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Proprioseiopsis penai Denmark and Evans Proprioseiopsis penai Denmark and Evans, in Denmark et al., 1999: 17. The biology of this species found only once in Honduras on Citrus sp. by Denmark et al. (1999) is unknown. Previous Records — Honduras (Denmark et al., 1999; Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, RiviŁre Corossol, Route du Col des Mamelles, 16°10’N, 61°41’W, alt. 221 m, 1 ♀ on an unknown small mauve flowering plant, Rault coll., 22 December 2008; Basse-Terre, Petit-Bourg, Domaine Duclos de l’INRA Antilles-Guyane, 16°12’N, 61°39’W, alt. 85 m, 2 ♀ on Centrosema pubescens, Kreiter coll., 31 December 2008 and 1 ♀ on Clidemia hirta, Kreiter coll., 3 January 2009. Remarks — this is the first record of this species in the West Indies. The measurements of three of the four specimens collected fit very well the measurements given by Denmark et al. (1999). The average measurements of these three adult females are as follow: dorsal shield sclerotized and reticulated, dorsal shield length 449 (439-464), width 332 (321-342), 5 solenostomes (gd 1, 2, 5, 8 and 9), j1 34 (33 – 35), j3 48 (47 – 49), j4 4, j5 6 (5 – 6), j6 8 (8 – 9), J2 5, J5 4, z2 26 (25 – 27), z4 22 (20 – 24), z5 5 (5 – 6), Z1 7 (6 – 8), Z4 93 (83 – 108), Z5 103 (102 – 105), s4 121 (121 – 122), S2 7 (5 – 8), S4 7 (6 – 7), S5 7 (7 – 8), r3 26 (19 – 33), R1 8 (7 – 9), all setae smooth, st1-st1 66, st2-st2 84 (84 – 85), st1-st3 64 (63 – 64), posterior margin of the sternal shield concave, metapodal 1 length 33 (32 – 35), width 6 (5 – 7), metapodal 2 length 16 (13 – 19), width 1, ventrianal shield 121 (120 – 123) long, 120 (119 – 120) wide at level of anterior corners and 99 (95 – 102) wide at level of anus, 5 poroids around the ventrianal shield and genital shield, JV5 68, spermatheca 20 (18 – 21) long and 11 (9 – 13) width, SgeI 32 (30 – 33), SgeII 34 (33 – 35), SgeIII 39 (38 – 40), StiIII 28, SgeIV 79 (78 – 79), StiIV 53 (50 – 58), StIV 66 (61 – 68), macrosetae whip-like, fixed digit of chelicerae 35 (33 – 37), mobile digit of chelicerae 35 (35 – 37), 13 denticles on fixed digit and 3 on mobile digit., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 291, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Quadromalus colombiensis Moraes, Denmark and Guerrero Quadromalus columbiensis Moraes, Denmark and Guerrero, 1982: 17 The biology of this species found only once in Colombia on Tanicum paniculatum (Moraes et al., 1982) remains totally unknown. Previous Records — Colombia (Moraes et al., 1982; Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Petit-Bourg, Domaine Duclos de l’INRA Antilles-Guyane, 16°12’N, 61°39’W, alt. 85 m, 3 ♀ + 1 ♂ on Cissus verticilata, Kreiter coll., 18 December 2008. Remarks — this is the first record of this species in the West Indies. The measurements and description of the three female specimens collected fit very well those given by Moraes et al. (1982). No measurement for the male was given in Moraes et al. (1982). Diagnosis proposed by Moraes et al. (1982) corresponds very well to the specimens collected in Guadeloupe: Z4 and Z5 not serrated, only one macroseta on leg IV, 3-4 teeth on fixed and 3 on movable digits, respectively, and moreover 4 extralong setaceous setae on the tarsus 1 (Moraes et al., 1982). The average measurements of three adult females presently found are as follow: dorsal shield smooth, dorsal shield length 370 (361 – 385), width 246 (239 – 2555), 6 solenostomes (gd 1, 2, 4, 6, 8 and 9), j1 29 (28 – 30), j3 38 (37 – 40), j4 13 (12 – 13), j5 14 (13 – 15), j6 12, J2 19 (17 – 20), J5 13, z2 37, z4 41 (40 – 42), z5 17 (15 – 20), Z1 27 (25 – 3), Z4 34 (33 – 35), Z5 33, s4 44 (42 – 45), S2 34 (33 – 35), S4 31 (28 – 32), S5 28 (25 – 30), r3 34 (33 – 35), R1 30, all setae smooth, st1-st1 56 (55 – 58), st2-st2 66 (60 – 75), st1-st3 70 (60 – 75), posterior margin of the sternal shield not straight, metapodal 1 length 25, width 7 (6 – 8), metapodal 2 not visible, ventrianal shield 103 (100 – 108) long, 57 (48 – 67) wide at level of anterior corners and 68 (60 – 75) wide at level of anus, 4 poroids around the ventrianal shield, JV5 41 (40 – 43), spermatheca 24 (20 – 28) long and 3 width, SgeI 25 (23 – 28), SgeII 22 (20 – 23), SgeIII 24 (23 – 25), StiIII 21 (20 – 23), SgeIV 29 (28 – 30), StiIV 26 (24 – 28), StIV 41 (40 – 42), macrosetae very slightly knobbed, fixed digit of chelicerae 31 (28 – 32), mobile digit of chelicerae 32 (30 – 35), 3 large strong denticles on fixed digit and 3 large strong denticles on mobile digit. The measurements of one adult male are: dorsal shield smooth, dorsal shield length 298, width 225, 6 solenostome (gd 1, 2, 4, 6, 8 and 9), j1 20, j3 25, j4 10, j5 9, j6 10, J2 15, J5 9, z2 25, z4 28, z5 12, Z1 23, Z4 25, Z5 25, s4 35, S2 25, S4 23, S5 23, r3 25, R1 23, all setae smooth, st1-st1 48, st2-st2 63, st1-st3 58, st1-st5 120, ventrianal shield 100 long, 167 wide at level of anterior corners and 60 wide at level of anus,JV5 25, SgeI 23, SgeII 23, SgeIII 15, StiIII 15, SgeIV 23, StiIV 24, StIV 28, schaft of the spermatodactyl 23.

Phytoseius woodburyi De Leon Phytoseius woodburyi De Leon, 1965: 130; De Leon, 1967: 12; Prasad, 1968: 1461; Denmark and Muma, 1975: 295; Denmark and Muma, 1978: 15; Moraes et al., 1991: 133; Kreiter and Moraes, 1997; 380; Moraes et al., 2000: 260. The biology of this species remains unknown. Previous Records: Brazil, Colombia, French Antilles (Guadeloupe, Marie-Galante, Martinique) (Moraes et al., 2000), Hawaii, India, Jamaica, Puerto Rico, Trinidad (Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Volcan La SoufriŁre, 16°02’45.19”N, 61°31’55.65”W, alt. 1300 m, 6 ♀ on Centropogon cornutus, Kreiter coll., 21 June 2008; Vieux-Habitants, Station CIRAD de Le Bouchu, 16°03’N, 61°45’W, alt. 21 m, 1 ♀ on Erythrina sp. and 1 ♀ on Psidium guajava, Kreiter and Rault coll., 19 December 2008; Les Saintes, Terre de Bas, L’Etang, 15°51’N, 61°37’W, alt. 239 m, 18 ♀ on Cordia alliodora, Kreiter coll., 2 January 2009; Martinique, Lamentin, 14°39’N, 60°58’W, alt. 46 m, 1 ♀ on Psidium guajava, Kreiter coll., 15 November 2010; Case-Pilote, 14°39’N, 61°07’W, alt. 176 m, 1 ♀ on Psidium guajava, Kreiter coll., 22 November 2010. Remarks — the measurements and description of the specimens collected fit very well those given by Kreiter and Moraes (1997)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 299, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["De Leon D. 1965 - A note on Neoseiulus Hughes 1948 and new synonymy (Acarina: Phytoseiidae) - Proceed. Entomol. Soc. Washington 67 (1): 23.","De Leon D. 1967 - Some mites of the Caribbean Area. Part I. Acarina on Plants in Trinidad, West Indies - Allen Press Inc., Lawrence, Kansas, 66 pp.","Prasad V. 1968 - Some phytoseiid mites from Hawaii - Ann. Entomol. Soc. America 61 (6): 1459 - 1462.","Denmark H. A., Muma M. H. 1975 - The Phytoseiidae (Acarina: Mesostigmata) of Puerto Rico - J. Agric. Univ. Puerto Rico 59, 279 - 304.","Denmark H. A., Muma M. H. 1978 - Phytoseiidae of Jamaica, an annoted list (Acari: Mesostigmata) - Intern. J. Acarol. 4 (1): 1 - 22.","Moraes G. J. de, Mesa N. C., Braun A. 1991 - Some phytoseiid mites of Latin America (Acari: Phytoseiidae) - Intern. J. Acarol. 17: 117 - 139.","Kreiter S., Moraes G. J. de, 1997 - Phytoseiidae mites (Acari: Phytoseiidae) from Guadeloupe and Martinique - Fla. Entomol. 80: 376 - 382.","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Arrenoseius crassipes (Denmark) Paraamblyseius crassipes Denmark, 1988: 37. The biology of this species found only once in Mexico on Eichomia crassipes by Denmark (1988) is unknown. Previous Records — Mexico (Denmark, 1988; Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Vernou, Route du Col des Mamelles, 16°11’N, 61°39’W, alt. 184 m, 2 ♀ and 1 ♂ on Ricinus communis, Kreiter coll., 26 December 2008. Remarks — this is the first record of this species in the West Indies. The measurements of the specimens collected fit well the measurements given by by Denmark (1988). Macrosetae are present on all legs but were not mentioned and measured in Denmark (1988). The average measurements of the two adult females presently found are as follow: dorsal shield sclerotized and reticulated, dorsal shield length 353- 422, width 300 – 325, 3 solenostomes (gd 1, 8 and 9), j1 20, j3 25, j4 4, j5 5, j6 5, J5 9 – 12, z2 8 – 10, z4 10 – 13, z5 5, Z1 8, Z4 85 – 100, Z5 95 – 100, s4 87, S2 12, S4 10, S5 10, r3 10, R1 12, all setae smooth, st1-st1 53 – 55, st2-st2 64 – 75, st1-st3 40 – 43, posterior margin of the sternal shield concave, metapodal 1 length 28, width 10, metapodal 2 length 8, width 5, ventrianal shield 150 – 153 long, 210 – 212 wide at level of anterior corners and 150 wide at level of anus, 2 poroids around the ventrianal shield, JV5 40, spermatheca 23 – 25 long and 5 – 7 width, StIV 40, macrosetae whip-like, fixed digit of chelicerae 38 – 40, mobile digit of chelicerae 38 – 40, 11 – 12 denticles on fixed digit and 2 on mobile digit., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 292, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Denmark H. A. 1988 - Revision of the genus Paraamblyseius Muma (Acari: Phytoseiidae) - Intern. J. Acarol. 14 (1): 23 - 40. doi: 10.1080 / 01647958808683801","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Typhloseiopsis pritchardi (Chant and Baker) Amblyseius pritchardi Chant and Baker, 1965: 15. Amblyseius pritchardi Chant and Yoshida-Shaul, 1983: 1037. Typhloseiopsis pritchardi Moraes et al., 2000: 259. The biology of this species remains unknown. Previous Records — Costa Rica (Moraes et al., 2004b), French Antilles (Gualeloupe, Marie- Galante, Martinique, Saint-Martin) (Moraes et al., 2000). Specimens examined — La DØsirade, Parc Eolien, 16°19’N, 61°02’W, 269 m, 3 ♀ and 1 ♂ on Coccoloba pubescens, Kreiter coll., 5 January 2009. Remarks — the measurements and description of the specimens collected fit very well those given by Moraes et al. (2000), Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 299, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Chant D. A., Baker, E. W. 1965 - The Phytoseiidae (Acarina) of Central America - Mem. Entomol. Soc. Canada, 41, 56 pp.","Chant D. A., Yoshida-Shaul E. 1983 - A world review of the simplex species group in the genus Typhlodromus Scheuten (Acarina: Phytoseiidae) - Can. J. Zool. 61: 1142 - 1151. doi: 10.1139 / z 83 - 151","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Amblydromalus higuilloae (Denmark and Muma) Typhlodromalus higuilloae Denmark and Muma, 1975: 292. Typhlodromalus higuilloae Denmark et al., 1999: 59. The biology of this species found only two times in Puerto Rico (Denmark and Muma, 1975) on "higuillo" leaves (Piper marginatum) and in Honduras by Denmark et al. (1999) on Calea urticifolia remains unknown. Previous Records — Puerto Rico (Denmark and Muma, 1975; Moraes et al., 2004b), Honduras (Denmark et al., 1999; Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Petit-Bourg, Domaine Duclos de l’INRA Antilles-Guyane, 16°12’N, 61°39’W, alt. 85 m, 1 ♀ on Vigna sp., Kreiter coll., 21 December 2008; Basse-Terre, Route ForestiŁre, Jules-Grosse Montagne, 16°12’N, 61°39’W, alt. 90 m, 1 ♀ on Vigna sp., Kreiter coll., 24 December 2008; Basse-Terre, riviŁre Corossol, Cascade des Ecrevisses, 16°11’N, 61°39’W, alt. 184 m, 1 ♀ on an unknown Melastomataceae, Kreiter coll., 21 December 2008; Basse- Terre, Vernou, Route du Col des Mamelles, 16°11’N, 61°39’W, alt. 184 m, 1 ♀ on Vigna sp., Kreiter coll., 26 December 2008; Grande-Terre, Porte d’Enfer, beach, 16°31’N, 61°28’W, alt. 2 m, 2 ♀ on Pluchea symphytifolia, Kreiter coll., 27 December 2008; Basse-Terre, 1 Łre Chute du Carbet, 16°03’N, 61°39’W, alt. 890 m, 1 ♂ on Clidemia umbrosa, Kreiter coll., 29 December 2008; Martinique, Basse-Pointe, Jardins ClØment, 14°50’N, 61°05’W, alt. 91 m, 2 ♀ on Panicum maximum, Kreiter coll., 16 November 2010. Remarks — this is the first record of this species in the French West Indies. The measurements of five (4 ♀ and 1 ♂) of the nine specimens collected fit rather well the measurements given by Denmark et al. (1999). All setae are however slightly shorter, between 1 and 5 for the greater, which represent less than 10 % of variation, which is less than the intraspecific of 20 % around the mean defined by Tixier (2012). The average measurements of the four adult females presently found are as follow: dorsal shield sclerotized and reticulated, dorsal shield length 351 (350 – 352), width 232 (223 – 250), 5 solenostomes (gd 1, 2, 6, 8 and 9), j1 25 (22 – 28), j3 42 (40 – 44), j4 12 (9 – 13), j5 11 (10 – 14), j6 11 (11 – 14), J2 13 (12 – 14), J5 11 (9 – 14), z2 18 (16 – 21), z4 31 (30 – 33), z5 9 (7 – 9), Z1 12 (11 – 13), Z4 43 (42 – 44), Z5 67 (65 – 71), s4 66 (63 – 68), S2 26 (21 – 30), S4 16 (13 – 18), S5 11 (11 – 17), r3 30 (27 – 34), R1 14, all setae smooth except Z4 and Z5, st1-st1 60 (58 – 61), st2-st2 63 (61 – 65), st1-st3 62 (58 – 64), posterior margin of the sternal shield with a convex lobe, metapodal 1 length 17 (16 – 18), width 4, metapodal 2 length 7 (6 – 8), width 1, ventrianal shield 98 (94 – 104) long, 58 (55 – 58) wide at level of anterior corners and 57 (55 – 58) wide at level of anus, 4 poroids around the ventrianal shield and genital shield, JV5 41, spermatheca 16 (13 – 22) long and 5 width, SgeI 31 (30 – 32), SgeII 26, SgeIII 27 (25 – 29), StiIII 19, SgeIV 59 (56 – 63), StiIV 31 (29 – 32), StIV 63 (61 – 67), macrosetae whip-like, fixed digit of chelicerae 32 (29 – 33), mobile digit of chelicerae 31, 12 denticles on fixed digit and 5 on mobile digit. The measurements of the single adult male presently found are as follow: dorsal shield sclerotized and reticulated, dorsal shield length 245, width 162, 2 solenostome (gd 1, 2, 6, 8 and 9), j1 15, j3 25, j4 7, j5 7, j6 9, J2 10, J5 8, z2 10, z4 12, z5 7, Z1 9, Z4 38, Z5 58, s4 32, S2 14, S4 12, S5 9, r3 13, R1 8, all setae smooth except Z4 and Z5, st1-st1 49, st2-st2 55, st1-st3 59, st1-st5 103., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 292-293, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Denmark H. A., Muma M. H. 1975 - The Phytoseiidae (Acarina: Mesostigmata) of Puerto Rico - J. Agric. Univ. Puerto Rico 59, 279 - 304.","Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494.","Tixier M. - S. 2012 - Approaches to assess intraspecific variations of morphological continuous characters: the case study of the family Phytoseiidae (Acari: Mesostigmata) - Cladistics sous presse, doi: 10.1111 / j. 1096 - 0031.2012.00394. x"]}

Typhlodromalus peregrinus (Muma) Typhlodromus peregrinus Muma, 1955: 270; Typhlodromalus peregrinus Muma and Denmark, 1970: 88; Moraes et al., 1986: 132, 2004: 202; Zacarias and Moraes, 2001: 582; Typhlodromus (Amblyseius) peregrinus Chant, 1959: 97; Amblyseius peregrinus McMurtry, 1983: 255. Moraes et al., 1991: 130; Typhlodromus (Amblyseius) robineae Chant, 1959: 98; Typhlodromus (Amblyseius) evansi Chant, 1959: 99; Typhlodromus (Amblyseius) primulae Chant, 1959: 99 (synonymy, according to Muma, 1964) This species is very common on citrus (Muma, 1955, 1967; Peaea, 1992; Childers, 1994; Villanueva and Childers, 2004, 2005; Fadamiro et al., 2008, 2009) and solanaceous plants (McMurtry, 1983; Fiaboe et al., 2007) in several countries and is very often reported as the most abundant species. Typhlodromalus peregrinus can be found at the underside of mature citrus leaves, inside tree canopy, under empty scale armor, clump and dead scale insects whitefly exuvia, sooty mold and mine of Phyllocnistis citrella Stainton (Muma, 1967; Childers, 1994; Villanueva and Childers, 2011). Muma (1969) reported that T. peregrinus was able to reproduce and develop on Panonychus citri (McGregor) but did perform better on eggs and crawlers of chaff scale, Parlatoria pergandii Comstock, and six-spotted spider mite, Eotetranychus sexmaculatus (Riley). This phytoseiid was also reported to feed on Phyllocoptruta oleivora (Ashmead), providing some degree of rust mite suppression on lime (Peaea, 1992). Thus, T. peregrinus seems to be a generalist species with the ability to reproduce and develop on the two key pests on Guadeloupe and Martinique citrus, P. citri and P. oleivora and probably several occasional pests. Its optimal preys were evaluated as Aleyrodidae, Coccidae, and Tetranychidae by Muma (1971). Fouly et al. (1995) have studied the biology of T. peregrinus in the laboratory at 26°C. Each of the following organisms was evaluated as suitable diet: all stages of T. urticae; immature stages of P. citri; and pollens of Malephora crocea, Quercus virginiana, and Typha latifolia. The combination of T. urticae with pollen was also tested. Total developmental time ranged between 5.73 and 7 days for females and between 5.67 and 6.93 days for males. The percentage of females in the total population ranged between 53 and 61 %. A diet of T. urticae provided the shortest generation time (T), greatest female longevity, and mean total fecundity (F) which resulted in the highest net reproductive rate (Ro) value (25.31 expected females per female), intrinsic rate of increase (rm = 0.224), and finite rate of increase (λ = 1.25) per day for T. peregrinus. Diets of only P. citri or M. crocea resulted in close values of T = 8.67, 8.91; F = 47.11, 49.47; R 0 = 24.00, 26.65; rm = 0.210, 0.219; and λ = 1.23, 1.24, respectively. Quercus virginiana and T. latifolia were the less favorable food sources, with results of T = 8.78, 9.41; F = 30.38, 24.25; R 0 = 14.20, 12.04; rm = 0.193, 0.170, and λ = 1.21, 1.18, respectively. The occurrence of high densities of this species on ground cover vegetation (weeds) is explained in Alabama citrus orchard (Fadamiro et al., 2008, 2009) by the possibility that grasses may serve as overwintering sites and alternative food sources, which is probably the most important factors in French West Indies citrus orchards as there is no overwintering in citrus crop in this tropical area. Typhlodromalus peregrinus was collected from 64 ground cover plants or vines in Florida citrus fields (Childers and Denmark, 2011) with highest numbers found on the following plants: Bidens alba, Solanum americanum (which is one plant of the ground cover on which T. peregrinus was collected in Guadeloupe), Amaranthus spinosus, Gnaphalium pensylvanicum, Lantana camara and Chenopodium ambrosioides). In Florida, the highest numbers of T. peregrinus in ground cover corresponded with peaks in thrips numbers, suggesting possible predation on one or more species of thrips occurring. Childers and Denmark (2011) suggest that this species should therefore be evaluated as a predator of thrips larvae and/or adults. Significant increases in numbers of T. peregrinus were also correlated with increased levels of several pollen species on citrus leaves (Villanueva and Childers, 2004). Thus, considering all these elements, it is possible that T. peregrinus may constitute a key species in citrus orchards in French West Indies. Previous Records — Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Hawaii, Honduras, Mexico, Nicaragua, Puerto Rico, Suriname, USA (Florida, Alabama, Missouri), Venezuela (Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Vieux-Habitants, Station Le Bouchu du CIRAD, 16°03’N, 61°45’W, alt. 21 m, 9 ♀ and 7 ♂ extracted froom a mixture of various herbaceous plants collected in an experimental citrus crop (Achyranthhes aspera, Centrosema sp., Chamaescye hypericifolia, Chloris inflata, Croton lobatus, Echinochloa colona, Ipomea hederifolia, Macroptilium lathyroides, Merremia umbellata, Neonotonia wightii, Rhynchosia minima, Solanum americanum, Tridax procumbens, Urena lobata, Vernonia cinerea), Mailloux coll., April to December 2008. Remarks — this is the first record of this species in the West Indies. The measurements and description of the specimens collected fit very well those given by Moraes and Mesa (1988). The average measurements of the nine adult females presently found are: dorsal shield strongy reticulated on the whole dorsum, 344 (329 – 350) long and 219 (184 – 285) wide, with 5 solenostomes (gd1, 2, 6, 8 and 9), 10 pairs of poroids, 17 pairs of dorsal setae and 2 pairs of sub-lateral setae: j1 25 (22 – 29), j3 37 (34 – 42), j4 15 (11 – 20), j5 17 (14 – 20), j6 20 (16 – 27), J2 20 (17 – 23), J5 8 (6 – 11), z2 21 (20 – 25), z4 33 (28 – 41), z5 17 (11 – 20), Z1 32 (29 – 35), Z4 48 (47 – 51), Z5 59 (57 – 61), s4 45 (38 – 50), S2 31 (28 – 34), S4 25 (20 – 30), S5 13 (10 – 19), r3 18 (12 – 22), R1 17 (13 – 20). All setae smooth, except Z4 and Z5 which are moderately serrated. Peritreme extending to the level of j1. Ventral shields smooth. Sternal shield large, with 3 pairs of setae and 2 pairs of pores; 1 pair (st4) out of the sternal shield, on a small metasternal shield; posterior margin with a convex lobe overlapping the genital shield. Distances between st1-st3 66 (63 – 71), st2-st2 63 (60 – 65), st5- st5 76 (73 – 78). Two pairs of metapodal shields 16 (14 – 18) long, 4 (3 – 6) wide for the largest, 5 – 8 long and very thin for the smallest one. Ventrianal shield with 3 pairs of pre-anal setae, JV1, JV2 and ZV2 and one pair of elliptical pre-anal pores. Membrane surrounding ventrianal shield with 4 pairs of setae ZV1, ZV3, JV4 and JV5 and 4 pairs of round to oblong poroids; ventrianal shield 93 (88 – 103) long, 64 (60 – 68) wide at level of anterior corners and 61 (57 – 74) wide at level of anus. JV5 42 (37 – 45) long and smooth. All legs with smooth macrosetae: SgeI 17 (13 – 23), SgeII 20 (18 – 25), SgeIII 30 (28 – 30), Sti- III 20 (15 – 25), SgeIV 42 (34 – 44), StiIV 3 (18 – 25), StIV 62 (58 – 65). All macrosetae are whip-like with a small bulbous tip. Chaetotactic formula of genu II: 2-2/0, 2/0-1; genu III: 1-2/1, 2/0-1. Length of legs I: 342 (278 – 369), II: 253 (226 – 282), III: 251 (214 – 286), IV: 341 (275 – 388). Chelicerae with fixed digit 32 (30 – 36) with 7 teeth and movable digit 30 (28 – 32) with 3-4 teeth. Calyx of spermatheca fundibular (Denmark et al. 1999) with the cervix elongate, 5 (3 – 7) wide and 16 (9 – 22) long, with a big atrium at the basis. The average measurements of the seven adult males presently found are: dorsal shield like in the female, 256 (243 – 275) long and 143 (134 – 149) wide. Setae j1 19 (18 – 20), j3 29 (25 – 30), j4 11 (7 – 13), j5 12 (8 – 16), j6 14 (12 – 16), J2 11 (7 – 16), J5 7 (5 – 8), z2 16 (13 – 17), z4 27 (25 – 29), z5 11 (10 – 12), Z1 18 (16 – 22), Z4 34 (31 – 37), Z5 39 (34 – 42), s4 35 (33 – 36), S2 17 (16 – 18), S4 14 (12 – 16), S5 9 (7 – 12), r3 17 (15 – 19), Rl 13 (12 – 16). Peritreme extending to the level of j1. All venter shields smooth. Sternogenital shield with 5 pairs of setae and 2 pairs of pores. Distances between st1- st3 58 (56 – 59), st2-st2 54 (50 – 56), st5-st5 37 (32 – 40). Ventrianal shield not fused with peritremal shields with 3 pairs of pre-anal setae, JV1, JV2 and ZV2, one pair of elliptical preanal solenostomes and 2 pairs of poroids. Membrane surrounding ventrianal shield with one pair of setae JV 5 in level with anal opening, and no visible pair of oblong poroids; ventrianal shield 93 (85 – 100) long, 132 (127 – 146) wide at level of anterior corners and 59 (54 – 65) at level of anus. Macrosetae on all legs: SgeI 15 (13 – 18), SgeII 16 (15 – 20), SgeIII 22 (20 – 25), StiIII 15, SgeIV 31 (27 – 36), StiIV 17 (15 – 21), StIV 46 (40 – 51). Chaetotactic formula of genu II and genu III are identical to the female. Length of leg I: 250 (228 – 276), II: 202 (182 – 228), III: 218 (193 – 253), IV: 304 (275 – 326). Spermatodactyl of cheliecerae with a large toe terminating the foot, shaft 19 (18 – 20) long., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 294-295, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Muma M. H. 1955 - Phytoseiidae (Acarina) associated with citrus in Florida - Ann. Entomol. Soc. Am. 48: 262 - 272.","Muma M. H., Denmark H. A. 1970 - Phytoseiidae of Florida - Arthropods of Florida and neighboring land areas, 6. Florida Department of Agriculture and Consumer Services, Division of Plant Industry, Gainesville, 150 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat - Embrapa ed. and Pub., Brasilia, 353 pp + VII.","Zacarias M. S., Moraes G. J. de, 2001 - Phytoseiid mites (Acari) associated with rubber trees and other euphorbiaceous plants in southeastern Brazil - Noetrop. Entomol. 30: 579 - 586.","Chant D. A. 1959 - Phytoseiid mites. Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of 38 new species - Can. Entomol. 91, suppl. 12: 1 - 166.","McMurtry J. A. 1983 - Phytoseiid mites from Guatemala, with descriptions of two new species and redefinitions of the genera Euseius, Typhloseiopsis, and the Typhlodromus occidentalis species group (Acari: Mesostigmata) - Intern. J. Entomol. 25 (4): 249 - 272.","Moraes G. J. de, Mesa N. C., Braun A. 1991 - Some phytoseiid mites of Latin America (Acari: Phytoseiidae) - Intern. J. Acarol. 17: 117 - 139.","Muma M. H. 1964 - Annoted list and keys to Phytoseiidae (Acarina: Mesostigmata) associated with Florida citrus - Univ. Fla. Agric. Exp. Sta. Bull. 685: 1 - 42.","Muma M. H. 1967 - Typhlodromalus peregrinus (Muma) (Acari: Phytoseiidae) on Florida citrus - Proceedings, 2 nd International Congress of Acarology. Sutton Bonington, 19 - 25 July 1967, England. Akade'miai Kiado', Budapest, Hungary: 135 - 148.","Peaea J. F. 1992 - Predator-prey interactions between Typhlodromalus peregrinus and Polyphagotarsonemus latus: effects of alternative prey and other food resources - Fla. Entomol. 75: 241 - 248. doi: 10.2307 / 3495626","Childers C. C. 1994 - Biological control of phytophagous mites on Florida citrus utilizing predatory arthropods - D. Rosen, F. Bennet, and J. Capinera [eds.], Pest management in the subtropics: biological control Florida perspective. Intercept, Andover, United Kingdom: 255 - 288.","Villanueva R. T., Childers C. C. 2004 - Phytoseiidae increase with pollen deposition on citrus leaves - Fla. Entomol. 4: 609 - 611. doi: 10.1653 / 0015 - 4040 (2004) 087 [0609: PIWPDO] 2.0. CO; 2","Villanueva R. T., Childers C. C. 2005 - Diurnal and spatial patterns of Phytoseiidae in the citrus canopy - Exp. Appl. Acarol 35 (4): 269 - 280. doi: 10.1007 / s 10493 - 004 - 5728 - 4","Fadamiro H. Y., Xiao Y., Hargroder T., Nesbitt M., Umeh V., Childers C. C. 2008 - Seasonal occurrence of key arthropod pests and associated natural enemies in Alabama satsuma citrus - Environ. Entomol. 2: 555 - 567. doi: 10.1603 / 0046 - 225 X (2008) 37 [555: SOOKAP] 2.0. CO; 2","Fadamiro H. Y., Xiao Y., Hargroder T., Nesbitt M., Childers C. C. 2009 - Diversity and seasonal abun- dance of predacious mites in Alabama Satsuma citrus - Ann. Entomol. Soc. Am. 102 (4): 617 - 628.","Fiaboe K. K. M., Gondim M. G. C. Jr., Moraes G. J. de, Ogol C. K. P. O., Knapp. M. 2007 - Surveys for natural enemies of the tomato red spider mite Tetranychus evansi (Acari: Tetranychidae) in the northeastern and southeastern Brazil - Zootaxa 1395: 33 - 58.","Villanueva R. T., Childers C. C. 2011 - Mine-damaged leaves by Phyllocnistis citrella Stainton provide refuge for phytoseiids on grapefruit in Florida and Texas - Zoosymposia 6: 118 - 123.","Muma M. H. 1969 - Biological control of various insects and mites on Florida citrus - Proceedings, 1 st International Citrus Symposium, 16 - 26 March 1969, Riverside, CA. University of California, Riverside: 863 - 870.","Muma M. H. 1971 - Food habits of Phytoseiidae (Acarina: Mesostigmata) including common species on Florida citrus - Fla Entomol. 54 (1): 21 - 34.","Fouly A. H., Abou-Setta M. M., Childers C. C. 1995 - Effects of diets on the biology and life tables of Typhlodromalus peregrines - Environ. Entomol. 24: 870 - 877.","Childers C. C., Denmark H. A. 2011 - Phytoseiidae (Acari: Mesostigmata) within citrus orchards in Florida: species distribution, relative and seasonal abundance within trees, associated vines and ground cover plants - Exp. Appl. Acarol. 54: 331 - 371.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494.","Moraes G. J. de, Mesa N. C. 1988 - Mites of the family Phytoseiidae (Acari) in Colombia, with descriptions ofthree new species - Int. J. Acarol. 14: 71 - 88.","Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp."]}

Transeius rufus (Denmark and Evans) Neoseiulus rufus Denmark and Evans, in Denmark et al., 1999: 73. The biology of this species found only once in Honduras by Denmark and Evans (in Denmark et al., 1999) on the giant thatching grass Hyparrhenia rufa (Poaceae), a hairy plant, is unknown. Previous Records — Honduras (Denmark et al., 1999). Specimens examined — Martinique, Montagne PelØe, Parking, 14°48’N, 61°09’W, alt. 825 m, 2 ♀ on Urena lobata, Kreiter coll., 21 November 2010. Remarks — this is the first record of this species in the West Indies. The measurements of the two specimens collected fit very well the measurements given by Denmark et al. (1999), except setae r3 and R1 (30 µm in our two specimens instead of 16 in Denmark et al. (1999) on a single specimen). The average measurements of the two adult females presently found are as follow: dorsal shield sclerotized, five solenostomes (gd 1, 2, 4, 8 and 9), j1 23 – 24, j3 27 – 35, j4 9 – 10, j5 9, j6 9 – 10, J2 10 – 14, J5 8 – 10, z2 11 – 15, z4 20 – 30, z5 8 – 9, Z1 11 – 15, Z4 63, Z5 71 – 75, s4 47 – 50, S2 24 – 27, S4 12 – 14, S5 13 – 14, r3 16, R1 16, Z4 and Z5 serrated, st1-st1 53 – 59, st2-st2 62 – 65, st1-st3 60 – 62, posterior margin of the sternal shield concave, metapodal 1 length 23 – 28, width 3, metapodal 2 length 10 – 13, width 1, ventrianal shield 98 – 115 long, 90 – 92 wide at level of anterior corners and 80-88 wide at level of anus, 5 poroids around the ventri-aan shield and the genital shield, JV5 55 – 60, spermatheca 19 – 20 long and 5 – 7 width, SgeIV 44 – 45, StiIV 18 – 25, StIV 47 – 50, macrosetae all whip-like, fixed digit of chelicerae 30 with 7 denticles, mobile digit of chelicerae 32 without denticle., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 289-290, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp."]}

Transeius mariae-angeae Kreiter n.sp. Description Adult Female (Figs. 1-3) (n = 2) Dorsum (Fig. 1) — Dorsal shield 305 – 313 long and 205 – 215 wide, strongly reticulated on the whole dorsum, with 5 solenostomes (gd1, 2, 6, 8 and 9), 9 pairs of poroids, 17 pairs of dorsal setae and 2 pairs of sub-lateral setae: j1 20 – 23, j3 36 – 38, j4 15 – 18, j5 16 – 19, j6 15 – 18, J2 13 – 14, J5 8 – 9, z2 33 – 26, z4 29 – 33, z5 9, Z1 16 – 18, Z4 54 – 55, Z5 74 – 75, s4 58 – 60, S2 28 – 30, S4 14 – 15, S5 14 – 16, r3 23 – 25, R1 21. All setae smooth except Z4 and Z5 which are moderately serrated. Peritreme (Fig. 1) — Extending to the level of j1. Venter (Fig. 2) — Sternal shield smooth. Other shields smooth. Sternal shield not very large, with 3 pairs of setae and 2 pairs of pores; 1 pair (st4) out of the sternal shield, on a small metasternal shield; posterior margin straight. Distances between st1- st3 55, st2-st2 64 – 65, st5-st5 55 – 56. Two pairs of metapodal shields 15 long and 1 – 3 wide for the largest, 6 – 8 long and very thin for the smallest one. Ventrianal shield with 3 pairs of pre-anal setae, JV1, JV2 and ZV2 and one pair of large elliptical pre-anal solenostomes. Membrane surrounding ventrianal shield with 4 pairs of setae ZV1, ZV3, JV4 and JV5 and 4 pairs of poroids; ventrianal shield 90 – 98 long, 63 – 64 wide at level of anterior corners and 58 – 60 wide at level of anus. JV5 43 – 50 long and smooth. Legs — Macrosetae on all legs: SgeI 16 – 18, SgeII 18, SgeIII 15 – 18, StiIII 23 – 25, SgeIV 36 – 40, StiIV 19 – 23, StIV 38. All macrosetae whip-like. Chaetotactic formula of genu II: 2-2/0, 2/0-1; genu III: 1-2/1, 2/0-1. Length of legs I: 310 – 318, II: 223 – 238, III: 235 – 245, IV: 320 – 325. Chelicera — Fixed digit 30 – 31 with 9 – 10 teeth and movable digit 33 – 34 with four teeth. Spermatheca (Fig. 3) — Calyx of spermatheca pocular (Denmark et al. 1999) with the cervix elongate, 4 – 6 wide and 6 – 8 long, with a big atrium at the basis and a visible long ductus minor on the paratype female. Adult Male: unknown. Holotype — 1 female (on one preparation), La DØsirade, Parc Eolien, 16°11’N, 61°39’W, alt. 184 m, on Coccoloba pubescens, Kreiter coll., 5 January 2008, deposited in the Montpellier SupAgro Acarology Collection in UMR CBGP. Paratype — 1 female (on a separate preparation), same location deposited in the same collection. Etymology — the name of the species refers to the first name of the first author’s wife of this paper and describer of this new species, Marie-Ange Burgell, to whom this species is dedicated. Diagnosis — the two specimens of Transeius mariae-angeae n. sp. found are close to Transeius bellotti (Moraes and Mesa) but differ in having: the shape of spermatheca very different (both pocular sensu denmark et al. (1999) but with an open cervix and with a small atrium at the basis of the cervix for T. bellotti and a more closed cervix, a strong large atrium and swollen edges of the cervix for T. mariaeangeae n. sp.); the presence of macrosetae on all legs and not only of leg IV; the setae length longer in the new species except Z4, S4, SgeIV and StIV; the dorsum smooth and not imbricate; the setae j3 and s4 not serrated and the number of teeth on movable/fixed digits = 9-10/4 (against 9/ 3 in T. bellotti). It resembles also to T. rufus (Denmark and Evans) but differs in having: several setae much longer except for j1, Z4, SgeIV and StIV which are longer in T. rufus; the dorsal shield not reticulated in the new species and slightly reticulated in T. rufus; the number of teeth on movable/fixed digits of chelicerae = 9-10/4 against 7/0. It is also close to T. sanblasensis (De Leon) but setae are longer in the new species except for some (j1, z4, s4, r3, S2) which are very much longer and all macrosetae of the four legs are smaller in the new species. It resembles also to T. aciculus (De Leon) but differs in having j4 shorter (15-18 instead of 36) and S5 longer (14-16 instead of 9); and to T. cristobalensis (De Leon) but differs in having a shorter StIV (38 instead of 88).

Neoseiulus paspalivorus (De Leon) Typhlodromus paspalivorus De Leon, 1957: 143 Neoseiulus paspalivorus Muma and Denmark, 1970: 110; Moraes et al., 2000: 248. Amblyseius paspalivorus Schicha, 1981: 210. The biology of this species was only recently studied. It seems to be common on various herbaceous plants (Moraes et al., 1986) and could be a Gondwanian species because of its currently known area of distribution: Caribbean, India, Oriental region and Africa (Ueckermann and Lawson- Balagbo, pers. comm.). This species predominates in the dry areas of states of CearAE and Pernambuco in Brazil on coconuts (Lawson-Balagbo et al., 2008a). This species thrived on the coconut eriophyid, Aceria guerreronis Keifer as primary food source resulting in shorter developmental time, higher ovipositon rate and higher intrinsic rate of increase than on any other diet (Lawson-Balagbo et al., 2007). Neoseiulus paspalivorus is dorso-ventrally flattened giving it an advantage in accessing the area under the bracts (Lawson-Balagbo et al., 2008b). This species is thus cited as a promising candidate for the biological control of the coconut eriophyid (Lawson- Balagbo et al., 2008a). This latter pest was recently introduced in Sri Lanka and southern India where it is causing considerable damage to coconut. Neoseiulus paspalivorus was found only on coconut in the area of production and on fruits, in association with A. guerreronis (Fernando et al., 2003; Moraes et al., 2004a). Its presence on coconut in French West Indies contaminated by A. guerreronis is thus not surprising. The specimens collected were found very close from coconuts. Previous Records — Guadeloupe (Moraes et al., 2000), India and Sri Lanka (Fernando et al., 2003; Moraes et al., 2004a), Jamaica, Philippines, USA (Florida) (Moraes et al., 2004b), Cuba (Cabrera et al., 2008), USA Florida (Muma and Denmark, 1970), Brazil (Lawson-Balagbo et al., 2008a). Specimens examined — Guadeloupe, Basse- Terre, Vieux-Habitants, Station Le Bouchu du CIRAD, 16°03’N, 61°45’W, alt. 21 m, 2 ♀ collected on various herbaceous plants covering the soil of an experimental citrus crop (Alysicarpus vaginalis, Chloris inflata, Cleome rutidosperma, Dicanthium annulatum, Echinochloa colona, Vernonia cinerea), Mailloux coll., April to December 2008. Remarks — the measurements and description of the specimens collected fit very well those given by Moraes et al. (2000)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 298-299, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["De Leon D. 1957 - Three new Typhlodromus from southern Florida - Fla Entomol. 40: 141 - 144.","Muma M. H., Denmark H. A. 1970 - Phytoseiidae of Florida - Arthropods of Florida and neighboring land areas, 6. Florida Department of Agriculture and Consumer Services, Division of Plant Industry, Gainesville, 150 pp.","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Schicha E. 1981 - A new species of Amblyseius from Australia compared with ten closely related species from Asia, America and Africa - Intern. J. Acarol. 7: 203 - 216. doi: 10.1080 / 01647958108683262","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat - Embrapa ed. and Pub., Brasilia, 353 pp + VII.","Lawson-Balagbo L. M., Gondim Jr. M. G. C., Moraes G. J. de, Hanna R., Schausberger P. 2008 a - Exploration of the acarine fauna on coconut palm in Brazil with emphasis on Aceria guerreronis (Acari: Eriophyidae) and its natural enemies - Bull. Entomol. Res. 98: 83 - 96. doi: 10.1017 / S 0007485307005421","Lawson-Balagbo L. M., Gondim Jr. M. G. C., Moraes G. J. de, Hanna R., Schausberger P. 2007 - Life history of the predatory mites Neoseiulus paspalivorus and Proctolaelaps bickleyi, candidates for biological control of Aceria guerreronis - Exp. Appl. Acarol. 43: 49 - 61.","Lawson-Balagbo L. M., Gondim Jr. M. G. C., Moraes G. J. de, Hanna R., Schausberger P. 2008 b - Compatibility of Neoseiulus paspalivorus and Proctolaelaps bickleyi, candidate biocontrol agents of the coconut mite Aceria guerreronis: spatial niche use and intraguild predation - Exp. Appl. Acarol. 45: 1 - 13. doi: 10.1007 / s 10493 - 008 - 9156 - 8","Fernando L. C. P., Aratchige N. S., Peiris T. S. G. 2003 - Distribution patterns of cocounut mite, Aceria guerreronis, and its predator Neoseiulus aff. paspalivorus in coconut palms - Exp. Appl. Acarol. 31: 71 - 78.","Moraes G. J. de, Lopes P. C., Fernando L. C. P. 2004 a - Phytoseiid mites (Acari: Phytoseiidae) of coconut growing areas in Sri Lanka, with descriptions of three new species - J. Acarol. Soc. Japan 13 (2): 141 - 160.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494.","Cabrera R. I., Cueto Rodriguez J. R., Otero Colina G. 2008 - Natural enemies of mite Aceria guerreronis Keifer (Acari: Eriophyidae) in Cuba and their perspectives for pest management - Fitosanidad 12 (2): 99 - 107."]}

Transeius mariae-angeae Kreiter n.sp. Description Adult Female (Figs. 1-3) (n = 2) Dorsum (Fig. 1) — Dorsal shield 305 – 313 long and 205 – 215 wide, strongly reticulated on the whole dorsum, with 5 solenostomes (gd1, 2, 6, 8 and 9), 9 pairs of poroids, 17 pairs of dorsal setae and 2 pairs of sub-lateral setae: j1 20 – 23, j3 36 – 38, j4 15 – 18, j5 16 – 19, j6 15 – 18, J2 13 – 14, J5 8 – 9, z2 33 – 26, z4 29 – 33, z5 9, Z1 16 – 18, Z4 54 – 55, Z5 74 – 75, s4 58 – 60, S2 28 – 30, S4 14 – 15, S5 14 – 16, r3 23 – 25, R1 21. All setae smooth except Z4 and Z5 which are moderately serrated. Peritreme (Fig. 1) — Extending to the level of j1. Venter (Fig. 2) — Sternal shield smooth. Other shields smooth. Sternal shield not very large, with 3 pairs of setae and 2 pairs of pores; 1 pair (st4) out of the sternal shield, on a small metasternal shield; posterior margin straight. Distances between st1- st3 55, st2-st2 64 – 65, st5-st5 55 – 56. Two pairs of metapodal shields 15 long and 1 – 3 wide for the largest, 6 – 8 long and very thin for the smallest one. Ventrianal shield with 3 pairs of pre-anal setae, JV1, JV2 and ZV2 and one pair of large elliptical pre-anal solenostomes. Membrane surrounding ventrianal shield with 4 pairs of setae ZV1, ZV3, JV4 and JV5 and 4 pairs of poroids; ventrianal shield 90 – 98 long, 63 – 64 wide at level of anterior corners and 58 – 60 wide at level of anus. JV5 43 – 50 long and smooth. Legs — Macrosetae on all legs: SgeI 16 – 18, SgeII 18, SgeIII 15 – 18, StiIII 23 – 25, SgeIV 36 – 40, StiIV 19 – 23, StIV 38. All macrosetae whip-like. Chaetotactic formula of genu II: 2-2/0, 2/0-1; genu III: 1-2/1, 2/0-1. Length of legs I: 310 – 318, II: 223 – 238, III: 235 – 245, IV: 320 – 325. Chelicera — Fixed digit 30 – 31 with 9 – 10 teeth and movable digit 33 – 34 with four teeth. Spermatheca (Fig. 3) — Calyx of spermatheca pocular (Denmark et al. 1999) with the cervix elongate, 4 – 6 wide and 6 – 8 long, with a big atrium at the basis and a visible long ductus minor on the paratype female. Adult Male: unknown. Holotype — 1 female (on one preparation), La DØsirade, Parc Eolien, 16°11’N, 61°39’W, alt. 184 m, on Coccoloba pubescens, Kreiter coll., 5 January 2008, deposited in the Montpellier SupAgro Acarology Collection in UMR CBGP. Paratype — 1 female (on a separate preparation), same location deposited in the same collection. Etymology — the name of the species refers to the first name of the first author’s wife of this paper and describer of this new species, Marie-Ange Burgell, to whom this species is dedicated. Diagnosis — the two specimens of Transeius mariae-angeae n. sp. found are close to Transeius bellotti (Moraes and Mesa) but differ in having: the shape of spermatheca very different (both pocular sensu denmark et al. (1999) but with an open cervix and with a small atrium at the basis of the cervix for T. bellotti and a more closed cervix, a strong large atrium and swollen edges of the cervix for T. mariaeangeae n. sp.); the presence of macrosetae on all legs and not only of leg IV; the setae length longer in the new species except Z4, S4, SgeIV and StIV; the dorsum smooth and not imbricate; the setae j3 and s4 not serrated and the number of teeth on movable/fixed digits = 9-10/4 (against 9/ 3 in T. bellotti). It resembles also to T. rufus (Denmark and Evans) but differs in having: several setae much longer except for j1, Z4, SgeIV and StIV which are longer in T. rufus; the dorsal shield not reticulated in the new species and slightly reticulated in T. rufus; the number of teeth on movable/fixed digits of chelicerae = 9-10/4 against 7/0. It is also close to T. sanblasensis (De Leon) but setae are longer in the new species except for some (j1, z4, s4, r3, S2) which are very much longer and all macrosetae of the four legs are smaller in the new species. It resembles also to T. aciculus (De Leon) but differs in having j4 shorter (15-18 instead of 36) and S5 longer (14-16 instead of 9); and to T. cristobalensis (De Leon) but differs in having a shorter StIV (38 instead of 88)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 286-289, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp."]}

Neoseiulus longispinosus (Evans) Typhlodromus longispinosus Evans, 1952: 413; Evans, 1953: 465; Womersley, 1954: 177; Ehara, 1958: 55. Typhlodromus (Amblyseius) longispinosus, Chant, 1959: 74. Amblyseius longispinosus, Corpuz and Rimando, 1966: 129; Schicha, 1975: 103. Neoseiulus longispinosus, Moraes et al., 2000: 245. This species was already mentioned from Guadeloupe and other Islands of the French Antilles (Moraes et al., 2000) but only in very few localities on various host plants. It is distributed in many countries of the world, mainly in tropical areas. The biology of this species has been studied for pest control purposes including side effects of acaricides (Bin Ibrahim and Tan, 2000). The activity, feeding, development, predation, cannibalism, intra-guild predation and behaviour have been extensively studied by several authors (Schausberger and Croft, 1999a, b; Croft et al., 1999a, b; Schausberger and Croft, 2000 a, b; Blackwood et al., 2001). Previous Records — French Antilles (Guadeloupe, Les Saintes, Marie-Galante, Martinique, Saint-BarthØlØmy) (Moraes et al., 2000), Australia, China, Egypt, Hawaii, Hong-Kong, India, Indonesia, Japan, Malaysia, New Zealand, Pakistan, Papua New Guinea, Philippines, Russia, South Korea, Taïwan, Thailand (Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Vieux-Habitants, Station Le Bouchu du CIRAD, 16°03’N, 61°45’W, alt. 21 m, 14 ♀ and 1 ♂ on Neonotonia wightii and Tridax procumbens within an experimental citrus crop, Mailloux coll., April to December 2008; Basse-Terre, Petit-Bourg, Domaine Duclos de l’INRA Antilles-Guyane, 16°12’N, 61°39’W, alt. 85 m, 1 ♂ on Vigna sp., Kreiter coll., 21 December 2008. Remarks — the measurements and description of the specimens collected fit very well those given by Moraes et al. (2000)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 298, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Evans G. O. 1952 - On a new predatory mite of economic importance - Bull. Entomol. Res. 43: 397 - 401.","Evans G. O. 1953 - On some mites of the genus Typhlodromus Scheuten, 1857, from S. E. Asia - Ann. Mag. Nat. Hist. 6: 449 - 467. doi: 10.1080 / 00222935308654444","Womersley H. 1954 - Species of the subfamily Phytoseiinae (Acarina: Laelaptidae) from Australia - Austral. J. Zool. 2: 169 - 191. doi: 10.1071 / ZO 9540169","Ehara S. 1958 - Three predatory mites of the genus Typhlodromus from Japan (Phytoseiidae) - Annot. Zool. Japonenses 31: 53 - 57.","Chant D. A. 1959 - Phytoseiid mites. Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of 38 new species - Can. Entomol. 91, suppl. 12: 1 - 166.","Corpuz L. A., Rimando L. 1966 - Some Philippine Amblyseiinae (Phytoseiidae: Acarina) - Philippine Agriculture 50: 114 - 136.","Schicha E. 1975 - Predacious mites (Acarina: Phytoseiidae) on sprayed apple trees at Bathurst (N. S. W.) - J. Austral. Entomol. Soc. 14: 217 - 219.","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Bin Ibrahim Y., Tan S. Y. 2000 - Influence of sublethal exposure to abamectin on the biological performance of Neoseiulus longispinosus (Acari: Phytoseiidae) - J. Econ. Entomol. 93 (4): 1085 - 1089. doi: 10.1603 / 0022 - 0493 - 93.4.1085","Schausberger P., Croft B. A. 1999 a - Predation on and discrimination between con- and heterospecific eggs among specialist and generalist phytoseiid mites (Acari: Phytoseiidae) - Environ. Entomol. 28: 523 - 528.","Croft B. A., Luh H. - K., Schausberger P. 1999 a - Larval size relative to larval feeding, cannibalism of larvae, egg, or adult female size and larval-adult setal patterns among thirteen phytoseiid mite species - Exp. Appl. Acarol. 23: 599 - 610.","Schausberger P., Croft B. A. 2000 a - Nutritional benefits of intraguild predation and cannibalism among generalist and specialist phytoseiid mites - Ecol. Entomol. 25: 1 - 8. doi: 10.1046 / j. 1365 - 2311.2000.00284. x","Blackwood J. S., Schausberger P., Croft B. A. 2001 - Prey stage preferences in generalist and specialist phytoseiid mites (Acari: Phytoseiidae) when offered Tetranychus urticae (Acari: Tetranychidae) eggs and larvae - Environ. Entomol. 30: 1103 - 1111.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Amblyseius segregans De Leon Amblyseius segregans De Leon, 1966: 90; McMurtry, 1983: 252; Denmark and Muma, 1989: 126; Denmark et al., 1999: 30; Moraes et al., 2000: 241. The biology of this species found only four times in British Guyana (2 females), Guatemala and Honduras (2 females), Guadeloupe (1 female) and Martinique (1 female) on various plants (De Leon, 1966; McMurtry, 1983; Denmark and Muma, 1989; Moraes et al., 2000; Moraes et al., 2004b) remains unknown. Previous Records — British Guyana (De Leon, 1966), Guadeloupe and Martinique (Moraes et al., 2000), Honduras and Guatemala (Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, RiviŁre Corossol, Cascade des Ecrevisses, 16°11’N, 61°39’W, alt. 177 m, 4 ♀ on a small schrub, Kreiter coll., 26 Dec. 2008; Basse-Terre, 1 Łre Chute du Carbet, 16°03’N, 61°39’W, alt. 890 m, 2 ♀ on Miconia furfuracea, Kreiter coll., 30 December 2008; Grande-Terre, Porte d’Enfer, beach, 16°31’N, 61°28’W, alt. 2 m, 1 ♀ on Pluchea symphytifolia, Kreiter coll., 27 December 2008. Remarks — the measurements and description of the specimens collected fit very well those mentioned by Moraes et al. (2000)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 297, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["De Leon D. 1966 - Phytoseiidae of British Guyana with keys to species (Acarina: Mesostigmata) - Studies on the Fauna of Suriname and other Guyanas 8: 81 - 102.","McMurtry J. A. 1983 - Phytoseiid mites from Guatemala, with descriptions of two new species and redefinitions of the genera Euseius, Typhloseiopsis, and the Typhlodromus occidentalis species group (Acari: Mesostigmata) - Intern. J. Entomol. 25 (4): 249 - 272.","Denmark H. A., Muma M. H. 1989 - A revision of the genus Amblyseius Berlese, 1914 (Acari: Phytoseiidae) - Occasional Papers of the Florida State Collection of Arthropods, USA, 4, 149 pp.","Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp.","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Amblyseius sakalava Blommers Amblyseius sakalava Blommers, 1976: 96; suspected junior synonym of Amblyseius largoensis (according to Ueckermann and Loots, 1988). Biology — Amblyseius sakalava was first found in Madagascar on Corchorus trilocularis (Blommers, 1976) and then in a search for native natural enemies of Tetranychus urticae Koch in Sri-Lanka, collected on Manihot esculenta and identified by Dr Viebergen (Wijesekara, 2006). Its life history, reproductive performance on different foods and functional response to the prey were studied in the laboratory. At an average temperature of 29°C, when fed on pollen, A. sakalava completed its life cycle (egg to adult) in 4.5 days on average. The mean preovipositional period is 4.1 days and a female lays an average of 2.2 eggs per day when fed on T. urticae and 1.9 eggs per day when fed on pollen of Tridax procumbens (Asteraceae). The predatory mite reproduces equally when fed on this pollen and a mixture of T. procumbens pollen and the prey mite. But the reproductive performance was significantly lower when the predatory mite was fed on Tetranychus urticae Koch mite alone or pollen of Tithonia diversifolia or a mixture of T. diversifolia pollen and the latter prey mite. Study of functional response showed a typical type 2 response (Wijesekara, 2006). Previous Records — Madagascar (Blommers, 1976; Moraes et al., 2004b), Sri Lanka (Viebergen, in Wijesekara, 2006). Specimens examined — Guadeloupe, Basse- Terre, RiviŁre Corossol, Route des Mamelles, 16°10’N, 61°41’W, alt. 221 m, 1 ♀ on Syngonium podophyllum, Kreiter coll., 22 Dec. 2008. Remarks — this is the first record of this species in the West Indies. The measurements of the single specimen collected fit rather well the measurements given by Blommers (1976). This species was considered as a junior synonym of Amblyseius largoensis (Muma) by Ueckermann and Loots (1988). We fully disagree with this claimed synonymy as to our knowledge it was not demonstrated and as several characters are different between the two species: A. sakalava has 30 % longer measurements for several setae, especially Z5, nearly two times longer cervix of the spermatheca, at least 7 poroids on the dorsal shield to up to 15 compared to none to three maximum in A. largoensis, and the posterior limit of the sternal concave instead of straight in A. largoensis. However as we have collected only one specimen, all these differences have to be examined carefully on more individuals and the potential synonymy studied with modern analysis means. The measurements of the adult female presently found are as follow: dorsal shield smooth, dorsal shield length 365, width 258, 7 solenostomes (gd 1, 2, 3, 4, 5, 8 and 9), j1 35, j3 52, j4 5, j5 8, j6 8, J2 8, J5 13, z2 15, z4 13, z5 8, Z1 10, Z4 100, Z5 264, s4 93, S2 10, S4 8, S5 8, r3 10, R1 8, all setae smooth, st1- st1 65, st2-st2 70, st1-st3 68, posterior margin of the sternal shield straigth, metapodal 1 length 20, width 5, metapodal 2 length 15, width 1, ventrianal shield 108 long, 55 wide at level of anterior corners and 68 wide at level of anus, 4 poroids around the ventrianal shield and genital shield, JV5 63, spermatheca 33 long and 5 width, SgeI 45, SgeII 38, SgeIII 43, StiIII 40, SgeIV 118, StiIV 83, StIV 70, macrosetae whip-like, fixed digit of chelicerae 33, mobile digit of chelicerae 33, 11 denticles on fixed digit and 5 on mobile digit., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 290-291, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Blommers L. 1976 - Some Phytoseiidae (Acarina: Mesostigmata) from Madagascar, with descriptions of eight new species and notes on their biology - Bijdragen tot Dierkunde 46 (1): 80 - 106.","Ueckermann E. A., Loots G. C. 1988 - The African species of the subgenera Anthoseius De Leon and Amblyseius Berlese (Acari: Phytoseiidae) - Entomol. Mem. Depart. Agric. Water Supply, RSA 73: 1 - 168.","Wijesekara G. A. W. 2006 - Life history, reproductive performance and functional response of Amblyseius sakalava, a potential biocontrol agent of the twospotted spider mite - Cey. J. Sci. (Bio. Sci.) 35 (2): 137 - 140.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Typhlodromips amilus De Leon Typhlodromips amilus De Leon, 1967: 28, senior synonym of Typhlodromips bhoraii De Leon, in Denmark et al., 1999: 37-38. The biology of this species found only two times, in Trinidad Island on Cedrela sp. and on an unknown Bromeliaceae by De Leon (1967) and on Hevea brasiliensis by Ferla and Moraes (2002) is unknown. This species is mentioned and indicated as a senior synonym of T. bhoraii De Leon in the catalogue of Denmark et al. (1999). Previous Records — Brazil, Trinidad (De Leon, 1967; Denmark et al., 1999; Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Petit-Bourg, Domaine Duclos de l’INRA Antilles-Guyane, 16°12’N, 61°39’W, alt. 85 m, 1 ♀ on Spathodea campanulata, Kreiter coll., 3 January 2009. Remarks — this is the first record of this species in the West Indies. The measurements of the single specimen collected fit rather well the measurements given by De Leon (1967) and by Denmark et al. (1999). All setae are however slightly shorter, between 1 and 5 µm, which represent less than 10 % of variation, which is less than the intraspecific variation of 20 % around the mean defined by Tixier (2012). In addition, Denmark et al. (1999) mentioned that T. amilus is a species submitted to variation in lengths of setae. The measurements of the single adult female presently found are as follow: dorsal shield reticulated, dorsal shield length 355, width 183, six solenostomes (gd 1, 2, 3, 5, 8 and 9), j1 18, j3 20, j4 10, j5 13, j6 10, J2 13, J5 8, z2 15, z4 15, z5 11, Z1 13, Z4 33, Z5 53, s4 20, S2 16, S4 15, S5 10, r3 10, R1 13, Z4 and Z5 serrated, st1-st1 52, st2-st2 63, st1-st3 58, posterior margin of the sternal shield straight, metapodal 1 length 15, width 5, metapodal 2 length 10, width 2, ventrianal shield 100 long, 90 wide at level of anterior corners and 80 wide at level of anus, 3 poroids around the ventrianal shield, JV5 28, spermatheca 18 long and 8 width, SgeI 15, SgeII 13, SgeIII 13, StiIII 20, SgeIV 28, StiIV 13, StIV 30, macrosetae all knobbed, fixed digit of chelicerae 28, mobile digit of chelicerae 28, 8-9 denticles on fixed digit and 3 on mobile digit., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 296, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["De Leon D. 1967 - Some mites of the Caribbean Area. Part I. Acarina on Plants in Trinidad, West Indies - Allen Press Inc., Lawrence, Kansas, 66 pp.","Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp.","Ferla N. J., Moraes G. J. de, 2002 - Acaros predadores (Acari) em plantas nativas e cultivadas do Estado do Rio Grannde do Sul, Brasil - Rev. Brasil. Zool. 19 (4): 1001 - 1031.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494.","Tixier M. - S. 2012 - Approaches to assess intraspecific variations of morphological continuous characters: the case study of the family Phytoseiidae (Acari: Mesostigmata) - Cladistics sous presse, doi: 10.1111 / j. 1096 - 0031.2012.00394. x"]}

Euseius ovaloides (Blommers) Amblyseius (Amblyseius) ovaloides Blommers, 1974: 147. Amblyseius ovaloides Schicha and McMurtry, 1986: 177; Gutierrez and Etienne, 1986: 88. Euseius ovaloides Quilici et al., 1997: 284; Moraes et al., 2000; Quilici et al., 2000: 100; 242; Moraes et al., 2001: 43. The biology of this species found only a few times in Madagascar (Blommers, 1974), Papua-New Guinea (Schicha and Gutierrez 1985), Seychelles (Schicha, 1987), Reunion Island, (Quilici et al., 1997, 2000), Guadeloupe, Martinique and Marie-Galante (Moraes et al., 2000; Kreiter et al., 2006) on various plants remains unknown. It was suspected to be a poorly active predator of tetranychid mites (Gutierrez and Etienne, 1986). Previous Records — French Antilles (Guadeloupe and Marie-Galante) (Moraes et al., 2000), Madagascar, Papua New Guinea, Reunion Island (Kreiter et al., 2000), Seychelles (Moraes et al., 2004b). Specimens examined — Guadeloupe, Basse- Terre, Vieux-Habitants, Station Le Bouchu du CIRAD, 16°03’N, 61°45’W, alt. 21 m, 1 ♀ on Spondias monbim, Kreiter coll., 19 June 2008; Grande-Terre, Vieux Bourg, Port, 16°21’N, 61°31’W, alt. 2m, 1 ♀ on Carica papaya, Kreiter coll., 29 december 2008; Basse- Terre, Viard, beach, 16°10’N, 61°35’W, alt. 4 m, 1 ♀ on Ricinus communis, Kreiter and Rault coll., 1 January 2009; Martinique, Saint-Esprit, Quartier RØgal, 14°32’N, 60°58’W, alt. 112 m, 1 ♀ on Spondias dulcis, Kreiter coll., 15 November 2010; Le Marin, 14°28’N, 60°51’W, alt. 112 m, 3 ♀ on Annona muricata, Kreiter coll., 16 November 2010; Le Lorrain, Morne DØgras, 14°48’N, 61°05’W, alt. 355 m, 2 ♀ on Carica papaya, Kreiter coll., 22 November 2010. Remarks — the measurements and description of the specimens collected fit very well those given by Moraes et al. (2000)., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on pages 297-298, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Blommers L. 1974 - Species of the genus Amblyseius Berlese, 1914, from Tamatave, east Madagascar (Acarina: Phytoseiidae) - Bull. Zool. Museum Univ. van Amsterdam 3: 143 - 155.","Schicha E., McMurtry J. A. 1986 - Two new and two known species of Typhlodromus Scheuten (Acari: Phytoseiidae) from Australia - J. Austral. Entomol. Soc. 25: 177 - 183. doi: 10.1111 / j. 1440 - 6055.1986. tb 01100. x","Gutierrez J., Etienne J. 1986 - Les Tetranychidae de La ROunion et quelques-uns de leurs prOdateurs - Agronomie trop. 41 (1): 84 - 91.","Quilici S., Kreiter S., Ueckermann E. A., Vincenot D. 1997 - Predatory mites (Acari) from various crops on ROunion Island - Intern. J. Acarol. 23: 283 - 291.","Moraes G. J. de, Kreiter S., Lofego A. C. 2000 - Plant mites of the French Antilles. 3. Phytoseiidae - Acarologia 40: 237 - 264.","Quilici S., Ueckermann E. A., Kreiter S., VayssiLres J. - F. 2000 - Phytoseiidae (Acari) of ROunion Island - Acarologia 41: 95 - 106.","Moraes G. J. de, Ueckermann E. A., Oliveira A. R., Yaninek J. S. 2001 - Phytoseiidae mites of the genus Euseius (Acari: Phytoseiidae) from Sub-Saharan Africa - Zootaxa 3: 1 - 70.","Schicha E., Gutierrez J. 1985 - Phytoseiidae of Papua New Guinea, with three new species, and new records of Tetranychidae (Acari) - Intern. J. Acarol. 11: 173 - 181. doi: 10.1080 / 01647958508683412","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494."]}

Amblyseius fernandezi Chant and Baker Amblyseius fernandezi Chant and Baker, 1965: 19. The biology of this species, found in Central America, is unknown. Previous Records — Honduras, Nicaragua and Costa Rica (Chant and Baker, 1965; Denmark et al., 1999; Moraes et al., 2004b; Castro et al., 2010), El Salvador, Mexico and Venezuela (Castro et al., 2010). Specimens examined — Guadeloupe, Basse- Terre, RiviŁre Corossol, Cascade des Ecrevisses, 16°11’N, 61°39’W, alt. 177 m, 1 ♀ on an unknown Melastomataceae, Kreiter coll., 26 Dec. 2008. Remarks — this is the first record of this species in the West Indies. The measurements of the single specimen collected fit rather well the measurements given by Denmark and Muma (1989). All the setae are however slightly longer, but at maximum less than 10 % of variation, which is less than the threshold of 20 % around the mean defined by Tixier (2012) to characterize intraspecific variations. The measurements of the adult female presently found are as follow: dorsal shield smooth, dorsal shield length 368, width 288, 1 solenostome visible (gd 9), j1 30, j3 41, j4 3, j5 5, j6 5, J2 5, J5 8, z2 15, z4 12, z5 3, Z1 10, Z4 112, Z5 190, s4 90, S2 10, S4 10, S5 10, r3 12, R1 10, all setae smooth, st1-st1 65, st2-st2 75, st1-st3 63, posterior margin of the sternal shield concave, metapodal 1 length 25, width 3, metapodal 20 length 10, width 2, ventrianal shield 115 long, 78 wide at level of anterior corners and 78 wide at level of anus, 4 poroids around the ventrianal shield and genital shield, JV5 80, spermatheca 17 long and 5 width, SgeI 40, SgeII 40, SgeIII 50, StiIII 35, SgeIV 75, StiIV 62, StIV 75, macrosetae whip-like, fixed digit of chelicerae 37, mobile digit of chelicerae 40, 13-14 denticles on fixed digit and 4 on mobile digit., Published as part of Kreiter, S., Mailloux, J., Tixier, M. - S., Le Bellec, F., Douin, M., Guichou, S. & Etienne, J., 2013, New Phytoseiid Mites Of The French West Indies, With Description Of A New Species, And New Records (Acari: Mesostigmata), pp. 285-303 in Acarologia 53 (3) on page 290, DOI: 10.1051/acarologia/20132095, http://zenodo.org/record/4667233, {"references":["Chant D. A., Baker, E. W. 1965 - The Phytoseiidae (Acarina) of Central America - Mem. Entomol. Soc. Canada, 41, 56 pp.","Denmark H. A., Evans G. A., Aguilar H., Vargas C., Ochoa R. 1999 - Phytoseiidae of Central America - Indira Publishing House, West Bloomfield, Michigan, USA, 125 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 b - A revised catalog of the mite family Phytoseiidae - Zootaxa 434: 1 - 494.","Castro T. M. M. G. de, Moraes G. J. de, McMurtry J. A. 2010 - New Phytoseiidae (Acari: Mesostigmata) from Costa Rica, with description of two new species and additional information on other species - Intern. J. Acarol. 36 (1): 35 - 48. doi: 10.1080 / 01647950903506718","Denmark H. A., Muma M. H. 1989 - A revision of the genus Amblyseius Berlese, 1914 (Acari: Phytoseiidae) - Occasional Papers of the Florida State Collection of Arthropods, USA, 4, 149 pp.","Tixier M. - S. 2012 - Approaches to assess intraspecific variations of morphological continuous characters: the case study of the family Phytoseiidae (Acari: Mesostigmata) - Cladistics sous presse, doi: 10.1111 / j. 1096 - 0031.2012.00394. x"]}

Neoseiulella splendida Ferragut and Peaea-Estevez (Figure 39) Neoseiulella splendida Ferragut and Peaea-Estevez 2003: 161-164; Moraes et al. 2004: 295; Moraza and Peaea-Estevez 2006: 58; Chant and McMurtry 2007: 147. Adult female (Figure 39a – e) Dorsal shield (Figure 39a) — Dorsal shield heavily reticulated: length 361 (352 – 369); width 189 (181 – 196) (at level of s4) and 195 (193 – 197) (at level of Z1). Seven pairs of solenostomes on the dorsal shield: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. Ferragut and Peaea-Estevez (2003) noted the presence of only six pairs of solenostomes (gd1, gd2, gd5, gd6, gd8, and gd9). However, on their drawings a pair of solenostomes corresponding to the position of gd4 (according Athias-Henriot [1975] and Swirski et al. [1998]) is present. One pair of poroids. Sublateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except Z5 slightly serrated: j1 26 (23 – 28); j3 34 (32 – 35); j4 16; j5 16 (15 – 16); j6 18 (17 – 19); J2 19 (18 – 20); J5 17; z2 17 (16 – 17); z3 34 (33 – 35); z4 24; z5 16 (15 – 17); Z1 20 (19 – 20); Z4 62 (61 – 62); Z5 78; s4 34 (32 – 35); s6 40 (37 – 43); S2 41 (37 – 45); S4 27 (26 – 28); S5 26 (24 – 28); sub-lateral setae r3 30 (29 – 30) and R1 27 (25 – 29). Peritreme extending anteriorly to the level between j1-j3. Ventral shields (Figure 39b) — Sternal shield 69 (68 – 70) long and 69 (68 – 70) wide (at level of ST2), smooth with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 on separate platelets and a pair of small poroids accompanying ST4. Genital shield 125 long, 68 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla (folded in the specimen examined) between genital shield and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 1 pair of poroids around the genital shield. Ventrianal shield elongated with conspicuous waist, much longer than wide, 134 (124 – 143) long and 86 (82 – 89) wide (at level of ZV2), faintly striate, with four pairs of preanal setae (JV1, JV2 JV3 and ZV2) and with a pair of circular solenostomes gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 50 (44 – 56) long, smooth. Primary metapodal plate or inguinal sigillum 31 (26 – 35) long and 6 (5 – 7) wide. Spermatheca (Figure 39c) — Cervix 4 (3 – 4) long, U-shaped. Chelicera (Figure 39d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 36 (34 – 38) long, unidentate. Legs (Figure 39e) — Measurements of legs: leg I 332 (331 – 333); leg II 286 (279 – 292); leg III 284 (276 – 292); leg IV 392 (383 – 401). Seven setae (2- 2/0, 2/0-1) on the genu II. One pointed macroseta, 47 (46 – 48) long, on the basitarsus IV. Material examined — Four female paratypes deposited in the Department of Agroforestal Ecosystems, Universidad PolitØcnica de Valencia, Spain. Adult male (Figure 39f, g) Dorsal shield — dorsal shield similar to but smaller than the female. Sub-lateral seta R 1 inserted on the dorsal shield (different from the female, on which R 1 inserted on lateral margin). Ventrianal shield 140 long and 135 wide, distinctly reticulated, bearing seven pairs of preanal setae with a pair of poroids (Figure 39 f). Spermatodactyl Lshaped (Figure 39 g). This description is based on a paratype specimen. Previous reports — N. splendida is only known from Canary Islands. Plants on which this species was reported are: Argyranthemum lidii Humphries, A. escarrei (Svent.) Humphries, Cynara cardunculus, Nauplius stenophyllus (Link) Webb and Berth., Schizogyne glaberrima DC., Sonchus brachylobus Webb and Berthel (Asteraceae); Hypericum reflexum L.f. (Hypericaceae); Pennisetum setaceum (Forssk.) Chiov. (Poaceae); Suaeda vera J.F.Gmel., S. vermiculata Forssk. ex J.F.Gmel. (Chenopodiaceae). Remarks — N. splendida is similar to N. litoralis. However these two species differ by the following characters: gd4 presence/ absence (present on N. splendida; absent on N. litoralis); j3, z3, Z4, Z5, s4, and s6 lengths; the length of the peritreme (extending between j1-j3 on N. splendida; extending between z2-j3 on N. litoralis). Moreover, the males of these two species differ in the number of setae on the ventrianal shield (seven pairs on N. splendida, six on N. litoralis). These two species seem thus to be separate entities. However, further experiments would be useful in order to conclude on the reliability of these morphological characters in species identification., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 332-334, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Ferragut F., Peaea-Estevez M. A. 2003 - Phytoseiid mites of the Canary Islands (Acari: Phytoseiidae): 1. Gran Canaria Island - Intern. J. Acarol., 29 (2): 149 - 170.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Moraza M. L., Peaea-Estevez M. A .. 2006 - A new species of Neoseiulella (Acari: Phytoseiidae) from the Macaronesian Region, Canary Islands - Zootaxa, 1366: 55 - 59.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Athias-Henriot C. 1975 - Nouvelles notes sur les Amblyseiini. II. Le relevO organotaxique de la face dorsale adulte (Gamasides protoadOniques, Phytoseiidae) - Acarologia, 17 (1): 20 - 29.","Swirski E., Ragusa S., Tsolakis H. 1998 - Keys to the phytoseiid mites (Parasitiformes: Phytoseiidae) of Israel - Phytophaga, 8: 85 - 154."]}

Neoseiulella spaini (Collyer) (Figure 38) Typhlodromus spaini Collyer 1982: 189; Schicha 1987: 134-135; Chant and Yoshida-Shaul 1989: 1024-1026. Neoseiulella (Typhloctona) spaini (Collyer) Denmark and Rather 1996: 73-74. Neoseiulella spaini (Collyer) Moraes et al. 1986: 202; Moraes et al. 2004: 295; Chant and McMurtry 2007: 147. Adult female (Figure 38a – d) Dorsal shield (Figure 38a) — Dorsal shield distinctly reticulated along the lateral area: length 412; width 209 (at level of s4) and 218 (at level of Z1). Six pairs of solenostomes on the dorsal shield: gd1, gd2, gd4, gd5, gd6, and gd9. One pair of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z5 slightly serrated: j1 25; j3 26; j4 11; j5 11; j6 15; J2 16; J5 12; z2 10; z3 16; z4 17; z5 11; Z1 15; Z4 31; Z5 48; s4 23; s6 23; S2 20; S4 17; S5 17; sublateral setae r3 20 and R1 16. Peritreme extending anteriorly to the level between j1-j3. Ventral shields (Figure 38b) — Sternal shield 84 long and 91 wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. Metasternal setae (ST4) inserted on separate platelets with a pair of small poroids. Genital shield 126 long and 72 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield subtriangular-shaped, 140 long and 118 wide (at level of ZV2), is smooth, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of small and circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 44 long, smooth. Primary metapodal plate or inguinal sigillum 35 long and 8 wide. Spermatheca (Figure 38c) — Cervix 8 long, cupshaped. Chelicera (Figure 38d) — Five teeth and a pilus dentilis on the fixed digit. Movable digit 33 long, bidentate. Legs (Figure 38e) — Measurements of legs: leg I 368; leg II 322; leg III 320; leg IV 412. Eight setae (2- 2/1, 2/0-1) on the genu II. Three knobbed macrosetae, 30, 28 and 54 long, on the genu, tibia and basitarsus IV, respectively. Material examined — The female holotype deposited in the New Zealand Arthropod Collection (NZAC), Landcare Research, Auckland, New Zealand. Adult male Described by Collyer (1982) and Denmark and Rather (1996). We were not able to borrow the male type specimen of this species. Previous reports — N. spaini is only known from New Zealand (Collyer 1982; Schicha 1987). The female holotype, paratypes and other specimens of this species are reported on Olearia colensoi (Asteraceae) at Ruahine Range and Urewera National Park., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 331-332, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella ferraguti Moraza and Peaea-Estevez (Figure 28) Neoseiulella ferraguti Moraza and Peaea-Estevez 2006: 56-58. Adult female (Figure 28a – e) Dorsal shield (Figure 28a) — Dorsal shield smooth: length 365; width 226 (at level of s4), 264 (at level of Z1), with seven pairs of solenostomes: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. Seven pairs of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except Z4 and Z5 which are slightly serrated: j1 28; j3 49; j4 31; j5 32; j6 38; J2 34; J5 19; z2 15; z3 54; z4 47; z5 29; Z1 34; Z4 77; Z5 94; s4 62; s6 64; S2 71; S4 54; S5 33; sub-lateral setae r3 28 and R1 35. Most dorsal setae of opisthosomal area (S2, S4, S5, Z4, Z5, and J5) on tubercles. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 28 b) — Sternal shield 67 long and 68 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of distinct poroids. ST3 and ST4 on separate platelets. A pair of small poroids accompanies ST4. Genital shield 130 long and 66 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under genital shield. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield subtriangularshaped, 127 long and 112 wide (at level of ZV2), faintly striate, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and a pair of large and circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 66 long, lightly serrated. Primary metapodal plate or inguinal sigillum 22 long and 11 wide. Spermatheca (Figure 28c) — Cervix 4 long, Ushaped. Chelicera (Figure 28d) — Seven teeth and a pilus dentilis on the fixed digit. Movable digit 37 long, bidentate. Legs (Figure 28e) — Measurements of legs: leg I 422; leg II 313; leg III 314; leg IV 362. Eight setae (2-2/1, 2/0-1) on the genu II. One macroseta (not reported on the original description), 33 long, on the genu III. Three other pointed macrosetae, 54, 34, 67 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female holotype deposited in Museum of Zoology of University of Navarra (MZUNAV), Spain. Adult male (Figure 28f, g) Male specimen of this species similar to but smaller than the female. Moraza and Peaea-Estevez (2006) described the male of N. ferraguti. However, these authors did not give a complete setal and body measurements. We thus present a complete description of this male. Dorsal shield — Dorsal shield chaetotaxy similar to the female, with smooth surface: length 322; width 163 (at level of s4), 200 (at level of Z1), with seven pairs of solenostomes: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. Dorsal shield bearing 21 pairs of setae (sub-lateral setae, r3 and R 1 on the dorsal shield). All dorsal setae smooth, except Z5 which is slightly serrated: j1 29; j3 36; j4 22; j5 22; j6 25; J2 25; J5 13; z2 11; z3 43; z4 33; z5 24; Z1 25; Z4 55; Z5 68; s4 49; s6 48; S2 50; S4 39; S5 29; sub-lateral setae r3 23 and R 1 32. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 28f) — Sternogenital shield is smooth. Ventrianal shield 135 long and 170 wide, lightly reticulated, bearing seven pairs of preanal setae with two pairs of poroids. Caudoventral seta JV5 42 long, smooth, on the integument surrounding the ventrianal shield. Chelicera (Figure 28g) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 31 long, unidentate. Spermatodactyl L-shaped. Legs — Measurements of legs: leg I 375; leg II 306; leg III 283; leg IV 355. Three macrosetae, 35, 35 and 50 long, on genu, tibia, and basitarsus IV, respectively. The male specimen of this species is different from the female by the position of both sub-lateral setae (r3 and R 1) (on dorsal shield on the male; on lateral margin on the female), and the number of solenostomes on the ventrianal shield (two pairs on the male; one pair on the female). Material examined — A male paratype specimen. Previous reports — N. ferraguti is only known from Tenerife (Canary Islands). It was reported from lichens on dead log, and from soil and litter of Cheirolophus canariensis var. subexpinnatus (Burch.) A. Hansen and Sunding (Asteraceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 313-316, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Moraza M. L., Peaea-Estevez M. A .. 2006 - A new species of Neoseiulella (Acari: Phytoseiidae) from the Macaronesian Region, Canary Islands - Zootaxa, 1366: 55 - 59."]}

Neoseiulella neoviniferae Basha, Mahrous and Mostafa (Figure 33) Neoseiulella grapevini Basha et al., Chant and Mc- Murtry 2007: 147. Adult female (Figure 33a – d) Dorsal shield (Figure 33a) — Dorsal shield distinctly reticulated throughout: length 386 (381 – 390), width 175 (171 – 178) (at level of s4) and 191 (187 – 195) (at level of Z1). Seven pairs of large solenostomes on the dorsal shield: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. Five pairs of poroids. Sublateral setae (r3 and R1) on the lateral margin. Like us, Basha et al. (2004) noted the presence of seven pairs of solenostomes. However, on their drawings gd4 is absent (present on the specimens examined). Dorsal shield bearing 19 pairs of setae, all smooth: j1 23 (21 – 25); j3 24 (22 – 26); j4 14 (12 – 15); j5 12 (10 – 13); j6 16 (15 – 17); J2 15 (14 – 15); J5 15 (14 – 15); z2 10 (9 – 11); z3 24 (22 – 25); z4 19 (17 – 20); z5 14 (13 – 15); Z1 17 (16 – 17); Z4 47 (44 – 49); Z5 62 (59 – 64); s4 23 (22 – 24); s6 26 (25 – 27); S2 35 (34 – 36); S4 33 (31 – 35); S5 28 (26 – 30); sub-lateral setae r3 28 (26 – 30) and R1 25 (23 – 27). Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j3. Ventral shields (Figure 33b) — Sternal shield 62 (60 – 63) long and 67 (66 – 67) wide (at level of ST2), is smooth with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 on separate platelets. A pair of small poroids accompanying ST4. Genital shield 128 (125 – 131) long and 66 (64 – 67) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subtriangularshaped, 130 (125 – 134) long and 98 (95 – 101) wide (at level of ZV2), distinctly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 48 (46 – 50) long, smooth. Primary metapodal plate or inguinal sigillum 39 (38 – 40) long and 5 (4 – 6) wide. Spermatheca (Figure 33c) — Cervix 4 (3 – 4) long, cup-shaped. Chelicera (Figure 33d) — Two large teeth and a pilus dentilis on the fixed digit. Movable digit 38 (36 – 39) long, unidentate. Basha et al. (2004) mentioned that the fixed digit bears three teeth. Legs (Figure 33e) — Measurements of legs: leg I 341 (333 – 349); leg II 286 (285 – 286); leg III 285 (284 – 286); leg IV 381 (369 – 393). Seven setae (2- 2/0, 2/0-1) on the genu II. One macroseta 42 (37 – 47) long, on the basitarsus IV. Material examined — Four female paratypes deposited in the collection of Plant Protection Department, Faculty of Agriculture, Zagazig University, Egypt. Adult male The male of this species is unknown. Previous reports — N. neoviniferae is only known from Al-Sharkia Governorate, El-Khattara District (Egypt) on Vitis vinifera L. (Vitaceae). Remarks — N. neoviniferae is similar to N. montforti but differs by the presence/ absence of gd5 (present on N. neoviniferae; absent on N. montforti), the shape of the spermatheca (c-shaped on N. neoviniferae; U-shaped on N. montforti) and measurements of some dorsal setae (j6, J2, z2, Z1 and R 1)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 322-324, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090

Neoseiulella dachanti (Collyer) (Figure 25) Typhlodromus dachanti Collyer 1964: 638-640. Schicha 1980: 22-24; Collyer 1982: 189; Schicha 1987: 139, 141; Chant and Yoshida-Shaul 1989: 1032-1034. Neoseiulella (Neoseiulella) dachanti (Collyer) Denmark and Rather 1996: 54-55. Neoseiulella dachanti (Collyer) Moraes et al. 1986: 201; Moraes et al. 2004: 293; Chant and McMurtry 2007: 147. Adult female (Figure 25a – e) Dorsal shield (Figure 25a) — Dorsal shield smooth: length 394; width 225 (at level of s4), 226 (at level of Z1). Four pairs of solenostomes: gd2, gd4, gd6, and gd9. Four pairs of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all dorsal and sub-lateral setae smooth, except Z5 slightly serrated: j1 27; j3 41; j4 7; j5 8; j6 12; J2 13; J5 7; z2 11; z3 39; z4 13; z5 8; Z1 13; Z4 48; Z5 111 (106 – 115); s4 42; s6 17; S2 23; S4 12; S5 10; r3 14 and R1 15. Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 25b) — Sternal shield 83 long and 80 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 104 long and 68 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under genital shield. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield 121 long and 88 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), with a pair of large solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 39 long, smooth. Primary metapodal plate or inguinal sigillum 20 long and 6 wide. Spermatheca (Figure 25c) — Cervix 3 long, shallowly cup-shaped. Chelicera (Figure 25d) — Twelve teeth, without pilus dentilis, on the fixed digit. Movable digit, 35 long, bearing three teeth. Legs (Figure 25e) — Measurements of legs: leg I 372; leg II 323 (318 – 328); leg III 323 (318 – 328); leg IV 395 (391 – 400). Seven setae (2-2/0, 2/0-1) on the genu II. Two macrosetae, 30 and 27 long, on genu and the tibia III, respectively. Three other macrosetae, 46, 48 and 48 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female lectotype deposited in the BMNH, the British Museum of Natural History, Cromwell Road, London, UK. Adult male Described by Collyer (1964) and Schicha (1980). We were not able to borrow the male type specimen of this species. Previous reports — The type specimens of N. dachanti were collected at Waitakeres, near Auckland, New Zealand, on "native bush". This species is only known from the Australasian area: New Zealand (Collyer 1964, 1982) and Australia (Schicha 1987). Plant supports on which this species was reported are: Brachyglottis repanda J. R. and G. Forst. (Asteraceae); Coprosma sp. (Rubiaceae); Cytisus sp. (Fabaceae); Elaeocarpus dentatus (G. R. and G. Forster) Vahl (Elaeocarpaceae); Erica sp. (Ericaceae); Melicytus ramiflorus G. R. and G. Forster (Violaceae); Nothofagus sp. (Nothofagaceae); Nothopanax sp., Pseudopanax sp. (Araliaceae); Podocarpus sp. (Podocarpaceae); Eucalyptus sp. (Myrtaceae); Rubus sp. (Rosaceae) and tree ferns., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 310, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1964 - The occurrence of some mites of the family Phytoseiidae in New Zealand, and descriptions of seven new species - Acarologia, 6: 632 - 646.","Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella longiseta Moraza, Peaea-Estevez and Ferragut (Figure 30) Neoseiulella longiseta Moraza, Peaea-EstØvez and Ferragut 2005: 107-111; Moraza and Peaea-Estevez 2006: 59; Chant and McMurtry 2007: 147. Adult female (Figure 30a – e) Dorsal shield (Figure 30a) — Dorsal shield faintly striate: length 405 (401 – 408); width 202 (196 – 207) (at level of s4), 238 (234 – 242) (at level of Z1), with seven pairs of solenostomes: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. No poroid visible. Sublateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z4 and Z5 serrated. Most lateral setae on tubercles: j1 31; j3 41 (37 – 45); j4 12 (10 – 14); j5 16 (15 – 16); j6 26 (25 – 26); J2 16 (15 – 16); J5 14 (13 – 15); z2 10; z3 46 (41 – 51); z4 25 (23 – 27); z5 15 (14 – 15); Z1 15; Z4 88 (84 – 91); Z5 117 (116 – 118); s4 67 (63 – 70); s6 71 (70 – 72); S2 75 (73 – 76); S4 63 (62 – 63); S5 45 (41 – 48); sub-lateral setae r3 24 (23 – 25) and R1 34 (33 – 35). Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 30 b) — Sternal shield 75 (74 – 76) long and 75 (74 – 76) wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and poroids. ST3 and ST4 on separate platelets with a pair of small poroids accompanying ST4. Genital shield 140 long and 75 (73 – 76) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. No pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield subcircular-shaped, reticulate with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and a pair of large and circular solenostomes gv3 posteromediad to JV3. Length 136 (135 – 137); wide 129 (127 – 130) (at level of ZV2), 142 (137 – 147) (at level of JV3), 99 (97 – 101) (al level of anus). Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 83 (80 – 85) long, smooth. Primary metapodal plate or inguinal sigillum 26 (24 – 28) long and 8 (6 – 7) wide. Spermatheca (Figure 30c) — Cervix 5 (4 – 6) long, U-shaped. Chelicera (Figure 30d) — Six teeth and a pilus dentilis on the fixed digit. Movable digit 46 (42 – 48) long, bidentate. Legs (Figure 30e) — Measurements of legs: leg I 401; leg II 306 (287 – 325); leg III 313 (300 – 325); leg IV 438 (430 – 445). Eight setae (2-2/1, 2/0-1) on the genu II. Three pointed macrosetae, 63 (62 – 64), 49 (40 – 58) and 76 (72 – 78) long, on the genu, tibia and basitarsus IV, respectively. Material examined — Two female paratypes deposited in Museum of Zoology of University of Navarra, Spain. Adult male Described by Moraza et al. (2005). We were not able to borrow the male type specimen of this species. Previous reports — N. longiseta is only known from the Canary Islands. It was collected from dry soil, moss and litter under the following plants: Aeonium sp., Monanthes sp. (Crassulaceae), Asplenium sp. (Aspleniaceae), Juniperus turbinata Guss. (Cupressaceae), Cistus monspeliensis L. (Cistaceae), and from lichens (Moraza et al. 2005)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 318, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Moraza M. L., Peaea-Estevez M. A .. 2006 - A new species of Neoseiulella (Acari: Phytoseiidae) from the Macaronesian Region, Canary Islands - Zootaxa, 1366: 55 - 59.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Moraza M. L., Peaea-Estevez M. A., Ferragut F .. 2005 - Two new species of Neoseiulella Muma of the Canary Islands (Acari: Phytoseiidae) - Intern. J. Acarol., 31 (2): 107 - 112. doi: 10.1080 / 01647950508683659"]}

Neoseiulella montforti (Rivnay and Swirski) (Figure 31) Typhloctonus montforti Rivnay and Swirski 1980: 179-182; Swirski and Amitai 1984: 77; Denmark and Rather 1984: 169-170; Moraes et al. 1986: 233. Typhlodromus montforti (Rivnay and Swirski) Chant and Yoshida-Shaul 1989: 1020-1023. Neoseiulella montforti (Rivnay and Swirski) Swirski and Amitai 1997b: 37; Moraes et al. 2004: 294; Chant and Mc- Murtry 2007: 147. Neoseiulella (Typhloctona) montforti (Rivnay and Swirski) Denmark and Rather 1996: 66- 68. Adult female (Figure 31a – d) Dorsal shield (Figure 31a) — Dorsal shield heavily reticulate: length 383 (381 – 384); width 193 (191 – 194) (at level of s4), 196 (194 – 197) (at level of Z1), with six pairs of solenostomes: gd1, gd2, gd4, gd6, gd8, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae. On one of specimens examined, one seta of the J1 pair is present. All dorsal and sub-lateral setae smooth, except Z5 serrated: j1 30 (28 – 31); j3 36 (34 – 37); j4 25 (23 – 26); j5 23 (22 – 24); j6 31 (30 – 31); J1 36; J2: 37 (36 – 37); J5 13 (12 – 13); z2 24 (22 – 25); z3 32 (31 – 32); z4 34 (33 – 34); z5 23; Z1 41 (39 – 43); Z4 48 (46 – 50); Z5 58 (56 – 59); s4 38 (37 – 39); s6 43; S2 45 (43 – 46); S4 43 (41 – 45); S5 40 (38 – 41); sub-lateral setae r3 35 (33 – 36) and R1 41 (39 – 42). Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j3 or between j1-j3. Ventral shields (Figure 31b) — Sternal shield weakly sclerotized, 74 (73 – 74) long and 80 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 on separate platelets with a pair of small poroids accompanying ST4. Genital shield 150 long and 76 (74 – 78) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subtriangular-shaped, 133 (131 – 135) long and 116 (112 – 119) wide (at level of ZV2), reticulate with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and a pair of small circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 52 (51 – 53) long, is smooth. Primary metapodal plate or inguinal sigillum 38 (35 – 40) long and 5 (4 – 6) wide. Spermatheca — Spermatheca not discernible in the two type specimens examined. Rivnay and Swirski (1980) mentioned that the cervix has a Ushaped type. Chelicera (Figure 31c) — Two or three teeth and a pilus dentilis on the fixed digit. Movable digit 39 (38 – 40) long, bears one or two teeth. Legs (Figure 31d) — Measurements of legs: leg I 357 (350 – 363); leg II 302 (297 – 307); leg III 312 (302 – 321); leg IV 422 (411 – 432). Seven setae (2-2/0, 2/0-1) on the genu II. One macroseta, 38 (34 – 41) long, on the basitarsus IV. Material examined — Two female paratypes deposited in the Division of Entomology, Agricultural Research Organization, Bet Dagan, Israel. Adult male The male of this species is unknown. Previous reports — N. montforti is only known from Israel (Swirski and Amitai 1984; Swirski and Amitai 1997a). The type specimens were collected at Monfort (Western Galilee, Israel) on Quercus calliprinos (Fagaceae). It has since been reported on Artemisia monosperma, Limbarda crithmoides (Asteraceae); at Haifa (Western Carmel) on Q. calliprinos; and on Platanus orientalis L. (Platanaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 319-321, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Rivnay T., Swirski E. 1980 - Four new species of phytoseiid mites (Acarina: Mesostigmata) from Israel - Phytoparasitica, 8: 173 - 187.","Swirski E., Amitai S. 1984 - Notes on phytoseiid mites (Mesostigmata: Phytoseiidae) from the Mediterranean littoral zone of Israel, with a description of a new species of Typhloctonus - Israel J. Entomol., 18: 71 - 82.","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Swirski E., Amitai S. 1997 b - Annotated list of phytoseiid mites (Mesostigmata: Phytoseiidae) in Israel - Israel J. Entomol., 31: 21 - 46.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Swirski E., Amitai S. 1997 a - Notes on phytoseiid mites (Mesostigmata: Phytoseiidae) of Mt. Carmel (Israel), with descriptions of two new species - Israel J. Entomol., 31: 1 - 20."]}

Neoseiulella squamiger (Wainstein) (Figure 3) Typhlodromus squamiger Wainstein 1960: 689-690; Chant and Yoshida-Shaul 1989: 1013. Typhlodromus (Nesbitteius) squamiger (Wainstein) Arutunjan 1970: 19. Typhloctonus squamiger (Wainstein) Wainstein 1973: 176; Chaudhri et al. 1974, 231; Kolodochka 1974a: 27; Arutunjan 1977: 55; Kolodochka 1978: 50-51; Denmark and Rather 1984: 173-174; Moraes et al. 1986: 233. Pegodromus squamiger (Wainstein) Athias-Henriot and Fauvel 1981: 74. Neoseiulella (Typhloctona) squamiger (Wainstein) Denmark and Rather 1996: 61-62. Neoseiulella squamiger (Wainstein) Moraes et al. 2004: 291; Chant and McMurtry 2007: 147. Adult female (Figure 3a – d) Dorsal shield (Figure 3a) — Dorsal shield distinctly reticulated throughout: 342 long, 184 (at level of s4) and 210 (at level of Z1). Five pairs of small solenostomes on the dorsal shield: gd1, gd2, gd6, gd8, and gd9 and no poroid visible. Sub-lateral setae r3 on the lateral margin and R1 on the dorsal shield. Dorsal shield bearing 20 pairs of setae (R1 on dorsal shield) all serrated: j1 14; j3 20; j4 17; j5 16; j6 21; J2 29; J5 13; z2 20; z3 25; z4 28; z5 15; Z1 28; Z4 35; Z5 42; s4 29; s6 30; S2 34; S4 27; S5 17; sublateral setae r3 20 and R1 20. Peritreme extending anteriorly to the level of j3. Ventral shields (Figure 3b) — Sternal shield 37 long and 53 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids (posterior to ST1; anterior to ST3). ST3 on an elongate projection of the sternal shield. A pair of metasternal setae (ST4) on separate platelets with a pair of small poroids. Genital shield 130 long and 60 wide (at level of ST5), smooth. Four elongate platelets or sigilla between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield subquadrateshaped 103 long and 86 wide (at level of ZV2), distinctly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of solenostomes gv3 posterior to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 17 long, smooth. Primary metapodal plate or inguinal sigillum 30 long and 3 wide. Spermatheca (Figure 3c) — Cervix 4 long, cupshaped, with an enlarged atrium. Chelicera (Figure 3d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 25 long, is edentate. Legs — Measurements of legs: leg I 227; leg II 209; leg III 211; leg IV 239. Seven setae (2-2/0, 2/0- 1) on the genu II. No macroseta on legs. Material examined — The female holotype deposited in the ASU, Institute of Zoology, Academy of Sciences, Kiev, Ukraine. Adult male (Figure 3e, f) Described by Denmark and Rather (1984, 1996). Dorsal shield chaetotaxy is similar to the female. Ventrianal shield 103 long and 142 wide, distinctly reticulated and with five pairs of preanal setae and three pairs of poroids. Spermatodactyl with a terminal foot, and a slightly enlarged toe. This description is based on one of the male specimens of our collection (Montpellier SupAgro collection, UMR CBGP), collected in Kiev (Ukraine) on Norway maple (Acer platanoides L.). Previous reports — N. squamiger is only known from the West-Palearctic area. Countries from which this species is reported are: Armenia (Arutunjan 1970, 1971); Croatia (Tixier et al. 2010); Greece (Papadoulis and Emmanouel 1990); Hungary (Bozai 1996, 1997; Ripka 1998, 2006); Moldova (Wainstein 1960, 1973); Ukraine (Kolodochka 1973, 1974b, 1978). Plant supports on which N. squamiger was collected are: Acer campestre, A. platanoides, Acer sp. (Aceraceae); Carpinus betulus, Corylus avellana, Ostrya carpinifolia Scopoli (Betulaceae); Cornus sp. (Cornaceae); Fraxinus ornus L. (Oleaceae); J. regia (Juglandaceae); Picea sp. (Pinaceae); Prunus (Cerasus) sp., Prunus sp. (Rosaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 271, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Wainstein B. A. 1960 - New species and subspecies of the genus Typhlodromus Scheuten (Parasitiformes, Phytoseiidae) of the USSR fauna - Zool. Zh., 39: 683 - 690.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Arutunjan E. S. 1970 - Phytoseiid mites (Phytoseiidae) on agricultural crops in the Armenian SSR - Akademii Nauk Armyanskoi SSR, Otdelenie Biologicheskikh Nauk, Dissertatsii na Soiskanie Uchenoi Stepeni Candidata Biologrcheskikh Nauk, Zooliya, Armenia, 97: 1 - 31.","Chaudhri W. M., Akbar S., Rasool A. 1974 - Taxonomic studies of the mites belonging to the families Tenuipalpidae, Tetranychidae, Tuckerellidae, Caligonellidae, Stigmaeidae and Phytoseiidae - University of Agriculture Technical Bulletin, Lyallpur, Pakistan. 1: 204 - 233.","Kolodochka L. A. 1974 a - The predaceous phytoseiid mites (Parasitiformes: Phytoseiidae) from the forest steppe of the Ukranian SSR. Part II. Species of the genera Kampimodromus, Paraseiulus, Typhlodromus, Typhloctonus, Anthoseius, Phytoseius - Vest. Zool., (1): 25 - 29.","Arutunjan E. S. 1977 - Identification manual of phytoseiid mites of agricultural crops of the Armenian SSR - Akademiya Nauk Armyanskoi SSR, Zoologicheskii Institut, Erevan, 1 - 177","Kolodochka L. A. 1978 - Handbook on identifying of plant inhabiting phytoseiid mites - Nauka Dumka, Kiev. 78 pp.","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Athias-Henriot C., Fauvel. G. 1981 - Pegodromus crassipilis, n. g., n. sp., Typhlodromini nouveau du sud de la France (Parasitiformes: Phytoseiidae) - Intern. J. Acarol., 7: 71 - 74. doi: 10.1080 / 01647958108683246","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Arutunjan E. S. 1971 - Confinement of phytoseiid mites to biotopes and their morphological adaptations - Biolgicheskii Zhurnal Armenii, Akadrmiya Nauk Armyanskoi SSR, 24: 41 - 47.","Tixier M. - S., Klaric V., Kreiter S., Duso C. 2010 - Phytoseiid Mite Species From Croatia, With Description of a New Species of the Genus Typhlodromus (Typhlodromus) - Entomol. Soc. Amer., 103 (2): 165 - 180.","Papadoulis G. T., Emmanouel N. G. 1990 - Phytoseiid mites of Greece: new records of species and description of the male and immature stages of Typhlodromus erymanthii Papadoulis and Emmanouel - Biol. Gallo- Hellen., 17 (1): 13 - 26.","Bozai J. 1996 - Data to the knowledge of predaceous mites of Hungary (Acari: Phytoseiidae, Phytoseiinae) - NovOnyvOdelem., 32: 521 - 525.","Bozai J. 1997 - Data to the fauna of predaceous mites of Hungary with the description of four new species (Acari: Phytoseiidae) - Fol. Entomol. Hungar., 58: 35 - 43.","Ripka G. 1998 - New data to the knowledge on the phytoseiid fauna in Hungary (Acari: Mesostigmata) - Acta Phytopathol. Entomol. Hung., 33: 395 - 405.","Ripka G. 2006 - Checklist of the Phytoseiidae of Hungary (Acari: Mesostigmata) - Folia Entomol., 67: 229 - 260.","Kolodochka L. A. 1973 - Predaceous phytoseiid mites (Parasitiformes: Phytoseiidae) from the forest steppe of the Ukrainian SSR. Part I. Species of the genus Amblyseius - Vest. Zool., (5): 78 - 81.","Kolodochka L. A. 1974 b - Predaceous phytoseiid mites (Parasitiformes, Phytoseiidae) from the forest steppe of the Ukrainian SSR. Part III. A key to genera and species - Vest. Zool., (3): 87 - 89."]}

Neoseiulella manukae (Collyer) (Figure 12) Typhlodromus manukae Collyer 1964: 637-638; Schicha 1980: 16-18; Collyer 1982: 190; Schicha 1987: 140-143; Chant and Yoshida-Shaul 1989: 1034-1036. Neoseiulella manukae (Collyer) Moraes et al. 1986: 201; Moraes et al. 2004: 294; Chant and McMurtry 2007: 147. Neoseiulella (Neoseiulella) manukae (Collyer) Denmark and Rather 1996: 55-56. Adult female (Figure 12a – e) Dorsal shield (Figure 12a) — Dorsal shield smooth: length 440 (439 – 440); width 226 (225 – 226) (at level of s4), 224 (223 – 224) (at level of Z1), with five pairs of solenostomes: gd2, gd4, gd5, gd6, and gd9. three pairs of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except Z5 which is slightly serrated: j1 34 (33 – 34); j3 46 (45 – 46); j4 15 (14 – 15); j5 14; j6 16; J2 16; J5 14; z2 17; z3 30 (29 – 30); z4 31 (30 – 31); z5 15; Z1 15; Z4 55; Z5 134 (133 – 134); s4 36 (35 – 36); s6 25 (24 – 25); S2 28; S4 17; S5 21; sublateral setae r3 20 and R1 17. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 12b) — Sternal shield 92 (91 – 92) long and 88 (86 – 89) wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 133 (132 – 133) long and 74 (73 – 74) wide (at level of ST5), smooth. Elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield longer than wide, 143 (142 – 143) long and 110 (109 – 110) wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), and a pair of small circular solenostomes gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 50 (49 – 50) long, smooth. Primary metapodal plate or inguinal sigillum 23 (22 – 23) long and 7 wide. Spermatheca (Figure 12c) — Cervix 5 long, cupshaped. Chelicera (Figure 12d) — Eleven teeth and a pilus dentilis on the fixed digit. Movable digit 33 (32 – 33) long, bearing three teeth. Legs (Figure 12e) — Measurements of legs: leg I 396 (395 – 396); leg II 330 (329 – 330); leg III 345 (344 – 346); leg IV 437 (436 – 437). Seven setae (2-2/0, 2/0-1) on the genu II. Two macrosetae, 41 (40 – 41) and 32 (31 – 32) long, on genu and tibia III, respectively. Three other macrosetae, 81, 67 and 50 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female lectotype and one female paralectotype deposited in the BMNH, the British Museum of Natural History, Cromwell Road, London, UK. Adult male The male of this species is unknown. Previous reports — N. manukae is only known from the Australasian area. The type specimens were collected at Waitakeres, near Auckland, New Zealand, on Leptospermum scoparium J.R. Forst. and G. Forst. (Myrtaceae) (Collyer 1964). This species has been also found on L. ericoides A. Rich. at Awanui Inlet, New Zealand (Collyer 1982); and on Eucalyptus parivolia Cambage (Myrtaceae) in New South Wales, Australia (Schicha 1987)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 288-290, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1964 - The occurrence of some mites of the family Phytoseiidae in New Zealand, and descriptions of seven new species - Acarologia, 6: 632 - 646.","Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080"]}

Neoseiulella nesbitti (Womersley) (Figure 14) Typhlodromus nesbitti Womersley 1954: 179-180; Hirschmann 1962: 12; Narayanan and Ghai 1963: 541; Prasad 1974: 174; Schicha 1978: 5-7; Schicha 1987: 132-133; Chant and Yoshida-Shaul 1989: 1043- 1044. Typhlodromus (Typhlodromus) nesbitti (Womersley) Chant 1959: 65-66. Neoseiulella nesbitti (Womersley) Muma 1961: 295; Moraes et al. 1986: 201; Chant and McMurtry 1994: 249; Moraes et al. 2004: 294; Chant and McMurtry 2007: 147. Typhlodromus (Nesbittieus) nesbitti (Womersley) Wainstein 1962: 23. Typhlodromus (Seiulus) nesbitti (Womersley) Ehara 1966: 16-17. Typhloctonus (Neoseiulellus) nesbitti (Womersley) Wainstein 1977: 1415. Typhlodromus (Typhloctonus) nesbitti (Womersley) Gupta 1985: 403-404. Neoseiulella (Neoseiulella) nesbitti (Womersley) Denmark and Rather 1996: 46-48. textitAdult female (Figure 14a – e) Dorsal shield (Figure 14a) — Dorsal shield lightly reticulated: length 399; width 228 (at level of s4) and 233 (at level of Z1). Five pairs of solenostomes: gd2, gd4, gd5, gd6, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z4 and Z5 which are slightly serrated: j1 25; j3 27; j4 18; j5 19; j6 20; J2 21; J5 12; z2 19; z3 28; z4 28; z5 18; Z1 20; Z4 38; Z5 57; s4 26; s6 27; S2 31; S4 25; S5 23; sub-lateral setae r3 21 and R1 20. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 14b) — Sternal shield 86 long and 82 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 126 long and 73 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield much longer than wide, 144 long and 110 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), with a pair of small circular solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5, JV3 absent) on the integument surrounding the ventrianal shield. JV5 41 long, smooth. Primary metapodal plate or inguinal sigillum 28 long and 7 wide. Spermatheca (Figure 14c) — Cervix 6 long, shallowly cup-shaped. Chelicera (Figure 14d) — Nine teeth and a pilus dentilis on the fixed digit. Movable digit 35 long, bearing three teeth. Legs (Figure 14e) — Measurements of legs: leg I 353; leg II 322; leg III 302; leg IV 395. Seven setae (2- 2/0, 2/0-1) on the genu II. Three pointed macrosetae, 31, 33 and 49 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female holotype deposited in the South Australian Museum, Adelaide, Australia. Adult male Described by Schicha (1978). We were not able to borrow the male type specimens of this species. Previous reports — The type specimens of N. nesbitti were collected from Australia (New South Wales) on Medicago arborea L. (Fabaceae). This species was reported from two biogeographical areas: Australasian and Oriental areas. Countries in which N. nesbitti was recorded are: Australia (New South Wales) (Womersley 1954; Schicha 1975; Wainstein 1977; Schicha 1978). Australia (Tasmania) (Schicha 1987); India (Delhi) (Narayanan and Ghai 1961). Plants on which this species was collected are: Actinidia chinensis Planch. (Actinidiaceae); Banksia integrifolia L.f. (Proteaceae); Eucalyptus parivolia (Myrtaceae); Casuarina sp. (Casuarinaceae); Hydrangea macrophylla (Thunb.) Ser. (Hydrangeaceae); Malus domestica (Rosaceae); Mangifera indica L. (Anacardiaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 292, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Womersley H. 1954 - Species of the subfamily Phytoseiinae (Acarina: Laelaptidae) from Australia - Austral. J. Zool., 2: 169 - 191. doi: 10.1071 / ZO 9540169","Hirschmann W. 1962 - Gangystematik der Parasitiformes - Acarologie Schriftenreihe fur Vergleichende Milbenkunde, Hirschmann-Verlag, Furth / Bay. 5 (5 - 6): 80 pp + 32 plates.","Narayanan E. S., Ghai S. 1963 - Some new records and a new species of mites associated with malformation of mango trees in India - Proceed. National Institute Science India, 29 (b): 535 - 546.","Prasad V. 1974 - A catologue of mites of India - Indira Acarology Publishing House, Ludhiana, Punjab. 320 pp.","Schicha E. 1978 - Typhlodromus nesbitti Womersly redescribed (Acari: Phytoseiidae) - Austral. Entomol. Mag., 5 (1): 5 - 7.","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Chant D. A. 1959 - Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species - Can. Entomol., Supplement 12, 166 pp.","Muma M. H. 1961 - Sub-familes, genera, and species of Phytoseiidae (Acarina: Mesostigmata) - Flor. St. Mus. Bull., 5 (7): 267 - 302.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Wainstein B. A. 1962 - ROvision du genre Typhlodromus Scheuten, 1857 et systematique de la famille des Phytoseiidae (Berlese 1916) (Acarina: Parasitiformes) - Acarologia, 4: 5 - 30.","Ehara S. 1966 - A tentative catalogue of predatory mites of Phytoseiidae known from Asia, with descriptions of five new species from Japan - Mushi, 39: 9 - 30.","Wainstein B. A. 1977 - A contribution to the fauna of the family Phytoseiidae (Parasitiformes) in Australia - Zool. Zh., 56: 1413 - 1416.","Gupta S. K. 1985 - Plant mites of India - Zooogical Survey of India Handbook Series, Calcutta. 5: 1 - 520.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Schicha E. 1975 - Predacious mites (Acarina: Phytoseiidae) on sprayed apple trees at Bathurst (N. S. W.) - J. Austral. Entomol. Soc., 14: 217 - 219."]}

Neoseiulella cassiniae (Collyer) (Figure 20) Typhlodromus cassiniae Collyer 1982: 189; Chant and Yoshida-Shaul 1989: 1024-1026. Typhloctonus cassiniae (Collyer) Denmark and Rather 1984: 171; Moraes et al. 1986: 232. Neoseiulella (Typhloctona) cassiniae (Collyer) Denmark and Rather 1996: 70. Neoseiulella cassiniae (Collyer) Moraes et al. 2004: 292; Chant and McMurtry 2007: 147. Adult female (Figure 20a – e) Dorsal shield (Figure 20a) — Dorsal shield slightly reticulated throughout: length 473; width 235 (at level of s4) and 260 (at level of Z1). Six pairs of solenostomes on the dorsal shield: gd1, gd2, gd4, gd5, gd6 and gd9. No poroid visible. Sub-lateral setae (r3 and R 1) on the lateral margin. Dorsal shield bearing 19 pairs of setae all smooth: j1 20; j3 20; j4 11; j5 11; j6 12; J2 broken in the specimen examined (Denmark and Rather [1984, 1996] mentioned J2 11); J5 11; z2 11; z3 15; z4 14; z5 9; Z1 14; Z4 broken on the specimen examined (Denmark and Rather [1984, 1996] mentioned Z4 27); Z5 39; s4 20; s6 21; S2 19; S4 14; S5 14; sub-lateral setae r3 19 and R 1 18. Peritreme extending anteriorly to the level between j1-j3. Ventral shields (Figure 20b) — Sternal shield 71 long and 76 wide (at level of ST2), smooth with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 on separate platelets and a pair of small poroids accompanying ST4. Genital shield 167 long and 87 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla between genital shield and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield subtriangular-shaped, 157 long and 131 wide (at level of ZV2), smooth, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 36 long, smooth. Primary metapodal plate or inguinal sigillum 34 long and 7 wide. Spermatheca (Figure 20c) — Cervix 8 long, cupshaped. Chelicera (Figure 20d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 37 long, unidentate. Legs (Figure 20e) — Measurements of legs: leg I 391; leg II 323; leg III 337; leg IV 417. Eight setae (2-2/1, 2/0-1) on the genu II. Three macrosetae, 25, 30 and 32 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female holotype deposited in the New Zealand Arthropod Collection (NZAC), Landcare Research, Auckland, New Zealand. Adult male The adult of this species is unknown. Previous reports — N. cassiniae is only known from New Zealand, on Cassinia sp. (Asteraceae) and Hebe sp. (Plantaginaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 302-303, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Kanouh, M., Kreiter, S., Douin, M., Tixier, M.-S. (2012): Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females. Acarologia 52 (3): 259-348, DOI: 10.1051/acarologia/20122048, URL: http://dx.doi.org/10.1051/acarologia/20122048

Neoseiulella coreen Walter (Figure 21) Neoseiulella coreen Walter 1997: 336-337. Neoseiulella coreen Walter, Moraes et al. 2004: 292; Chant and McMurtry 2007: 147. Adult female (Figure 21 a – d) Dorsal shield (Figure 21a) — Dorsal shield is smooth: length 266 (261 – 270); width 132 (129 – 135) (at level of s4), 145 (139 – 150) (at level of Z1). Tuberculous ornements covering the lateral areas of the dorsal shield. Three pairs of large circular solenostomes (gd1, gd6, and gd9) and one pair of small and crescent-shaped solenostomes (gd8) on the dorsal shield. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae all smooth and rising to tubercles, except Z5 slightly serrated: j1 16 (12 – 20); j3 22 (20 – 24); j4 24 (23 – 25); j5 29 (28 – 30); j6 30 (28 – 31); J2 28 (26 – 30); J5 9 (8 – 10); z2 18; z3 25 (23 – 26); z4 25 (24 – 26); z5 16 (15 – 17); Z1 31 (30 – 32); Z4 25 (24 – 26); Z5 31 (30 – 32); s4 23; s6 29; S2 30 (29 – 31); S4 24 (22 – 25); S5 20 (19 – 21); sub-lateral setae r3 22 (21 – 23) and R1 14 (12 – 15). Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 21b) — Sternal shield weakly sclerotized, 58 (54 – 62) long and 56 (55 – 57) wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of small poroids. A pair of metasternal setae (ST4) inserted on separate platelets accompanied by a pair of small poroids. Genital shield 95 (93 – 96) long and 39 (35 – 42) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield 79 (74 – 83) long and 47 (44 – 49) wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 14 (13 – 15) long, smooth. Primary metapodal plate or inguinal sigillum 19 long and 2 (1.5 – 2) wide. Spermatheca (Figure 21c) — Cervix 4 (3 – 4) long, U-shaped. Chelicera (Figure 21d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 24 (23 – 25) long, edentate. Legs — Measurements of legs: leg I 270 (268 – 272); leg II 205 (195 – 215); leg III 191 (188 – 194); leg IV 271 (268 – 274). Eight setae (2-2/1, 2/0-1) observed on the genu II. No macroseta on legs. This species is easily distinghuished from the other species in the genus Neoseiulella by the tuberculous ornements on the dorsal shield, and by the small size and the shape of the ventrianal shield, which is narrowly vase-shaped. Material examined — The female holotype and five female paratypes deposited in the UQIC, Department of Entomology, University of Queensland, St Lucia, Australia. Adult male Described by Walter (1997). We were not able to borrow any male type specimens of this species. Previous reports — N. coreen is only known from Queensland (Australia) on rainforest trees., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 304-305, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Walter D. E. 1997 - Notes on Australian Typhlodrominae (Acari: Mesostigmata: Phytoseiidae), with descriptions of two new species of Neoseiulella Muma from Tropical Rainforests in far northern Queensland - Austral. J. Entomol., 36: 333 - 338. doi: 10.1111 / j. 1440 - 6055.1997. tb 01482. x","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella arinoi Moraza, Peaea-EstØvez and Ferragut (Figure 16) Neoseiulella arinoi Moraza, Peaea-EstØvez and Ferragut 2005: 109-112; Moraza and Peaea-Estevez 2006: 59; Chant and McMurtry 2007: 147. Adult female (Figure 16a – e) Dorsal shield (Figure 16a) — Dorsal shield distinctly reticulated throughout: length 409; width 191 (at level of s4) and 202 (at level of Z1). Seven pairs of solenostomes: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. No poroid visible. Sub-lateral setae (r3 and R 1) inserted on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except for Z4 and Z5 slightly serrated. Lateral setae on small tubercles: j1 34; j3 37; j4 18; j5 20; j6 36; J2 49; J5 12; z2 18; z3 30; z4 42; z5 24; Z1 46; Z4 75; Z5 80; s4 41; s6 52; S2 63; S4 60; S5 47; sub-lateral setae r3 41 and R 1 41. Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 16b) — Sternal shield 75 long and 78 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 on separate platelets, and a pair of small poroids accompanying ST4. Genital shield 122 long and 74 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded undethe genital shield. One pair of poroids close to the genital shield (ST 5) and 1 pair of poroids around the genital shield. Ventrianal shield longer than wide, 141 long and 110 wide (at level of ZV2), faintly striate, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 62 long, smooth. Dimensions of the primary metapodal plate or inguinal sigillum 37 long and 5 wide. Spermatheca (Figure 16c) — Cervix 4 long, Ushaped. Chelicera (Figure 16d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 33 long, unidentate. Legs (Figure 16e) — Measurements of legs: leg I 373; leg II 345; leg III 328; leg IV 416. Seven setae (2- 2/0, 2/0-1) on the genu II. One pointed macroseta, 82 long, on the basitarsus IV. On the specimen examined, two other macrosetae, 30 and 36, on genu and tibia IV, respectively. Even if Moraza et al. (2005) draw three macrosetae on the leg IV, they only mentioned in the text, the presence of one macroseta on this leg (on basitarsus). Material examined — A female paratype deposited in Museum of Zoology of University of Navarra, Spain. Adult male Described by Moraza et al. (2005). We were not able to borrow the male type specimen of this species. Previous reports — N. arinoi is only known from the Canary Islands. It was collected from lichens on dead log, from soil of "lapilli" and litter of Ficus carica (Moraceae), Castanea sativa (Fagaceae) and Echium virescens D. C. (Boraginaceae) (Moraza et al. 2005)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 295-297, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Moraza M. L., Peaea-Estevez M. A .. 2006 - A new species of Neoseiulella (Acari: Phytoseiidae) from the Macaronesian Region, Canary Islands - Zootaxa, 1366: 55 - 59.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Moraza M. L., Peaea-Estevez M. A., Ferragut F .. 2005 - Two new species of Neoseiulella Muma of the Canary Islands (Acari: Phytoseiidae) - Intern. J. Acarol., 31 (2): 107 - 112. doi: 10.1080 / 01647950508683659"]}

Neoseiulella elongata Ferragut and Peaea-Estevez (Figure 27) Neoseiulella elongata Ferragut and Peaea-Estevez 2003: 164; Moraes et al. 2004: 293; Moraza and Peaea- Estevez 2006: 58; Chant and McMurtry 2007: 147. Adult female (Figure 27a – e) Dorsal shield (Figure 27a) — Dorsal shield heavily reticulated: length 394; width 175 (at level of s4), 178 (at level of Z1), with four pairs of solenostomes: gd2, gd6, gd8, and gd9. Four pairs of poroids. Sublateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth: j1 20; j3 16; j4 12; j5 11; j6 14; J2 16; J5 11; z2 14; z3 16; z4 16; z5 11; Z1 20; Z4 23; Z5 29; s4 17; s6 19; S2 20; S4 21; S5 19; sub-lateral setae r3 23 and R1 24. Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of z2. It is noteworthy that the peritreme differs from the ones of the other species of the genus Neoseiulella as its anterior surface is striate. Ventral shields (Figure 27b) — Sternal shield 83 long and 73 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of small poroids. Two other pairs of sternal setae (ST3, ST4) on separate platelets. A pair of small poroids accompanies ST4. Genital shield 140 long and 66 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under genital shield. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subtriangular-shaped, 134 long and 93 wide (at level of ZV2), lightly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 20 long, smooth. Primary metapodal plate or inguinal sigillum 42 long and 4 wide. Spermatheca (Figure 27c) — Cervix 5 long, shallowly cup-shaped. Chelicera (Figure 27d) — One tooth and a pilus dentilis on the fixed digit. Movable digit 44 long, unidentate. Legs (Figure 27e) — Measurements of legs: leg I 335; leg II 281; leg III 275; leg IV 356. Seven setae (2- 2/0, 2/0-1) on the genu II. One macroseta, 61 long, on the basitarsus IV. Material examined — A female paratype deposited in the Department of Agroforestal Ecosystems, Universidad PolitØcnica of Valencia, Spain. Adult male The adult of this species is unknown. Previous reports — N. elongata is only known from the Canary Islands on Pinus canariensis C. Sm. (Pinaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 313, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Ferragut F., Peaea-Estevez M. A. 2003 - Phytoseiid mites of the Canary Islands (Acari: Phytoseiidae): 1. Gran Canaria Island - Intern. J. Acarol., 29 (2): 149 - 170.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Key to adult females of the genus Neoseiulella Muma 1. Ventrianal shield with three pairs of preanal setae (JV3 absent)...................................... 2 — Ventrianal shield with four pairs of preanal setae (JV3 present).................................... 11 2(1). Caudoventral setae JV4 present............. 3 — Caudoventral setae JV4 absent; dorsal shield with four pairs of solenostomes (gd2, gd4, gd5, gd6); spermatheca with U-shaped cervix; ventrianal shield with a pair of solenostomes; macrosetae on genu and tibia III, genu, tibia and basitarsus IV......................................... N. oleariae 3 (2). Presence of solenostomes on the ventrianal shield (gv3), genu II with seven setae (2-2/0, 2/0-1).................................................. 4 — Absence of solenostomes on the ventrianal shield (gv3); genu II with eight setae (2-2/1, 2/0-1); legs without macrosetae; tuberculous ornaments with large glands cover the dorsal shield....... N. coreen 4(3). Leg IV with three macrosetae, on genu, tibia and basitarsus................................... 5 — Leg IV with two thick and short macrosetae, on genu and basitarsus, tibia IV without macroseta; all dorsal and sub-lateral setae inserted to tubercles; spermatheca with shallowly cup-shaped......................................... N. elaeocarpi 5(4). Dorsal shield with four pairs of solenostomes: gd2, gd4, gd6, gd9 (gd1, gd5, and gd8 absent); fixed digit without pilus dentilis........................ 6 — Dorsal shield with more than four pairs of solenostomes: gd2, gd4, gd6 and gd9 always present; gd1, gd5 and gd8 present or absent...... 7 6(5). Most dorsal setae inserted on tubercles; spermatheca with U-shaped cervix; fixed digit with nine teeth; leg III with one macroseta, on genu......................................... N. corrugata All dorsal setae not inserted on tubercles; spermatheca with shallowly cup-shaped cervix; fixed digit with twelve teeth; leg III with two macrosetae, on genu and tibia.......................... N. dachanti 7(5). Dorsal shield with six pairs of solenostomes. 8 — Dorsal shield with five pairs of solenostomes.. 9 8(7). gd1 present, gd5 absent, gd2, gd4, gd6, gd8, and gd9 present; Z4 and Z5 very long (120 µm and 230 µm respectively), most of the other dorsal setae very short (5-15 µm long)................ N. cottieri — gd1 absent, gd5 present, gd2, gd4, gd6,gd8, gd9 present; Z4 and Z5 greatly shorter (about 60 and 90 µm respectively)........................ N. ashleyae 9(7). Leg III without macroseta.................. 10 — Leg III with macrosetae on genu and tibia....................... N. manuka e = N. glenfieldensis 10(9). Dorsal shield heavily reticulated; ventrianal shield with a large pair of solenostomes mediad to JV2..................... N. nesbitti = N. armidalensis — Dorsal shield lightly reticulated; ventrianal shield with a small pair of solenotomes posterior to JV2........................... N. novaezealandiae 11(1). Absence of macroseta on the leg IV....... 12 — Presence of at least one macroseta on the leg IV................................................. 19 12(11). Seven setae on the genu II............... 13 — Eight setae on the genu II.................... 18 13(12). One pair of solenostomes (gd8) on the dorsal shield................................ N. compta — More than one pair of solenostomes on the dorsal shield....................................... 14 14(13). Three pairs of solenostomes (gd2, gd6, gd9) on the dorsal shield.................. N. tuberculata — More than three pairs of solenostomes on the dorsal shield.................................... 15 15(14). Five solenostomes (gd1, gd2, gd6, gd8, gd9) on the dorsal shield...................................................... N. aceri = N. squamiger = N. aceris — Four solenostomes (gd2, gd5, gd6, gd9) on the dorsal shield.................................... 16 16(15). Peritreme reaching level of j1.... N. runiacus — Peritreme reaching level between j3 and z2... 17 17(16). Dorsal setae arising from tubercles, three teeth on the fixed digit of the chelicerae. N. sexapori — Dorsal setae not arising from tubercles, seven teeth on the fixed digit of the chelicerae........................................ N. arutunjani 18(12). Dorsal shield setae long (i.e. Z4 = 73; Z5 = 67; S4 = 75), peritreme reaching level of j1......................................... N. crassipilis — Dorsal shield setae shorter (i.e. Z4 = 38; Z5 = 44; S4 = 35), peritreme reaching level of z4.............................. N. tiliarum = N. formosa 19(11). One macroseta on the leg IV (on the basitarsus)............................................. 20 — Three macrosetae on the leg IV............... 29 20(19). Presence of solenostomes (gv3) on the ventrianal shield.................................... 21 — Absence of solenostomes (gv3) on the ventrianal shield........................................... 23 21(20). One pair of solenostome on the dorsal shield (gd9), peritreme reaching level of j1........ N. celtis — More than one pair of solenostomes on the dorsal shield, peritreme reaching level of j3......... 22 22(21). Six pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd6, gd8, gd9), spermatheca with U-shaped cervix, setae on the dorsal shield longer (i.e. R 1 = 41, S5 = 40, s6 = 43, s4 = 38, J2 = 36, j6 = 31)................................ N. montforti — Seven pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd8, gd9), spermatheca with cup-shaped cervix, setae on the dorsal shield shorter (i.e. R 1 = 25, S5 = 28, s6 = 26, s4 = 23, J2 = 15, j6 = 16).............................. N. neoviniferae 23(20). Six setae on the genu II........... N. carmeli — More than six setae on the genu II............ 24 24(23). Eight setae on the genu II................ 25 — Seven setae on the genu II.................... 26 25(24). No tooth on the mobile digit of the chelicerae, one tooth on the fixed digit, six pairs of solenostomes on the dorsal shield (gd1, gd2, gd5, gd6, gd8, gd9), some setae shorter (Z4 = 34, Z5 = 52, s6 = 26, S2 = 29)........................ N. perforata — Two teeth on the mobile digit of the chelicerae, six teeth on the fixed digit, seven pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd8, gd9), some setae longer (Z4 = 52, Z5 = 74, s6 = 43, S2 = 46).............................. N. steveni 26(24). Four pairs of solenostomes on the dorsal shield (gd2, gd6, gd8, gd9), spermatheca with cupshaped cervix, one tooth on the fixed digit of the chelicerae.............................. N. elongata — More than four pairs of solenostomes on the dorsal shield, spermatheca with U-shaped cervix, two teeth on the fixed digit of the chelicerae......... 27 27(26). Six pairs of solenostomes on the dorsal shield (gd1, gd2, gd5, gd6, gd8, gd9)..... N. litoralis — Seven pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd8, gd9)..... N. splendida 28(19). Absence of solenostomes (gv3) on the ventrianal shield........................... N. cassiniae — Presence of solenostomes (gv3) on the ventrianal shield........................................... 29 29(28). Presence of at least one macroseta on the leg III.............................................. 30 — Absence of macroseta on the leg III........... 32 30(29). One macroseta on the leg III (on the genu), spermatheca with U-shaped cervix, seven pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd8, gd9)..................... N. ferraguti — More than one macroseta on the leg III, spermatheca with cup-shaped cervix, five pairs of solenostomes on the dorsal shield............... 31 31(30). Three macrosetae on the leg III (genu, tibia, basitarsus), eight teeth on the fixed digit of the chelicerae, one tooth on the mobile digit, some setae longer (i.e. z4 = 29, z3 = 32, j3 = 52)...... N. myopori — Two macrosetae on the leg III (genu, tibia), ten teeth on the fixed digit of the chelicerae, 3 teeth on the mobile digit, some setae shorter (i.e. z4 = 18, z3 = 18, j3 = 36)............................... N. steeli 32(29). Eight setae on the genu II................ 33 — Seven setae on the genu II.................... 34 33(32). Dorsal seta lengths shorter (i.e. Z4 = 31, Z5 = 48, s4 = 23, s6 = 26, S2 = 20, S4 = 17, S5 = 17), six pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd9), spermatheca with cupshaped cervix............................. N. spaini — Dorsal seta lengths longer (i.e. Z4 = 88, Z5 = 117, s4 = 67, s6 = 71, S2 = 62, S4 = 63, S5 = 45), seven pairs of solenostomes on the dorsal shield (gd1, gd2, gd4, gd5, gd6, gd8, gd9), spermatheca with U-shaped cervix.................................. N. longiseta 34(32). Three pairs of solenostomes on the dorsal shield........................................... 35 — Seven pairs of solenostomes on the dorsal shield................................................. 36 35(34). Presence of solenostomes gd2, gd8, gd9, spermatheca with cup-shaped cervix, some dorsal setae shorter (i.e. Z1 = 22, Z4 = 31, S2 = 31, S4 = 30, S5 = 23, J2 = 24)................................................. N. vollsela = N. transitans = N. prunus — Presence of solenostomes gd5, gd8, gd9, spermatheca with U-shaped cervix, some dorsal setae longer (i.e. Z1 = 43, Z4 = 60, S2 = 62, S4 = 55, S5 = 50, J2 = 55)................................ N. elisae 36(34). Two teeth on the mobile digit of the chelicerae, six teeth on the fixed digit, some different dorsal seta lengths (J5 = 32, S4 = 41, S5 = 30)........................................ N. canariensis — One tooth on the mobile digit of the chelicerae, two teeth on the fixed digit, some different dorsal seta lengths (J5 = 12, S4 = 61, S5 = 47)...... N. arinoi, Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 337-340, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090

Neoseiulella formosa (Wainstein) (Figure 6) Typhlodromus formosus Wainstein 1958: 206; Abbasova 1966: 186. Typhlodromus (Seiulus) formosus (Wainstein) Ehara 1966: 16. Typhlodromus (Nesbitteius) formosus (Wainstein) Arutunjan 1970: 19. Typhloctonus formosus (Wainstein) Kolodochka 1974a: 26-27; Rivnay and Swirski 1980: 183. Neoseiulella formosa (Wainstein) Moraes et al. 2004: 297; Chant and McMurtry 2007: 147. Adult female (Figure 6a – d) Dorsal shield (Figure 6a) — Dorsal shield distinctly reticulated throughout: length 336; width 163 (at level of s4) and 180 (at level of Z1). Five pairs of large circular solenostomes: gd1, gd2, gd6, gd8, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except Z4 and Z5 slightly serrated: j1 19; j3 30; j4 19; j5 20; j6 18; J2 26; J5 11; z2 22; z3 28; z4 31; z5 17; Z1 33; Z4 38; Z5 44; s4 34; s6 34; S2 39; S4 35; S5 19; sub-lateral setae r3 28 and R1 21. Peritreme extending anteriorly to the level of z4. Ventral shields (Figure 6b) — Sternal shield 66 long and 65 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. ST3 on an elongate projection of the sternal shield. A pair of metasternal setae (ST4) on separate platelets with a pair of small poroids. Genital shield 117 long and 62 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla folded under the genital shield and situated between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield rectangular-shaped, 102 long and 66 wide (at level of ZV2), lightly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 24 long, smooth. Primary metapodal plate or inguinal sigillum 31 long and 3 wide. Spermatheca (Figure 6c) — Cervix 13 long, elongated and cup-shaped. Chelicera (Figure 6d) — One tooth and a pilus dentilis on the fixed digit. Movable digit 26 long, bearing one tooth. Legs — Measurements of legs: leg I 271; leg II 219; leg III 211; leg IV 275. Eight setae (2-2/1, 2/0-1) on the genu II. No macroseta on legs. Material examined — The female lectotype deposited in the ASU, Institute of Zoology, Academy of Sciences, Kiev, Ukraine. Adult male (Figure 6e, f) Dorsal shield chaetotaxy similar to the female, but sub-lateral seta R1 on the dorsal shield. Ventrianal shield 99 long and 139 wide, distinctly reticulated and bearing five pairs of preanal setae and four pairs of poroids. Spermatodactyl L-shaped with a terminal foot and a toe enlarged. This description is based on one of the paralectotype specimens (deposited in the ASU, Ukraine). Previous reports — N. formosa is only known from the West-Palaearctic area. Countries from which this species is reported are: Armenia (Arutunjan 1970, 1971); Caucasus region (Abbasova 1966); Georgia (Wainstein 1958; Samsoniya 1972; Wainstein and Vartapetov 1973); Hungary (Bozai 1996, 1997; Ripka 1998, 2006); Moldova (Wainstein 1973); Ukraine (Wainstein 1958; Kolodochka 1974b, 1978). Plants on which N. formosa was reported are: A. campestre, Acer sp. (Aceraceae); Cydonia sp., Malus sp., Mespilus germanica L., Mespilus sp., P. armeniaca, P. cerasifera, P. (Cerasus) avium, Prunus sp., P. spinosa, Pyrus sp., Rosa sp., Rubus sp., Sorbus sp. (Rosaceae); Alnus sp., C. avellana, Corylus sp. (Betulaceae); Cornus sp. (Cornaceae); Ficus carica L. (Moraceae); Juglans regia (Juglandaceae); Phaseolus sp. (Fabaceae); Q. cerris L., Quercus sp. (Fagaceae); U. laevis, U. scabra, Ulmus sp. (Ulmaceae). The examination of the type materials of both N. tiliarum and N. formosa shows that except for the position of sternal setae ST3 (on/off the sternal shield), all other morphological characters and measurements are similar. Furthermore, morphological and molecular analyses (Kanouh et al. 2010) showed that the position of seta ST3 is not a reliable diagnostic character and thus that N. tiliarum is a senior synonym of N. formosa. These results agree with the conclusions of Kolodochka (1986), Chant and Yoshida-Shaul (1989) and Denmark and Rather (1996)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 277-278, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Wainstein B. A. 1958 - New species of mites of the genus Typhlodromus (Parasitiformes: Phytoseiidae) from Georgia - Soobsh. Akad. Nauk. Gruz. SSR, 21 (2): 201 - 207.","Abbasova E. D. 1966 - Study on predators of tetranychid mites of Bol'shom Caucasia (Phytoseiidae) - Materialy Sessii Zakavkazskogo Soveta po Koordinatsii Nauchno-Issledovateliskikh Rabot po Zashchite Rasteniy, Baku, Azerbaijan, 184 - 187.","Ehara S. 1966 - A tentative catalogue of predatory mites of Phytoseiidae known from Asia, with descriptions of five new species from Japan - Mushi, 39: 9 - 30.","Arutunjan E. S. 1970 - Phytoseiid mites (Phytoseiidae) on agricultural crops in the Armenian SSR - Akademii Nauk Armyanskoi SSR, Otdelenie Biologicheskikh Nauk, Dissertatsii na Soiskanie Uchenoi Stepeni Candidata Biologrcheskikh Nauk, Zooliya, Armenia, 97: 1 - 31.","Kolodochka L. A. 1974 a - The predaceous phytoseiid mites (Parasitiformes: Phytoseiidae) from the forest steppe of the Ukranian SSR. Part II. Species of the genera Kampimodromus, Paraseiulus, Typhlodromus, Typhloctonus, Anthoseius, Phytoseius - Vest. Zool., (1): 25 - 29.","Rivnay T., Swirski E. 1980 - Four new species of phytoseiid mites (Acarina: Mesostigmata) from Israel - Phytoparasitica, 8: 173 - 187.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Arutunjan E. S. 1971 - Confinement of phytoseiid mites to biotopes and their morphological adaptations - Biolgicheskii Zhurnal Armenii, Akadrmiya Nauk Armyanskoi SSR, 24: 41 - 47.","Samsoniya T. I. 1972 - Species composition of predatory mites (Parasitiformes: Phytoseiidae) on stonefruit plants in eastern Georgia - Bull. Acad. Sc. Georg. SSR, 65 (1): 193 - 196.","Wainstein B. A., Vartapetov S. G. 1973 - Predatory mites of the family Phytoseiidae (Parasitiformes) of Adzharskaya ASSR - Akademiya Nauk Armyanskoy SSR, Biologicheskiy Zhurnal Armenii, 26 (2): 102 - 105.","Bozai J. 1996 - Data to the knowledge of predaceous mites of Hungary (Acari: Phytoseiidae, Phytoseiinae) - NovOnyvOdelem., 32: 521 - 525.","Bozai J. 1997 - Data to the fauna of predaceous mites of Hungary with the description of four new species (Acari: Phytoseiidae) - Fol. Entomol. Hungar., 58: 35 - 43.","Ripka G. 1998 - New data to the knowledge on the phytoseiid fauna in Hungary (Acari: Mesostigmata) - Acta Phytopathol. Entomol. Hung., 33: 395 - 405.","Ripka G. 2006 - Checklist of the Phytoseiidae of Hungary (Acari: Mesostigmata) - Folia Entomol., 67: 229 - 260.","Kolodochka L. A. 1974 b - Predaceous phytoseiid mites (Parasitiformes, Phytoseiidae) from the forest steppe of the Ukrainian SSR. Part III. A key to genera and species - Vest. Zool., (3): 87 - 89.","Kolodochka L. A. 1978 - Handbook on identifying of plant inhabiting phytoseiid mites - Nauka Dumka, Kiev. 78 pp.","Kanouh M., Tixier M. - S., Guichou S., Cheval B., Kreiter S. 2010 - Two synonymies within the genus Neoseiulella (Acari: Phytoseiidae): is the molecular evidence so evident? - Biol. J. Linn. Soc., 101: 323 - 344. doi: 10.1111 / j. 1095 - 8312.2010.01516. x","Kolodochka L. A. 1986 - On taxonomic status of two Typhloctonus species (Parasitiformes, Phytoseiidae) - Vest. Zool., (2): 26 - 34.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080"]}

Neoseiulella novaezealandiae (Collyer) (Figure 34) Typhlodromus novaezealandiae Collyer 1964: 635-637; Schicha 1980: 24-26; Collyer 1982: 188-189; Schicha 1987: 136-137; Chant and Yoshida-Shaul 1989: 1032- 1035. Neoseiulella novaezealandiae (Collyer) Moraes et al. 1986: 202; Moraes et al. 2004: 295; Chant and McMurtry 2007: 147. Neoseiulella (Neoseiulella) novaezealandiae (Collyer) Denmark and Rather 1996: 48-49. Adult female (Figure 34a – e) Dorsal shield (Figure 34a) — Dorsal shield heavily reticulated: length 400; width 235 (at level of s4), 231 (at level of Z1), and 262 (at level of S2). Five pairs of solenostomes on the dorsal shield: gd2, gd4, gd5, gd6, and gd9. One pair of poroids. Chant and Yoshida-Shaul (1989) draw and mentioned only four pairs of solenostomes (gd5 absent). However, the present examination of type material shows the presence of gd5, like in the drawings of Schicha (1987). Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z4 and Z5 serrated: j1 24; j3 27; j4 17; j5 15; j6 18; J2 21; J5 12; z2 17; z3 24; z4 23; z5 17; Z1 22; Z4 40; Z5 59; s4 24; s6 22; S2 25; S4 23; S5 21; sublateral setae r3 21 and R1 17. Presence of very few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 34b) — Sternal shield 85 long and 84 wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 121 long and 74 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields not discernible in the specimen examined. No pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield 139 long and 110 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent) and a pair of large solenostomes gv3 mediad to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 35 long, smooth. Primary metapodal plate or inguinal sigillum 30 long and 8 wide. Spermatheca (Figure 34c) — Cervix 5 long, shallowly cup-shaped. Chelicera (Figure 34d) — Nine teeth and a pilus dentilis on the fixed digit. Movable digit 33 (32 – 34) long, bearing three teeth. Legs (Figure 34e) — Measurements of legs: leg I 320; leg II 286; leg III 286; leg IV 391.. Seven setae (2- 2/0, 2/0-1) on the genu II. Three thick and pointed macrosetae, 30, 33 and 45 long, on the genu, tibia and basitarsus IV, respectively. Material examined — One female paralectotype deposited in the New Zealand Arthropod Collection (NZAC), Landcare Research, Auckland, New Zealand. Adult male Described by Collyer (1964) and Schicha (1980) and Denmark and Rather (1996). We were not able to borrow male type specimens of this species. Previous reports — N. novaezealandiae is only known from the Australasian area. The type specimens (30 females and 5 males) were collected at Kawau Island (New Zealand), on Metrosideros excelsa Gaertn (Myrtaceae). Collyer (1964, 1982,) reported that this species is distributed throughout New Zealand, on a wide range of native plants. Additional collections of this species have been reported from New South Wales and Queensland (Australia) (Schicha 1987; Walter 1999). Plants on which this species was collected are: Brachyglottis repanda (Asteraceae); Pseudopanax sp. (Araliaceae); Callicoma serratifolia Andrews (Cunoniaceae); Elaeocarpus dentatus (Elaeocarpaceae); Nothofagus sp., Quercus sp., Sophora tetraptera J.F.Mill. (Fagaceae); Eucalyptus sp., Leptospermum scoparium (Myrtaceae); Tacsonia mollissima Kunth, Passiflora sp. (Passifloraceae); Dacrydium cupressinum Sol. ex Lamb., Podocarpus sp. (Podocarpaceae); Eriobotrya sp., Malus domestica, Pyrus communis (Rosaceae); Elatostema rugosum A.Cunn. (Urticaceae); Lantana camara L., Vitex lucens T. Kirk (Verbenaceae). Remarks — N. novaezealandiae is very similar to N. nesbitti in setal and body measurements. However, differences are observed for the ornementation of dorsal shield (lightly reticulated on N. nesbitti, heavily reticulated on N. novaezealandiae) and the position and size of prenanal solenostomes (small and posterior to JV 2 in N. nesbitti, large and mediad to JV 2 in N. novaezealandiae). As several authors have showed the importance of such characters for species identification (Shicha 1980; Chant and Mc- Murtry 1994; Swirski et al. 1998; Chant and Mc- Murtry 2007; Okassa et al. 2009), we consider that these two species are separate specific entities., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 324-326, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1964 - The occurrence of some mites of the family Phytoseiidae in New Zealand, and descriptions of seven new species - Acarologia, 6: 632 - 646.","Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Walter D. E. 1999 - Cryptic inhabitants of a noxious weed: mites (Arachnida: Acari) on Lantana camara L. invading forests in Queensland - Austral. J. Entomol., 38: 197 - 200. doi: 10.1046 / j. 1440 - 6055.1999.00101. x","Swirski E., Ragusa S., Tsolakis H. 1998 - Keys to the phytoseiid mites (Parasitiformes: Phytoseiidae) of Israel - Phytophaga, 8: 85 - 154.","Okassa M., Tixier M. - S., Cheval B., Kreiter S. 2009 - Molecular and morphological evidence for a new species status within the genus Euseius (Acari: Phytoseiidae): consequences for taxonomy - Can. J. Zool., 87: 689 - 698. doi: 10.1139 / Z 09 - 057"]}

Neoseiulella armidalensis (Schicha and Elshafie) (Figure 15) Typhlodromus armidalensis Schicha and Elshafie 1980: 36; Schicha 1987: 140; Chant and Yoshida-Shaul 1989: 1042. Neoseiulella armidalensis (Schicha and Elshafie) Moraes et al. 1986: 201; Moraes et al. 2004: 291; Chant and McMurtry 2007: 147. Neoseiulella (Typhloctona) armidalensis (Schicha and Elshafie) Denmark and Rather 1996: 57. Adult female (Figure 15a – e) Dorsal shield (Figure 15a) — Dorsal shield smooth: length 416; width 214 (at level of s4) and 223 (at level of Z1). Five pairs of solenostomes: gd2, gd4, gd5, gd6, and gd9. No poroid visible. Sublateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z5 which is slightly serrated. Lateral setae S4 and S5 on small tubercles in the specimen examined: j1 22; j3 28; j4 13; j5 10; j6 12; J2 broken in the specimen examined (Chant and Yoshida-Shaul [1989] and Denmark and Rather [1996] mentioned J2 15); J5 11; z2 14; z3 21; z4 18; z5 10; Z1 16; Z4 33; Z5 65; s4 22; s6 17; S2 23; S4 22; S5 22; r3 14 and R1 15. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 15b) — Sternal shield 90 long and 83 wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 125 long and 73 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating the genital and ventrianal shields not discernible on the specimen examined. One pair of poroids close to the genital shield (ST 5) and no poroid visible around the genital shield. Ventrianal shield (in poor conditions in the specimen examined) 134 long and 117 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent) and a pair of small circular solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 42 long, smooth. Primary metapodal plate or inguinal sigillum 24 long and 7 wide. Spermatheca (Figure 15c) — Cervix 7 long, cupshaped. Chelicera (Figure 15d) — Eight or nine teeth and pilus dentilis on the fixed digit. Movable digit 35 long, bearing three teeth. Chant and Yoshida-Shaul (1989) mentioned that the pilus dentilis is not visible whereas Schicha (1980, 1987) and Denmark and Rather (1996) drew the pilus dentilis. Legs (Figure 15e) — Measurements of legs: leg I 390; leg II 315; leg III 305; leg IV 408. Seven setae (2- 2/0, 2/0-1) on the genu II. Three knobbed macrosetae, 37, 45 and 47 long, on genu, tibia and basitarsus IV, respectively. Material examined — One female paratype specimen (deposited in NSW Department of Primary Industries, Agricultural Scientific Collections Unit (Acarology), Orange Agricultural Institute, Australia. Adult male The male of this species is unknown. Previous reports — N. armidalensis is only known from New South Wales (Australia), on Malus sp. (Rosaceae) and Eucalyptus sp. (Myrtaceae). The present examination of the type materials of N. nesbitti and N. armidalensis shows that JV3 is absent; the fixed digit has a pilus dentilis, and that leg IV bears three macrosetae for the two species. Moreover, no difference in setal and body measurements between these two species is observed. We thus propose that N. nesbitti is a senior synonym of N. armidalensis. This conclusion does not agree with those of Chant and Yoshida-Shaul (1989) and Denmark and Rather (1996). The unique difference we observe between N. nesbitti and N. armidalensis is the nature of macrosetae on leg IV (pointed on N. nesbitti, knobbed on N. armidalensis). Further experiments would be consequently interesting to carry out in order to determine the reliabilty of such a difference for species diagnosis. From these latter considerations, we thus include 37 valid species in the genus Neoseiulella: N. aceri (Collyer) (senior synonym of N. squamiger [Wainstein], and provisionally of N. aceris [Lehman]); N. arinoi Moraza, Peaea-Estevez and Ferragut; N. arutunjani (Kuznetsov); N. ashleyae (Chant and Yoshida-Shaul); N. canariensis Ferragut and Pena-Estevez; N. carmeli (Rivnay and Swirski); N. cassiniae (Collyer); N. celtis (Denmark and Rather); N. compta (Corpuz-Raros) (suspected synonym of N. multispinosa [Tseng] [Chant and Yoshida-Shaul, 1989]); N. coreen Walter; N. corrugata (Schicha); N. cottieri (Collyer); N. crassipilis (Athias-Henriot and Fauvel); N. dachanti (Collyer); N. elaeocarpi (Schicha); N. elongata Ferragut and Pena-Estevez; N. elisae (Schicha and McMurtry), N. ferraguti Moraza and Pena-Estevez; N. litoralis (Swirski and Amitai); N. longiseta Moraza, Pena-Estevez and Ferragut; N. manukae (Collyer) (senior synonym of N. glenfieldensis [Schicha]); N. montforti (Rivnay and Swirski); N. myopori (Collyer); N. neoviniferae (Basha, Mahrous and Mostafa); N. nesbitti (Womersley) (senior synonym of N. armidalensis [Schicha and Elshafie]); N. novaezealandiae (Collyer); N. oleariae (Collyer); N. perforata (Athias-Henriot); N. runiacus (Kolodochka); N. spaini (Collyer); N. splendida Ferragut and Pena-Estevez; N. steeli (Shicha and Mc- Murtry); N. steveni (Schicha); N. tiliarum (Oudemans) (senior synonym of N. formosa [Wainstein]); N. tuberculata (Wainstein); N. sexapori (Karg and Edland); N. transitans (Gupta) (senior synonym of N. prunus [Denmark and Rather], and provisionally a junior synonym of N. vollsella [Chaudhri, Akbar and Rasool] [Chant and Yoshida-Shaul 1989; Denmark and Rather 1996)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 293-295, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E., Elshafie M. 1980 - Four new species of phytoseiid mites from Australia, and three species from America redescribed (Acari: Phytoseiidae) - J. Austral. Entomol. Soc., 19: 27 - 36. doi: 10.1111 / j. 1440 - 6055.1980. tb 00957. x","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080"]}

Neoseiulella corrugata (Schicha) (Figure 22) Typhlodromus corrugatus Schicha 1983: 120; Schicha 1987: 141-144; Chant and Yoshida-Shaul 1989: 1039- 1040. Seiulus corrugatus (Schicha) Moraes et al. 1986: 230. Neoseiulella (Neoseiulella) corrugata (Schicha) Denmark and Rather 1996: 50. Neoseiulella corrugata (Schicha) Moraes et al. 2004: 293; Chant and McMurtry 2007: 147. Adult female (Figure 22 a – e) Dorsal shield (Figure 22a) — Dorsal shield smooth: length 409; width 247 (at level of s4), 240 (at level of Z1) and 272 (at level of S2). Four pairs of solenostomes on the dorsal shield: gd2, gd4, gd6 and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all except j1 on tubercles. All dorsal and sub-lateral setae smooth, except Z4 and Z5 distinctly serrated: j1 29; j3 40; j4 28; j5 26; j6 30; J2 35; J5 12; z2 25; z3 37; z4 37; z5 24; Z1 37; Z4 66; Z5 117; s4 42; s6 40; S2 48; S4 36; S5 29; sub-lateral setae r3 23 and R1 22. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 22b) — Sternal shield 80 long and 92 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) inserted on separate platelets accompanied by a pair of small poroids. Genital shield 126 long and 83 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield 148 long and 120 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), with a pair of circular solenostomes gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 53 long, smooth. Primary metapodal plate or inguinal sigillum 29 long and 8 wide. Spermatheca (Figure 22c) — Cervix 6 long, Ushaped. Chelicera (Figure 22d) — Nine teeth, without pilus dentilis, on the fixed digit. Movable digit 33 long, bearing three teeth. Legs (Figure 22e) — Measurements of legs: leg I 443; leg II 351; leg III 361; leg IV 456. Seven setae (2-2/0, 2/0-1) on the genu II. One macroseta (32 long) on the genu III. Three other macrosetae, 47, 55 and 52 long, on genu, tibia and basitarsus IV, respectively. Material examined — The female holotype deposited in NSW Department of Primary Industries, Agricultural Scientific Collections Unit (Acarology), Orange Agricultural Institute, Australia. Adult male The male of this species is unknown. Previous reports — N. corrugata is only known from Australia on Malus domestica (Rosaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 305-307, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E. 1983 - New species, new records, and redescriptions of phytoseiid mites from Australia, Tahiti, and the African region (Acari: Phytoseiidae) - Intern. J. Entomol., 25 (2 - 3): 103 - 126.","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

(i). Neoseiulella schusteri (Youssef and El-Brollosy) Typhlodromus schusteri Youssef and El-Brollosy in Zaher 1986: 129. Neoseiulella schusteri (Youssef and El-Brollosy) Moraes et al. 2004: 295; Chant and McMurtry 2007: 147. This species was not included in the genus Neoseiulella in the previous revisions of Chant and Yoshida-shaul (1989) and Denmark and Rather (1984, 1996). However, it was considered a member of the genus Neoseiulella in the last world catalogue of the family Phytoseiidae (Moraes et al. 2004), and then in the revision of the family (Chant and Mc- Murtry 2007). As it has been impossible to borrow and thus to observe the type specimens (deposited in Plant Protection Department, Faculty of Agriculture, Cairo University, Egypt), an attentive examination of the dorsal chaetotaxy of the female, based on the original description, was carried out. We observe that the dorsal seta z3 is absent. This latter character with the presence of nine pairs of lateral setae on the dorsal shield (j3, z2, z4, s4, s6, S2, S4, S5, Z5) keys to the species Cydnoseius negevi (Swirski and Amitai), tribe Galendromimini (sub-family Typhlodrominae). El-Brollosy (personal comm. 2008) confirmed that N. schusteri had been misidentified, and that it is a junior synonym of C. negevi., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 266, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Zaher M. A. 1986 - Predaceous and nonphytophagous mites (Nile Valley and Delta) - Survey and ecological studies on phytophagous, predaceous and soil mites in Egypt. PL 480 Programme USA, Project EG-ARS- 30, Grant No. FG-EG- 139, 567 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080"]}

Neoseiulella runiacus (Kolodochka) (Figure 37) Typhloctonus runiacus Kolodochka 1980: 64-65; Denmark and Rather 1984: 75-76; Moraes et al. 1986: 233; Kolodochka 2009: 488-489. Typhlodromus runiacus (Kolodochka) Chant and Yoshida-Shaul 1989: 1015-1017. Neoseiulella (Typhloctona) runiaca (Kolodochka) Denmark and Rather 1996: 63-64. Neoseiulella runiaca (Kolodochka) Moraes et al. 2004: 295; Chant and McMurtry 2007: 147. Adult female (Figure 37a – d) Dorsal shield (Figure 37a) — Dorsal shield heavily reticulated throughout: length 382; width 220 (at level of s4), 271 (at level of Z1). Four pairs of large solenostomes on the dorsal shield: gd2, gd5, gd6, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except J5 and Z5 serrated. Most dorsal setae thick and on tubercles: j1 18; j3 23; j4 18; j5 18; j6 28; J2 32; J5 17; z2 17; z3 26; z4 25; z5 20; Z1 30; Z4 39; Z5 44; s4 27; s6 28; S2 31; S4 26; S5 24; sub-lateral setae r3 22 and R1 15. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 37b) — Sternal shield 51 long and 64 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 and ST4 are inserted on separates platelets and a pair of small poroids accompanying ST4. Genital shield 136 long and 67 wide (at level of ST5), smooth. Six elongate platelets or genital sigilla situated between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield subquadrate-shaped, 110 long and 108 wide (at level of ZV2), distinctly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of circular solenostomes gv3 posterior to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 16 long, smooth. Primary metapodal plate or inguinal sigillum 45 long and 4 wide. Spermatheca (Figure 37c) — Cervix 9 long, cupshaped. Chelicera (Figure 37d) — Four teeth and a pilus dentilis on the fixed digit. Movable digit 23 long, bidentate. Legs — Measurements of legs: leg I 250; leg II 230; leg III 240; leg IV 300. Seven setae (2-2/0, 2/0- 1) on the genu II. No macroseta on legs. Material examined — The female holotype deposited in the ASU, Institute of Zoology, Academy of Sciences, Kiev, Ukraine. Adult male (Figure 37 e-f) Dorsal shield similar to but smaller than female. Ventrianal shield 120 long and 142 wide, distinctly reticulated and bearing 5 pairs of preanal setae and three pair of poroids. Spermatodactyl L-shaped. This description is based on a paratype specimen. Previous reports — The type specimens of N. runiacus were collected from Zakarpattia region (Ukraine) on Alnus viridis (Chaix) DC. (Betulaceae) and Fagus silvatica L. (Fagaceae) (Kolodochka 1980, 2009). Chant and Yoshida-Shaul (1989) indicated that a specimen originally identified as Typhlodromus aceri, collected in the Lake District (UK), was in fact N. runiacus. Remarks — N. runiacus is similar to N. sexapori. Except for characteristics of dorsal setae (thick on N. runiacus; simple on N. sexapori), the nature of J5 (serrated on N. runiacus; smooth on N. sexapori), and the length of the peritreme (extending to the level of j1 on N. runiacus; extending to the level between z2-j3 on N. sexapori), all other morphological characters are similar. As several authors have already shown the importance of such characters for species differentiation (Athias-Henriot 1977; Chant and Yoshida- Shaul 1983; Chant and McMurtry 1994; Ragusa and Tsolakis 1998; Swirski et al. 1998; Chant and Mc- Murtry 2007), we consider that N. runiacus and N. sexapori are two valid species., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 329-331, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Kolodochka L. A. 1980 - New species of phytoseiid mites from the fauna of the USSR (Parasitiformes: Phytoseiidae) - Vest. Zool., (2), 64 - 70.","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Kolodochka L. A. 2009 - A review of predaceous mites of the genus Typhloctonus Muma (Parasitiformes, Phytoseiidae) in Ukraine with the description of unknown male of T. tuberculatus - Vest. Zool., 43 (6): 481 - 494.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Athias-Henriot C. 1977 - Nouvelles notes sur les Amblyseiini. III. Sur le genre Cydnodromus: RedOfinition, composition (Parasitiformes, Phytoseiidae) - Entomophaga, 22: 61 - 73. doi: 10.1007 / BF 02372991","Ragusa S., Tsolakis H. 1998 - Phytoseiid mites (Parasitiformes, Phytoseiidae) of Lesvos Island (Greece) with a description of a new species - Entomol. Hellen., 12: 55 - 64.","Swirski E., Ragusa S., Tsolakis H. 1998 - Keys to the phytoseiid mites (Parasitiformes: Phytoseiidae) of Israel - Phytophaga, 8: 85 - 154."]}

(ii). Neoseiulella eleglidus (Tseng) Typhlodromus (Typhlodromus) eleglidus Tseng 1983: 64-66. Typhlodromus eleglidus (Tseng) Wu et al. 1991: 82. Neoseiulella eleglidus (Tseng) Moraes et al. 2004: 293; Chant and McMurtry 2007: 147. This species was not included in the genus Neoseiulella in the revisions of Chant and Yoshida-shaul (1989) and Denmark and Rather (1984, 1996). However, it was included in the genus Neoseiulella in the last world catalogue (Moraes et al. 2004), and then in the revision of the family (Chant and Mc- Murtry 2007). It has been impossible to borrow and thus to observe the type specimens (deposited in Plant Quarantine Laboratory, Tainan Branch, Bureau of Commodity Inspection and Quarantine, Tainan, Taiwan). However, the original description of this species shows 18 pairs of setae on the dorsal shield (j1, j3, j4, j5, j6, J2, J5, z2, z3, z4, z5, Z4, Z5, s4, s6, S2, S4, S5) and 2 sub-lateral setae (r3 and R 1). Because of the absence of seta Z1 and presence of seta S5, this dorsal setal pattern (12A: 8B) keys to the genus Typhlodromus (Anthoseius) De Leon (tribe Typhlodromini, sub-family Typhlodrominae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 266, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Tseng Y. H. 1983 - Further study on phytoseiid mites from Taiwan (Acarina: Mesostigmata) - Chin. J. Entomol., 3: 33 - 74.","Wu W. N., Lan W. M. 1991 - A new species of the genus Iphiseius from Guangdong Province, China (Acari: Phytoseiidae) - Acta Zootax. Sin., 16 (2): 191 - 193.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080"]}

Neoseiulella litoralis (Swirski and Amitai) (Figure 29) Typhloctonus litoralis Swirski and Amitai 1984: 73-76. Neoseiulella (Typhloctona) litoralis (Swirski and Amitai) Denmark and Rather 1996: 71-72. Neoseiulella litoralis (Swirski and Amitai) 1997b: 37; Moraes et al. 2004: 294; Chant and McMurtry 2007: 147. Adult female (Figure 29a – e) Dorsal shield (Figure 29a) — Dorsal shield heavily reticulate: length 393 (383 – 403); width 184 (178 – 190) (at level of s4), 197 (188 – 205) (at level of Z1), with six pairs of solenostomes: gd1, gd2, gd5, gd6, gd8, and gd9. One pair of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae all smooth (in one paratype specimen, Z5 is slightly serrated): j1 23 (21 – 24); j3 24 (22 – 26); j4 15 (13 – 17); j5 13 (11 – 14); j6 15 (14 – 16); J2 18 (17 – 18); J5 14; z2 13 (11 – 15); z3 22 (20 – 24); z4 18 (17 – 19); z5 14 (12 – 15); Z1 19 (18 – 20); Z4 50 (48 – 51); Z5 63 (60 – 66); s4 20 (19 – 21); s6 28 (26 – 30); S2 37 (35 – 38); S4 29 (26 – 31); S5 29 (26 – 31); sub-lateral setae r3 23 and R1 23 (20 – 25). Peritreme extending anteriorly to the level of j3 or between z2 – j3. Ventral shields (Figure 29b) — Sternal shield 59 (55 – 63) long and 69 wide (at level of ST2), smooth, with two pairs of setae (ST1 and ST2) and two pairs of poroids. The third and fourth pairs of sternal setae (ST3, ST4) on separate platelets and a pair of small poroids accompanying ST4. Genital shield 130 (122 – 137) long and 67 (66 – 68) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. No pair of poroids close to the genital shield (ST 5) and 1 pair of poroids around the genital shield. Ventrianal shield subtriangular-shaped with conspicuous waist, much longer than wide, 140 (137 – 143) long and 98 (92 – 104) wide (at level of ZV2), reticulate with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 45 (37 – 52) long, smooth. Primary metapodal plate or inguinal sigillum 41 (37 – 44) long and 5 wide. Spermatheca (Figure 29c) — Cervix 4 long, Ushaped. Chelicera (Figure 29d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 37 (36 – 38) long, unidentate. Legs (Figure 29e) — Measurements of legs: leg I 338 (331 – 344); leg II 299 (295 – 303); leg III 296 (292 – 300); leg IV 395 (386 – 404). Seven setae (2-2/0, 2/0-1) on the genu II. One pointed macroseta 41 (36 – 45) long, on the basitarsus IV. Material examined — Three female paratypes deposited in the Division of Entomology, Agricultural Research Organization, Bet Dagan, Israel. Acarologia 52(3): 259–348 (2012) Adult male (Figure 29f, g) Dorsal shield chaetotaxy similar to but smaller than the female. Sub-lateral seta R1 inserted on the dorsal shield (different from the female, on which R1 on lateral margin). Six pairs of solenostomes on the dorsal shield. Ventrianal shield 124 long and 142 wide, distinctly reticulated, bearing six pairs of preanal setae and a pair of poroids. Spermatodactyl L-shaped. This description is based on a paratype specimen. Previous reports — N. litoralis is only known from the West- Palaearctic area. Countries from which this species was reported are: Israel (Swirski and Amitai 1984; Klein et al. 1994); Spain (Ferragut et al. 2008). Plant supports on which this species was collected are: Ammophila arenaria (L.) Link (Poaceae); Achillea maritima (L.) Ehrend and Y.P.Guo, Artemisia monosperma Delile, Limbarda crithmoides (L.) Dumort (Asteraceae); Crucianella maritima L. (Rubiaceae); Echiohilon fruticosum Desf. (Boraginaceae); Lotus creticus L., Retama raetam (Forssk.) Webb. (Fabaceae); Oenothera drummondi Hook (Onagraceae); Polygonum equisetiforme Sibth and Sm. (Polygonaceae); Scrophularia syriaca A.D.C. (Scrophulariaceae); Vitis vinifera subsp. sylvestris (C.C.Gmel.) Hegi (Vitaceae). This species was recently collected in Villeneuve-lŁs-Maguelone (HØrault, Mediterranean Coast) on red swampfire (Salicornia rubra Nelson) and sea purslane (Halimione portulacoides (L.) Aellen (Amaranthaceae) (Kanouh et al. 2010)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 316-318, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Swirski E., Amitai S. 1984 - Notes on phytoseiid mites (Mesostigmata: Phytoseiidae) from the Mediterranean littoral zone of Israel, with a description of a new species of Typhloctonus - Israel J. Entomol., 18: 71 - 82.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Klein Z., Kuslitzky W., Zarabi L. 1994 - List of Israeli mites (Acarina) - Ministry of Agriculture. Plant Protection and Inspection Services, Entomological Section. Bet Dagan, 51 pp.","Kanouh M., Tixier M. - S., Guichou S., Cheval B., Kreiter S. 2010 - Two synonymies within the genus Neoseiulella (Acari: Phytoseiidae): is the molecular evidence so evident? - Biol. J. Linn. Soc., 101: 323 - 344. doi: 10.1111 / j. 1095 - 8312.2010.01516. x"]}

Neoseiulella elisae (Schicha and McMurtry) (Figure 1) Typhlodromus elisae Schicha and McMurtry 1986: 177; Chant and Yoshida-Shaul 1989: 1023-1024; Schicha 1987: 152, 154. Typhlodromus (Anthoseius) elisae Schicha and McMurtry Moraes, 2004: 322. Adult female (Figure 1a – d) Dorsal shield (Figure 1a) — Dorsal shield strongly reticulated, more heavily sclerotized on dorsocentral area of the shield: length 485; width 300 (at level of s4 and at level of Z1). Three pairs of solenostomes: gd5, gd8, and gd9 and 1 pair of poroids clearly visible. Sub-lateral setae (r3 and R1) on the lateral margin. Bearing 19 pairs of setae, all dorsal and sub-lateral setae smooth: j1 22; j3 33; j4 18; j5 28; j6 42; J2 55; J5 18; z2 25; z3 25; z4 30; z5 30; Z1 43; Z4 60; Z5 55; s4 43; s6 50; S2 63; S4 55; S5 50; sub-lateral setae r3 38 and R1 40. Peritreme extending anteriorly to level of j1. Ventral shields (Figure 1b) — Sternal shield lightly reticulated, 70 long and 50 wide (at level of ST2), with three pairs of setae (ST1, ST2 and ST3) and two pairs of small solenostomes. The fourth pairs, the metasternal setae (ST4) on separate platelets. A pair of small pores accompanying ST4. Genital shield lightly reticulated, 145 long and 70 wide (at level of ST5). Four elongate platelets or genital sigilla separating the genital and ventrianal shields not discernible on the specimen examined. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subtriangular, distinctly reticulated throughout, 143 long and 133 wide (at level of ZV2), with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), and a pair of small circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 50 long, smooth. Spermatheca (Figure 1c) — Cervix 20 long, coneshaped. Chelicera — Not visible. Legs (Figure 1d) — Measurements of legs: leg I 356; leg II 270; leg III 288; leg IV 388. Seven setae (2-2/0, 2/0-1) on the genu II. Legs much shorter than the dorsal shield and macrosetae only on leg IV, SgeIV: 13, Sti IV: 40 with tip tapered, St IV: 28. Material examined — The female holotype deposited in the New South Wales Department of Primary Industries, Agricultural Scientific Collections Unit (Acarology), Orange Agricultural Institute, Forest Road, Orange NSW 2800, Australia. Adult male — The male of this species is unknown. Previous reports — N. elisae is only known from Western Australia on Eucalyptus sp. (Myrtaceae). We thus consider 44 valid species in the genus Neoseiulella in the followings parts of this paper., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on page 267, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E., McMurtry J. 1986 - Two new and two known species of Typhlodromus Scheuten (Acari: Phytoseiidae) from Australia - J. Austral. Entomol. Soc., 25: 177 - 183. doi: 10.1111 / j. 1440 - 6055.1986. tb 01100. x","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494."]}

Neoseiulella steveni (Schicha) (Figure 41) Typhlodromus steveni Schicha 1987: 135-136; Chant and Yoshida-Shaul 1989: 1026-1028. Neoseiulella (Typhloctona) steveni (Schicha) Denmark and Rather 1996: 64-66. Neoseiulella steveni (Schicha) Moraes et al. 2004: 296; Chant and McMurtry 2007: 147. Adult female (Figure 41a – e) Dorsal shield (Figure 41a) — Dorsal shield smooth, except for the lateral area of the podosoma lightly reticulated: length 359 (344 – 373); width 175 (166 – 183) (at level of s4) and 190 (183 – 197) (at level of Z1). Seven pairs of solenostomes: gd1, gd2, gd4, gd5, gd6, gd8, and gd9. No poroid visible. Sublateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z5 slightly serrated: j1 24 (34 – 24); j3 31 (31 – 32); j4 17 (15 – 18); j5 16 (15 – 17); j6 17 (16 – 18); J2 21 (20 – 22); J5 11 (10 – 12); z2 18 (16 – 19); z3 33 (30 – 35); z4 30 (29 – 30); z5 18 (17 – 18); Z1 20 (18 – 21); Z4 52 (50 – 53); Z5 74 (72 – 75); s4 38 (37 – 39); s6 43 (40 – 45); S2 46 (44 – 48); S4 25 (23 – 26); S5 25 (23 – 26); sub-lateral setae r3 28 (27 – 28) and R1 20 (19 – 21). Peritreme extending anteriorly to the level between j1-j3. Ventral shields (Figure 41b) — Sternal shield 73 (66 – 80) long and 77 (75 – 79) wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 128 (124 – 131) long and 63 (61 – 65) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields not discernible in the specimen examined. One pair of poroids close to the genital shield (ST 5) and 1 pair of poroids around the genital shield. Ventrianal shield 124 (121 – 126) long and 98 (94 – 101) wide (at level of ZV2), smooth to faintly striate, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 49 (47 – 51) long, smooth. Primary metapodal plate or inguinal sigillum 29 (28 – 30) long and 5 wide. In a paratype specimen: secondary metapodal plate or inguinal sigillum 21 long and 3 wide. Spermatheca (Figure 41c) — Cervix 4 long, Ushaped. Chelicera (Figure 41d) — Six teeth and a pilus dentilis observed on the fixed digit. Movable digit 29 (27 – 31) long, bidentate. Legs (Figure 41e) — Measurements of legs: leg I 369 (364 – 374); leg II 339 (328 – 350); leg III 298 (294 – 302); leg IV 434 (422 – 443). Eight setae (2-2/1, 2/0-1) on the genu II. One pointed macroseta 48 (42 – 54) long, on the basitarsus IV. Material examined — The female holotype and two female paratypes deposited in NSW Department of Primary Industries, Agricultural Scientific Collections Unit (Acarology), Orange Agricultural Institute, Australia. Adult male Described by Schicha (1987) and Denmark and Rather (1996). We were not able to borrow the male type specimens of this species. Previous reports — N. steveni is only known from New South Wales (Australia), on Banksia sp. (Proteaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 336-337, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella cottieri (Collyer) (Figure 23) Typhlodromus cottieri Collyer 1964: 640-641; Schicha 1980: 20-22; Collyer 1982: 189-190; Schicha 1987: 138-140; Chant and Yoshida-Shaul 1989: 1029-1032. Typhloctonus (Tasmanidromus) cottieri (Collyer) Wainstein 1977: 1415. Neoseiulella (Neoseiulella) cottieri (Collyer) Denmark and Rather 1996: 53. Neoseiulella cottieri (Collyer) Moraes et al. 1986: 248; Chant and McMurtry 1994: 248; Moraes et al. 2004: 293; Chant and McMurtry 2007: 147. Adult female (Figure 23a – e) Dorsal shield (Figure 23a) — Dorsal shield smooth: length 385; width 215 (at level of s4) and 255 (at level of Z1). Six pairs of solenostomes observed: gd1, gd2, gd4, gd6, gd8 and gd9. Four pairs of poroids. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except for Z4 and Z5 which are slightly serrated: j1 29; j3 43; j4 5; j5 6; j6 9; J2 8; J5 11; z2 6; z3 48; z4 7; z5 6; Z1 6; Z4 121; Z5 232; s4 67; s6 7; S2 9; S4 10; S5 11; sub-lateral setae r3 15 and R1 12. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 23b) — Sternal shield 77 long and 70 wide (at level of ST2), smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 120 long and 68 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla situated between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 2 pairs of poroids around the genital shield. Ventrianal shield, longer than wide 124 long and 101 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent), with a pair of circular solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) inserted on the integument surrounding the ventrianal shield. JV5 93 long, smooth. Primary metapodal plate or inguinal sigillum 25 long and 7 wide. Spermatheca (Figure 23c) — Cervix 11 long, Ushaped. Chelicera (Figure 23d) — Eight teeth and a pilus dentilis on the fixed digit. Movable digit 30 long, bearing three teeth. Legs (Figure 23e) — Measurements of legs: leg I 403; leg II 318; leg III 313; leg IV 427. Seven setae (2-2/0, 2/0-1) on the genu II. One pointed macroseta (38 long) on the genu III. Three other pointed macrosetae, 82, 65 and 77 long, on genu, tibia and basitarsus IV, respectively. This species is easily distinguished from all other species of this genus by the lengths of Z4 (121) and Z5 (232). Material examined — The female lectotype deposited in the BMNH, the British Museum of Natural History, Cromwell Road, London, UK. Adult male Described by Collyer (1964) and Schicha (1980). We were not able to borrow the male type specimens of this species. Previous reports — The type specimens of N. cottieri were collected from New Zealand on thistle (Asteraceae). This species is only known from the Australasian area: Australia (Wainstein 1977; Schicha 1980, 1987) and New Zealand (Collyer 1964, 1982). Plants on which this species was collected are: Astelia sp. (Asteliaceae); Cassiniae sp. (Asteraceae); Fragaria x ananassa Duchesne, Malus domestica (Rosaceae); Hebe sp. (Plantaginaceae); Humulus lupulus L. (Cannabaceae); Trifolium repens L. (Fabaceae) and on moss samples., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 307-308, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1964 - The occurrence of some mites of the family Phytoseiidae in New Zealand, and descriptions of seven new species - Acarologia, 6: 632 - 646.","Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Wainstein B. A. 1977 - A contribution to the fauna of the family Phytoseiidae (Parasitiformes) in Australia - Zool. Zh., 56: 1413 - 1416.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella aceri (Collyer) (Figure 2) Typhlodromus aceri Collyer 1957: 199-200; Chant 1958: 626; Hirschmann 1962: 12; Livshitz and Kuznetsov 1972: 20; Chant and Yoshida-Shaul 1989: 1013. Typhlodromus (Typhlodromus) aceri (Collyer) Chant 1959: 65; Westerboer and Bernhard 1963: 565- 568. Typhloctonus aceri (Collyer) Muma 1961: 299; Denmark and Rather 1984: 166-167; Kolodochka 1986: 30-31; Moraes et al. 1986: 232; Kolodochka 2009: 486-487. Typhlodromus (Nesbitteius) aceri (Collyer) Wainstein 1962: 23. Seiulus aceri (Collyer) Abbasova 1972: 22; Karg and Edland 1987: 387; Steeghs et al. 1993: 24. Seiulus (Typhloctonus) aceri (Collyer) Beglyarov 1981: 19. Neoseiulella (Typhloctona) aceri (Collyer) Denmark and Rather 1996: 60. Neoseiulella aceri (Collyer) Moraes et al. 2004: 290; Chant and McMurtry 2007: 147. Adult female (Figure 2a – d) Dorsal shield (Figure 2a) — Dorsal shield distinctly reticulated throughout: length 318 (312 – 326), width 170 (166 – 174) (at level of s4) and 201 (198 – 207) (at level of Z1). Five pairs of small solenostomes on the dorsal shield: gd1, gd2, gd6, gd8, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all serrated: j1 16 (15 – 17); j3 26 (25 – 26); j4 18 (17 – 18); j5 18 (17 – 18); j6 24 (23 – 25); J2 31 (30 – 31); J5 14 (13 – 14); z2 21 (20 – 22); z3 26; z4 27; z5 16 (15 – 16); Z1 30 (28 – 31); Z4 35; Z5 38 (37 – 39); s4 31; s6 31 (30 – 31); S2 31; S4 26 (25 – 26); S5 14 (13 – 15); sub-lateral setae r3 21 (20 – 21) and R1 20. A pair of dorsal setae J1 (32) on one of the syntype females examined. Peritreme extending anteriorly to the level of j3 or between z2-j3. Ventral shields (Figure 2b) — Sternal shield 37 (34 – 40) long and 53 (51 – 58) wide (at level of ST2), is smooth with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids (posterior to ST1; anterior to ST3). The pair of sternal setae ST3 on an elongate projection of the sternal shield. A pair of metasternal setae (ST4) on separate platelets with a pair of small poroids. Genital shield 126 (118 – 132) long and 60 (58 – 61) wide (at level of ST5), smooth. Four elongate platelets or genital sigilla between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subquadrate-shaped, 99 (96 – 101) long and 83 (82 – 86) wide (at level of ZV2), distinctly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and a pair of solenostomes gv3 posterior to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 14 (13 – 14) long, smooth. Primary metapodal plate or inguinal sigillum 30 (26 – 31) long and 3 (2 – 4) wide. Spermatheca (Figure 2c) — Cervix 4 (3 – 5) long, cup-shaped, with an enlarged atrium. Chelicera (Figure 2d) — Two teeth and a pilus dentilis on the fixed digit. Movable digit 22 (20 – 23) long, edentate. Legs. Measurements of legs: leg I 225 (210 – 231); leg II 204 (190 – 222); leg III 212 (190 – 225); leg IV 249 (233 – 270). Seven setae (2-2/0, 2/0-1) on the genu II. No macroseta on legs. Material examined. Four female syntypes deposited in the British Museum of Natural History = BMNH, Cromwell Road, London, UK. Adult male (Figure 2e, f) Described by Collyer (1957), Chant (1958) and Denmark and Rather (1984, 1996). Dorsal shield chaetotaxy similar to the female. Ventrianal shield 97 long and 138 wide, distinctly reticulated and with five pairs of preanal setae and three pairs of poroids. Spermatodactyl with a terminal foot, and with a non-enlarged toe. This description is based on one male syntype. Previous reports — N. aceri is only known from the West-Palaearctic and Nearctic areas. However, its occurrence in the Nearctic area (three records) might be due to an introduction. Countries from which this species is reported are: Azerbaijan (Abbasova 1972); Belgium (AndrØ 1986); Croatia (Tixier et al. 2010); Czech Republic (Kabicek 2003; Kabicek and Rehakova 2004; Kabicek 2005, 2010;); England (Collyer 1957); Finland (Tuovinen and Rokx 1991; Tuovinen 1993); France (Chant and Yoshida-Shaul 1989); Germany (Balder et al. 1999); Hungary (Szabo 1980; Ripka 1998, 2006); Italy (Ragusa and Paoletti 1985; Duso et al. 1993, 2004); Moldova (Beglyarov and Malov 1977); Norway (Karg and Edland 1987); Sweden (Steephs et al. 1993); Turkey (Cobanoglu 1996, 2000); Ukraine (Livshitz and Kuznetsov 1972; Akimov et al. 2007; Kolodochka and Omeri 2007; Kolodochka 2009); USA (Chant 1959 b; Chant and Yoshida-Shaul 1989; Congdon 2002). Neoseiulella aceri was collected from a wide range of plant supports: Acer campestre L., A. macrophyllum Pursh, A. platanoides L., A. pseudoplatanus L., Acer sp. (Aceraceae); Aesculus hippocastanum L. (Hippocastanaceae); Alnus incana (L.) Moench, Carpinus betulus L., Corylus avellana L. (Betulaceae); Juglans regia L. (Juglandaceae); Juniperus sabina L. (Cupressaceae); Morus alba L. (Moraceae); Prunus (Cerasus) sp., Prunus armeniaca L., Prunus sp., Rubus sp. (Rosaceae); Quercus ilex L. (Fagaceae); Trifolium pratense L. (Fabaceae); Zelkova sp. (Ulmaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 269-271, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1957 - Two new species of the genus Typhlodromus Scheuten, 1857 (Acarina: Phytoseiidae) - Annu. Magaz. Natur. Hist., 12: 199 - 203.","Chant D. A. 1958 - Immature and adult stages of some British Phytoseiidae Berl., 1916 (Acarina) - J. Linn. Soc. London, Zool., 43: 599 - 643.","Hirschmann W. 1962 - Gangystematik der Parasitiformes - Acarologie Schriftenreihe fur Vergleichende Milbenkunde, Hirschmann-Verlag, Furth / Bay. 5 (5 - 6): 80 pp + 32 plates.","Livshitz I. Z., Kuznetsov N. N. 1972 - Phytoseiid mites from Crimea (Parasitiformes: Phytoseiidae) [in Russian] - In: Pests and diseases of fruit and ornamental plants. Proceed. All-Union V. I. Lenin Academy of Agricultural Science, The State Nikita Botanical Gardens, 61: 13 - 64.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Chant D. A. 1959 - Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species - Can. Entomol., Supplement 12, 166 pp.","Westerboer I., Bernhard F. 1963 - Die Familie Phytoseiidae Berlese 1916 - In: Stammer, H. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina. Band II, Mesostigmata I. pp. 451 - 791.","Muma M. H. 1961 - Sub-familes, genera, and species of Phytoseiidae (Acarina: Mesostigmata) - Flor. St. Mus. Bull., 5 (7): 267 - 302.","Denmark H. A., Rather A. Q. 1984 - Revision of the genus Typhloctonus Muma, 1961 (Acarina: Mesostigmata) - Intern. J. Acarol., 10: 163 - 177.","Kolodochka L. A. 1986 - On taxonomic status of two Typhloctonus species (Parasitiformes, Phytoseiidae) - Vest. Zool., (2): 26 - 34.","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Kolodochka L. A. 2009 - A review of predaceous mites of the genus Typhloctonus Muma (Parasitiformes, Phytoseiidae) in Ukraine with the description of unknown male of T. tuberculatus - Vest. Zool., 43 (6): 481 - 494.","Wainstein B. A. 1962 - ROvision du genre Typhlodromus Scheuten, 1857 et systematique de la famille des Phytoseiidae (Berlese 1916) (Acarina: Parasitiformes) - Acarologia, 4: 5 - 30.","Abbasova E. D. 1972 - Phytoseiid mites (Parasitiformes: Phytoseiidae) of Azerbaijan - Avtoreferat Dissertatsii na Soiskanie Uchenoy Stepeni Kandidata Biologicheskikh Nauk. Akadrmiya Nauk Azerbaydzhanskoy SSR, Institut Zoologii, Baku, 1 - 34.","Karg W., Edland T. 1987 - Neue Raubmilbenarten der Phytoseiidae Berlese, 1916 - Deut. Entomol. Z., 34 (4 - 5), 387 - 395.","Steeghs N., Nedstam B., Lundqvist L. 1993 - Predatory mites of the family Phytoseiidae (Acari, Mesostigmata) from south Sweden - Entomol. Tidskrift, 114: 19 - 27.","Beglyarov G. A. 1981 - Keys to the determination of phytoseiid mites of the USSR [in Russian] - Information Bulletin International Organization for Biological Control of Noxious Animals and Plants, East Palaearctic Section, Leningrad. 2: 1 - 97.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","AndrO H. M. 1986 - Notes on the ecology of corticolous epiphyte dwellers. 4. Actinedida (especially Tydeidae) and Gamasida (especially Phytoseiidae) - Acarologia, 27: 107 - 115.","Tixier M. - S., Klaric V., Kreiter S., Duso C. 2010 - Phytoseiid Mite Species From Croatia, With Description of a New Species of the Genus Typhlodromus (Typhlodromus) - Entomol. Soc. Amer., 103 (2): 165 - 180.","Kabicek J. 2003 - Broad leaf trees as reservoirs for phytoseiid mites (Acari: Phytoseiidae) - Plant Protec. Sci., 39: 65 - 69.","Kabicek J., Rehakova M. 2004 - Phytoseiid mite community on Aesculus hippocastanum in the parks - Acta Fytotech. Zootech., 7: 114 - 115.","Kabicek J. 2005 - Intraleaf distribution of the phytoseiid mites (Acari, Phytoseiidae) on several species of wild broad leaf trees - Biologia, 60: 523 - 528.","Kabicek J. 2010 - Scarceness of Phytoseiid Species cooccurrence (Acari: Phytoseiidae) on Leaflets of Juglans regia - Plant Protec. Sci., 46 (2): 79 - 82.","Tuovinen T., Rokx J. A. H. 1991 - Phytoseiid mites (Acari: Phytoseiidae) on apple trees and in surrounding vegetation in southern Finland. Densities and species composition - Exp. Appl. Acarol., 12: 35 - 46.","Tuovinen T. 1993 - Identification and occurrence of phytoseiid mites (Gamasina: Phytoseiidae) in Finnish apple plantations and their surroundings - Entomol. Fenn., 4: 95 - 114.","Balder H., Jackel B., Pradel B. 1999 - Investigations on the existence of beneficial organisms on urban trees in Berlin - In Proc. Internatonal Symposium on Urban Tree Health., 496: 189 - 194.","Szabo P. 1980 - Faunisztikai vizsgalatok Toserdo atkain (Acari) - Fol. Entomol. Hung., 33 (2): 377 - 378.","Ripka G. 1998 - New data to the knowledge on the phytoseiid fauna in Hungary (Acari: Mesostigmata) - Acta Phytopathol. Entomol. Hung., 33: 395 - 405.","Ripka G. 2006 - Checklist of the Phytoseiidae of Hungary (Acari: Mesostigmata) - Folia Entomol., 67: 229 - 260.","Ragusa S., Paoletti M. G. 1985 - Phytoseiid mites (Parasitiformes, Phytoseiidae) of corn and soybean agroecosystems in the low-laying plain of Veneto (N-E Italy) - Redia, 68: 69 - 89.","Duso C., Torresan L., Vettorazzo E. 1993 - La vegetazione spontanea come riserva di ausiliari: considerazioni sulla diffusione degli Acari fitoseidi (Acari Phytoseiidae) in un vigneto e sulle piante spontanee contigue - Boll. Zool. Agr. Bachic., Serie II, 25 (2): 183 - 203.","Duso C., Fontana P., Malagnini V. 2004 - Diversity and abundance of phytoseiid mites (Acari: Phytoseiidae) in vineyards and in the surrounding vegetation in northeastern Italy - Acarologia, 41: 31 - 47.","Beglyarov G. A., Malov N. A. 1977 - Predaceous phytoseiid mites (Phytoseiidae: Parasitiformes) of the fauna of Moldavia and north Bukovina (Report II) - Entomofagi v Zashchite Rasteniy. Vsesoyuznaya Ordena Lenina Akademiya Seliskokhozyaystvennyukh Nauk Imeni V. I. Lenina. Kishinev, 3 - 11.","Cobanoglu S. 1996 - Typhloctonus Muma, 1961 (Acarina: Phytoseiidae) species, from the Thrace region of Turkey - Turk. J. Agric. Forest., 20: 353 - 357.","Akimov I. A., Kolodochka L. A., Zhovnerchuk O. V., Omeri I. D., Samojlova T. P. 2007 - Species composition and ecological peculiarities of mites of the superfamily Tetranychoidea (Acariformes, Trombidiformes) and family Phytoseiidae (Parasitiformes, Gamasina), inhabiting plants in botanical gardens of Kyiv (Ukraine) - Vestn. Zool., 41: 521 - 534.","Kolodochka L. A., Omeri I. D. 2007 - Species diversity and distribution of plant-inhabiting phytoseiid mites (Parasitiformes, Phytoseiidae) in Kaniv Reserve - Vest. Zool., 41: 35 - 46.","Congdon B. D. 2002 - The family Phytoseiidae (Acari) in western Washington State with descriptions of three new species - Intern. J. Acarol., 28 (1): 3 - 27."]}

Neoseiulella arutunjani (Kuznetsov) (Figure 9) Seiulus (Typhloctonus) arutunjani Kuznetsov 1984: 54-57. Neoseiulella arutunjani (Kuznetsov) Chant and McMurtry 1994: 248; Moraes et al. 2004: 292; Chant and McMurtry 2007: 147. Adult female (Figure 9a – d) Dorsal shield (Figure 9a) — Dorsal shield heavily reticulated throughout: length 313; width 155 (at level of s4), 160 (at level of Z1). Four pairs of large solenostomes: gd2, gd5, gd6, and gd9. No poroid visible. Sub-lateral seta r3 on the dorsal shield. The other sub-lateral seta R 1 on the lateral margin. Dorsal shield bearing 20 pairs of setae, all dorsal and sub-lateral setae smooth, except Z5 slightly serrated: j1 13; j3 19; j4 12; j5 12; j6 19; J2 20; J5 10; z2 12; z3 19; z4 20; z5 13; Z1 20; Z4 24; Z5 38; s4 22; s6 20; S2 22; S4 20; S5 16; sub-lateral setae r3 13 and R 1 24. Peritreme extending anteriorly to the level between z2-j3. Ventral shields (Figure 9b) — Sternal shield 63 long and 51 wide (at level of ST2), smooth with two pairs of setae (ST1 and ST2) and two pairs of poroids. The third and fourth pairs of sternal setae (ST3, ST4) on separate platelets. A pair of small poroids accompanying ST4. Genital shield 105 long and 49 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield subquadrateshape, 110 long and 78 wide (at level of ZV2), distinctly reticulated throughout, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2), with a pair of circular solenostomes gv3 posteromediad to JV3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. NB: one seta of the JV4 pair is present (reported absent on the original description) supporting Kolodochka (2009). JV5 13 long, smooth. Primary metapodal plate or inguinal sigillum 35 long and 4 wide. Spermatheca (Figure 9c) — Cervix short, 4 long, shallowly cup-shaped. Chelicera (Figure 9d) — Seven teeth and a pilus dentilis on the fixed digit. Movable digit 29 long, bidentate. Legs — Measurements of legs: leg I 248; leg II 213; leg III 213; leg IV 283. Seven setae (2-2/0, 2/0- 1) on the genu II. No macroseta on legs. Material examined — The female holotype deposited in the collection of Nikita State Botanic Garden, Yalta, Crimea, Ukraine. This material was examined in the ASU, Institute of Zoology, Academy of Sciences, Kiev, Ukraine. Adult male The male of this species is unknown. Previous reports — N. arutunjani is only known from Ukraine, on Artemisia sp. (Asteraceae). Remarks — It is noteworthy that in the original description, two females mounted on one slide were mentioned. Kolodochka (2009) showed that only one female actually belonged to this species. The other female was Paraseiulus incognitus Wainstein and Arutunjan. Our examination of these two females supports Kolodochka’s observations. The present examination of the type materials of these three latter species could emphasize the following conclusions: Neoseiulella tuberculata does not differ from N. sexapori in setal and body measurements. Even if the shape of the spermatheca of these two species is slightly different (cup-shaped cervix in N. sexapori and a disc-shaped [shallowly cup-shaped] cervix in N. tuberculata), these two types of shapes are very close and such differences could be simply due to specimen mounting artefacts. The pair of solenostomes gd5 is observed on the four examined specimens of N. sexapori, and only on one of the three type specimens of N. tuberculata. At last, we observe other differences not mentioned by Chant and Yoshida-Shaul (1989): all dorsal setae are arising from tubercles in N. sexapori but not in N. tuberculata; the peritreme is shorter (at level between z2-j3) for N. sexapori than for N. tuberculata (at level of j1 or between j1-j3); the number of teeth on the fixed digit is different (three for N. sexapori, five for N. tuberculata); the ventrianal shield is heavily reticulated in N. sexapori, and lightly in N. tuberculata. As some authors have already shown the importance of such morphological characters for species differentiation (Chant and McMurtry 1994; Tixier et al. 2006a, b; Chant and McMurtry 2007; Okassa et al. 2009), we conclude that N. tuberculata and N. sexapori seem to be separate specific entities. These results agree with the conclusions of Karg and Edland (1987), but not with Chant and Yoshida-Shaul (1989). Small differences in setal and body measurements are observed between N. tuberculata and N. arutunjani. According to intraspecific variations already observed within the family Phytoseiidae for continued characters, these differences seem to be too tiny for separating N. tuberculata from N. arutunjani. However, other differences are observed: the peritreme is shorter in N. arutunjani (reaching between z2-j3) than in N. tuberculata (reaching j1 or between j1-j3); the ventrianal shield is heavily reticulated on N. arutunjani, and lightly on N. tuberculata; fixed digit of chelicera bear seven teeth on N. arutunjani, and five on N. tuberculata; sub-lateral seta r3 is inserted on the dorsal shield on N. arutunjani and on lateral margin on N. tuberculata. However, as the position of sub-lateral seta (R 1) was demontrasted to be a non reliable diagnostic character to differentiate between N. aceri and N. squamiger (Kanouh et al. 2010), the position of r3 could be also assumed to be a non reliable diagnostic character as well. Though, the other morphological differences between N. tuberculata and N. arutunjani seem sufficiently discriminant to conclude that these two species are not synonyms. These results do not agree with the conclusion of Kolodochka (2009). Small differences in setal and body measurements are observed between N. arutunjani and N. sexapori. However, all dorsal setae are arising from tubercles on N. sexapori, but not on N. arutunjani; the number of teeth on the fixed digit is different (three on N. sexapori, seven on N. arutunjani); spermatheca has a cup-shaped cervix in N. sexapori, but a discshaped (shallowly cup-shaped) cervix in N. arutunjani. These morphological differences between N. sexapori and N. arutunjani seem also sufficiently discriminant to conclude that these two species are not synonyms. It seems thus that N. tuberculata, N. arutunjani and N. sexapori present sufficiently different characters to support their specific validity. These conclusions did not agree with Chant and Yoshida-Shaul (1989) and Kolodochka (2009)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 283-285, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Kuznetsov N. N. 1984 - New species of predacious mites of the family Phytoseiidae (Parasitiformes) in the USSR - Entomol. Obozrenie, 63 (2): 384 - 397.","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Kolodochka L. A. 2009 - A review of predaceous mites of the genus Typhloctonus Muma (Parasitiformes, Phytoseiidae) in Ukraine with the description of unknown male of T. tuberculatus - Vest. Zool., 43 (6): 481 - 494.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Tixier M. - S., Kreiter S., Ferragut F., Cheval B. 2006 a - The suspected synonymy of Kampimodromus hmiminai and Kampimodromus adrianae (Acari: Phytoseiidae): morphological and molecular investigations - Can. J. Zool., 84 (8): 1216 - 1222. doi: 10.1139 / z 06 - 108","Okassa M., Tixier M. - S., Cheval B., Kreiter S. 2009 - Molecular and morphological evidence for a new species status within the genus Euseius (Acari: Phytoseiidae): consequences for taxonomy - Can. J. Zool., 87: 689 - 698. doi: 10.1139 / Z 09 - 057","Karg W., Edland T. 1987 - Neue Raubmilbenarten der Phytoseiidae Berlese, 1916 - Deut. Entomol. Z., 34 (4 - 5), 387 - 395.","Kanouh M., Tixier M. - S., Guichou S., Cheval B., Kreiter S. 2010 - Two synonymies within the genus Neoseiulella (Acari: Phytoseiidae): is the molecular evidence so evident? - Biol. J. Linn. Soc., 101: 323 - 344. doi: 10.1111 / j. 1095 - 8312.2010.01516. x"]}

Neoseiulella tiliarum (Oudemans) (Figure 5) Typhlodromus tiliarum Oudemans 1930: 51-52; Chant 1958: 622. Typhlodromus (Typhlodromus) tiliarum (Oudemans) Chant 1959: 65. Typhloctonus tiliarum (Oudemans) Muma 1961: 299; Denmark and Rather 1984: 165; Kolodochka 1986: 26-27; Moraes et al. 1986: 233; Kolodochka 2009: 485-486. Typhlodromus (Nesbitteius) tiliarum (Oudemans) Wainstein 1962: 23. Seiulus tiliarum Abbasova 1972: 21; Karg 1982: 205; Karg and Edland 1987: 387. Seiulus (Typhloctonus) tiliarum (Oudemans) Beglyarov 1981: 19. Neoseiulella (Typhloctona) tiliarum (Oudemans) Denmark and Rather 1996: 58-59. Neoseiulella tiliarum (Oudemans) Chant and McMurtry 1994: 248; Moraes et al. 2004: 296; Chant and McMurtry 2007: 147. Adult female (Figure 5a – d) Dorsal shield (Figure 5a) — Dorsal shield distinctly reticulated throughout: length 350; width 164 (at level of s4) and 182 (at level of Z1). Five pairs of large circular solenostomes: gd1, gd2, gd6, gd8, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except Z4 and Z5 slightly serrated: j1 20; j3 30; j4 19; j5 21; j6 20; J2 26; J5 7; z2 25; z3 31; z4 32; z5 20; Z1 26; Z4 38; Z5 44; s4 34; s6 36; S2 40; S4 35; S5 18; sub-lateral setae r3 29 and R1 21. Peritreme extending anteriorly to the level of z4. Ventral shields (Figure 5b) — Sternal shield 82 long and 84 wide (at level of ST2), smooth with two pairs of setae (ST1 and ST2) and two pairs of poroids. ST3 on a separate platelet close to the sternal shield. A pair of metasternal setae (ST4) on separate platelets with a pair of small poroids. Genital shield 123 long and 62 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla folded under the genital shield and between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 1 pair of poroids around the genital shield. Ventrianal shield rectangular-shaped, 103 long and 60 wide (at level of ZV2), slightly reticulated, with four pairs of preanal setae (JV1, JV2, JV3 and ZV2) and without solenostome gv3. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 26 long, smooth. Primary metapodal plate or inguinal sigillum 28 long and 3 wide. Spermatheca (Figure 5c) — Cervix 14 long, elongated and cup-shaped. Chelicera (Figure 5d) — One tooth and a pilus dentilis on the fixed digit. Movable digit 27 long, unidentate. Legs — Measurements of legs: leg I 274; leg II 222; leg III 219; leg IV 281. Eight setae (2-2/1, 2/0-1) on the genu II. No macroseta on legs. Material examined — The female holotype deposited in the Rijksmuseum van Natuurlijke Historie, Leiden, the Netherlands. As this specimen was in bad conditions, the leg chaetotaxy and lengths were observed on one specimen of our collection (Montpellier SupAgro, UMR CBGP), collected in Valleraugues (Gard, South of France) on small leaved linden (Tilia cordata Miller). Adult male (Figure 5e, f) Described by Livshitz and Kuznetsov (1972) and Denmark and Rather (1984, 1996). Dorsal shield chaetotaxy similar to the female, but the sub-lateral seta R1 on the dorsal shield. Ventrianal shield 104 long and 131 wide, distinctly reticulated and bearing five pairs of preanal setae and three pairs of poroids. Spermatodactyl Lshaped with a terminal foot and a toe slightly enlarged. This description is based on a specimen of our collection (Montpellier SupAgro, UMR CBGP), collected in Valleraugues (Gard, South of France) on T. cordata. Previous reports — N. tiliarum is only known from the West- Palaearctic and Nearctic areas. However, its occurrence in the Nearctic area (15 reports in Canada and three in USA) might be due to introductions. The three reports in Massachusetts (USA) were on insects: Graphiphora haruspica (Grote) and Spaelotis clandestine (Harris) (Noctuidae, Lepidoptera) (Treat 1975). Countries from which this species is reported are: Algeria (Athias-Henriot, 1958 b); Austria (Bohm 1960; Ragusa and Ragusa 1997); Azerbaijan (Abbasova 1972); Canada (Nesbitt 1951; Herbert 1952; Chant et al. 1974); Croatia (Tixier et al. 2010); Czech Republic (Kabicek 2003; Kabicek and Rehakova 2004; Kabicek 2008, 2010); Denmark (Hansen and Johnsen 1986); England (Nesbitt 1951; Chant 1955, 1956; Collyer 1956); France (Gunthart 1957, 1960; Rambier 1974; Kreiter and Brian 1987; Kreiter et al. 2000; Tixier et al. 2000; Arnault et al. 2008); Georgia (Samsoniya 1977); Germany (Oudemans 1930; Dosse 1956a, b, 1957; Westerboer and Bernhard 1963; Thill 1964; Karg 1970, 1971, 1972, 1975); Greece (Swirski and Ragusa 1976; Ragusa and Tsolakis 1998); Hungary (Kropczy´nska and Jenser 1968; Bozai 1980; Dellei and Szendrey 1991; Sarospataki et al. 1992; Bream and Bozai 1995; Bozai 1996, 1997; Ripka 1998, 2006); Iran (Daneshvar 1987); Italy (Gunthart 1957, 1960; Coiutti 1993; Duso et al. 1993; Michelatti et al. 1994; Nicotina and Cioffi 1998; Duso et al. 2004); Moldova (Beglyarov and Malov 1977); Montenegro (Mijuskovic and Tomasvic 1975); Netherlands (Nesbitt 1951; Van de Vrie and Fluiter 1958; Van de Vrie 1963, 1972; Miedema 1987); Norway (Karg and Edland 1987); Poland (Wiackowski and Suski 1963; Boczek 1964; Dabrowski 1968; Dabrowski 1970; Skorupska 1981; Chant and Yoshida-Shaul 1989; Jaworski 2000; Kropczy´nska et al. 2009); Russia (Beglyarov 1962); Slovak Republic (Praslicka and Bartekova 2008; Praslicka et al. 2009); Spain (Chant and Yoshida-Shaul 1989; Villaronga and Garcia- Mari 1988, 1992; Minarro et al. 2005); Switzerland (Gunthart 1957, 1960; Genini et al. 1983; Klay 1987); Tunisia (Kreiter et al. 2002); Turkey (Swirski and Amitai 1982; Düzgüne¸ s and Kiliç 1983; Cobanoglu 1991, 1992, 1996, 2000); Ukraine (Akimov and Kolodochka 1970; Livshitz and Kuznetsov 1972; Beglyarov and Malov 1977; Kolodochka and Omeri 2007; Kolodochka 2009; Omeri 2009); USA (Treat 1975). Neoseiulella tiliarum was collected on a wide range of plant supports: A. campestre, Acer sp. (Aceraceae); Actinidia deliciosa (Chev.) Liang and Ferguson (Actinidiaceae); A. hippocastanum (Hippocastanaceae); Agrimonia sp., Malus baccata (L.) Borkh, M. pumila Miller, Malus sp., Mespilus sp., P. armeniaca, P. cerasifera Ehrh, P. (Cerasus) avium (L.) L., P. domestica L., Prunus sp., P. spinosa L., Pyrus aucuparia L., P. communis L., Rosa sp., Rubus sp. (Rosaceae); Alnus sp., Carpinus betulus, Corylus avellana, Corylus sp. (Betulaceae); Castanea sativa Miller, Fagus sylvatica L., Quercus sp. (Fagaceae); Cercis canadensis L. (Fabaceae); Citrus sp. (Rutaceae); Cornus sanguinea L., Cornus sp. (Cornaceae); Ficus carica L. (Moraceae); Frangula alnus Miller (Rhamnaceae); Fraxinus angustifolia Vahl, Fraxinus sp., Olea sp. (Oleaceae); Dittrichia viscosa ( L.) Greuter (Asteraceae); Juglans regia (Juglandaceae); Pteridium aquilinum (L.) Kuhn (Hypolepidaceae); Ribes nigrum L., Ribes sp. (Grossulariaceae); Tilia americana L., T. cordata Miller, T. miqueliana Maxim., T. platyphyllos Scopoli, Tilia sp. (Tiliaceae); Ulmus campestris L., U. laevis Pallas, U. pumila L., U. scabra (Miller), Ulmus sp., U. suberosa Moench, Zelkova carpinifolia (Pall.) Dippel., Zelkova sp. (Ulmaceae); Urtica dioica L. (Urticaceae); Vitis sp., V. vinifera L. (Vitaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 275-277, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Oudemans A. C. 1930 - Acarologische Aanteekeningen. CI - Entomol. Berich., 8: 48 - 53.","Chant D. A. 1958 - Immature and adult stages of some British Phytoseiidae Berl., 1916 (Acarina) - J. Linn. Soc. London, Zool., 43: 599 - 643.","Chant D. 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Lenin Academy of Agricultural Science, The State Nikita Botanical Gardens, 61: 13 - 64.","Treat A. E. 1975 - Mites of moths and butterflies - Comstock Publishing Associates, Cornell Univ. Press, Ithaca, USA, 362 pp.","Athias-Henriot C. 1958 - Phytoseiidae et Aceosejidae (Acarina: Gamasina) d'AlgOrie. II. Phytoseiidae. ClO des genres Amblyseius Berlese (Suite) et Seiulus Berlese - Bull. Soc. Hist. Natur. Afr. N., 49: 23 - 43.","Bohm H. 1960 - Untersuchungen uber Spinnmilbenfeinde in Osterreich - Pflanzens; Ber., 25: 23 - 46.","Ragusa S., Ragusa E. 1997 - On Some Phytoseiid Mites (Parasitiformes, Phytoseiidae) from Styria (Austria) - Mitt. Naturw. Ver. Steiermarkt, 127: 137 - 145.","Nesbitt H. H. J. 1951 - A taxonomic study of the Phytoseiidae (Family Laelaptidae) predaceous upon Tetranychidae of economic importance - Zool. Verhand., 12: 64 pp. + 32 plates.","Herbert H. J. 1952 - Progress report on predacious mite investigations in Nova Scotia (Acarina, Phytoseiidae) - Ann. Rep. Entomol. Soc. Ontario, 83, 27 - 29.","Chant D. A., Hansell R. I. C., Yoshida-Shaul E. 1974 - The genus Typhlodromus Scheuten (Acarina: Phytoseiidae) in Canada and Alaska - Can. J. Zool., 52: 1265 - 1291. doi: 10.1139 / z 74 - 168","Tixier M. - S., Klaric V., Kreiter S., Duso C. 2010 - Phytoseiid Mite Species From Croatia, With Description of a New Species of the Genus Typhlodromus (Typhlodromus) - Entomol. Soc. Amer., 103 (2): 165 - 180.","Kabicek J. 2003 - Broad leaf trees as reservoirs for phytoseiid mites (Acari: Phytoseiidae) - Plant Protec. Sci., 39: 65 - 69.","Kabicek J., Rehakova M. 2004 - Phytoseiid mite community on Aesculus hippocastanum in the parks - Acta Fytotech. Zootech., 7: 114 - 115.","Kabicek J. 2008 - Cohabitation and intraleaf distribution of phytoseiid mites (Acari, Phytoseiidae) on leaves of Corylus avellana - Plant Prot. Sci., 44 (1): 32 - 36.","Kabicek J. 2010 - Scarceness of Phytoseiid Species cooccurrence (Acari: Phytoseiidae) on Leaflets of Juglans regia - Plant Protec. Sci., 46 (2): 79 - 82.","Hansen E. W., Johnsen S. 1986 - Predatory mites of the family Phytoseiidae in Denmark (Acarina, Gamasina) - Entomol. Meddelelser., 53 (3): 137 - 142.","Chant D. A. 1955 - Notes on mites of the genus Typhlodromus Scheuten, 1857 (Acarina: Laelaptidae), with descriptions of the males of some species and the female of a new species - Can. Entomol .., 87 (11): 496 - 503. doi: 10.4039 / Ent 87496 - 11","Chant D. A. 1956 - Some mites of the subfamily Phytoseiinae (Acarina: Laelaptidae) from southeastern England, with descriptions of new species - Can. Entomol., 88: 26 - 37. doi: 10.4039 / Ent 8826 - 1","Collyer E. 1956 - Notes on the biology of some predacious mites on fruit trees in south-eastern England - Bull. Entomol. Res., 47 (2): 205 - 214.","Gunthart E. 1957 - Neues uber Auftreten und Bekampfung der Spinnmilben an Reben - Schweizer. Z. Obst- Weinbau, 66: 231 - 236.","Gunthart E. 1960 - Uber das Auftreten von Spinnmilben (Tetranychidae) und Raubmilben (Phytoseiinae, Acari) in der Schweiz - Proceed. IV Internat. Cong. Crop Protection, Hamburg, 1957, 927 - 928.","Rambier A. 1974 - Relations entre les acariens nuisibles et leurs ennemis naturels - In: Les organismes auxiliares en verger de pommiers - IOBC / WPRS Bull., 3: 107 - 109.","Kreiter S., Brian F. 1987 - Les Phytoseiidae de la vigne en France: aspects faunistique, biogOographiques et agronomique - Proceed. Intern. Conf. Pests Agric., 6 (3): 241 - 249.","Kreiter S., Tixier M. - S., Auger P., Muckensturm N., Sentenac G., Doublet B., Weber M. 2000 - Phytoseiid mites of vineyards in France (Acari: Phytoseiidae) - Acarologia, 41 (1 - 2): 77 - 96.","Tixier M. - S., Kreiter S., Auger P., Sentenac G., Salva G., Weber M. 2000 - Phytoseiidae mite species located in uncultivated areas surrounding vineyards in three French regions - Acarologia, 41 (1 - 2): 127 - 140.","Samsoniya T. I. 1977 - Zonal-vertical distribution of Phytoseiidae in eastern Georgia on pip fruit culture - Bull. Acad. Sc. Georg. SSR, 87 (1): 181 - 183.","Dosse G. 1956 a - Ueber die Bedeutung der Raubmilben innerhalb der Spinnmilbenbiozone auf Apfel. I. Grundsatzliches aus der Biologie rauberischer Milben - Mitt. Biol. Bund. Land- Forstwirschaft, 85: 40 - 44.","Westerboer I., Bernhard F. 1963 - Die Familie Phytoseiidae Berlese 1916 - In: Stammer, H. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina. Band II, Mesostigmata I. pp. 451 - 791.","Thill H. 1964 - Beobachtungen an raubmilben auf Zwetschenbaumen - Entomophaga, 9: 239 - 242.","Karg W. 1970 - Uber die Moglichkeiten von integrierten Pflanzenschutzmassnahmen bei der Spinnmilbenbekampfung im Obstbau - Nachrich. Deut. Pflanzen., 24 (8): 166 - 171.","Karg W. 1971 - Untersuchungen uber die Acarofauna in Apfelanlagen im hinblick auf den ubergang von standardspritzprogrammen zu integrierten Behandlungsmassnahmen - Archiv. Pflanzen., 7 (4): 243 - 279.","Karg W. 1972 - Untersuchungen uber die Korrelation zwischen dominierenden Raubmilben arten und ihrer moglichen Beute in Apfelanlagen - Archiv. Pflanzen., B 8 (1): 29 - 52.","Karg W. 1975 - Nutzlingsschonende Spinnmilbenbekampfung im Obstbau - Gartenbau, 22: 51 - 54.","Swirski E., Ragusa S. 1976 - Notes on predacious mites of Greece, with a description of five new species (Mesostigmata: Phytoseiidae) - Phytoparasitica, 4: 101 - 122. doi: 10.1007 / BF 02980341","Ragusa S., Tsolakis H. 1998 - Phytoseiid mites (Parasitiformes, Phytoseiidae) of Lesvos Island (Greece) with a description of a new species - Entomol. Hellen., 12: 55 - 64.","Kropczy ' nska D., Jenser G. 1968 - Data to the knowledge of the Phytoseiidae in Hungarian orchards - Fol. Entomol. Hungar., 21: 321 - 323.","Bozai J. 1980 - Data to the Phytoseiidae fauna of Hungary (Acari) - Fol. Entomol. Hungar., 41 (33): 193 - 194.","Dellei A., Szendrey L. 1991 - The mite infestation of different grapevine varieties in the vineyards of Heves County - Novenyvedelem, 27 (2): 55 - 61.","Sarospataki G., Szendrey L., Mikulas J. 1992 - Raubmilben in der Weingarten von Ungarn - Med. Fac. Landbouww. Univ. Gent, 57: 965 - 967.","Bream A. S., Bozai J. 1995 - Data to the knowledge about mite population on linden trees - NovOnyvOdelem., 31 (12): 585 - 588.","Bozai J. 1996 - Data to the knowledge of predaceous mites of Hungary (Acari: Phytoseiidae, Phytoseiinae) - NovOnyvOdelem., 32: 521 - 525.","Bozai J. 1997 - Data to the fauna of predaceous mites of Hungary with the description of four new species (Acari: Phytoseiidae) - Fol. Entomol. Hungar., 58: 35 - 43.","Ripka G. 1998 - New data to the knowledge on the phytoseiid fauna in Hungary (Acari: Mesostigmata) - Acta Phytopathol. Entomol. Hung., 33: 395 - 405.","Ripka G. 2006 - Checklist of the Phytoseiidae of Hungary (Acari: Mesostigmata) - Folia Entomol., 67: 229 - 260.","Daneshvar H. 1987 - Some predatory mites from Iran, with descriptions of one new genus and six new species (Acari: Phytoseiidae, Ascidae) - Entomol. Phytopathol. 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J. de. 1958 - Some observations on the effect of insecticides and acaricides on the population of the European red spider mite (Metatetranychus ulmi Koch) and its principal predators in commercial orchards in the Netherlands - Proceed. 10 th Intern. Cong. Entomol., Montreal, Aug. 17 - 25, 1956, Canada. 4: 603 - 606.","Van de Vrie M. 1963 - Observations on the ecology of predatory mites (Phytoseiidae) on fruit trees - Mitt. Schweizer. Entomol. Gesell., 36: 54 - 55.","Van de Vrie M. 1972 - Phytoseiid mites on tree crops, ornamental and wild plants in the Netherlands - Entomol. Berich., 32: 13 - 20.","Miedema E. 1987 - Survey of phytoseiid mites (Acari: Phytoseiidae) in orchards and surrounding vegetation of northwestern Europe, especially in the Netherlands. Keys, descriptions and figures - Nether. J. Plant Pathol., 93 (2): 1 - 64.","Wiackowski S. K. Suski Z. W. 1963 - Preliminary investigations on the predacious mites (Acarina: Phytoseiidae) in Poland - Prace Instytutu Sadownictwa, 7: 233 - 237.","Boczek J. 1964 - Studies on mites (Acarina) living on plants in Poland. IV - Bull. Acad. Pol. Sc., SOrie Sciences Biologiques, 12: 365 - 369 + 4 plates.","Dabrowski Z. T. 1968 - Studies on the toxicity of pesticides commonly used in the orchards in Poland on predatory mites (Phytoseiidae) - Roczniki Nauk Rolniczych., 93 (A), 655 - 670.","Dabrowski Z. T. 1970 - Density of spider mites (Tetranychidae) and predatory mites (Phytoseiidae) in apple orchards treated and not treated with pesticides - Ekol. Polska, 18: 111 - 136.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Jaworski S. 2000 - Occurrence of phytoseiid mites (Acari: Phytoseiidae) on blackcurrant plantations and in surrounding vegetation in Southern Poland - IOBC / WPRS Bull., 23 (11): 57 - 62.","Beglyarov G. A. 1962 - On the fauna of predaceous mites of the family Phytoseiidae in Krasnodar region - In: Zimina, L. S. and Shchepetilinikovoiy, V. A. (Eds.), Biologicheskiy Metod Bor'by s Vreditelyami i Boleznyami Cel'skokhozyaystvennykh kul'tur, 1: 198 - 203.","Praslicka J., Bartekova A. 2008 - Occurrence of predatory mites of the Phytoseiidae family on apple-trees in integrated and ecological orchards - Plant Prot. 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Neoseiulella oleariae (Collyer) (Figure 35) Typhlodromus oleariae Collyer 1982: 190-191; Schicha 1987: 137-138; Chant and Yoshida-Shaul 1989: 1035- 1039. Amblydromella oleariae (Collyer) Moraes et al. 1986: 168-169. Neoseiulella (Neoseiulella) oleariae (Collyer) Denmark and Rather 1996: 52-53. Neoseiulella oleariae (Collyer) Moraes et al. 2004: 295; Chant and McMurtry 2007: 147. Adult female (Figure 35a – e) Dorsal shield (Figure 35a) — Dorsal shield smooth, except for the lateral area of the podosoma lightly reticulated: length 418 (417 – 418); width 258 (257 – 259) (at level of s4), 269 (at level of Z1), and 280 (279 – 280) (at the level of S2). Five pairs of solenostomes on the dorsal shield: gd2, gd4, gd5, gd6, and gd9. No poroid visible. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z4 and Z5 slightly serrated: j1 38 (37 – 39); j3 56; j4 31 (30 – 31); j5 23 (22 – 23); j6 30 (29 – 30); J2 37 (35 – 39); J5 11 (10 – 11); z2 28; z3 50 (45 – 55); z4 38 (37 – 39); z5 11; Z1 46; Z4 87 (86 – 88); Z5 112 (110 – 113); s4 57 (55 – 58); s6 54 (51 – 56); S2 58 (57 – 59); S4 33 (32 – 34); S5 35 (33 – 36); sub-lateral setae r3 40 (38 – 42) and R1 31. Presence of few visible muscles marks (sigilles) on the dorsal shield. Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 35b) — Sternal shield 92 long and 94 wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 121 (119 – 123) long and 76 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under the genital shield. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield 152 (150 – 153) long and 123 (121 – 124) wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent) and a pair of small circular solenostomes gv3 posterior or posterolateral to JV2. Three pairs of caudoventral setae (ZV1, ZV3 and JV5, JV4 absent) on the integument surrounding the ventrianal shield. Absence of the caudoventral seta JV4 allows to easily distinguish this species from all the. JV5 53 (51 – 55) long, smooth. Primary metapodal plate or inguinal sigillum 35 (34 – 35) long and 11 (10 – 11) wide. Spermatheca (Figure 35c) — Cervix 10 (9 – 10) long, U-shaped. Chelicera (Figure 35d) —1 Nine teeth and a pilus dentilis on the fixed digit. Movable digit 38 (36 – 39) long, bearing three teeth. Legs (Figure 35e) — Measurements of legs: leg I 391 (387 – 395); leg II 334 (324 – 343); leg III 340 (331 – 348); leg IV 439. Seven setae (2-2/0, 2/0-1) on the genu II. Two macrosetae 34 (34 – 35) and 33 (32 – 33) long on the genu and tibia III, respectively. Three other knobbed macrosetae, 53 (52 – 54), 52 (50 – 54) and 54 (51 – 57) long, on the genu, tibia and basitarsus IV, respectively. Material examined — Two female paratypes deposited in the New Zealand Arthropod Collection (NZAC), Landcare Research, Auckland, New Zealand. Adult male Described by Collyer (1982) and Denmark and Rather (1996). We were not able to borrow male type specimens of this species. Previous reports — N. oleariae is only known from New Zealand. The female holotype and two paratype specimens (1 female and 1 male) of N. oleariae were collected on Olearia colensoi Moench. (Asteraceae) at Ruahine Range. Other paratype specimens (2 females and 1 male) were also collected on O. colensoi at Magister Ridge (Westland), Strachan Ridge (South Westland), and Pillans Pass (Manapouri). Other specimens (excluded from the type series) were collected from both North and South islands of New Zealand. Plants on which this species was collected are: Carmichaelia sp. (Fagaceae); Carpodetus serratus J.R. and G.Forst. (Carpodetaceae); Coprosma polymorpha W.R.B. Oliver (Rubiaceae); Dacrophyllum sp. (Podocarpaceae); Olearia lacunosa Moench. (Asteraceae); Pseudowintera axillaris J.R. and G.Forst. (Winteraceae); Vitex lucens Kirk (Verbenaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 326-327, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Moraes G. J. de, McMurtry J. A., Denmark H. A. 1986 - A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat - EMBRAPA-DDT, Brasilia. 353 pp.","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}

Neoseiulella ashleyae (Chant and Yoshida-Shaul) (Figure 17) Typhlodromus oleariae sensu Schicha (1987, 187, plate 92), non sensu Collyer (1982). Typhlodromus ashleyae Chant and Yoshida-Shaul 1989: 1039-1041. Neoseiulella (Neoseiulella) ashleyae (Chant and Yoshida- Shaul) Denmark and Rather 1996: 51. Neoseiulella ashleyae (Chant and Yoshida-Shaul) Moraes et al. 2004: 292; Chant and McMurtry 2007: 147. Adult female (Figure 17a – e) Dorsal shield (Figure 17a) — Dorsal shield smooth: length 464 (458 – 469); width 302 (299 – 304) (at level of s4) and 285 (280 – 290) (at level of Z1). Six pairs of small solenostomes: gd2, gd4, gd5, gd6, gd8 and gd9. Four pairs of poroids. In the original description of this species, Chant and Yoshida- Shaul (1989) noted the presence of only five pairs of solenostomes (gd2, gd4, gd5, gd6, and gd9). However, on their drawings a large pair of solenostomes corresponding to the position of gd8 (according to Athias-Henriot [1975] and Swirski et al. [1998]) is present. We observed this pair of solenostomes on the four type specimens examined, we thus considered that gd8 is present and that N. ashleyae bears six solenostomes on the dorsal shield. Sub-lateral setae (r3 and R1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth, except Z5 slightly serrated: j1 35 (33 – 37); j3 53 (51 – 56); j4 17 (15 – 18); j5 13 (10 – 16); j6 20; J2 24 (21 – 27); J5 11 (10 – 11); z2 18 (16 – 20); z3 48 (45 – 50); z4 26 (25 – 27); z5 11 (10 – 11); Z1 28 (26 – 29); Z4 61 (59 – 62); Z5 91 (88 – 94); s4 51 (49 – 53); s6 38 (36 – 39); S2 51 (50 – 52); S4 31 (28 – 33); S5 27 (26 – 27); sub-lateral setae r3 24 (20 – 28) and R1 22 (19 – 24). Peritreme extending anteriorly to the level of j1. Ventral shields (Figure 17b) — Sternal shield 93 (91 – 94) long and 101 (98 – 103) wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. A pair of metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 147 long, 78 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla between genital and ventrianal shields. One pair of poroids close to the genital shield (ST 5) and 4 pairs of poroids around the genital shield. Ventrianal shield subtriangularshaped, 152 (147 – 156) long and 128 (122 – 133) wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent) and a pair of small solenostomes gv3 posterior to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 46 (41 – 51) long, smooth. Primary metapodal plate or inguinal sigillum 26 (24 – 28) long and 8 (6 – 9) wide. Spermatheca (Figure 17c) — Cervix 14 (12 – 16) long, thick and U-shaped. Chelicera (Figure 17d) — Nine teeth and a pilus dentilis on the fixed digit. Movable digit 42 (39 – 44) long, bears three teeth. Legs (Figure 17e) — Measurements of legs: leg I 435 (430 – 440); leg II 388 (386 – 390); leg III 364 (361 – 366); leg IV 441 (436 – 445). Seven setae (2-2/0, 2/0-1) on the genu II. One macroseta (28 long) on the genu III. Three other macrosetae, 42 (39 – 45), 42 (40 – 44) and 34 (32 – 35) long, on genu, tibia and basitarsus IV, respectively. Material examined — The female holotype and three female paratypes deposited in the Canadian National Collection, Department of Zoology, University of Toronto, Toronto, Canada. A specimen of this species labeled as Typhlodromus oleariae (Schicha 1987) (deposited in the New Zealand Arthropod Collection [NZAC], Landcare Research, Auckland, New Zealand) was also presently examined. Our examination of this specimen shows that, as stated by Chant and Yoshida-Shaul (1989), it belongs to N. ashleyae, but not to N. oleariae. Adult male The male of this species is unknown. Previous reports — N. ashleyae is only known from New Zealand (Nelson Province) (Chant and Yoshida-Shaul 1989). Plants on which this species was collected are: Elaeocarpus hookerianus Raoul (Elaeocarpaceae), Citrus sp. (Rutaceae), Nothofagus menziesii (Hook.f.) Oerst. (Nothofagaceae) and Pinus uncinata Ramond ex D. C. (Pinaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 297-299, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E. 1987 - Phytoseiidae of Australia and neighboring areas - Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.","Collyer E. 1982 - The Phytoseiidae of New Zealand (Acarina) 1. The genera Typhlodromus and Amblyseius - keys and new species - New Zealand J. Zool., 9: 185 - 206. doi: 10.1080 / 03014223.1982.10423848","Chant D. A., Yoshida-Shaul. E. 1989 - A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae) - Can. J. Zool., 67 (4): 1006 - 1046. doi: 10.1139 / z 89 - 144","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp.","Athias-Henriot C. 1975 - Nouvelles notes sur les Amblyseiini. II. Le relevO organotaxique de la face dorsale adulte (Gamasides protoadOniques, Phytoseiidae) - Acarologia, 17 (1): 20 - 29.","Swirski E., Ragusa S., Tsolakis H. 1998 - Keys to the phytoseiid mites (Parasitiformes: Phytoseiidae) of Israel - Phytophaga, 8: 85 - 154."]}

Neoseiulella elaeocarpi (Schicha) (Figure 26) Typhlodromus elaeocarpi Schicha, in Schicha and O’Dowd 1993: 304-305. Neoseiulella (Neoseiulella) elaeocarpi (Schicha) Denmark and Rather 1996: 49. Neoseiulella elaeocarpi (Schicha) Moraes et al. 2004: 293; Chant and McMurtry 2007: 147. Adult female (Figure 26a – e) Dorsal shield (Figure 26a) — Dorsal shield heavily reticulated throughout: length 356; width 201 (at level of s4) and 226 (at level of Z1). Six pairs of solenostomes on the dorsal shield: gd1, gd4, gd5, gd6, gd8, and gd9. One pair of poroids. Sublateral setae (r3 and R 1) on the lateral margin. Dorsal shield bearing 19 pairs of setae, all smooth except for Z4 (slightly serrated) and Z5 (distinctly serrated), and on tubercles: j1 18; j3 20; j4 16; j5 16; j6 18; J2 22; J5 9; z2 16; z3 20; z4 20; z5 17; Z1 21; Z4 25; Z5 42; s4 19; s6 21; S2 22; S4 23; S5 25; sub-lateral setae r3 18 and R 1 18. Peritreme extending anteriorly to the level j1. Ventral shields (Figure 26b) — Sternal shield 67 long and 72 wide (at level of ST2), smooth, with three pairs of setae (ST1, ST2 and ST3) and two pairs of poroids. The metasternal setae (ST4) on separate platelets accompanied by a pair of small poroids. Genital shield 128 long and 67 wide (at level of ST5), smooth. Four elongate platelets or genital sigilla separating genital and ventrianal shields folded under genital shield. One pair of poroids close to the genital shield (ST 5) and 3 pairs of poroids around the genital shield. Ventrianal shield subgquadrateshaped, 122 long and 92 wide (at level of ZV2), smooth, with three pairs of preanal setae (JV1, JV2 and ZV2; JV3 absent) and a pair of large solenostomes gv3 posteromediad to JV2. Four pairs of caudoventral setae (ZV1, ZV3, JV4 and JV5) on the integument surrounding the ventrianal shield. JV5 22 long, smooth. Primary metapodal plate or inguinal sigillum 30 long and 6 wide. Spermatheca (Figure 26c) — Cervix 4 long, shallowly cup-shaped. Chelicera (Figure 26d) — Seven or eight teeth and a pilus dentilis on the fixed digit. Movable digit 31 long, bearing three teeth. Legs (Figure 26e) — Measurements of legs: leg I 299; leg II 270; leg III 264; leg IV 332. Seven setae (2-2/0, 2/0-1) on the genu II. Two thick and short macrosetae, 6 and 22 long, on genu and basitarsus IV, respectively. Material examined — The female holotype deposited in NSW Department of Primary Industries, Agricultural Scientific Collections Unit (Acarology), Orange Agricultural Institute, Australia. Adult male The male of this species is unknown. Previous reports — N. elaeocarpi is only known from Queensland (Australia), on Elaeocarpus coorangooloo J. Bailey and C. White (Elaeocarpaceae)., Published as part of Kanouh, M., Kreiter, S., Douin, M. & Tixier, M. - S., 2012, Revision Of The Genus Neoseiulella Muma (Acari: Phytoseiidae). Re-Description Of Species, Synonymy Assessment, Biogeography, Plant Supports And Key To Adult Females, pp. 259-348 in Acarologia 52 (3) on pages 311-313, DOI: 10.1051/acarologia/20122048, http://zenodo.org/record/4667090, {"references":["Schicha E., O'Dowd D. J. 1993 - New Australian species of Phytoseiidae (Acarina) from leaf domatia - J. Austral. Entomol. Soc., 32: 297 - 305. doi: 10.1111 / j. 1440 - 6055.1993. tb 00589. x","Denmark H. A., Rather A. Q. 1996 - Revision of the genus Neoseiulella Muma (Acari: Phytoseiidae) - Intern. J. Acarol., 22 (1): 43 - 77. doi: 10.1080 / 01647959608684080","Moraes G. J. de, McMurtry J. A., Denmark H. A., Campos C. B. 2004 - A revised catalog of the mite family Phytoseiidae - Zootaxa, 434: 1 - 494.","Chant D. A .., McMurtry J. A. 2007 - Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata) - Michigan, Indira Publishing House, 220 pp."]}