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Thirty-eight species of Isopoda, belonging to 13 families and 29 genera, are listed from Kuwait based on previous literature records (of 17 species) and collections carried out along Kuwait’s coastal and subtidal zones during the present study. The majority of species belongs to the suborder Cymothoida (23), followed by Sphaeromatidea (9), Oniscidea (3), Valvifera (2), and Asellota (1). In total, 25 species were collected and identified from 12 families and 22 genera from Kuwaiti coastal and subtidal areas. These include eight families, 15 genera, and 21 species recorded for the first time from Kuwait. Isopod diversity was highest in the sandy rock areas, including southern Kuwait, particularly in Al-Khiran and Al-Nuwaiseeb, and in mixed habitat (muddy, rocky, and sandy) intertidal transects such as in Failaka Island. The species number increased from the subtidal and lowest zones into the high tidal zone. Isopods were found in sandy substrata, among shells, cobbles, rocks, dead corals, and algae.

An overview of the Estonian terrestrial isopod fauna is given, based on literature data and material collected from 1984 to 2021. The identified material consisted of 10915 specimens belonging to 14 species and collected from 172 localities throughout Estonia. In combination with previous data from the literature data, there are now reliable records of 16 species of woodlice from Estonia. Two species, viz. Platyarthrus hoffmannseggii Brandt, 1833 and Hyloniscus riparius (C. Koch, 1838), are new for the fauna. The latter has probably colonised Estonia recently and range expansions have been reported elsewhere. The data on Philoscia muscorum (Scopoli, 1763) are dubious, and this species is currently excluded from the Estonian list.

Ligia baudiniana Milne-Edwards, 1840 Fig. 1 Ligia baudiniana Milne-Edwards, 1840: 155. Ligia (Hirtiligia) baudiniana ��� Van Name 1936: 58, figs 5a, 14. Ligia baudiniana ��� R��os & Ramos 1990: 93, fig. 7. ��� Schmalfuss 2003: 124. ��� Lazarus-Agudelo & Cantera-Kintz 2007: 226. ��� L��pez-Orozco et al. 2014: 196, figs 1���3. Additional references Jackson (1922); Schultz (1984); Kensley & Schotte (1989); Leistikow (1997); Leistikow & W��gele (1999); Espinosa-P��rez & Hendrickx (2001); Santamaria et al. (2014). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���34.8��� N, 75��44���26.6��� W; 24 Aug. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 141 ��� 3 ♂♂, 2 ♀♀; same collection data as for preceding; 10��10���35��� N, 75��44���26.4��� W; CUDC-CRU 142 ��� 1 juv.; same collection data as for preceding; 10��10���35.4��� N, 75��44���26.2��� W; CUDC-CRU 143 ��� 1 ♂, 1 ♀; same collection data as for preceding; 10��10���34.3��� N, 75��44���26.7��� W; CUDC-CRU 144 ��� 1 ♂; same locality and collectors as for preceding; 10��10���32.9��� N, 75��44���27.3��� W; 25 Aug. 2017; CUDC-CRU 145 ��� 1 ♀; same collection data as for preceding; 10��10���32.3��� N, 75��44���27.2��� W; CUDC-CRU 146 ��� 1 ♂; same collection data as for preceding; 10��10���32.5��� N, 75��44���27.1��� W; CUDC-CRU 147 ��� 1 ♀; Ca��o Rat��n, Laguna Palmar; 10��10���24.5��� N, 75��44���39.7��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 148 ��� 2 ♂♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���35.4��� N, 75��44���26��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 149 ��� 1 ♀; same locality and collectors as for preceding; 10��10���48��� N, 75��43���39.3��� W; 25 Aug. 2017; CUDC-CRU 150 ��� 1 ♂; same collection data as for preceding; 10��10���34��� N, 75��44���26.8��� W; CUDC-CRU 151 ��� 1 juv.; same collection data as for preceding; 10��10���47.74��� N, 75��43���39.93��� W; CUDC-CRU 152 ��� 1 ♂, 1 ♀; same collection data as for preceding; 10��10���35��� N, 75��44���26.9��� W; ICN- CR-is 260 ��� 1 ♂, 2 ♀♀; Ca��o Rat��n; 10��10���44.91��� N, 75��44���53.01��� W; 24 Nov. 2017, same collectors as for preceding; ICN-CR-is 261 ��� 1 juv.; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018, C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 222. Previous records Valle del Cauca: Bah��a M��laga (Pacific); Bol��var: Cartagena de Indias (Caribbean); Magdalena: Santa Marta (R��os & Ramos 1990; Lazarus-Agudelo & Cantera-Kintz 2007; L��pez-Orozco et al. 2014; Santamaria et al. 2014). Distribution This species was originally described from Veracruz, Mexico (but see Discussion). Molecular phylogenetic analyses, however, revealed that what has been denominated as this species corresponds to a clade composed of highly divergent lineages (probably a complex of cryptic species), distributed in southern Florida, across the Caribbean Sea, Bahamas, Bermuda, and probably in northern Brazil, and the Pacific coast from Costa Rica, Panam��, Colombia, probably to Ecuador (Santamaria et al. 2014). Although this species has been reported in the Gulf of California and California, extensive sampling for phylogeographic studies conducted by some of us suggest that these previous records are likely misidentifications (Hurtado et al. 2010; Eberl et al. 2013; Hurtado et al. 2018)., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 7, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Milne-Edwards M. 1840. Ordre des isopodes. In: Histoire Naturelle des Crustaces, comprenant l'Anatomie, la Physiologie et la Classficiation de ces Animaux. Tome Troisieme. 115 - 283. Librairie Encyclopedique de Roret, Paris.","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Rios R. & Ramos G. E. 1990. Los isopodos (Crustacea: Isopoda) de Bahia Malaga, Colombia. Revista de Ciencias, Universidad del Valle 2: 83 - 96.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Lazarus-Agudelo J. F. & Cantera-Kintz J. R. 2007. Crustaceos (Crustacea: Sessilia, Stomatopoda, Isopoda, Amphipoda, Decapoda) de Bahia Malaga, Valle del Cauca (Pacifico Colombiano). Biota Colombiana 8 (2): 221 - 239.","Lopez-Orozco C. M., Bermudez A. & Navas G. R. 2014. Primer registro de Ligia baudiniana (Crustacea: Isopoda: Oniscidea) para el Caribe Colombiano. Boletin de Investigaciones marinas y costeras 43 (1): 195 - 200. https: // doi. org / 10.25268 / bimc. invemar. 2014.43.1.41","Jackson H. G. 1922. A revision of the isopod genus Ligia (Fabricius). Proceedings of the Zoological Society of London 92 (3): 683 - 703. https: // doi. org / 10.1111 / j. 1096 - 3642.1922. tb 02164. x","Kensley B. & Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022].","Leistikow A. 1997. Terrestrial isopods from Costa Rica and a redescription of Ischioscia variegata (Dollfus, 1893) (Crustacea: Isopoda: Oniscidea). Canadian Journal of Zoology 75 (9): 1415 - 1464. https: // doi. org / 10.1139 / z 97 - 768","Leistikow A. & Wagele J. W. 1999. Checklist of terrestrial isopods of the New World (Crustacea, Isopoda, Oniscidea). Revista brasileira de Zoologia 16: 1 - 72. https: // doi. org / 10.1590 / S 0101 - 81751999000100001","Espinosa-Perez M. C. & Hendrickx M. E. 2001. Checklist of isopods (Crustacea: Peracarida: Isopoda) from the Eastern Tropical Pacific. Belgian Journal of Zoology 131 (1): 43 - 55.","Santamaria C. A., Mateos M. C. & Hurtado L. A. 2014. Diversification at the narrow sea-land interface in the Caribbean: phylogeography of endemic supralittoral Ligia isopods. Frontiers in Ecology and Evolution 2: e 42. https: // doi. org / 10.3389 / fevo. 2014.00042","Hurtado L. A., Mateos M. & Santamaria C. A. 2010. Phylogeography of supralittoral rocky intertidal Ligia isopods in the Pacific region from Central California to Central Mexico. PLoS ONE 5 (7): e 11633. https: // doi. org / 10.1371 / journal. pone. 0011633","Eberl R., Mateos M., Grosberg R. K., Santamaria C. A. & Hurtado L. A. 2013. Phylogeography of the supralittoral isopod Ligia occidentalis around the Point Conception marine biogeographic boundary. Journal of Biogeography 40: 2361 - 2372. https: // doi. org / 10.1111 / jbi. 12168","Hurtado L. A., Mateos M., Wang C., Santamaria C. A., Jung J., Khalaji-Pirbalouty V. & Kim W. 2018. Out of Asia: mitochondrial evolutionary history of the globally introduced supralittoral isopod Ligia exotica. PeerJ 6: e 4337. https: // doi. org / 10.7717 / peerj. 4337"]}

In this study, we report the results of a taxonomic survey of terrestrial isopods from Isla Grande, a ca 2 km2 island located in the north of Cartagena de Indias, Colombia. We found a total of 17 species belonging to nine families and 10 genera. Eight of these species have been reported only from the Caribbean region, nine are recorded for the first time in Colombia, and three are new to science and described here: Tylos negroi López-Orozco, Carpio-Díaz & Campos-Filho sp. nov., Stenoniscus nestori López-Orozco, Taiti & Campos-Filho sp. nov. and Armadilloniscus luisi Carpio-Díaz, Taiti & Campos-Filho sp. nov. Our revision also determined that the genus Microphiloscia is a junior synonym of Halophiloscia; and moreover, Halophiloscia trichoniscoides comb. nov. is redescribed. We also provide illustrations for Armadilloniscus caraibicus and Armadilloniscus ninae. Most (16) of the species were found in coastal habitats (i.e., beaches, coastal lagoons and mangroves), whereas the tropical dry forest harbored only two species. Molecular phylogenetic inferences supported the presence of three species of Tylos in Isla Grande (i.e., one new species and a new lineage within each of two known species). Our work on Tylos highlights the importance of combining taxonomic and molecular analyses to support taxonomic decisions and uncover cryptic diversity. Due to the multiple threats to Caribbean coastal habitats, taxonomic and molecular genetic research are urgently needed to understand biodiversity patterns of oniscideans in the Caribbean, which will inform strategies for their protection. Such studies will also contribute to our knowledge of their evolution, ecology, and potential uses, as well as the factors that have shaped the remarkable Caribbean biodiversity.

Eoligiiscus, new genus TYPE SPECIES: Eoligiiscus tarraconensis, new species. DIAGNOSIS: Sex unknown. An oniscidean with the following unique combination of characters: Body size small (Eos), goddess of the dawn; Ligia Fabricius, type genus of the family; and the Greek suffix - iskos, a diminutive commonly applied as a suffix to woodlice generic names (as in Oniscus: itself a combination of Greek ὄνος [onos], meaning, “woodlouse”), and - iskos). The gender of the name is masculine., Published as part of Sánchez-García, Alba, Peñalver, Enrique, Delclòs, Xavier & Engel, Michael S., 2021, Terrestrial Isopods from Spanish Amber (Crustacea: Oniscidea): Insights into the Cretaceous Soil Biota, pp. 1-32 in American Museum Novitates 2021 (3974) on page 5, DOI: 10.1206/3974.1, http://zenodo.org/record/5356138

Autrigoniscus, new genus TYPE SPECIES: Autrigoniscus resinicola, new species. DIAGNOSIS: Male. An oniscidean with the following combination of characters: Body size small (Female. Unknown. ETYMOLOGY: The new genus group name is a combination of the pre-Roman tribe Autrigones (who lived in northern Spain near the region of the amber localities along with the Cantabri) and the Greek suffix – iskos, denoting a diminutive. The gender of the name is masculine., Published as part of Sánchez-García, Alba, Peñalver, Enrique, Delclòs, Xavier & Engel, Michael S., 2021, Terrestrial Isopods from Spanish Amber (Crustacea: Oniscidea): Insights into the Cretaceous Soil Biota, pp. 1-32 in American Museum Novitates 2021 (3974) on page 13, DOI: 10.1206/3974.1, http://zenodo.org/record/5356138

[EN] Terrestrial isopods (Crustacea: Oniscidea) are a model group for studying the colonization of land. However, their fossil record is remarkably scarce and restricted to amber inclusions, and therefore amber deposits represent valuable windows to their past diversity and morphology. Here we present a new collection of 11 terrestrial isopod specimens preserved in Albian-aged amber from the Peñacerrada I outcrop, northern Spain, which collectively represent the most thoroughly documented fauna of Mesozoic Oniscidea. The three new genera and species identified belong to three of five major groups of the Oniscidea: Eoligiiscus tarraconensis, new genus and species (Ligiidae), Autrigoniscus resinicola, new genus and species (Synocheta: Trichoniscidae), and Heraclitus helenae, new genus and species (Crinocheta: Detonidae?). These taxa significantly expand the known fossil record of Oniscidea and demonstrate that considerable cladogenesis had already transpired by the Albian. The assemblage represents the earliest-known diversification of Oniscidea, extending direct evidence of terrestrialization in the group back to the late Early Cretaceous. These new taxa exhibit some characteristics that may inform hypotheses relating to general patterns of terrestrial isopod evolution. A discussion is provided about different aspects of the paleoecology and biology of the fossils compared to the Recent fauna. The new species indicate that Cretaceous isopods were a group of considerable adaptive diversity, exhibiting innovations analogous to what Recent isopods would exhibit 105 million years later., A.S.-G. is funded by an APOSTD2019 Research Fellowship from the Generalitat Valenciana and the European Social Fund (ESF). This study is a contribution to the Spanish AEI/FEDER, UE Project CRE (CGL2017-84419).

Nine species of terrestrial isopods are reported for the Polynesian island of Rapa Nui (Easter Island) based upon museum materials and recent collections from field sampling. Most of these animals are non-native species, but two are new to science: Styloniscus manuvaka sp. n. and Hawaiioscia rapui sp. n. Of these, the former is believed to be a Polynesian endemic as it has been recorded from Rapa Iti, Austral Islands, while the latter is identified as a Rapa Nui island endemic. Both of these new species are considered ‘disturbance relicts’ and appear restricted to the cave environment on Rapa Nui. A short key to all the oniscidean species presently recorded from Rapa Nui is provided. We also offered conservation and management recommendations for the two new isopod species.

Thirty-four species of terrestrial isopods (Crustacea: Oniscidea) from the Oued Laou basin, in the Rif area of north-eastern Morocco, are recorded. One genus (Paractenoscia) and seven species (Trichoniscus microphthalmus, Paractenoscia cavernicola, Bathytropa rifensis, Soteriscus gibbosus, S. laouensis, Porcellio pseudornatus, and Eluma praticola) are described as new. The genus Soteriscus, unavailable according to article 13.3 of the International Commission on Zoological Nomenclature (ICZN), is here revalidated by choosing S. gaditanus as type species. Six species (Graeconiscus thermophilus, Ctenoscia minima, Platyarthrus parisii, Porcellio humberti, Porcellio flavocinctus and Eluma caelata) have been fully illustrated to facilitate their identifications. Ctenoscia dorsalis Verhoeff is considered to be a junior synonym of C. minima Dollfus. Porcellio ornatus from southern Spain is also figured for comparison with P. pseudornatus sp. nov. The composition and origin of the oniscidean fauna of the Rif region is briefly discussed.

Twenty-seven species of terrestrial Isopoda are recorded from Socotra Island (Yemen). One genus (Dumetoniscus) in the family Bathytropidae and three species (Serendibia cavernicola, Dumetoniscus graniticus and Platyarthrus alticolus) are described as new. A key to the 41 species of Oniscidea presently known from the Socotra Archipelago is given. The new records confirm the high number of endemics present on this archipelago.

Ligia italica Fabricius, 1798 Material examined 1 ♂, 2 ♀♀, St. 1, on rocks near sea, leg. S. Taiti and C. Rossano, 28 September 2005 (MZUF 9449). Distribution Coasts of Black Sea and Mediterranean Sea, Atlantic coasts of the Iberian Peninsula and northern Africa down to Cape Vert, and Macaronesian islands. New record for the Rif region. Remarks For diagnostic characters of L. italica see Vandel (1960a, p. 122, Figures 54 and 55)., Published as part of Taiti, Stefano & Rossano, Claudia, 2015, Terrestrial isopods from the Oued Laou basin, north-eastern Morocco (Crustacea: Oniscidea), with descriptions of two new genera and seven new species, pp. 2067-2138 in Journal of Natural History 49 (33) on page 2069, DOI: 10.1080/00222933.2015.1009512, http://zenodo.org/record/3999869, {"references":["Fabricius JC. 1798. Supplementum Entomologiae Systematicae. Hafniae: CG Proft & Storch; p. 296 - 306.","Vandel A. 1960 a. Isopodes terrestres (Premiere partie). Faune Fr. 64: 1 - 416."]}

Mead, A., Carlton, J.T., Griffiths, C.L., Rius, M. (2011): Introduced and cryptogenic marine and estuarine species of South Africa. Journal of Natural History 45 (39-40): 2463-2524, DOI: 10.1080/00222933.2011.595836, URL: http://dx.doi.org/10.1080/00222933.2011.595836

Ligia dioscorides Taiti & Ferrara, 2004 Ligia dioscorides Taiti & Ferrara, 2004: 217, figs 1���4, pl. 2. Material examined . Socotra: 2 ������, 8 ♀♀, Qaryah, 12��38'05.6"N 54��12'35.9"E, along wadi, near bridge, 14.I.2003, leg. S. Taiti & A. K. Nasher, MZUF-9197; 10 ♀♀, same data, NHCY (MZUF); 2 ������, 4 ♀♀, Ras Momi, wadi Erher, 12��33'02.7"N 54��27'35.7"E, 16.I.2003, leg. S. Taiti, MZUF-9198; 1 ���, 5 ♀♀, 2 juvs, same data, NHCY (MZUF); 1���, 3 ♀♀, Qalansiya Lagoon, Detwah Protected Area, 12��41'55.4"N 53��29'58.1"E, 20.I.2003, leg. S. Taiti, MZUF-9195; 1 ���, 2 ♀♀, between Didum and Cabho, 12��34'42"N 54��24'20"E, 16.I.2003, leg. S. Taiti & A. K. Nasher, MZUF- 9196; 2 ♀♀, same data, NHCY (MZUF); 1 ���, 3 ♀♀, 4 juvs, Erisseyl, 12��32'18.3"N 54��30'58.1"E, under seaweed on rocks, 17.I.2003, leg. S. Taiti, MZUF-9194. Distribution. Littoral species endemic to the Socotra Archipelago. It has been recorded from Socotra, Samha and Darsa islands. Family Tylidae, Published as part of Taiti, Stefano & Checcucci, Ilaria, 2009, New species and records of terrestrial Isopoda (Crustacea, Oniscidea) from Socotra Island, Yemen, pp. 73-103 in ZooKeys 31 (3) on page 78, DOI: 10.3897/zookeys.31.140, http://zenodo.org/record/576582, {"references":["Taiti S, Ferrara F (2004) The terrestrial Isopoda (Crustacea: Oniscidea) of the Socotra Archipelago. Fauna of Arabia 20: 211 - 325."]}

Genus Ligia Fabricius, 1798 Type-species: Oniscus oceanicus Linn��, 1767 by subsequent designation., Published as part of Taiti, Stefano & Checcucci, Ilaria, 2009, New species and records of terrestrial Isopoda (Crustacea, Oniscidea) from Socotra Island, Yemen, pp. 73-103 in ZooKeys 31 (3) on page 78, DOI: 10.3897/zookeys.31.140, http://zenodo.org/record/576582, {"references":["Fabricius JC (1798) Supplementum Entomologiae Systematicae. Proft & Storch, Hafniae, pp. 1 - 572.","Linne C (1767) Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio duodecima, reformata. Regnum Animale. Tom. I. Pars II. Vol. 1. Laurentii Salvii, Holmiae, pp. 533 - 1327."]}

LIGIIDAE Ligia Weber, 1795 Ligia (Megaligia) exotica Roux, 1828. Van Name, 1936:48, Published as part of Perez-Gelabert, Daniel E., 2008, Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography, pp. 1-530 in Zootaxa 1831 (1) on page 32, DOI: 10.11646/zootaxa.1831.1.1

11. Ligidium germanicum Verh. 3 junge Tiere am 28. X. in der Smoganica bei St. Lucia gefunden. Sie d��rften durch Regeng��sse hinabgetrieben worden sein., Published as part of K. W. Verhoeff, 1929, Arthropoden aus s��dostalpinen H��hlen., pp. 41-55 in Mitteilungen ��ber H��hlen- und Karstforschung 1929 on page 53