: animals with exoantennal conglobation; cephalon with triangular frontal shield and profons having complex depressions, and pereonite 1 epimera with dorsolateral furrow to fit antennae during conglobation (reduced or absent in cave-dwellers or endogean species); epimera of pereonites 1–3 with small ventral lobes or schisma on pereonite 1 (reduced or absent in cave-dwellers or endogean species); pleon outline continuous to that of pereonite 7, neopleurae 3–5 well developed and rectangular, telson triangular or trapezoidal with proximal portion wider than distal portion; antennal flagellum of three articles, second and third divided by suture; uropod protopod subquadrangular, exopod inserted distally, endopod not surpassing the distal margin of telson; and pleopod exopods with respiratory structures. Loretoniscus gen. nov. is included in the family Pudeoniscidae by sharing most of these mentioned characters. The absence of dorso-lateral furrow and schisma on pereonite 1 epimera, and absence of ventral lobes on pereonites 1–3 epimera are closely related with the conglobation ability, an ecomorphological strategy of oniscideans used against predation and to reduce water loss (Edney 1951; Schmalfuss 1984). The new genus includes one new species with endogean habits, and probably the absence of these structures is related with habitat selection., Published as part of Ocampo-Maceda, Ana Teresa, López-Orozco, Carlos Mario, López-Tejeda, Evaristo Luciano & Campos-Filho, Ivanklin Soares, 2023, First record of Pudeoniscidae Lemos de Castro, 1973 (Oniscidea) from Peru, with the description of a new genus and species, pp. 65-77 in European Journal of Taxonomy 861 on pages 67-68, DOI: 10.5852/ejt.2023.861.2067, http://zenodo.org/record/7728898, {"references":["Campos-Filho I. S., Lisboa J. T. & Cardoso G. M. 2018 b. A new genus and two new species of Pudeoniscidae Lemos de Castro, 1973 (Crustacea: Isopoda: Oniscidea) from Brazil. Journal of Natural History 52 (7 - 8): 457 - 482. https: // doi. org / 10.1080 / 00222933.2018.1437229","Edney E. B. 1951. The evaporation of water from woodlice and the millipede Glomeris. Journal of Experimental Biology 28: 91 - 115.","Schmalfuss H. 1984. Eco-morphological strategies in terrestrial isopods. Symposia of the Zoological Society of London 53: 339 - 368."]}

Family Halophilosciidae Verhoeff, 1908 Remarks The family Halophilosciidae comprises 34 species distributed in three genera, Halophiloscia Verhoeff, 1908 (9 spp.), Littorophiloscia Hatch, 1947 (22 spp.), and Stenophiloscia Verhoeff, 1908 (3 spp.) (WoRMS 2020c). In Colombia, Pearse (1915) recorded Philoscia culebrae Moore, 1901 [= Littorophiloscia c.] in coffee plantations from Hacienda Cincinati and Hacienda La Rosa, Sierra Nevada de Santa Marta (ca 1400 m a.s.l.), which is considered to be a misidentification by Van Name (1936). Littorophiloscia culebrae (Moore, 1901) is recorded from coastal areas of Angola, Brazil, Canary Islands, Caribbean islands, Florida (USA), Hawaii, Iraq, Madagascar, Socotra Archipelago and Yemen (Schmalfuss 2003; Taiti & Ferrara 2004; Araujo & Taiti 2007; Taiti & Lopez 2008; Naser et al. 2015; Lisboa et al. 2017; Campos-Filho et al. 2018). Recent expeditions to the north coast of Bol��var revealed the existence of this species in Colombia, inhabiting the coastal areas of the Bar�� Peninsula, Cartagena de Indias (C. L��pez-Orozco & Y. Carpio-D��az, pers. obs.)., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 29, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Pearse A. S. 1915. An account of the Crustacea collected by the Walker Expedition to Santa Marta, Colombia. Proceedings of the United States National Museum 49: 531 - 556. https: // doi. org / 10.5479 / si. 00963801.49 - 2123.531","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. & Ferrara F. 2004. The terrestrial Isopoda (Crustacea: Oniscidea) of the Socotra Archipelago. Fauna of Arabia 20: 211 - 326.","Araujo P. B. & Taiti S. 2007. Terrestrial isopods (Crustacea, Oniscidea) from Rocas Atoll, northeastern Brazil. Arquivos do Museu Nacional 65: 347 - 355.","Taiti S. & Lopez H. 2008. New records and species of Halophilosciidae (Crustacea, Isopoda, Oniscidea) from the Canary Islands (Spain). In: Zimmer M., Charfi-Cheikhrouha F. & Taiti S. (eds) Proceedings of the International Symposium of Terrestrial Isopod Biology ISTIB- 07: 43 - 58. Shaker-Verlag, Aachen, Germany.","Naser M. D., Khalaf T. A. & Yasser A. G. 2015. First record of the terrestrial isopod Littorophiloscia culebrae (H. F. Moore, 1901) (Isopoda, Oniscidea) from Khor Al-Zubair, Iraq and the Persian Gulf. Crustaceana 88 (5): 611 - 615. https: // doi. org / 10.1163 / 15685403 - 00003432","Lisboa J. T., Campos-Filho I. S., Couto E. C. G. & Araujo P. B. 2017. Distribution of terrestrial isopods of the genus Littorophiloscia (Isopoda, Halophilosciidae) along the Brazilian coast. North-Western Journal of Zoology 13: e 167301.","Campos-Filho I. S., Cardoso G. M. & Aguiar J. O. 2018. Catalogue of terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Brazil: an update with some considerations. Nauplius 26: e 2018038. https: // doi. org / 10.1590 / 2358 - 2936 e 2018038"]}

Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. Figs 1, 2G, 15–17 Microphiloscia trichoniscoides Vandel, 1973: 170, figs 19–21. Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂; Paraíso Secreto; 10°10′15.9″ N, 75°44′38.8″ W; 4 Apr. 2018; C.M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 218, 220, 221 • 1 ♂, 1 ♀ (parts in micropreparations); same locality as for preceding; 6 Sep. 2017; C.M. López-Orozco, Y.M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 219. CUBA – Matanzas, Bahia de Conchinos • 4 ♂♂, 10 ♀♀; near Cueva de los Peces, between Playa Larga and Playa Giron; 10 Apr. 2002; S. Taiti leg.; MZUF 9607. – Pinar del Rio • 2 ♂♂, 3 ♀♀; Penisula Guanahacabibes, La Bajada; 7 Apr. 2002; S. Taiti leg.; MZUF 9608. Redescription MEASUREMENTS. Maximum body length: male 2.8 mm and female 4 mm. BODY. Color light brown (Fig. 2G), mottled with the usual pale muscle spots; posterior margins of pereonites 4–7, pleonites 3–5 and uropod branches more pigmented. Body outline as in Fig. 15A. Back smooth with numerous triangular scale-setae (Fig. 15B). One line of noduli laterales per side on the pereonites inserted at same distance from lateral margins (Fig. 15A, C). CEPHALON. Without frontal line, suprantennal line straight on median portion; eyes consisting of 10 ommatidia (Fig. 15D–E). PLEON. Narrower than pereon; epimera of pleonites 3–5 not developed. Telson with lateral sides slightly concave, distal margin broadly rounded (Fig. 15A). ANTENNULA. Composed of three articles subequal in length, distal article with two lateral aesthetascs plus apical pair (Fig. 15F). ANTENNA. When extended posteriorly reaches posterior margin of pereonite 5; flagellum as long as fifth article of peduncle, first article slightly longer than second and third, second and third articles with three and two lateral aesthetascs, respectively (Fig. 15G). MOUTH. Mandibles with molar penicil consisting of at least five branches, left mandible with 2+1 penicils (Fig. 16A), right mandible with 1+1 penicils (Fig. 16B). Maxillula (Fig. 16C) inner endite with two long penicils inserted transversely; outer endite of 5+5 teeth, inner set apically cleft. Maxilla (Fig. 16D) with bilobate apex, inner lobe rounded, bearing thick setae, outer lobe rounded, covered with thin setae plus three thick setae. Maxilliped (Fig. 16E) base bearing sparse setae, proximal article of palp with two stout setae slightly distinct in length; endite subrectangular, inner and distal margins bearing several thin setae, medial seta not surpassing distal margin, distal margin with one small hairy penicil. UROPOD. Protopod and exopod grooved on outer margin, bearing many glandular pores, exopod almost three times as long as endopod (Fig. 16F). PEREOPODS. Rather slender, pereopod 1 carpus bearing transverse antennal grooming brush and distal seta apically cleft; dactylus with ungual seta simple and surpassing outer claw, dactylar seta simple, not surpassing outer claw. Male PEREOPODS 1 AND 7. Pereopod 1 (Fig. 17A) propodus slightly flattened, carpus with transverse antennal grooming brush and distal seta hand-like. Pereopod 7 (Fig. 17B) without any sexual dimorphism. GENITAL PAPILLA. Ventral shield ovoid, papilla bifurcate with genital orifices opening at apex (Fig. 17C). PLEOPODS. Pleopod 1 (Fig. 17D) exopod triangular, inner margin almost straight, outer margin sinuous, distal portion triangular with distal margin rounded; endopod with thickest distal part with sides almost parallel, inner distal part with long pointed process concave on outer margin, prominent subquadrangular apical lobe on outer distal portion bearing thin setae. Pleopod 2 (Fig. 17E) exopod triangular, outer margin concave, bearing seven stout setae; endopod robust, distinctly longer than exopod. Exopods of pleopods 3–5 as in Fig. 17F–H. Remarks Vandel (1973) erected the genus Microphiloscia to allocate the new species M. trichoniscoides from Cueva de la Colorada, Provincia de Oriente, Sierra Maestra, Colorada del Maso, Cuba. After the examination of material belonging to this species, M. trichoniscoides shows the shape of the genital papilla and male pleopod 1 endopod typical of members of Halophiloscia (see also Taiti & Lopez 2008). Halophiloscia includes 9 halophilic species mainly distributed along the Mediterranean and Atlantic coasts of Europe and Africa (Schmalfuss 2003; Taiti & Lopez 2008; Taiti & Argano 2009, 2011). In America, only H. couchii (Kinahan, 1858) was recorded, from Argentina, Bermuda, and USA (see Schmidt 2003). The genus is mainly characterized by the runner-type habitus (sensu Schmalfuss 1984), epimera of pereonites 1–7 with one or more lines of noduli laterales, male pereopod 1 and sometimes pereopod 2 carpus and propodus enlarged and covered with several scales on frontal side, male genital papilla distally bifurcated, and male pleopod 1 endopod stout bearing a long, pointed process on the apex (see Schmidt 2003; Taiti & Lopez 2008). The species is included within the genus since it shows all the previously mentioned characters, except that the male pereopods 1 and 2 do not have the carpus and propodus enlarged. Thus, Microphiloscia is considered to be a junior synonym of Halophiloscia. Distribution Cuba (Vandel 1973). First record for Colombia.

Key to species of terrestrial isopods from Isla Grande 1. Eyes present....................................................................................................................................... 2 ��� Eyes absent............................... Stenoniscus nestori L��pez-Orozco, Taiti & Campos-Filho sp. nov. 2. Animal with conglobation ability...................................................................................................... 3 ��� Animal without conglobation ability................................................................................................. 6 3. Pereon dorsally smooth; pereonite 5 epimera with ventral phylacomera.......................................... 4 ��� Pereon dorsally tuberculate; pereonite 5 epimera without phylacomera..................................................................................................................................... Ctenorillo tuberosus (Budde-Lund, 1904) 4. Phylacomera 5 paddle-shaped, extended anteriorly.......................................................................... 5 ��� Phylacomera 5 sickle-shaped.................................................. Tylos marcuzzii Giordani Soika, 1954 5. Inner lobe of schisma extending beyond posterior margin of outer lobe.............................................................................................. Tylos negroi L��pez-Orozco, Carpio-D��az & Campos-Filho sp. nov. ��� Inner lobe of schisma not extending beyond posterior margin of outer lobe................................................................................................................................................ Tylos niveus Budde-Lund, 1885 6. Flagellum of the antennae with 4 or more articles............................................................................ 7 ��� Flagellum of the antennae with 2���3 articles.................................................................................... 10 7. Flagellum of the antennae with 4 articles.......................................................................................... 8 ��� Flagellum of the antennae with more than 10 articles......... Ligia baudiniana Milne-Edwards, 1840 8. Dorsal surface almost smooth or with small tubercles...................................................................... 9 ��� Dorsal surface of pereon with 4 prominent longitudinal ribs..................................................................................................................................... Armadilloniscus caraibicus Paoletti & Stinner, 1989 9. Dorsal surface almost smooth.................................................. Armadilloniscus ninae Schultz, 1984 ��� Dorsal surface with small tubercles....................................................................................................................................................... Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. 10. Flagellum of the antennae with 3 articles.........................................................................................11 ��� Flagellum of the antennae with 2 articles........................................................................................ 14 11. Genital papilla with truncate apex................................................................................................... 12 ��� Genital papilla with bifurcated apex....... Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. 12. Male pleopod 1 endopod without lobe on outer margin of apical part............................................ 13 ��� Male pleopod 1 endopod with a lobe on outer margin of apical part.................................................................................................................................. Littorophiloscia tropicalis Taiti & Ferrara, 1986 13. Male pleopod 1 endopod with apex divided into three points........................................................................................................................................ Littorophiloscia amphindica Taiti & Ferrara, 1986 ��� Male pleopod 1 endopod with a long spine and 2 small distinct denticles on the inner margin of the apical part.......................................................... Littorophiloscia denticulata (Ferrara & Taiti, 1982) 14. Pleopod exopods with lungs............................................................................................................ 15 ��� Pleopod exopods without lungs....................................................................................................... 16 15. Pleopodal lungs present in first two pairs of pleopods...... Porcellionides pruinosus (Brandt, 1833) ��� Pleopodal lungs present in all five pairs of pleopods.......................................................................................................................................................... Agnara madagascariensis (Budde-Lund, 1885) 16. Colorless body; eyes composed of two unequal ommatidia........................................................................................................................................ Trichorhina heterophthalma Lemos de Castro, 1964 ��� Pigmented body; eyes with 10 ommatidia........................................................................................................................... Trichorhina bermudezae Carpio-D��az, L��pez-Orozco & Campos-Filho, 2018, Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 40-41, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1904. A Revision of Crustacea Isopoda Terrestria, with Additions and Illustrations. Pt 1 Eubelum, Pt 2 Spherilloninae, Pt 3 Armadillo. H. Hagerup, Copenhagen, Denmark.","Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83.","Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Milne-Edwards M. 1840. Ordre des isopodes. In: Histoire Naturelle des Crustaces, comprenant l'Anatomie, la Physiologie et la Classficiation de ces Animaux. Tome Troisieme. 115 - 283. Librairie Encyclopedique de Roret, Paris.","Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Vandel A. 1973. Les isopodes terrestres et cavernicoles de l'ile de Cuba. In: Orghidan T., Nunez A., Botosaneanu L., Decou V., Negrea S. & Vina N. (eds) Resultats des Expeditions biospeologiques cubanoroumaines a Cuba, vol. 1: 153 - 188. Editura Academiei Republicii Socialiste Romania, Bucharest.","Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Ferrara F. & Taiti S. 1982. Isopodi terrestri delle Isole Andamane. Bollettino del Museo civico di Storia naturale di Verona 8: 459 - 492.","Brandt J. F. 1833. Conspectus Monographiae Crustaceorum Oniscodorum Latreillii. Byulleten Moskovskogo Obshchestva Ispytatelei Prirody 6: 171 - 193.","Lemos de Castro A. 1964. Trichorhina heterophthalma nueva especie de isopodo terrestre cavernicola de Cuba. Poeyana 2 A: 1 - 7.","Carpio-Diaz Y. M., Lopez-Orozco C. M., Campos-Filho I. S. & Navas G. R. 2018. Terrestrial isopods (Isopoda: Oniscidea) of the Botanical Garden of Cartagena \" Guillermo Pineres \", Colombia, with the description of three new species. Arthropoda Selecta 27 (4): 301 - 318. https: // doi. org / 10.15298 / arthsel. 27.4.05"]}

Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. Figs 1, 2G, 15���17 Microphiloscia trichoniscoides Vandel, 1973: 170, figs 19���21. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 218, 220, 221 ��� 1 ♂, 1 ♀ (parts in micropreparations); same locality as for preceding; 6 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 219. CUBA ��� Matanzas, Bahia de Conchinos ��� 4 ♂♂, 10 ♀♀; near Cueva de los Peces, between Playa Larga and Playa Giron; 10 Apr. 2002; S. Taiti leg.; MZUF 9607. ��� Pinar del Rio ��� 2 ♂♂, 3 ♀♀; Penisula Guanahacabibes, La Bajada; 7 Apr. 2002; S. Taiti leg.; MZUF 9608. Redescription MEASUREMENTS. Maximum body length: male 2.8 mm and female 4 mm. BODY. Color light brown (Fig. 2G), mottled with the usual pale muscle spots; posterior margins of pereonites 4���7, pleonites 3���5 and uropod branches more pigmented. Body outline as in Fig. 15A. Back smooth with numerous triangular scale-setae (Fig. 15B). One line of noduli laterales per side on the pereonites inserted at same distance from lateral margins (Fig. 15A, C). CEPHALON. Without frontal line, suprantennal line straight on median portion; eyes consisting of 10 ommatidia (Fig. 15D���E). PLEON. Narrower than pereon; epimera of pleonites 3���5 not developed. Telson with lateral sides slightly concave, distal margin broadly rounded (Fig. 15A). ANTENNULA. Composed of three articles subequal in length, distal article with two lateral aesthetascs plus apical pair (Fig. 15F). ANTENNA. When extended posteriorly reaches posterior margin of pereonite 5; flagellum as long as fifth article of peduncle, first article slightly longer than second and third, second and third articles with three and two lateral aesthetascs, respectively (Fig. 15G). MOUTH. Mandibles with molar penicil consisting of at least five branches, left mandible with 2+1 penicils (Fig. 16A), right mandible with 1+1 penicils (Fig. 16B). Maxillula (Fig. 16C) inner endite with two long penicils inserted transversely; outer endite of 5+5 teeth, inner set apically cleft. Maxilla (Fig. 16D) with bilobate apex, inner lobe rounded, bearing thick setae, outer lobe rounded, covered with thin setae plus three thick setae. Maxilliped (Fig. 16E) base bearing sparse setae, proximal article of palp with two stout setae slightly distinct in length; endite subrectangular, inner and distal margins bearing several thin setae, medial seta not surpassing distal margin, distal margin with one small hairy penicil. UROPOD. Protopod and exopod grooved on outer margin, bearing many glandular pores, exopod almost three times as long as endopod (Fig. 16F). PEREOPODS. Rather slender, pereopod 1 carpus bearing transverse antennal grooming brush and distal seta apically cleft; dactylus with ungual seta simple and surpassing outer claw, dactylar seta simple, not surpassing outer claw. Male PEREOPODS 1 AND 7. Pereopod 1 (Fig. 17A) propodus slightly flattened, carpus with transverse antennal grooming brush and distal seta hand-like. Pereopod 7 (Fig. 17B) without any sexual dimorphism. GENITAL PAPILLA. Ventral shield ovoid, papilla bifurcate with genital orifices opening at apex (Fig. 17C). PLEOPODS. Pleopod 1 (Fig. 17D) exopod triangular, inner margin almost straight, outer margin sinuous, distal portion triangular with distal margin rounded; endopod with thickest distal part with sides almost parallel, inner distal part with long pointed process concave on outer margin, prominent subquadrangular apical lobe on outer distal portion bearing thin setae. Pleopod 2 (Fig. 17E) exopod triangular, outer margin concave, bearing seven stout setae; endopod robust, distinctly longer than exopod. Exopods of pleopods 3���5 as in Fig. 17F���H. Remarks Vandel (1973) erected the genus Microphiloscia to allocate the new species M. trichoniscoides from Cueva de la Colorada, Provincia de Oriente, Sierra Maestra, Colorada del Maso, Cuba. After the examination of material belonging to this species, M. trichoniscoides shows the shape of the genital papilla and male pleopod 1 endopod typical of members of Halophiloscia (see also Taiti & Lopez 2008). Halophiloscia includes 9 halophilic species mainly distributed along the Mediterranean and Atlantic coasts of Europe and Africa (Schmalfuss 2003; Taiti & Lopez 2008; Taiti & Argano 2009, 2011). In America, only H. couchii (Kinahan, 1858) was recorded, from Argentina, Bermuda, and USA (see Schmidt 2003). The genus is mainly characterized by the runner-type habitus (sensu Schmalfuss 1984), epimera of pereonites 1���7 with one or more lines of noduli laterales, male pereopod 1 and sometimes pereopod 2 carpus and propodus enlarged and covered with several scales on frontal side, male genital papilla distally bifurcated, and male pleopod 1 endopod stout bearing a long, pointed process on the apex (see Schmidt 2003; Taiti & Lopez 2008). The species is included within the genus since it shows all the previously mentioned characters, except that the male pereopods 1 and 2 do not have the carpus and propodus enlarged. Thus, Microphiloscia is considered to be a junior synonym of Halophiloscia. Distribution Cuba (Vandel 1973). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 29-32, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Vandel A. 1973. Les isopodes terrestres et cavernicoles de l'ile de Cuba. In: Orghidan T., Nunez A., Botosaneanu L., Decou V., Negrea S. & Vina N. (eds) Resultats des Expeditions biospeologiques cubanoroumaines a Cuba, vol. 1: 153 - 188. Editura Academiei Republicii Socialiste Romania, Bucharest.","Taiti S. & Lopez H. 2008. New records and species of Halophilosciidae (Crustacea, Isopoda, Oniscidea) from the Canary Islands (Spain). In: Zimmer M., Charfi-Cheikhrouha F. & Taiti S. (eds) Proceedings of the International Symposium of Terrestrial Isopod Biology ISTIB- 07: 43 - 58. Shaker-Verlag, Aachen, Germany.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. & Argano R. 2009. New species of terrestrial isopods from Sardinia (Isopoda: Oniscidea). Zootaxa 2318 (1): 38 - 55. https: // doi. org / 10.11646 / zootaxa. 2318.1.5","Taiti S. & Argano R. 2011. Oniscidea di Sardegna (Crustacea, Isopoda). Conservazione Habitat Invertebrati 5: 163 - 222.","Schmidt C. 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2 (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Zoologischen Museum in Berlin, Zoologische Reihe 79: 3 - 179. https: // doi. org / 10.1002 / mmnz. 20030790102","Schmalfuss H. 1984. Eco-morphological strategies in terrestrial isopods. Proceedings of the Zoological Society of London 53: 49 - 63."]}

Tylos marcuzzii Giordani Soika, 1954 Figs 1, 2A Tylos marcuzzii Giordani Soika, 1954: 79, fig. 10/1. Additional references Schultz (1974); Kensley & Schotte (1989). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♀; Playa Libre; 10��10���52.4��� N, 75��43���56.1��� W; 7 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio- D��az leg.; CUDC-CRU 136 ��� 11 ♂♂, 39 ♀♀, 1 juv.; La Punta, Hotel Majagua; 10��10���56.41��� N, 75��43���33.51��� W; 6 Oct. 2017; same collectors as for preceding; CUDC-CRU 137. Distribution Florida, Bahamas, Leeward Islands, Belize, Venezuela and Cuba (Schultz 1974; Kensley & Schotte 1989; Schmalfuss & Vergara 2000; Schmidt 2001; Schmalfuss 2003; Hurtado et al. 2014). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 9, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83.","Schultz G. A. 1974. Terrestrial isopod crustaceans (Oniscoidea) mainly from the West Indies and adjacent regions: 1. Tylos and Ligia. Studies on the fauna of Cura ҫao and other Caribbean islands 149: 162 - 173. Available from http: // www. repository. naturalis. nl / document / 549820 [accessed 7 Jan. 2022].","Kensley B. & Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022].","Schmalfuss H. & Vergara K. 2000. The isopod genus Tylos (Oniscidea: Tylidae) in Chile, with bibliographies of all described species of the genus. Stuttgarter Beitrage zur Naturkunde A 612: 1 - 42.","Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Hurtado L. A., Lee E. J., Mateos M. & Taiti S. 2014. Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PloS ONE 9 (4): e 94081. https: // doi. org / 10.1371 / journal. pone. 0094081"]}

Littorophiloscia amphindica Taiti & Ferrara, 1986 Figs 1, 2H Littorophiloscia amphindica Taiti & Ferrara, 1986: 1369, fig. 16. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 5 ♂♂, 4 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96��� W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 223 ��� 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���34��� N, 75��44���26.8��� W; 25 Aug. 2017; leg. Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja-Arrieta; CUDC- CRU 224 ��� 1 ♂, 4 ♀♀; Laguna La Charca; 10��10���47.8��� N, 75��44���0.4��� W; 7 Sep. 2017; same collectors as for preceding; CUDC-CRU 225 ��� 2 ♂♂, 3 ♀♀; Hotel La Pola; 10��10���19.4��� N, 75��44���48.9��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 226 ��� 1 ♂, 1 ♀; Ca��o Rat��n; 10��10���42.6��� N, 75��44���52.6��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 227 ��� 1 ♂, 1 ♀; 10��10���33.4��� N, 75��44���52.4��� W; 24 Nov. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC- CRU 228 ��� 1 ♂, 3 ♀♀; same collection data as for preceding; 10��10���44.91��� N, 75��44���53.01��� W; CUDC- CRU 229 ��� 1 ♂, 1 ♀; La Punta, Laguna Encantada; 10��10���48.3��� N, 75��43���38.9��� W; 25 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 230 ��� 5 ♂♂, 6 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���39.1��� W; CUDC-CRU 231 ��� 9 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; CUDC-CRU 232 ��� 1 ♀; same collection data as for preceding; 10��10���48.4��� N, 75��43���38.4��� W; CUDC-CRU 233 ��� 1 ♀; same collection data as for preceding; 10��10���48��� N, 75��43���39.3��� W; CUDC-CRU 234 ��� 1 ♂, 5 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; 7 Sep. 2017; CUDC-CRU 235 ��� 3 ♂♂, 1 ♀; Para��so Secreto; 10��10���15.9���N, 75��44���38.8��� W; 24 Nov.2017; C.M. L��pez-Orozco leg.; CUDC-CRU236 ��� 13 ♂♂, 13 ♀♀, 8 juvs; same locality and collector as for preceding, 6 Sep. 2017; CUDC-CRU 237 ��� 14 ♂♂, 32 ♀♀, 3 juvs; same locality as for preceding; 4Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 238. Distribution Tanzania (Zanz��bar), Maldives, Comoro Islands, Indonesia (Bali) and Taiwan (Schmalfuss 2003; Taiti 2014). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 34, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. 2014. The terrestrial Isopoda (Crustacea, Oniscidea) of the Maldives. Tropical Zoology 27 (1): 9 - 33. https: // doi. org / 10.1080 / 03946975.2014.894397"]}

Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. urn:lsid:zoobank.org:act: 88600E5C-BFF9-49BA-976C-8853E33A6E58 Figs 1, 2E, 10���12 Etymology The new species is named after Mr Luis Felipe Arrieta Z����iga, for his interest in nature and support to Ricardo Borja-Arrieta as a biologist. Material examined Holotype COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� ♂; Laguna del Silencio, El Silencio; 10��10���35.4��� N, 75��44���26��� W; 24 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 193. Paratypes COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 194 ��� 10 ♂♂, 5 ♀♀; same collection data as for holotype; CUDC-CRU 195 ��� 1 ♀; 10��10���32.9��� N, 75��44���27.3��� W; same date and collectors as for holotype; CUDC-CRU 206 ��� 1 ♀; 10��10���33.5��� N, 75��44���26.9��� W; same date and collectors as for holotype; CUDC-CRU 207 ��� 1 ♂; 10��10���35��� N, 75��44���26.4��� W; same date and collectors as for holotype; CUDC-CRU 208 ��� 1 ♂; 10��10���34.8��� N, 75��44���26.6��� W; same date and collectors as for holotype; CUDC-CRU 209 ��� 6 ♂♂, 12 ♀♀; same collection data as for holotype; 10��10���35��� N, 75��44���16.4��� W; CUDC-CRU 210 ��� 14 ♂♂, 10 ♀♀; 10��10���34��� N, 75��44���26.8��� W; 25 Aug. 2017; same locality and collectors as for holotype; CUDC-CRU 211 ��� 3 ♂♂; La Punta, Laguna Encantada; 10��10���48.4��� N, 75��43���38.4��� W; 25 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja- Arrieta leg.; CUDC-CRU 196 ��� 1 ♂; same locality as for preceding; 10��10���47.72��� N, 75��43���40.09��� W; same date and collectors as for holotype; CUDC-CRU 197 ��� 1 ♂, 1 ♀; same locality as for preceding; 10��10���48��� N, 75��43���39.3��� W; same date and collectors as for holotype; CUDC-CRU 198 ��� 1 ♀; same locality as for preceding; 10��10���47.7��� N, 75��43���39.93��� W; same date and collectors as for holotype; CUDC-CRU 199 ��� 2 ♂♂, 3 ♀♀; same locality as for preceding; 10��10���48.3��� N, 75��43���38.9��� W; same date and collectors as for holotype; CUDC-CRU 200 ��� 3 ♀♀; same locality as for preceding; 10��10���48.1��� N, 75��43���39.1��� W; same date and collectors as for holotype; CUDC-CRU 201 ��� 2 ♀♀; same locality as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; same date and collectors as for holotype; CUDC-CRU 202 ��� 4 ♂♂, 4 ♀♀; same locality as for preceding; 10��10���47.72��� N, 75��43���40.09��� W; same date and collectors as for holotype; CUDC-CRU 203 ��� 2 ♂♂, 2 ♀♀; same locality as for preceding; 7 Sep. 2017; same date and collectors as for holotype; CUDC-CRU 204 ��� 3 ♂♂, 4 ♀♀; Laguna La Charca; 10��10���47.8��� N, 75��44���0.4��� W; 7 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC- CRU 205. Description MEASUREMENTS. Maximum length: male 2.5 mm; female 3.1 mm. BODY. Color brown with usual pale muscle spots; antennae brown with pale flagellum (Fig. 2E). Body elongated and elliptical-shaped (Fig. 10A, H). Dorsal surface bearing elongated scale-setae (Fig. 10B). Dorsum covered with broad tubercles (Fig. 10A, C���E, H): cephalon bearing seven tubercles in two rows, anterior row with four and posterior row with three; ♂ pereonites 1���3 with one row of 10 tubercles, 4���7 with one row of eight tubercles; ♀ pereonites 1, 5 and 6 with one row of 10 tubercles, 2���4 and 7 with one row of eight tubercles; ♂ pleonites 1���5 and telson with two paramedian tubercles, pleonite 2 with two broad tubercles at sides, pleonite 3 with two small additional tubercles; ♀ pleonites 1���5 and telson with two paramedian tubercles, pleonites 2 and 3 with two additional small tubercles at sides. CEPHALON. Lateral lobes well developed and directed outwards, median lobe triangular, slightly surpassing distal margin of lateral lobes, frontal and suprantennal lines absent; eyes consisting of four ommatidia (Fig. 10C���D). PLEON. Outline continuous with that of pereonite 7, epimera of pleonites 3���5 rectangular (Fig. 10A, E, H). Telson (Fig. 10E) triangular, lateral margins almost straight, distal part truncate and not surpassing distal margin of pleonite 5 epimera. ANTENNULA. Composed of two articles, second article bearing a few lateral setae, distal margin bearing one flagellar seta and two aesthetascs (Fig. 10F). ANTENNA. When extended posteriorly slightly surpassing pereonite 1, flagellum of four articles (Fig. 10G). MOUTH. Mandibles with molar penicil of several plumose setae, left mandible with 2+1 penicils (Fig. 11A), right with 1+1 penicils (Fig. 11B). Maxillula (Fig. 11C) inner endite with two penicils; outer endite of 4+8 teeth, five of them pectinated, and one slender stalk between outer and inner set, outer distal margin bearing fringe of long thin setae. Maxilla (Fig. 11D) with lobes fused, inner middle portion with five long setae, distal margin rounded, bearing several thin and thick setae. Maxilliped (Fig. 11E) palp bearing two tufts of setae on first article, distal articles of palp fused, bearing several thin and thick setae; endite rectangular, inner margin bearing fringe of long thin setae plus four long thick setae, distal margin bilobed, bearing broad rounded penicil with long thick setae. UROPOD. Protopod enlarged with distal margin rounded, exopod not surpassing protopod, endopod longer than exopod (Fig. 11F). PEREOPODS. Pereopods 1���7 stout, merus to propodus bearing sparse setae on sternal margin, carpus 1 with strong distal seta bearing long sensilla, dactylus with very long ungual seta, dactylar seta elongated, apically cleft and plumose. Male PEREOPODS 1 AND 7. Without any sexual modifications (Fig. 12A���B). GENITAL PAPILLA. Ventral shield triangular, papilla slightly cleft at apex, bearing small setae (Fig. 12C). PLEOPODS. Pleopod 1 (Fig. 12D) exopod ovoid; endopod three times as long as exopod, apical portion bent outwards. Pleopod 2 (Fig. 12E) exopod subtriangular; endopod with distal article flagelliform. Exopods of pleopods 3���5 as in Fig. 12F���H. Remarks In the Caribbean region, four species of Armadilloniscus have been recorded: A. caraibicus from Venezuela, A. ellipticus (Harger, 1878) from the Atlantic coasts of USA and Bermuda, A. ninae Schultz, 1984 from Belize, and A. steptus Schotte & Heard, 1991 from Turks and Caicos Islands (British West Indies) (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schotte & Heard 1991). For a definition of the genus see Arcangeli (1957), Vandel (1962), Taiti & Ferrara (1989), and Schmidt (2002). Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. is readily distinguished from all the other species of the genus in the number and arrangement of the dorsal tubercles on the cephalon, pereon and pleon. Moreover, it can be distinguished from the other four Caribbean species in having two broad tubercles on pleonite 2 (vs small in A. caraibicus and A. steptus, absent in A. ninae), the cephalon with the median lobe triangular (vs rounded in A. caraibicus) and lateral lobes subquadrangular (vs subtriangular in A. ninae and A. steptus), eyes composed of four ommatidia (vs 4���5 in A. caraibicus, 6 in A. ellipticus and 5 in A. ninae), telson with distal margin truncate (vs rounded in A. caraibicus and A. ellipticus, triangular in A. ninae), and ♂ pleopod 1 exopod ovoid (vs subtriangular in A. caraibicus, A. ellipticus and A. ninae, almost rounded in A. steptus) (Paoletti & Stinner 1989; Schultz 1984; Schotte & Heard 1991; Garthwaite et al. 1992; Schmidt 2002). Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 21-26, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Schotte M. & Heard R. W. 1991. Studies on the Crustacea of the Turks and Caicos Islands, British West Indies. II. Armadilloniscus steptus, n. sp. (Isopoda: Oniscidea: Scyphacidae) from Pine Cay. Gulf Research Reports 8 (3): 247 - 250. https: // doi. org / 10.18785 / grr. 0803.04","Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Taiti S. & Ferrara F. 1989. New species and records of Armadilloniscus Uljanin 1875 (Crustacea Isopoda Oniscidea) from the coasts of the Indian and Pacific oceans. Tropical Zoology 2: 59 - 88. https: // doi. org / 10.1080 / 03946975.1989.10539427","Arcangeli A. 1957. Il genere Armadilloniscus Ulj. e gli Scyphacidae (crostacei isopodi terrestri). Atti della reale Accademia delle Scienze di Torino 91: 399 - 428.","Vandel A. 1962. Isopodes terrestres (Deuxieme Partie). Faune de France 66: 417 - 931.","Schmidt C. 2002. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 1 (Olibrinidae to Scyphacidae s. str.). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 78: 275 - 352. https: // doi. org / 10.1002 / mmnz. 20020780207","Garthwaite R., Lawson R. & Taiti S. 1992. Morphological and genetical relationships among four species of Armadilloniscus Uljanin, 1875 (Isopoda: Oniscidea: Scyphacidae). Journal of Natural History 26: 327 - 338. https: // doi. org / 10.1080 / 00222939200770171"]}

Genus Halophiloscia Verhoeff, 1908 Microphiloscia Vandel, 1973. Syn. nov., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 29, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Vandel A. 1973. Les isopodes terrestres et cavernicoles de l'ile de Cuba. In: Orghidan T., Nunez A., Botosaneanu L., Decou V., Negrea S. & Vina N. (eds) Resultats des Expeditions biospeologiques cubanoroumaines a Cuba, vol. 1: 153 - 188. Editura Academiei Republicii Socialiste Romania, Bucharest."]}

Armadilloniscus caraibicus Paoletti & Stinner, 1989 Figs 1, 2D, 9 Armadilloniscus caraibicus Paoletti & Stinner, 1989: 73, figs 6���12. Additional reference Schmidt (2002). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 2 ♀♀; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 189 ��� 4 ♂♂, 10 ♀♀; same locality as for preceding; 6 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 190 ��� 1 ♀; Ca��o Rat��n; 10��10���33.4��� N, 75��44���52.4��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 191 ��� 2 ♀♀; 10��10���42.6��� N, 75��44���52.6��� W; same collection data as for preceding; CUDC-CRU 192. Remarks Paoletti & Stinner (1989) described A. caraibicus from Parque Marrocoy, Falc��n State, Venezuela. In the description, no sexual dimorphism was noted in the dorsal ornamentation. However, the male specimens show four dorsal tubercles on pleonite 2, while females have only two (see Fig. 9 and Paoletti & Stinner 1989: fig. 11). Distribution Venezuela: Coralline cays of the Parque Natural Morrocoy (Paoletti & Stinner 1989), and north coast of the Pen��nsula de Paria (Schmidt 2001). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 19-20, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Schmidt C. 2002. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 1 (Olibrinidae to Scyphacidae s. str.). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 78: 275 - 352. https: // doi. org / 10.1002 / mmnz. 20020780207","Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12."]}

Ligia baudiniana Milne-Edwards, 1840 Fig. 1 Ligia baudiniana Milne-Edwards, 1840: 155. Ligia (Hirtiligia) baudiniana ��� Van Name 1936: 58, figs 5a, 14. Ligia baudiniana ��� R��os & Ramos 1990: 93, fig. 7. ��� Schmalfuss 2003: 124. ��� Lazarus-Agudelo & Cantera-Kintz 2007: 226. ��� L��pez-Orozco et al. 2014: 196, figs 1���3. Additional references Jackson (1922); Schultz (1984); Kensley & Schotte (1989); Leistikow (1997); Leistikow & W��gele (1999); Espinosa-P��rez & Hendrickx (2001); Santamaria et al. (2014). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���34.8��� N, 75��44���26.6��� W; 24 Aug. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 141 ��� 3 ♂♂, 2 ♀♀; same collection data as for preceding; 10��10���35��� N, 75��44���26.4��� W; CUDC-CRU 142 ��� 1 juv.; same collection data as for preceding; 10��10���35.4��� N, 75��44���26.2��� W; CUDC-CRU 143 ��� 1 ♂, 1 ♀; same collection data as for preceding; 10��10���34.3��� N, 75��44���26.7��� W; CUDC-CRU 144 ��� 1 ♂; same locality and collectors as for preceding; 10��10���32.9��� N, 75��44���27.3��� W; 25 Aug. 2017; CUDC-CRU 145 ��� 1 ♀; same collection data as for preceding; 10��10���32.3��� N, 75��44���27.2��� W; CUDC-CRU 146 ��� 1 ♂; same collection data as for preceding; 10��10���32.5��� N, 75��44���27.1��� W; CUDC-CRU 147 ��� 1 ♀; Ca��o Rat��n, Laguna Palmar; 10��10���24.5��� N, 75��44���39.7��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 148 ��� 2 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���35.4��� N, 75��44���26��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 149 ��� 1 ♀; same locality and collectors as for preceding; 10��10���48��� N, 75��43���39.3��� W; 25 Aug. 2017; CUDC-CRU 150 ��� 1 ♂; same collection data as for preceding; 10��10���34��� N, 75��44���26.8��� W; CUDC-CRU 151 ��� 1 juv.; same collection data as for preceding; 10��10���47.74��� N, 75��43���39.93��� W; CUDC-CRU 152 ��� 1 ♂, 1 ♀; same collection data as for preceding; 10��10���35��� N, 75��44���26.9��� W; ICN- CR-is 260 ��� 1 ♂, 2 ♀♀; Ca��o Rat��n; 10��10���44.91��� N, 75��44���53.01��� W; 24 Nov. 2017, same collectors as for preceding; ICN-CR-is 261 ��� 1 juv.; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018, C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 222. Previous records Valle del Cauca: Bah��a M��laga (Pacific); Bol��var: Cartagena de Indias (Caribbean); Magdalena: Santa Marta (R��os & Ramos 1990; Lazarus-Agudelo & Cantera-Kintz 2007; L��pez-Orozco et al. 2014; Santamaria et al. 2014). Distribution This species was originally described from Veracruz, Mexico (but see Discussion). Molecular phylogenetic analyses, however, revealed that what has been denominated as this species corresponds to a clade composed of highly divergent lineages (probably a complex of cryptic species), distributed in southern Florida, across the Caribbean Sea, Bahamas, Bermuda, and probably in northern Brazil, and the Pacific coast from Costa Rica, Panam��, Colombia, probably to Ecuador (Santamaria et al. 2014). Although this species has been reported in the Gulf of California and California, extensive sampling for phylogeographic studies conducted by some of us suggest that these previous records are likely misidentifications (Hurtado et al. 2010; Eberl et al. 2013; Hurtado et al. 2018)., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 7, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Milne-Edwards M. 1840. Ordre des isopodes. In: Histoire Naturelle des Crustaces, comprenant l'Anatomie, la Physiologie et la Classficiation de ces Animaux. Tome Troisieme. 115 - 283. Librairie Encyclopedique de Roret, Paris.","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Rios R. & Ramos G. E. 1990. Los isopodos (Crustacea: Isopoda) de Bahia Malaga, Colombia. Revista de Ciencias, Universidad del Valle 2: 83 - 96.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Lazarus-Agudelo J. F. & Cantera-Kintz J. R. 2007. Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico Colombiano). Biota Colombiana 8 (2): 221 - 239.","Lopez-Orozco C. M., Bermudez A. & Navas G. R. 2014. Primer registro de Ligia baudiniana (Crustacea: Isopoda: Oniscidea) para el Caribe Colombiano. Boletin de Investigaciones marinas y costeras 43 (1): 195 - 200. https: // doi. org / 10.25268 / bimc. invemar. 2014.43.1.41","Jackson H. G. 1922. A revision of the isopod genus Ligia (Fabricius). Proceedings of the Zoological Society of London 92 (3): 683 - 703. https: // doi. org / 10.1111 / j. 1096 - 3642.1922. tb 02164. x","Kensley B. & Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022].","Leistikow A. 1997. Terrestrial isopods from Costa Rica and a redescription of Ischioscia variegata (Dollfus, 1893) (Crustacea: Isopoda: Oniscidea). Canadian Journal of Zoology 75 (9): 1415 - 1464. https: // doi. org / 10.1139 / z 97 - 768","Leistikow A. & Wagele J. W. 1999. Checklist of terrestrial isopods of the New World (Crustacea, Isopoda, Oniscidea). Revista brasileira de Zoologia 16: 1 - 72. https: // doi. org / 10.1590 / S 0101 - 81751999000100001","Espinosa-Perez M. C. & Hendrickx M. E. 2001. Checklist of isopods (Crustacea: Peracarida: Isopoda) from the Eastern Tropical Pacific. Belgian Journal of Zoology 131 (1): 43 - 55.","Santamaria C. A., Mateos M. C. & Hurtado L. A. 2014. Diversification at the narrow sea-land interface in the Caribbean: phylogeography of endemic supralittoral Ligia isopods. Frontiers in Ecology and Evolution 2: e 42. https: // doi. org / 10.3389 / fevo. 2014.00042","Hurtado L. A., Mateos M. & Santamaria C. A. 2010. Phylogeography of supralittoral rocky intertidal Ligia isopods in the Pacific region from Central California to Central Mexico. PLoS ONE 5 (7): e 11633. https: // doi. org / 10.1371 / journal. pone. 0011633","Eberl R., Mateos M., Grosberg R. K., Santamaria C. A. & Hurtado L. A. 2013. Phylogeography of the supralittoral isopod Ligia occidentalis around the Point Conception marine biogeographic boundary. Journal of Biogeography 40: 2361 - 2372. https: // doi. org / 10.1111 / jbi. 12168","Hurtado L. A., Mateos M., Wang C., Santamaria C. A., Jung J., Khalaji-Pirbalouty V. & Kim W. 2018. Out of Asia: mitochondrial evolutionary history of the globally introduced supralittoral isopod Ligia exotica. PeerJ 6: e 4337. https: // doi. org / 10.7717 / peerj. 4337"]}

Littorophiloscia denticulata (Ferrara & Taiti, 1982) Figs 1, 2I Bilawrencia denticulata Ferrara & Taiti, 1982: 469, figs 6���7. Additional reference Leistikow (2000). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂, 2 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96������ W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja- Arrieta leg.; CUDC-CRU 239 ��� 3 ♂♂, 3 ♀♀; Para��so Secreto 10��10���15.9��� N, 75��44���38.8��� W; 6 Sep. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 240 ��� 13 ♂♂, 18 ♀♀; same locality as for preceding, 4 Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 241. Distribution Andaman Islands, Brazil and Guatemala (Schmalfuss 2003; Lisboa et al. 2017). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 34-35, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Ferrara F. & Taiti S. 1982. Isopodi terrestri delle Isole Andamane. Bollettino del Museo civico di Storia naturale di Verona 8: 459 - 492.","Leistikow A. 2000. Terrestrial Isopoda from Guatemala and Mexico (Crustacea: Oniscidae: Crinocheta). Revue suisse de Zoologie 107: 283 - 323. https: // doi. org / 10.5962 / bhl. part. 80131","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Lisboa J. T., Campos-Filho I. S., Couto E. C. G. & Araujo P. B. 2017. Distribution of terrestrial isopods of the genus Littorophiloscia (Isopoda, Halophilosciidae) along the Brazilian coast. North-Western Journal of Zoology 13: e 167301."]}

Stenoniscus nestori L��pez-Orozco, Taiti & Campos-Filho sp. nov. urn:lsid:zoobank.org:act: 045F7904-E6DE-4912-9073-1473A522FAC7 Figs 1, 2C, 6���8 Etymology The new species is named after Dr Nestor Hernando Campos, for his contribution to the knowledge of Crustacea from Colombia. Material examined Holotype COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� ♂ (parts in micropreparations); La Punta, Laguna Encantada; 10��10���48.1��� N, 75��43���37.5��� W; 7 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 185. Paratypes COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 186 ��� 2 ♀♀; same collection data as for holotype; CUDC-CRU 187 ��� 1 ♀; same locality and collectors as for holotype; 24 Aug. 2017; CUDC-CRU 188. Description MEASUREMENTS. Maximum body length: male 1.8 mm; female 2.4 mm. BODY. Body pigments absent (Fig. 2C). Body (Fig. 6A���C) slender, about four times as long as wide, parallel sides. Dorsal surface bearing ribs and tubercles (Fig. 6A���C, E���G), as follows: cephalon with two tubercles on anterior portion and three on posterior portion; pereonite 1 with two transversal ribs; pereonites 1���4 with three longitudinal ribs (middle rib split in two on pereonites 2 and 3), one lateral tubercle and one tubercle between two lateral ribs per side; pereonites 5���7 with two longitudinal ribs and one lateral tubercle per side; pleonites 2���5 and telson with two paramedian tubercles. Dorsum bearing rectangular scale-setae with fringed apex (Fig. 6D). CEPHALON. Sub-rectangular (Fig. 6E���F), median lobe triangular, lateral lobes well-developed, directed frontwards, not surpassing median lobe; eyes absent. PEREON. Epimera of pereonites 2���7 laterally grooved (Fig. 6B). Pereonites 4���5 ventrally with lateral sternal thickenings small, triangular; pereonites 6 and 7 ventrally with sub-rectangular sternal plates (Fig. 6C). PLEON. Pleonite 2 dorsally visible, 3���5 with rectangular epimera (Fig. 6A���C). Telson (Fig. 6G) trapezoidal, distal margin broadly rounded. ANTENNULA. Composed of two articles (Fig. 6H), proximal article stout with shield-like projection on distal margin, distal article comma-shaped bearing two distal aesthetascs. ANTENNA. Short (Fig. 6I), peduncle articles more or less subequal in length; flagellum of two articles, distal article twice as long as proximal article, bearing two lateral aesthetascs. MOUTH. Mandibles with molar penicil consisting of two plumose setae, left mandible (Fig. 7A) with two penicils on incisor process, right mandible (Fig. 7B) with one penicil. Maxillula (Fig. 7C) inner endite with two short apical penicils; outer endite bearing 4+4 teeth, inner set with two teeth apically cleft. Maxilla (Fig. 7D) of two rounded lobes, inner lobe bearing three thick setae. Maxilliped (Fig. 7E) basis with sparse setae; endite stout, distal margin rounded, bearing thin setae, one large penicil inserted near medial margin. UROPOD. Protopod stout and conical, exopod short, bearing two long apical setae, endopod inserted proximally (Fig. 7F). PEREOPODS. Pereopods 1���7 short, stout, merus to propodus bearing sparse setae on sternal margins; pereopod 1 carpus with distal seta fringed at apex; dactylus of two claws, inner claw very short, ungual seta simple, not surpassing outer claw, dactylar seta stout, reaching distal margin of outer claw, apex bearing many small setae. Male PEREOPODS 1 AND 7. Without sexual dimorphism (Fig. 8A���B). PLEOPODS. Pleopod 1 exopod absent; endopod (Fig. 8C) enlarged on proximal portion, distal portion tapering. Pleopod 2 (Fig. 8D) exopod triangular, inner margin grooved, bearing one strong seta; endopod about twice as long as exopod. Pleopod 3 and 4 exopods (Fig. 8E���F) sub-quadrangular, inner margin grooved, distal margin slightly serrate. Pleopod 5 exopod (Fig. 8G) sub-rectangular, inner margin slightly serrate, distal margin straight, bearing one seta. Remarks Presently, Stenoniscus comprises four species (Schmalfuss 2003): S. pleonalis Aubert & Dollfus, 1890 from the northern coasts of the Mediterranean east to the Aegean, Bermuda, Bulgaria, Madeira and Mexico; S. carinatus Silvestri, 1897 from coastal areas of Croatia, Italy, Portugal and Spain; S. aenariensis (Verhoeff, 1942) and S. plutonis (Verhoeff, 1942) from Ischia Island, Italy, both most probably junior synonyms of S. pleonalis according to the short descriptions by Verhoeff (1942). The genus can be defined by animals of reduced size (up to 3.5 mm), body slender with lateral sides parallel, dorsum bearing longitudinal ribs and tubercles, sometimes reduced, epimera of pereonites 2���7 with lateral margins grooved, pereonites 4���5 ventrally with lateral sternal thickenings and 6���7 with sternal plates, antennula of two articles, pleonite 1 absent, uropods of conical shape, pleopod exopods without respiratory organs, and pleopod 1 absent on females (see Vandel 1962; Schmidt 2003). Stenoniscus nestori L��pez-Orozco, Taiti & Campos-Filho sp. nov. is morphologically similar to Stenoniscus carinatus, from which it differs mainly in the number and disposition of the dorsal ribs and tubercles on the cephalon and pereonites (compare Fig. 6A���B with Schmidt 2003: fig. 67 and Taiti & Ferrara 1982: fig. 6a for S. carinatus). The new species is readily distinct from Stenoniscus pleonalis in having a strong dorsal ornamentation (vs very weak) and lacking the tomentose appearance due to long dorsal scale-setae (see Vandel 1944: fig. 1a���c). Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 14-19, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Verhoeff K. W. 1942. Landisopoden der Insel Ischia, systematisch, morphologisch, phanologisch, okologisch, geographisch beurteilt. Zeitschrift fur Morphologie und Okologie der Tiere 38: 435 - 482.","Vandel A. 1962. Isopodes terrestres (Deuxieme Partie). Faune de France 66: 417 - 931.","Schmidt C. 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2 (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Zoologischen Museum in Berlin, Zoologische Reihe 79: 3 - 179. https: // doi. org / 10.1002 / mmnz. 20030790102","Ferrara F. & Taiti S. 1982. Isopodi terrestri delle Isole Andamane. Bollettino del Museo civico di Storia naturale di Verona 8: 459 - 492.","Vandel A. 1944. Isopodes terrestres recoltes par M. Remy au cours de son voyage en Corse. II. La famille de Stenoniscidae. Archives de Zoologie experimentale et generale 84: 23 - 47."]}

Trichorhina bermudezae Carpio-D��az, L��pez-Orozco & Campos-Filho, 2018 Fig. 1 Trichorhina bermudezae Carpio-D��az, L��pez-Orozco & Campos-Filho in Carpio-D��az et al., 2018: 307, figs 29���50. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂; La Punta (TDF); 10��10���48.42��� N, 75��43���41.36��� W; 8 Sep. 2017; Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja- Arrieta leg.; CUDC-CRU 267 ��� 1 ♂, 1 ♀; same collection data as for preceding; 10��10���47.36��� N, 75��43���42.5��� W; CUDC-CRU 268 ��� 4 ♂♂, 6 ♀♀; same collection data as for preceding; 10��10���47.55��� N, 75��43���42.09��� W; CUDC-CRU 269 ��� 2 ♂♂, 7 ♀♀; same collection data as for preceding; 10��10���46.79��� N, 75��43���43.51��� W; CUDC-CRU 270 ��� 2 ♂♂, 6 ♀♀; same collection data as for preceding; 10��10���48.02��� N, 75��43���41.58��� W; CUDC-CRU 271 ��� 1 ♂, 6 ♀♀; same collection data as for preceding; 10��10���47.74��� N, 75��43���41.8��� W; CUDC-CRU 272 ��� 1 ♂, 6 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; CUDC-CRU 273 ��� 1 ♂; La Punta (TDF); 10��10���47.76������ N, 75��43���40.27��� W; 25 Aug. 2017; Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 274 ��� 1 ♂, 1 ♀; La Punta, Casa Sra. Miriam; 10��10���46.91��� N, 75��43���46.73��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 275 ��� 2 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96��� W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC- CRU 276 ��� 4 ♂♂, 4 ♀♀, 3 juvs; El Silencio (TDF); 10��10���31.91��� N, 75��44���23.68��� W; 24 Aug. 2017; A. Figueroa and B. Valencia leg.; CUDC-CRU 277 ��� 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���32.3��� N, 75��44���27.2��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 278 ��� 7 ♂♂, 12 ♀♀; El Paraiso (TDF); 10��10���22.9��� N, 75��44���32.8��� W; 6 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 279 ��� 3 ♂♂; Ca��o Rat��n; 10��10���42.9��� N, 75��44���52.8��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 280. Previous records Botanical Garden ���Guillermo Pi��eres���, Turbaco, Bol��var (Carpio-D��az et al. 2018). Distribution Colombia. First record for continental islands of the Colombian Caribbean., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 35-36, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Carpio-Diaz Y. M., Lopez-Orozco C. M., Campos-Filho I. S. & Navas G. R. 2018. Terrestrial isopods (Isopoda: Oniscidea) of the Botanical Garden of Cartagena \" Guillermo Pineres \", Colombia, with the description of three new species. Arthropoda Selecta 27 (4): 301 - 318. https: // doi. org / 10.15298 / arthsel. 27.4.05"]}

Ctenorillo tuberosus (Budde-Lund, 1904) Figs 1, 2L Armadillo tuberosus Budde-Lund, 1904: 109, pl. x, figs 1���4. Additional reference Campos-Filho et al. (2017). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂; Ca��o Rat��n, Casa Comunal; 10��10���39.06��� N, 75��44���50.4��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta, Y.M. Carpio-D��az and A. Figueroa leg.; CUDC-CRU 255 ��� 1 ♂; Orika; 10��10���39��� N, 75��44���26.8��� W; 25Aug. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 256 ��� 2 ♂♂, 5 ♀♀; La Punta, Casa Sra. Miriam; 10��10���46.91��� N, 75��43���46.73��� W; 8 Sep. 2017; same collectors as for preceding; CUDC-CRU 257 ��� 4 ♂♂; El Silencio, Laguna del Silencio; 10��10���33.5��� N, 75��44���26.9��� W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 258 ��� 2 ♂♂; same collection data as for preceding; 10��10���32.9��� N, 75��44���27.3��� W; CUDC-CRU 259 ��� 3 ♂, 11 ♀♀; same collection data as for preceding; 10��10���34.3��� N, 75��44���26.7��� W; ICN-CR-is 266. Distribution Brazil and Haiti (Schmalfuss 2003; Campos-Filho et al. 2017). First record for Colombia. Molecular phylogenetic analyses of Tylos Audouin, 1826 Sequences obtained for the three species of Tylos from Isla Grande and the specimen from Florida have been deposited under GenBank Accession Numbers MW532964 ��� MW532970 and MW533069 ��� MW533075. Only the 12S rDNA sequence was obtained for the specimen from Isla Grande assigned to T. marcuzzii. Phylogenetic analyses using only 12S rDNA group this sequence with the two specimens of T. marcuzzii from Cuba reported in Hurtado et al. (2014); see Fig. 18. The 12S rDNA p-distance between T. marcuzzii from Isla Grande and T. marcuzzii from Cuba is 2.0���2.7%, whereas the distance between the specimens from the two Cuban localities is 2.0% (Table 2). Only the 16S rDNA sequence was obtained for the specimen of T. niveus from the Florida Keys, the type locality of this species. Phylogenetic analyses of 16S rDNA show that specimens identified as T. niveus from Isla Grande are most closely related to the T. niveus specimen from the Florida Keys. The 16S rDNA p-distance between T. niveus from Isla Grande and the Florida Keys is 2.7% (Table 2). The concatenated (12S rDNA and 16S rDNA) dataset was comprised of 43 taxa and 740 characters (excluding characters with ambiguous homology), 236 of which were parsimony-informative. The specimens assigned to T. marcuzzii were used as outgroup. Five main lineages were identified for the ingroup taxa in the concatenated phylogenetic analyses, which formed a basal polytomy (Fig. 18): (1) T. niveus from Puerto Rico; (2) Tylos sp. from Yaguanabo, Cuba; (3) T. niveus from Florida and Isla Grande; (4) T. negroi sp. nov.; and (5) Tylos punctatus sensu lato from the Northeastern Pacific. The range of 16S rDNA and 12S rDNA p-distances between the four basal lineages excluding T. punctatus is 11.1���13.6% and 9.7���15.5%, respectively. The range of 16S rDNA and 12S rDNA p-distances between these ingroup lineages and T. marcuzzii is 19.4���22.2% and 19.5���23.8%, respectively., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 38, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1904. A Revision of Crustacea Isopoda Terrestria, with Additions and Illustrations. Pt 1 Eubelum, Pt 2 Spherilloninae, Pt 3 Armadillo. H. Hagerup, Copenhagen, Denmark.","Campos-Filho I. S., Montesanto G., Araujo P. B. & Taiti S. 2017. New species and new records of terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Brazil. Iheringia, Serie Zoologia 107: e 2017034. https: // doi. org / 10.1590 / 1678 - 4766 e 2017034","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Hurtado L. A., Lee E. J., Mateos M. & Taiti S. 2014. Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PloS ONE 9 (4): e 94081. https: // doi. org / 10.1371 / journal. pone. 0094081"]}

Littorophiloscia tropicalis Taiti & Ferrara, 1986 Figs 1, 2J Littorophiloscia tropicalis Taiti & Ferrara, 1986: 1361, fig. 9. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� many ♂♂ and ♀♀; Playa Libre; 10��10���52.3��� N, 75��43���55.7��� W; 7 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 242 ��� 1 ♂, 6 ♀♀; same locality as for preceding; 10��10���52.3��� N, 75��43���54.8��� W; 5 Oct. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 243 ��� 2 ♂♂; La Punta, Hotel Majagua; 10��10���56.41��� N, 75��43���33.51��� W; 6 Oct. 2017; C.M. L��pez-Orozco and L. Vides leg.; CUDC- CRU 244 ��� 1 ♂, 2 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96��� W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 245 ��� 1 ♂, 2 ♀♀, 1 juv.; El Silencio, Playa La Cocotera; 10��10���49.3��� N, 75��44���16.7��� W; 6 Oct. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 246 ��� 1 ♂, 2 ♀♀; Ca��o Rat��n; 10��10���44.91��� N, 75��44���53.01��� W; 24 Nov. 2017; C.M. L��pez-Orozco, R. Borja- Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 247 ��� 2 ♂♂, 2 ♀♀; same locality, date and collectors; as for preceding; 10��10���33.4��� N, 75��44���52.4��� W; CUDC-CRU 248 ��� 1 ♂, 3 ♀♀; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; ICN-CR-is 265. Distribution Circumtropical (Schmalfuss 2003; Taiti & Ferrara 2004; Taiti & Checcucci 2009; Taiti 2014; Lisboa et al. 2017). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 35, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. & Ferrara F. 2004. The terrestrial Isopoda (Crustacea: Oniscidea) of the Socotra Archipelago. Fauna of Arabia 20: 211 - 326.","Taiti S. & Checcucci I. 2009. New species and records of terrestrial Isopoda (Crustacea, Oniscidea) from Socotra Island, Yemen. ZooKeys 31: 73 - 103. https: // doi. org / 10.3897 / zookeys. 31.140","Taiti S. 2014. The terrestrial Isopoda (Crustacea, Oniscidea) of the Maldives. Tropical Zoology 27 (1): 9 - 33. https: // doi. org / 10.1080 / 03946975.2014.894397","Lisboa J. T., Campos-Filho I. S., Couto E. C. G. & Araujo P. B. 2017. Distribution of terrestrial isopods of the genus Littorophiloscia (Isopoda, Halophilosciidae) along the Brazilian coast. North-Western Journal of Zoology 13: e 167301."]}

Tylos negroi L��pez-Orozco, Carpio-D��az & Campos-Filho sp. nov. urn:lsid:zoobank.org:act: DFCDBD2C-AEBD-4754-951A-1D927E81E6DC Figs 1, 2B, 3���5 Etymology The new species is named after Mr Hernando G��mez Molina, also known as Mr Negro, for his efforts to conserve the biodiversity of the Rosario Islands. Material examined Holotype COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� ♂; Playa Libre; 10��10���52.3��� N, 75��43���54.8��� W; 5 Oct. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio- D��az leg.; CUDC-CRU 127. Paratypes COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 128 ��� 2 ♀♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 129 ��� many ♂♂ and ♀♀; same collection data as for holotype; CUDC-CRU 130 ��� 7 ♂♂, 13 ♀♀; El Silencio, Playa La Cocotera; 10��10���49.3��� N, 75��44���16.7��� W; 6 Oct. 2017; same collectors as for holotype; CUDC-CRU 131 ��� 8 ♂♂, 10 ♀♀; La Punta, El Terminal; 10��10���55.75��� N, 75��43���36.57��� W; same collectors as for holotype; CUDC-CRU 132 ��� 5 ♀♀; Playa Libre; 10��10���52.3��� N, 75��43���55.7��� W; 7 Sep. 2017; same collectors as for holotype; CUDC-CRU 133 ��� 1 ♀; same collection data as for holotype; CUDC-CRU 134 ��� 4 ♂♂, 3 ♀♀; same collection data as for holotype; 5 Oct. 2017; ICN-CR-is 262 ��� 1 ♂; La Punta, Hotel Cocoliso; 10��10���42.19��� N, 75��43���32.88��� W; 6 Oct. 2017; same collectors as for holotype; CUDC-CRU 135. Description MEASUREMENTS. Maximum body length: male 10 mm; female 8.1 mm. BODY. Colour yellowish with many irregular dark spots (Fig. 2B). Body outline as in Fig. 3A. Endoantennal conglobation. Dorsum covered with lanceolate scale-setae (Fig. 3B). CEPHALON. Frontal lamina triangular (Fig. 3C���D); eyes consisting of 26���27 ommatidia. PEREON. Pereonite 1 epimera grooved on lateral margins, schisma with inner lobe triangular, slightly surpassing outer lobe (Fig. 3E); pereonite 2 with ventral lobe triangular, shorter than outer lobe (Fig. 3E); epimera of pereonites 2���4 subtriangular, 5���7 subquadrangular (Fig. 3A). PLEON. Epimera of pleonites 3���5 with rounded distal margin (Fig. 3F). Ventral phylacomera present on pleonites 3���5; phylacomera 5 oar-shaped, with wide distal part, medial margin oblique and slightly sinuous (Fig. 3G). Telson rectangular, posterior margin slightly folded upwards (Fig. 3F). ANTENNULA. Composed of one article, flattened and triangular, several aesthetascs inserted on slightly rounded lobe (Fig. 3H). ANTENNA. Short and thickset, flagellum of three articles, apical article as long as first and second articles (Fig. 4A). MOUTH. Mandibles with strong molar process, left mandible (Fig 4B) with two penicils on molar process, two on hairy lobe and five free penicils; right mandible (Fig. 4C) with two penicils on molar process, one on hairy lobe and five free penicils. Maxillula (Fig. 4D) inner endite with three long penicils densely covered with thin setae, apical penicil longest; outer endite composed of 5+5 simple teeth plus one sensory seta, inner set bearing two teeth slightly serrate, median portion bearing three short setae. Maxilla (Fig. 4E) of two lobes, inner lobe rounded, covered with thick setae, outer lobe covered with thin setae. Maxilliped (Fig. 4F) basis with rounded outer margin, palp proximal article triangular without setae; endite rectangular, bearing five apical and two subapical penicils. UROPOD. As in Fig 3G. PEREOPODS. Pereopod 1 (Fig. 5A) basis with triangular latero-apical process. Pereopod 7 (Fig. 5B) propodus slightly inflated; dactylus of one claw, dactylar seta stout, not surpassing claw and bearing several small sensory setae on median to apical portion. Male PEREOPODS 1 AND 7. Without any sexual dimorphism (Fig. 5A���B). PLEOPODS. Pleopod 2 exopod rectangular (Fig. 5C) with respiratory folds; endopod with distal portion slightly directed inwards, apex covered with cleft scales of 2���4 branches (Fig. 5C). Remarks Presently, the genus Tylos has a worldwide distribution and comprises 21 coastal species (Schmalfuss & Vergara 2000; Schmalfuss 2003). In Colombia, only T. niveus Budde-Lund, 1885 was recorded from the Caribbean region (Carpio-D��az et al. 2016). Tylos negroi sp. nov. can be distinguished from all species of the genus by the shape of the ventral phylacomera 5. Moreover, it differs from T. niveus in the pereonite 1 epimeron having the inner lobe of the schisma surpassing the outer lobe (vs inner lobe not surpassing outer lobe) and the pereopod 7 propodus being inflated (vs not inflated). Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 11-13, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Schmalfuss H. & Vergara K. 2000. The isopod genus Tylos (Oniscidea: Tylidae) in Chile, with bibliographies of all described species of the genus. Stuttgarter Beitrage zur Naturkunde A 612: 1 - 42.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Carpio-Diaz Y. M., Lopez-Orozco C. M., Herrera-Medina Y., Navas G. R. & Bermudez A. 2016. Primer registro de Tylos niveus y nuevo reporte de Porcellionides pruinosus (Oniscidea: Tylidae y Porcellionidae) para Colombia. Revista de la Academia colombiana de Ciencias exactas fisicas y naturales 40 (156): 433 - 437. https: // doi. org / 10.18257 / raccefyn. 343"]}

Agnara madagascariensis (Budde-Lund, 1885) Figs 1, 2K Metoponorthus Madagascariensis Budde-Lund, 1885: 189. Additional references Schmalfuss & Ferrara (1978); Ferrara & Taiti (1986). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 5 ♂♂, 6 ♀♀; Ca��o Rat��n, Casa Comunal; 10��10���39.06��� N, 75��44���50.4��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja- Arrieta, Y.M. Carpio-D��az and A. Figueroa leg.; CUDC-CRU 138 ��� 2 ♀♀; Orika; 10��10���37.2��� N, 75��44���27.59��� W; 8 Sep. 2017; A. Figueroa leg.; CUDC-CRU 139 ��� 13 ♂♂, 49 ♀♀; La Punta, Casa Sra. Miriam; 10��10���46.91��� N, 75��43���46.73��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 140. Distribution Ascension Island, Senegal, Guinea Bissau, Arabian Peninsula, Madagascar, southern China, Taiwan and Venezuela (Ashmole & Ashmole 2000; Schmidt 2001; Schmalfuss 2003). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 37, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Schmalfuss H. & Ferrara F. 1978. Terrestrial isopods from West Africa, Part 2: families Tylidae, Ligiidae, Trichoniscidae, Styloniscidae, Rhyscotidae, Halophilosciidae, Philosciidae, Platyarthridae, Rhyscotidae, Trachelipidae, Porcellionidae, Armadillidiidae. Monitore zoologico italiano, nuova serie, Supplemento 11: 15 - 9. https: // doi. org / 10.1080 / 03749444.1978.10736575","Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Ashmole P. & Ashmole M. 2000. St Helena and Ascension Island: a Natural History. Anthony Nelson, Oswestry, England.","Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341."]}

Porcellionides pruinosus (Brandt, 1833) Fig. 1 Porcellio pruinosus Brandt, 1833: 19. Porcellionides pruinosus ��� Carpio-D��az et al. 2016: 434, figs 1���2. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂, 7 ♀♀; Orika; 10��10���37.2��� N, 75��44���27.59��� W; 8 Sep. 2017; A. Figueroa leg.; CUDC-CRU 249 ��� 2 ♀♀; El Silencio, Playa La Cocotera; 10��10���49.3��� N, 75��44���16.7��� W; 6 Oct. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 250 ��� 1 ♂, 11 ♀♀; El Silencio, Laguna del Silencio; 10��10���33.5��� N, 75��44���26.9��� W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 251 ��� 1 ♀; same collection data as for preceding; 10��10���32.9��� N, 75��44���27.3��� W; CUDC-CRU 252 ��� 18 ♂♂, 15 ♀♀, 1 juvenile; same collection data as for preceding; 10��10���35��� N, 75��44���26.4��� W; CUDC-CRU 253 ��� 3 ♀♀; La Punta, Laguna Encantada; 10��10���47.76��� N, 75��43���40.27��� W; 25 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio- D��az and R. Borja-Arrieta leg.; CUDC-CRU 254. Previous records San Andr��s (Van Name 1936); Angel��polis, Cafetal La Camelia Antioquia Dept, and near Nevado del Ru��z, Caldas Dept (Richardson 1912); Cartagena de Indias, Bar��, Bol��var Dept (Caribbean region) (Carpio-D��az et al. 2016). Distribution Cosmopolitan species (Schmalfuss 2003). First record for continental islands of the Colombian Caribbean., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 36-37, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Brandt J. F. 1833. Conspectus Monographiae Crustaceorum Oniscodorum Latreillii. Byulleten Moskovskogo Obshchestva Ispytatelei Prirody 6: 171 - 193.","Carpio-Diaz Y. M., Lopez-Orozco C. M., Herrera-Medina Y., Navas G. R. & Bermudez A. 2016. Primer registro de Tylos niveus y nuevo reporte de Porcellionides pruinosus (Oniscidea: Tylidae y Porcellionidae) para Colombia. Revista de la Academia colombiana de Ciencias exactas fisicas y naturales 40 (156): 433 - 437. https: // doi. org / 10.18257 / raccefyn. 343","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Richardson H. 1912. Terrestrial isopods of Colombia. Memoires de la Societe des Sciences naturelles Neuchatel 5: 29 - 32.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341."]}

Trichorhina bermudezae Carpio-Díaz, López-Orozco & Campos-Filho, 2018 Fig. 1 Trichorhina bermudezae Carpio-Díaz, López-Orozco & Campos-Filho in Carpio-Díaz et al., 2018: 307, figs 29–50. Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂; La Punta (TDF); 10°10′48.42″ N, 75°43′41.36″ W; 8 Sep. 2017; Y.M. Carpio-Díaz, C.M. López-Orozco and R. Borja- Arrieta leg.; CUDC-CRU 267 • 1 ♂, 1 ♀; same collection data as for preceding; 10°10′47.36″ N, 75°43′42.5″ W; CUDC-CRU 268 • 4 ♂♂, 6 ♀♀; same collection data as for preceding; 10°10′47.55″ N, 75°43′42.09″ W; CUDC-CRU 269 • 2 ♂♂, 7 ♀♀; same collection data as for preceding; 10°10′46.79″ N, 75°43′43.51″ W; CUDC-CRU 270 • 2 ♂♂, 6 ♀♀; same collection data as for preceding; 10°10′48.02″ N, 75°43′41.58″ W; CUDC-CRU 271 • 1 ♂, 6 ♀♀; same collection data as for preceding; 10°10′47.74″ N, 75°43′41.8″ W; CUDC-CRU 272 • 1 ♂, 6 ♀♀; same collection data as for preceding; 10°10′48.1″ N, 75°43′37.5″ W; CUDC-CRU 273 • 1 ♂; La Punta (TDF); 10°10′47.76″′ N, 75°43′40.27″ W; 25 Aug. 2017; Y.M. Carpio-Díaz, C.M. López-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 274 • 1 ♂, 1 ♀; La Punta, Casa Sra. Miriam; 10°10′46.91″ N, 75°43′46.73″ W; 8 Sep. 2017; C.M. López-Orozco, R. Borja-Arrieta and Y.M. Carpio-Díaz leg.; CUDC-CRU 275 • 2 ♀♀; El Silencio, Laguna Caracolí; 10°10′28.4″ N, 75°44′29.96″ W; 6 Oct. 2017; Y.M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC- CRU 276 • 4 ♂♂, 4 ♀♀, 3 juvs; El Silencio (TDF); 10°10′31.91″ N, 75°44′23.68″ W; 24 Aug. 2017; A. Figueroa and B. Valencia leg.; CUDC-CRU 277 • 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10°10′32.3″ N, 75°44′27.2″ W; 24 Nov. 2017; C.M. López-Orozco leg.; CUDC-CRU 278 • 7 ♂♂, 12 ♀♀; El Paraiso (TDF); 10°10′22.9″ N, 75°44′32.8″ W; 6 Sep. 2017; C.M. López-Orozco, R. Borja-Arrieta and Y.M. Carpio-Díaz leg.; CUDC-CRU 279 • 3 ♂♂; Caño Ratón; 10°10′42.9″ N, 75°44′52.8″ W; 24 Nov. 2017; C.M. López-Orozco leg.; CUDC-CRU 280. Previous records Botanical Garden “Guillermo Piñeres”, Turbaco, Bolívar (Carpio-Díaz et al. 2018). Distribution Colombia. First record for continental islands of the Colombian Caribbean.

Trichorhina heterophthalma Lemos de Castro, 1964 Fig. 1 Trichorhina heterophthalma Lemos de Castro, 1964: 2, figs 1���2. Trichorhina heterophthalma ��� Carpio-D��az et al. 2018: 303, fig. 3. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂, 9 ♀♀; La Punta, Casa Sra. Miriam; 10��10���46.91��� N, 75��43���46.73��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 260 ��� 3 ♀♀; Ca��o Rat��n; 10��10���42.9��� N, 75��44���52.8��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 261 ��� 3 ♀♀; El Silencio, Laguna del Silencio; 10��10���31.91��� N, 75��44���23.68��� W; 25 Aug. 2017; Alison Figueroa leg.; CUDC-CRU 262 ��� 1 ♂, 1 ♀; La Punta (TDF), Laguna Encantada; 10��10���47.36��� N, 75��43���42.5��� W; 8 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 263 ��� 2 ♂♂, 3 juvs; same locality, date and collectors as for preceding; 10��10���47.55��� N, 75��43���42.89��� W; CUDC-CRU 264 ��� 2 ♀♀; same locality and collectors as for preceding; 10��10���47.8��� N, 75��43���37.5��� W; 25 Aug. 2017; CUDC-CRU 265 ��� 2 ♂♂, 2 ♀♀; La Punta; 10��10���47.74��� N, 75��43���41.85��� W; 8 Sep. 2017, same collectors as for preceding; CUDC-CRU 266. Previous records Botanical Garden ���Guillermo Pi��eres���, Turbaco, Bol��var (Carpio-D��az et al. 2018). Distribution Pantropical (Schmalfuss 2003). First record for continental islands of the Colombian Caribbean., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 36, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Lemos de Castro A. 1964. Trichorhina heterophthalma nueva especie de isopodo terrestre cavernicola de Cuba. Poeyana 2 A: 1 - 7.","Carpio-Diaz Y. M., Lopez-Orozco C. M., Campos-Filho I. S. & Navas G. R. 2018. Terrestrial isopods (Isopoda: Oniscidea) of the Botanical Garden of Cartagena \" Guillermo Pineres \", Colombia, with the description of three new species. Arthropoda Selecta 27 (4): 301 - 318. https: // doi. org / 10.15298 / arthsel. 27.4.05","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341."]}

Tylos niveus Budde-Lund, 1885 Fig. 1 Tylos niveus Budde-Lund, 1885: 278. Tylos niveus ��� Carpio-D��az et al. 2016: 435, fig. 3. Additional references Boone (1934); Van Name (1936); Schultz (1970); Kensley & Schotte (1989); Silva & Alves (2000); Hurtado et al. (2014); Taiti et al. (2018). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂, 1 ♀; La Punta, Laguna Encantada; 10��10���48.1��� N, 75��43���37.5��� W; 24 Aug. 2017; Y.M. Carpio-D��az, C.M. L��pez- Orozco and R. Borja-Arrieta leg.; CUDC-CRU 153 ��� 4 ♂♂, 5 ♀♀; same locality and collectors as for preceding; 10��10���48.4��� N, 75��43���38.4��� W; 25 Aug. 2017; CUDC-CRU 154 ��� 2 ♂♂, 1 ♀; same collection data as for preceding; 10��10���48.3��� N, 75��43���38.9��� W; CUDC-CRU 155 ��� 5 ♂♂, 7 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���39.1��� W; CUDC-CRU 156 ��� 4 ♂♂, 1 ♀; same collection data as for preceding; 10��10���48��� N, 75��43���39.3��� W; CUDC-CRU 157 ��� 5 ♂♂, 2 ♀♀; same collection data as for preceding; 10��10���47.72��� N, 75��43���40.09��� W; CUDC-CRU 158 ��� 2 ♂♂, 1 ♀; same collection data as for preceding; 10��10���47.76��� N, 75��43���40.27��� W; CUDC-CRU 159 ��� 5 ♂♂, 2 ♀♀; same collection data as for preceding; 10��10���47.7��� N, 75��43���39.93��� W; CUDC-CRU 160 ��� 9 ♂♂, 17 ♀♀; El Silencio, Laguna del Silencio; 10��10���32.9��� N, 75��44���27.3��� W; 24 Aug. 2017, same collectors as for preceding; CUDC-CRU 161 ��� 3 ♂♂, 5 ♀♀; same collection data as for preceding; 10��10���35��� N, 75��44���26.4��� W; CUDC-CRU 162 ��� 10 ♂♂, 6 ♀♀; same locality and collectors as for preceding; 10��10���48.3��� N, 75��43���38.9��� W; 25 Aug. 2017; CUDC-CRU 163 ��� 15 ♂♂, 12 ♀♀; La Punta, El Terminal; 10��10���55.75��� N, 75��43���36.57��� W; 6 Oct. 2017; C.M. L��pez-Orozco leg.; ICN-CR-is 263 ��� 2 ♀♀; La Punta, Laguna Encantada; 10��10���47.8��� N, 75��43���39.5��� W; 25 Aug. 2017; Y.M. Carpio- D��az, C.M. L��pez-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 164 ��� 2 ♂♂, 2 ♀♀; La Punta, Hotel Cocoliso; 10��10���42.19��� N, 75��43���32.88��� W; 6 Oct. 2017; same collectors as for preceding; CUDC- CRU 165 ��� 4 ♀♀; Hotel La Pola; 10��10���19.4��� N, 75��44���48.9��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 166 ��� 1 ♂, 2 ♀♀; Ca��o Rat��n, Laguna Palmar; 10��10���24.5��� N, 75��44���39.7��� W; same date and collectors as for preceding; CUDC-CRU 167 ��� 4 ♂♂, 5 ♀♀; Playa Libre; 10��10���52.3��� N, 75��43���55.7��� W; same date as for preceding; C.M. L��pez-Orozco leg.; CUDC-CRU 168 ��� 3 ♂♂, 2 ♀♀; El Silencio, Laguna del Silencio; 10��10���34.3��� N, 75��44���26.7��� W; 24 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 169 ��� 1 ♀; El Silencio, Laguna del Silencio; 10��10���35��� N, 75��44���26.4��� W; 25 Aug. 2017; same collectors as for preceding; CUDC-CRU 170 ��� 3 ♂♂, 6 ♀♀, 1 juv.; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 171 ��� 1 ♂, 6 ♀♀; El Silencio, Laguna del Silencio; 10��10���31.9��� N, 75��44���26.9��� W; 7 Sep. 2017; same collector as for preceding; CUDC-CRU 172 ��� 4 ♂♂, 6 ♀♀; La Punta, Laguna Encantada; 10��10���48.1��� N, 75��43���37.5��� W; 7 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja- Arrieta leg.; CUDC-CRU 173 ��� 9 ♂♂, 17 ♀♀; El Silencio, Mirador del Silencio; 10��10���33.36��� N, 75��44���24.14��� W; 5 Oct. 2017; same collectors as for preceding; CUDC-CRU 174 ��� 4 ♂♂, 6 ♀♀, 1 juv.; Ca��o Rat��n; 10��10���44.91��� N, 75��44���53.01��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC- CRU 175 ��� 29 ♂♂, 38 ♀♀; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 6 Sep. 2017; same collector as for preceding; CUDC-CRU 176 ��� 18 ♂♂, 17 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96��� W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 177 ��� many ♂♂ and ♀♀; Ca��o Rat��n; 10��10���42.6��� N, 75��44���52.6��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC- CRU 178 ��� 1 ♂; La Punta, Laguna Encantada; 10��10���48.2��� N, 75��43���38��� W; 24 Aug. 2017; Y.M. Carpio- D��az, C.M. L��pez-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 179 ��� 5 ♂♂, 8 ♀♀; El Silencio, Laguna del Silencio; 10��10���35.4��� N, 75��44���26��� W; same date and collectors as for preceding; CUDC- CRU 180 ��� 4 ♂♂, 6 ♀♀; same collection data as for preceding; 10��10���34.8��� N, 75��44���26.6��� W; CUDC- CRU 181 ��� 10 ♂♂, 5 ♀♀; 10��10���32.3��� N, 75��44���27.2��� W; 25 Aug. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 182 ��� 19 ♂♂, 21 ♀♀; La Punta, Laguna Encantada; 10��10���47.72��� N, 75��43���40.09��� W; 25 Aug. 2017; Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja-Arrieta leg.; CUDC-CRU 183 ��� 14 ♂♂, 7 ♀♀; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 184 ��� 10 ♂♂, 17 ♀♀; El Silencio, Laguna del Silencio; 10��10���35.4��� N, 75��44���26.2��� W; 24 Aug. 2017; Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja- Arrieta leg.; ICN-CR-is 264. Previous records Bol��var: Bar��, Cartagena de Indias (Caribbean) (Carpio-D��az et al. 2016). Distribution Widely distributed in Atlantic coastal areas of Mexico (Kensley & Schotte 1989); Florida, Cuba (Van Name 1936); Bahamas, Tobago, Bonaire (Schultz 1974); Dominica, Curacao, Virgin Islands, Belize, Venezuela (Kensley & Schotte 1989); Puerto Rico (Hurtado et al. 2014); Colombia (Carpio-D��az et al. 2016); and Brazil (Silva & Alves 2000). It is also present on the Pacific coast of Costa Rica (Taiti et al. 2018). First record of the species for continental islands of the Colombian Caribbean., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 8-9, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Carpio-Diaz Y. M., Lopez-Orozco C. M., Herrera-Medina Y., Navas G. R. & Bermudez A. 2016. Primer registro de Tylos niveus y nuevo reporte de Porcellionides pruinosus (Oniscidea: Tylidae y Porcellionidae) para Colombia. Revista de la Academia colombiana de Ciencias exactas fisicas y naturales 40 (156): 433 - 437. https: // doi. org / 10.18257 / raccefyn. 343","Boone L. 1934. New and rare Cuban and Haitian terrestrial Isopoda. Bulletin of the American Museum of Natural History 66: 567 - 598.","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Schultz G. A. 1970. A review of the species of the genus Tylos Latreille from the New World (Isopoda. Oniscidea). Crustaceana 19: 297 - 30. https: // doi. org / 10.1163 / 156854070 X 00383","Kensley B. & Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022].","Silva J. & Alves E. 2000. Tylos niveus Budde-Lund, 1885 (Crustacea: Isopoda: Oniscidea: Tylidae): redescricao e nova ocorrencia para a praia de Taquaras, Santa Catarina, Brasil. Acta Biologica Paranaense 29 (1 - 4): 265 - 285. https: // doi. org / 10.5380 / abpr. v 29 i 0.595","Hurtado L. A., Lee E. J., Mateos M. & Taiti S. 2014. Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PloS ONE 9 (4): e 94081. https: // doi. org / 10.1371 / journal. pone. 0094081","Taiti S., Montesanto G. & Vargas J. 2018. Terrestrial Isopoda (Crustacea, Oniscidea) from the coasts of Costa Rica, with descriptions of three new species. Revista de Biologia tropical 66 (1): S 187 - S 210. https: // doi. org / 10.15517 / RBT. V 66 I 1.33296","Schultz G. A. 1974. Terrestrial isopod crustaceans (Oniscoidea) mainly from the West Indies and adjacent regions: 1. Tylos and Ligia. Studies on the fauna of Cura ҫao and other Caribbean islands 149: 162 - 173. Available from http: // www. repository. naturalis. nl / document / 549820 [accessed 7 Jan. 2022]."]}

Family Detonidae Budde-Lund, 1904 Remarks The family Detonidae Budde-Lund, 1904 comprises 39 littoral species in the genera Armadilloniscus Uljanin, 1875 (32 spp.), Deto Gu��rin-M��neville, 1836 (5 spp.) and Detonella Lohmander, 1927 (2 spp.), including two dubious species, Armadilloniscus minutus Uljanin, 1875 and Deto spinicornis Brandt, 1851 (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schmidt 2002; Schmalfuss 2003; Schmidt & Leistikow 2004; WoRMS 2020b). The family can be characterized by animals having a clinger habitus (sensu Schmalfuss 1984), cephalon with well-developed lateral lobes, dorsal surface more or less tuberculated or with longitudinal ridges, exopods of pleopods 1���5 without respiratory structures and lateral margins covered with thin setae conferring a fringe-like appearance (see Taiti & Ferrara 1989; Schmidt 2002). Recently, Reinoso-Fl��rez et al. (2016) recorded the family from P��ramo de Estambul (Tolima), Colombia. However, after the re-examination of the material deposited at Colecci��n Zool��gica de la Universidad del Tolima (CZUT-IT 1423���1426, see Reinoso-Fl��rez et al. 2016), the specimens belong to the families Styloniscidae and Philosciidae, and therefore, our work constitutes the first record of the family in the country., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 19, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1904. A Revision of Crustacea Isopoda Terrestria, with Additions and Illustrations. Pt 1 Eubelum, Pt 2 Spherilloninae, Pt 3 Armadillo. H. Hagerup, Copenhagen, Denmark.","Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Taiti S. & Ferrara F. 1989. New species and records of Armadilloniscus Uljanin 1875 (Crustacea Isopoda Oniscidea) from the coasts of the Indian and Pacific oceans. Tropical Zoology 2: 59 - 88. https: // doi. org / 10.1080 / 03946975.1989.10539427","Schmidt C. 2002. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 1 (Olibrinidae to Scyphacidae s. str.). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 78: 275 - 352. https: // doi. org / 10.1002 / mmnz. 20020780207","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Schmidt C. & Leistikow A. 2004. Catalogue of genera of the terrestrial Isopoda (Crustacea: Isopoda: Oniscidea). Steenstrupia 28 (1): 1 - 118.","Schmalfuss H. 1984. Eco-morphological strategies in terrestrial isopods. Proceedings of the Zoological Society of London 53: 49 - 63.","Reinoso-Florez G., Villa-Navarro F. A. & Losada-Prado S. 2016. Artropofauna epigea del paramo Estambul (Tolima), Colombia. Biota Colombiana 17 (2): 39 - 51. Available from http: // hdl. handle. net / 20.500.11761 / 9381 [accessed 6 Jan. 2022]."]}

Armadilloniscus luisi Carpio-Díaz, Taiti & Campos-Filho sp. nov. urn:lsid:zoobank.org:act: 88600E5C-BFF9-49BA-976C-8853E33A6E58 Figs 1, 2E, 10–12 Etymology The new species is named after Mr Luis Felipe Arrieta Zúñiga, for his interest in nature and support to Ricardo Borja-Arrieta as a biologist. Material examined Holotype COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • ♂; Laguna del Silencio, El Silencio; 10°10′35.4″ N, 75°44′26″ W; 24 Aug. 2017; C.M. López-Orozco, Y.M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 193. Paratypes COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 194 • 10 ♂♂, 5 ♀♀; same collection data as for holotype; CUDC-CRU 195 • 1 ♀; 10°10′32.9″ N, 75°44′27.3″ W; same date and collectors as for holotype; CUDC-CRU 206 • 1 ♀; 10°10′33.5″ N, 75°44′26.9″ W; same date and collectors as for holotype; CUDC-CRU 207 • 1 ♂; 10°10′35″ N, 75°44′26.4″ W; same date and collectors as for holotype; CUDC-CRU 208 • 1 ♂; 10°10′34.8″ N, 75°44′26.6″ W; same date and collectors as for holotype; CUDC-CRU 209 • 6 ♂♂, 12 ♀♀; same collection data as for holotype; 10°10′35″ N, 75°44′16.4″ W; CUDC-CRU 210 • 14 ♂♂, 10 ♀♀; 10°10′34″ N, 75°44′26.8″ W; 25 Aug. 2017; same locality and collectors as for holotype; CUDC-CRU 211 • 3 ♂♂; La Punta, Laguna Encantada; 10°10′48.4″ N, 75°43′38.4″ W; 25 Aug. 2017; C.M. López-Orozco, Y.M. Carpio-Díaz and R. Borja- Arrieta leg.; CUDC-CRU 196 • 1 ♂; same locality as for preceding; 10°10′47.72″ N, 75°43′40.09″ W; same date and collectors as for holotype; CUDC-CRU 197 • 1 ♂, 1 ♀; same locality as for preceding; 10°10′48″ N, 75°43′39.3″ W; same date and collectors as for holotype; CUDC-CRU 198 • 1 ♀; same locality as for preceding; 10°10′47.7″ N, 75°43′39.93″ W; same date and collectors as for holotype; CUDC-CRU 199 • 2 ♂♂, 3 ♀♀; same locality as for preceding; 10°10′48.3″ N, 75°43′38.9″ W; same date and collectors as for holotype; CUDC-CRU 200 • 3 ♀♀; same locality as for preceding; 10°10′48.1″ N, 75°43′39.1″ W; same date and collectors as for holotype; CUDC-CRU 201 • 2 ♀♀; same locality as for preceding; 10°10′48.1″ N, 75°43′37.5″ W; same date and collectors as for holotype; CUDC-CRU 202 • 4 ♂♂, 4 ♀♀; same locality as for preceding; 10°10′47.72″ N, 75°43′40.09″ W; same date and collectors as for holotype; CUDC-CRU 203 • 2 ♂♂, 2 ♀♀; same locality as for preceding; 7 Sep. 2017; same date and collectors as for holotype; CUDC-CRU 204 • 3 ♂♂, 4 ♀♀; Laguna La Charca; 10°10′47.8″ N, 75°44′0.4″ W; 7 Sep. 2017; C.M. López-Orozco, Y.M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC- CRU 205. Description MEASUREMENTS. Maximum length: male 2.5 mm; female 3.1 mm. BODY. Color brown with usual pale muscle spots; antennae brown with pale flagellum (Fig. 2E). Body elongated and elliptical-shaped (Fig. 10A, H). Dorsal surface bearing elongated scale-setae (Fig. 10B). Dorsum covered with broad tubercles (Fig. 10A, C–E, H): cephalon bearing seven tubercles in two rows, anterior row with four and posterior row with three; ♂ pereonites 1–3 with one row of 10 tubercles, 4–7 with one row of eight tubercles; ♀ pereonites 1, 5 and 6 with one row of 10 tubercles, 2–4 and 7 with one row of eight tubercles; ♂ pleonites 1–5 and telson with two paramedian tubercles, pleonite 2 with two broad tubercles at sides, pleonite 3 with two small additional tubercles; ♀ pleonites 1–5 and telson with two paramedian tubercles, pleonites 2 and 3 with two additional small tubercles at sides. CEPHALON. Lateral lobes well developed and directed outwards, median lobe triangular, slightly surpassing distal margin of lateral lobes, frontal and suprantennal lines absent; eyes consisting of four ommatidia (Fig. 10C–D). PLEON. Outline continuous with that of pereonite 7, epimera of pleonites 3–5 rectangular (Fig. 10A, E, H). Telson (Fig. 10E) triangular, lateral margins almost straight, distal part truncate and not surpassing distal margin of pleonite 5 epimera. ANTENNULA. Composed of two articles, second article bearing a few lateral setae, distal margin bearing one flagellar seta and two aesthetascs (Fig. 10F). ANTENNA. When extended posteriorly slightly surpassing pereonite 1, flagellum of four articles (Fig. 10G). MOUTH. Mandibles with molar penicil of several plumose setae, left mandible with 2+1 penicils (Fig. 11A), right with 1+1 penicils (Fig. 11B). Maxillula (Fig. 11C) inner endite with two penicils; outer endite of 4+8 teeth, five of them pectinated, and one slender stalk between outer and inner set, outer distal margin bearing fringe of long thin setae. Maxilla (Fig. 11D) with lobes fused, inner middle portion with five long setae, distal margin rounded, bearing several thin and thick setae. Maxilliped (Fig. 11E) palp bearing two tufts of setae on first article, distal articles of palp fused, bearing several thin and thick setae; endite rectangular, inner margin bearing fringe of long thin setae plus four long thick setae, distal margin bilobed, bearing broad rounded penicil with long thick setae. UROPOD. Protopod enlarged with distal margin rounded, exopod not surpassing protopod, endopod longer than exopod (Fig. 11F). PEREOPODS. Pereopods 1–7 stout, merus to propodus bearing sparse setae on sternal margin, carpus 1 with strong distal seta bearing long sensilla, dactylus with very long ungual seta, dactylar seta elongated, apically cleft and plumose. Male PEREOPODS 1 AND 7. Without any sexual modifications (Fig. 12A–B). GENITAL PAPILLA. Ventral shield triangular, papilla slightly cleft at apex, bearing small setae (Fig. 12C). PLEOPODS. Pleopod 1 (Fig. 12D) exopod ovoid; endopod three times as long as exopod, apical portion bent outwards. Pleopod 2 (Fig. 12E) exopod subtriangular; endopod with distal article flagelliform. Exopods of pleopods 3–5 as in Fig. 12F–H. Remarks In the Caribbean region, four species of Armadilloniscus have been recorded: A. caraibicus from Venezuela, A. ellipticus (Harger, 1878) from the Atlantic coasts of USA and Bermuda, A. ninae Schultz, 1984 from Belize, and A. steptus Schotte & Heard, 1991 from Turks and Caicos Islands (British West Indies) (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schotte & Heard 1991). For a definition of the genus see Arcangeli (1957), Vandel (1962), Taiti & Ferrara (1989), and Schmidt (2002). Armadilloniscus luisi Carpio-Díaz, Taiti & Campos-Filho sp. nov. is readily distinguished from all the other species of the genus in the number and arrangement of the dorsal tubercles on the cephalon, pereon and pleon. Moreover, it can be distinguished from the other four Caribbean species in having two broad tubercles on pleonite 2 (vs small in A. caraibicus and A. steptus, absent in A. ninae), the cephalon with the median lobe triangular (vs rounded in A. caraibicus) and lateral lobes subquadrangular (vs subtriangular in A. ninae and A. steptus), eyes composed of four ommatidia (vs 4–5 in A. caraibicus, 6 in A. ellipticus and 5 in A. ninae), telson with distal margin truncate (vs rounded in A. caraibicus and A. ellipticus, triangular in A. ninae), and ♂ pleopod 1 exopod ovoid (vs subtriangular in A. caraibicus, A. ellipticus and A. ninae, almost rounded in A. steptus) (Paoletti & Stinner 1989; Schultz 1984; Schotte & Heard 1991; Garthwaite et al. 1992; Schmidt 2002). Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia.

Ischioscia perijaensis Carpio-D��az, L��pez-Orozco & Campos-Filho n. sp. (Figs 4-6) urn:lsid:zoobank.org:act: DC17E8EB-202D-4FBE-BB4A-40F721BD516A TYPE MATERIAL. ��� Holotype. Colombia ��� ♂ (parts in micropreparations); Quebrada El Jord��n, El Contento Village, San Jos�� de Oriente, La Paz, Cesar; 10��17���40.4���N, 72��55���52.1���W; 14.VIII.2015; C. M. L��pez-Orozco leg.; CUDC-CRU 40. ETYMOLOGY. ��� The new species is named after the locality where the specimens were collected: Serran��a de Perij��. DESCRIPTION Body size: 6 mm �� 2.9 mm. Body as in Fig. 4A. Colour dark brown: antennal peduncle with first and second articles slightly pigmented, third to fifth article and flagellum strongly pigmented; cephalon with irregular unpigmented spots; pereon with longitudinal unpigmented spots on paramedian region, epimera with one unpigmented spot; pleon and telson strongly pigmented; uropods slightly pigmented (Fig. 4A). Dorsal surface bearing short triangular scale-setae (Fig. 4B). Cephalon (Fig. 4C, D) without lateral lobes, frontal line and supra-antennal slightly bent downwards medially; vertex dorsoventrally flattened with prominent compound eyes on lateral protrusions; eyes with 25 ommatidia arranged in four rows. Pleon (Fig. 4A) narrower than pereon, neopleurae 3-5 well developed and directed backwards. Telson (Fig. 4E) with lateral margins concave, apex rounded. Antennula (Fig. 4F) of three articles, distal article bearing six aesthetascs in three sets plus apical pair. Antenna (Fig. 4G) when extended posteriorly reaching anterior margin of pereonite 3; flagellum of three articles subequal in length, apical organ very long. Mandibles with molar penicil composed of 5 to 6 branches, left mandible (Fig. 5A) with 2+1 penicils, right mandible (Fig. 5B) with 1+1 penicils. Maxillula (Fig. 5C) inner endite with distal margin rounded bearing two stout hairy penicils; outer endite of 4+6 teeth, two of them with cleft apex. Maxilla (Fig. 5D) outer lobe twice as wide as inner lobe, distal margin rounded and covered with thin setae, two thick setae on inner portion; inner lobe rounded bearing thick setae. Maxilliped (Fig. 5E) basis rectangular bearing sparse setae; palp with two setae on proximal article; endite sub-rectangular, median seta surpassing distal margin, strong seta on distal outer portion, ventral surface setose. Uropod (Fig. 5F) protopod and exopod grooved on lateral margin, exopod longer than endopod, endopod inserted proximally. Pereopods 1-7 long and slender, carpus 1 with antennal grooming brush transverse; dactylus of two claws, inner claw not surpassing outer claw, ungual seta long, dactylar seta apically plumose. Pleopod exopods without respiratory areas. Male Pereopod 1 (Fig. 6A) with merus, carpus and propodus, and pereopod 2 merus and carpus bearing dense field of setae. Pereopod 7 (Fig. 6B) ischium with rounded depression on rostral surface, sternal margin bearing brush of thin setae on central part. Genital papilla (Fig. 6C) truncated apically, triangular ventral shield and two apical orifices. Pleopod 1 (Fig. 6D) exopod cordiform, inner margin almost straight, outer margin without lateral incision, distal margin rounded, lateral process short; endopod stout, almost twice as long as exopod, distal portion straight bearing row of small setae on medial margin and subapical lobe on outer margin. Pleopod 2 (Fig. 6E) exopod triangular, outer margin concave bearing several setae; endopod stout, slightly longer than exopod. Pleopods 3 and 4 as in Fig. 6F and G, respectively. Pleopod 5 (Fig. 6H) exopod rhomboid slightly wider than long, transverse rows of scales, outer margin sinuous bearing four setae, inner and outer margins densely covered with thin setae. REMARKS Ischioscia perijaensis Carpio-D��az, L��pez-Orozco & Campos- Filho n. sp. is readily distinguishable from their congeners in having the male pereopod 1 carpus not enlarged, the male pereopod 7 ischium with a rounded depression on rostral surface, and the male pleopod 1 exopod cordiform with lateral process short and outer margin lacking the lateral incision. In addition, I. perijaensis Carpio-D��az, L��pez-Orozco & Campos-Filho n. sp. differs from I. herrerai L��pez-Orozco, Carpio-D��az & Campos-Filho n. sp. in having the male pereopod 1 carpus not enlarged, the male pereopod 7 ischium without lobe and in the shape of the male pleopod 1 exopod., Published as part of Campos-Filho, Ivanklin Soares, L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., ��guiar, Jos�� Ot��vio & Navas, Gabriel R., 2020, Three new species of Ischioscia Verhoeff, 1928 (Isopoda, Oniscidea, Philosciidae) from Serran��a de Perij��, Andean Cordillera, Colombian Caribbean, pp. 115-130 in Zoosystema 42 (8) on page 120, DOI: 10.5252/zoosystema2020v42a8, http://zenodo.org/record/3726211

Ischioscia herrerai López-Orozco, Carpio-Díaz & Campos-Filho n. sp. (Figs 1-3) urn:lsid:zoobank.org:act: 84046969-EDD0-4A8E-8175-7C645A1BB873 TYPE MATERIAL. — Holotype. Colombia • ♂; Quebrada El Jordán, El Contento Village, San José de Oriente, La Paz, Cesar; 10°17’40.4”N, 72°55’52.1”W; 13.VIII.2015; Y. Herrera-Medina leg.; CUDC- CRU 33. Paratypes. Colombia • 1 ♂ (part in micropreparations); same locality and date as for holotype; C. M. López-Orozco leg.; CUDC-CRU 45 • 1 ♀ (part in micropreparations); same locality and date as for holotype; C. M. López-Orozco leg.; CUDC-CRU 47 • 2 ♂, 3 ♀; same locality and date as for holotype; C. M. López-Orozco leg.; ICN-CI-86 • 12 ♂, 37 ♀; same locality and date as for holotype; C. M. López-Orozco leg.; CUDC-CRU-37 • 1 ♂, 13 ♀; Cerro Pintao, La Jagua del Pilar, La Guajira; 10°25’41.8”N, 72°56’8.3”W; 10.VIII.2015; C. M. López-Orozco leg.; CUDC-CRU-36. ETYMOLOGY. — The new species is named after Yeison Herrera- Medina who collected part of the specimens used in this study. DESCRIPTION Maximum body size: male 9 mm × 3.5 mm (holotype); female 9.8 mm × 3.8 mm. Body as in Fig. 1A. Colour dark brown: antennal peduncle with first and second articles slightly pigmented, third to fifth articles and flagellum strongly pigmented; cephalon with irregular unpigmented spots; pereon with longitudinal unpigmented areas on paramedian region, pereonites 4-7 with one longitudinal unpigmented spot on median portion, epimera with two distinct unpigmented spots near posterior portions; pleon, telson and uropod exopods strongly pigmented (Fig. 1A). Dorsal surface bearing short triangular scale-setae (Fig. 1B). Cephalon (Fig. 1C, D) lateral lobes and frontal line absent, supraantennal line almost straight; vertex dorsoventrally flattened with prominent compound eyes on lateral protrusions; eyes with 27 ommatidia arranged in four rows. Pleon (Fig. 1A) narrower than pereon, neopleurae 3-5 well developed and directed backwards. Telson (Fig. 1E) with lateral margins concave, apex rounded. Antennula (Fig. 1F) of three articles, distal article bearing six superimposed aesthetascs plus apical pair. Antenna (Fig. 1G) when extended posteriorly reaching anterior portion of pereonite 3; flagellum of three articles subequal in length, apical organ very long. Mandibles with molar penicil composed of seven branches, left mandible (Fig. 2A) with 2+1 penicils, right mandible (Fig. 2B) with 1+1 penicils. Maxillula (Fig. 2C) inner endite with distal margin rounded bearing two hairy penicils; outer endite of 4+5 teeth, two of them with cleft apex. Maxilla (Fig. 2D) outer lobe almost three times as wide as inner lobe, distal margin rounded and covered with thin setae, one thick seta on inner portion; inner lobe rounded bearing thick setae. Maxilliped (Fig. 2E) basis rectangular bearing sparse setae; palp with two setae on proximal article; endite sub-rectangular, rostral and caudal surfaces hairy, medial seta not surpassing distal margin. Uropod (Fig. 2F) protopod and exopod grooved on lateral margin, exopod as long as endopod, endopod inserted proximally. Pereopods 1-7 long and slender; carpus 1 with antennal grooming brush transverse and distal setae apically cleft; dactylus of two claws, inner claw not surpassing outer claw, ungual seta long, dactylar seta apically plumose. Pleopod exopods without respiratory areas. Male Pereopods 1-3 with carpus enlarged, rostral portion bearing dense field of setae. Pereopod 1 and 2 propodus and pereopods 1-3 merus bearing dense field of setae on rostral and sternal portions (Fig. 3A). Pereopod 7 (Fig. 3B) ischium with rounded lobe bearing one strong seta on caudal surface, and distal sternal portion prominent. Genital papilla (Fig. 3C) truncated apically, triangular ventral shield and two apical orifices. Pleopod 1 (Fig. 3D) exopod cordiform, inner margin slightly sinuous, outer margin with distinct lateral incision, apical lobe triangular, directed outwards; endopod stout, almost twice as long as exopod, distal portion straight with row of small setae on medial portion, apex rounded with subapical lobe on outer margin. Pleopod 2 (Fig. 3E) exopod triangular, outer margin concave bearing several setae; endopod stout, longer than exopod, apex pointed. Pleopods 3 and 4 as in Fig. 3F and G, respectively. Pleopod 5 (Fig. 3H) exopod triangular, transverse rows of scales, outer margin straight bearing five setae. REMARKS In the shape of the male pleopod 1 exopod Ischioscia herrerai López-Orozco, Carpio-Díaz & Campos-Filho n. sp. resembles I. martinae Leistikow, 1997 and I. muelleri Leistikow, 1997, but it differs in the shape of the male pereopod 1-2 carpus and the presence of a lobe on the male pereopod 7 ischium. In having a lobe on the male pereopod 7 ischium I. herrerai López-Orozco, Carpio-Díaz & Campos-Filho n. sp. is similar to I. variegata (Dollfus, 1893), I. panamaensis Leistikow, 1999, I. marmorata Leistikow, 2000, I. hirsuta Leistikow, 2001, I. colorata Leistikow, 2001, I. pariae Leistikow, 2001, I. guamae Leistikow, 2001, and I. trifasciata Leistikow, 2001; however, it differs from all these species in the colour pattern, shape of the male pereopods 1-3 carpus, of the lobe on the male pereopod 7 ischium, and of the male pleopod 1 exopod.

Ischioscia herrerai L��pez-Orozco, Carpio-D��az & Campos-Filho n. sp. (Figs 1-3) urn:lsid:zoobank.org:act: 84046969-EDD0-4A8E-8175-7C645A1BB873 TYPE MATERIAL. ��� Holotype. Colombia ��� ♂; Quebrada El Jord��n, El Contento Village, San Jos�� de Oriente, La Paz, Cesar; 10��17���40.4���N, 72��55���52.1���W; 13.VIII.2015; Y. Herrera-Medina leg.; CUDC- CRU 33. Paratypes. Colombia ��� 1 ♂ (part in micropreparations); same locality and date as for holotype; C. M. L��pez-Orozco leg.; CUDC-CRU 45 ��� 1 ♀ (part in micropreparations); same locality and date as for holotype; C. M. L��pez-Orozco leg.; CUDC-CRU 47 ��� 2 ♂, 3 ♀; same locality and date as for holotype; C. M. L��pez-Orozco leg.; ICN-CI-86 ��� 12 ♂, 37 ♀; same locality and date as for holotype; C. M. L��pez-Orozco leg.; CUDC-CRU-37 ��� 1 ♂, 13 ♀; Cerro Pintao, La Jagua del Pilar, La Guajira; 10��25���41.8���N, 72��56���8.3���W; 10.VIII.2015; C. M. L��pez-Orozco leg.; CUDC-CRU-36. ETYMOLOGY. ��� The new species is named after Yeison Herrera- Medina who collected part of the specimens used in this study. DESCRIPTION Maximum body size: male 9 mm �� 3.5 mm (holotype); female 9.8 mm �� 3.8 mm. Body as in Fig. 1A. Colour dark brown: antennal peduncle with first and second articles slightly pigmented, third to fifth articles and flagellum strongly pigmented; cephalon with irregular unpigmented spots; pereon with longitudinal unpigmented areas on paramedian region, pereonites 4-7 with one longitudinal unpigmented spot on median portion, epimera with two distinct unpigmented spots near posterior portions; pleon, telson and uropod exopods strongly pigmented (Fig. 1A). Dorsal surface bearing short triangular scale-setae (Fig. 1B). Cephalon (Fig. 1C, D) lateral lobes and frontal line absent, supraantennal line almost straight; vertex dorsoventrally flattened with prominent compound eyes on lateral protrusions; eyes with 27 ommatidia arranged in four rows. Pleon (Fig. 1A) narrower than pereon, neopleurae 3-5 well developed and directed backwards. Telson (Fig. 1E) with lateral margins concave, apex rounded. Antennula (Fig. 1F) of three articles, distal article bearing six superimposed aesthetascs plus apical pair. Antenna (Fig. 1G) when extended posteriorly reaching anterior portion of pereonite 3; flagellum of three articles subequal in length, apical organ very long. Mandibles with molar penicil composed of seven branches, left mandible (Fig. 2A) with 2+1 penicils, right mandible (Fig. 2B) with 1+1 penicils. Maxillula (Fig. 2C) inner endite with distal margin rounded bearing two hairy penicils; outer endite of 4+5 teeth, two of them with cleft apex. Maxilla (Fig. 2D) outer lobe almost three times as wide as inner lobe, distal margin rounded and covered with thin setae, one thick seta on inner portion; inner lobe rounded bearing thick setae. Maxilliped (Fig. 2E) basis rectangular bearing sparse setae; palp with two setae on proximal article; endite sub-rectangular, rostral and caudal surfaces hairy, medial seta not surpassing distal margin. Uropod (Fig. 2F) protopod and exopod grooved on lateral margin, exopod as long as endopod, endopod inserted proximally. Pereopods 1-7 long and slender; carpus 1 with antennal grooming brush transverse and distal setae apically cleft; dactylus of two claws, inner claw not surpassing outer claw, ungual seta long, dactylar seta apically plumose. Pleopod exopods without respiratory areas. Male Pereopods 1-3 with carpus enlarged, rostral portion bearing dense field of setae. Pereopod 1 and 2 propodus and pereopods 1-3 merus bearing dense field of setae on rostral and sternal portions (Fig. 3A). Pereopod 7 (Fig. 3B) ischium with rounded lobe bearing one strong seta on caudal surface, and distal sternal portion prominent. Genital papilla (Fig. 3C) truncated apically, triangular ventral shield and two apical orifices. Pleopod 1 (Fig. 3D) exopod cordiform, inner margin slightly sinuous, outer margin with distinct lateral incision, apical lobe triangular, directed outwards; endopod stout, almost twice as long as exopod, distal portion straight with row of small setae on medial portion, apex rounded with subapical lobe on outer margin. Pleopod 2 (Fig. 3E) exopod triangular, outer margin concave bearing several setae; endopod stout, longer than exopod, apex pointed. Pleopods 3 and 4 as in Fig. 3F and G, respectively. Pleopod 5 (Fig. 3H) exopod triangular, transverse rows of scales, outer margin straight bearing five setae. REMARKS In the shape of the male pleopod 1 exopod Ischioscia herrerai L��pez-Orozco, Carpio-D��az & Campos-Filho n. sp. resembles I. martinae Leistikow, 1997 and I. muelleri Leistikow, 1997, but it differs in the shape of the male pereopod 1-2 carpus and the presence of a lobe on the male pereopod 7 ischium. In having a lobe on the male pereopod 7 ischium I. herrerai L��pez-Orozco, Carpio-D��az & Campos-Filho n. sp. is similar to I. variegata (Dollfus, 1893), I. panamaensis Leistikow, 1999, I. marmorata Leistikow, 2000, I. hirsuta Leistikow, 2001, I. colorata Leistikow, 2001, I. pariae Leistikow, 2001, I. guamae Leistikow, 2001, and I. trifasciata Leistikow, 2001; however, it differs from all these species in the colour pattern, shape of the male pereopods 1-3 carpus, of the lobe on the male pereopod 7 ischium, and of the male pleopod 1 exopod., Published as part of Campos-Filho, Ivanklin Soares, L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., ��guiar, Jos�� Ot��vio & Navas, Gabriel R., 2020, Three new species of Ischioscia Verhoeff, 1928 (Isopoda, Oniscidea, Philosciidae) from Serran��a de Perij��, Andean Cordillera, Colombian Caribbean, pp. 115-130 in Zoosystema 42 (8) on pages 116-120, DOI: 10.5252/zoosystema2020v42a8, http://zenodo.org/record/3726211, {"references":["LEISTIKOW A. 1997. - Terrestrial isopods from Costa Rica and a redescription of Ischioscia variegata (Dollfus, 1893) (Crustacea: Isopoda: Oniscidea). Canadian Journal of Zoology 75 (9): 1415 - 1464. https: // doi. org / 10.1139 / z 97 - 768","LEISTIKOW A. 1999. - New species of the genera Ischioscia and Andenoniscus, terrestrial isopods from Panama (Crustacea, Isopoda). Studies on Neotropical Fauna and Environment 34 (3): 156 - 175. https: // doi. org / 10.1076 / snfe. 34.3.156.8911","LEISTIKOW A. 2000. - The terrestrial isopod genus Ischioscia in Costa Rica: new species and records, and an analysis of its phylogeny (Crustacea, Isopoda, Oniscidea). Mitteilungen aus dem Museum fur Naturkunde in Berlin 76 (1): 19 - 49. https: // doi. org / 10.1002 / mmnz. 20000760104","LEISTIKOW A. 2001 a. - Phylogeny and Biogeography of South American Crinocheta, traditionally placed in the family \" Philosciidae \" (Crustacea: Isopoda: Oniscidea). Organism Diversity and Evolution, 1 (Electr. Suppl. 4): 1 - 85. http: // www. senckenberg. uni-frankfurt. de / odes / 01 - 04. htm"]}