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Dudgeodes palnius Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 59���77. Materials examined. Holotype: Male larva, India, Tamil Nadu, Palni hills, Perumal Malai stream, 10 o 16 ��� 15.39 N, 77 o 33 ��� 15.83 E, 1484 m; 28.IX. 2013, Colls. C. Selvakumar& K. G. Sivaramakrishnan [ZSI-I/E 13]. Paratypes: Four larvae, India, Tamil Nadu, Upper Kodaiyar, 08�� 31 ' 50.2 " N, 77 �� 21 ' 33.0" E, 1299 m; 06.IX. 2010, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 14]. Two larvae, same data as holotype [UM]; One larva, India, Tamil Nadu, Tirunelveli, Gadana river, Kallar, 0 8 o 48 '04.5" N, 77 o 18 '05.3" E, 144 m; 20.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Three larvae, India, Tamil Nadu, Tirunelveli, Puliyarai, Kannupulimedu, 0 8 o 56 ' 20.35 " N, 77 o 12 ' 25.74 " E, 164 m; 17.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. One larva, India, Tamil Nadu, Tirunelveli, Ramanathi river, 0 8 o 50 ' 53.4 "N, 77 o 18 ' 51.2 " E, 237 m; 21.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Description. Larva. Body length up to 3.5 mm and 3.0 mm, without cerci, in female and male larvae respectively; cerci length subequal to body length. General colouration highly variable, ranging from light to dark brown; young specimens generally paler; legs light to medium brown, with femora bearing 4 characteristic maculae (Fig. 59). Head: Outer margin of head fringed with row of short, basally forked setae from in front of eyes to labrum insertion. Antennae length 1.2 times head width, flagellum with 15 segments. Dorsal part of male eyes blackish. Labrum compact, ca. 2 times wider than long, with smooth anterior emargination; dorsal face covered medially by scattered simple and long setae; anterior margin with row of small, thin setae (Fig. 60). Hypopharynx with superlinguae oval with row of long, simple setae at apex (Fig. 61). Mandibles slender with one thin seta in middle of outer margin; left mandible (Fig. 64) with outer incisor consisting of one slightly concave teeth; inner incisor with two teeth inserted transversely, one smaller and pointed, one large and rounded, prostheca small; no setae below mola; right mandible (Fig. 65) with outer incisor composed of one large tooth, slightly indented medially; inner incisor with two teeth; prostheca reduced, with appearance of cluster of thin setae; row of 7 long and thin setae below mola and several short setae above mola. Maxilla slender, with well-developed canina, two indented dentisetae and three long setae on inner apical region and cluster of long, simple setae at crown; inner margin at base of lacinia, with two feathered and long setae, one dorsally and one ventrally; maxillary palp highly reduced (Fig. 62). Submentum well-developed laterally; glossae and paraglossae partially fused; paraglossae larger than glossae; labial palp three-segmented, segments 1 and 2 clearly visible, without constriction; segment 3 ca. 2.5 times as long as wide (Fig. 63). Thorax: Prothorax and mesothorax with 4 rounded tubercles on dorsal surface; outer margin fringed with row of short, basally-forked setae. Forefemur dilated, ca. 1.5 times longer than wide; outer margin covered by stout and long setae, meeting transverse row of both long and stout pointed setae across dorsal face; without thin setae at apex; inner margin with short row of long and thin setae proximally, reaching distally to transverse row (Figs. 66, 67). Middle and hind femora similar, more slender, ca. 2 times longer than wide; dorsal and inner margins each with row of long and stout setae (Figs. 69, 70). Tibia with row of long and stout setae on inner margin, and row of long and thin setae on outer margin (Fig. 68). Tarsal claw hooked, bearing three or four blunt teeth medially, and two pointed teeth subapically; outer tooth well developed, inner one smaller; apex of claw with two rows of three or four thin setae laterally (Fig. 71). Abdomen: Terga each with moderately developed median tubercle on segments I and X, more highly developed on segments II���IX (Fig. 72). No posterolateral projections on segments II���IV, but distinct on segments V���IX (Fig. 73). Lateral margins of terga with long thin setae. Gills on segments II���V (Figs. 74���77); gill II with dorsal lamella operculate, oval and with margin entire; gills III���V with dorsal lamella incised medially; ventral lobe flabelliform, well-developed and purple on gills II���IV. Cerci with stout setae every two or three segments; setae longer than length of corresponding segment. Winged stages. Unknown. Etymology. This species is named after the Palni hills of the Western Ghats from which type specimens were collected. Diagnosis. Dudgeodes palnius, n. sp., can be distinguished from other species of Dudgeodes and from the apparently similar species of D. hutanis by the following combination of characters: (i) apex of forefemur without seta (Fig. 67); (ii) a single ventral long seta on inner margin of galea-lacinia (Fig. 62); (iii) labial palp segments 1 and 2 clearly visible, without constriction (Fig. 63); (iv) left mandible with outer incisor consisting of one slightly concave teeth (Fig. 64) and (v) right mandible with outer incisor composed of one large tooth, slightly indented medially, with a row of seven long and thin setae below mola (Fig. 65)., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on pages 95-99, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885

Indoganodes Selvakumar, Sivaramakrishnan & Jacobus, n. gen. Figs. 21���39. Type species: Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n.sp. Description. Larva (Fig. 21). Head: Fringe or row of setae absent. Labrum subquadrate, approximately twice as broad as long, with short, scattered setae over entire dorsal surface(Fig. 22). Hypopharynx with lingua convex with minute setae at apical margin; superlingua oval with a row of long, simple setae at apex (Fig. 23). Mandibles robust; inner and outer incisors divergent; long mediolateral setae absent; medioapical outer incisor with at least one sharp denticle and apicomedial patch of setae absent on left mandible (Figs. 24, 25). Maxillae slender, with a well-developed canine and two dentisetae; two clusters of long setae apically; inner margin at middle with row of simple long setae; outer margin at base with minute hairs laterally; maxillary palp reduced to articulated palp (Fig. 26). Submentum welldeveloped laterally; glossae and paraglossae deeply divided; paraglossae somewhat pointed; labial palp 3 - segmented, with segments 1 and 2 subequal in length; segment 3 small and conical (Figs. 27, 28). Thorax: Prosternumwithout bilobular, spinous process medially (Fig. 29) (see Fig. 65 of McCafferty & Wang (1997) for prosternum with medial, bilobular, spiny process). Pro- and mesothorax without tubercles on dorsal surface. Forefemur relatively narrow, length 2.17 times width; outer margin covered by stout setae; scattered stout setae on apical region; inner margin with short thin setae (Fig. 30). Middle and hind femora similar, slightly more slender, length 2.3 times width (Figs. 31, 32); dorsal and inner margins with row of long, stout setae and irregular small thin setae. Fore tibia without setae on outer margin and irregular short thin setae on inner margin; middle and hind tibia with row of long stout setae and irregular short thin setae on inner and outer margins. Hind leg somewhat larger than fore and mid legs. Tarsal claw hooked, bearing 4 small denticles medially (Fig. 33). Abdomen: Gills absent on abdominal segment I; gills present on abdominal segments II���VI with margins entire (Figs. 34���38); gill II with dorsal lamella semi-operculate; gills II���V with ventral lobes flabelliform (Figs. 34���37); gill VI simple (Fig. 38). Poorly developed median tergal tubercles on segments I���IX; posterior margins of terga VI���IX each with 2���3 small spines flanking each side of median tubercle (total of 4���6 such posteromarginal spines on each tergum VI, VII, VIII or IX); posterolateral processes poorly developed on segments I���V and well developed on segments VI���IX (Fig. 39). Lateral margins of terga with thin stout setae. Caudal filaments banded; median caudal filament subequal in length to cerci. Winged stages. Unknown. Etymology. The new genus name is an arbitrary combination of letters, referring to the country of holotype origin, India, followed by the suffix ��� ganodes with reference to its inclusion in the family Teloganodidae. Diagnosis. The larvae of Indoganodes, n. gen.,can be distinguished from other genera of Teloganodidae, and from the apparently similar genus Ephemerellina (see McCafferty & Wang, 1997), by the following combination of characters: (i) prosternum without bi-lobular, spinous process medially (Fig. 29); (ii) abdominal posterolateral processes poorly developed on segments I���V and well developed on segments VI���IX (Fig. 39); (iii) tarsal claw hooked, bearing 4 small denticles medially (Fig. 33); (iv) labrum subquadrate, approximately twice as broad as long, with short, scattered setae over entire dorsal surface (Fig. 22); (v) superlinguae of hypopharynx moderately developed (Fig. 23) and (vi) left mandible without medioapical setal patch (Fig. 24). Species composition. Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n. sp. (India) Distribution. India (restricted to the Western Ghats)., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on pages 91-93, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885, {"references":["McCafferty, W. P. & Wang, T. - Q. (1997) Phylogenetic systematics of the family Teloganodidae (Ephemeroptera: Pannota). Annals of the Cape Provincial Museums (Natural History), 19, 387 - 437."]}

Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 21–39. Materials examined. Holotype: Male larva, India, Kerala, Silent Valley, Poochipara, 11 °06’ 49.5 ” N, 76 ° 25 ’ 52.4 ” E, 935 m, 18.IV. 2013, Coll. Jobin C. Tharian [ZSI-I/E 10]. Paratypes: Two larvae, same data as holotype [UM]. Description. Larva. Body 10–12 mm long. Cerci 12–14 mm long. General colouration very variable, ranging from grey to brown. Head: Dorsal part of male eyes blackish. Antennae length 2 times head width, pedicel dark brown. Mandibles with 2 or 3 pointed incisors with molar region consisting of 5 or 6 ridges; outer margin with minute hairs at base; right mandible with two rows of 6–8 long and thin setae below mola. Thorax: Pro- and mesothorax brownish; sternum yellowish. Fore, mid and hind legs yellowish. Abdomen: Terga brownish; anterior paler, posterior darker. Cerci brownish with stout setae shorter than length of corresponding segment. Etymology. This species is named in honour of Jobin C. Tharian for having magnanimously donated the specimens from his collection for the present study.

Teloganodes sartorii Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 40���58. Materials examined. Holotype: Male larva, India, Tamil Nadu, Salem, Yercaud, Killiyur falls, 11 o 47 ��� 40.05 ��� N, 78 o 11 ��� 59.62 ��� E, 1226 m; 20.VIII. 2010, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 11]. Paratypes: Five larvae, same data as holotype [ZSI-I/E 12]; 15 larvae, same data as holotype [UM]. Description. Larva. Body length 4���6 mm without cerci. Cerci slightly longer than body length. General dorsal colouration of head, thorax and abdomen darkbrown; sterna and legs uniformly yellowish. Eyes of male larva yellow and black (Fig. 40). Head: Outer margin of head fringed with regular row of setae from behind eyes to labrum insertion. Antennae long, 1.2 times head width, flagellum with 19���20 segments. Mouthparts similar to those of T. kodai (Figs. 41���46). Thorax: Pro and meso thorax with 6 rounded tubercles on dorsal face. Legs not similar in shape and without ornamentation; femoral submarginal row of short and blunt setae present. Forefemur moderately broad, ca. 1.5 times longer than wide; outer margin covered by thick, long setae and pointed setae (Fig. 47); two clusters of thin setae on apex (Fig. 50); 6���8 short and blunt setae on middle of dorsal surface; inner margin with row of long and thin setae, row continuing to dorsal face, close to articulation with trochanter. Mid and hind femora similar, slender, ca. 2 times longer than wide; dorsal margins each with row of long and stout setae and inner margins each with row of short, stout setae (Figs. 48, 49). Tibia and tarsal claw as in T. kodai (Fig. 51). Abdomen: No posterolateral projections on segments I���II, poorly developed on segments on segments II���VI, moderately developed on segments on segments VII���IX (Fig. 52). Tergal tubercles as in T. kodai (Fig. 53). Lateral margins of terga with short and thin setae. Gills on abdominal segments II���VI as in T. kodai (Figs. 54���58). Cerci with whorl of spineson every segment; spines shorter than length of corresponding segment. Winged stages. Unknown. Etymology. This species is named after Dr. Michel Sartori in recognition of his major contribution to our knowledge of Oriental Teloganodidae. Diagnosis. The larva of T. sartorii, n. sp., can be distinguished from the apparently similar species T. kodai by the following combination of characters: (i) legs not similar in shape and without ornamentation (Figs 47���49); (ii) outer margin of the forefemora having a row of thick setae and two clusters of thin setae on apex; (Fig. 50); (iii) antennae long, 1.2 times head width, flagellum with 19���20 segments; and (iv) cerci with whorl of spines on every segment; spines shorter than length of corresponding segment., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on page 94, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885

Janohyphella Selvakumar, Sivaramakrishnan & Jacobus, n. gen. Figs. 2���20. Type species: Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Description. Larva (Fig. 2). Head: Outer margin of head fringed with row of simple setae from behind eyes to labrum insertion (Fig. 3). Labrum short and broad; width less than 3 times length; distal margin broadly emarginate, with dense, fine, moderately long feathered setae; dorsal surface with scattered simple setae forming irregular row (Fig. 4). Hypopharynx with lingua and superlinguae subequal in length; lingua distally somewhat truncate to slightly coneshaped, not notched at apex; superlinguae broadly rounded distally (Fig. 5). Mandibles narrow, outer incisors separate and divergent; inner incisor of left mandible pointed, that of right mandible absent; with sparse mediolateral setae; otherwise with many slender long hair-like setae evenly distributed (Figs. 6, 7). Maxillae slender, with well-developed canine and two dentisetae; two clusters of long setae apically; inner margin smooth; outer margin at base with minute hairs laterally; maxillary palp reduced to articulated palp (Fig. 8). Labial palp segment 3 reduced in size; with relatively well-developed glossae and paraglossae; paraglossae somewhat bluntly pointed medioapically (Fig. 9). Submentum evenly rounded and lateral margin with sparse row of fine, short, hair-like setae. Thorax: Prosternum without medial, bilobular, spiny process (Fig. 3) (see Fig. 65 of McCafferty & Wang (1997) for prosternum with medial, bilobular, spiny process). Forefemur very broad and without transverse row of setae (Fig. 10); midfemur broad with transverse row of spatulate setae (Figs. 11, 12); hindfemur slender and without transverse row of setae (Fig. 13). Tarsal claws sharply curved, without denticles. Forewing pads with slight basal fusion (Fig. 14). Abdomen: Gills I small, single, two segmented and fibrilliform (Fig. 14); lamellate gills on abdominal segments II���V (Figs. 15���18); gills II operculate, appearing broadly rounded with median horizontal cleft on dorsal lamella (Fig. 15). Terga with two small tubercles separated by concave margin on segmentsII���VIII (Figs. 2, 19, 20), and with posterolateral processes well-developed on segments II���IX but not segment III (Fig. 20). Caudal filaments banded, fringed with lateral hairs. Median caudal filament length subequal to cerci. Winged stages. Unknown. Etymology. Janohyphella, n. gen., is an arbitrary combination of letters, with reference to Dr. Janice Peters, in honour of her significant contribution to the study of Oriental Ephemeroptera. Diagnosis. Janohyphella, n. gen., can be distinguished from all other genera of the Teloganodidae by the following combination of characters. In the larvae: (1) no reduction of median caudal filament (Fig. 2); (2) single, fibrilliform gills on abdominal segment I coupled with presence of lamellate gills on abdominal segment II���V (Figs. 14���18); (3) terga with two small tubercles separated by concave margin on segments III���VIII and with posterolateral processes well developed on segments II���IX, remarkably absent on segment III (Figs. 19, 20); (4) only midfemur has a transverse row of setae (Fig. 12); (5) tarsal claws sharply curved, without denticles (Fig. 13) and (6) mandible with sparse medio-lateral cluster of setae (Figs. 6, 7). Janohyphella, n. gen., is similar to the genus Manohyphella, but the new genus can be distinguished from it (see McCafferty & Benstead, 2002) by the following combination of characters in the larva: (1) terga with two small tubercles separated by concave margin on segments III���VIII and with posterolateral processes well developed on segments II���IX except segment III (Figs. 19, 20); (2) transverse row of setae only in midfemora (Fig. 12); (3) tarsal claws sharply curved, without denticles (Fig. 13) and (4) mandible with relatively sparse mediolateral setae (Figs. 6, 7). Species Composition. Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. (India) Distribution. India (restricted to the Western Ghats)., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on pages 88-89, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885, {"references":["McCafferty, W. P. & Wang, T. - Q. (1997) Phylogenetic systematics of the family Teloganodidae (Ephemeroptera: Pannota). Annals of the Cape Provincial Museums (Natural History), 19, 387 - 437.","McCafferty, W. P. & Benstead, J. P. (2002) Cladistic resolution and ecology of the Madagascar genus Manohyphella Allen (Ephemeroptera: Teloganodidae). Annales de Limnologie, 38, 41 - 52. http: // dx. doi. org / 10.1051 / limn / 2002005"]}

Derlethina tamiraparaniae Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 78���92. Materials examined. Holotype: Male larva, India, Tamilnadu, Tamiraparani river, Papanasam, 8 �� 42 ��� 37.1 ��� N, 77 �� 22 ���03.1��� E, 108 m, 15.V. 2013, Colls. C. Selvakumar& K. G. Sivaramakrishnan [ZSI-I/E 15]. Paratypes: Two larvae same data as holotype [UM]. One larva, India, Karnataka, Agumbe, Jogi Gunndi falls, 13 �� 29 ��� 55.95 ��� N, 75 ��06��� 10.85 ��� E, 514 m, 02.V. 2013, Colls. C. Selvakumar& K. G. Sivaramakrishnan [UM]. Description. Larva. Body length up to 3.5 ���4.0 mm and 4.0��� 4.5 mm in male and female respectively, cerci slightly longer than body length. Body flattened, eyes in dorsal position brownish in male larvae (Fig. 78). General colouration greyish black dorsally; scape and flagellum of the antennae whitish, pedicel blackish; legs yellowish, femora darker, with four dark brown maculae on dorsal face, maculae sometimes merging; apex of tibiae brownish. Head: Outer margin of head fringed with row of forked setae from behind eyes to labrum insertion. Antennae length 0.9 ���1.0 times head width, flagellum with 15 or 16 segments. Labrum with wide anteromedian emargination; anterior margin covered by thin setae; dorsal face with row of long and simple setae (Fig. 79). Hypopharynx with lingua broad, superlinguae quadrangular, outer margin almost straight, with row of simple and long setae at apex (Fig. 80). Mandibles slender with one thin seta in middle of outer margin; outer incisor of left mandible (Fig. 81) composed of one tooth indented basally, inner incisor with two teeth inserted transversally, one small and pointed, one large and rounded; prostheca reduced with several well-developed seta-like projections; no setae below mola; right mandible (Fig. 82) with outer incisor composed of one tooth, slightly indented medially; inner incisor with two teeth; prostheca reduced to seta-like cluster; row of three or four long, thin setae below mola and several short setae above mola. Maxilla moderately slender, with well-developed canine (Fig. 83);, two dentisetae and three long setae on inner apical portion and group of long, simple setae at crown; inner margin at base of lacinia with one feathered and long seta dorsally and two short, feathered setae ventrally (Fig. 83); maxillary palp highly reduced. Labial palp three-segmented, but articulation between segments 1 and 2 obscure; segment 3 ca. 2.0 times as long as wide at base (Fig. 84). Thorax: Prothorax and mesothorax without tubercles on dorsal face. Forefemur dilated, ca. 1.5���1.6 times longer than wide; outer margin covered by stout and long setae, and dorsal face broad and dilated with transverse row of stout and blunt setae (Fig. 85) almost reaching inner margin; inner margin with row of long and thin setae almost reaching apex of femur. Foretibia with one visible row of long and thin setae on inner margin, and another on outer margin. Middle femur more slender, ca. 2.0��� 2.2 times longer than wide; dorsal and inner margins with row of long and stout setae (Fig. 86). Tibia with a row of long and stout setae on inner margin, and row of long, thin setae on outer margin. Hind leg similar, ca. 2.5 times longer than wide, outer margin not concave (Fig. 87). Tarsal claw hooked, with three or four pointed teeth medially, and two pointed teeth subapically, with inner tooth reduced; apex of claw with two rows of three or four thin setae laterally. Abdomen: Terga without median tubercles on segments I���IV, but poorly developed on segments V���IX, and absent on segment X (Fig. 88). Posterolateral projections absent from segments II���IV, present and well-developed on segments V���IX; lateral margins of terga with very long and thin setae (Fig. 89). Gills on abdominal segments II���IV (Figs. 90���92), gills II���III with ventral lobes, gill IV without ventral lobe; gills II with dorsal lamella operculate, oval and with margin entire; gills III with dorsal lamella incised medially; gills IV with dorsal lamella incised medially; ventral lobe flabelliform, well developed and purple on gills II and III. Cerci with stout setae longer than length of corresponding segment. Terminal filament highly reduced. Winged stages. Unknown. Etymology. This species named after the type locality, the Tamiraparani River. Diagnosis. Derlethina tamiraparaniae, n. sp., can be distinguished from D. eloisae by the following combination ofcharacters: (i) gill IV incised (Fig. 92); (ii) outer margin of hind femora not concave (Fig. 87) and (iii) prostheca of left mandible reduced, but with several well-developed seta-like projections (Fig. 81)., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on pages 99-101, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885

Teloganodes sartorii Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 40–58. Materials examined. Holotype: Male larva, India, Tamil Nadu, Salem, Yercaud, Killiyur falls, 11 o 47 ’ 40.05 ” N, 78 o 11 ’ 59.62 ” E, 1226 m; 20.VIII. 2010, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 11]. Paratypes: Five larvae, same data as holotype [ZSI-I/E 12]; 15 larvae, same data as holotype [UM]. Description. Larva. Body length 4–6 mm without cerci. Cerci slightly longer than body length. General dorsal colouration of head, thorax and abdomen darkbrown; sterna and legs uniformly yellowish. Eyes of male larva yellow and black (Fig. 40). Head: Outer margin of head fringed with regular row of setae from behind eyes to labrum insertion. Antennae long, 1.2 times head width, flagellum with 19–20 segments. Mouthparts similar to those of T. kodai (Figs. 41–46). Thorax: Pro and meso thorax with 6 rounded tubercles on dorsal face. Legs not similar in shape and without ornamentation; femoral submarginal row of short and blunt setae present. Forefemur moderately broad, ca. 1.5 times longer than wide; outer margin covered by thick, long setae and pointed setae (Fig. 47); two clusters of thin setae on apex (Fig. 50); 6–8 short and blunt setae on middle of dorsal surface; inner margin with row of long and thin setae, row continuing to dorsal face, close to articulation with trochanter. Mid and hind femora similar, slender, ca. 2 times longer than wide; dorsal margins each with row of long and stout setae and inner margins each with row of short, stout setae (Figs. 48, 49). Tibia and tarsal claw as in T. kodai (Fig. 51). Abdomen: No posterolateral projections on segments I–II, poorly developed on segments on segments II–VI, moderately developed on segments on segments VII–IX (Fig. 52). Tergal tubercles as in T. kodai (Fig. 53). Lateral margins of terga with short and thin setae. Gills on abdominal segments II–VI as in T. kodai (Figs. 54–58). Cerci with whorl of spineson every segment; spines shorter than length of corresponding segment. Winged stages. Unknown. Etymology. This species is named after Dr. Michel Sartori in recognition of his major contribution to our knowledge of Oriental Teloganodidae. Diagnosis. The larva of T. sartorii, n. sp., can be distinguished from the apparently similar species T. kodai by the following combination of characters: (i) legs not similar in shape and without ornamentation (Figs 47–49); (ii) outer margin of the forefemora having a row of thick setae and two clusters of thin setae on apex; (Fig. 50); (iii) antennae long, 1.2 times head width, flagellum with 19–20 segments; and (iv) cerci with whorl of spines on every segment; spines shorter than length of corresponding segment.

Dudgeodes palnius Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 59–77. Materials examined. Holotype: Male larva, India, Tamil Nadu, Palni hills, Perumal Malai stream, 10 o 16 ’ 15.39 N, 77 o 33 ’ 15.83 E, 1484 m; 28.IX. 2013, Colls. C. Selvakumar& K. G. Sivaramakrishnan [ZSI-I/E 13]. Paratypes: Four larvae, India, Tamil Nadu, Upper Kodaiyar, 08º 31 ' 50.2 " N, 77 º 21 ' 33.0" E, 1299 m; 06.IX. 2010, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 14]. Two larvae, same data as holotype [UM]; One larva, India, Tamil Nadu, Tirunelveli, Gadana river, Kallar, 0 8 o 48 '04.5" N, 77 o 18 '05.3" E, 144 m; 20.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Three larvae, India, Tamil Nadu, Tirunelveli, Puliyarai, Kannupulimedu, 0 8 o 56 ' 20.35 " N, 77 o 12 ' 25.74 " E, 164 m; 17.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. One larva, India, Tamil Nadu, Tirunelveli, Ramanathi river, 0 8 o 50 ' 53.4 "N, 77 o 18 ' 51.2 " E, 237 m; 21.VII. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Description. Larva. Body length up to 3.5 mm and 3.0 mm, without cerci, in female and male larvae respectively; cerci length subequal to body length. General colouration highly variable, ranging from light to dark brown; young specimens generally paler; legs light to medium brown, with femora bearing 4 characteristic maculae (Fig. 59). Head: Outer margin of head fringed with row of short, basally forked setae from in front of eyes to labrum insertion. Antennae length 1.2 times head width, flagellum with 15 segments. Dorsal part of male eyes blackish. Labrum compact, ca. 2 times wider than long, with smooth anterior emargination; dorsal face covered medially by scattered simple and long setae; anterior margin with row of small, thin setae (Fig. 60). Hypopharynx with superlinguae oval with row of long, simple setae at apex (Fig. 61). Mandibles slender with one thin seta in middle of outer margin; left mandible (Fig. 64) with outer incisor consisting of one slightly concave teeth; inner incisor with two teeth inserted transversely, one smaller and pointed, one large and rounded, prostheca small; no setae below mola; right mandible (Fig. 65) with outer incisor composed of one large tooth, slightly indented medially; inner incisor with two teeth; prostheca reduced, with appearance of cluster of thin setae; row of 7 long and thin setae below mola and several short setae above mola. Maxilla slender, with well-developed canina, two indented dentisetae and three long setae on inner apical region and cluster of long, simple setae at crown; inner margin at base of lacinia, with two feathered and long setae, one dorsally and one ventrally; maxillary palp highly reduced (Fig. 62). Submentum well-developed laterally; glossae and paraglossae partially fused; paraglossae larger than glossae; labial palp three-segmented, segments 1 and 2 clearly visible, without constriction; segment 3 ca. 2.5 times as long as wide (Fig. 63). Thorax: Prothorax and mesothorax with 4 rounded tubercles on dorsal surface; outer margin fringed with row of short, basally-forked setae. Forefemur dilated, ca. 1.5 times longer than wide; outer margin covered by stout and long setae, meeting transverse row of both long and stout pointed setae across dorsal face; without thin setae at apex; inner margin with short row of long and thin setae proximally, reaching distally to transverse row (Figs. 66, 67). Middle and hind femora similar, more slender, ca. 2 times longer than wide; dorsal and inner margins each with row of long and stout setae (Figs. 69, 70). Tibia with row of long and stout setae on inner margin, and row of long and thin setae on outer margin (Fig. 68). Tarsal claw hooked, bearing three or four blunt teeth medially, and two pointed teeth subapically; outer tooth well developed, inner one smaller; apex of claw with two rows of three or four thin setae laterally (Fig. 71). Abdomen: Terga each with moderately developed median tubercle on segments I and X, more highly developed on segments II–IX (Fig. 72). No posterolateral projections on segments II–IV, but distinct on segments V–IX (Fig. 73). Lateral margins of terga with long thin setae. Gills on segments II–V (Figs. 74–77); gill II with dorsal lamella operculate, oval and with margin entire; gills III–V with dorsal lamella incised medially; ventral lobe flabelliform, well-developed and purple on gills II–IV. Cerci with stout setae every two or three segments; setae longer than length of corresponding segment. Winged stages. Unknown. Etymology. This species is named after the Palni hills of the Western Ghats from which type specimens were collected. Diagnosis. Dudgeodes palnius, n. sp., can be distinguished from other species of Dudgeodes and from the apparently similar species of D. hutanis by the following combination of characters: (i) apex of forefemur without seta (Fig. 67); (ii) a single ventral long seta on inner margin of galea-lacinia (Fig. 62); (iii) labial palp segments 1 and 2 clearly visible, without constriction (Fig. 63); (iv) left mandible with outer incisor consisting of one slightly concave teeth (Fig. 64) and (v) right mandible with outer incisor composed of one large tooth, slightly indented medially, with a row of seven long and thin setae below mola (Fig. 65).

Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S., Arumugam, M. (2014): Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India. Zootaxa 3846 (1): 87-104, DOI: http://dx.doi.org/10.11646/zootaxa.3846.1.4

Janohyphella Selvakumar, Sivaramakrishnan & Jacobus, n. gen. Figs. 2–20. Type species: Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Description. Larva (Fig. 2). Head: Outer margin of head fringed with row of simple setae from behind eyes to labrum insertion (Fig. 3). Labrum short and broad; width less than 3 times length; distal margin broadly emarginate, with dense, fine, moderately long feathered setae; dorsal surface with scattered simple setae forming irregular row (Fig. 4). Hypopharynx with lingua and superlinguae subequal in length; lingua distally somewhat truncate to slightly coneshaped, not notched at apex; superlinguae broadly rounded distally (Fig. 5). Mandibles narrow, outer incisors separate and divergent; inner incisor of left mandible pointed, that of right mandible absent; with sparse mediolateral setae; otherwise with many slender long hair-like setae evenly distributed (Figs. 6, 7). Maxillae slender, with well-developed canine and two dentisetae; two clusters of long setae apically; inner margin smooth; outer margin at base with minute hairs laterally; maxillary palp reduced to articulated palp (Fig. 8). Labial palp segment 3 reduced in size; with relatively well-developed glossae and paraglossae; paraglossae somewhat bluntly pointed medioapically (Fig. 9). Submentum evenly rounded and lateral margin with sparse row of fine, short, hair-like setae. Thorax: Prosternum without medial, bilobular, spiny process (Fig. 3) (see Fig. 65 of McCafferty & Wang (1997) for prosternum with medial, bilobular, spiny process). Forefemur very broad and without transverse row of setae (Fig. 10); midfemur broad with transverse row of spatulate setae (Figs. 11, 12); hindfemur slender and without transverse row of setae (Fig. 13). Tarsal claws sharply curved, without denticles. Forewing pads with slight basal fusion (Fig. 14). Abdomen: Gills I small, single, two segmented and fibrilliform (Fig. 14); lamellate gills on abdominal segments II–V (Figs. 15–18); gills II operculate, appearing broadly rounded with median horizontal cleft on dorsal lamella (Fig. 15). Terga with two small tubercles separated by concave margin on segmentsII–VIII (Figs. 2, 19, 20), and with posterolateral processes well-developed on segments II–IX but not segment III (Fig. 20). Caudal filaments banded, fringed with lateral hairs. Median caudal filament length subequal to cerci. Winged stages. Unknown. Etymology. Janohyphella, n. gen., is an arbitrary combination of letters, with reference to Dr. Janice Peters, in honour of her significant contribution to the study of Oriental Ephemeroptera. Diagnosis. Janohyphella, n. gen., can be distinguished from all other genera of the Teloganodidae by the following combination of characters. In the larvae: (1) no reduction of median caudal filament (Fig. 2); (2) single, fibrilliform gills on abdominal segment I coupled with presence of lamellate gills on abdominal segment II–V (Figs. 14–18); (3) terga with two small tubercles separated by concave margin on segments III–VIII and with posterolateral processes well developed on segments II–IX, remarkably absent on segment III (Figs. 19, 20); (4) only midfemur has a transverse row of setae (Fig. 12); (5) tarsal claws sharply curved, without denticles (Fig. 13) and (6) mandible with sparse medio-lateral cluster of setae (Figs. 6, 7). Janohyphella, n. gen., is similar to the genus Manohyphella, but the new genus can be distinguished from it (see McCafferty & Benstead, 2002) by the following combination of characters in the larva: (1) terga with two small tubercles separated by concave margin on segments III–VIII and with posterolateral processes well developed on segments II–IX except segment III (Figs. 19, 20); (2) transverse row of setae only in midfemora (Fig. 12); (3) tarsal claws sharply curved, without denticles (Fig. 13) and (4) mandible with relatively sparse mediolateral setae (Figs. 6, 7). Species Composition. Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. (India) Distribution. India (restricted to the Western Ghats).

Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 2–20. Materials examined. Holotype: Female larva, India, Tamilnadu, Tirunelveli, Nambiyar river, Nambikovil, 8 ° 26 ’01.22” N, 77 ° 29 ’ 55.07 ” E, 412 m, 23.II. 2012, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 9]. Paratype: One larva, India, Karnataka, Sringeri, Nanthinihole, 13 ° 23 ’ 23.52 ” N, 75 ° 10 ’ 47.02 ” E, 640 m, 03.V. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Description. Larva. Body length 4.0 – 4.5 mm; cerci length 2.0 mm. Ratio of body length versus cerci> 1.3. General colouration yellowish brown to dark brown markings dorsally, pale ventrally. Head: Markings variable, in general yellowish brown, paler between compound eyes and immediately anterior to median ocellus. Ratio of head width to antenna length Thorax: Femora with dark maculae dorsally; marginal armature consisting of moderately long, sharp, spinelike setae; fore femora with single macula dorsally (Fig. 10); mid- and hind femora with two maculae dorsomedially and two maculae dorsodistally (Fig. 11, 13); transverse row of setae only present on midfemur, pattern of arrangement as in Fig. 12. Tibiae with maculae; tarsi with median transverse band; claws yellowish brown, sharply curved, without denticles. Abdomen: Terga reddish brown, paler medially with paired black round maculae on tergum 6 anterolaterally (Fig. 20). Etymology. The new species is named after the country of holotype origin, India.

Janohyphella indica Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 2���20. Materials examined. Holotype: Female larva, India, Tamilnadu, Tirunelveli, Nambiyar river, Nambikovil, 8 �� 26 ���01.22��� N, 77 �� 29 ��� 55.07 ��� E, 412 m, 23.II. 2012, Colls. C. Selvakumar & K. G. Sivaramakrishnan [ZSI-I/E 9]. Paratype: One larva, India, Karnataka, Sringeri, Nanthinihole, 13 �� 23 ��� 23.52 ��� N, 75 �� 10 ��� 47.02 ��� E, 640 m, 03.V. 2013, Colls. C. Selvakumar & K. G. Sivaramakrishnan [UM]. Description. Larva. Body length 4.0 ��� 4.5 mm; cerci length 2.0 mm. Ratio of body length versus cerci> 1.3. General colouration yellowish brown to dark brown markings dorsally, pale ventrally. Head: Markings variable, in general yellowish brown, paler between compound eyes and immediately anterior to median ocellus. Ratio of head width to antenna length Etymology. The new species is named after the country of holotype origin, India., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on pages 89-91, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885

Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n. sp. Figs. 21���39. Materials examined. Holotype: Male larva, India, Kerala, Silent Valley, Poochipara, 11 ��06��� 49.5 ��� N, 76 �� 25 ��� 52.4 ��� E, 935 m, 18.IV. 2013, Coll. Jobin C. Tharian [ZSI-I/E 10]. Paratypes: Two larvae, same data as holotype [UM]. Description. Larva. Body 10���12 mm long. Cerci 12���14 mm long. General colouration very variable, ranging from grey to brown. Head: Dorsal part of male eyes blackish. Antennae length 2 times head width, pedicel dark brown. Mandibles with 2 or 3 pointed incisors with molar region consisting of 5 or 6 ridges; outer margin with minute hairs at base; right mandible with two rows of 6���8 long and thin setae below mola. Thorax: Pro- and mesothorax brownish; sternum yellowish. Fore, mid and hind legs yellowish. Abdomen: Terga brownish; anterior paler, posterior darker. Cerci brownish with stout setae shorter than length of corresponding segment. Etymology. This species is named in honour of Jobin C. Tharian for having magnanimously donated the specimens from his collection for the present study., Published as part of Selvakumar, C., Sivaramakrishnan, K. G., Jacobus, Luke M., Janarthanan, S. & Arumugam, M., 2014, Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India, pp. 87-104 in Zootaxa 3846 (1) on page 94, DOI: 10.11646/zootaxa.3846.1.4, http://zenodo.org/record/249885

Indoganodes Selvakumar, Sivaramakrishnan & Jacobus, n. gen. Figs. 21–39. Type species: Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n.sp. Description. Larva (Fig. 21). Head: Fringe or row of setae absent. Labrum subquadrate, approximately twice as broad as long, with short, scattered setae over entire dorsal surface(Fig. 22). Hypopharynx with lingua convex with minute setae at apical margin; superlingua oval with a row of long, simple setae at apex (Fig. 23). Mandibles robust; inner and outer incisors divergent; long mediolateral setae absent; medioapical outer incisor with at least one sharp denticle and apicomedial patch of setae absent on left mandible (Figs. 24, 25). Maxillae slender, with a well-developed canine and two dentisetae; two clusters of long setae apically; inner margin at middle with row of simple long setae; outer margin at base with minute hairs laterally; maxillary palp reduced to articulated palp (Fig. 26). Submentum welldeveloped laterally; glossae and paraglossae deeply divided; paraglossae somewhat pointed; labial palp 3 - segmented, with segments 1 and 2 subequal in length; segment 3 small and conical (Figs. 27, 28). Thorax: Prosternumwithout bilobular, spinous process medially (Fig. 29) (see Fig. 65 of McCafferty & Wang (1997) for prosternum with medial, bilobular, spiny process). Pro- and mesothorax without tubercles on dorsal surface. Forefemur relatively narrow, length 2.17 times width; outer margin covered by stout setae; scattered stout setae on apical region; inner margin with short thin setae (Fig. 30). Middle and hind femora similar, slightly more slender, length 2.3 times width (Figs. 31, 32); dorsal and inner margins with row of long, stout setae and irregular small thin setae. Fore tibia without setae on outer margin and irregular short thin setae on inner margin; middle and hind tibia with row of long stout setae and irregular short thin setae on inner and outer margins. Hind leg somewhat larger than fore and mid legs. Tarsal claw hooked, bearing 4 small denticles medially (Fig. 33). Abdomen: Gills absent on abdominal segment I; gills present on abdominal segments II–VI with margins entire (Figs. 34–38); gill II with dorsal lamella semi-operculate; gills II–V with ventral lobes flabelliform (Figs. 34–37); gill VI simple (Fig. 38). Poorly developed median tergal tubercles on segments I–IX; posterior margins of terga VI–IX each with 2–3 small spines flanking each side of median tubercle (total of 4–6 such posteromarginal spines on each tergum VI, VII, VIII or IX); posterolateral processes poorly developed on segments I–V and well developed on segments VI–IX (Fig. 39). Lateral margins of terga with thin stout setae. Caudal filaments banded; median caudal filament subequal in length to cerci. Winged stages. Unknown. Etymology. The new genus name is an arbitrary combination of letters, referring to the country of holotype origin, India, followed by the suffix – ganodes with reference to its inclusion in the family Teloganodidae. Diagnosis. The larvae of Indoganodes, n. gen.,can be distinguished from other genera of Teloganodidae, and from the apparently similar genus Ephemerellina (see McCafferty & Wang, 1997), by the following combination of characters: (i) prosternum without bi-lobular, spinous process medially (Fig. 29); (ii) abdominal posterolateral processes poorly developed on segments I–V and well developed on segments VI–IX (Fig. 39); (iii) tarsal claw hooked, bearing 4 small denticles medially (Fig. 33); (iv) labrum subquadrate, approximately twice as broad as long, with short, scattered setae over entire dorsal surface (Fig. 22); (v) superlinguae of hypopharynx moderately developed (Fig. 23) and (vi) left mandible without medioapical setal patch (Fig. 24). Species composition. Indoganodes jobini Selvakumar, Sivaramakrishnan & Jacobus, n. sp. (India) Distribution. India (restricted to the Western Ghats).