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Parachernes (Parachernes) nitidimanus (Ellingsen, 1905) Material examined. 1 female, Pernambuco, Brazil: Furna do Morcego cave, 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR010). Diagnosis. According to Ellingsen (1905) and Beier (1932b) can be easily distinguished from the other species of the genus Parachernes Chamberlin (1931) by the carapace as long as broad 1.14×, gradually narrowing forwards, the short frontal margin straightened. Metazone distinctly granulated, yet a little glossy before the first furrow. Two distinct transverse furrows, the anterior one about in the middle, the posterior one somewhat nearer to the hinder margin than to the anterior furrow. Tergite divided (except XI), slightly granulated, al setae rather long, slightly clavate. Chaetotaxy tergal: 10: 10: 10: 8(1): 8(2): 6(2): 8(2): 8(2): 8(2): 10(2): 4(2ST):2. Genital region opercula with a central group of 18 short, three setae on each side, and ten marginal setae. Chelicerae, female with galea very slender and short, with some fine teeth in and near the extremity on the underside, male with about 6 terminal branches; paired spermatheca, typical of the genus. Pedipalps moderately strong, all segments smooth and glossy. The setae on the inner side of the trochanter and femur are slightly clavate, the other setae of the palps are slightly dentate in the extremity, on the fingers simple with some longer ones intermixed; trochanter and the proximal part of femur slightly shagreened; femur 2.35×, strongly granulated; patella 2.02×, more slightly, rather shagreened. chela with pedicel 3.18×, chela without pedicel 2.47×, hand round about smooth and very glossy, fingers as general smooth and glossy. Measurements (mm). Body length: 2.359. Carapace: 0.771 / 0.672. Pedipalps: trochanter: 0.347 / 0.197, femur: 0.544 / 0.235, patella: 0.589 / 0.266, chela with pedicel: 1.064 / 0.384, chela without pedicel: 0.950 / 0.384, movable finger: 0.538. Chelicera: 0.277 / 0.141, movable finger length:0.202. Leg I: femur: 0.105 / 0.127, patella: 0.317 / 0.137, tibia: 0.272 / 0.091, tarsus: 0.308 / 0.071. Leg IV: femur+patella: 0.569 / 0.158, tibia: 0.318 / 0.103, tarsus: 0.255 / 0.052. Distribution. Brazil, Saint Vincent, and the Grenadines (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 320, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Ellingsen, E. (1905) Pseudoscorpions from South America collected by Dr. A. Borelli, A. Bertoni de Winkelried, and Prof. Goeldi. Bollettino dei Musei di Zoologia e di Anatomia Comparata della R. Universita di Torino, 20 (500), 1 - 17. https: // doi. org / 10.5962 / bhl. part. 9304","Beier, M. (1932 b) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i - xxi + 1 - 294.","Chamberlin, J. C. (1931) The arachnid order Chelonethida. Stanford University Publications, Biological Science, 7 (1), 1 - 284.","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

Ceriochernes foliaceosetosus Beier, 1974 Material examined. 1 female, Pernambuco, Brazil: Meu Rei cave, 08°29’14.1”S, 37° 16’48.8”W, 777 m a.s.l., 19- I-2016, Catimbau National Park, E. Barbier leg. (LECA; Ps-012). Diagnosis. According to the description made by Beier (1974), the specimens of Ceriochernes foliaceosetosus Beier, 1974 can be easily distinguished by the carapace reddish-brown, slightly longer than wide, without eyes and eyespots, strongly granulated, the granules of the metazone covered with an epicuticle, reticulate in the form of a rough honeycomb, both transverse furrow narrow and not very deep, but distinct, the anterior curved laterally, the posterior closer to the rear margin, with six setae on the anterior margin, and eight on the posterior margin. All tergites divided, setae clavate and broadly foliated, rounded at the tip, curved and almost sessile, partially covered by epicuticle, with midrib, pleural membrane very densely granulated. Chaetotaxy tergal: 6: 8: 8: 6: 6: 8: 8: 8: 8: 8: 8: 2. Genital region operculum with a set of 12 setae, and ten setae on posterior margin. Chelicerae with six to seven setae on hand, es very short, three blades on the rallum, Serrula with 17 blades, galea with 5 very short branches distally. Pedipalps strongly reticulated, trochanter 1.0×, femur 2.58×, patella 2.15×, fingers not as long as the hand without pedicel, each with about 35–40 marginal teeth, the accessory teeth small and inconspicuous, present on the medial side only on the distal part of the finger, trichobothrium ist midway between isb and it, est proximal of ist; trichobothrium st closest to t. Measurements (mm). Body length: 2.021. Carapace: 0.638 / 0.538. Pedipalps: trochanter: 0.344 / 0.244, femur: 0.522 / 0.204, patella: 0.569 / 0.204, chela with pedicel: 0.864 / 0.346, chela without pedicel: 0.833 / 0.346, movable finger: 0.383. Leg I: trochanter: 0.124 / 0.105, femur: 0.127 / 0.093, patella: 0.255 / 0.093, tibia: 0.275 / 0.072, tarsus: 0.283 / 0.053. Leg IV: trochanter: 0.183 / 0.140, femur+patella: 0.488 / 0.152, tibia: 0.377 / 0.109, tarsus: 0.385 / 0.088. Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 321, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Beier, M. (1974) Brasilianische Pseudoscorpione aus dem Museum in Genf. Revue Suisse de Zoologie, 81, 899 - 909. https: // doi. org / 10.5962 / bhl. part. 76050","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

Petterchernes brasiliensis Heurtault, 1986 Material examined. 1 male, Pernambuco, Brazil: Furna do Morcego cave, 8°34’14.1”S; 37°22’55.6”W, 556 m a.s.l., 09-V-2019; Catimbau National Park, E. Barbier leg. (LECA; Ps-SCR011). Diagnosis. According to Heurtault (1986) and Mahnert (1994) the males of Petterchernes brasiliensis Heurtault, 1986 can be easily distinguished by the carapace without eyes and without eyespots, dark brown, metazone finely granulated, setae of the carapace and tergites strongly clavate, with two deep transverse furrows, for setae on anterior margin; tergites divided, granulate. Chaetotaxy tergal: 9: 9: 9: 10: 10: 10: 12: 12: 10: 10: 10: 2. Genital region operculum with 32 setae and ten setae on posterior margin. Chelicerae, galea with multiple branches, and three blades in the rallum. Pedipalps coarsely granulate, setae stout and enlarged, trochanter 1.67×, with high dorsal hump, femur 2.22×, abruptly enlarged, patella 2.15×, chela with pedicel 2.28×, chela without pedicel 2.09×, trichobothrium isb, ib, and est at same level, hand with well pronounced internal hump bearing some longer and stouter setae. Legs, tarsus IV without tactile setae, claws smooth and simple, subterminal seta smooth, curve. Measurements (mm). Body length: 1.251. Carapace: 0.554 / 0.473. Pedipalps: trochanter: 0.269 / 0.161, femur: 0.477 / 0.214, patella: 0.415 / 0.193, chela with pedicel: 0.766 / 0.335, chela without pedicel: 0.7020 / 0.335, movable finger: 0.418. Leg I: trochanter: 0.137 / 0.085, femur: 0.131 / 0.115, patella: 0.106 / 0.076, tibia: 0.127 / 0.078, tarsus: 0.202 / 0.047. Leg IV: trochanter: 0.153 / 0.121, femur+patella: 0.380 / 0.121, tibia: 0.164 / 0.086, tarsus: 0.276 / 0.058. Distribution. Brazil (Harvey 2013; World Pseudoscorpiones Catalog 2022)., Published as part of Bedoya-Roqueme, Edwin, Tizo-Pedroso, Everton, Barbier, Eder & Lira, André Felipe De Araújo, 2023, Two new cave-dwelling pseudoscorpion species (Arachnida: Pseudoscorpiones) from Northeastern Brazil, pp. 317-332 in Zootaxa 5293 (2) on page 321, DOI: 10.11646/zootaxa.5293.2.6, http://zenodo.org/record/7960232, {"references":["Heurtault, J. (1986) Petterchernes brasiliensis, genre et espece nouveaux de Pseudoscorpions du Bresil (Arachnides, Pseudoscorpionida, Chernetidae). Bulletin du Museum National d'Histoire Naturelle, Paris, 8, 351 - 355.","Mahnert, V. (1994) New chernetid pseuuoscorpions (Pseudoscorpionida: Chemetidae) from Venezuela and Brazil, with remarks on the genus Ancalochernes Beier. Revue suisse de Zoologie, 101, 829 - 838. https: // doi. org / 10.5962 / bhl. part. 79929","Harvey, M. S. (2013) Pseudoscorpions of the World. Version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 20 February 2023)","World Pseudoscorpiones Catalog (2022) World Pseudoscorpiones Catalog. Natural History Museum Bern. Available from: http: // wac. nmbe. ch (accessed 20 February 2022)"]}

In a study of pseudoscorpions in caves from the northeastern Brazil, eight species of Pseudoscorpiones were identified. The known distribution of the species Pseudochthonius biseriatus Mahnert, 2001, Cryptoditha aff. elegans, Parachernes (P.) nitidimanus (Ellingsen, 1905), Petterchernes brasiliensis Heurtault, 1986, Ceriochernes foliaceosetosus Beier, 1974, and Progarypus setifer Mahnert, 2001 is extended to include the State of Pernambuco. Additionally, Geogarypus gollumi sp. nov. (Geogarypidae) and Progarypus smaugi sp. nov. (Hesperolpiidae) are described as new species.

Rhopalochernes luiscarlosi sp. nov. Figs. 1A–F, 2A–G, 3A–G, 7A ZooBank Registration: urn:lsid:zoobank.org:pub: 82D786A8-5B1A-4CF5-8194-64EF3B303633 Type material. female holotype: COLOMBIA, Bolívar, Municipio de San Jacinto, Vereda La Flecha, Finca Amanecer Gaitero (9°51′09.74″N; 75°10′32.32″W), 250 masl. 10.XI.2019, K. Marimon & E. Villarreal leg. (ICN.APs841). Paratypes: 1 male and 2 females, with same data as for holotype. (IAvH-I-3866 [1♀]; (IAvH-I-3876 [1♀]; ICN-APs-842 [1♂]), Etymology. The specific epithet is a patronym in honor of the late professor Luis Carlos Gutierrez Moreno, who was a lead researcher in the Caribbean region of Colombia, as well as founder of the research group Biodiversidad del Caribe Colombiano at Universidad del Atlántico. Diagnosis. Rhopalochernes luiscarlosi sp. nov. is characterized by the following combination of characters: cheliceral hand with four setae (Figs. 2F, 3F); serrula exterior with 11 blades; eyeless (Figs. 1A, 1C, 1D, 1F); carapace with two deep transverse furrows (Figs. 1C, 1F); nodus ramosus proximal to st (Figs. 2E, 3E); and trichobothria st closer to t than sb (Figs. 2E, 3E). Differential diagnosis: Rhopalochernes luiscarlosi sp. nov. can be differentiated from R. chamberlini, R. panamensis and R. ohausi by the lack of eyespots. It also differs from R. chamberlini by the position of the nodus ramosus which is midway between sb and st (Figs. 2E, 3E), but at the same level as st in R. chamberlini. It can be separated from R. panamensis by the serrula exterior which has 11‒12 blades in R. luiscarlosi sp. nov. and 14‒15 blades in R. panamensis. From the type species, R. ohausi, R. luiscarlosi sp. nov. is distinguished by having gaping chelal fingers (not gaping chelal fingers in R. ohausi) and the position of nodus ramosus which is proximal to or at same level of st in R. ohausi (see Mahnert, 2001 Fig. 110). Rhopalochernes luiscarlosi sp. nov. is similar to R. insulanus and R. catalinae sp. nov. by the absence of eyes, but can be differentiated from R. insulanus by the presence of 4 setae on the cheliceral hand (6 in R. insulanus), a trait that is also present in R. chamberlini, R. panamensis and R. catalinae sp. nov. (see Heurtault, 1998, Figs. 4, 9). It can be distinguished from R. catalinae sp. nov. by the furrows on the surface of the carapace, which are deep and conspicuous in R. luiscarlosi sp. nov. (Figs. 1C, 1F), but shallow and inconspicuous in R. catalinae sp. nov. (Figs. 4C, 4F). Additionally, they differ by the shape of the spermathecae, which have elongated and narrow ducts in R. luiscarlosi sp. nov. (Fig. 7A) but have short, thick ducts in R. catalinae sp. nov. (Fig. 7B). Finally, R. luiscarlosi sp. nov. has a slender tarsus IV (4.00–4.25 × longer than deep) while that of R. catalinae sp. nov. is more robust (2.75–3.75 × longer than deep). Description of adult: Colour: Carapace and pedipalps reddish brown. Tergites and legs yellowish brown (Fig. 1A–F). Carapace: heavily granulated; 1.15‒1.23 (♂), 1.09‒1.27 (♀) × longer than broad, eyes or eye-spots absent; with 2 deep transverse furrows, posterior furrow closer to posterior margin than anterior furrow; posterior margin of carapace straight; anterior margin with 4 setae (♂, ♀), posterior margin with 8 (♂), 10 (♀) setae (Fig. 3C, F). Chelicera: 1.67‒1.80 (♂), 1.29‒1.83 (♀) × longer than broad; with 4 setae on hand (sbs absent) and 1 distal seta (gs) on movable finger, is, bs and es short and acuminate, ls longer and thicker (Figs. 2F, 3F); galea with 4 rami (♀) or simple (♂) (Figs. 2D, 3G); rallum with 3 blades (Figs. 2G, 3D), anterior blade with several ramifications while others smooth; serrula exterior with 11‒12 blades; lamina exterior present. Pedipalp: trochanter, femur, patella and chelal hand heavily granulated, with short, feathered and acuminate setae (Figs. 2C, 3C); chelal fingers smooth; trochanter 1.50‒1.62 (♂), 1.40‒1.71 (♀), femur 2.13‒2.19 (♂), 2.31‒2.67 (♀), patella 2.00‒2.20 (♂), 1.88‒2.20 (♀), chela (with pedicel) 2.60‒2.74 (♂), 2.80‒3.95 (♀), chela (without pedicel) 2.40‒2.53 (♂), 2.45‒2.63 (♀), hand 1.10‒1.21 (♂), 1.20‒1.26 (♀) × longer than broad, movable finger 1.09‒ 1.18 (♂), 1.04‒1.27 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, accessory teeth not visible. Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria: eb and esb placed basally, est medially at equal distance from et and eb, and et distally near the tip of the fixed finger; ib at the same level as esb and close to ist, which is placed midway in the first half of the fixed finger, isb slightly more distal than est and it closer to et than isb; b, sb, st and t about an equal distance from each other, sb slightly ahead of esb, st in the middle of movable finger slightly closer to t than sb, t placed distally and closer to st than tip of the finger. Venom apparatus present only in movable chelal finger with nodus ramosus proximal to st (Figs. 2E, 3E). Coxal region: maxilla smooth except for the antero-lateral region; coxae smooth; manducatory process triangular in shape with 2 apical setae, 1 prolateral suboral seta and 12 (♂), 9 (♀) additional setae on maxilla. Median maxillary lyrifissure rounded and situated submedially. Chaetotaxy of coxae I‒IV, ♂, 9: 9: 8: 15; ♀, 10: 13: 11: 20. Legs: granulate; feathered setae on retrolateral margin and dentate setae on prolateral margins. Femur + patella of leg IV 2.45 ‒ 2.60 (♂), 2.73‒2.80 (♀) × longer than deep; tibia 3.14‒3.50 (♂), 3.14‒3.83 (♀) × longer than deep; tarsus 4.00‒4.25 (♂), 4.00‒4.50 (♀) × longer than deep. Tarsi III and IV with short tactile seta, located subdistally. Arolium undivided, slightly shorter than claws; claws slender and simple, not modified (Figs. 2A‒B, 3A‒B). Abdomen: tergites I‒X and sternites IV ‒X with a clear medial suture line (Figs. 7C). Tergal chaetotaxy: ♂, 10: 12: 12: 14: 14: 14: 15: 14: 14: 12: 9 (2 T): 2; ♀, 14: 12: 11: 15: 17: 16: 15: 14: 12: 13: 10 (2 T): 2; Sternal chaetotaxy: ♂, 17:(1) 8 (1): (1) 10 (1): 13: 14: 14: 12: 11: 9: 9: 2; ♀, 16: (1) 10 (1): (1) 9 (1): 14: 14: 15: 12: 10: 9: 4: 2. Genitalia: female: spermathecae paired, with long ducts ending in oval sacs (Fig. 7A). Male: genital opercula containing 9 setae on anterior operculum (sternite II), 8 setae clustered together just anterior to the aperture, 4 in posterior margin of aperture, and 4 in distal margin of posterior operculum (sternite III). Dimensions: holotype female (ICN APs-841), with range of 4 female paratype in parentheses: Body length 1.36 (1.12‒1.36). Pedipalps: trochanter 0.22/0.15 (0.21‒0.24/0.14‒0.15), femur 0.35/0.15 (0.33‒0.38/0.14‒0.16), patella 0.31/0.15 (0.31‒0.34/0.15‒0.17), chela (with pedicel) 0.56/0.2 (0.53‒0.56/0.18‒0.20), chela (without pedicel) 0.49 (0.49‒0.50), hand (without pedicel) length 0.24 (0.22‒0.24), movable finger length 0.27 (0.26‒0.28). Carapace 0.5/0.46 (0.49‒0.52/0.41‒0.46). Leg IV: femur + patella 0.28/0.10 (0.28‒0.30/0.22‒0.23), tibia 0.22/0.07 (0.22‒0.23/0.06‒0.07), tarsus 0.17/0.04 (0.16‒0.18/0.04). Allotype male (ICN APs-842), with other male in parentheses: Body length 1.18 (1.03). Pedipalps: trochanter 0.21/0.13 (0.21/0.14), femur 0.34/0.16 (0.35/0.16), patella 0.32/0.16 (0.33/0.15), chela (with pedicel) 0.52/0.19 (0.52/0.20), chela (without pedicel) 0.48 (0.48), hand (without pedicel) length 0.23 (0.22), movable finger length 0.25 (0.26). Carapace 0.47/0.41 (0.48‒0.39). Leg IV: femur + patella 0.27/0.11 (0.26/0.10), tibia 0.22/0.07 (0.21/0.06), tarsus 0.17/0.04 (0.16/0.04).

Rhopalochernes catalinae sp. nov. Figs. 4A–F, 5A–G, 6A–G, 7B ZooBank Registration: urn:lsid:zoobank.org:pub: 33CE2DF6-A490-4524-89DF-D6C5D5A884A7 Type material. female holotype: COLOMBIA, Atlántico, Municipio de Usiacurí, Reserva Campesina La Montaña (10°46’02,06” N, 75°02’34” W), 150m, 08.VI.2019, K. Marimon and E. Villarreal leg. (ICN-APs-843). Paratypes: 1 female and 3 males with the same data as the holotype (IAvH-I-3856 [2♂]; ICN-Aps-844 [1♂, 1♀]); 2 females and 2 males, Colombia, Atlántico, Municipio de Juan de Acosta, Corregimiento de Chorrera (10°47′48.34″ N, 75°00′45.37″ W), 270 masl. 09. VI.2018, K. Marimon, D. Tilano and E. Villarreal leg (IAvH-I-3846 [1♀]; ICN-Aps- 845 [1♀, 2♂]); 1 female, Colombia, Bolívar, Municipio de San Jacinto, Vereda La Flecha, Finca Amanecer Gaitero (9°51′9.74″ N; 75°10′32.32″ W), 250 masl. 11-12.IX.2019, K. Marimon & E. Villarreal leg. (ICN-APs-846). Etymology. This species is named for our colleague Catalina Romero-Ortiz, in appreciation of her research on Colombian pseudoscorpions. Diagnosis: Rhopalochernes catalinae sp. nov. can be distinguished from other species of the genus by the following combination of characters: cheliceral hand with four setae (Figs. 5E, 6E); serrula exterior with 11-12 blades; eyeless (Figs. 4C, 4F); carapace with shallow transverse furrows (only visible in lateral view); nodus ramosus at the same level or slightly close to st (Figs. 5G, 6G); and trichobothria st midway between t and sb (Figs. 5G, 6G). Differential diagnosis: Rhopalochernes catalinae sp. nov. can be differentiated from R. chamberlini, R. panamensis and R. ohausi by the lack of eyespots (Figs. 4C, 4F). It also differs from R. chamberlini by the thicker femur + patella of leg IV, 2.60‒2.89 versus 3.73 × longer than deep in R. chamberlini. It can be distinguished from R. panamensis and R. ohausi by the serrula exterior which has 11‒12 blades, but 14‒15 blades in R. panamensis and 16 blades in R. ohausi. It can also be distinguished from R. ohausi by having gaping chelal fingers (versus not gaping in R. ohausi, see Mahnert, 2001, Fig. 110). Rhopalochernes catalinae sp. nov. most closely resembles R. insulanus and R. luiscarlosi sp. nov. in the absence of eyes (Figs. 4C, 4F), but can be differentiated from R. insulanus by the number of setae on the cheliceral hand, 6 in R. insulanus versus 4 in R. catalinae sp. nov. and R. luiscarlosi sp. nov., a trait also found in R. chamberlini and R. panamensis (see Heurtault, 1998, Figs. 4 and 9). Rhopalochernes catalinae sp. nov. can also be distinguished from R. luiscarlosi sp. nov. by the shape of the furrows on the surface of the carapace, which are shallow and inconspicuous in R. catalinae sp. nov. (Figs. 4C, 4F), but are deep and conspicuous in R. luiscarlosi sp. nov. (Figs. 1C, 1F). They also differ in other features such as: the shape of the spermathecae, with short and thicker ducts (Fig. 7B) versus elongated and narrow ducts in R. luiscarlosi sp. nov. (Fig. 7A); the shape of tarsus IV, which is robust (2.75–3.75 × longer than deep) while that of R. luiscarlosi sp. nov. is slender (4.00–4.25 × longer than deep). Description of adult: Colour: Carapace reddish brown towards the anterior margin and lighter towards posterior margin, pedipalps reddish brown. Tergites and legs yellowish brown (Fig. 4). Carapace: heavily granulated; 1.24‒1.41 (♂) 1.24‒1.37 (♀) × longer than broad, eyes or eye‒spot absent; with 2 inconspicuous transverse furrows slightly visible on lateral view, posterior furrow closer to posterior margin than anterior furrow; posterior margin of carapace slightly v shaped; anterior margin with 6 setae (♂, ♀), posterior margin with 8‒9 setae (♂, ♀). (Fig. 4C, F) Chelicera: 1.43‒2.00 (♂), 1.33‒1.80 (♀) × longer than broad; hand with 4 setae (sbs absent) and 1 distal seta (gs) on movable finger, is, bs and es short and acuminate, ls longer and broad (Figs. 5E, 6E), galea with 4 rami (♀) or simple (♂) (Figs. 5C, 6D); rallum with 3 blades (Figs. 5D, 6C), anterior blade with several ramifications while others smooth; serrula exterior with 11‒12 blades (♂, ♀), lamina exterior present. Pedipalp: trochanter, femur, patella and hand heavily granulated, with short, feathered and acuminate setae (Figs. 5F, 6F), chelal fingers smooth; trochanter 1.20‒1.57 (♂), 1.20‒1.64 (♀), femur 2.07‒2.77 (♂), 2.43‒3.08 (♀), patella 1.62‒2.36 (♂), 1.93‒2.43 (♀), chela (with pedicel) 2.94‒3.27 (♂), 2.89‒3.24 (♀), chela (without pedicel) 2.40‒2.53 (♂), 2.45‒2.63 (♀). Hand 1.24‒1.40 (♂), 1.25‒1.41 (♀) × longer than broad, movable finger 1.10‒1.24 (♂), 1.09‒1.27 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, accessory teeth not visible. Fixed chelal finger with 8 trichobothria. movable chelal finger with 4 trichobothria: eb, esb and est in the first half of fixed finger, esb closer to eb than est, which is closer to the midpoint of the finger, et distal and close to the fingertip; ib at the same level of eb, ist slightly aligned to esb, isb at the same level as st, close to est, it closer to et than isb; b and sb grouped basally, st and t in the middle of the movable finger, st slightly closer to t than sb, t much closer to st than tip of the finger. Venom apparatus present only in movable chelal finger with nodus ramosus at the same level or slightly close to st (Figs. 5G, 6G). Coxal region: maxillae smooth except for the antero-lateral region, coxae smooth, manducatory process triangular in shape with 2-3 apical setae, 1 prolateral suboral seta and 14-15 (♂), 11 (♀) additional setae on maxilla. Median maxillary lyrifissure rounded and situated submedially. Chaetotaxy of coxae I-IV, ♂, 7-8: 8-10: 8: 12; ♀, 9: 9: 12: 24. Legs: granulate, feathered setae on retrolateral margin and dentate setae on prolateral margins. Femur + patella of leg IV 2.25‒2.70 (♂), 2.60‒2.89 (♀) × longer than depth, as well as tibia and tarsus 2.71‒3.33 (♂), 3.17‒3.67 (♀) and 2.75‒3.75 (♂), 3.25‒3.75 (♀) × longer than broad, respectively. Tarsi III and IV with tactile seta, located subdistally. Arolium undivided, slightly shorter than claws, claws slender and simple, not modified (Figs. 5A‒B, 6A‒B). Abdomen: tergites I-X and sternites IV-X with a clear medial suture line (Fig. 7D). Tergal chaetotaxy: ♂, 12: 11-12: 11-13:15-16: 14-15: 15-16: 15-16: 14-16: 13-14: 12-13: 9-10: 2, ♀, 12: 14: 13: 16: 17: 18: 19: 16: 16: 13: 9: 2. Sternal chaetotaxy: ♂, (1) 4-6 (1): (1) 7-9 (1): 13-14: 12-13: 13: 13: 12: 10: 4-6: 2, ♀, 20: (1) 5 (1): (1) 15 (1): 15: 15: 14: 14: 14: 9: 16: 2. Tergites with setae acuminate, sternites with setae leaf-like shape. Pleural membrane striated and without setae. Genitalia: female: spermathecae paired, with short ducts ending in oval sacs (Fig. 7A). Male: genital opercula containing 18 setae on the anterior operculum and 4 in distal margin of posterior operculum. Dimensions: holotype female (ICN APs-843), with a range of 5 females paratype in parentheses: Body length 1.4 (1.04‒1.50). Pedipalps: trochanter 0.21/0.14 (0.21‒0.23/0.13‒0.15), femur 0.33/0.13 (0.33‒0.37/0.12‒0.14), patella 0.27/0.14 (0.27‒0.34/0.14‒0.15), chela (with pedicel) 0.51/0.16 (0.51‒0.55/0.16‒0.18), chela (without pedicel) 0.47 (0.46‒0.50), hand (without pedicel) length 0.2 (0.20‒0.24), movable finger length 0.25 (0.25‒0.28). Carapace 0.45/0.36 (0.45‒0.49/0.36‒0.39). Leg IV: femur + patella 0.25/0.09 (0.25‒0.27/0.09‒0.1), tibia 0.19/0.06 (0.19‒ 0.22/0.06), tarsus 0.14/0.04 (0.13‒0.15/0.04). Paratype male (ICN APs-844), with a range of 7 male in parentheses: Body length 1.34 (1.11‒1.35). Pedipalps: trochanter 0.21/0.14 (0.18‒0.22/0.14‒0.15), femur 0.36/0.13 (0.29‒0.38/0.13‒0.16), patella 0.33/0.15 (0.21‒ 0.34/0.13‒0.16), chela (with pedicel) 0.51/0.17 (0.48‒0.55/0.15‒0.18), chela (without pedicel) 0.46 (0.43‒0.49), hand (without pedicel) length 0.21 (0.21‒0.23), movable finger length 0.26 (0.23‒0.27). Carapace 0.47/0.35 (0.46‒ 0.48/0.34‒0.37). Leg IV: femur + patella 0.26/0.11 (0.25‒0.27/0.10‒0.12), tibia 0.19/0.07 (0.17‒0.22/0.06‒0.07), tarsus 0.14/0.04 (0.11‒0.15/0.04)., Published as part of Marimon, Karla, Blanco, Eduardo Villarreal & Harvey, Rk. S., 2022, Two new species of Rhopalochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) from Colombia, pp. 397-410 in Zootaxa 5150 (3) on pages 403-407, DOI: 10.11646/zootaxa.5150.3.5, http://zenodo.org/record/6623085, {"references":["Heurtault, J. (1998) Pseudoscorpions of the genus Rhopalochernes (Chernetidae) from Panama and Venezuela. Journal of Arachnology, 26, 442 - 446."]}

Marimon, Karla, Blanco, Eduardo Villarreal, Harvey, Rk. S. (2022): Two new species of Rhopalochernes Beier, 1932 (Pseudoscorpiones: Chernetidae) from Colombia. Zootaxa 5150 (3): 397-410, DOI: https://doi.org/10.11646/zootaxa.5150.3.5

Genus Nudochernes Beier, 1935 Nudochernes Beier, 1935: 122; Beier, 1959: 48; Harvey, 1991: 607. Type species: Nudochernes montanus Beier, 1935, by original designation. Key to the Chinese species of Nudochernes 1. Pedipalpal femur 3.20 times longer than broad............................................ N. lipsae Mahnert, 2003 - Pedipalpal femur 2.40���2.90 times longer than broad.......................................................... 2 2. Trichobothria st closer to t than to sb; pedipalpal chela (with pedicel) 3.40���3.80 times longer than broad............................................................................................... N. troglobius Mahnert, 2009 - Trichobothria s t situated midway between sb and t; pedipalp chela (with pedicel) 3.10 (♂) 2.62���2.68 (♀) times longer than broad.......................................................................... N. pseudptroglobius sp. nov., Published as part of Xu, Hongru, Gao, Zhizhong & Zhang, Feng, 2022, Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 from Guizhou, China (Pseudoscorpiones: Chernetidae), pp. 581-592 in Zootaxa 5105 (4) on pages 582-587, DOI: 10.11646/zootaxa.5105.4.7, http://zenodo.org/record/6333898, {"references":["Mahnert, V. (2003) Four new species of pseudoscorpions (Arachnida, Pseudoscorpiones: Neobisiidae, Chernetidae) from caves in Yunnan Province, China. Revue suisse de Zoologie, 110 (4), 739 - 748. https: // doi. org / 10.5962 / bhl. part. 80209","Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492"]}

Nudochernes pseudotroglobius sp. nov. (Figs 1–4) http://zoobank.org/NomenclaturalActs/ 20C7BD03-E125-4379-8D91-A3111A273DDF Type material. Holotype ♀ (Ps.- MHBU-GZXS202401): China, Guizhou Province, Xishui County, Daozuo Cave [28.511389°N, 107.153333°E], elev. 1601m a.s.l., 31 August 2020, Zegang Feng & Yanmeng Hou leg. Paratypes: 1♀ (Ps.- SWUC-GZXS202402), 1♂ (Ps.- MHBU-GZXS202403), same data as holotype. Etymology. The name refers to its similar habitat with Nudochernes troglobius Mahnert, 2009. Diagnosis. This species can be distinguished from other Nudochernes species by the presence of dentate setae on carapace and pedipalp, spermathecae with a long unpaired tube and two short apical tubules, st situated midway between sb and t, the presence of elongate setae on the posterior corners of coxae IV, and the tarsus IV with a tactile seta in middle of tarsus (TS=0.55 (♂) 0.57 (♀)). Description. (Fig. 2). Males smaller than females with 2.77 (♂) 3.11–3.16 (♀), carapace and pedipalps reddish brown, chelicera, sternites, tergites and legs yellowish brown. Carapace (Fig. 3A): evenly granulate; 1.12 (♂) 0.97–0.98 (♀) times longer than broad; eyes or eye-spots absent; with two distinct deep transverse furrows; subbasal transverse furrow situated closer to the posterior margin than to the median furrow; posterior area with a small longitudinal furrow; including 16 (♂) 10 (♀) setae on anterior margin and 15 (♂) 16 (♀) setae on posterior margin. Abdomen: all tergites and sternites narrowly divided (except the last one). Tergal chaetotaxy (I–XI): ♂, 7–7: 10–8: 8–7: 9–9: 8–8: 9–8: 9–9: 9–7: 7–7: 6–9: 11(2T); ♀, 7–7: 9–10: 9–8: 11–10: 10–10: 9–10: 9–10: 8–9: 7–8: 7–7: 10(2T); all setae clavate. Sternal chaetotaxy (IV–XI): ♂, 7–9: 11–11: 11–10: 9–10: 9–8: 8–8: 8–7: 10; ♀, 4–4: 6–6: 12–13: 13–12: 12–13: 13–13: 9–8: 11; all setae acuminate. Sternite II of ♀ with 18 setae arranged as an inverted “U” pattern (Figs 3E, 4E). Chelicerae (Figs 3D, 4D): 5 setae on hand and 1 subdistal seta on movable finger; sbs, bs, es dentate apically, ls, is, gs acuminate; fixed finger with 6 proximad-directed teeth, movable finger with broad tooth-like subapical lobe; galea with 6 long branchlets in distal part; rallum with 3 dentate blades (Fig. 4C); serrula exterior with 20 (♂, ♀) blades. Pedipalp (Figs 3B, 4A): pedipalp robust; trochanter, femur, patella and chelal hand coarsely granulate, with dentate setae; chelal fingers with short setae; trochanter 2.24 (♂) 1.50–1.62 (♀), femur 2.24 (♂) 2.33–2.61 (♀), patella 2.30 (♂) 2.12–2.38 (♀), chela (without pedicel) 2.71 (♂) 2.33–2.47 (♀), hand (without pedicel) 1.29 (♂) 1.21–1.26 (♀) times longer than broad, movable finger 1.27 (♂) 1.12–1.28 (♀) times as long as hand (without pedicel). Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria (Figs 3C, 4F): eb situated basally; est situated midway between esb and et; ib and esb situated sub-basally, it situated closer to tip of finger than to ib; s t situated midway between sb and t, sb situated closer to b than to s t, and t situated midway between tip of finger and st; fixed finger with 49 teeth, plus 5 prolateral accessory teeth and 5 retrolateral accessory teeth; movable finger with 47 teeth, plus 6 prolateral accessory teeth and 5 retrolateral accessory teeth. Venom apparatus only present in movable chelal finger, venom duct slender, nodus ramosus closer to t than to st. Coxae: with numerous short setae over entire ventral surface, with longer setae on posterior margin of coxae IV. Genitalia (Fig. 4B): ♂: typical of Chernetidae; ♀: spermathecae with a long unpaired tube and two short apical tubules. Legs I (Figs 3F, 4G): trochanter, femur, patella, tibia with numerous clavate setae; tarsus with numerous acuminate setae; proportions: trochanter 1.09 (♂) 1.20–1.31 (♀), femur 1.46 (♂) 1.51–1.84 (♀), patella 2.07 (♂) 2.09– 2.41 (♀), tibia 3.86 (♂) 3.33–3.70 (♀), tarsus 3.98 (♂) 4.30–5.01 (♀) times longer than wide; claws simple, arolium shorter than claws. Legs IV (Figs 3H, 4H): femur, patella, tibia with numerous clavate setae; trochanter and tarsus with numerous acuminate setae; proportions: trochanter 1.69 (♂) 1.65–1.77 (♀), femur + patella 5.50 (♂) 5.46–5.50 (♀), tibia 4.24 (♂) 4.58–5.10 (♀), tarsus 4.32 (♂) 4.20–4.87 (♀) times longer than deep, a tactile seta near middle of tarsus (Figs 3I, 4I) (TS=0.55 (♂) 0.57 (♀)); claws simple, arolium shorter than claws. Dimensions (length/breadth or depth, in mm). ♀: Total length 3.11–3.16. Carapace 0.70–0.76/0.71–0.78. Pedipalps: trochanter 0.41–0.42/0.25–0.31, femur 0.64–0.68/0.25–0.27, patella 0.64–0.68/0.25–0.29, chela (with pedicel) 1.14–1.31/0.36–0.42, chela (without pedicel) length 1.02–1.18, hand (without pedicel) length 0.48–0.53, movable finger length 0.54–0.68. Leg I: trochanter 0.17–0.18/0.13–0.15, femur 0.21–0.24/0.13–0.14, patella 0.23– 0.29/0.11–0.12, tibia 0.30–0.37/0.09–0.10, tarsus 0.30–0.35/0.07. Leg IV: trochanter 0.28–0.32/0.17–0.18, femur + patella 0.71/0.13–0.15, tibia 0.51–0.55/0.10–0.12, tarsus 0.39–0.42/0.08–0.10, TS=0.57. ♂: Total length 2.77. Carapace 0.72/0.64. Pedipalps: trochanter 0.40/0.22, femur 0.63/0.28, patella 0.65/0.28, chela (with pedicel) 1.15/0.37, chela (without pedicel) length 1.01, hand (without pedicel) length 0.48, movable finger length 0.61. Leg I: trochanter 0.15/0.14, femur 0.21/0.14, patella 0.25/0.12, tibia 0.33/0.09, tarsus 0.30/0.08. Leg IV: trochanter 0.26/0.15, femur + patella 0.59/0.11, tibia 0.42/0.10, tarsus 0.38/0.09, TS=0.55. Remarks. This new species is close related to N. troglobius Mahnert, 2009 (Hubei, China), but can be distinguished by the stouter pedipalp (chela (with pedicel) 3.10 (♂) 2.62–2.68 (♀) times longer than board versus 3.40–3.80 (♂, ♀) in N. troglobius) and the position of trichobothrium (s t situated midway between sb and t in N. pseudotroglobius sp. nov.; st closer to t than to sb in N. troglobius) (Mahnert 2009). This new species can be distinguished from N. lipsae Mahnert, 2003 (Yunnan, China) by the stouter legs I (patella 2.07 (♂) 2.09–2.41 (♀) times longer than deep vs 3.50 (♀) in N. lipsae; tibia 3.86 (♂) 3.33–3.70 (♀) times longer than deep vs 5.00 (♀) in N. lipsae), and by the position of trichobothrium (s t situated midway between sb and t in N. pseudotroglobius sp. nov., whereas s t closer to t than to sb in N. lipsae) (Mahnert 2003). Geographically, only one species, N. spalacis Beier, 1955, is described in Asia (Israel). The new species can be distinguished from N. spalacis by having slender legs IV (femur + patella 5.50 (♂) 5.46–5.50 (♀) times longer than deep versus 3.70 (♂) in N. spalacis) and the different position of trichobothrium (est situated midway between et and esb in N. pseudotroglobius sp. nov., whereas est closer to et than to esb in N. spalacis) (Beier 1955). Distribution. Known only from the type locality in Guizhou, China. Habitat. All specimens were collected under a rock within the dark zone of the cave.

Megachernes biyunensis sp. nov. (Figs 5–7) http://zoobank.org/NomenclaturalActs/31be2f78-81ae-4bb7-aa18-076eb4722f89 Type material. Holotype ♀ (Ps.- MHBU-GZ19080501): China, Guizhou Province, Pan County, Biyun Park [25.775000°N, 104.638333°E], 5 August 2019, Zegang Feng & Zhaoyi Li leg. Paratypes: 1♀ (Ps.-SWUC-GZ19080502); 1♂ (Ps.- MHBU-GZ19080503), same data as holotype. Etymology. The specific name refers to the type locality, Biyun Park, Pan County, Guizhou Province, China. Diagnosis. This new species can be recognized by the following combined characters: subbasal transverse furrow on carapace located midway between posterior margin and central furrow, the trichobothria st situated closer to sb than to t, t situated midway between tip of finger and st. Description. (Fig. 5). Carapace, chelicera, tergites and sternites yellowish brown, pedipalps reddish brown, colour of chelal hand darker than that of trochanter, femur and patella; soft parts pale. Carapace (Fig. 6A): evenly granulate; 1.07 (♂) 1.02 (♀) times longer than broad; eyes or eye-spots absent; with two distinct deep transverse furrows, subbasal transverse furrow situated midway between posterior margin and central furrow; posterior area with a small longitudinal furrow; with numerous setae including 16 (♂) 20 (♀) acuminate setae on anterior margin and 10 (♂) 14 (♀) on posterior margin. Abdomen: all tergites and sternites narrowly divided. Tergal chaetotaxy (I–XI): ♂, 7–6: 10–8: 7–8: 7–8: 7–8: 9– 8: 9–9: 7–9: 8–6: 7–8: 6(2 T)–7(2T); ♀, 7–7: 9–8: 8–8: 9–9: 9–8: 8–9: 8–9: 8–9: 9–7: 7–8: 7(2 T)–7(2T); I–X tergites with paired dark spot. Sternal chaetotaxy (IV–XI): ♂, 4–5: 8–9: 13–11: 13–12: 10–11: 11–19: 11–9: 6(2 T)–7(2T); ♀, 4–3: 10–12: 14–12: 14–14: 13–14: 12–14: 12–12: 6(2 T)–6(2 T); all sternal setae acuminate. Sternite II of ♀ with 33 setae arranged in a cluster (Figs. 6E, 7F). Chelicerae (Figs. 6D, 7D): 7 setae on hand and 1 subdistal seta on movable finger; basal 5 setae finely dentate apically, ls, is and gs acuminate; fixed finger with 3 proximad-directed teeth; galea comprising 2 parts, first part with 5 branches and the second with 4 branches (Fig. 7B). Serrula exterior with 27 (♂, ♀) blades; rallum with 3 dentate blades, the apex of the first one slightly twisted and with anterior margin dentate, the other two blades apically dentate (Fig. 7C). Pedipalp (Figs 6B, 7A): trochanter, femur, patella and chelal hand coarsely granulate, with acuminate setae; chelal fingers with short setae; trochanter 1.39 (♂) 1.67 (♀), femur 2.51 (♂) 2.31 (♀), patella 2.26 (♂) 2.23 (♀), chela (without pedicel) 3.02 (♂) 2.81 (♀), hand (without pedicel) 1.58 (♂) 1.51 (♀) times longer than board, movable finger 0.91 (♂) 0.96 (♀) times as long as hand (without pedicel). Fixed finger with 8 trichobothria, movable finger with 4 trichobothria (Figs 6C, 7E): eb situated basally; est situated closer to esb than to et; ib and esb situated sub-basally; it situated closer to tip of finger than to ib; st situated closer to sb than to t, sb situated closer to s t than to b, and t situated midway between tip of finger and st, fixed finger with 68 teeth, plus 13 prolateral accessory teeth, and 8 retrolateral accessory teeth; movable finger with 59 teeth, plus 12 prolateral accessory teeth, and 7 retrolateral accessory teeth. Venom apparatus only present in movable chelal finger, venom duct slender, nodus ramosus closer to t than to st. Coxae: with numerous short setae over entire ventral surface and with long setae on posterior margin of coxae IV; posterior corner of coxae IV slightly protruding. Genitalia (Figs 6E, 7H): ♂: typical of Chernetidae; ♀: spermathecae T-shaped and with extremely elongated ends. Leg I (Figs 6F, 7G): with long acuminate setae; proportions: trochanter 1.11 (♂, ♀), femur 1.50 (♂) 1.51 (♀), patella 2.60 (♂) 2.17 (♀), tibia3.71 (♂) 3.50 (♀), tarsus 4.41 (♂) 4.10 (♀) times longer than deep; claws simple, arolium shorter than claws. Legs IV (Figs 6H, 7I): with long acuminate setae; proportions: trochanter 1.76 (♂) 1.44 (♀), femur + patella 3.96 (♂) 3.43 (♀), tibia 5.01 (♂) 4.71 (♀), tarsus 3.80 (♂) 3.83 (♀) times longer than deep; tactile seta in middle tarsus (TS=0.51 (♂) 0.54 (♀)), length of tactile seta 2/3 of tarsus (Fig. 7J); claws simple, arolium shorter than claws. Dimensions: (length/breadth or depth, in mm). ♀: Total length 4.22. Carapace 1.20/1.18. Pedipalps: trochanter 0.65/0.39, femur 0.99/0.43, patella 1.07/0.48, chela (with pedicel) 1.95, chela (without pedicel) 1.80, hand (without pedicel) length 0.97, movable finger length 0.94. Leg I: trochanter 0.21/0.18, femur 0.30/0.21, patella 0.39/0.18, tibia 0.49/0.14, tarsus 0.41/0.10. Leg IV: trochanter 0.36/0.25, femur + patella 0.79/0.23, tibia 0.80/0.17, tarsus 0.46/0.12, TS=0.54. ♂: Total length 4.10. Carapace 1.14/1.05. Pedipalps: trochanter 0.59/0.43, femur 1.05/0.41, patella 1.06/0.47, chela (with pedicel) 1.93/0.64, chela (without pedicel) length 1.87, hand (without pedicel) length 1.01, movable finger length 0.92. Leg I: trochanter 0.22/0.20, femur 0.36/0.24, patella 0.52/0.2, tibia 0.51/0.14, tarsus 0.48/0.11. Leg IV: trochanter 0.36/0.21, femur + patella 1.06/0.27, tibia 0.81/0.16, tarsus 0.57/0.15, TS=0.51. Remarks. Based on the key of Megachernes species (Mahrad & Reza 2017), this new species is similar to M. limatus Hoff & Parrack, 1958 from Oceania, but can be distinguished by the character of carapace (the subbasal transverse furrow situated closer to the posterior margin than to the median furrow in M. limatus, whereas the subbasal transverse furrow situated midway in M. biyunensis sp. nov.); and by the different trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. limatus) (Hoff & Parrack 1958). It can be distinguished from M. himalayensis (Ellingsen, 1914) (China (Fujian); India; Nepal) by the number of accessory teeth on the chelal fingers (movable finger with 12 accessory teeth and fixed finger with 13 accessory teeth in M. biyunensis sp. nov., whereas movable finger with 6 accessory teeth and fixed finger with 9 accessory teeth in M. himalayensis); and by the stouter legs IV (tibia 5.01 (♂) 4.71 (♀) times longer than deep versus 6.20 (♀) in M. himalayensis; tarsus 3.80 (♂) 3.83 (♀) times longer than deep vs 5.00 (♀) in M. himalayensis) (Ellingsen 1914). It can be distinguished from M. titanius Beier, 1951 (China (Sichuan, Guangxi); Southeast Asia) by the trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. titanius); and by the stouter legs IV (5.01 (♂) 4.71 (♀) times longer than wide versus 6.20 (♀) in M. titanius; tarsus 3.80 (♂) 3.83 (♀) times longer than deep versus 5.20 (♀) in M. titanius) (Beier 1951). It differs from M. tuberosus Mahnert, 2009 (Sichuan, China) by different the trichobothrial pattern (st situated closer to sb than to t in M. biyunensis sp. nov., whereas st is closer to t than to sb in M. tuberosus); and by the morphology of the rallum (all three blades denticulate in M. biyunensis sp. nov., blades terminally scarcely dentate and apex of first blade slightly twisted in M. tuberosus) (Mahnert 2009). It differs from M. vietnamensis Beier, 1967 (China (Sichuan, Hubei, Yunnan); Vietnam) by the number of accessory teeth on the chelal fingers (movable finger with 12 accessory teeth and fixed finger with 13 accessory teeth in M. biyunensis sp. nov., whereas fixed finger with 21 accessory teeth in M. vietnamensis); and by the cheliceral hand setae (basal 5 setae with terminally dentateand acuminate in M. biyunensis sp. nov., whereas all setae with acuminate in M. vietnamensis) (Beier 1967). Distribution. Known only from the type locality, Guizhou, China. Habitat. All specimens were collected in wet leaf litter under Populus (Salicaceae).

Genus Megachernes Beier, 1932 Megachernes Beier, 1932: 128; Beier, 1933: 518; Beier, 1948: 476; Morikawa, 1960: 144; Harvey, 1991: 599. Type species: Chernes grandis Beier, 1930, by original designation. Key to the Chinese species of Megachernes 1. Leg IV femur + patella with numerous glandular microsetae........................... M. glandulosus Mahnert, 2009 - Leg IV femur + patella without glandular microsetae......................................................... 2 2. Male pedipalpal patella with hump................................................. M. tuberosus Mahnert, 2009 - Male pedipalpal patella without hump..................................................................... 3 3. Pedipalpal femur less than 4.00 times longer than board....................................................... 4 - Pedipalpal femur more than 4.00 times longer than board..................................... M. titanius Beier, 1951 4. Leg IV femur + patella more than 4.00 times longer than deep.................................................. 5 - Leg IV femur + patella less than 4.00 times longer than deep................................................... 6 5. Movable chelal finger with 9 retrolateral accessory teeth; all blades of rallum smooth..... M. himalayensis (Ellingsen, 1914) - Movable chelal finger with 21 retrolateral accessory teeth; only distal blade of rallum denticulate.................................................................................................... M. vietnamensis Beier, 1967 6. Only distal blade of rallum denticulate; st situated closer to t than to sb........................ M. grandis (Beier, 1930) - All blades of rallum denticulate; st situated closer to sb than to t................................ M. biyunensis sp. nov., Published as part of Xu, Hongru, Gao, Zhizhong & Zhang, Feng, 2022, Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 from Guizhou, China (Pseudoscorpiones: Chernetidae), pp. 581-592 in Zootaxa 5105 (4) on pages 587-591, DOI: 10.11646/zootaxa.5105.4.7, http://zenodo.org/record/6333898, {"references":["Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, i - xxi + 1 - 294. https: // doi. org / 10.1515 / 9783111385402","Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492","Beier, M. (1951) Die Pseudoscorpione Indochinas. Memoires du Museum National d'Histoire Naturelle, Paris, Nouvelle Serie, 147 - 123.","Beier, M. (1967) Pseudoscorpione vom kontinentalen Sudost-Asien. Pacific Insects, 9, 341 - 369."]}

Two new species of the pseudoscorpion subfamily Lamprochernetinae Beier, 1932 are described from Guizhou Province, China, Nudochernes pseudotroglobius sp. nov. and Megachernes biyunensis sp. nov., with detailed diagnoses, descriptions and illustrations.

Pachychernes zehorum Muchmore, 1997 Figs. 4A–D, 5A–G, 6A–G, 7B Pachychernes zehorum Muchmore, 1997: 19–21, figs 1–4; Ceballos, 2004: 429; Villegas-Guzman, 2006: 133; Harvey 2013. Diagnosis. This species differs from all other Pachychernes species by the first leg of the males, which not only has long and short modified setae on the tibia, but also a conspicuous depression on the dorsal side of the tarsus (Muchmore 1997) (Fig. 4). Examined material. COLOMBIA, Bolívar, Municipio de San Jacinto, La Flecha, Finca Amanecer Gaitero (9°51'9.74" N; 75°10'32.32" W), 250 m, 4 ♀ (ICN-Aps-835) and 2 ♂ (IAvH-I-4079), 11–12.IX.2019, K. Marimon and E. Villarreal leg. Description (adults): Colour: carapace red-brown with a lighter tone below the transversal furrow. Tergites light brown, pedipalps reddish-brown. Legs light yellow-brown toned. Carapace: heavily granulated, subrectangular with broad posterior margin; 0.78–0.93 (♂) 0.86–1.13 (♀) × longer than broad, with two eye spots; anterior margin with 8 setae (♂) 7 setae (♀), posterior margin with 6 setae (♂, ♀). Chelicera: 2.13–2.54 (♂), 1.44–2.40 (♀) × longer than broad; with 5 setae on hand and 1 distal seta on movable finger, es, is and ls acuminate, bs and sbs dentate; galea with 5 rami; rallum with 3 blades, posterior blade with several ramifications while others smooth; serrula exterior with 24 blades (♂, ♀); lamina exterior present (Figs. 5, 6). Pedipalp: with robust trochanter, femur, patella and granulated hand with several denticulated setae and chelal fingers smooth; trochanter 1.30–1.36 (♂), 1.27–1.39 (♀), femur 2.70–2.92 (♂), 2.10–2.42 (♀), patella 2.38–2.50 (♂), 2.12–2.37 (♀), chela (with pedicel) 2.81–3.05 (♂), 2.61–2.89 (♀), chela (without pedicel) 2.63–2.86 (♂), 2.61–2.89 (♀), hand 1.53–1.63 (♂), 1.54–1.68 (♀) × longer than broad, movable finger 0.76–0.77 (♂), 0.59–0.86 (♀) × longer than hand. All marginal teeth rounded except the terminal ones, which are pointed, external accessory teeth present on both fingers, with 6 (♂), 7 (♀) on the movable finger and 9 (♂), 11(♀) on the fixed finger. Fixed chelal finger with 8 trichobothria, movable chelal finger with 4 trichobothria: eb, esb, est and et slightly L-shaped, eb and esb situated basally, est located medially and at an equal distance from esb and et, which is located terminally; ib and ist basally located, with ist, eb and esb aligned, isb located in a medial position and closer to it and ib; b, sb and st placed in the first half of the movable finger with b and sb much closer to each other than to st, t located subterminally, midway from st and tip of finger. Venom apparatus present only in movable chelal finger with nodus ramosus located next to t (Figs. 5, 6). Coxal region: maxillae smooth except for the antero-lateral region, coxae smooth, manducatory process triangular in shape; regarding the setae, these individuals have one apical, one subapical and one sub oral while there are 25 additional setae (♂). Maxillary lyrifissure rounded and placed submedially. Chaetotaxy of coxae I–IV in ♂: 14: 12: 16: 44; ♀: 13: 11: 15: 36. Legs: leg I sexually dimorphic, with males having a tibia with a large number of setae plus a distinct depression on the surface of the tarsus while that of the females are chernetid in structure. Femur + patella of leg IV 2.67–2.77 (♂), 3.56–4.17 (♀) × longer than broad, as well as tibia and tarsus 3.79–4.11 (♂), 3.56–4.17 (♀) and 4.27–4.64 (♂), 3.89–4.67 (♀) × longer than broad, respectively. Tarsi III and IV with tactile seta, located subbasally, TS= 0.18 in both males and females approximately. Claws and arolium are equal in length. Abdomen: tergites I–X and sternites I–X with a clear medial suture line. Tergal chaetotaxy: ♂, 8: 7: 9: 8: 11: 13: 12: 12: 13: 12: 11: 2; ♀, 10: 10: 10: 10: 12: 12: 12: 13: 14: 13: 11: 2, all setae clavate. Sternal chaetotaxy: ♂, 47, (3) 16 (3): (1) 8 (1): 17: 19: 19: 23: 20: 17: 15: 2; ♀, 38: (2) 8 (2): (1) 9 (1): 20: 19: 18: 18: 18: 17: 16: 4. Spiracle with helix. Pleural membrane striated and without setae. Genitalia: spermatheca somewhat H-shaped, with four lobes, the anterior ones, oval in shape while the posterior ones have a tubular form with several cribriform plates along the ducts (Fig. 7B). Male genitalia consisting of 47 setae on the anterior operculum and 16 on the posterior operculum. Dimensions (mm): males, followed by females in parentheses: Body length 3.05–3.10 (2.44–3.30). Carapace 0.86–1.09/1.10–1.17 (0.81–1.05/0.82–1.22) Chelicera 0.32–0.33/0.13–0.15 (0.23–0.31/0.10–0.16), movable finger length 0.23–0.28 (0.20–0.25. Pedipalps: trochanter 0.48–0.49/0.36–0.37 (0.33–0.46/0.26–0.36), femur 1.10–1.14/0.37–0.39 (0.61–0.87/0.29–0.39), patella 1.00–1.15/0.42–0.46 (0.72–0.97/0.34–0.45), chela (with pedicel) 1.74/0.57–0.62 (1.39–1.89/0.46–0.62), chela (without pedicel) 1.63 (1.31–1.79), hand (without pedicel) length 0.93–0.95 (0.71–1.00), movable finger length 0.72 (0.57–0.85). Leg IV: femur 0.35–0.37/0.27–0.30 (0.26– 0.32/0.19–0.25), patella 0.77–0.8/0.35–0.36 (0.51–0.73/0.20–0.33), tibia 0.72–0.78/0.19 (0.50–0.72/0.12–0.18) tarsus 0.47–0.51/0.11 (0.35–0.49/0.09–0.11). Distribution and habitat. This species has been previously recorded from Mexico and Panama (Muchmore 1997; Ceballos 2004; Villegas-Guzman 2006), and according to Muchmore (1997) it is probably widespread through Central America. Here we expand the distribution to South America, with a population found in the Montes de María in northern Colombia. The new records demonstrate that this species is not necessarily restricted to environments such as rain forest (type locality) but also occurs in tropical dry forest (Figs. 8, 9)., Published as part of Marimon, Karla, Villarreal-Blanco, Eduardo, Romero-Ortiz, Catalina & Gutierrez, Luis C., 2021, A new species and new records of Pachychernes Beier, 1932 from Colombia (Pseudoscorpiones, Chernetidae), pp. 363-376 in Zootaxa 4999 (4) on pages 369-375, DOI: 10.11646/zootaxa.4999.4.6, http://zenodo.org/record/5119308, {"references":["Muchmore, W. (1997) An unusual new Pachychernes from Panama and Mexico (Pseudoscorpionida: Chernetidae). Entomological News, 108, 19 - 23.","Harvey, M. (2013) Pseudoscorpions of the world. Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues / pseudoscorpions (accessed 25 February 2021)"]}

In Colombia, the status of the Neotropical genus Pachychernes Beier, 1932 is unknown. The only available data in literature to support the presence of this genus is an adult of P. aff. subrobustus, possibly female, from copal resin without specific locality data, and a record at generic level from Northern Colombia. In this study, we extend the distribution of Pachychernes by describing a new species, P. florezi sp. n. and several reporting individuals of P. zehorum Muchmore, 1997 from Montes de María and the Caribbean coast of the country. These species are clearly differentiated from one another by the sexual dimorphism in the latter species, with a depression on the tarsus I in males. We also present an updated key for all species of the genus.

Americhernes incertus Mahnert, 1979 Americhernes incertus Mahnert, 1979: 772���775. Type locality: R��o Tarum-Mir��m, Manaus, Amazonas, Brazil (-2.963377; -60.195382). Distribution: Brazil and Colombia. Distribution in Colombia: C��rdoba: San Carlos (8.7457; -75.6795) (Bedoya-Roqueme 2019). Repository of Colombian material: LEUC-Ps-024 (1 ♀, 2 ♂) (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 121, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810. https: // doi. org / 10.5962 / bhl. part. 82338"]}

Americhernes bethaniae Mahnert, 1979 Americhernes bethaniae Mahnert, 1979: 769���772. Type locality: R��o Tarum-Mir��m, Manaus, Amazonas, Brazil (-2.963377; -60.195382). Distribution: Brazil and Colombia. Distribution in Colombia: Sucre: Sincelejo (9.3150; -75.3885) (Bedoya-Roqueme 2019) Repository of Colombian material: LEUC-Ps-025 (2 ♂) (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 121, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810. https: // doi. org / 10.5962 / bhl. part. 82338"]}

Dasychernes inquilinus Chamberlin, 1929 Dasychernes inquilinus Chamberlin 1929b: 51. Type locality: Rio Frio, Magdalena, Colombia (10.9181; -74.1665) Distribution: Colombia. Distribution in Colombia: Magdalena: Rio Fr��o (10.9181; -74.1665); Valle del Cauca: Sevilla (4.2743; -75.9310) (Chamberlin 1929b); Tolima: Cunday (4.0605; -74.6918) (Gonzalez et al. 2007). Repositories of Colombian material: holotype in CAS, MHNG (1♀, 1 ♂ and a tritonymph), MPUJ-ENT (2 ♀, 2 ♂, 2 tritonymphs, 4 deutonymphs and 3 protonymphs), (Gonzalez et al. 2007). Although Gonzalez et al. (2007) stated that some specimens are in ICN, they were never deposited there. Genus Epactiochernes Muchmore, 1974, Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on pages 124-125, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Chamberlin, J. C. (1929 b) Dasychernes inquilinus from the nest of meliponine bees in Colombia (Arachnida: Chelonethida). Entomological News, 40, 49 - 51. Available from: https: // www. biodiversitylibrary. org / item / 20333 page / 459 / mode / 1 up (Accessed 12 Dec. 2019)","Gonzalez, V. H., Mantilla, B. & Mahnert, V. (2007) A new host record for Dasychernes inquilinus (Arachnida, Pseudoscorpiones, Chernetidae), with an overview of pseudoscorpion-bee relationships. Journal of Arachnology, 35 (3), 470 - 474. https: // doi. org / 10.1636 / H 06 - 62.1","Muchmore, W. B. (1974) Pseudoscorpions from Florida. 3. Epactiochernes, a new genus based upon Chelanops tumidus Banks (Chernetidae). Florida Entomologist, 57, 397 - 407. https: // doi. org / 10.2307 / 3493504"]}

Parachernes (Parachernes) pallidus Beier, 1959 Parachernes pallidus Beier 1959: 211���212. Parachernes (Parachernes) pallidus Beier: Harvey 1991: 618. Type locality: without a precise location, ��� Colombia ���. Distribution in Colombia: without a precise location ( Beier 1959) Repository of Colombian material: NHMW, Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 127, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Beier, M. (1959) Zur Kenntnis der Pseudoscorpioniden-Fauna des Andengebietes. Beitrage zur Neotropischen Fauna, 1 (3), 185 - 228. https: // doi. org / 10.1080 / 01650525909380614","Harvey, M. S. (1991) Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester, 720 pp."]}

Incachernes brevipilosus (Ellingsen, 1910) Chelifer (Lamprochernes) brevipilosus Ellingsen 1910: 364–365. Neochernes brevipilosus (Ellingsen): Beier 1932d: 166. Incachernes brevipilosus (Ellingsen): Beier 1933: 95. Type locality: Central Colombia (without a precise location) (Ellingsen 1910). Distribution: known only from the type locality. Distribution in Colombia: without a precise location. Repository of Colombian material: Syntype in ZMB., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 125, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Ellingsen, E. (1910) Die Pseudoskorpione des Berliner Museums. Mitteilung aus dem Zoologischen Museum in Berlin, 4, 357 - 423.","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114","Beier, M. (1933) Pseudoskorpione aus Mexiko. Zoologischer Anzeiger, 104, 91 - 101."]}

Parachernes (Parachernes) adelaidae Bedoya-Roqueme, 2019 Parachernes (Parachernes) adelaidae Bedoya-Roqueme 2019: 24 ���25. Type locality: Punta Nisperal, San Antero, C��rdoba, Colombia (9.3931; -75.7879). Distribution: known only from the type locality. Distribution in Colombia: known only from the type locality (Bedoya-Roqueme 2019). Repository of Colombian material: holotype (♀) and paratypes (4 ♂, 2 ♀) in LEUC-Ps-026 (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 126, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488

Parachernes (Parachernes) plumosus (With, 1908) Chelifer plumosus With 1908: 279–280. Parachernes (Argentochernes) plumosus (With): Beier 1932d: 120–121. Parachernes plumosus (With): Beier 1959: 208. Parachernes (Parachernes) plumosus (With): Harvey 1991: 618. Type locality: without a precise location, “ Moka, Venezuela ”. Distribution: Colombia and Venezuela. Distribution in Colombia: Sucre: Colosó (9.5292, -75.3578) (Bedoya-Roqueme 2019) Repository of Colombian material: LEUC-Ps-C011 (5 ♀) (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 127, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["With, C. J. (1908) An account of the South American Cheliferinae in the collections of the British and Copenhagen Museums. Transactions of the Zoological Society of London, 18 (1), 217 - 340. https: // doi. org / 10.1111 / j. 1469 - 7998.1907. tb 07826. x","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114","Beier, M. (1959) Zur Kenntnis der Pseudoscorpioniden-Fauna des Andengebietes. Beitrage zur Neotropischen Fauna, 1 (3), 185 - 228. https: // doi. org / 10.1080 / 01650525909380614","Harvey, M. S. (1991) Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester, 720 pp."]}

Parachernes (Parachernes) albomaculatus (Balzan, 1892) Chelifer (Trachychernes) albomaculatus Balzan 1892: 526–527. Chelifer albomaculatus Balzan: With 1908: 269–272. Parachernes (Argentochernes) albomaculatus (Balzan): Beier 1932d: 119. Parachernes (Parachernes) albomaculatus (Balzan): Harvey 1991: 613. Type locality: Colonia Tovar, Aragua, Venezuela. (10.4166; -67.2833). Distribution: Brazil, Colombia-? and Venezuela. Distribution in Colombia: without precise locality. Repository of Colombian material: BMNH-? Remarks: The only record of this species is indicated by With (1908) referring the locality as “ New Granada ”, thus the presence in Colombia is uncertain (see discussion)., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 126, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Balzan, L. (1892) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societ Entomologique de France, 60, 497 - 552.","With, C. J. (1908) An account of the South American Cheliferinae in the collections of the British and Copenhagen Museums. Transactions of the Zoological Society of London, 18 (1), 217 - 340. https: // doi. org / 10.1111 / j. 1469 - 7998.1907. tb 07826. x","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114","Harvey, M. S. (1991) Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester, 720 pp."]}

Parazaona ellingsenii (With, 1908) Chelifer ellingsenii With 1908: 287–289. Parazaona ellingsenii (With): Beier 1932d: 144. Type locality: without a precise location: “ New Granada.” Distribution: Colombia? Distribution in Colombia: Bogotáa (With, 1908). Repository of Colombian material: BMNH., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on pages 127-128, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["With, C. J. (1908) An account of the South American Cheliferinae in the collections of the British and Copenhagen Museums. Transactions of the Zoological Society of London, 18 (1), 217 - 340. https: // doi. org / 10.1111 / j. 1469 - 7998.1907. tb 07826. x","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114"]}

Lustrochernes sp. Distribution in Colombia: Santander: Valle de San Jos�� (6.4481; -73.1437) (Murienne et al. 2008)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 131, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Murienne, J., Harvey, M. S. & Giribet, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49 (1), 170 - 184. https: // doi. org / 10.1016 / j. ympev. 2008.06.002"]}

Lustrochernes argentinus (Thorell, 1877) Chelifer argentinus Thorell 1877: 216–218. Chelifer capreolus Balzan 1888: unpaginated. Chelifer (Lamprochernes) argentinus (Thorell): Balzan 1892: 516–517. Lamprochernes argentinus (Thorell): Beier 1930: 211. Lustrochernes argentinus (Thorell): Beier 1932c: 259. Type locality: Córdoba, Argentina (-31.4200; -64.1887). Distribution: Argentina, Brazil, Colombia, Ecuador, Mexico, Paraguay, Peru, Puerto Rico, Trinidad and Tobago and Venezuela. Distribution in Colombia: Valle del Cauca: Buenaventura (3.8830; -77.0197) (Beier 1959). Repository of Colombian material: CAS., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on pages 125-126, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Thorell, T. (1877) Sobre algunos Aracnidos de la Republica Argentina. 1. Scorpiones, Opiliones y Pseudoscorpiones. Periodico Zoologico, 2, 201 - 218.","Balzan, L. (1888) Chernetidae nonnullae Sud-Americanae, III. Privately published, Asuncion. [unpaginated]","Balzan, L. (1892) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societ Entomologique de France, 60, 497 - 552.","Beier, M. (1930) Die Pseudoskorpione des Wiener Naturhistorischen Museums. III. Annalen des Naturhistorischen Museums in Wien, 44, 199 - 222.","Beier, M. (1932 c) Zur Kenntnis der Lamprochernetinae (Pseudoscorp.). Zoologischer Anzeiger, 97, 258 - 267.","Beier, M. (1959) Zur Kenntnis der Pseudoscorpioniden-Fauna des Andengebietes. Beitrage zur Neotropischen Fauna, 1 (3), 185 - 228. https: // doi. org / 10.1080 / 01650525909380614"]}

Neochernes sp. Distribution in Colombia: Nari��o: Barbacoas (1.6733; -78.1385) (Romero-Ortiz 2015c). Colombian material: ICN-APs-479, 485., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 131, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Romero-Ortiz, C. (2015 c) Orden Pseudoscorpiones (Chelonethi). In: Florez, E., Romero-Ortiz, C. & Lopez-Cubillos, S. (Eds.), Los artropodos de la Reserva Natural Rio Nambi. Universidad Nacional de Colombia, Bogota, pp. 89 - 108."]}

Semeiochernes armiger (Balzan, 1892) [NEW RECORD] Chelifer (Trachychernes) armiger Balzan 1892: 527–528. Chelifer armiger Balzan: With 1908: 261. Semeiochernes armiger (Balzan): Beier 1932d: 181–182. Type locality: Pebas, Loreto, Peru (-3.3224; -71.8617). Distribution: Colombia, Ecuador, Panama, Peru, Trinidad and Tobago and Venezuela. Distribution in Colombia: Nariño: Tumaco (1.8077; -78.7705). Repository of Colombian material: ICN-APs-646 (2 ♀). Remarks: Collection data as in label: 2 masl; 10-ago-2017; William Gavis leg.; phoretic on a Pantophthalmus fly. The association of Semeiochernes with giant flies has been previously recorded (Zeh and Zeh 1992a). Since Zeh and Zeh (1992b) work on sexual dimorphisms and phenotypic variation, Semeiochernes species are in strong doubt and waiting for a further taxonomic analysis. This record is the first in Colombia of S. armiger and agrees with the distribution of the species recorded from Mexico to Brazil including all Colombia’s neighbor countries (Christophoryová et al. 2018), Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 128, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Balzan, L. (1892) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societ Entomologique de France, 60, 497 - 552.","With, C. J. (1908) An account of the South American Cheliferinae in the collections of the British and Copenhagen Museums. Transactions of the Zoological Society of London, 18 (1), 217 - 340. https: // doi. org / 10.1111 / j. 1469 - 7998.1907. tb 07826. x","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114","Zeh, D. W. & Zeh, J. A. (1992 a) Emergence of a giant fly triggers phoretic dispersal in the neotropical pseudoscorpion, Semeiochernes armiger (Balzan) (Pseudoscorpionida: Chernetidae). Bulletin of the British Arachnological Society, 9 (2), 43 - 46.","Zeh, J. A. & Zeh, D. W. (1992 b) Are sexually-selected traits reliable species characters? Implications of intra-brood variability in Semeiochernes armiger (Balzan) (Pseudoscorpionida: Chernetidae). Bulletin of the British Arachnological Society, 9 (2), 61 - 64.","Christophoryova, J., Vidlicka, L., Kautman, V. & Krajcovicov, K. (2018) First record of neotropical genus Semeiochernes Beier, 1932 in Ecuador (Pseudoscorpiones, Chernetidae). Spixiana, 41 (1), 26."]}

Sphenochernes attazi Bedoya-Roqueme, 2019 Sphenochernes attazi Bedoya-Roqueme 2019: 29 -30. Type locality: Patio Bonito-Santa Isabel, Monter��a, C��rdoba, Colombia (8.6116; -75.7422). Distribution: known only from the type locality. Distribution in Colombia: known only from the type locality (Bedoya-Roqueme 2019). Repository of Colombian specimens: holotype (♀) and paratypes (13 ♂, 10 ♀) in LEUC-Ps-034 (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 128, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488

Parachernes (Parachernes) crassimanus (Balzan, 1887) Chelifer crassimanus Balzan 1887b: unpaginated. Chelifer (Trachychernes) crassimanus Balzan: Balzan 1892: 548. Parachernes (Argentochernes) crassimanus (Balzan): Beier 1932d: 121–122. Parachernes (Parachernes) crassimanus (Balzan): Harvey 1991: 614. Type localities: Resistencia, Chaco, Argentina (-27.45300; -58.9891), Asuncion, Paraguay (-25.2639; -57.5878) and “ Mato Grosso, Brazil ”. Distribution: Argentina, Brazil, Colombia, Ecuador, Paraguay, Peru and Venezuela. Distribution in Colombia: without a precise location (Beier 1932d). Repository of Colombian material: not specified in the paper (Beier 1932d)., Published as part of Romero-Ortiz, Catalina, García, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 126, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Balzan, L. (1887 b) Chernetidae nonnullae Sud-Americanae, II. Privately published, Asuncion. [unpaginated]","Balzan, L. (1892) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societ Entomologique de France, 60, 497 - 552.","Beier, M. (1932 d) Pseudoscorpionidea II: Subord. C. Cheliferinea. Das Tierreich, 58, 1 - 294. https: // doi. org / 10.1515 / 9783111435114","Harvey, M. S. (1991) Catalogue of the Pseudoscorpionida. Manchester University Press, Manchester, 720 pp."]}

Colombia is a mega-biodiverse country and rich in ecosystems as different as the Amazon and the Andes. Much is known of the vertebrate fauna, however there is still an information-gap for many arthropod groups including the arachnids. Here, we compile all the information available for pseudoscorpions (Arachnida: Pseudoscorpiones) recorded from Colombia and include several new records and distribution extensions. For each described species, we present information on taxonomic history, type localities, global and local distributions, repository of Colombian specimens, and collection numbers when available. We document 12 families, 45 genera and 65 species of pseudoscorpions for Colombia and most species belong to the families Chernetidae (27 species) and Olpiidae (8 species). We record Beierolpium venezuelense Heurtault, 1982, Geogarypus amazonicus Mahnert, 1979, Sathrochthonius venezuelanus Muchmore, 1989, and Semeiochernes armiger (Balzan, 1892) for the first time for Colombia. We further extend the known ranges of Parachernes melonopygus Beier, 1959 and Paratemnoides nidificator (Balzan, 1888). The data suggest that the Caribbean region of Colombia has the highest number of records. Total numbers are not complete and many other new pseudoscorpion species are expected.

Incachernes sp. Distribution in Colombia: Santander: Curit�� (6.6045; -73.0692) (Murienne et al. 2008)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 131, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Murienne, J., Harvey, M. S. & Giribet, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49 (1), 170 - 184. https: // doi. org / 10.1016 / j. ympev. 2008.06.002"]}

Pachychernes sp. Distribution in Colombia: Bol��var: San Jacinto (9.8527; -75.1756) (Villarreal et al. 2019) Repository of Colombian material: ICN-APs-611 (3 individuals), Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 132, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Villarreal, E., Martinez, N. & Romero-Ortiz, C. (2019) Diversity of Pseudoscorpiones (Arthropoda: Arachnida) in two fragments of dry tropical forest in the colombian Caribbean region. Caldasia, 41 (1), 139 - 151. https: // doi. org / 10.15446 / caldasia. v 41 n 1.72189"]}

Pachychernes aff. subrobustus Repository of Colombian material: BMNH and (1 ♀?) in PCJW. Remarks: The specimen reported in Judson (2010) is a pseudofossil inside of light-yellow piece of Colombian copal (59 x 30 x 14 mm) of the J��rg Wunderlich private collection (F2209). The exact location is unknown., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 126, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Judson, M. L. (2010) Redescription of Chelifer eucarpus Dalman (Arachnida, Chelonethi, Withiidae) and first records of pseudoscorpions in copal from Madagascar and Colombia. Palaeodiversity, 3, 33 - 42."]}

Gomphochernes volkeri Bedoya-Roqueme, 2019 Gomphochernes volkeri Bedoya-Roqueme 2019: 23 ���24. Type locality: Tuis Tuis, Tierra Alta, C��rdoba (8.0340; -76.0916). Distribution: known only from the type locality. Distribution in Colombia: known only from the type locality (Bedoya-Roqueme 2019). Repository of Colombian material: holotype (♂) and paratypes (4 ♂) in LEUC-Ps-023 (Bedoya-Roqueme 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 125, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488

Parachernes sp. Distribution in Colombia: Atl��ntico: Usiacur��-Juan de Acosta (10.7672; -75.0427) (Villarreal et al. 2019); Boyac��: Arcabuco, Santuario de Flora y Fauna Iguaque (5.6722; -73.4747) (Romero-Ortiz 2017). Repository of Colombian material: ICN-APs-084 (1♂) (Romero-Ortiz 2017) and 17 individuals in ICN-APs-677 (Villarreal et al. 2019)., Published as part of Romero-Ortiz, Catalina, Garc��a, Fabian & Villarreal, Eduardo, 2019, Checklist of the false scorpions (Arachnida: Pseudoscorpiones) of Colombia, with new records and a key to the identification of the families, pp. 107-139 in Zootaxa 4711 (1) on page 132, DOI: 10.11646/zootaxa.4711.1.4, http://zenodo.org/record/3573488, {"references":["Villarreal, E., Martinez, N. & Romero-Ortiz, C. (2019) Diversity of Pseudoscorpiones (Arthropoda: Arachnida) in two fragments of dry tropical forest in the colombian Caribbean region. Caldasia, 41 (1), 139 - 151. https: // doi. org / 10.15446 / caldasia. v 41 n 1.72189","Romero-Ortiz, C. (2017) Pseudoescorpiones (Arachnida: Pseudoscorpiones) del nororiente andino de Colombia. Biota Colombiana, 18 (1), 198 - 205. https: // doi. org / 10.21068 / c 2017. v 18 n 01 a 12"]}

Tenebriochernes gen. nov. urn:lsid:zoobank.org:act: A4181424-D1A0-4A5A-8025-1EFC040DF730 Type species: Tenebriochernes concavus n. sp. Etymology. The generic name refers to the presence of the type species within caves (tenebrio, Latin, shunner of light, lover of darkness), which is combined with the genus name Chernes (Greek, labourer). It is masculine in gender. Diagnosis: Tenebriochernes n. gen. can be distinguished from all other chernetid genera by the following combination of characters: presence of 7 denticulate and thickened setae (Sd) proximal of the base of fixed chelal finger; the concavity on the retrolateral margin of the pedipalpal patella; the chelal hand shape and the strong concavity on the base of the movable chelal finger proximal of the condyle; trichobothrium ib, ist, esb and eb grouped at the base of the fixed finger in a diagonal row, trichobothrium st equidistant between t and sb, proximal of nodus ramosus and approximately at the same level as est; females spermathecae with a distally expanded and less globose sacs with darkened tips, and two long tubes which not leading to a single medium chamber with a cribriform plate; tarsus IV without tactile seta. Remarks: The genus Tenebriochernes is referred to the subfamily Chernetinae by lacking T-shaped spermathecae as occurs in the Lamprochernetinae and the angular posterior margin of the carapace, exclusive of the Goniochernetinae. Like a number of other chernetine genera, Tenebriochernes has four blades in the cheliceral rallum and lacks a tactile seta on tarsi III and IV. Tenebriochernes differs from the genera Atherochernes Beier, 1955 from Venezuela, Austrochernes Beier, 1932 and Balgachernes Harvey, 2018 from Australia (Harvey 2018) and Eumechochernes Beier, 1932 from Hawaii (With 1905; Beier 1932, 1954; Harvey 2013) by the pattern of trichobothrium ib, ist, esb and eb grouped at the base of the fixed finger in a diagonal row; trichobothrium st equidistant between t and sb, trichobothrium st almost at the same level as est; venom duct long, nodus ramosus midway between trichobothria t and st; the chelal hand shape and the strong concavity on the base of movable chelal finger proximal of the condyle, the robust pedipalps and the concavity on the retrolateral margin of the pedipalpal patella. It differs from the genus Marachernes Harvey, 1992 from Australia by the absence of an internolateral mound on the male movable chelal finger and the spermathecae with two long tubes and distally expanded sacs with darkened tips. It differs from the genera Mirochernes Beier, 1930 from United States and Semeiochernes Beier, 1932 from Costa Rica (Beier 1930, 1932; Harvey 2013) by the robust and granulated pedipalps highly sexually dimorphic, the males possess greatly enlarged chelae and chelal peg-like processes, that are absent in females (Chrystophoryov�� et al. 2018), as well as the distribution of the trichobothria, with ist distal of est on the fixed chelal finger and st closer to t than to sb on the movable chelal finger. It differs from the genera Dinocheirus Chamberlin, 1929 from American region, Asia and Europe, that bears only five setae on the cheliceral hand (b and es being acuminate), tarsus with an acuminate tactile seta distal of midpoint, position of the trichobothrium ib at the same level of est, and trichobothrium isb proximal of it (Chamberlin 1929; Hoff 1947; Muchmore 1974). The morphology of the spermatheca of females has been found to provide reliable characters for the delimitation of genera in Chernetidae (e.g. Muchmore 1974, 1975; Muchmore & Hentschel 1982, Hentschel & Muchmore, 1989; Mahnert 1979, 1985; Harvey 1995; Harvey & Volschenk 2007). The spermathecae of Tenebriochernes have two long tubes, opening distally into expanded sacs with darkened tips, which distinguish it from the genera Chernes Menge, 1855 in the Holarctic region, Ceriochernes Beier, 1937 from Asia and South America, Cocinachernes Hentschel & Muchmore, 1989 from Mexico, Illinichernes Hoff, 1949, Neochelanops Beier, 1964, Semeiochernes Beier, 1932; Hesperochernes Chamberlin, 1924 from the American region; Maxchernes Feio, 1960 from South America; Austrochernes Beier, 1932, Paraustrochernes Beier, 1966, and Satrapanus Harvey and Volschenk, 2007 from Australia (Beier 1932; 1937; 1966; Hoff 1949; Hentschel & Muchmore 1989; Muchmore 1999; Harvey & Volschenk 2007; Harvey 2013). Unfortunately, the spermathecae of many chernetine genera remain undescribed and others have yet to be studied in detail, which hinders our understanding of the group. In this sense, a further review of the spermathecal morphology is necessary for each of the genera of Chernetidae family, to clarify and establish phylogenetic relationships within this large and complex family (e.g. Harvey 1991, 1995, 2013; Muchmore 1974, 1975; Hentschel & Muchmore 1989). Finally, Tenebriochernes resembles Neoallochernes Hoff, 1947 from the United States, Central America, and some Caribbean islands in having spermatheca with dark tips. The genus Neoallochernes is characterized by the spermathecae in form of elongated sacs with short slender tubules leading into a single median chamber with a cribriform plate; in this sense, the unusual spermathecal morphology is clearly diagnostic for this genus (e.g. Hoff 1957; Muchmore 1975, 1992, 1996). Tenebriochernes can be distinguished from Neoallochernes by the spermathecae with distally expanded and less globose sacs, and two long tubes which are not leading to a single medium chamber with a cribriform plate. Likewise, Tenebriochernes has the setae sbs on the cheliceral hand (absent in Neoallochernes), four blades on the rallum (three blades in Neoallochernes), and also differs by the concavity on the prolateral margin of the pedipalpal patella; further in the shape of the chelal hand: the strong concavity at the base of the movable chelal finger proximal of the condyle, and the distribution of trichobothria ib, ist, esb and eb which are grouped at the base of the fixed finger in a diagonal row. Due to these characters, the new genus can be distinguished from all other genera of the family Chernetidae by the shape of the spermathecae, the chelal hand shape with a strong concavity near the base of the movable finger, the form of the retrolateral accessory teeth of the movable chelal finger, proximal of fingertip of fixed chelal finger, the position of the trichobothria, and the concavity on the retrolateral margin of the pedipalpal patella., Published as part of Bedoya-Roqueme, Edwin & Torres, Richard A., 2019, Tenebriochernes, a new genus and species of Chernetidae (Arachnida: Pseudoscorpiones) from north-western Colombia, with ecological observations, pp. 87-107 in Zootaxa 4624 (1) on pages 89-90, DOI: 10.11646/zootaxa.4624.1.6, http://zenodo.org/record/3258482, {"references":["Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294.","Harvey, M. S. (2018) Balgachernes occultus, a new genus and species of pseudoscorpion (Pseudoscorpiones: Chernetidae) associated with balga (Xanthorrhoea preissii) in south-western Australia, with remarks on Austrochernes and Troglochernes. Records of the Western Australian Museum, 33 (1): 115 - 130.","With, C. J. (1905) On Chelonethi, chiefly from the Australian region, in the collection of the British Museum, with observations on the \" coxal sac \" and on some cases of abnormal segmentation. Annals and Magazine of Natural History, 15 (7), 94 - 143, 328.","Beier, M. (1954) Eine Pseudoscorpioniden-Ausbeute aus Venezuela. Memorie del Museo Civico di Storia Naturale di Verona, 4, 131 - 142.","Harvey, M. S. (2013) Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues-beta / pseudoscorpions (accessed 15 May 2018)","Harvey, M. S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata). Invertebrate Taxonomy, 6, 1373 - 1435. https: // doi. org / 10.1071 / IT 992137 3","Beier, M. (1930) Die Pseudoskorpione des Wiener Naturhistorischen Museums. III. Annalen des Naturhistorischen Museums in Wien, 44, 199 - 222.","Chamberlin, J. C. (1929) Dinocheirus tenoch, an hitherto undescribed genus and species of false scorpion from Mexico (Arachnida - Chelonethida). Pan-Pacific Entomologist, 5, 171 - 173.","Hoff, C. C. (1947) The species of the pseudoscorpion genus Chelanops described by Banks. Bulletin of the Museum of Comparative Zoology, 98, 471 - 550.","Muchmore, W. B. (1974). Clarification of the genera Hesperochernes and Dinocheirus (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 2, 25 - 36","Muchmore, W. B. (1975) Use of the spermathecae in the taxonomy of chernetid pseudoscorpions. Proceedings of the International Arachnological Congress, Amsterdam, 6, 17 - 20.","Muchmore, W. B. & Hentschel, E. (1982) Epichernes aztecus, a new genus and species of pseudoscorpion from Mexico (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 10, 41 - 45.","Hentschel, E. & Muchmore, W. B. (1989) Cocinachernes foliosus, a new genus and species of pseudoscorpion (Chernetidae) from Mexico. Journal of Arachnology, 17, 345 - 349.","Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810.","Mahnert, V. (1985) Pseudoscorpions (Arachnida} recoltes durant la mission speologique espagnole au Perou en 1977. Revue Arachnologique, 6, 17 - 28.","Harvey, M. S. (1995) Barbaraella gen. nov. and Cacoxylus Beier CPseudoscorpionida: Chernetidae), two remarkable sexually dimorphic pseudoscorpions from Australasia. Records ofthe Western Australian Museum Supplement, 52, 199 - 208.","Harvey, M. S. & Volschenk, E. S. (2007) Review of some Australasian Chernetidae: Sundochernes, Troglochernes and a new genus (Pseudoscorpiones). The Journal of Arachnology, 35, 238 - 277.","Hoff, C. C. (1949) The pseudoscorpions of Illinois. Bulletin of the Illinois Natural History Survey, 24, 407 - 498.","Beier, M. (1964) Further records of Pseudoscorpionidea from the Solomon Islands. Pacific lnsects, 6, 592 - 598.","Beier, M. (1966). On the Pseudoscorpionidea of Australia. Australian Journal of Zoology, 14, 275 - 303.","Muchmore, W. B. (1999) Redefinition of the genus Chelanops Gervais (Pseudoscorpionida: Chernetidae). Pan-Pacific Entomologist, 75, 103 - 111.","Hoff, C. C. (1957) Tejachernes (Arachnida - Chelonethida, Chernetidae - Chernetinae), a new genus of pseudoscorpion based on Dinocheirus stercoreus. Southwestern Naturalist, 2, 83 - 88.","Muchmore, W. B. (1992) Cavernicolous pseudoscorpions from Texas and New Mexico (Arachnida: Pseudoscorpionida). Texas Memorial Museum, Speleological Monographs, 3, 127 - 153.","Muchmore, W. B. (1996) A new species of Neoallochernes from Antigua (Pseudoscorpionida: Chernetidae). Caribbean Journal of Science, 32, 387 - 389."]}

Tenebriochernes pilosus n. sp. Figs 7–8; 40–54; 55. urn:lsid:zoobank.org:act: 409B7802-A553-431F-AACA-0C2940FD2E6C Material examined. Female holotype: Montería, Córdoba department, Colombia: Patio Bonito-Santa Isabel, 8°36´42” N, 75°44´32” W, elev. 192 m, 2 December 2013, E. Bedoya-Roqueme leg. (LEUC; Ps-040). Two male paratypes, same data and repository as holotype (LEUC; Ps-041). Diagnosis. Tenebriochernes pilosus n. sp., can be easily distinguished by the trichobothria et and it located distally near fingertip, est proximal of isb, ib distal of ist and distinctly proximal of isb, trichobothrium ib, ist, esb and eb grouped at the base of the fixed finger in a diagonal row, st proximal to the level of est and close to the margin of the finger; nodus ramosus proximal of t and distal of st; length/ width of the femur, female 2.8 × (males 2.5–2.6 ×); patella 2.6 × (males, 2.5–2.6 ×); hand with pedicel 1.5 × (males, 1.6–1.7 ×); chela with pedicel 2.3 × (males, 2.5–2.6 ×); chelal fixed finger of female with 34 teeth offset in saw-like arrangement, two retrolateral accessory teeth and one prolateral accessory tooth (males, 30-32 teeth, two retrolateral accessory teeth, without prolateral accessory tooth); movable chelal finger of female with 38 teeth, two retrolateral accessory teeth and two prolateral accessory teeth (males, 34-36 teeth, two retrolateral accessory teeth and without prolateral accessory teeth). Spermatheca consisting of distally well-expanded and less globose sacs with darkened tips, and two shorts tubes which do not lead to a single medium chamber. Adult. Carapace and pedipalps red-brownish, legs yellow-brownish, heavily granulate (Figs 40–41). Carapace (Fig. 42) with granules of mesozone hexagon-shaped, large and spaced (Figs 12–13); with two indistinct transverse furrows which are granulate; subbasal transverse furrow closer to medial furrow than to posterior margin; without eyes or eye-spots; female with about 105 setae (males with about 95-98 setae), each denticulate, short and strongly clavate; female with six setae on anterior margin and 16 on posterior margin (males with nine setae on anterior margin and 16-18 setae on posterior margin), each short and strongly clavate; 1.3× as long as broad (males 1.1-1.2× as long as broad). Pedipalpal coxa of female and males with three subdistal setae, indistinctly granulate, all setae simple and acuminate, coxa I with 30-32 setae, coxa II 28-30 (of which two suboral microsetae), coxa III 34 –38, and coxa IV 36 –48 setae. Pleural membrane longitudinally striated (Figs 40–41). All the tergites divided (except the XI), male with tergite I apparently not divided, and XI divided (Figs 40–41); tergites with scaly sculpture, strongly granulate; all the setae broadly clavate setae and on tubercles. Chaetotaxy of tergites I–XII: female holotype: 14: 16: 18: 20: 20: 22: 20: 20: 16: 14: 10: 2; male paratypes: 14–16: 16-18: 18-20: 18–20: 20-22: 20-22: 14-16: 14-16: 14-16: 12-14: 10-12: 2. Tergites IV-IX each with two lateral and two medial setae, without tactile setae on tergite XI. Sternites divided, with scaly sculpture, all the setae broadly clavate. Sternal chaetotaxy of female holotype, sternites II–XII: 32: (3) 14 (3): (3) 18 (3): 18: 18: 16: 16: 16: 16: 10: 2; males, 28(30): (3)30(3):(3)16–18(3): 16-18: 16-18: 16-18: 14-16: 14- 16: 14-16: 10-12: 2, no tactile setae. Anterior genital operculum of a female with 32 setae on each side and 14 setae along the posterior margin (Fig. 53). Spermatheca consisting of distally well-expanded and less globose sacs with darkened tips, and two shorts tubes which do not lead to a single medium chamber (Fig. 53). Anterior genital operculum of male with a set of 30 microsetae grouped on each side of operculum on the anterior region, and a set of 28 long setae located on the center, posterior genital operculum with 28-30 setae along the posterior margin (Fig. 54). Chelicera: hand with 6 setae, setae es, bs 1 bs 2 and sbs denticulate, ls and is long and acicular (Fig. 43); fixed finger with 3 large and 2 small teeth, movable finger with a subapical tooth–like lobe; galea of female and males with the tips bifid and a subdistal ramus (Figs 45–46); serrula exterior 20–21 blades; rallum with 4 blades, distal blade dentate, others smooth (Fig. 44). Pedipalps (Figs 47–48): elongated and robust, strongly granulate, with broad clavate setae on tubercles, setae on chelal hand more slender, those on both fingers simple; trochanter with a rounded dorsal hump, female holotype: 1.8× (males, 1.7–1.8×); femur robust, basally pedicelled, 2.8× (males 2.5–2.6×); patella robust, retrolateral margin with a concavity near midpoint of segment, 2.6× (males, 2.5–2.6×); hand with pedicel 1.5× (males, 1.6–1.7×); chela with pedicel 2.3× (males, 2.5–2.6×); 7 long denticulate and thickened setae (Sd) proximal of base of fixed finger chelal; base of movable chelal finger with strong concavity proximal of condyle and four short denticulate and thickened setae (Sd) (Fig. 49). Fixed finger of female with 34 teeth offset in saw-like arrangement, giving heterodonty shape, including 22 flattened proximal teeth and 12 acute, spaced teeth; two retrolateral accessory teeth and one prolateral accessory tooth (males, with 30-32 teeth, giving heterodonty shape, including 20 flattened proximal teeth and 12 acute spaced teeth; two retrolateral accessory teeth, without prolateral accessory teeth) (Fig. 49). Fingertip of fixed chelal finger with four apical teeth in saw-shape; movable chelal finger of female with 38 teeth, two retrolateral accessory teeth and two prolateral accessory teeth; two retrolateral accessory teeth in saw-shape near to fingertips on internal margin of movable finger (males, 34-36 teeth, two retrolateral accessory teeth and without prolateral accessory teeth; 3 retrolateral accessory teeth in saw-shape near to fingertips on internal margin of movable finger). Trichobothria: et and it located distally near fingertip, est proximal of isb, ib distal of ist and distinctly proximal of isb, trichobothrium ib, ist, esb and eb grouped at the base of the fixed finger in a diagonal row, st proximal to the level of est and close to the dental margin of the finger; nodus ramosus proximal of t and distal of st (Fig. 49). All legs yellowish, surface granulate, setae long and multidentate, hence pilose appearance on the legs (Figs 50–51), tactile setae on tarsus IV absent (Fig. 51). Leg I: trochanter female holotype 1.2× (males, 1.3–1.4×), femur 1.5× (1.7–1.8×), patella 2.4× (2.0-2.1×), tibia 4.4× (4.1–4.3×), tarsus 5.1× (5.0–5.2×). Leg IV: trochanter female holotype 1.8× (males 1.6–1.7×), femur + patella 2.3× (2.6–2.7×), tibia 5.3× (5.5–5.7×), tarsus 5.5× (5.1–5.2×), subterminal setae toothed and curved; claws extremely toothed and longer than the undivided arolia (Fig. 52). Measurements (mm): Female holotype: body length 3.9; carapace 1.04/0.85. Chelicera 0.30/0.16. Pedipalp: trochanter 0.45/0.27; femur 0.96/0.35; patella 0.90/0.36; hand (with pedicel) 0.93/0.66; chela (with pedicel) 1.55/0.66; chela (without pedicel) 1.48/0.66; movable finger length 0.69. Leg I: trochanter 0.21/0.18; femur 0.26/0.18; patella 0.36/0.16; tibia 0.48/0.11; tarsus 0.46/0.09. Leg IV: trochanter 0.29/0.17; femur + patella: 0.47/0.19; tibia 0.58/0.11; tarsus 0.50/0.09. Male paratypes: Body length 3.5-3.6; carapace 0.85–0.86/0.78–0.79. Chelicera 0.30–0.31/0.15–0.16. Pedipalp: trochanter 0.45–0.46/0.25–0.26; femur 0.87–0.89/0.30–0.31; patella 0.86–0.87/0.33–0.34; hand (with pedicel) 0.85–0.86/0.67–0.68; chela (with pedicel) 1,50–1,51/0,67–0,68; chela (without pedicel) 1.45–1.46/0.67–0.58; movable finger length 0.61–0.63. Leg I: trochanter 0.18–0.19/0.12–0.14; femur 0.26–0.27/0.13–0.14; patella 0.33– 0.34/0.14–0.15; tibia 0.44–0.45/0.10–0.11; tarsus 0.41–0.42/0.08–0.09. Leg IV: trochanter 0.26–0.27/0.15–0.16; femur + patella 0.48–0.49/0.20–0.21; tibia 0.60–0.61/0.11–0.12; tarsus 0.50–0.51/0.10–0.11. Etymology. The specific epithet (pilosus) is a Latin adjective meaning “pilosus”, emphasizing the hairy legs in both male and female. Distribution (Fig. 55).: Only known from the type locality.

Genus Orochernes Beier, 1968 Orochernes Beier, 1968: 17. Type species: Orochernes nepalensis Beier, 1968 b, by original designation., Published as part of Gao, Zhizhong & Zhang, Feng, 2019, First record of the genus Orochernes (Pseudoscorpiones: Chernetidae) from China, with description of a new species, pp. 126-132 in Zootaxa 4612 (1) on page 127, DOI: 10.11646/zootaxa.4612.1.9, http://zenodo.org/record/3233671, {"references":["Beier, M. (1968) Ein neues Chernetiden-Genus (Pseudoscorp.) aus Nepal. Khumbu Himal, 3, 17 - 18."]}

The chernetid genus Orochernes Beier, 1968 is reported from China for the first time. Orochernes ganziensis sp. n. is described from the Sichuan Province and characterized through illustrations and genetic data. The spermathecae of the genus Orochernes are illustrated for the first time.

The pseudoscorpion genus Verrucachernes Chamberlin, 1947 is widely distributed in the Old World tropics, with three named Australasian and West Pacific species and two from Africa. A review of some pseudoscorpions described from India has revealed that Withius parvus Beier, 1930 (currently in the genus Metawithius) and Pselaphochernes indicus Beier, 1974 are misplaced and actually belong to the genus Verrucachernes Chamberlin, 1947, forming the new combinations V. parvus (Beier, 1930) comb. nov. and V. indicus (Beier, 1974) comb. nov., respectively. Both species possess the single, large, rounded spermatheca and other features typical of Verrucachernes.