1,729 results on '"Gadallah, Neveen S."'
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2. Tachysphex nitidior de Beaumont 1940
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Tachysphex ,Arthropoda ,Animalia ,Biodiversity ,Tachysphex nitidior ,Hymenoptera ,Taxonomy - Abstract
Tachysphex nitidior de Beaumont, 1940 Fig. 15A–I Tachysphex nitidior de Beaumont, 1940: 175, ♀, ♁. Diagnosis. Body length: 6.5 mm. Male: Body wholly black (including antennae and legs) (Fig. 15A, B); mandible yellowish red mesally in most specimens (Fig. 15D); scutal and mesopleural sutures well-defined; fore femur with relatively large, glabrous and shiny notch (Fig. 15I); T 1–3 silvery fasciate posteriorly (Fig. 15G); gastral sterna with well-defined punctures. Material examined: 1♁, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Distribution: Azerbaijan, Bulgaria, Croatia, Egypt, France, Germany, Greece, Iran, Israel, Italy, Kazakhstan, Libya, Malta, Mongolia, Morocco, Portugal, Russia, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine, United Arab Emirates, Uzbekistan (Pulawski 2022), Saudi Arabia (new record). Comments. The characters of our male specimen agree well with de Beaumont’s key (1940: 162, couplet 30). It also agrees with Pulawski’s recognition (2007: 442).
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- 2023
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3. Gastrosericus sanctus Pulawski in de Beaumont, Bytinski-Salz & Pulawski 1973
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Gastrosericus ,Animalia ,Biodiversity ,Hymenoptera ,Gastrosericus sanctus ,Taxonomy - Abstract
Gastrosericus sanctus Pulawski in de Beaumont, Bytinski-Salz & Pulawski, 1973 Fig. 7A–G Gastrosericus sanctus Pulawski in de Beaumont, Bytinski-Salz & Pulawski, 1973: 16, ♀, ♁. Diagnosis. Male: body length: 7.0 mm. Body colour: see Fig. 7 (A–F). The male of this species shares with that of G. moricei the appressed genal (Fig. 7D) and propodeal pubescence (Fig. 7C); sternal fimbriae of S 3, S 4 noticeably long, slightly extending beyond posterior margin of sterna (Fig. 7E); gastral terga without yellow bands (Fig. 7A). It differs from that of G. moricei in the following: clypeal lobe broad, with arcuate to slightly sinuate free margin (Fig. 7B) and well-defined corner (obtusely pointed in G. moricei, see Pulawski 1995: 94, fig. 78f, h); S 3 and S 4 depressed medially (Fig. 7E), not fimbriate laterally (fimbriae of S 3 and S 4 present side to side, sterna not depressed medially, see Pulawski 1995: 95, fig. 79e, f). Material examined: 2 ♁, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Previous Saudi Arabian records: Abu Arish, Riyadh (Pulawski 1995). Distribution: Burkina Faso, Ghana, Israel, Ivory Coast, Kenya, Mali, Mauritania, Namibia, Niger, Oman, Pakistan, Saudi Arabia, Senegal, Somalia, South Africa, Tanzania, Togo, United Arab Emirates, Yemen, Zimbabwe. Comments. Characters of the collected specimens of this species agree with Pulawski’s key (1995: 23, couplet 24). It also agrees with Schmid-Egger & van Harten’s key (2022: 547, couplet 9), but gastral terga of male black, reddish brown posteriorly.
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- 2023
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4. Miscophus Jurine 1807
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Miscophus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Miscophus Jurine, 1807 Miscophus Jurine, 1807: 206. Type species: Miscophus bicolor Jurine, 1807, by monotypy.
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- 2023
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5. Gastrosericus Spinola 1839
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Gastrosericus ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Gastrosericus Spinola, 1839 Gastrosericus Spinola, 1839: 480. Type species: Gastrosericus waltlii Spinola, 1839, by monotypy.
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- 2023
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6. Solierella insidiosa de Beaumont 1964
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Solierella insidiosa ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy ,Solierella - Abstract
Solierella insidiosa de Beaumont, 1964 Fig. 11A–G Solierella insidiosa de Beaumont, 1964: 59, ♀, ♁. Diagnosis. Body length: 4.0 mm. Body colour: see Fig. 11 (A–F). Metanotum yellow; apex of median lobe of clypeus roundly projected, smooth and shiny (Fig. 11B); scutum densely finely punctate; punctures somewhat more spaced on scutellum, with more than a puncture diameter apart (Fig. 11C); metanotum smooth (Fig. 11C); depressed sides of scutellum densely finely pubescent; propodeum finely reticulate (Fig. 11A, C); mesepimeron with small rounded area smooth and shiny (Fig. 11D); fore and midfemora all black (Fig. 11E); fore and middle tibiae with small yellowish to whitish spot, hind tibia with whitish band along basal half dorsally. Material examined: 1♀, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Distribution: Saudi Arabia (new record), Spain, Syria, United Arab Emirates. Comments. This species is a new record for Saudi Arabia. Characters of the specimens collected from Farasan agree with de Beaumont’s key (1964: 55, couplet 5). It differs from the Syrian specimens of de Beaumont (1964: 59) in having the pale areas of body yellowish (whitish in the Syrian specimens). It also agrees well with Schmid-Egger’s key (2011: 582, couplet 4).
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- 2023
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7. Trypoxylon Latreille 1796
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Trypoxylon ,Hymenoptera ,Taxonomy - Abstract
Genus Trypoxylon Latreille, 1796 Trypoxylon Latreille, 1796: 121, no included species. Type species: Sphex figulus Linnaeus, 1758, designated by Latreille, 1802.
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8. Gastrosericus moricei E. Saunders 1910
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Gastrosericus moricei ,Gastrosericus ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Gastrosericus moricei E. Sauders, 1910 Fig. 6A–F Gastrosericus moricei E. Saunders, 1910: 529, ♁. Diagnosis. Body length: 7.5 mm. Body colour: see Fig. 6 (A–F). The female of this species shares with that of Gastrosericus sanctus the entire pygidial plate covered with dense, stout setae (Fig. 6F); appressed genal (Fig. 6C) and propodeal pubescence (Fig. 6D); the apical tarsomeres with one or more basoventral spines (spines lacking in some sanctus); gastral terga without yellow bands (Fig. 6A). It differs from G. sanctus by the following: free margin of clypeal lobe arcuate (Fig. 6B) (somewhat sinuate in most G. sanctus, see Pulawski 1995: 119, fig. 102a, b); the lobe free margin and the lip are simple (free margin slightly projecting close to lateral corners in G. sanctus, see Pulawski 1995: 120, fig. 103a, b); apical tarsomeres with one to four basomedian spines ventrally (0–2 spines in G. sanctus). Material examined: 1♀, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Previous Saudi Arabian records: Asir (Wadi Lasaba), Bahra, Riyadh (Pulawski 1995), Jeddah, Haddat Asham (Pulawski 1995; Gadallah & Assery 2004), Hutet Beni Tamim (10 km south of Riyadh) (Gadallah et al. 2013). Distribution: Algeria, Burkina Faso, Egypt, Gambia, India, Israel, Kazakhstan, Libya, Mali, Mauritania, Morocco, Niger, Oman, Pakistan, Saudi Arabia, Senegal, Somalia, Sri Lanka, Sudan, Thailand, Tajikistan, Togo, Tunisia, United Arab Emirates, Uzbekistan, Yemen. Comments. Characters of the only specimen collected agree with Pulawski’s key (1995: 19, couplet 18). It differs from the Arabian specimens in having femora black, pale yellow apically (in the Arabian specimens, femora reddish, pale yellowish apically, Pulawski 1995), but it resembles the Somalian females in having all femora black, yellow at tip (Pulawski 1995). This may be attributed to the fact that Farasan Archipelago belongs to the Afrotropical region.
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- 2023
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9. Nysson arabicus Gadallah & Edmardash 2023, sp. nov
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Nysson ,Nysson arabicus ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Nysson arabicus Gadallah & Edmardash sp. nov. Figs 3A–D, 4A, B, 5A–C Type material: SAUDI ARABIA. Holotype (♁), Sala Mountain, 1186 m [17°04’10.5”N 43°07’43.3”E] (Abdul Rahman Al Jabri farm), 15.xi.2022, Malaise trap, on Coffea arabica, collector Usama Abu El-Ghiet. Description of male holotype: body length: 6.0 mm. Colouration (Figs 3A–C, 5A–C): Head and thorax black (except posterior margin of pronotum, tegula, and basal area of scutellum dark ferruginous), antenna with scape (except yellow ventrally), pedicel and the following 4–5 flagellomeres dark reddish brown, rest dark brown; mandible ferruginous with dark tip; abdomen: T 1 mostly dark ferruginous, with two yellow postero-lateral oval markings, posterior margin black; T 2 with thin, medially interrupted yellow band posteriorly; T 3 with broad band posteriorly, narrowly interrupted medially; T 4 and T 5 with complete broad yellow bands posteriorly, T 6 and T 7 entirely reddish; metasomal sterna mostly ferruginous, but S 1 mostly black, ferruginous at base; legs all ferruginous, with some yellow at tip of fore femur, and along whole length of hind tibia dorsally, hind tibia reddish ventrally, with black spurs. Fore wing slightly darkened, fumigated at apex, with dark brown veins. Head (Figs 3C, D, 4A). In dorsal view as broad as mesoscutum, densely rugulose-lacunose, with dense fine silvery pubescence laterally; vertex thin and sharp; frons densely foveolate, with sharp carina between antennal bases, with coarser silvery setae; clypeus with dense silvery setae, with about 4–5 rounded teeth at apex; antennae placed at an elevated tubercle, scape longer than wide, distinctly notched apically, pedicle cup-shape, all flagellomeres transverse, wider than long, last flagellomere longer than wide, 1.5× as long as its basal width, obliquely truncate at apex. Thorax (Figs 3A, B, 4B). Pronotum distinctly angulate antero-laterally, posterior margin coarsely punctate along ferruginous area; mesoscutum densely areolate, scutellum and metanotum densely areolate (except scutellum basally nearly smooth and shiny); scutellum with two indistinct teeth laterally, hardly seen because of the coarse foveolation of the integument; propodeum broadly strigate dorsally, with two pale sharp teeth postero-laterally; tegula with dense fine and shallow punctures; mesopleuron densely foveate, metapleuron transversely strigate, shiny; mesosternum densely foveolate. Legs normal, simple. Wings as in other Nysson species. Gaster (Figs 3A, B, 5B, C). Terga double-edged, T 1 irregularly punctate, with distinct spaces between punctures, punctures coarser and larger in the yellow areas; T 2 and T 3 with dense shallow punctures, alutaceous in between punctures; T 3 slightly elevated and smooth at base, together with the following two terga with punctures more distinct along yellow bands; terga narrowed towards apex; T 6 and T 7 densely foveolate along their whole areas, T 7 with two short spine-like processes, with rounded area in between; S 1 densely punctate with large spaces between punctures, sparsely setose; S 2 with large rounded hump (not tubercle) near to the middle, posterior margins of S 2–4 with fine punctures arranged in two or three rows; S 5 with dense coarse punctures throughout. Female: Unknown. Etymology. This species is named after the Arabian area where the specimen was collected. Distribution: Saudi Arabia (Alridah, Jazan). Comments. On the generic level, the double-edged gastral terga of the new species agree with Synnevrus A. Costa, 1859 (Guichard 1986; Nemkov 2001), however, in a cladistic analysis of the subfamily Bembicinae (based on morphological characters) carried out by Nemkov & Lelej (2013), this genus was synonymized with Nysson Latreille.Another opinion was that of W. Pulawski (pers. comm.), who attributed this to the presence of intermediate cases in some species. The genus is a new record to the Saudi Arabian fauna. The new species agrees with N. barrei (= Synnevrus barrei) in Nemkov’s key (2001: 4, couplet 4) in having two lateral small teeth on scutellum; pronotum sharply angulate antero-laterally (Fig. 4A, B); however, it differs from N. barrei in having thorax mainly black, with few areas ferruginous (Figs 3A, 4B) (thorax mainly ferruginous in N. barrei). Based on Radoszkowski (1893: 71), our male specimen differs from the male N. barrei in having posterior margin of prothorax dark ferruginous, not interrupted medially (Fig. 4A) (with ivory white band, interrupted medially in N. barrei); mesoscutum, scutellum (except ferruginous at base) entirely black (Fig. 4B) (reddish in N. barrei); T 1 with two postero-lateral oval yellow spots, posterior margins of T 2–5 with yellow bands, interrupted medially on T 2 (Figs 3A, 5C) (posterior margins of T 1–4 with ivory-white bands, interrupted medially in N. barrei); gaster ventrally mostly dark ferruginous, except basal two thirds of S 1 black (Fig. 5B) (S 2 reddish in N. barrei); fore wing slightly darkened, fumigated at apex (Fig. 5A) (transparent in N. barrei); only hind tibia along its whole dorsal side (all tibiae ivory white dorsally in N. barrei). On the other hand, the male N. arabicus somewhat agrees with the male N. ohli (Schmid-Egger 2011: 592) in having the sharply angulate antero-lateral sides of pronotum (Fig. 4A); the same body sculpturing (Fig. 3A); the presence of a median bulge between spines at apex of T 7 (Fig. 5C). However, it differs from N. ohli in the following combination of characters: body length 6.0 mm (4.0 mm in N. ohli); antenna with scape yellow ventrally (Fig. 3C) (whole antenna reddish, darkened apically in N. ohli); pronotum dark ferruginous posteriorly reaching dark ferruginous pronotal lobes (Fig. 4A) (light yellow in N. ohli reaching the dark reddish pronotal lobe); scutellum dark ferruginous at base (Fig. 4B) (clear yellow in N. ohli); T 2–5 with yellow bands posteriorly that is interrupted medially on T 2 (Figs 3A, 5C) (T 2 & T 3 with narrower lateral yellow spots, rest of terga black in N. ohli); sternal colour differs in the two species; terga medially without medial bulges (with small medial bulges in N. ohli); S 2 with a distinct, large rounded hump near to middle (absent in N. ohli); hind tibia broadly yellow dorsally (Fig. 3A) (all tibia with narrow light yellow over whole length dorsally in N. ohli). T 6 & T 7 entirely reddish (Fig. 5C) (T 6 partly and T 7 reddish in N. ohli).
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10. Solierella nigridorsum Pulawski 1964
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Solierella nigridorsum ,Taxonomy ,Solierella - Abstract
Solierella nigridorsum Pulawski, 1964 Fig. 12A–F Solierella nigridorsum Pulawski, 1964: 136, ♀. Diagnosis. Body length: 3.6 mm. This species is characterized by the entirely black thoracic dorsum (Fig. 12A, C). Material examined: 1♀, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Distribution: Egypt, Iran, Saudi Arabia (new record), United Arab Emirates. Comments. This species is a new record for Saudi Arabia. Characters of the only collected specimen agree well with Pulawski’s description of the Egyptian females (1964: 136). It also agrees with the Arabian S. nigridorsum in having the metanotum completely black (Schmid-Egger 2011). Furthermore, it agrees with the Iranian female in having the pronotum and metanotum black (Schmid-Egger et al. 2021).
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- 2023
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11. Further addition to the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae), with new genera and species records, and the description of two new species
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., Edmardash, Yusuf A. (2023): Further addition to the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae), with new genera and species records, and the description of two new species. Zootaxa 5319 (2): 151-177, DOI: 10.11646/zootaxa.5319.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5319.2.1
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12. Solierella Spinola 1851
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy ,Solierella - Abstract
Genus Solierella Spinola, 1851 Solierella Spinola, 1851: 349. Type species: Solierella miscophoides Spinola, 1851, by monotypy. This is the first record of the genus for the Saudi Arabian fauna.
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- 2023
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13. Tachysphex Kohl 1883
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Tachysphex ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Tachysphex Kohl, 1883 Tachysphex Kohl, 1883: 166. Type species: Tachysphex filicornis Kohl, 1883 [= Tachytes fugax Radoszkowski, 1877], designated by Bingham, 1897: 192.
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14. Tachysphex consocius Kohl 1892
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Tachysphex ,Arthropoda ,Animalia ,Biodiversity ,Tachysphex consocius ,Hymenoptera ,Taxonomy - Abstract
Tachysphex consocius Kohl, 1892 Fig. 13A–G Tachysphex consocius Kohl, 1892: 217, ♀. Diagnosis. Body length: 7.0 mm. Body, including legs and antennae, black (Fig. 13A); pygidial plate reddish apically in some females; mandible reddish medially, black at base and apex (Fig. 13B); labrum flat, concealed under clypeus; middle clypeal section of female convex (Fig. 13B), with one lateral incision, free margin sinuate; postocellar area with erect setae; mid tarsomere II more than twice as long as apically wide (Fig. 13A); mesopleuron distinctly punctate, punctures minute; propodeal dorsum irregularly rugose or irregularly ridged (Fig. 13C); gastral T 1–4 silvery fasciate apically (Fig. 13A); dorsal length of female flagellomere I about 2.3 × apical width; outer apical spine of male foretarsomere II longer than foretarsomere III. Material examined: 2♀, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Previous Saudi Arabian records: Fayfa (Pulawski, 2007). Distribution: Widely distributed in Africa, southern Europe to Central Asia and India and Sri Lanka. Comments. Characters of this species agree with Pulawski’s key (2007: 49, couplet 103), as well as the diagnosis recognizing the species in Pulawski (2007: 208).
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15. Tachysphex fugax
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Tachysphex ,Arthropoda ,Tachysphex fugax ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Tachysphex fugax (Radoszkowski, 1877) Fig. 14A–F Tachytes fugax Radoszkowski, 1877: 30, ♁. Diagnosis. Body length: 7.0 mm. Body entirely black (including antennae and legs) (Fig. 14A–C, E); clypeal lobe flat, with at most a rudimentary bevel in female, angulate laterally in male; labrum greatly concealed by clypeus; antennal flagellomeres elongate, distinctly longer than broad, without sensory areas (Fig. 14B, C); length of flagellomere IV 4.0–4.5× apical width in female and 2.2–2.8× in most males; frons punctatorugose; mesoscutum and mesopleuron densely punctate (Fig. 14C, D), with puncture one diameter apart, much denser posteriorly at junction with scutellum; propodeum longitudinally ridged, setae directed obliquely anterad (Fig. 14D); hind coxa more or less expanded at base dorsally; bottom of male fore femoral notch glabrous, microscopically ridged; outer margin of fore basitarsus in most males with 3–5 rake spines; outer apical spine of male tarsomere II no longer than tarsomere III; T 1–3 with apical silvery fasciae (Fig. 14A). Material examined: 1♀, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022. Previous Saudi Arabian records: Adriyah, El Riyadh, Fayfa, South Jeddah, Wadi Maraba (Pulawski 2007). Distribution: Algeria, Angula, Austria, Azerbaijan, Botswana, Bulgaria, Burkina Faso, Burundi, Canary Islands, Central African Republic, Comoro Islands, Croatia, Cyprus, Democratic Republic of the Congo, Egypt, Ethiopia, France, Georgia, Greece, Guinea, Hungary, Israel, Italy, Ivory Coast, Kazakhstan, Kenya, Lesotho, Madagascar, Mali, Malta, Morocco, Mozambique, Namibia, Portugal, Romania, Russia, Saudi Arabia, Senegal, Serbia, Seychelles, Sierra Lione, Somalia, South Africa, Spain,? Switzerland, Syria, Tanzania, Tunisia, Turkey, Ukraine, Uzbekistan, Zimbabwe. Comments. Characters of the only female collected from Farasan agrees well with Pulawski’s key (2007: 48, couplet 98).
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16. Ammatomus yemenensis Schmid-Egger 2019
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Ammatomus ,Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Ammatomus yemenensis ,Taxonomy - Abstract
Ammatomus yemenensis Schmid-Egger, 2019 Fig. 2A–F Ammatomus yemenensis Schmid-Egger, 2019: 448, ♀, ♁. Diagnosis. Body length: 8.0 mm. This species is diagnosed by the following characters: gastral T 1 distinctly petiolate, with distinct constriction between T 1 and T 2 (Fig. 2A); metanotum ivory, smooth; T 2–4 with complete ivory bands (Fig. 2A); T 1 about 1.5× as long as apical width, with two large posterolateral ivory markings a short distance apart, remainder black, posterior margin translucent; S 2 with two small triangular ivory markings posterolaterally (Fig. 2E); hind tarsomeres I–III all yellowish to whitish (Fig. 2A, E); hind claws black (Fig. 2E); T 1 without or with indistinct lateral carina. Male: Not available. Material examined: 1♀, Farasan Island Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022 Distribution: Yemen (Schmid-Egger 2019), Saudi Arabia (new record). Comments: Ammatomus yemenensis Schmid-Egger from the Farasan Archipelago is a new record for Saudi Arabia. The characters of the only specimen collected agree well with A. yemenensis in Schmid-Egger’s key (2019: 440, couplet 11). It differs from the Yemeni female in having the pale parts of body ivory white (whitish yellow in Yemeni specimens); antennal flagellum entirely black except last two flagellomeres reddish beneath (AS 1–3 yellowish white in Yemeni specimens); T 1 1.29× as long as mesoscutum (1.1× mesoscutum in Yemeni specimens). It also agrees with Schmid-Eggers’ (2019: 456, figs 41–44).
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- 2023
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17. Nysson Latreille 1802
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Nysson ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Nysson Latreille, [1802] Nysson Latreille, [1802: 340]. Type species: Crabro spinosus Fabricius, 1775 [= Sphex spinosus J. Forster, 1771], designated by Schuckard, 1837. This is the first record of the genus for the Saudi Arabian fauna.
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- 2023
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18. Ammatomus A. Costa 1859
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Ammatomus ,Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Ammatomus A. Costa, 1859 Ammatomus A. Costa, 1859: 36. Type species: Gorytes coarctatus Spinola, 1808, by monotypy. This is the first record of the genus for the Saudi Arabian fauna.
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19. Trypoxylon splendidum Antropov 2011
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Trypoxylon splendidum ,Animalia ,Biodiversity ,Trypoxylon ,Hymenoptera ,Taxonomy - Abstract
Trypoxylon splendidum Antropov, 2011 Fig. 16A–G Trypoxylon splendidum Antropov, 2011: 610, ♀, ♁. Diagnosis. Body (including antennae and legs) entirely black (Fig. 16A); frontal shield in female and male relatively long, pointed, a short distance above antennal bases, rounded basally and angulate apico-laterally, coarsely rugose in female (Fig. 16B), densely punctate in male (Fig. 16C), with an area of silvery pubescence extending from middle ocellus but not reaching ventral end of shield (Fig. 16B, C); last antennal segment in male relatively long, narrowly rounded apically, about as long as preceding three antennal segments or slightly shorter (Fig. 16G); scutum with dense superficial punctures (Fig. 16D), mesopleuron densely punctate, with interspaces (Fig. 16E); metapleuron finely transversely striate (Fig. 16E); dorsal enclosure of propodeum coarsely rugose, with dense, upturned fine pale setae posteriorly (Fig. 16D); first gastral segment distinctly long in both sexes (Fig. 16F); S 8 of male completely as in Antropov (2011: 615, fig. 10). Material examined: 2♀, 1♁, Farasan, Abdulmajeed farm [16 Q 42.07970’N; 42 Q 08.5781’E], 23–30.ix.2022; 1♀, Farasan (Abdulmajeed Farm), 30.ix–7x.2022. Previous Saudi Arabian records: Maha’il (Antropov 2011), Jazan (Gadallah et al. 2021). Distribution: Egypt, Israel, Lebanon, Oman, Palestine, Qatar, Saudi Arabia, United Arab Emirates. Comments. Characters of this species fully agree with Antropov’s key (2001: 616, couplet 3), as well as his diagnosis (2011: 610).
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20. Miscophus sauditus Gadallah & Edmardash 2023, sp. nov
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Abu El-Ghiet, Usama M., Gadallah, Neveen S., Gasib, Abdulmajeed M., Al-Fifi, Zarrag I. A., and Edmardash, Yusuf A.
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Crabronidae ,Miscophus ,Insecta ,Arthropoda ,Animalia ,Miscophus sauditus ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Miscophus sauditus Gadallah & Edmardash sp. nov. Figs 8A–E, 9A–D, 10A–E Type material: SAUDI ARABIA. Holotype (♁), Farasan [16° 41.8’ N; 42° 7.3333’ E], 23–30.ix.2022. Description of male holotype: body length: 7.0 mm. Colouration: head and thorax black with cuprous to dark green luster (Figs 8A, B, 9A, B); red: gaster (Figs 8A, 9C, D), tegula (Figs 8A, 9A) and legs (except coxae externally black) (Fig. 8A); scape and pedicel yellowish, darker above (Fig. 8B–D); flagellum wholly black (Fig. 8B–D); apical margin of clypeus ferruginous (Fig. 8C), mandible ferruginous with black tip; forewing membrane hyaline to subhyaline, with a transverse fumigate area subapically (not reaching wing apex), veins dark brown (Fig. 10E). Head (Fig. 8A–E): face above antennal sockets and clypeus densely clothed with silvery pubescence concealing underlying integument (Fig. 8C); upper face with less pubescence, not concealing integument, densely punctaterugose (Fig. 8C); eye orbit slightly concave above, convex below (Fig. 8C); clypeus medially with apex slightly arcuate (Fig. 8C); last three flagellomeres depressed dorsally (Fig. 8E); middle ocellus distinctly larger than lateral ocelli (Fig. 8B); AS 3 2.5× as long as its apical width (Fig. 8E), distinctly longer than pedicel. Thorax (Fig. 9A, B): pronotum well-developed, with slight longitudinal sulcus medially (Fig. 9A); mesoscutum densely punctate, with indistinct short sulcus antero-medially (Fig. 9A); scutellum densely punctate, glabrous (Fig. 9A); propodeum with dense silvery pubescence especially laterally, densely granulate (Fig. 9A), transversely punctate posteriorly; propleuron with dense silvery pubescence concealing underlying integument (Fig. 9B); mesopleuron with such pubescence anteriorly and medially, remainder smooth and shiny (Fig. 9B); legs (Figs 8A, 9B): fore coxa with two longitudinal carinae externally, depressed in between; fore basitarsus long and slender, as long as following tarsomeres combined, with four black unequal spines, of which basal one is very short, apical spine is the longest, about as long as following tarsomere; spines of tarsal comb thin (not spatulate); fore femur with wavy silvery pubescence ventrally; mid and hind tibiae irregularly spinose. Gaster (Figs 9C, D, 10A–C) impunctate, terga with fine silvery, short pubescence laterally (Figs 9C, 10A); T 6 conical, broad and convex at base, with numerous short, black spines along lateral margins (Fig. 10A, B); S 3–6 each with a row of black spines posteriorly, increasing in number toward apex (Fig. 9D), S 8 with two long spines curved along almost their whole length, parallel-sided at apex (Fig. 10C); gonoforceps (Fig. 10D) narrowly rounded apically, with dense, long setae at basal two-thirds, distinctly bent medially (Fig. 10D). Female: Unknown. Etymology: This species is named after the Kingdom of Saudi Arabia where the specimen was collected. Distribution: Known only from Saudi Arabia (Farasan Archipelago). Comments. The new species belongs to Miscophus helveticus group based on Schmid-Egger & El-Jahdhami’s key to males species-groups (2022: 324, couplet 3) in having erect whitish pilosity on gena (Fig. 8D, E), and S 8 bidentate (Fig. 10C). It greatly resembles the male of M. papyrus de Andrade, but differs from it in the following: clypeus slightly arcuate (Fig. 8C) (clypeus mesally pointed at apex in M. papyrus, Pulawski 1964: 124, fig. 62); propodeal dorsum densely granulate (striate in M. papyrus; Pulawski 1964); gaster entirely red (T 4 and T 5 clearly brownish in M. papyrus; Pulawski 1964); S 8 with two long spines curved along almost their whole length, parallel-sided at apex (Fig. 10C) (spines parallel-sided throughout in M. papyrus, see Pulawski 1964: 123, fig. 63); gonoforceps with dense long setae that are distinctly bent medially (Fig. 10D) (bent at apex in M. papyrus; see Pulawski 1964: 123, fig. 64). It also resembles M. ctenopus Kohl in having posterior ocelli closer to each other than the distance between each of them and the posterior ocellus (Fig. 8B); radial cell distinctly larger than the petiolate SMC 2 (Fig. 10E). However, it differs from M. ctenopus in the following (based on Kohl’s key to males, 1885: 221, couplet 2): mesoscutum dorsally with noticeable side corners in M. ctenopus (side corners gently rounded in M. sauditus, Fig. 9A); fore wing in M. ctenopus light, heavily clouded at tip margin (in M. sauditus fore wing more or less subhyaline, heavily clouded subapically, Fig. 10E); gaster bright red, except brownish medially in M. ctenopus (in M. sauditus gaster entirely red, Figs 8A, 9C, D). Based on de Andrade (1954): M. sauditus differs from the male M. ctenopus in the following aspects: free margin of clypeal lobe normal (Fig. 8C) (depressed medially and slightly raised at the sides in M. ctenopus); spines of tarsal comb thin (not spatulate) (spatulate in M. ctenopus); lower edge of the apical zone of gonoforceps distinctly curved (Fig. 10D) (not markedly curved in M. ctenopus, see de Andrade (1954: 11, fig. 25)); gonoforceps with dense long setae that are distinctly bent or curved medially (Fig. 10D) (somewhat irregularly wavy in M. ctenopus; see de Andrade 1954: 11, fig. 25).
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21. Philanthinus de Beaumont 1949
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthinus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Genus Philanthinus de Beaumont, 1949 Philanthinus de Beaumont, 1949: 194 (as subgenus of Philanthus). Type species: Philanthus integer de Beaumont, 1949, by original designation and monotypy. The genus Philanthinus is represented in Egypt by a single species, P. integer de Beaumont., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 159, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216."]}
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22. Philanthus ammochrysus W. Schulz 1905
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Philanthus ammochrysus ,Hymenoptera ,Taxonomy - Abstract
Philanthus ammochrysus W. Schulz, 1905 Fig. 4A–E Philanthus ammochrysus W. Schulz, 1905: 59, ♁. Diagnosis. Body length: 11 mm (male). Body generally bright yellow with the following black: ocellar triangle extending to inner eye margin above emargination, large spot above antennal sockets, spot at posterior eye margin that narrows posteriorly (Fig. 4A– E); F3–11 (except F3 ventrally) (Fig. 4B), clypeal moustache (Fig. 4B), three longitudinal bands on mesoscutum connected basally and apically (Fig. 4D), scutellum and postscutellum laterally (Fig. 4C), basal triangle of propodeum (Fig. 4D), mesosternum laterally, large irregular area on metapleuron, dorsal basal streak on hind femur (Fig. 4E), apical margins of gastral terga. Gastral T 3 yellow with two black markings baso-laterally, T 6 black with narrow yellow band apically (Fig. 4E), S 2 with black bands apically and basally, with two small black markings basolaterally, S 3 –S 5 with black band basally, S 6 black. Face densely punctate (Fig. 4B); clypeal moustache long and dense, meeting medially (Fig. 4B); mesoscutum sparsely punctate (Fig. 4D), scutellum and postscutellum smooth and shiny (Fig. 4D); propodeal dorsum smooth and shiny, with median rugose suture (Fig. 4D); upper mesopleuron smooth, lower mesopleuron densely punctate (Fig. 4C); mesosternum coarsely punctate; hind coxa without keel; gastral T 1–3 densely punctate, with large interspaces, punctures sparser and superficial on rest of terga (Fig. 4E); gastral S 2 mostly smooth, dull; S 3–6 coarsely punctate. Male genitalia (see Fig. 23A). Female: Not available. Material examined: 1♁, Wadi Degla, 21.vi.2022, on Ochradenus baccatus Delile (Resedaceae), Alyaa A. Malash collector (EFC). Distribution in Egypt: Wadi Degla (present study). Extralimital distribution: Algeria, Jordan, Libya, Morocco, Tunisia. Comment: This species is a new record for Egypt. Characters of the only specimen collected agree with de Beaumont’s key (1949: 180, couplet 9). It also agrees with de Beaumont’s (1949: 178, fig. 5) except in the absence of the two median black spots between antennal sockets. It differs from the Tunisian male in having apical margins of gastral terga black, and T 3 with two black markings laterally (in Tunisian male, apical markings of terga are ferruginous, T 3 has no black markings laterally, see Ben Khedher et al. 2021: 7, fig. 1B), but agrees with fig. 3B (Ben Khedher et al. 2021: 8)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 160-162, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Schulz, W. A. (1905) Hymenopteren - Studien. Wilhelm Engelmann, Leipzig, 147 pp.","de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216.","Ben Khedher, H., Yildirim, E. & Braham, M. (2021) A checklist of Philanthini and Pseudoscoliini (Hymenoptera: Crabronidae) with new records from Tunisia. Transactions of the American Entomological Society, 147, 1 - 10. https: // doi. org / 10.3157 / 061.147.0101"]}
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23. Philanthus minor Kohl 1891
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Philanthus minor ,Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus minor Kohl, 1891 Figs 9A–D, 10A–E Philanthus minor Kohl, 1891: 358, ♀. Diagnosis. Body length: 12 mm (female), 12 mm (male). Female with head and thorax black and following parts yellow: postero-lateral margin of head, lower face (not reaching inner eye emargination laterally, and forming trifid marking medially between antennal bases, far from median ocellus), clypeus, basal half of mandible, first three antennal segments, and inner side of fourth segment, pronotum, scutellum, postscutellum, tegula, upper mesopleuron; gaster with all terga yellow banded with ferruginous and black; legs yellow and ferruginous (Figs 9A–C, 10A). The yellow is more developed in male, extending to middle ocellus, and with two small, rounded spots on head posterior margin; mesoscutum with two middle as well as lateral yellow makings, yellow of mesoscutum interrupted medially; first three gastral terga largely yellow, surrounded with light ferruginous, T 4–6 mainly black, each with narrow, medially interrupted yellow band posteriorly (Figs 9D, 10B–E). Male clypeal moustache dense, meeting medially (Fig. 9D); pronotum distinctly emarginate medially (Fig. 9B, C); propodeal enclosure with coarsely sculptured, broad median sulcus, sculpture covering whole basal area of propodeum as inverted triangle (Fig. 10A, E); upper mesopleuron smooth and shiny (Fig. 10C, D); hind femur with relatively long fine setae, about as long as greatest femur width (Figs 9A, 10A). Material examined: 1♀, Dahshour, 15.iii.1927 (AUCE); 1♁, Burg, 4.V. 1955, Sh. M. collector (EFC). Previous Egyptian Records: Dahshour (Kohl 1891; Honoré 1942; de Beaumont 1949; Roche 2007), no specific locality (Dalla Torre 1897; de Beaumont 1953), Marsa Matruh (Guichard 1994; Gadallah 1996; Roche 2007). Extralimital distribution: Algeria, Chad, Mali, Mauritania, Morocco, Tunisia, Sudan, United Arab Emirates. Remarks. Characters of Egyptian specimens agree with Guichard’s key (1994: 208, couplet 8), however, in the male, the ferruginous extends along the first three gastral terga (Fig. 10E). They also agree with the North African specimens of de Beaumont (1949: 179, couplet 8)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 165, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Kohl, F. F. (1891) Zur Kenntnis der Hymenopteren - Gattung Philanthus Fabr. (sens. lat.). Annalen des k. k. Naturhistorischen Hofmuseums, 6, 345 - 370.","Honore, A. (1942) Introduction a l'etude des Sphegides en Egypte (Hymenoptera: Aculeata). Bulletin de la Societe Fouad Ier d'Entomologie, 26, 25 - 80.","de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149.","Dalla Torre, K. W. (1897) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], 749 pp.","de Beaumont, J. (1953) Hymenopteres recoltes par une mission suisse au Maroc (1947). Sphecidae 2. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 32, 107 - 131.","Guichard, K. M. (1994) The genera Philanthus and Philanthinus (Hymenoptera: Sphecidae) in Arabia. Fauna of Saudi Arabia, 14, 207 - 212.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80."]}
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24. Philanthus pallidus Klug 1845
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus pallidus ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus pallidus Klug, 1845 Figs 11A–E, 12A, B. Philanthus pallidus Klug, 1845: [28], plate 47, fig. 8, ♁. Diagnosis. Body length: 11–13 mm (female); 10–13 mm (male). Body entirely yellow, except the following black in the male: flagellomeres 2–11, a thin streak above eye emargination, posterior margins of mesoscutum and scutellum, scutellum laterally; median sulcus of propodeum; posterior margin of T 1 and T 2 ferruginous (Figs 11A–E, 12A, B). Lower face sparsely punctate, while deeply and densely punctate above eye emargination (Fig. 11B, D, E); malar space very short, but distinct (Fig. 12A); clypeal moustache long, meeting medially (Fig. 11D, E); clypeus with but few fine setae directed above; upper mesopleuron (mesepimeron) with sparse microscopic punctures to nearly smooth, lower mesopleuron densely coarsely punctate (Fig. 12A, B); propodeal enclosure smooth and shiny, with rugose or shagreened median sulcus (Fig. 11A, C). Material examined: 1♀, Gebel Elba (Wadi Aideb), 28–29.ii.1938 (PPDD); 5♁, 2♀, Gebel Elba (Wadi Aideb), 5.iii.1938 (PPDD); 3♁, Gebel Elba (Wadi Aideb), 7.iii.1938 (PPDD); 1♀, Kafr Asfar, 4.vi.1937 (AUCE); 1♁, EL Gabal El Asfar, 1.viii.1937; 1♁, Kafr Farouk, 9.vii.1939 (AUCE); 1♁, Giza (without date) (AUCE); 2♀, El Gabal El Asfar, June (no year) (AUCE). Previous Egyptian Records: Abbasyiah (Kohl 1891; Dalla Torre 1897), no specific locality (Arnold 1925; Mochi 1939; de Beaumont 1949, 1953, 1956; Bohart & Menke 1976; Guichard 1994), El Gabal El Asfar (Honoré 1942), Gebel Elba (Wadi Aideb) (Gadallah 1996), Kerdasa, Kom Oshiem (Roche 2007), Wadi El Tih (Dollfuss 2017). Extralimital distribution: Algeria, Ethiopia, Eritrea, Iran, Israel-Palestine, Mauritania, Morocco, Oman, Saudi Arabia, Spain, Sudan, United Arab Emirates. Remarks. The characters of Egyptian specimens agree with the North African specimens of de Beaumont (1949: 179, couplet 9). They also agree with the Arabian species of Guichard (1994, couple 8, p. 208). However, they differ from those of Guichard’s in having the malar space very short but significant (Fig. 12A). In some specimens, face and thorax laterally are whitish (Figs 11B, E, 12A)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 170-172, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Klug, F. (1845) Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex Itinere per Africam Borealem et Asiam Occidentalem Frederici Guilelmi Hemprich et Christiani Godofredi Ehrenberg Medicinae et Chirurgiae Doctorum Studio Novae aut Illustratae redierunt. Decas Quinta. Officina Academica, Berolini, 41 pp., pls. 41 - 50.","Kohl, F. F. (1891) Zur Kenntnis der Hymenopteren - Gattung Philanthus Fabr. (sens. lat.). Annalen des k. k. Naturhistorischen Hofmuseums, 6, 345 - 370.","Dalla Torre, K. W. (1897) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], 749 pp.","Arnold, G. (1925) The Sphegidae of South Africa. Part VI. Annals of the Transvaal Museum, 11, 137 - 175.","Mochi, A. (1939) Revisione delle Specie egiziane dei Generi Philanthus F. e Nectanebus Spin. (Hymenoptera: Sphegidae - Philanthinae). Bulletin de la Societe Fouad Ier d'Entomologie, 23, 86 - 139.","de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216.","de Beaumont, J. (1953) Hymenopteres recoltes par une mission suisse au Maroc (1947). Sphecidae 2. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 32, 107 - 131.","de Beaumont, J. (1956) Sphecidae (Hym.) recoltes en Libye et au Tibesti par M. Kenneth M. Guichard. Bulletin of the British Museum (Natural History). Entomology, 4, 165 - 215.","Bohart, R. M. & Menke, A. S. (1976) Sphecid wasps of the world. A generic revision. University of California Press, Berkeley, Los Angeles and London, 1 color pl., IX + 695 pp. https: // doi. org / 10.1525 / 9780520309548","Guichard, K. M. (1994) The genera Philanthus and Philanthinus (Hymenoptera: Sphecidae) in Arabia. Fauna of Saudi Arabia, 14, 207 - 212.","Honore, A. (1942) Introduction a l'etude des Sphegides en Egypte (Hymenoptera: Aculeata). Bulletin de la Societe Fouad Ier d'Entomologie, 26, 25 - 80.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149.","Dollfuss, H. (2017) The Philanthinae wasps of the \" Biologiezentrum - Linz \" - collection in Linz, Austria, including the genera Aphilanthops Patton, Clypeadon Patton, Odontosphex Arnold, Philanthinus de Beaumont, Philanthus Fabricius, Pseudoscolia Radoszkowski and Trachypus Klug. Linzer Biologische Beitrage, 49, 1245 - 1274."]}
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25. Philanthus sulphureus F. Smith 1856
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Philanthus sulphureus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus sulphureus F. Smith, 1856 Philanthus sulphureus F. Smith, 1856: 469, ♁. Remarks. This species was described from northern India. In his list of insects probably collected by J. Lord in Egypt and determined by F. Walker, Innes Bey (1912 [1911]) recorded P. sulphureus from Egypt based on Walker’s label as sulfureus collected from Wadi Feiran (as Wady Ferran, South Sinai). Walker’s specimen was destroyed by dermestids, therefore, it must be regarded as being a doubtful record., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 184, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Smith, F. (1856) s. n. In: Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part IV. Sphegidae, Larridae and Crabronidae. Taylor and Francis, London, pp. 207 - 497.","Innes Bey, W. (1912 [1911]) Une liste d'especes recueillis probablement par S. Lord en Egypte et determinees par F. Walker. Bulletin de la Societe Fouad Ier d'Entomologie, 1911, 97 - 115."]}
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26. Philanthus variegatus subsp. ecoronatus Dufour 1854
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus variegatus ,Animalia ,Biodiversity ,Philanthus variegatus ecoronatus dufour, 1854 ,Hymenoptera ,Taxonomy - Abstract
Philanthus variegatus ecoronatus Dufour, 1854 Figs 20A–C, 21A–C Philanthus ecoronatus Dufour, 1854: 380, ♁. Diagnosis. Body length: 14 mm (female); 11.6 mm (male). Philanthus variegatus ecoronatus differs from the nominotypical subspecies in the following: mesonotum shiny black (Fig. 21B) (shiny ferruginous in P. variegatus variegatus (Fig. 19D)); scutellum and postscutellum ferruginous (Fig. 21B), propodeum black with two lateral yellow patches (Fig. 21B) (rest of thorax yellow in P. variegatus variegatus, except black median sulcus (Fig. 19C–E)); mesoscutum sparsely punctate, punctures becoming dense posteriorly (Fig. 21B) (nearly smooth, impunctate in P. variegatus variegatus, densely punctate at junction with scutellum (Fig. 19D)); gastral T 2 densely punctate (Fig. 21C) (sparsely punctate in P. variegatus variegatus (Fig. 19E)). Material examined: 1♀, Wadi El-Natroun, 12.v.1990, Neveen Gadallah collector (EFC); 1♀, Wadi El Natroun (Beo Ar Ashra), (18/94?) (EFC); 1♁, Marsa Matruh, 28.ix.1954, Aly collector (ASUE). Previous Egyptian records: Wadi El-Natroun (Gadallah 1996); Marsa Matruh (present study). Extralimital distribution: Algeria, Armenia, Chad, Italy, Libya, Morocco, Palestine, Tunisia, Comment: This subspecies was recorded for the first time from Egypt by Gadallah (1996). The characters agree with de Beaumont’s key (1949: 179, couplet 6) in having mesopleuron and propodeum greatly marked with yellow (Fig. 21A, B); upper mesopleuron densely punctate (Fig. 21A); gaster mostly yellow, narrowly black medially on T 1–4, with terminal depressions ferruginous (Fig. 21C); T 2 more or less densely, finely punctate (Fig. 21C); face entirely pale up to level of inner eye emargination in female (Fig. 20B), and dark reddish brown to ferruginous in male (Fig. 20C); scutellum and postscutellum ferruginous (Fig. 21B). It differs from de Beaumont description in having the pale colour of the face ivory white and extending very close to the middle ocellus (in de Beaumont’s specimens, 1949: 178, the pale colour of the face is yellow, and touching the middle ocellus, see fig. 1). The Tunisian specimen (Ben Khedher et al. 2021: 8, fig. 3J) is identical to de Beaumont’s. This may be a sexual difference (the Egyptian specimen is a female, while de Beaumont’s 1949 and Ben Khedher et al. 2021, specimens are males). The specimen of Wadi El Natroun, 12.v.1990 (EFC) was misidentified by Giles Roche as P. dufouri Lucas. However, it differs from P. dufouri in having the scutellum ferruginous (Fig. 21B) (clearly black in P. dufouri); gastral terga predominantly yellow, narrowly black medially (Fig. 21C) (black in P. dufouri, with yellow markings clearly delimited)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 179-182, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Dufour, L. (1854) Signalements de quelques especes nouvelles ou peu connues d'Hymenopteres algeriens. Annales de la Societe Entomologique de France, Serie 3, 1, 375 - 382.","Lepeletier de Saint Fargeau, A. L. M. (1845) Histoire naturelle des Insectes. Hymenopteres. Vol. 3. Librairie Encyclopedique de Roret, Paris, 646 + 4 pp., pls. 25 - 36.","Spinola, M. (1839) Compte-rendu des Hymenopteres recueillis par M. Fischer pendant son voyage en Egypte, et communiques par M. le Docteur Waltl a Maximilien Spinola. Annales de la Societe Entomologique de France, 7, 437 - 546.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80.","Ben Khedher, H., Yildirim, E. & Braham, M. (2021) A checklist of Philanthini and Pseudoscoliini (Hymenoptera: Crabronidae) with new records from Tunisia. Transactions of the American Entomological Society, 147, 1 - 10. https: // doi. org / 10.3157 / 061.147.0101","de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216."]}
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27. Philanthus schulthessi Maidl 1924
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus schulthessi ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus schulthessi Maidl, 1924 Figs 15A–C, 16A, B, 17A–D Philanthus schulthessi Maidl, 1924: 242, ♀. Diagnosis. Body length 11–13.5 mm (female). Greatly resembles female of P. minor, but differs from that species in having the body punctures conspicuously denser (Figs 15A–C, 16A, B, 17A–D); metapleuron densely pilose (inconspicuous in P. minor) (Fig. 16A); colour pattern of gaster (Fig. 16C, 17B); antenna whitish basally and pale reddish at the rest (Figs 15B, 17B) (basal segments yellowish and the rest of flagellum black in P. minor (Fig. 9C)); gastral terga with conspicuously denser punctures, interspaces smaller than puncture diameter, thus appearing dull (Figs 16C, 17B) (less dense, with larger interspaces and shiny in P. minor (Fig. 9A)). Material examined: 1♀, Marsa Matruh, 15.ix.1957, Aly collector (ASUA); 1♀, Marsa Matruh, vii.1952, Dr. T. S. collector (ASUA); 1♀, Marsa Matruh, 20.viii.1955, Aly collector (ASUA) (all were mistakenly identified as P. minor); a photograph from the OÖLM: 1♀, Marsa Matruh, 22–29.viii.1954, collector? (det. W.J. Pulawski, 1978). Previous Egyptian records: Marsa Matruh (Dollfuss 2017; present study). Extralimital distribution: Chad, Libya Mauritania, Sudan. Remarks. In his Mauritanian paper, de Beaumont (1953) has mistakenly identified P. schulthessi as P. minor. However, because of some differences between the specimens examined by him and the true minor (the ocelloocular distance; the degree of emargination of pronotum; the dense punctures; and the colour of gastral T 1 and T 2) he treated it as a subspecies of P. minor. In 1956, comparing his Mauritanian specimens with schulthessi from Wadi Wour (an island near Khartoum, Sudan), he found them conspecific, and thus he corrected the Mauritanian minor subsp. of 1953 to be P. schulthessi. The specimens examined from the ASUA collection were identified as P. minor, by comparing them with the photos of P. schulthessi received from OÖLM, they were found to be conspecific. Their characters also agree well with the Libyan specimen of de Beaumont’s P. schulthessi (1956:184) and to a large extent with figs 15, 16 (de Beaumont, 1956:184). All specimens under study as well as that photographed by OÖLM were collected from Marsa Matruh between May and September (1952–1957)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 172-178, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Maidl, F. (1924) Wissenschaftliche Ergebnisse der mit Unterstutzung der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Agyptischen Sudan (Kordofan) 1914. XV. Hymenoptera E. Scoliidae et Sphegidae. Denkschriften der Akademie der Wissenschaften in Wien. MathematischNaturwissenschaftliche Klasse, Abteilung I, 99, 233 - 246.","Spinola, M. (1839) Compte-rendu des Hymenopteres recueillis par M. Fischer pendant son voyage en Egypte, et communiques par M. le Docteur Waltl a Maximilien Spinola. Annales de la Societe Entomologique de France, 7, 437 - 546.","Dollfuss, H. (2017) The Philanthinae wasps of the \" Biologiezentrum - Linz \" - collection in Linz, Austria, including the genera Aphilanthops Patton, Clypeadon Patton, Odontosphex Arnold, Philanthinus de Beaumont, Philanthus Fabricius, Pseudoscolia Radoszkowski and Trachypus Klug. Linzer Biologische Beitrage, 49, 1245 - 1274.","de Beaumont, J. (1953) Hymenopteres recoltes par une mission suisse au Maroc (1947). Sphecidae 2. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 32, 107 - 131.","de Beaumont, J. (1956) Sphecidae (Hym.) recoltes en Libye et au Tibesti par M. Kenneth M. Guichard. Bulletin of the British Museum (Natural History). Entomology, 4, 165 - 215."]}
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28. Philanthus rutilus Spinola 1839
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Philanthus rutilus ,Taxonomy - Abstract
Philanthus rutilus Spinola, 1839 Figs 13A–C, 14A–C. Philanthus rutilus Spinola, 1839: 488, ♀, ♁. Diagnosis. Body length: 12.4 mm (female); 12.5 mm (male). Body relatively large, with body in female predominantly black, with tegula, scutellum, posterior margins of T 3–5 and legs ferruginous; clypeus and lower face yellow not reaching middle ocellus (Figs 13A, 14A). Male with clypeus and face whitish, whitish extending to middle ocellus; thorax entirely black (tegula yellow), gaster with T 1 ferruginous, while T 2 and T 3 yellow, with intersegments and base of T 1 black; T 4 &T 5 almost black (with very thin, hardly seen yellow streaks postero-laterally), F3–11 black; face and clypeus ivory white (Figs 13B, 14B, C). Face and clypeus with dense, fine and long pilosity (Fig. 14A–C), male clypeal moustache dense, ending very close to each other but not, or hardly joining medially (Fig. 14B); malar space insignificant (Fig. 14C); mandible coarsely longitudinally ridged (Fig. 14A); thorax with dense pilosity but shorter than on head (Fig. 14C); thoracic sterna with dense and longer fine pilosity; upper mesopleuron (mesepimeron) densely punctate, sculpture on lower mesopleuron (mesepisternum) greatly masked by long and dense pilosity (Fig. 14C); propodeum with median, somewhat broad median groove, lateral surface shiny but with some points (Fig. 13C). Material examined: 1♁, Wadi Al Awsag, 16.12.1930, det. Mochi (PPDD); 1♁, Mataria, 1.iv.1917 (AUCE); 1♁, Wadi Morrah, 7.iv.1925 (AUCE); 1♁, Kafr Hakim, 20.iii.1926 (AUCE); 1♁, Kafr Farouk, 12.ii.1929 (AUCE); 1♀, Pyramids, 4.iii.1931 (AUCE); 1♁, Kafr Gammus, 17.ii.1935 (AUCE); 2♁, El Gabal El Asfar, 7.ii.1937 (AUCE); 3♁, 1♀, Kafr Farouk, 26.ii.1939 (AUCE); 1♁, Kafr Farouk, 20.ii.1939 (AUCE) Previous Egyptian Records: No specific locality (Spinola 1839, 1843 as P. byssinus; Dalla Torre 1897; Mellor 1928 as P. rutilans; Mochi 1939; de Beaumont 1949, 1956, 1960, 1961 all as P. rutilus rutilus; Bohart & Menke 1976 as P. rutilus rutilus), near Giza (Pyramids) (A. Costa 1875), Maadi (Storey 1916 as P. rutilans), El Gabal El Asfar, Kafr Farouk (Honoré 1942; Roche 2007), Abu Rawash, Ameriah, Dahshour, El Marg, Wadi Awsag (Gadallah 1996 as P. rutilus rutilus), Gebel Elba (Gadallah 1996; Dollfuss 2017), Beni Yussef (Roche 2007). Extralimital distribution: Algeria, Chad, Israel-Palestine, Libya, Mauritania, Morocco, Spain, United Arab Emirates, Yemen. Remarks. The characters of our specimens agree with the North African specimens of de Beaumont (1949: 178, couplet 3), and with the Arabian specimens of Guichard (1994: 208, couplet 2)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 172, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Spinola, M. (1839) Compte-rendu des Hymenopteres recueillis par M. Fischer pendant son voyage en Egypte, et communiques par M. le Docteur Waltl a Maximilien Spinola. Annales de la Societe Entomologique de France, 7, 437 - 546.","Spinola, M. (1843) Sur la synonymie des Hymenopteres. Annales de la Societe Entomologique de France, Bulletin Entomologique, 11, XXXV - XXXVIII.","Dalla Torre, K. W. (1897) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. VIII. Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae [Leipzig], 749 pp.","Mellor, J. E. M. (1928) Notes on certain Hymenoptera, a coleopterous and two dipterous parasites, 1926 - 1927. Bulletin de la Societe Royale Entomologique d'Egypte, 11, 22 - 132.","Mochi, A. (1939) Revisione delle Specie egiziane dei Generi Philanthus F. e Nectanebus Spin. (Hymenoptera: Sphegidae - Philanthinae). Bulletin de la Societe Fouad Ier d'Entomologie, 23, 86 - 139.","de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216.","de Beaumont, J. (1956) Sphecidae (Hym.) recoltes en Libye et au Tibesti par M. Kenneth M. Guichard. Bulletin of the British Museum (Natural History). Entomology, 4, 165 - 215.","de Beaumont, J. (1960) Sphecidae de l'ile de Rhodes (Hym.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 33, 1 - 26.","de Beaumont, J. (1961) Notes sur les Philanthus palearctiques (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 33, 201 - 212.","Bohart, R. M. & Menke, A. S. (1976) Sphecid wasps of the world. A generic revision. University of California Press, Berkeley, Los Angeles and London, 1 color pl., IX + 695 pp. https: // doi. org / 10.1525 / 9780520309548","Costa, A. (1875) Relazione di un viaggio per l'Egitto, la Palestina et le coste della Turchia asiatica per ricerche zoologiche. Atti della R. Accademia delle Scienze Fisiche e Matematiche, 7 (2), 1 - 40.","Storey, G. (1916) List of the Hymenoptera Tubulifera and Aculeata in the collection of the Ministry of Agriculture of Egypt. Bulletin de la Societe Entomologique d'Egypte, 4, 100 - 117.","Honore, A. (1942) Introduction a l'etude des Sphegides en Egypte (Hymenoptera: Aculeata). Bulletin de la Societe Fouad Ier d'Entomologie, 26, 25 - 80.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80.","Dollfuss, H. (2017) The Philanthinae wasps of the \" Biologiezentrum - Linz \" - collection in Linz, Austria, including the genera Aphilanthops Patton, Clypeadon Patton, Odontosphex Arnold, Philanthinus de Beaumont, Philanthus Fabricius, Pseudoscolia Radoszkowski and Trachypus Klug. Linzer Biologische Beitrage, 49, 1245 - 1274.","Guichard, K. M. (1994) The genera Philanthus and Philanthinus (Hymenoptera: Sphecidae) in Arabia. Fauna of Saudi Arabia, 14, 207 - 212."]}
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29. Philanthinus integer
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthinus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Philanthinus integer ,Taxonomy - Abstract
Philanthinus integer (de Beaumont, 1949) Fig. 3A–D Philanthus integer de Beaumont, 1949: 194, ♀, ♁. Diagnosis. Body length: 7.0 mm (female). Body generally black, with the following parts pale yellow: clypeus (except ferruginous spot centrally that does not extend to apex), mandible medially, scape beneath, F1–3 beneath, pronotum, two thin median longitudinal lines, lateral sides of mesoscutum, tegula, upper mesopleuron, two large subquadrate markings on scutellum and smaller lateral spots, posterior margin of scutellum and all postscutellum; T 1 almost ferruginous, remaining terga and sterna with broad pale yellow bands; legs tricolored, with black (especially on femora), yellow and ferruginous; wings hyaline, with veins pale brown to yellowish (Fig. 3A–D). Head in frontal view (Fig. 3C) with inner eye margins slightly concave, margins slightly convergent above, face with dense, short erect pale brown to yellow pilosity extending to middle ocellus; clypeus broad, densely punctate, with arcuate lamella at apex, concave medially, bearing small tooth; propodeum finely reticulate throughout (Fig. 3A, D); mesepimeron smooth and shiny, rest of mesopleuron as well as lateral surface of propodeum densely setose (Fig. 3B). Male: Not available. Material examined: A photograph from CAS: 1♀, Ain El Sokhna road (29–43 km E Maadi), 13.vii.1991, C.G. Roche collector, det. W. Pulawski, 2009. Previous Egyptian Records: No specific locality (de Beaumont 1961; Bohart & Menke 1976; Dollfuss 2017), Ain El Sokhna road (present study). Extralimital distribution: Algeria, Israel, Morocco, Spain, Tunisia, United Arab Emirates. Remarks. The characters of the Egyptian specimen agree with de Beaumont’s description (1949: 194)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 159-160, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["de Beaumont, J. (1949) Les Philanthus et Philiponidea de l'Afrique du N. - O. (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 22, 173 - 216.","de Beaumont, J. (1961) Notes sur les Philanthus palearctiques (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 33, 201 - 212.","Bohart, R. M. & Menke, A. S. (1976) Sphecid wasps of the world. A generic revision. University of California Press, Berkeley, Los Angeles and London, 1 color pl., IX + 695 pp. https: // doi. org / 10.1525 / 9780520309548","Dollfuss, H. (2017) The Philanthinae wasps of the \" Biologiezentrum - Linz \" - collection in Linz, Austria, including the genera Aphilanthops Patton, Clypeadon Patton, Odontosphex Arnold, Philanthinus de Beaumont, Philanthus Fabricius, Pseudoscolia Radoszkowski and Trachypus Klug. Linzer Biologische Beitrage, 49, 1245 - 1274."]}
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30. Philanthus venustus
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Philanthus venustus ,Taxonomy - Abstract
Philanthus venustus (Rossi, 1790) Crabro venustus Rossi, 1790: 94, sex not stated. Remarks. This species is cited to occur in Egypt by Innes Bey (1912 [1911]) as P. melliniformis F. Smith, that was synonymized with P. venustus by Kohl (1891). This record was also based on Walker’s specimens from Sinai and Tadjoura. The species is known to occur in Israel, but has not been otherwise recorded from Egypt. Therefore, this record should be treated with suspicion until otherwise confirmed., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 186, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Rossi, P. (1790) Fauna Etrusca sistens insecta quae in Provinciis Florentina et Pisana praesertim collegit Petrus Rossius. Vols. 1 & 2. Thomae Masi & Sociorum, Liburni [Livorno], XXII + 272 pp. & 348 pp., X pls.","Innes Bey, W. (1912 [1911]) Une liste d'especes recueillis probablement par S. Lord en Egypte et determinees par F. Walker. Bulletin de la Societe Fouad Ier d'Entomologie, 1911, 97 - 115.","Kohl, F. F. (1891) Zur Kenntnis der Hymenopteren - Gattung Philanthus Fabr. (sens. lat.). Annalen des k. k. Naturhistorischen Hofmuseums, 6, 345 - 370."]}
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31. Philanthus variegatus Spinola 1839
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus variegatus ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus variegatus Spinola, 1839 Fig. 19A–E Philanthus variegatus Spinola, 1839: 484, ♀, ♁. Diagnosis. Body length: 10–12 mm (female), 12–13 mm (male). Body generally yellow, with some areas black and ferruginous; in female clypeus and face ivory white extending to eye emargination laterally, and reaching middle ocellus above; in male ivory white is replaced by clear yellow, with small black spot just above antennal sockets; head posteriorly ferruginous, the colour not reaching head posterior margin; mesoscutum ferruginous, black posteriorly; gastral terga with ferruginous intersegments (Fig. 19A–E); malar space inconspicuous (Fig. 19A, B); male clypeal moustache dense, with incurved setae, not joining medially (Fig. 19B); apex of clypeus with three inconspicuous teeth in male (Fig. 19B); pronotum with slight but distinct emargination anteromedially (Fig. 19A); both mesepimeron and mesepisternum densely punctate (Fig. 19C); propodeal dorsum smooth and shiny, with rugose middle sulcus (Fig. 19D, E); gastral T 2 finely, superficially punctate (Fig. 19E). Male genitalia (see Fig. 23D). Material examined: 1♁, 1♀, Marsa Matruh, 22–29.viiii.1954 (FFC); 1♁, Marsa Matruh, 10.v.1930, HC. E. & M.I. collectors (EFC); 1♀, Burg, 16–20.iv.1956 (EFC); 1♀, without collecting data (EFC); 2♁, Wadi Degla, 16. iv.1952, Shefix & Pries. collectors (EFC); 2♁, Burg, 16–20.iv.1956, Sh. H. collector (EFC); 1♁, Kafr Hakim, 19.vii.1933, on grasses (PPDD); 1♀, Manshiet Radwan, 9.v.1934, on flowers (PPDD); 1♁, Kafr Hakim, 15.vi.1935 (PPDD); 1♀, Berkash, 24.iii.1935, collector? on Lupinus (PPDD); 1♀, Kerdasa, 23.iv.1936 (PPDD); 1♀, Abu Rawash, 18.vii.1925 (PPDD); 1♀, 24.vi.1934 (PPDD); 3♀, 5♁, Marsa Matruh, 15.ix.1957, Aly collector (ASUA); 2♁, Marsa Matruh, 20.viii.1955, Aly collector (ASUA). Previous Egyptian Records: Abu Rawash, Berkash, Burg, Kafr Hakim, Kerdasa, Manshiet Radwan, Marsa Matruh, El Seuf, Wadi Degla, Wadi El Natroun (Gadallah 1996 as P. variegatus variegatus), Dahshour, Saqqara (Honoré 1942), El Gabal El Asfar (Roche 2007 as P. variegatus variegatus). Extralimital distribution: Algeria, Azerbaijan, China, Iran, Israel-Palestine, Kazakhstan, Kuwait, Kyrgyzstan, Libya, Mali, Morocco, Oman, Russia, Saudi Arabia, Spain, Sudan, Syria, Tajikistan, Turkey, Turkmenistan, United Arab Emirates. Remarks. Our specimens agree well with de Beaumont’s key (1961: 211, couple 8)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 179, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Spinola, M. (1839) Compte-rendu des Hymenopteres recueillis par M. Fischer pendant son voyage en Egypte, et communiques par M. le Docteur Waltl a Maximilien Spinola. Annales de la Societe Entomologique de France, 7, 437 - 546.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80.","Honore, A. (1942) Introduction a l'etude des Sphegides en Egypte (Hymenoptera: Aculeata). Bulletin de la Societe Fouad Ier d'Entomologie, 26, 25 - 80.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149."]}
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32. Philanthus pulawskii Malash & Edmardash & Gadallah 2023, sp. nov
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
- Subjects
Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus pulawskii ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus pulawskii sp. nov. Fig. 22A–D Type material: Holotype (♀) (EFC), Egypt: North Coast, El Hammam (Marsa Matrouh Governorate) [30 Q 51′21′′N 29 Q 21′18′′E], 28.viii.2022, sweep net, Alyaa A. Malash collector (with head, thorax and gaster separated, head and gaster glued on a triangular card). Description of female holotype: Body length: 12 mm, forewing length: 5.6 mm. Colouration: Body partly ivory white (with yellow hue in some areas), partly black: face whitish below antennal sockets extending laterally to eye emargination and mesally to middle ocellus, except for two small black spots slightly above antennal sockets, black above emargination laterally; first three antennal segments ivory, rest of antenna reddish brown ventrally and black dorsally; mandible ivory with brownish outer margin and darkened tip; pronotal collar, scutellum and postscutellum ivory, pronotal lobe ivory, mesoscutum black (except just before tegula), propodeum black (except for relatively large lateral whitish to yellowish marking), humeral plate and tegula ivory white (Fig. 22A–D), wing membrane hyaline, pterostigma pale brown with dark brown border, veins brownish; propleuron (except for small whitish marking above) and metapleuron black, mesopleuron black, ivory anterodorsally and dorsally (Fig. 22C), legs whitish (except mid and hind coxae yellow and hind femur black, white distally) (Fig. 22C), T 1 ferruginous apically with thin ivory band basally, T 2, T 4 and T 6 ivory, T 3 ivory, narrowly interrupted medially and black basally, with small, rounded black spot laterally on each of lateral spots, T 5 entirely black (Fig. 22D), S 2 – S 3 ivory with black bands apically, S 4 – S 5 ivory, pygidial plate brownish. Head (Fig. 22A): face (below antennal sockets) including clypeus mostly smooth, clypeus shiny, simple, with median lobe slightly bulging medially, with two lateral rounded teeth apically; mandible mostly smooth, face (above antennal sockets) densely punctate in black area, coarsely and densely punctate in yellow area, malar space insignificant; F1-6 distinctly wider than long. Thorax (Fig. 22B, C): pronotal collar smooth, shiny with rounded shoulders, straight anteriorly; mesoscutum shiny, with deep, sparse punctures, scutellum smooth, with few superficial punctures; black area of propodeum including dorsal enclosure shiny, smooth with thin, inconspicuously shagreened median sulcus, lateral whitish area of propodeum with dense coarse punctures, propleuron finely striated transversely, upper mesopleuron nearly smooth, lower mesopleuron densely punctate. hind tibia with 5 spines along its length arranged in two longitudinal rows, hind basitarsus with 6 spines along its length. Gaster (Fig. 22D): T 1 – T 3 with coarse, dense punctures, denser on T 1 and T 2 (especially among whitish bands), black basal area of T 2 nearly smooth, T 4 – T 6 with sparse moderate punctures, all sterna with very few sparse, fine punctures, last (epipygium) tergum parallel-sided, covered with fine setae. Male: Unknown. Etymology: This species is named in honour of the sphecid expert W. J. Pulawski, who confirmed that the species is an undescribed one. Egyptian localities: Known only from El Hammam, North Coast of Egypt (present study). Comment: The new species does not agree with any of the North African species. It moves to couple 12 in de Beaumont’s key of the African species (1961: 212) between P. ammochrysus and P. schulthessi. It differs from both in having lower face nearly smooth and shiny (distinctly punctate in P. ammochrysus (densely) and schulthessi (less densely)). It resembles P. schulthessi and P. minor in having hind femur with long setae ventrally, however, differs from both in having a nearly smooth linear sulcus of propodeal dorsum (sulcus coarsely rugose and extending to the base in the form of a triangle in P. minor and P. schulthessi), in addition to a different colour pattern.
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33. Philanthus genalis Kohl 1891
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Philanthus genalis ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Philanthus genalis Kohl, 1891 Figs 7A, B, 8A–E Philanthus genalis Kohl, 1891: 349, ♀, ♁. Diagnosis. Body length: 11−12 mm (female); 9.0− 12 mm (male). Body predominantly yellow with some areas black (Figs 7A, B, 8A–E); yellow face area in female ending far from middle ocellus, while in male it extends to middle ocellus; head with long fine silvery setae extending to level of middle ocellus in frontal view (Fig. 8A, B); malar space relatively long, about 2/3 length of F 1 in male (Fig. 8D) and half length of F 1 in female (Fig. 8C); clypeal moustache of male meeting medially (Fig. 8B); pronotum straight anteriorly (Fig. 7A, B); propodeal enclosure smooth and shiny, with smooth median sulcus (Fig. 7A, B); upper mesopleuron nearly smooth and shiny, lower mesopleuron with microscopic superficial punctures (Fig. 8C, D). Material examined: 1♁, 1♀, Wadi Degla, 4.iv.1928 (AUCE); 1♁, Suez Road, 15.ix.1929 (AUCE); 1♁, Fayoum, 8.iii.1935 (AUCE); 1♁, 1♀, Marsa Matruh, viii.1953, Dr. T. S. collector (ASUA); 1♁, Kom Oshiem, 4.iv.1953, Aly collector (ASUA); 1♁, Wadi El Natroun, 2.iv.1951, Roh. collector (ASUA). Previous Egyptian Records: Tor (S. Sinai) (Kohl 1891), Fayoum (El Khawagat), Kerdasa, Hammamat Faraoun (S. Sinai), Wadi Degla (Mochi 1939; Honoré 1942; Roche 2007), Sinai Peninsula (Roche & Zalat 1994), Dakhla Oasis, Kom Oshiem, Wadi El Natroun (Gadallah 1996), Dahshour, Wadi El Tih (Dollfuss 2017). Extralimital distribution: Jordan, Kuwait, Libya, Sudan, United Arab Emirates. Remarks. Characters of Egyptian specimens agree well with Guichard’s key (1994: 208, couplet 4). They also agree with de Beaumont’s description (1961: 202) as well as couplet 4 (de Beaumont 1961: 211), but differ in the malar space length: in our specimens the male malar space equals 2/3 the length of F1, and about half length of F 1 in female (in male, equal to F1, and in female 2/3 length of F 1 in de Beaumont’s key). Our specimens agree also with de Beaumont’s figs 4, 5 (1961: 207)., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 165, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Kohl, F. F. (1891) Zur Kenntnis der Hymenopteren - Gattung Philanthus Fabr. (sens. lat.). Annalen des k. k. Naturhistorischen Hofmuseums, 6, 345 - 370.","Mochi, A. (1939) Revisione delle Specie egiziane dei Generi Philanthus F. e Nectanebus Spin. (Hymenoptera: Sphegidae - Philanthinae). Bulletin de la Societe Fouad Ier d'Entomologie, 23, 86 - 139.","Honore, A. (1942) Introduction a l'etude des Sphegides en Egypte (Hymenoptera: Aculeata). Bulletin de la Societe Fouad Ier d'Entomologie, 26, 25 - 80.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149.","Roche, C. G. & Zalat, S. M. (1994) The Sphecidae (order: Hymenoptera) of the Sinai Peninsula. In: Proceedings of the First Egyptian-Hungarian Conference on Environment, St. Catherine, Sinai, Egypt, 5 - 7 April 1993. Suez Canal University, Ismailia, pp. 113 - 118.","Gadallah, N. S. (1996) Review of the genus Philanthus F. in Egypt with notes on little known species (Hymenoptera: Sphecidae). Bulletin of the Entomological Society of Egypt, 74, 68 - 80.","Dollfuss, H. (2017) The Philanthinae wasps of the \" Biologiezentrum - Linz \" - collection in Linz, Austria, including the genera Aphilanthops Patton, Clypeadon Patton, Odontosphex Arnold, Philanthinus de Beaumont, Philanthus Fabricius, Pseudoscolia Radoszkowski and Trachypus Klug. Linzer Biologische Beitrage, 49, 1245 - 1274.","de Beaumont, J. (1961) Notes sur les Philanthus palearctiques (Hym. Sphecid.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 33, 201 - 212."]}
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34. Philanthus triangulum subsp. obliteratum Pic. A 1917
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Philanthus triangulum ,Philanthus triangulum obliteratum pic, 1917 ,Taxonomy - Abstract
Philanthus triangulum obliteratum Pic, 1917 Philanthus triangulum v. nov. obliteratum Pic, 1917: 2, sex not stated. Remarks. The syntypes of this species from Algeria and Egypt (with no specific locality) are cited to be deposited in M. Pic’s collection, which may be now in the MNHN. Since the status of this subspecies has never been firmly established, we do not include it in the list of Egyptian Philanthus., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on page 186, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Pic, M. (1917) Notes hymenopterologiques. L'Echange, Revue Linneenne, 381, 1 - 4."]}
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35. Philanthus triangulum subsp. abdelcader Lepeletier de Saint Fargeau 1845
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Malash, Alyaa A., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Crabronidae ,Philanthus ,Insecta ,Arthropoda ,Philanthus triangulum abdelcader lepeletier de saint fargeau, 1845 ,Animalia ,Biodiversity ,Hymenoptera ,Philanthus triangulum ,Taxonomy - Abstract
Philanthus triangulum abdelcader Lepeletier de Saint Fargeau, 1845 Fig. 18A–E Philanthus abdelcader Lepeletier, 1845: 33, ♀, ♁. Diagnosis. Body length: 9–12 mm (female); 12–17 mm (male). This subspecies varies greatly in body size and colour. Body generally black and yellow; head with clypeus and face ivory white extending to eye emargination in both sexes (Fig. 18A, B); ivory white area just above clypeus bifurcate in females (Fig. 18A), trifurcate in males (Fig. 18B); head with two ferruginous bands postero-laterally (Fig. 18C, D); antenna thickened medially (Fig. 18A, C); malar space very short (Fig. 18A, C); male clypeal moustache long and dense, joining medially (Fig. 18B); female clypeus with 4–5 distinct teeth at apex, with two teeth in male and fringed with long reddish setae; mesopleuron entirely densely punctate (Fig. 18C); propodeum densely punctate dorsally (Fig. 18D, E); gastral terga punctate (Fig. 18E), while sterna almost smooth. Male genitalia (see Fig. 23C). Material examined: 12♁, Maadi, 19.iv.1912, LHS collector (PPDD); 5♀, 2♁; Giza, 1.iii.1913 (PPDD); 10♁, Maadi, 6.iii.1913, LHS collector (PPDD); 2♁, Pyramids, 17.iv.1914 (PPDD); 1♁, Hadaiq El Kuba, 16.v.1914, Naguib collector (PPDD); 4♁, Abu Rawash, 4.x.1914 (AUCE); 2♁, Banha, 30.v.1914 (PPDD); 1♁, Wadi Nesour, 23.x.1925 (PPDD); 1♀, El Gabal El Asfar, 25.x.1925 (PPDD); 3♁, 1♀, Kafr Hakim, 30.i.1926 (PPDD); 1♁, Zinein, 6.vii.1930, on sugarcane (PPDD); 1♁, Nubia, 23.vii.1930, on sugarcane (PPDD); 1♀, Embaba, 13.viii.1930, on sugarcane (PPDD); 1♁, Gebel Elba, 30.i.1933 (PPDD); 2♀, 1♁, Abu Rawash, 23.vii.1933 (PPDD); 1♀, Helwan, 1.iii.1934, Farag collector (EFC); 1♁, Wadi Garawi (Helwan), 10.iii.1934 (PPDD); 1♀, Helwan, 27.xii.1934, Farag collector (EFC); 1♀, Helwan, 22.i.1935, Farag collector (EFC); 2♁, Abu Rawash, 24.ii.1935, on flowers (PPDD); 1♀, Helwan, 21.xii.1935, Farag collector (EFC); 1♀, Helwan, 21.i.1936, Farag collector (EFC); 1♁, Abu Rawash, 11.iv.1939, Shaaban collector (EFC); 1♁, Abu Rawash, 12.v.1951, Roh. collector (ASUA); 1♀, Embaba, 13.xii.1952, Aly collector (ASUA); 2♁, Fayoum, 5.v.1952, Saleh collector (ASUA); 1♁, 1♀, El Gabal El Asfar, 1.i.1953, Aly collector (ASUA); 1♁, 1♀, Cairo, 2.i.1953, Aly collector (ASUA); 1♀, Ain Shams, 5.i.1953 (EFC); 2♁, Giza, 5.i.1953 (EFC); 1♁, Cairo, 13.i.1953, Aly collector (ASUA); 2♀, 1♁, Giza, 15.i.1953 (EFC); 2♀, 5♁, Pyramids, 19.i.1953, Aly collector (ASUA); 1♁, Giza, 7.ii.1953 (EFC); 1♀, 10.ii.1953, El Kharga (New Valley), Aly collector (ASUA); 1♀, Giza, 14.ii.1953 (EFC); 1♁, Barrage, 2.iii.1953 (EFC); 1♀, Giza, 15.iii.1953 (EFC); 1♀, 1♁, 3.v.1953 (EFC); 2♁, Girza (Giza), 9.v.1953 (EFC); 1♀, 1♁, Giza, 13.v.1953 (EFC); 1♁, Berkash, 10.x.1953, Aly collector (ASUA); 1♁, Giza, 2.i.1955, Dawlatly collector (EFC); 1♀, Burg El Arab, 12.ix.1955, Abdalla collector (EFC); 1♀, Giza, 8.xi.1955, Abdalla collector (EFC); 21♀, 2♁, Burg El Arab, 16–20.vi.1956, Shafix collector (EFC); 3♀, Burg El Arab, 10.vi.1960, Abdalla collector (EFC); 1♁, Burg El Arab, 10.iii.1961, Shafix collector (EFC); 1♀, Suez, 14.ii.1962, Dawlatli collector (EFC); 1♁, Magadlah, 7.iii.1978 (PPDD); 1♁, Mansoura, 16.ii.1986 (PPDD); 1♁, Wadi El Natroun (rest house), 2.vii.1996 (EFC); 1♀, Giza, 26.iii, ??? on flowers (EFC); 1♁, Maadi, 7.v.1913 (AUCE); 1♁, Ezbet Hulayil, 29.v.1913 (AUCE); 1♀, Kerdasa, 12.iv.1914 (AUCE); 3♁, Helwan, 29.x.1920 (AUCE); 3♁, Abu Rawash, 4.x.1924 (AUCE); 2♁, 1♀, Kerdasa, 8.ix.1926 (AUCE); 1♀, El Gabal El Asfar, 14.i.1931 (AUCE); 1♁, El Gabal El Asfar (AUCE); 2♁, El Gabal El Asfar, 4.ii.1934 (AUCE); 2♁, Fayoum, 29.iv.1935, A. Rabinovitch collector (AUCE); 1♁, Magadlah, 24.iii.1936 (AUCE); 2♁, El Gabal EI Asfar, 14.i.1936 (AUCE); 1♁, El Gabal El Asfar, 29.i.1936 (AUCE); 1♁, El Gabal El Asfar, 19.ii.1936 (AUCE); 2♁, Kafr Farouk, 19.ii.1939 (AUCE); 2♁, 1♀, without collecting data (EFC); 1♀, Giza, v.2022, Yusuf Edmardash collector (EFC). Previous Egyptian Records: Abu Rawash, Abu Sir, Dakhla Oasis (Tineida), Fayoum (Hawarah, Karanis, Kom El Atl), El Gabal El Asfar, Helwan, Ismailia, Kerdasa, Kom Oshiem, El Minya, Saqqara, Rafah (N. Sinai), (Roche 2007), Abukir (de Beaumont 1966), between Fayoum and Nile (A. Costa 1875 as P. Abdelkader), Heliopolis and Hor Tamanib (Walker 1871 as P. Abdelkader), Ramla near Alexandria (Lucas 1880 as P. abdelcader), Sinai Peninsula (Roche & Zalat 1994), Embaba, Pyramids, Berkash, Kharga, Wadi El Natroun, Giza, Wadi Nesour, Nubia, Mansoura, Wadi Garawi (10 km south east Helwan), Zinein, Banha, Maadi, Hadaiq El Kuba, Magadlah, Gebel Elba (present study). Extralimital distribution: Algeria, Cameroon, Iraq, Israel-Palestine, Italy, Libya, Madagascar, Morocco, Oman, Spain, Tunisia, Turkmenistan, Ukraine., Published as part of Malash, Alyaa A., Edmardash, Yusuf A. & Gadallah, Neveen S., 2023, The genera Philanthus Fabricius, 1790 and Philanthinus de Beaumont, 1949 (Hymenoptera: Crabronidae: Philanthinae) in Egypt, with a new record and the description of a new species, pp. 151-189 in Zootaxa 5249 (2) on pages 178-179, DOI: 10.11646/zootaxa.5249.2.1, http://zenodo.org/record/7687292, {"references":["Lepeletier de Saint Fargeau, A. L. M. (1845) Histoire naturelle des Insectes. Hymenopteres. Vol. 3. Librairie Encyclopedique de Roret, Paris, 646 + 4 pp., pls. 25 - 36.","Roche, C. G. (2007) Conspectus of the Sphecid wasps of Egypt (Hymenoptera: Ampulicidae, Sphecidae, Crabronidae). Egyptian Journal of Natural History, 4, 12 - 149.","de Beaumont, J. (1966) Quelques Sphecidae de la faune d'Egypte (Hym.). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 38, 203 - 212.","Costa, A. (1875) Relazione di un viaggio per l'Egitto, la Palestina et le coste della Turchia asiatica per ricerche zoologiche. Atti della R. Accademia delle Scienze Fisiche e Matematiche, 7 (2), 1 - 40.","Walker, F. (1871) A list of Hymenopterous insects collected by J. K. Lord, Esq. in Egypt, in the neighbourhood of the Red Sea, and in Arabia. With descriptions of the new species. E. W. Janson, London, 59 + IV pp. https: // doi. org / 10.5962 / bhl. title. 8818","Lucas, H. (1880) Une note relative a un Hymenoptere apivore. Annales de la Societe Entomologique de France, Serie 5, 10, CXXXVII - CXXXVIII.","Roche, C. G. & Zalat, S. M. (1994) The Sphecidae (order: Hymenoptera) of the Sinai Peninsula. In: Proceedings of the First Egyptian-Hungarian Conference on Environment, St. Catherine, Sinai, Egypt, 5 - 7 April 1993. Suez Canal University, Ismailia, pp. 113 - 118."]}
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36. Aleiodes subpallidator Edmardash & Gadallah 2022, sp. nov
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Arthropoda ,Animalia ,Aleiodes ,Biodiversity ,Hymenoptera ,Aleiodes subpallidator ,Taxonomy - Abstract
Aleiodes subpallidator Edmardash & Gadallah sp. nov. Figs 15 A, B, 16 A–D, 17 A–E urn:lsid:zoobank.org:act: FABD3356-A9CD-4B1A-B2DC-D1C389A9CDF1 Type material: Holotype, ♀ (EFC), Saudi Arabia (Farasan Archipelago, Al-Sajid), 16.86026N, 41.932564E, 17.xi.2017, light trap, leg. Usama Abu El-Ghiet & Tarek M. El-Sheikh. Diagnosis. Head (except black stemmaticum), mesosoma and posterior end of metasoma brownish in colour (Figs 15A, B, 17D); scapus slightly, but distinctly oblique apically, longer dorsally than ventrally (Fig. 16A); OOL about as long as diameter of posterior ocellus; occipital carina well seen ventrally, weak dorsally, reaching hypostomal carina; notauli absent on mesoscutum (Fig. 16D); vein 2–CU1 of fore wing 2.9 × as long as 1–CU1 (Fig. 17A); vein 1–SR very short, connected with vein 1–M at the same level (Fig. 17A), vein 1–M slightly curved posteriorly (Fig. 17A); metasomal T4 superficially granulate to nearly smooth (Fig. 17D); fore wing 2.8mm; malar space 0.2 × height of eye (Fig. 16A); marginal cell of hind wing parallel-sided apically (Fig. 17B); metatibia with inner comb of blunt bristles apically (Fig. 17C). Description. Holotype, ♀, length of body 3.5 mm; length of fore wing 2.8 mm. Colour (Figs 15A, B, 16B). Head and mesosoma (except propodeum), as well as end tergites brownish, propodeum and legs straw yellow; tegula pale yellow; stemmaticum black; wings (Figs 17A, B) hyaline, with pale brownish to yellowish veins, pterostigma bright yellow; antenna yellow, with apical half of flagellum brownish (apices broken). Head (Figs 16A–C).Antennae with broken flagella, scapus oblique apically, longer dorsally than ventrally; face superficially punctulate, shiny; OOL about as long as posterior ocellus diameter, finely rugose; vertex granulaterugose; clypeus somewhat convex, small, finely coriaceous, ventral margin concave; width of hypoclypeal depression 0.3× minimal width of face; face very finely granulate; occipital carina complete, but weak dorsally, meeting hypostomal carina ventrally; clypeus located slightly above lower level of eye; malar space 0.2× as long as height of eye laterally; eye distinctly bulged, with slight, but distinct emergination at uppermost part of inner orbits. Mesosoma (Fig. 16D). Mesoscutum very finely coriaceous throughout, notauli absent, posterior margin straight; mesopleuron and metapleuron very finely coriaceous-rugose throughout, rather shiny; scutellum somewhat convex, finely and superficially dotted to coriaceous, without lateral carina; propodeum finely rugose, with a complete median longitudinal carina. Wings (Figs 17A, B). Fore wing: vein r 0.45× vein 3SR; 1–CU1 slightly oblique, 0.34× as long as 2–CU1; vein r–m 0.54× vein 2–SR, and 0.34× vein 3–SR; SMC2 elongate, trapezoidal; vein 1–SR short, straight with 1–M; cu–a rather oblique, somewhat reduced ventrally; vein CU1a distinctly curved at base; vein 1–M slightly curved posteriorly. Hind wing: apical half of marginal cell parallel-sided; vein 2–SC+R very short; vein m-cu greatly reduced, as being just an unpigmented fold; M+CU 1.77× as long as 1–M; 1r–m 0.5× 1–M. Legs (Figs 15A, B, 17C). Claws simple, setose; hind coxa superficially coriaceous; femora thin and slender; hind trochantellus 2.9× as long as ventrally as apical width; metatibia with an inner transparent comb of blunt bristles apically; inner metatibia spur 0.18× as long as hind basitarsus; two metatibia spurs nearly equal in size. Metasoma (Figs 17D, E). Metasomal T1 0.84× as long as its posterior width, flattened and lamellate laterally; T1, T2 and base of T3 finely longitudinally rugose, T1 and T2 with strong longitudinal median carina; mediobasal smooth area of T2 absent; remainder of metasomal tergites superficially coriaceous to nearly smooth. Lateral sides of metasomal tergites with fine, short whitish setae. Ovipositor sheath short, slender, tapering at end, nearly smooth. Distribution. So far Saudi Arabia (Farasan Islands, Jazan). Remarks. The new species closely resembles Aleiodes pallidator (Thunberg) (based on the description of van Achterberg & Shaw, 2016), but differs from A. pallidator in the following combination of characters: notauli completely absent (Fig. 17D) (in A. pallidator notauli present, narrow, shallow and crenulate (Fig. 15C)); inner apex of metatibia with comb of transparent, blunt bristles (Fig. 17C) (in A. pallidator inner apex of metatibia without comb of bristles); ovipositor sheath not densely setose, very short (hardly seen in lateral view), tapering apically (Fig. 17E) (in A. pallidator ovipositor sheath densely setose, well-seen in lateral view, rounded apically). Measurements of fore wing veins (Fig. 17A): vein r 0.45× vein 3–SR (0.3× in A. pallidator); 1–CU1 slightly oblique (horizontal in A. pallidator); vein r–m 0.54× vein 2–SR (0.7× in A. pallidator), and 0.34× vein 3–SR (0.4× in A. pallidator); vein 1–SR short, straight with 1–M (angled with 1–M in A. pallidator); cu–a rather oblique, somewhat reduced ventrally; vein CU1a distinctly curved at base; vein 1–M slightly curved posteriorly. Etymology. The name subpallidator is named with reference to the similarity of this species to Aleiodes pallidator.
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37. Braconella minor Szepligeti 1906
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconella ,Braconidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Braconella minor ,Hymenoptera ,Taxonomy - Abstract
Braconella minor Sz��pligeti, 1906 Figs 2 A���D Braconella minor Sz��pligeti, 1906: 587, ♀. Description of the male: Body length: 5.0 mm; fore wing length: 4.2 mm. Colour & vestiture (Figs 2A���D): Body generally black (including antennae), metasoma dark ferruginous, with T1, base of T2 and posterior margins of T2���4 ivory white to pale yellowish; temple and lower face just behind antennal bases and inner orbits of eyes, clypeus, mandibles (except black tip), and palpi are honey yellow; first and middle legs (except black coxae) yellowish, hind legs mostly black, except for the apex of femur and basal third of tibia yellowish, tibial spurs yellowish; sides of pronotum ferruginous yellow; tegula ferruginous to orange. Wings hyaline, with pterostigma and veins black. Body densely clothed with short whitish to silvery setae especially on mesosoma and hind legs. Head (Fig. 2C): Transverse, more than 1.5�� as wide as mesosoma; occiput finely alutaceous, shiny; frons coarser than occiput; eyes distinctly convergent behind, slightly emarginate to concave opposite to antennal base; temple 0.88�� eye length in dorsal view; malar space distinctly short, 0.17�� as long as eye length in front view. Antenna 22-segmented, scape 1.55�� as long as wide, F1 1.3�� F2, flagellomeres of middle third of flagellum subequal, last flagellomere tapered apically. Mesosoma (Figs 2A, B): mesoscutum densely finely punctate, clothed with dense fine seta; notauli present, convergent and meeting at apex of mesoscutum; scutellum with simple, linear scutellar sulcus, propodeum nearly smooth and shiny, without median carina, lateral sides densely finely punctate, with fine whitish setae, median line shiny and smooth; pronotum laterally finely rugose; meso- and metapleura densely finely punctate, with dense fine whitish hairs as those covering mesoscutum and scutellum. Fore wing (Fig. 2A): about as long as body; with vein SR1 nearly reaching wing apex; vein r arising from middle of pterostigma, about 0.3�� as long as vein 3-SR; SR1 longer than veins r and 3���SR combined; vein cu-a almost interstitial, resulting in the absence of vein 1���CU1; vein 1���M distinctly straight; vein 2���SR+M present, relatively long; pterostigma 2.6�� as long as its maximum width, 0.7�� as long as 1���R1. Legs: with somewhat flattened femora; hind femur 2.7�� as long as broad, with an oblique, sharp triangular keel in the middle ventrally (Fig. 2D); hind basitarsus distinctly shorter than rest of tarsomeres combined (Fig. 2D). Metasoma (Figs 2A, B): T1 parallel-sided, with a V-shaped depression basally, slightly elevated towards apex; T2 strongly emarginate baso-medially, greatly widened posteriorly, 0.6�� as long as apical width; T3 and the following tergites are in the form of a transverse band, much wider than long, 2.2�� as wide as median length; T2 and T3 are separated by a crenulate suture; all tergites finely rugose. Material examined. 1♂, KSA: Jazan (Farasan Islands, Al-Sajid before Khotp), 16.883482N 41.909728E, 17. ii. 2017, sweep net, leg. Usama M. Abu El-Ghiet & Tarek, M. El-Sheikh. General distribution. South Africa, Tanzania, Zimbabwe (Yu et al. 2016), Saudi Arabia (Farasan Islands, Jazan) (new record). Remarks. The genus Braconella is a first record for the Arabian Peninsula. Masi (1944) saw a male of Braconella and suggested that it was B. minor, however, he had some doubts as it differed substantially from the female he examined. Here we considered our specimen as being the unknown male of B. minor and describe it for the first time. The specimen examined here resembles the Tanzanian female described by Brues (1924), except for the following: metasoma with T1, base of T2 broadly, and posterior margin of T2���4 thin ivory white to pale yellowish (Fig. 2B) (in female the metasoma to apex of T5 ferruginous); vein r 0.3�� as long as vein 3���SR (Fig. 2A) (two-thirds as long as 3���SR in female); vein SR1 distinctly longer than veins r and 3���SR combined (Fig. 2A) (as long as both veins combined in the female)., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on pages 52-54, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Szepligeti, G. (1906) Braconiden aus der Sammlung des Ungarischen National-Museums, 1. Annales Historico-Naturales Musei Nationalis Hungarici, 4, 547 - 618.","Yu, D. S. K., van Achterberg, C. & Horstmann, K. (2016) Taxapad 2016, Ichneumonoidea 2015. Taxapad database on flash drive, Ottawa.","Masi, L. (1944) Missione biologica Sagan-Omo firetta dal Prof. E. Zavattari. Nuove species d'imenotteri Braconidi. Annali del Museo Civico di Storia Naturale di Giacomo Doria, 62, 11 - 215.","Brues, C. T. (1924) Some South African parasitic Hymenoptera of the families Evaniidae, Braconidae, Alysiidae, and Plumariidae in the South African Museum with a catalogue of the known species. Annals of the South African Museum, 19, 1 - 150."]}
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38. Hormius undetermined
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Hormius ,Hormius undetermined ,Hymenoptera ,Taxonomy - Abstract
Hormius sp. Figs 5A���E Diagnosis. Body generally reddish yellow (Figs 5A, B, D), with lateral lobes of mesoscutum brownish; propodeum posteriorly and T1 brownish; legs yellowish; fore wing hyaline (Fig. 5B), with clear yellow pterostigma, veins pale brown. Mesoscutum smooth and shiny above, rugose along its posterior margin between notauli (Fig. 5D); notauli deep, convergent posteriorly; scutellar sulcus broad, crenulate (Fig. 5D); scutellum smooth and shiny (Fig. 5D); propodeum strongly and densely areolate (Fig. 5C); mesopleuron shiny, with shallow precoxal groove, present close to ventral side (Fig. 5A); epicranial carina present, thin (Fig. 5E). Fore wing fringed with fine setae especially at apex and ventrally (Fig. 5B); vein SR1 reaching wing apex; pterostigma 5.7�� as long as its maximum width, about as long as 1���R1; vein r 0.8�� as long as vein 3���SR; vein 3-SR shorter than 2���SR; r: 3-SR: SR1= 20: 25: 73; vein r���m present, membranous (non-tubular); vein cu���a interstitial; vein 1���M straight to slightly curved (Fig. 5B). Metasoma T1 coarsely rugose, narrow, nearly parallel-sided; T2 is the largest tergite, occupying a large part of metasoma; T2���5 wider than long, coarsely rugose, each with elevated, sharp posterior margin; T6 small, nearly smooth; ovipositor 0.35�� as long as metasomal length. Material examined. 1♀, KSA: Jazan (Farasan Islands, Al ���Sajid), 16.860626N 41.932564E, 17. ii. 2017, light trap, leg. Usama M. Abu El-Ghiet & Tarek M. El-Sheikh (specimen with broken head). Remarks. The subfamily Hormiinae is recorded here for the first time for the fauna of the Arabian Peninsula. From the generic point of view, our specimen is very close to Parahormius Nixon, but differs from it by the presence of an epicnemial carina (Fig. 5E), that can be used to differentiate between the genera Hormius and Parahormius (based on Sharkey et al., 2021 key, p. 310; M. Sharkey, pers. comm.). Because of the bad condition of the specimen, we are not able to identify it to the specific level., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on page 57, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Sharkey, M. J., Janzen, D. H., Hallwachs, W., Chapman, E. G., Smith, M. A., Dapkey, T., Brown, A., Ratnasingham, S., Naik, S., Manjunath, R., Perez, K., Milton, M., Hebert, P., Shaw, S. R., Kittel, R. N., Solis, M. A., Metz, M. A., Goldstein, P. Z., Brown, J. W., Quicke, D. L. J., van Achterberg, C., Brown, B. V. & Burns, J. M. (2021) Minimalist revision and description of 403 new species in 11 subfamilies of Costa Rican braconid parasitoid wasps including host records for 219 species. ZooKeys, 1013, 1 - 665. https: // doi. org / 10.3897 / zookeys. 1013.55600"]}
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39. Avga saudita Gadallah & Edmardash 2022, sp. nov
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Arthropoda ,Animalia ,Avga ,Biodiversity ,Avga saudita ,Hymenoptera ,Taxonomy - Abstract
Avga saudita Gadallah & Edmardash, sp. nov. Figs 10 A–C, 11 A–C urn:lsid:zoobank.org:act: 5D58F822-D1CB-45C3-997C-574C62CC8B25 Type material: Holotype, ♀ (EFC), KSA: Jazan (Farasan Islands, Al-Sajid), 10.xi.2017, light trap, leg. Usama Abu El Ghiet; Paratypes: 2 ♀, same data. (paratypes with broken flagella) Diagnosis. Body yellowish to yellowish brown (including legs) (Figs 10A–C), antenna with scape, pedicel and F1–3 clear yellow (Fig. 10B), then darkened in the following segments; metasoma waxy white in middle of T1 (Fig. 10A) and ventrally. Fore wing hyaline (Fig. 11C), with veins pale brown, except veins r, 1–SR, 1–M, apical half of M + CU, 1–CU1 and 2–CU1 are dark brown; pterostigma mostly pale brown, and dark brown apically. Head and mesosoma finely and closely granulate; mesopleuron and metapleuron are smooth and shiny (Fig. 10C); sternaulus deep and linear ventrally (Fig. 10C); propodeum nearly smooth, without distinct fields, with a short median carina, extending from base (Fig. 10A); vein r of fore wing about as long as or slightly longer than vein 3–SR; vein SR1 straight, but slightly concave apically, reaching wing apex; vein r–m present but not tubular (Fig. 11C); ovipositor sheath dark brown, about 0.3× as long as metasoma (Fig. 10B). Description. Holotype ♀, length of body (excluding ovipositor): 4.1mm.; length of fore wing: 4.2mm. Colour. Body yellowish to yellowish brown (including legs), antenna with scape, pedicel, and F1–3 clear yellow, rest of flagellum (unbroken part) darkened; mandible yellowish, with black tip; palpi whitish; metasoma waxy white ventrally. Fore wing hyaline, with veins pale brown, except veins r, 1–SR, 1-M, apical half of M + CU, 1–CU1 and 2–CU1 are dark brown; pterostigma mostly pale brown, and dark brown apically. Head (Figs 11A, B). Finely granulate dorsally; frons medially, just behind antennal bases, slightly swollen and shiny; area between stemmaticum and antennal bases distinctly depressed; inner orbits of eyes slightly emarginate above; eye in dorsal view 2.6× as long as temple; OOL: OD: POL = 35: 29: 17; malar space 1.3× as long as basal width of mandible; scape equal both ventrally and dorsally; pedicel slightly more swollen than F1, slightly shorter than scape; F1 about 5.0× as long as wide, 1.3× as long as F2; F2–4 about equal sized, 3.8× as long as wide; mandible with two subequal, sharp teeth. Mesosoma (Figs 10A–C) 2.0× as long as its height. Mesoscutum and mesoscutellum finely granulate; mesoscutum clothed with dense, fine whitish setae, becoming few along lateral and posterior margins of mesoscutellum; notauli very weak to indistinct; propodeum nearly smooth, with a short median longitudinal carina at base not reaching posterior margin; mesopleuron smooth and shiny, with a deep linear sternaulus ventrally. Fore wing (Fig. 11C). Vein r arising very near to the middle of pterostigma; SR1 straight, slightly concave near to apex, reaching wing apex; 2–SR+M very short, hardly seen; vein r 0.75× as long as pterostigma width, about slightly longer than vein 3–SR; pterostigma about as long as or slightly longer than vein 1–R1; vein 1–M nearly straight; vein cu–a postfurcal; r: 3–SR: SR1 = 15: 13: 90. Legs moderately long and slender; metatibia distinctly longer than hind femur, 1.3×; hind basitarsus slightly shorter than following tarsomeres combined. Metasoma (Figs 10A–C). Pear-shaped, T1 conical, slightly widened posteriorly, about 0.8× as long as its posterior width, finely rugose to nearly smooth; T2 relatively large, with sclerotized inverted U laterally, membranous and skin-like medially, rest of tergites band-like, distinctly wider than long; T3 membranous medially, sclerotized areas of tergites smooth and shiny; ovipositor sheath 0.3× as long as metasoma in lateral view. Remarks. The subfamily Pambolinae is recorded here for the first time for the fauna of the Arabian Peninsula. Etymology. The name saudita is derived from the country locality (Saudi Arabia) from which the specimen was collected and described. Distribution. Saudi Arabia (Farasan Islands, Jazan).
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40. Adelius aridus
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Adelius aridus ,Arthropoda ,Animalia ,Adelius ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Adelius aridus (Tobias, 1967) Figs 3A���D Myriola arida Tobias, 1967: 392. Acaelius arida Shenefelt, 1973: 670. Adelius arida Tobias, 1986: 591. Diagnosis. Body with head and thorax yellowish brown, metasoma waxy white, with the last three to four tergites dark brown (Figs 3A, B, D); legs yellowish; antenna yellowish basally, dark brown at apical half (Fig. 3A); fore wing hyaline, with slight infuscation behind pterostigma, pterostigma dark brown, colourless or very pale at extreme base and apex (Fig. 3C). Head transverse, 1.9�� as wide as its length, slightly wider than mesosoma (Fig. 3B); temple rounded, 0.6�� as long as eye in dorsal view (Fig. 3B); antenna filiform, somewhat longer than body (Fig. 3A). Mesoscutum densely finely punctate, uniformly setose throughout (Fig. 3B); scutellar sulcus crenulate (Fig. 3B); scutellum with sparse setae; propodeum areolate. With three rectangular areolae at base, and a narrow one apically; mesopleuron finely coriaceous, separated from metapleuron with a crenulate sulcus; metapleuron strigate (Fig. 3D). Wings slightly shorter than body (Fig. 3A); pterostigma enlarged, 2.4�� as long as its maximum width, with vein 1���R1 reduced; veins 2���SR and 3���SR+SR1 issued from the same point; vein 3���SR+ SR1 only sclerotized at base, vein 1���SR+M nearly straight, vein 1���M slightly curved, to nearly straight (Fig. 3C). Ovipositor sheath short, rounded apically, with few long fine setae (Fig. 3A). Material examined. 1♀, KSA: Jazan (Farasan Islands, Al-Sajid), 16. 860626N 41.932564E, 25.i. 2017, light trap, leg. Usama M. Abu El-Ghiet & Tarek M. El-Sheikh. General distribution. Iran, Turkmenistan (Derafshan et al. 2019), Saudi Arabia (Farasan Islands, Jazan) (new record). Remarks. This is the first record of the genus Adelius for the Arabian Peninsula. The Saudi Arabian specimen agrees with the Iranian specimens of A. aridus (see Derafshan et al. 2019). It only differs from the Iranian specimen in having the antennae slightly longer than the body (slightly shorter in the Iranian specimens. 0.9�� measured from Fig. 5B of Derafshan et al. (2019))., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on page 54, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Tobias, V. I. (1967) [Middle Asian species of braconids (Hymenoptera, Braconidae) taken with light quartz lamp.]. Trudy Zoologicheskogo Instituta, Leningrad, 38, 382 - 396.","Shenefelt, R. D. (1973) Braconidae 6. Cheloninae. Hymenopterorum Catalogus (novo editio), 10, 813 - 936.","Tobias, V. I. (1986) Cheloninae. In: Medvedev G. S. (ed.) ' Opredelitel Nasekomych Evrospeiskoi Tsasti SSSR. 3, Peredpontdakrylye 4. Opr. Faune SSSR ' 145, 1 - 501. Pp. 293 - 335."]}
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41. Diolcogaster mayae
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Diolcogaster ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Diolcogaster mayae ,Hymenoptera ,Taxonomy - Abstract
Diolcogaster mayae (Shestakov, 1932) Figs 6 A, B, 7 A���D, 8A���D Microgaster mayae Shestakov, 1932: 260. Description of female (Saudi Arabia; Farasan Islands, form D). Body length: 3.2mm; fore wing length: 3.0mm; length of antenna: 0.82mm. Colour (Figs 6 A, B, 7 A, C, 8 A���D). Head black, with face (behind antennal bases) dark reddish brown, with a median yellow, longitudinal streak, extending a distance behind antennal bases to base of clypeus; clypeus and mandible yellow (except tip of mandible black); eyes in dorsal view lined with thin yellow; scapus and pedicle yellow, flagellum dark brown; mesosoma and metasoma (except whitish T1 and T2) yellowish; legs clear yellow; ovipositor sheath dark brown. Fore wing hyaline, with pterostigma clear yellow; veins r, areola, 1���M, 1���CU1, 2���CU1, cu-a are dark brown; while veins 1���SR+M, m���cu, 2���SR+M, M+CU1 are non-tubular. Head (Figs 7A���C, 8A). Densely finely punctate; occiput smooth and shiny; head height 7.7�� temple length; preapical flagellomeres longer than wide, slender (not moniliform), F2, F12, F13, F14, penultimate, and apical flagellomeres: 2.8, 3, 3, 3, 2, 2.5�� as long as wide, respectively. Gena distinctly smaller than eye width, 0.3�� as wide as eye width; malar space extremely short; maximum width of interocular line 9.2�� as long as malar space; eye very large, distinctly bulged, densely setose, its height 1.6�� its width, its height 8.1�� temple length; eye height 13.9�� malar space; OD 3.6�� OOL, POL 5.0�� OOL. Face medially, under antennal bases with distinct longitudinal carina, not reaching clypeal base. Mesosoma (Figs 7D, 8A, C). Mesoscutum densely finely punctate, densely setose (except anteriorly); scutellum densely punctate as mesoscutum, punctures somewhat superficial, elevated medially, slightly darker medially, with inwardly directed whitish setae, 0.9�� as long as basal width; propodeum smooth and shiny, with a median branched carina, shortly bifurcated apically, in the form of an inverted U; as well as a lateral, short, curved irregular carina, appearing as if forming closed areola postero-laterally; SOS with four short, unequal rugae posteriorly; mesopleuron with precoxal area smooth and shiny; metapleuron smooth and shiny anteriorly, rugose posteriorly. Legs (Figs 6A, B). Metacoxa extremely large, extending to anterior margin of metasomal T3, together with metafemur are densely finely punctate along outer surfaces; metafemur 3.4�� as long as wide, 0.9�� as long as metatibia; inner metatibial apical spur 0.6�� as long as basitarsus. Wings (Fig 8B). Vein R1 of fore wing 1.3�� as long as pterostigmal length; pterostigma 2.9�� as long as its width; vein r 1.5�� as long as vein 2RS; vein r���m 1.4 as long as 3RS. Hind wing with vein cu���a straight. Metasoma (Figs 7D, 8D). Metasomal T1 with strong medio-longitudinal sulcus; medial length of T1 2.65�� as long as anterior width, and 2.75�� its posterior width; T2 transverse, with a median field, 1.8�� as wide as its length; T3 distinctly longer than T2, smooth, without any trace of median field; rest of tergites smooth and setose; ovipositor sheath dark brown, with fine whitish setae, 0.13�� as long as metatibia. Material examined. 1♀, Jazan, Farasan Islands (Al-Sajid), 16.860626N 41.932564E, 25.I.2017 (light trap), leg. Usama Abu El-Ghiet & Tarek El-Sheikh. Distribution. Afghanistan, Algeria, Armenia, Azerbaijan, Iran, Israel-Palestine, Kazakhstan, Mongolia, Romania, Russia, Tajikistan, Turkmenistan, Uzbekistan, Yemen (Ghafouri Moghaddam et al. 2019). Remarks. In the present study, the genus Diolcogaster is recorded for the first time for the fauna of Saudi Arabia. Characters of the Saudi Arabian specimen agree with those of D. mayae in the key of Ghafouri Moghaddam et al. (2019, alternative of couple 3, p. 100), as well as Table (3) (p. 106). The species is known by its high intraspecific variability (colour and measurements), in which three forms (A���C) were recorded in the Iranian fauna (see Ghafouri Moghaddam et al. 2019), as well as other hot countries including Yemen. We designate this Saudi Arabian specimen of D. mayae as form D. It is closest to form B, but differs from form B in the following: head black, with face behind antennal bases ferruginous, with a yellow longitudinal streak extending a distance from antennal bases to clypeal base (Fig. 7A) (entirely black in form B of Iran, see Ghafouri Moghaddam et al. (2019), Fig. 11D, p. 113); mesosoma yellow, slightly darker on mesoscutum and middle of scutellum (Fig. 7D), while propodeum honey yellowish (Fig. 8C) (brownish on mesosoma and dark brown on propodeum of Iranian specimens, see Ghafouri Moghaddam et al. (2019), Figs 12A, B, p. 114); pterostigma clear honey yellow (Fig. 8B) (dark brown in the Iranian specimens, see Ghafouri Moghaddam et al. (2019), Fig. 12F, p. 114); metatibia clear yellow, somewhat whitish at basal third (Fig. 6B) (reddish yellow in the Iranian form); antenna shorter than body, 0.82�� as long as body (Fig. 6A) (longer than body in the Iranian form); metasomal T3 without any trace of median field (Fig. 8D) (with weak median field in the Iranian form). These are in addition to morphometric differences provided in Table 1. On the other hand, giving the proximity of Farasan Islands (Jazan) with Yemen, and that form A of D. mayae is the only form collected from Yemen (Ghafouri Moghaddam, pers. comm.), a comparison of our specimen with form A (Ghafouri Moghaddam et al. 2019), shows that they are completely different by the predominance of black on the body; the pale spot on pterostigma of fore wing; and the bicoloured metatibia (see Figs 7���10, Ghafouri Moghaddam et al. 2019). Extensive studies should be carried out on this species as a great deal of morphological variation is seen among samples studied from the Arabian Peninsula (Fernandez-Triana, pers. comm.), and is likely to be a species complex. Due to the scarcity of specimens, we considered our specimen to be another form of D. mayae. There are many differences in morphometric measurements (see Table 1) between our Saudi Arabian specimen and the forms in Ghafouri Moghaddam et al. (2019)., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on pages 59-60, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Shestakov, A. (1932) Zur Kenntnis der asiatischen Braconiden. Zoologische Annalen. Wurzburg, 99, 255 - 263.","Ghafouri Moghaddam, M., Rakhshani, E., van Achterberg, C. & Mokhtari, A. (2019) A taxonomic review of the genus Diolcogaster Ashmead (Hymenoptera: Braconidae: Microgastrinae) in Iran, distribution and morphological variability. Zootaxa, 4590 (1), 95 - 124. https: // doi. org / 10.11646 / zootaxa. 4590.1.4"]}
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42. Gnamptodon undetermined
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Arthropoda ,Gnamptodon ,Animalia ,Gnamptodon undetermined ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Gnamptodon sp. Figs 4 A���D Colour (Figs 4A���C, E): Head, mandible (except tip black), scape of antenna, two basal metasomal segments, legs, tegula clear yellowish, mesosoma and rest of metasomal tergites shiny black; antennal pedicel and basal two flagellomeres reddish brown, rest of flagellomeres black (broken); palpi whitish. Wings hyaline, pterostigma brown, veins partly pale brown (Fig. 4D). Head (Figs 4B, C). With scattered fine, whitish, appressed setae; antenna filiform, F1 about 1.29�� F2; F1 and F2 about 4.0�� and 3.0�� as long as wide, respectively; face convex when seen in lateral view; vertex coriaceous, rest of head smooth; malar space 1.3�� as long as basal width of mandible. Mesosoma (Fig. 4A). Length of mesosoma 1.5�� as long as its maximum width; all smooth and shiny, with few scattered whitish setae; mesoscutum without median sulcus, notauli short, not reaching middle of mesoscutum; scutellar suture narrow, linear; propodeum smooth, without median longitudinal carina, with slightly roundly protruding postero-lateral corners. Fore wing (Fig. 4D) hyaline, with brownish pterostigma and pale brown veins, radial cell short, distance between fore wing apex and apex of marginal cell 2.2�� length of 1R1; vein r very short, slightly shorter than vein 3���SR; vein SR1 7.0�� as long as vein 3���SR; pterostigma 1.7�� as long as 1R1; 1CU1:2CU1= 8: 62; vein SR1 slightly curved. Metasoma (Figs 4A, E). Smooth and shiny, with few scattered setae; T1 smooth, widened posteriorly, apical width 2.0�� basal width, 0.6�� as long as its apical width; T2 with a deep, linear transverse sulcus, with a transverse elevation close to basal margin, median length of basal elevation 0.4�� median length of the rest of tergite; T3 shorter than T2 (0.8��). Material examined. 1♂, KSA: Jazan (Farasan Islands, Park 2), 16.715789N 42.080407E, 5. iii. 2017, sweep net, leg. Usama M. Abul El-Ghiet & Tarek M. El-Sheikh. Remarks. In the present study, the subfamily Gnamptodontinae is recorded for the first time for the Arabian Peninsula from Farasan Islands (Jazan, Kingdom of Saudi Arabia). The specimen does not agree with any of the Afrotropical species included in van Achterberg���s key (1983), or those of Fischer���s (1987). However, because of the bad condition of the specimen, we have refrained from allocating a species name until more specimens are collected., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on pages 56-57, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Fischer, M. (1987) Hymenoptera Braconidae (Opiinae III) - athiopische, orientalische, australische und ozeanische Region. Das Tierreich, 104, 1 - 734."]}
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43. Aleiodes farasanensis Gadallah & Edmardash 2022, sp. nov
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Aleiodes farasanensis ,Braconidae ,Insecta ,Arthropoda ,Animalia ,Aleiodes ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Aleiodes farasanensis Gadallah & Edmardash sp. nov. ( apicalis -gp.) Figs 12 A���F, 13 A���C, 14 A, B urn:lsid:zoobank.org:act: BEAF16FD-9CCF-4EE7-AFB1-7DD8E6638C0C Type material: Holotype, ♀ (EFC), Saudi Arabia (Farasan Archipelago, Al-Sajid), 16.86026N, 41.932564E, 25.i.2017, light trap, leg. Usama Abu El-Ghiet & Tarek M. El-Sheikh. Diagnosis. Body pale reddish to orange, with metasomal T1 and T2 reddish, rest of metasoma black (Figs 12A, 14A). Antenna with 64 antennomeres (Fig. 12F); maximum width of hypoclypeal depression 0.54�� as long as maximum width of face (Fig. 12B); OOL 0.9 �� as long as diameter of median ocellus, finely punctate; F4 distinctly longer than wide (1.66��) (Fig. 12F); clypeus in the form of a narrow transverse band, with the same width medially and laterally (Figs 12B, E); mesoscutum reticulate-rugose (Fig. 13A); tarsal claws without pectens; marginal cell of fore wing ending rather close to wing apex (Fig. 13C); vein 1���CU1 of fore wing 0.26�� as long as 2���CU1 (Fig. 13C); SMC2 medium-sized (Fig. 13C); marginal cell of hind wing gradually widened to apex, apical width about 1.8�� its width at level of hamuli (Fig. 13C); distal end of hind femur with black hue (Fig. 12A); palpi and tegula pale red to yellowish, antenna dark brown to black (scape reddish) (Fig. 12F). Description. Holotype, ♀, length of body 10mm; length of fore wing 6.5mm. Colour (Figs 12A���C; 13A, B, 14A, B). Head dark ferruginous to black, with clypeus, mandible (except black tip), and palpi reddish brown; antenna dark brown to black, with scapus dark reddish brown; mesosoma and first two metasomal segments reddish brown to orange, remainder of metasoma black. Legs as body colour, with hind femur darker, with black tint apically, tarsal claws yellowish. Wings (Fig. 13C) hyaline, with black pterostigma and veins (except bases of veins paler). Head (Figs 12B���F). Antenna with 64 antennomeres, F4 distinctly longer than wide, 1.66�� as long as wide, scapus oblique apically, longer dorsally than ventrally; face (behind antennal bases) bulged (when seen in lateral view), with a middle longitudinal carina,densely finely punctate;face just above antennal bases until behind ocelli strongly depressed, obliquely striated; OOL 1.3�� as long as posterior ocellus diameter, reticulate rugose; vertex finely or superficially rugose, shiny; clypeus coarsely punctate, with interspaces between punctures equal to puncture diameter, with sharp, nearly straight ventral margin; width of hypoclypeal depression 0.54�� as wide as maximum width of face; eye 1.6�� as long as temple in dorsal view; vertex behind stemmaticum with radiated rugae; clypeus located below lower level of eye; mandible bidentate, with two blunt teeth; malar space 0.22�� length of eye in lateral view; occipital carina sharp, not reaching genal carina. Mesosoma (Figs 13A, B, 14B). Mesoscutum reticulate rugose; precoxal area of mesopleuron rugose-foveolate above, punctate ventrally, remainder of mesopleuron as well as metapleuron densely foveolate; scutellum flat, superficially punctate, without lateral carina; propodeum irregularly foveolate, foveola small, with distinct median longitudinal carina, not reaching posterior declivity. Wings (Fig. 13C). Fore wing: vein r 0.6�� 3���SR; marginal cell ends close to wing apex; 1���CU1 nearly horizontal, 0.26�� as long as 2���CU1; r-m 0.58�� as long as 3���SR; SMC2 medium-sized; 3-SR 1.6�� as long as 2-SR; cu-a slightly oblique, straight; vein 1-M slightly curved posteriorly. Hind wing: marginal cell distinctly gradually widened to apex, apical width about 1.8�� its width at level of hamuli; vein M +CU 1.6�� as long as 1-M; 1r���m 1.1�� as long as 1���M; vein m���cu very short, more or less interstitial with vein 1r���m. Legs (Fig. 12A). Tarsal claws simple (not pectinate), swollen at base; metacoxa densely superficially punctate dorsally; hind trochantellus slender, sparsely punctate; metafemur 1.8�� as long as hind basitarsus; length of inner metatibial spur 0.34 �� as long as hind basitarsus. Metasoma (Figs 12A, 14A). Metasomal T1 rather flattened, 1.1�� as long as its apical width; T1 and T2 with medio-longitudianl carina not reaching apical margin on T2, both tergites with coarse, longitudinal rugosity, that is somewhat weaker on T2; medio-basal area of T2 with wide, smooth triangle; basal half of T3 finely longitudinally striate, remainder of metasoma nearly smooth and shiny, with short, fine whitish setae; ovipositor sheath finger-like, very short, with fine setae throughout, rounded apically, 0.1�� as long as metatibia. Remarks. The species belongs to apicalis -sp. group as its characters strongly agrees with van Achterberg and Shaw���s key (2016; first part of couplet 3, p. 12). It is very similar to the formerly known species (now invalid species, van Achterberg, pers. comm.) Aleiodes africanus (Enderlein, 1920). Based on van Achterberg���s brief description (1991), the new species differs from A. africanus in the following combination of characters: head dark ferruginous, with some parts red (Figs 12B���D), metasoma (except first two segments red) black (Fig. 14A) (in A. africanus body reddish); 64 antennomeres (Fig. 12F) (70 antennomeres in A. africanus). Etymology. The name farasanensis is derived from the locality (Farasan Islands) from which the specimen was collected and described., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on pages 68-72, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Enderlein, G. (1920) Zur Kenntnis aussereuropaischer Braconiden. Archiv fur Naturgeschichte, 84 (A), 11 (1918), 51 - 224. https: // doi. org / 10.5962 / bhl. part. 13627"]}
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44. Yelicones vojnitsi Papp 1992
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Abu El-Ghiet, Usama M., Edmardash, Yusuf A., and Gadallah, Neveen S.
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Braconidae ,Insecta ,Arthropoda ,Yelicones ,Animalia ,Biodiversity ,Hymenoptera ,Yelicones vojnitsi ,Taxonomy - Abstract
Yelicones aff. vojnitsi Papp, 1992 Figs 9A���F Yelicones vojnitsi Papp, 1992: 152, 153. Diagnosis. elongated species (Fig. 9A), with head and mesosoma dark reddish brown (Figs 9B, C), metasoma (Fig. 9F), hind leg and basal half of antenna yellowish red; stemmaticum black; base of hind tibia and tarsus pale yellowish; fore wing slightly infuscate, pterostigma dark brown, clear yellow at base (Fig. 9D), veins pale brown. Head transverse, about 1.5�� as wide as long; vertex to ocellar area extending medially (narrowly) to base of clypeus peculiarly wrinkled (Fig. 9B); face 1.75�� as wide as its height; temple short and constricted behind eye, 0.3�� as long as eye height; 29 antennomeres, last flagellomeres submoniliform, last one with fine spine apically; mesoscutum densely punctate that are denser and closer on lateral lobes (Fig. 9C); propodeum coarsely rugose, transversely wrinkled posteriorly (Fig. 9C); mesopleuron shiny, nearly smooth to superficially punctate (Fig. 9A); fore wing with pterostigma 0.77�� as long as 1R1; marginal cell ending closer to wing apex; cell 2a relatively large, vein 2���SR+M non tubular medially; vein r relatively long and oblique, issued from near to the middle of pterostigma to base, r vein of fore wing 1.5�� as long as 3-SR, r: 3���SR: SR1 = 29: 19: 76; vein 1���SR linearly connected with 1���M; vein 1���M slightly curved to nearly straight; vein M+CU distinctly curved; cu-a postfurcal, vein 1���CU1 0.5�� as long as 2���CU1 (Fig. 9D); metasomal T1 distinctly widened posteriorly, its posterior margin 2.4�� as wide as its basal margin, not protruding posteriorly; all T1 and most of T2 (except posteriorly) longitudinally rugose; rest of tergites smooth and shiny, with few setae (Fig. 9F). Ovipositor sheaths very short, hardly exserted from tip of metasoma. Material examined. 1♀, KSA: Jazan (Farasan Islands, Al-Sajid), 16.860626N 41.932564E, 5. iii. 2017, light trap, leg. Usama M. Abu El-Ghiet & Tarek M. El-Sheikh. General distribution. Egypt, Oman (Quicke & Chishti, 1997), Saudi Arabia (Farasan Islands, Jazan) (new record). Remarks. The genus Yelicones is a first record for the fauna of Saudi Arabia. Characters of the Saudi Arabian specimen agree well with the Omani specimen in Quicke & Chishti���s key (1997, couplet 7, p. 78), except for: shortest distance between hind ocelli 0.9�� to nearly as long as distance between hind ocellus and eye (in Omani specimens, 1.8-2.5��); length of head behind eye 0.52�� as long as horizontal length of eye (measured perpendicular to plane of face) (In Omani specimens, 0.65��); the general body colour which is uniformly reddish yellow in the Omani specimens (in Saudi Arabian specimen, head and mesosoma dark reddish yellow with some parts ferruginous, metasoma orange). It also agrees with Papp���s key (1992, couplet 6, p. 155), as well as Papp���s description (1992, p. 152). It differs from the Tanzanian specimens (Papp 1992) in the following: temple 0.3�� as long as eye height (0.5�� in Tanzanian specimens); pterostigma 3.0�� as long as wide (2.64�� in Tanzanian specimen); vein r 1.6�� as long as pterostigmal width (slightly longer than pterostigmal width in Tanzanian specimens); SR1 4.0�� as long as 3���SR (4.5-5.0�� in the Tanzanian specimens); posterior width of T1 2.4�� as wide as its basal width (twice in the Tanzanian specimens). In addition to the above-mentioned references. Our specimen is compared with coloured photos of Y. vojnitsi from Egypt, Gambia and Namibia (British Museum of Natural History (London) (NHML)). All specimens are in bad condition, however, the main difference that could be seen is the general body colour, which is yellowish red (except black stemmaticum), while in our specimen, head and mesosoma are dark red, with some parts ferruginous, and metasoma light yellowish red., Published as part of Abu El-Ghiet, Usama M., Edmardash, Yusuf A. & Gadallah, Neveen S., 2022, New species and records of braconid wasps from Farasan Archipelago (Jazan) Saudi Arabia, with notes on Diolcogaster mayae Shestakov, pp. 49-79 in Zootaxa 5104 (1) on page 64, DOI: 10.11646/zootaxa.5104.1.3, http://zenodo.org/record/6257841, {"references":["Papp, J. (1992) New braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 3. Annales Musei historico-naturalis hungarici, 84, 129 - 160.","Quicke, D. L. J. & Chishti, M. J. K. (1997) A revision of the Yelicones species (Hymenoptera: Braconidae: Rogadinae) from Africa and The Arabian Peninsula, with description of four new species. African Entomology, 5 (1), 77 - 91"]}
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45. Tachysphex buyssoni Morice 1897
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Tachysphex ,Arthropoda ,Animalia ,Tachysphex buyssoni ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Tachysphex buyssoni Morice, 1897 (Figure 12 a-d) Tachysphex buyssoni Morice, 1897: 307, ♀, ♂. Diagnosis Gaster and legs (except black coxae and trochanters) red (Figure 12 (a,b)); labrum distinctly convex, protruding well beyond clypeus (Figure 12 (d)); mandible with large mandibular notch (Figure 12 (c))., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on page 1722, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026, {"references":["Morice FD. 1897. New or little-known Sphegidae from Egypt. Trans Entomol Soc Lon. 1897: 301 - 316, pl. VI."]}
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46. Cerceris vagula Kohl 1906
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Cerceris vagula ,Insecta ,Arthropoda ,Masaridae ,Cerceris ,Animalia ,Biodiversity ,Hymenoptera ,Taxonomy - Abstract
Cerceris vagula Kohl, 1906 (Figure 3 e-h) Cerceris vagula Kohl, 1906: 209, ♀. Diagnosis Black and white species (Figure 3 (e–h). Characterised by the following combination of characters: the male last flagellomere slightly obliquely truncate (Figure 3 (e)); clypeus with three small teeth at apex of median lobe (that are dark reddish brown and smooth) (Figure 3 (e)); clypeal moustache dense, pale, far apart, restricted to lateral side of clypeus (Figure 3 (e)); propodeum with a small V-shaped smooth and shiny dorsal enclosure, surrounded by a simple sulcus, with simple longitudinal sulcus medially (Figure 3 (f)), with setiferous punctures laterally; T2 with traingular yellowish white marking at base (Figure 3 (g)), T6 largely whitish (Figure 3 (h)); S2 with small, raised platform basally, S3 with two relatively large latero-apical whitish markings. Records from Saudi Arabia Wadi Majarish (below Taif) (Guichard 1993), Jazan (present study). General distribution Oman, Saudi Arabia, United Arab Emirates, Yemen. Comments Based on Guichard (1993), this species resembles Cerceris constricta Guichard, 1993 (as constrictus) in having T1 equally narrow from base to apex. However, it differs from C. constricta in having T3 with apical pale yellowish band that is narrowed medially and dilated laterally (narrow throughout in C. constricta); T2 with small triangular pale yellowish marking basally (absent in C. constricta); T6 almost whitish (crescent-like in C. constricta). Material examined 1 ♂, Jazan (Samba), 17º02 ʹ 44.8 ʺ N 42º39 ʹ 31.4 ʺ E, 18.xii.2020, on Citrus aurantiifolia (Christmas) [Malaise trap], leg. Usama Abu El-Ghiet., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on pages 1704-1705, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026, {"references":["Kohl FF. 1906. Zoologische Ergebnisse der Expedition der Kaiserlichen Akademie der Wissen schaften nach Sudarabien und Sokotra im Jahre 1898 - 1899. Hymenopteren. Denkenschriften der kaiserlichen Akademie der Wissenschaften in Wien Mathematisch-Naturwissenschaftliche Klasse. 71: 169 - 301.","Guichard KM. 1993. The genus Cerceris (Hymenoptera: Sphecidae) in Arabia. Fauna Saudi Arabia. 13: 152 - 169."]}
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47. Oryttus pulawskii Gadallah & Edmardash 2021, sp. nov
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Oryttus pulawskii ,Arthropoda ,Animalia ,Biodiversity ,Hymenoptera ,Oryttus ,Taxonomy - Abstract
Oryttus pulawskii Gadallah & Edmardash, sp. nov. (Figures 21 (a-d), 22(a-d), 23(a-d)) http://www.zoobank.org/urn:lsid:zoobank.org:act: D24522 D3-0705-45D0-9753- 8E9786CE746A Type material Male (holotype), Jazan (Damad), 17º08 ʹ 05.1 ʺ N 42º45 ʺ 58.8 ʺ E, 20–25.xii.2020, on Moringa oleifera Lam. (Moringaceae) [Malaise trap], leg. Usama Abu El-Ghiet; Paratype: 1 ♂, same data of holotype (with broken head, glued on a card). Diagnosis Oryttus pulawskii is well characterised by the following characters (Figures 21 (a–c)), 23(a)): body mainly black and yellow, with some ferrugineous and reddish brown areas; T1 not globular apically, slightly broader apically; T2 entirely dull black, superficially punctate with large distances between punctures; F8-10 deeply excavated beneath, excavation extremely large on F8; gastral S1 with median longitudinal carina (Figure 22 (d)); S7 very narrow, parallel-sided, deeply emarginate apically in the form of inverted V (Figure 23 (b)); fore wing subhyaline, with dark brown patches in marginal cell, apex of SMC1, all SMC2 and 2nd discoidal cell (Figure 22 (a)); middle tibia with two spurs (Figure 21 (d)). Description of male holotype Body length: 12 mm., fore wing length: 9.0 mm. Colouration (Figures 21 (a–c), 22(a) and 23(a,b)). Body generally black and yellow, with the following reddish brown to ferruginous: head (dark ferruginous), inner margins of eyes reddish above and yellowish below; clypeus reddish brown (except yellow laterally); pronotum and scape above ferruginous. Yellow are: scape beneath, maxillary and labial palps, scutellum (except at base), postscutellum, apical third of gastral T1, T3 (except at base), posterior two-thirds of T4, almost all T5 and T6, T7 (except brownish at apex), small postero-lateral triangular areas on S4-6. Pedicle dark brown, all flagellum orange to bright red. Legs black except: front femur apico-ventrally, inner surface of front tibia, an inner streak on mid tibia, ventral side of front tarsus yellow, all tarsi dark reddish brown; tegula dark brown. fore wing subhyaline, with dark brown patches in the marginal cell, apex of SMC1, all SMC2 and 2nd discoidal cell, pterostigma and wing veins dark brown. Head (Figure 21 (a,b)). Posteriorly and face densely finely punctate, sparser on posterior margin of head in dorsal view. Covered with very fine pubescence not concealing integument. Clypeus smooth, with dense fine, silvery pubescence not concealing integument, glabrous at apex, convex medially, with straight apical margin. Inner eye margins slightly convergent towards clypeus. Mandible with one subapical, small tooth. Antenna with scape cylindrical, slightly shorter than F1; F1-6 slender, distinctly longer than wide, last flagellomere long, narrowly rounded at apex; F8-10 deeply excavated beneath, excavation very large on F8. Thorax (Figure 21 (a,c)). Pronotum swollen medially, posterior margin with 2–3 transverse ridges. Mesoscutum densely coarsely punctate, with irregular, finely rugose interspaces. Scutellum and postscutellum superficially foveolate, ridged between foveoli especially on scutellum. Propodeum with V-shaped enclosure, with middle sulcus, coarsely wrinkled around sulcus, lateral sides around enclosure densely coarsely punctate. Lateral panel of pronotum with few oblique striae; mesopleuron densely areolated throughout, metapleuron coarsely rugose. Mesosternum dull, densely superficially punctate. Middle tibia with two spurs. Gaster (Figures 22 (b–d) and 23(a,b)). Gastral T1 somewhat widened posteriorly, nearly shiny and smooth yellow band, sparsely punctate above; T2 dull, superficially punctate, with large, finely alutaceous interspaces, remaining terga nearly smooth, impunctate. Gastral S1 with median longitudinal carina; S2, S3, and S6 sparsely punctate, sterna smooth; S7 very narrow, parallel-sided, with deep, inverted V-shaped emargination apically. Male genitalia: Figure 23 (c,d). Distribution Known only from Jazan (southwestern Saudi Arabia). Etymology The species is named in honour of the sphecid expert W. J. Pulawski, who helped in recognising that the species is an undescribed one., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on pages 1736-1740, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026
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48. Cerceris rubida
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Insecta ,Arthropoda ,Masaridae ,Cerceris ,Animalia ,Biodiversity ,Cerceris rubida ,Hymenoptera ,Taxonomy - Abstract
Cerceris rubida (Jurine, 1807) (Figure 2 c-e) Philanthus rubidus Jurine, 1807: 202, ♀. Diagnosis Small slender species, with the whitish gastral markings of irregular type (Figure 2 (c)); male clypeus with three inconspicuous teeth apically (Figure 2 (d)); forewing with second submarginal cell clearly petiolate; tegula smooth (Figure 2 (e)); T1 without a fossette apicomesally (Figure 2 (c)); S2 without elevated basomedian platform; propodeal enclosure coarsely punctate, punctures as surroundings but sparser (Figure 2 (e))., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on page 1702, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026, {"references":["Jurine L. 1807. Nouvelle methode de classer les Hymenopteres et les Dipteres. Hymenopteres. Tome premier. Vol. 1 - 4. Geneve: J. J. Paschoud; p. 1 - 319 1 unnumbered, 1 - 14 pls."]}
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49. Liris maidli
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Liris ,Biodiversity ,Hymenoptera ,Taxonomy ,Liris maidli - Abstract
Liris maidli (Arnold, 1929) (Figure 4 a-g) Notogonidea maidli Arnold, 1929: 397. Substitute name for Notogonia minima Maidl, 1924. Diagnosis Body (including antennae and legs) entirely black; fore wing without transverse infumated band (Figure 4 (e)); apical margin of clypeus shiny dark reddish brown, conspicuously arched (Figure 4 (b)); scape with sharp longitudinal carina along its whole ventral surface (Figure 4 (b)), flagellomeres distinctly longer than wide; mesoscutum very finely rugose, with dense fine silvery pubescence along its anterolateral half (Figure 4 (a)); propodeum distinctly elongate (Figure 4 (d)), dull, finely coriaceous, with short silvery pubescence laterally on dorsal upper surface, posterior surface with a conspicuous, relatively wide median sulcus, coarsely transversely striated, and clothed with dense silvery pubescence; mesopleuron and metapleuron densely finely coriaceous throughout hardly visible because of dense pubescence (Figure 4 (c)); gaster with inconspicuous broad, incomplete bands of silvery pubescence on posterior margins of T1-3, hardly visible on T4 (Figure 4 (f)); S1 with longitudinal carina postero-medially., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on page 1705, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026, {"references":["Arnold G. 1929. The Sphegidae of South Africa. Part XII. Ann Transvaal Mus. 13: 217 - 319."]}
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50. Trypoxylon Latreille 1796
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Gadallah, Neveen S., Abu El-Ghiet, Usama M., and Edmardash, Yusuf A.
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Crabronidae ,Insecta ,Arthropoda ,Animalia ,Biodiversity ,Trypoxylon ,Hymenoptera ,Taxonomy - Abstract
Genus Trypoxylon Latreille, 1796 Trypoxylon Latreille, 1796, p. 121, no included species. Type species: Sphex figulus Linnaeus, 1758, designated by Latreille 1802., Published as part of Gadallah, Neveen S., Abu El-Ghiet, Usama M. & Edmardash, Yusuf A., 2021, New data on the crabronid fauna of Saudi Arabia (Hymenoptera: Apoidea (Spheciformes): Crabronidae) from Jazan Province, with the description of three new species and the hitherto unknown male of Miscophus chrysis Kohl, 1894, pp. 1697-1748 in Journal of Natural History 55 (27 - 28) on page 1727, DOI: 10.1080/00222933.2021.1955993, http://zenodo.org/record/5530026, {"references":["Latreille PA. 1796. Precis de caracteres generiques des Insectes, dispose dans une ordre naturel. Paris et Brive: F. Bourdeaux; pp. I - XIV + 1 - 201 + 7 unnumbered, 1 pl.","Latreille PA. 1802. Histoire naturelle Generale et particuliere des Crustaces et des Inscetes. Ouvrage faisant suite a l'Histoire Naturelle Generale et particuliere, composee par Leclercq de Buffon et redigee par C. S. Sonnini, member de plusier F. Paris: Dufart; p. I - XII, 13 - 467."]}
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