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Troglocoelotes Zhao & S. Li, 2019 is the only known genus of Coelotinae of which all species have deep morphological adaptations to the subterranean environment, such as depigmentation of body, degenerated or absent eyes and, frequently, with attenuated bodies and/or appendages. Four species of Troglocoelotes have been reported from Guizhou Province, China.A new funnel-web spider of the genus Troglocoelotes is described and illustrated on the basis of a single female specimen from Tongren City, Guizhou: Troglocoelotes sinanensis sp. nov. Additionally, photos of the collection site and a distribution map are also provided.

Vappolotes hei sp. n. Figs 5–7 LSIDurn:lsid:zoobank.org:act: C1707500-5E8B-4DEF-B424-D5E6724F4C80 Type material. Holotype: ♁ (IZCAS-Ar44387, YX371): China: Sichuan Prov.: Mianyang City: Jiangyou Co.: Wudu Town, Tuanshan Vil., Li Bai Reading Cave, (N31.9085°, E104.7808°, 604 m), 1.XII.2018, L. He leg. Paratypes: 2♀♀ (IZCAS-Ar44388, Ar44389, YX371): same data as holotype. Etymology. The new species is named after Li He, the collector of the specimens used in this study; noun (name) in the genitive case. Diagnosis. The male of Vappolotes hei sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 (i.e., the type species of Vappolotes) by tibia ca. 1.5 times longer than the length of the patella (Fig. 5C), but subequal in length in the latter (Fig. 1C); lateral tibial apophysis small, inconspicuous, and squared (Fig. 5C), but conspicuous and finger-shaped in the latter (Fig. 1C); anterior conductor ca. 2 times wider than the posterior conductor, dorsal conductor lamellar (Fig. 5B), but 3 times wider than the posterior conductor, dorsal conductor finger-shaped in the latter (Fig. 1B); posterior conductor pointed towards the posterior diagonally, with shallow apical concavity (Fig. 5A, B), but almost horizontally, with deeper apical concavity in the latter (Fig. 1A–C). The female of V. hei sp. n. can be distinguished from those of its congeners by the thin and long spermathecae, 5 times longer than wide (Fig. 6B), but ca. 3 times in V. jianpingensis Zhao & Li, 2019 and V. tianjiayu, ca. 2 times in V. longshan and V. ganlongensis Zhao & Li, 2019. Description. Male (holotype). Abdomen with a xiphoid pattern, rest as for the genus (Fig. 6C). Total length 5.55. Carapace 3.02 long, 1.99 wide. Abdomen 2.53 long, 1.46 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.02, AME–ALE 0.03, AME–PME 0.04, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.04. Leg measurements: I 12.11 (3.24, 0.81, 3.17, 2.82, 2.07); II 11.01 (3.00, 0.92, 2.54, 2.71, 1.84); III 10.44 (2.92, 0.59, 2.58, 2.98, 1.37); IV 13.98 (3.65, 1.03, 3.11, 4.12, 2.07). Palp as in Fig. 5A–C: femur 3 times longer than tibia, while tibia slightly longer than patella; patellar apophysis finger-shaped, ca. 2 times longer than wide; retrolateral tibial apophysis ca. 2 times longer than wide; lateral tibial apophysis small, squared, and inconspicuous; cymbium more than 2 times longer than bulb; cymbial furrow short, ca. 1/3 the length of cymbium; anterior conductor enlarged, ca. 1.5 times longer and 2 times wider than the posterior conductor; dorsal conductor expanding and lamellar; posterior conductor pointed towards the posterior diagonally, the proximal end with a finger-shaped outgrowth, the distal end with shallow median concavity; embolus originates around a 5 o’clock position. Female (IZCAS -Ar44388): Abdomen light brownish gray and without any pattern, rest as for the genus (Fig. 6D, E). Total length 5.78. Carapace 2.78 long, 1.84 wide. Abdomen 3.00 long, 1.83 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.07, PME 0.07, PLE 0.08; AME–AME 0.02, AME–ALE 0.03, AME–PME 0.03, ALE– PLE 0.03, PME–PME 0.05, PME–PLE 0.05. Leg measurements: leg I 9.08 (2.37, 0.92, 1.81, 2.31, 1.67); II 8.49 (2.31, 0.83, 1.72, 2.11, 1.52); III 8.23 (2.08, 0.71, 1.67, 2.37, 1.40); IV 10.79 (2.81, 0.86, 2.66, 2.65, 1.81). Epigyne as in Fig. 6A, B: ca.1.5 times longer than wide; atrium large, occupying ca. 3/5 of epigynal field, ca. 1.4 times longer than wide, its posterior margin conspicuous, ca. 1/5 the length of the atrium, as the same width as its lateral margin; copulatory ducts located posteriorly, ca. 1.5 times longer than spermathecae; spermathecae thin and long, 5 times longer than wide, occupying 1/7 of epigynal field, spermathecal head long and finger-shaped; fertilization ducts broad, ca. 3 times longer than wide. Variation. Total length of females: 5.78–6.02 (n=2). Distribution. Known only from Sichuan Province, China (Fig. 7).

Vappolotes hei sp. n. Figs 5–7 LSIDurn:lsid:zoobank.org:act: C1707500-5E8B-4DEF-B424-D5E6724F4C80 Type material. Holotype: ♁ (IZCAS-Ar44387, YX371): China: Sichuan Prov.: Mianyang City: Jiangyou Co.: Wudu Town, Tuanshan Vil., Li Bai Reading Cave, (N31.9085°, E104.7808°, 604 m), 1.XII.2018, L. He leg. Paratypes: 2♀♀ (IZCAS-Ar44388, Ar44389, YX371): same data as holotype. Etymology. The new species is named after Li He, the collector of the specimens used in this study; noun (name) in the genitive case. Diagnosis. The male of Vappolotes hei sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 (i.e., the type species of Vappolotes) by tibia ca. 1.5 times longer than the length of the patella (Fig. 5C), but subequal in length in the latter (Fig. 1C); lateral tibial apophysis small, inconspicuous, and squared (Fig. 5C), but conspicuous and finger-shaped in the latter (Fig. 1C); anterior conductor ca. 2 times wider than the posterior conductor, dorsal conductor lamellar (Fig. 5B), but 3 times wider than the posterior conductor, dorsal conductor finger-shaped in the latter (Fig. 1B); posterior conductor pointed towards the posterior diagonally, with shallow apical concavity (Fig. 5A, B), but almost horizontally, with deeper apical concavity in the latter (Fig. 1A–C). The female of V. hei sp. n. can be distinguished from those of its congeners by the thin and long spermathecae, 5 times longer than wide (Fig. 6B), but ca. 3 times in V. jianpingensis Zhao & Li, 2019 and V. tianjiayu, ca. 2 times in V. longshan and V. ganlongensis Zhao & Li, 2019. Description. Male (holotype). Abdomen with a xiphoid pattern, rest as for the genus (Fig. 6C). Total length 5.55. Carapace 3.02 long, 1.99 wide. Abdomen 2.53 long, 1.46 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.02, AME–ALE 0.03, AME–PME 0.04, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.04. Leg measurements: I 12.11 (3.24, 0.81, 3.17, 2.82, 2.07); II 11.01 (3.00, 0.92, 2.54, 2.71, 1.84); III 10.44 (2.92, 0.59, 2.58, 2.98, 1.37); IV 13.98 (3.65, 1.03, 3.11, 4.12, 2.07). Palp as in Fig. 5A–C: femur 3 times longer than tibia, while tibia slightly longer than patella; patellar apophysis finger-shaped, ca. 2 times longer than wide; retrolateral tibial apophysis ca. 2 times longer than wide; lateral tibial apophysis small, squared, and inconspicuous; cymbium more than 2 times longer than bulb; cymbial furrow short, ca. 1/3 the length of cymbium; anterior conductor enlarged, ca. 1.5 times longer and 2 times wider than the posterior conductor; dorsal conductor expanding and lamellar; posterior conductor pointed towards the posterior diagonally, the proximal end with a finger-shaped outgrowth, the distal end with shallow median concavity; embolus originates around a 5 o’clock position. Female (IZCAS -Ar44388): Abdomen light brownish gray and without any pattern, rest as for the genus (Fig. 6D, E). Total length 5.78. Carapace 2.78 long, 1.84 wide. Abdomen 3.00 long, 1.83 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.07, PME 0.07, PLE 0.08; AME–AME 0.02, AME–ALE 0.03, AME–PME 0.03, ALE– PLE 0.03, PME–PME 0.05, PME–PLE 0.05. Leg measurements: leg I 9.08 (2.37, 0.92, 1.81, 2.31, 1.67); II 8.49 (2.31, 0.83, 1.72, 2.11, 1.52); III 8.23 (2.08, 0.71, 1.67, 2.37, 1.40); IV 10.79 (2.81, 0.86, 2.66, 2.65, 1.81). Epigyne as in Fig. 6A, B: ca.1.5 times longer than wide; atrium large, occupying ca. 3/5 of epigynal field, ca. 1.4 times longer than wide, its posterior margin conspicuous, ca. 1/5 the length of the atrium, as the same width as its lateral margin; copulatory ducts located posteriorly, ca. 1.5 times longer than spermathecae; spermathecae thin and long, 5 times longer than wide, occupying 1/7 of epigynal field, spermathecal head long and finger-shaped; fertilization ducts broad, ca. 3 times longer than wide. Variation. Total length of females: 5.78–6.02 (n=2). Distribution. Known only from Sichuan Province, China (Fig. 7)., Published as part of Li, Bing, Zhao, Zhe & Li, Shuqiang, 2023, Three new species of the genus Vappolotes Zhao & Li, 2019 (Araneae, Agelenidae) from southwest China, pp. 325-336 in Zootaxa 5270 (2) on pages 332-335, DOI: 10.11646/zootaxa.5270.2.8, http://zenodo.org/record/7865018

Vappolotes tianjiayu sp. n. Figs 4, 7 LSIDurn:lsid:zoobank.org:act: FD59BB0A-7E3B-4CBE-B406-1D3AB94ECE7E Type material. Holotype: ♀ (IZCAS-Ar44376, XQ270): China: Hunan Prov.: Changde City: Shimen Co.: Hupingshan Town, Tianjiayu Vil., an unnamed cave, (N29.9065°, E110.8087°, 448 m), 14.IV.2016, Y.L. Li and Z.G. Chen leg. Paratypes: 1♀ (IZCAS-Ar44382, XQ263): same town as holotype, Qingshanxi Vil., an unnamed cave, (N29.9869°, E110.8630°, 442 m), 12.IV.2016, same leg.; 5♀♀ (IZCAS-Ar44377–Ar44381, XQ270), same data as holotype; 1♀ (IZCAS-Ar44383, XQ256): same vil. as holotype, another unnamed cave, (N29.9066°, E110.8088°, 455 m), 14.IV.2016, same leg.; 3♀♀ (IZCAS-Ar44384–Ar44386, XQ245): same town as holotype, Guluoping Vil., Guluo Cave, (N29.9362°, E110.8574°, 395 m), 14.IV.2016, same leg. Etymology. The new species is named after the type locality; noun in apposition. Diagnosis. The female of Vappolotes tianjiayu sp. n. can be distinguished from other species of the genus by the small copulatory ducts and the large spermathecae (Fig. 4B). The length of copulatory ducts far less than the length of spermathecae (Fig. 4B), but longer than 1.5 times in others (Figs 1E, 3B, 6B). Description. Female (holotype): Abdomen light brownish gray, four chevrons present, rest as for the genus (Fig. 4C–E). Total length 7.16. Carapace 3.08 long, 2.04 wide. Abdomen 4.08 long, 2.81 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.11, PME 0.09, PLE 0.10; AME–AME 0.03, AME–ALE 0.04, AME–PME 0.03, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.06. Leg measurements: leg I 9.23 (2.51, 0.79, 2.39, 1.80, 1.74); II 8.59 (2.33, 0.73, 2.03, 1.86, 1.64); III 8.09 (2.49, 0.78, 1.86, 1.94, 1.02); IV 11.62 (3.13, 0.84, 2.97, 2.92, 1.76). Epigyne as in Fig 4A, B: ca. 1.4 times wider than long; atrium large, occupying ca. 3/5 of epigynal field, as long as wide; its posterior margin narrowed in the middle, ca. 1/7 the length of the atrium, at least 1.4 times longer than the width of its lateral margin; copulatory ducts cumuliform, located posteriorly, occupying less than one tenth of spermathecae, covered by spermathecae in dorsal view; spermathecae enlarged, occupying 1/2 of epigynal field; fertilization ducts thin, ca. 5 times longer than wide. Male. Unknown. Variation. Total length of females: 6.52–7.16 (n=11). Distribution. Known only from Hunan Province, China (Fig. 7).

Genus Vappolotes Zhao & Li, 2019 Type species. Vappolotes ganlongensis Zhao & Li, 2019, from Guizhou Province. Remarks. The genus Vappolotes originally was described with only two species: V. ganlongensis Zhao & Li, 2019 (♀ ³) and V. jianpingensis Zhao & Li, 2019 (♀). Some diagnostic characters of this genus are amended here, eg. conductor with dorsal and ventral apophyses in Li et al. (2019a) corrected to dorsal conductor and anterior conductor; tegular apophysis in Li et al. (2019a) corrected to posterior conductor; retrolateral tibial apophysis in Li et al. (2019a) corrected to lateral tibial apophysis; ventrolateral tibial apophysis in Li et al. (2019a) corrected to retrolateral tibial apophysis (Fig. 1A–C); atrium with longitudinal setae plexus, copulatory ducts and spermathecae located anteriorly, medially or posteriorly (Fig. 1D, E). Li et al. (2019a) and Okumura & Zhao (2022) proposed Vappolotes, Longicoelotes Wang, 2002, Platocoelotes Wang, 2002, Papiliocoelites Zhao & Li, 2016 and Spiricoelotes Wang, 2022 are sister groups belonging to lacking epigynal teeth clade, as: {[(Vappolotes + Papiliocoelotes) + Spiricoelotes] + Longicoelotes} + Platocoelotes., Published as part of Li, Bing, Zhao, Zhe & Li, Shuqiang, 2023, Three new species of the genus Vappolotes Zhao & Li, 2019 (Araneae, Agelenidae) from southwest China, pp. 325-336 in Zootaxa 5270 (2) on page 326, DOI: 10.11646/zootaxa.5270.2.8, http://zenodo.org/record/7865018, {"references":["Li, B., Zhao, Z., Chen, Y., Chen, H., Wu, Z. & Li, S. (2019 a) Vappolotes, a new genus of coelotine spiders (Araneae, Agelenidae) from Guizhou, China. Zootaxa, 4701 (5), 434 - 442. https: // doi. org / 10.11646 / zootaxa. 4701.5.3","Okumura, K. & Zhao, Z. (2022) Taxonomic revision of six species of the subfamily Coelotinae (Araneae: Agelenidae) from Japan accompanied with the description of Nesiocoelotes gen. n. Acta Arachnologica, 71 (2), 93 - 103. https: // doi. org / 10.2476 / asjaa. 71.93","Wang, X. (2002) A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 269, 1 - 150. https: // doi. org / 10.1206 / 0003 - 0090 (2002) 269 E 2.0. CO; 2","Zhao, Z. & Li, S. (2016) Papiliocoelotes gen. n., a new genus of Coelotinae (Araneae, Agelenidae) spiders from the Wuling Mountains, China. ZooKeys, 585, 33 - 50. https: // doi. org / 10.3897 / zookeys. 585.8007"]}

Vappolotes longshan sp. n. Figs 2–3, 7 LSIDurn:lsid:zoobank.org:act: D15A6701-396A-4679-BE12-E6961B690E0D Type material. Holotype: ♁ (IZCAS-Ar44326, XQ191): China: Hunan Prov.: Xiangxi Tujia and Miao Autonomous Pref.: Longshan Co.: Xichehe Town, Ganxi Vil., Da Cave, (N29.1842°, E109.5623°, 444 ± 14 m), 5.I.2016, Z.G. Chen and J.Y. Wang leg. Paratypes: 1³ 6♀♀ (IZCAS-Ar44335– Ar 44341, XQ202), same town as holotype, Xinjian Vil., Xianren Cave, (N29.0855°, E109.5110°, 477 ± 28 m), 4.I.2016, same leg.; 1³ 11♀♀ (IZCAS- Ar 44342– Ar 44353, XQ211), same town as holotype, Xinjian Vil., Guanjia Cave, (N29.0866°, E109.5104°, 413 ± 10 m), 4.I.2016, same leg.; 4³³ 4♀♀ (IZCAS-Ar44327– Ar 44334, XQ191), same data as holotype; 1³ 6♀♀ (IZCAS-Ar44354– Ar 44360, XQ203), same town as holotype, the wayside of Zhijia Vil., (N29.1504°, E109.5516°, 455 ± 15 m), 5.I.2016, same leg.; 5♀♀ (IZCAS-Ar44361– Ar 44365, LB204), same co. as holotype: Hongyanxi Town, Daxi Vil., Feng Cave, (N29.2576°, E109.6086°, 389 ± 13 m), 9.I.2016, same leg.; 4♀♀ (IZCAS-Ar44366– Ar 44369, XQ218), same co. as holotype: Hongyanxi Town, Daxi Vil., Xiangjiayan Cave, (N29.2527°, E109.6051°, 427 ± 13 m), 9.I.2016, same leg.; 3³³ 3♀♀ (IZCAS-Ar44370– Ar 44375, YX004), same town as holotype, Shuitong Vil., Yangjia Cave, (N29.0879°, E109.4945°, 431 m), Shuitong Village, the same town as holotype, 26.X.2018, Z.G. Chen and X.Q. Zhang leg. Etymology. The new species is named after the type locality; noun in apposition. Diagnosis. The males of Vappolotes longshan sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 (i.e., the type species of Vappolotes) by tibia longer than the length of patella (Fig. 2C), but subequal in the latter (Fig. 1C); patellar apophysis thick and dark, with a concave distal end (Fig. 2C), but thin, with a pointed and dark distal end in the latter (Fig. 1C); lateral tibial apophysis rectangular (Fig. 2C), but finger-like in the latter (Fig. 1C); posterior conductor ca. 2 times longer than the anterior conductor, with a forked distal end (Fig. 2B), but ca. 1/2, with unbifurcated distal end in the latter (Fig. 1B); dorsal conductor with a blunt distal end (Fig. 2A, B), but with a pointed distal end in the latter (Fig. 1A, B). The females of V. longshan sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 by atrium longer than wide (Fig. 3A), but its length subequal width in the latter (Fig. 1D); copulatory ducts arched (Fig. 3B), but wing-like in the latter (Fig. 1E); anterior spermathecae contiguous (Fig. 3B), but not touching each other in the latter (Fig. 1E). Description. Male (holotype). Abdomen light yellow, dark gray sides, with four chevrons, rest as described for the genus (Fig. 3C). Total length 6.27. Carapace 3.38 long, 2.34 wide. Abdomen 2.89 long, 1.79 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.10, PME 0.08, PLE 0.10; AME–AME 0.04, AME–ALE 0.02, AME–PME 0.04, ALE–PLE 0.02, PME–PME 0.05, PME–PLE 0.04. Leg measurements: I 12.11 (3.26, 0.72, 2.86, 3.09, 2.18); II 11.19 (2.81, 1.06, 2.58, 2.79, 1.95); III 10.61 (2.73, 0.83, 2.29, 2.67, 2.09); IV 12.67 (3.13, 0.88, 3.31, 3.54, 1.81). Palp as in Fig. 2A–C: femur subequal the length of the tibia, ca. 1.3 times longer than patella; patellar apophysis long, ca. 2 times longer than wide, with a concave distal end; retrolateral tibial apophysis rectangular, ca. 4 times longer than wide; lateral tibial apophysis rectangular, ca. 3 times wider than long; cymbium longer than 2.5 times the length of the bulb; cymbial furrow short, ca. 1/2 the length of cymbium; anterior conductor ca. 1/2 the length of the posterior conductor; dorsal conductor large, folded backward; posterior conductor enlarged and pointed diagonally towards the posterior, proximal end with a lamellar and large outgrowth, distal end with shallow median concavity; embolus originates around a 5 o’clock position. Female (IZCAS -Ar44331): Colour and abdominal pattern as in male (Fig. 3D, E). Total length 5.31. Carapace 2.79 long, 1.84 wide. Abdomen 2.52 long, 1.59 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.03, AME–ALE 0.02, AME–PME 0.03, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.04. Leg measurements: leg I 8.73 (2.08, 0.68, 2.11, 2.22, 1.64); II 8.15 (2.26, 0.82, 1.73, 1.92, 1.42); III 7.97 (2.16, 0.73, 1.81, 1.91, 1.36); IV 10.61 (2.66, 0.76, 2.48, 3.07, 1.64). Epigyne as in Fig. 3A, B: ca. 1.2 times wider than long; atrium large, occupying ca. 1/4 of epigynal field, ca. 1.4 times longer than wide; its posterior margin conspicuous, 1/2 the length of atrium, 2 times longer than width of its lateral margin; copulatory ducts enlarged, ca. 2 times longer than spermathecae; spermathecae small, occupying 1/5 of epigynal field, ca. 2 times longer than wide, with long stalks and finger-shaped heads; fertilization ducts broad, ca. 2 times longer than wide. Variation. Total length of males: 5.44–6.29 (n=11), and of females: 5.09–5.69 (n=39). Distribution. Known only from Hunan Province, China (Fig. 7)., Published as part of Li, Bing, Zhao, Zhe & Li, Shuqiang, 2023, Three new species of the genus Vappolotes Zhao & Li, 2019 (Araneae, Agelenidae) from southwest China, pp. 325-336 in Zootaxa 5270 (2) on pages 328-330, DOI: 10.11646/zootaxa.5270.2.8, http://zenodo.org/record/7865018

Vappolotes longshan sp. n. Figs 2–3, 7 LSIDurn:lsid:zoobank.org:act: D15A6701-396A-4679-BE12-E6961B690E0D Type material. Holotype: ♁ (IZCAS-Ar44326, XQ191): China: Hunan Prov.: Xiangxi Tujia and Miao Autonomous Pref.: Longshan Co.: Xichehe Town, Ganxi Vil., Da Cave, (N29.1842°, E109.5623°, 444 ± 14 m), 5.I.2016, Z.G. Chen and J.Y. Wang leg. Paratypes: 1³ 6♀♀ (IZCAS-Ar44335– Ar 44341, XQ202), same town as holotype, Xinjian Vil., Xianren Cave, (N29.0855°, E109.5110°, 477 ± 28 m), 4.I.2016, same leg.; 1³ 11♀♀ (IZCAS- Ar 44342– Ar 44353, XQ211), same town as holotype, Xinjian Vil., Guanjia Cave, (N29.0866°, E109.5104°, 413 ± 10 m), 4.I.2016, same leg.; 4³³ 4♀♀ (IZCAS-Ar44327– Ar 44334, XQ191), same data as holotype; 1³ 6♀♀ (IZCAS-Ar44354– Ar 44360, XQ203), same town as holotype, the wayside of Zhijia Vil., (N29.1504°, E109.5516°, 455 ± 15 m), 5.I.2016, same leg.; 5♀♀ (IZCAS-Ar44361– Ar 44365, LB204), same co. as holotype: Hongyanxi Town, Daxi Vil., Feng Cave, (N29.2576°, E109.6086°, 389 ± 13 m), 9.I.2016, same leg.; 4♀♀ (IZCAS-Ar44366– Ar 44369, XQ218), same co. as holotype: Hongyanxi Town, Daxi Vil., Xiangjiayan Cave, (N29.2527°, E109.6051°, 427 ± 13 m), 9.I.2016, same leg.; 3³³ 3♀♀ (IZCAS-Ar44370– Ar 44375, YX004), same town as holotype, Shuitong Vil., Yangjia Cave, (N29.0879°, E109.4945°, 431 m), Shuitong Village, the same town as holotype, 26.X.2018, Z.G. Chen and X.Q. Zhang leg. Etymology. The new species is named after the type locality; noun in apposition. Diagnosis. The males of Vappolotes longshan sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 (i.e., the type species of Vappolotes) by tibia longer than the length of patella (Fig. 2C), but subequal in the latter (Fig. 1C); patellar apophysis thick and dark, with a concave distal end (Fig. 2C), but thin, with a pointed and dark distal end in the latter (Fig. 1C); lateral tibial apophysis rectangular (Fig. 2C), but finger-like in the latter (Fig. 1C); posterior conductor ca. 2 times longer than the anterior conductor, with a forked distal end (Fig. 2B), but ca. 1/2, with unbifurcated distal end in the latter (Fig. 1B); dorsal conductor with a blunt distal end (Fig. 2A, B), but with a pointed distal end in the latter (Fig. 1A, B). The females of V. longshan sp. n. can be distinguished from those of V. ganlongensis Zhao & Li, 2019 by atrium longer than wide (Fig. 3A), but its length subequal width in the latter (Fig. 1D); copulatory ducts arched (Fig. 3B), but wing-like in the latter (Fig. 1E); anterior spermathecae contiguous (Fig. 3B), but not touching each other in the latter (Fig. 1E). Description. Male (holotype). Abdomen light yellow, dark gray sides, with four chevrons, rest as described for the genus (Fig. 3C). Total length 6.27. Carapace 3.38 long, 2.34 wide. Abdomen 2.89 long, 1.79 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.10, PME 0.08, PLE 0.10; AME–AME 0.04, AME–ALE 0.02, AME–PME 0.04, ALE–PLE 0.02, PME–PME 0.05, PME–PLE 0.04. Leg measurements: I 12.11 (3.26, 0.72, 2.86, 3.09, 2.18); II 11.19 (2.81, 1.06, 2.58, 2.79, 1.95); III 10.61 (2.73, 0.83, 2.29, 2.67, 2.09); IV 12.67 (3.13, 0.88, 3.31, 3.54, 1.81). Palp as in Fig. 2A–C: femur subequal the length of the tibia, ca. 1.3 times longer than patella; patellar apophysis long, ca. 2 times longer than wide, with a concave distal end; retrolateral tibial apophysis rectangular, ca. 4 times longer than wide; lateral tibial apophysis rectangular, ca. 3 times wider than long; cymbium longer than 2.5 times the length of the bulb; cymbial furrow short, ca. 1/2 the length of cymbium; anterior conductor ca. 1/2 the length of the posterior conductor; dorsal conductor large, folded backward; posterior conductor enlarged and pointed diagonally towards the posterior, proximal end with a lamellar and large outgrowth, distal end with shallow median concavity; embolus originates around a 5 o’clock position. Female (IZCAS -Ar44331): Colour and abdominal pattern as in male (Fig. 3D, E). Total length 5.31. Carapace 2.79 long, 1.84 wide. Abdomen 2.52 long, 1.59 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.09, PME 0.08, PLE 0.09; AME–AME 0.03, AME–ALE 0.02, AME–PME 0.03, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.04. Leg measurements: leg I 8.73 (2.08, 0.68, 2.11, 2.22, 1.64); II 8.15 (2.26, 0.82, 1.73, 1.92, 1.42); III 7.97 (2.16, 0.73, 1.81, 1.91, 1.36); IV 10.61 (2.66, 0.76, 2.48, 3.07, 1.64). Epigyne as in Fig. 3A, B: ca. 1.2 times wider than long; atrium large, occupying ca. 1/4 of epigynal field, ca. 1.4 times longer than wide; its posterior margin conspicuous, 1/2 the length of atrium, 2 times longer than width of its lateral margin; copulatory ducts enlarged, ca. 2 times longer than spermathecae; spermathecae small, occupying 1/5 of epigynal field, ca. 2 times longer than wide, with long stalks and finger-shaped heads; fertilization ducts broad, ca. 2 times longer than wide. Variation. Total length of males: 5.44–6.29 (n=11), and of females: 5.09–5.69 (n=39). Distribution. Known only from Hunan Province, China (Fig. 7).

Vappolotes tianjiayu sp. n. Figs 4, 7 LSIDurn:lsid:zoobank.org:act: FD59BB0A-7E3B-4CBE-B406-1D3AB94ECE7E Type material. Holotype: ♀ (IZCAS-Ar44376, XQ270): China: Hunan Prov.: Changde City: Shimen Co.: Hupingshan Town, Tianjiayu Vil., an unnamed cave, (N29.9065°, E110.8087°, 448 m), 14.IV.2016, Y.L. Li and Z.G. Chen leg. Paratypes: 1♀ (IZCAS-Ar44382, XQ263): same town as holotype, Qingshanxi Vil., an unnamed cave, (N29.9869°, E110.8630°, 442 m), 12.IV.2016, same leg.; 5♀♀ (IZCAS-Ar44377–Ar44381, XQ270), same data as holotype; 1♀ (IZCAS-Ar44383, XQ256): same vil. as holotype, another unnamed cave, (N29.9066°, E110.8088°, 455 m), 14.IV.2016, same leg.; 3♀♀ (IZCAS-Ar44384–Ar44386, XQ245): same town as holotype, Guluoping Vil., Guluo Cave, (N29.9362°, E110.8574°, 395 m), 14.IV.2016, same leg. Etymology. The new species is named after the type locality; noun in apposition. Diagnosis. The female of Vappolotes tianjiayu sp. n. can be distinguished from other species of the genus by the small copulatory ducts and the large spermathecae (Fig. 4B). The length of copulatory ducts far less than the length of spermathecae (Fig. 4B), but longer than 1.5 times in others (Figs 1E, 3B, 6B). Description. Female (holotype): Abdomen light brownish gray, four chevrons present, rest as for the genus (Fig. 4C–E). Total length 7.16. Carapace 3.08 long, 2.04 wide. Abdomen 4.08 long, 2.81 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.11, PME 0.09, PLE 0.10; AME–AME 0.03, AME–ALE 0.04, AME–PME 0.03, ALE–PLE 0.01, PME–PME 0.04, PME–PLE 0.06. Leg measurements: leg I 9.23 (2.51, 0.79, 2.39, 1.80, 1.74); II 8.59 (2.33, 0.73, 2.03, 1.86, 1.64); III 8.09 (2.49, 0.78, 1.86, 1.94, 1.02); IV 11.62 (3.13, 0.84, 2.97, 2.92, 1.76). Epigyne as in Fig 4A, B: ca. 1.4 times wider than long; atrium large, occupying ca. 3/5 of epigynal field, as long as wide; its posterior margin narrowed in the middle, ca. 1/7 the length of the atrium, at least 1.4 times longer than the width of its lateral margin; copulatory ducts cumuliform, located posteriorly, occupying less than one tenth of spermathecae, covered by spermathecae in dorsal view; spermathecae enlarged, occupying 1/2 of epigynal field; fertilization ducts thin, ca. 5 times longer than wide. Male. Unknown. Variation. Total length of females: 6.52–7.16 (n=11). Distribution. Known only from Hunan Province, China (Fig. 7)., Published as part of Li, Bing, Zhao, Zhe & Li, Shuqiang, 2023, Three new species of the genus Vappolotes Zhao & Li, 2019 (Araneae, Agelenidae) from southwest China, pp. 325-336 in Zootaxa 5270 (2) on pages 330-332, DOI: 10.11646/zootaxa.5270.2.8, http://zenodo.org/record/7865018

Vappolotes ganlongensis Zhao et S. Li, sp. n. Figs 2, 3, 5 Type material. Holotype: ♂ (IZCAS-Ar39637, ZZ550): Chenjia Cave (N28.38187º, E108.67368º, 792 m), Ganlong Town, Miao Autonomous County of Songtao, Guizhou Province, China, 9.III.2013, H.F. Zhao and J.C. Liu leg. Paratypes: 7♀♀ (IZCAS-Ar39638–Ar39644, ZZ550): same data as holotype. Etymology. The specific name refers to the type locality, Ganlong Town; adjective. Diagnosis. Vappolotes ganlongensis sp. n. can be distinguished from V. jianpingensis sp. n. by the following: the posterior margin of the atrium is concave medially (Fig. 3A), whereas it is arciform in V. jianpingensis sp. n. (Fig. 4A); the copulatory ducts are large, about 2 times larger as the spermathecae (Fig. 3B), whereas the copulatory ducts are nearly the same size as the spermathecae in V. jianpingensis sp. n. (Fig. 4B). Description. Male. Clypeus brown, abdomen gray, other parts are the same color as for the genus (Fig. 3C). Total length 5.76. Carapace 3.04 long, 2.19 wide. Abdomen 2.72 long, 1.75 wide. Eye sizes and interdistances: AME 0.11, ALE 0.16, PME 0.15, PLE 0.16; AME–AME 0.04, AME–ALE 0.02, AME–PME 0.04, ALE–PLE 0, PME–PME 0.06, PME–PLE 0.05. Leg measurements: I 12.24 (3.15, 4.06, 2.94, 2.09); II 11.52 (3.19, 3.52, 2.93, 1.88); III 11.11 (2.67, 3.46, 3.05, 1.93); IV 14.41 (3.71, 4.39, 4.12, 2.19). Palp: Femur 4.5 times longer than wide; patella with a long anterodorsal macroseta, almost 2.3 times longer than the patella, patellar apophysis about 1/2 the patella length, with blunt tip; tibia and base of cymbium with long dorsal and prolateral macrosetae; VTA and RTA spine-shaped, VTA about 3 times longer than RTA, extending beyond the distal margin of the tibia; cymbium 2.2 times longer than bulb, cymbial furrow short, less than 1/2 the cymbial length; dorsal apophysis of the conductor pointed, and the large lamella of conductor spoon-shaped; tegular apophysis shorter than 1/2 the cymbial furrow, with bifurcated tip; embolus originates around the 6 o’clock position (Fig. 2 A–C). Female (IZCAS-Ar39638). Same color as male except abdomen yellow-brown (Fig. 3D, E). Total length 5.95. Carapace 2.56 long, 1.81 wide. Abdomen 3.39 long, 2.34 wide. Eye sizes and interdistances: AME 0.10, ALE 0.14, PME 0.12, PLE 0.13; AME–AME 0.05, AME–ALE 0.03, AME–PME 0.05, ALE–PLE 0, PME–PME 0.07, PME–PLE 0.06. Leg measurements: I 9.04 (2.59, 2.89, 2.11, 1.45); II 8.77 (2.54, 2.78, 2.06, 1.39); III 8.68 (2.41, 2.72, 2.24, 1.31); IV 11.98 (3.35, 3.67, 3.31, 1.65). Epigynum: about 1.7 times wider than long; atrium occupying 3/4 of epigynum, length equal to width, posterior lateral margin of atrium about 1/2 the width of the lateral margin, concave medially; copulatory ducts large, about 3 times larger than spermathecae; spermathecae small, occupying 1/4 of epigynum, over 2 times longer than wide; fertilization ducts 2 times longer than wide (Fig. 3A, B). Variation. Total length of females 5.72–5.96 (n=7). Distribution. Guizhou, China (Fig. 5).

Li, Bing, Zhao, Zhe, Chen, Yuanxue, Chen, Haifeng, Wu, Zhiyan, Li, Shuqiang (2019): Vappolotes, a new genus of coelotine spiders (Araneae, Agelenidae) from Guizhou, China. Zootaxa 4701 (5): 434-442, DOI: https://doi.org/10.11646/zootaxa.4701.5.3

Vappolotes ganlongensis Zhao et S. Li, sp. n. Figs 2, 3, 5 Type material. Holotype: ♂ (IZCAS-Ar39637, ZZ550): Chenjia Cave (N28.38187��, E108.67368��, 792 m), Ganlong Town, Miao Autonomous County of Songtao, Guizhou Province, China, 9.III.2013, H.F. Zhao and J.C. Liu leg. Paratypes: 7♀♀ (IZCAS-Ar39638���Ar39644, ZZ550): same data as holotype. Etymology. The specific name refers to the type locality, Ganlong Town; adjective. Diagnosis. Vappolotes ganlongensis sp. n. can be distinguished from V. jianpingensis sp. n. by the following: the posterior margin of the atrium is concave medially (Fig. 3A), whereas it is arciform in V. jianpingensis sp. n. (Fig. 4A); the copulatory ducts are large, about 2 times larger as the spermathecae (Fig. 3B), whereas the copulatory ducts are nearly the same size as the spermathecae in V. jianpingensis sp. n. (Fig. 4B). Description. Male. Clypeus brown, abdomen gray, other parts are the same color as for the genus (Fig. 3C). Total length 5.76. Carapace 3.04 long, 2.19 wide. Abdomen 2.72 long, 1.75 wide. Eye sizes and interdistances: AME 0.11, ALE 0.16, PME 0.15, PLE 0.16; AME���AME 0.04, AME���ALE 0.02, AME���PME 0.04, ALE���PLE 0, PME���PME 0.06, PME���PLE 0.05. Leg measurements: I 12.24 (3.15, 4.06, 2.94, 2.09); II 11.52 (3.19, 3.52, 2.93, 1.88); III 11.11 (2.67, 3.46, 3.05, 1.93); IV 14.41 (3.71, 4.39, 4.12, 2.19). Palp: Femur 4.5 times longer than wide; patella with a long anterodorsal macroseta, almost 2.3 times longer than the patella, patellar apophysis about 1/2 the patella length, with blunt tip; tibia and base of cymbium with long dorsal and prolateral macrosetae; VTA and RTA spine-shaped, VTA about 3 times longer than RTA, extending beyond the distal margin of the tibia; cymbium 2.2 times longer than bulb, cymbial furrow short, less than 1/2 the cymbial length; dorsal apophysis of the conductor pointed, and the large lamella of conductor spoon-shaped; tegular apophysis shorter than 1/2 the cymbial furrow, with bifurcated tip; embolus originates around the 6 o���clock position (Fig. 2 A���C). Female (IZCAS-Ar39638). Same color as male except abdomen yellow-brown (Fig. 3D, E). Total length 5.95. Carapace 2.56 long, 1.81 wide. Abdomen 3.39 long, 2.34 wide. Eye sizes and interdistances: AME 0.10, ALE 0.14, PME 0.12, PLE 0.13; AME���AME 0.05, AME���ALE 0.03, AME���PME 0.05, ALE���PLE 0, PME���PME 0.07, PME���PLE 0.06. Leg measurements: I 9.04 (2.59, 2.89, 2.11, 1.45); II 8.77 (2.54, 2.78, 2.06, 1.39); III 8.68 (2.41, 2.72, 2.24, 1.31); IV 11.98 (3.35, 3.67, 3.31, 1.65). Epigynum: about 1.7 times wider than long; atrium occupying 3/4 of epigynum, length equal to width, posterior lateral margin of atrium about 1/2 the width of the lateral margin, concave medially; copulatory ducts large, about 3 times larger than spermathecae; spermathecae small, occupying 1/4 of epigynum, over 2 times longer than wide; fertilization ducts 2 times longer than wide (Fig. 3A, B). Variation. Total length of females 5.72���5.96 (n=7). Distribution. Guizhou, China (Fig. 5)., Published as part of Li, Bing, Zhao, Zhe, Chen, Yuanxue, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, Vappolotes, a new genus of coelotine spiders (Araneae, Agelenidae) from Guizhou, China, pp. 434-442 in Zootaxa 4701 (5) on pages 436-437, DOI: 10.11646/zootaxa.4701.5.3, http://zenodo.org/record/3559385

Draconarius budanlaensis Z. Zhao & S. Li, sp. n. Figs 2, 8 Type material. Holotype ♀ (IZCAS-Ar39617): Budanla Mountain (N29.02890��, E92.35338��, 4930 m), Nyalam County, Shigatse, Tibet, China, 10. VIII. 2014, Y.C. Li leg. Etymology. The specific name refers to the type locality, Budanla Mountain; adjective. Diagnosis. Draconarius budanlaensis sp. n. is most similar to D. yadongensis (Hu & Li, 1987). The epigyne of both species has broad teeth that are widely separated; atrium flat and wide; spermathecae close together medially. The two species differ by the locations of the epigynal teeth (laterally in D. budanlaensis sp. n. in Fig. 2A and centrally in D. yadongensis in fig. 20.1 in Hu & Li 1987); the shape of the epigynal teeth (broad and short in D. budanlaensis sp. n. in Fig. 2A and as wide as long in D. yadongensis in fig. 20.1 in Hu & Li 1987), and the length of the posterior margin of the atrium (1/3 as long as wide in D. budanlaensis sp. n. in Fig. 2A and 5 times wider than long in D. yadongensis in fig. 20.1 in Hu & Li 1987). Description. Female (IZCAS-Ar39617). Carapace and legs brown. Clypeus, endites, labium and sternum black-brown. Abdomen grey-brown. The colour of spinnerets equals the carapace. Abdomen with five chevronshaped patterns. Total length 14.10. Carapace 6.41 long, 4.16 wide. Abdomen 7.69 long, 3.95 wide. Eye sizes and interdistances: AME 0.16, ALE 0.20, PME 0.18, PLE 0.19; AME���AME 0.21, AME���ALE 0.17, AME���PME 0.22, ALE���PLE 0, PME���PME 0.29, PME���PLE 0.28. Leg measurements: I 16.79 (4.49, 5.77, 3.92, 2.61); II 15.69 (4.16, 5.13, 3.80, 2.60); III 15.54 (4.01, 4.81, 4.16, 2.56); IV 19.48 (4.80, 6.09, 5.38, 3.21). Epigyne: about 1.5 times wider than long, with 2 broad teeth that are about 3 times wider than long, situated anteriorly and widely separated, hoods distinct, atrium flat and wide, about 1.7 times wider than long, posterior margin of atrium 3 times wider than long; spermathecae longer than wide, spermathecal heads originating anteriorly from the spermathecae; fertilization ducts 3 times wider than long (Fig. 2 A���B). Male. Unknown. Distribution. Known only from Tibet, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, Four new species of the genus Draconarius Ovtchinnikov, 1999 (Araneae, Agelenidae) from the Tibetan Plateau, China, pp. 141-154 in Zootaxa 4648 (1) on page 143, DOI: 10.11646/zootaxa.4648.1.7, http://zenodo.org/record/3353959, {"references":["Hu, J. & Li, A. (1987) The spiders collected from the fields and the forests of Xizang Autonomous Region, China. (II). Agricultural Insects, Spiders, Plant Diseases and Weeds of Xizang, 2, 247 - 353."]}

Draconarius yingbinensis Z. Zhao & S. Li, sp. n. Figs 7���8 Type material. Holotype ♀ (IZCAS-Ar29625): Yingbin Road (N27.98375��, E85.98287��, 2450 m), Zhangmu Township, Nyalam County, Shigatse, Tibet, China, 29. VII. 2014, Y.C. Li leg. Paratype: 1♀ (IZCAS-Ar39626): same data as holotype. Etymology. The specific name refers to the type locality, Yingbin Road; adjective. Diagnosis. Draconarius yingbinensis sp. n. is most similar to D. nanyuensis (Peng & Yin, 1998). The epigynal teeth of both species are located anteriorly, close together, bent at the end, and have the same length to width ratio; atrium small, close to the posterior margin of the epigyne; spermathecae spherical, spermathecal heads originating anteriorly from the spermathecae. The two species differ by the shape of the epigynal teeth at the end (blunt in D. yingbinensis sp. n. in Fig. 7A and pointed in D. nanyuensis in fig. 8 in Peng & Yin 1998); the shape of the atrium (rectangular in D. yingbinensis sp. n. in Fig. 7A and with a triangular septum with atrium on both sides of septum in D. nanyuensis in fig. 8 in Peng & Yin 1998). Description. Female (IZCAS-Ar39625). Carapace, sternum, legs, abdomen and spinnerets brown; clypeus, endites and labium darker than carapace.Abdomen with five chevron-shaped patterns and covered by blueish-green hairs. Total length 6.34. Carapace 3.48 long, 2.23 wide. Abdomen 2.86 long, 2.01 wide. Eye sizes and interdistances: AME 0.11, ALE 0.20, PME 0.11, PLE 0.16; AME���AME 0.08, AME���ALE 0.06, AME���PME 0.14, ALE���PLE 0, PME���PME 0.12, PME���PLE 0.16. Leg measurements: I 8.03 (2.34, 2.66, 1.72, 1.31); II 7.11 (2.13, 2.25, 1.56, 1.17); III 6.72 (1.88, 2.03, 1.72, 1.09); IV 8.72 (2.38, 2.66, 2.38, 1.30). Epigyne: more than 1.5 times wider than long, epigynal teeth about 3 times wider than long, located anteriorly and close together, hoods distinct, atrium small, close to the posterior margin of the epigynue, about 2.5 times wider than long, epigyne 3 times wider than atrium, copulatory openings near the center of the atrium, posterior margin of atrium 3.5 times wider than long; spermathecae spherical, as long as wide, spermathecal heads originating anteriorly from the spermathecae, copulatory ducts wide; fertilization ducts 4 times longer than wide (Fig. 7 A���B). Male. Unknown. Variation. Total length of females 6.34, 6.53 (n=2). Distribution. Known only from Tibet, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, Four new species of the genus Draconarius Ovtchinnikov, 1999 (Araneae, Agelenidae) from the Tibetan Plateau, China, pp. 141-154 in Zootaxa 4648 (1) on pages 149-152, DOI: 10.11646/zootaxa.4648.1.7, http://zenodo.org/record/3353959, {"references":["Peng, X. & Yin, C. (1998) Four new species of the genus Coelotes (Araneae, Agelenidae) from China. Bulletin of the British Arachnological Society, 11, 26 - 28."]}

Draconarius baibaensis Z. Zhao & S. Li, sp. n. Figs 1, 8 Type material. Holotype ♀ (IZCAS-Ar39616): Baiba Township (N29.79867��, E93.84720��, 3191 m), Bayi County, Nyingchi City, Tibet, China, 31. VII. 2012, Z.Y. Yao and Z. Zhao leg. Etymology. The specific name refers to the type locality, Baiba Township; adjective. Diagnosis. Draconarius baibaensis sp. n. is most similar to D. altissimus (Hu, 2001). The epigyne of both species have a small atrium located posteriorly, and the epigynal teeth are pointed at the end, with spermathecal heads long and thin. However, the two species differ by the location of the epigynal teeth (medially in D. baibaensis sp. n. in Fig. 1A and posteriorly in D. altissimus in fig. 43.1 in Hu 2001 and fig. 112A in Zhu et al. 2017) and the shape of the posterior margin of the atrium (rectangular in D. baibaensis sp. n. in Fig. 1A and trapezoidal in D. altissimus in fig. 43.1 in Hu 2001 and fig. 112A in Zhu et al. 2017). Description. Female (IZCAS-Ar39616). Cephalothorax brown. The clypeus, endites, labium darker than carapace but colour of sternum and legs the same as carapace. Abdomen light-brown, covered by gray hairs. The colour of spinnerets same as carapace. Total length 11.29. Carapace 5.20 long, 3.61 wide. Abdomen 6.09 long, 3.96 wide. Eye sizes and interdistances: AME 0.12, ALE 0.21, PME 0.15, PLE 0.18; AME���AME 0.12, AME���ALE 0.11, AME���PME 0.13, ALE���PLE 0, PME���PME 0.14, PME���PLE 0.04. Leg measurements: I 15.39 (4.84, 4.99, 3.36, 2.20); II 13.86 (4.18, 4.49, 3.21, 1.98); III 12.78 (3.91, 3.80, 3.16, 1.91); IV 16.74 (4.97, 5.04, 4.49, 2.24). Epigyne: subequal to 2 times wider than long with 2 triangular teeth, pointed at end, medially situated and widely separated, hoods distinct, atrium small and located posteriorly, posterior margin of atrium 4 times wider than long; spermathecae 1.5 times longer than wide, anteriorly wider than posteriorly, spermathecal heads long and thin, originating medially from the spermathecae; the width of fertilization ducts subequal to its length (Fig. 1 A���B). Male. Unknown. Distribution. Known only from Tibet, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, Four new species of the genus Draconarius Ovtchinnikov, 1999 (Araneae, Agelenidae) from the Tibetan Plateau, China, pp. 141-154 in Zootaxa 4648 (1) on page 143, DOI: 10.11646/zootaxa.4648.1.7, http://zenodo.org/record/3353959, {"references":["Hu, J. (2001) Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp.","Zhu, M., Wang, X. & Zhang, Z. (2017) Fauna Sinica: Invertebrata Vol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp."]}

Draconarius yigongensis Z. Zhao & S. Li, sp. n. Figs 5–6, 8 Type material. Holotype ♂ (IZCAS-Ar39622): Yigong Village (N30.12525º, E95.03213º, 2072 m), Bomi County, Nyingchi City, Tibet, China, 12. VIII. 2014, Y.C. Li leg. Paratypes: 1♀ (IZCAS-Ar39623): same data as holotype; 1♀ (IZCAS-Ar39624): Yigong Village (N30.26715º, E94.77855º, 2183 m), Bomi County, Nyingchi City, Tibet, China, 13. VIII. 2014, Y.C. Li leg. Etymology. The specific name refers to the type locality, Yigong Village; adjective. Diagnosis. Draconarius yigongensis sp. n. is most similar to D. subdissitus Zhu, Wang & Zhang, 2017. Males of these species can be separated by the length of the patellar apophysis (short, 1/2 width and equal to length of RTA in D. yigongensis sp. n. in Fig. 5C and 3 times longer than RTA in D. subdissitus in fig. 243E in Zhu et al. 2017), the cymbial furrow (about 2/3 cymbial length in D. yigongensis sp. n. in Fig. 5C and 1 /2 cymbial length in D. subdissitus in fig. 243E in Zhu et al. 2017) and the end of the conductor (extending forward in D. yigongensis sp. n. in Fig. 5C but not in D. subdissitus; see fig. 243E in Zhu et al. 2017). Females of similar species can be separated by the position of the epigyne teeth (originating from the center of the epigyne in D. yigongensis sp. n. in Fig. 6A and nearly the lateral margin of the atrium in D. subdissitus in fig. 243A in Zhu et al. 2017), the shape of the posterior margin of the atrium (nearly rectangular in D. yigongensis sp. n. in Fig. 6A and uplifted centrally in D. subdissitus in fig. 243A in Zhu et al. 2017); and the size of the spermathecal heads (short in D. yigongensis sp. n. in Fig. 6B and long in D. subdissitus in fig. 243B in Zhu et al. 2017). Description. Male (IZCAS -Ar39622). Carapace, clypeus, endites, labium, sternum and spinnerets brown, legs and abdomen light-brown. Abdomen with five chevron-shaped patterns. Total length 5.81. Carapace 2.84 long, 1.87 wide. Abdomen 2.97 long, 1.91 wide. Eye sizes and interdistances: AME 0.06, ALE 0.15, PME 0.12, PLE 0.14; AME–AME 0.06, AME–ALE 0.11, AME–PME 0.10, ALE–PLE 0, PME–PME 0.10, PME–PLE 0.06. Leg measurements: I 8.28 (2.34, 2.81, 1.88, 1.25); II 7.11 (2.19, 2.19, 1.56, 1.17); III 6.72 (2.03, 1.88, 1.72, 1.09); IV 8.98 (2.65, 2.66, 2.34, 1.33). Palp: Femur 4.2 times longer than wide. Patellar apophysis almost entirely reduced, RTA small, VTA spine-like, long, about 3 times longer than RTA. Patella with anterodorsal long macrosetae, almost 1.5 times longer than patella, and tibia and base of cymbium with long dorsal and prolateral macrosetae. Cymbium 1.6 times longer than bulb, cymbial furrow long, about 2/3 cymbial length; conductor short, deeply grooved, with large basal lamella and triangular dorsal apophysis; embolus originates at about the 6 o'clock position (Fig. 5 A–C). Female (IZCAS -Ar39623). The colour and the abdominal pattern as the same as the males. Total length 6.33. Carapace 3.12 long, 2.03 wide. Abdomen 3.21 long, 2.19 wide. Eye sizes and interdistances: AME 0.09, ALE 0.14, PME 0.12, PLE 0.15; AME–AME 0.09, AME–ALE 0.13, AME–PME 0.12, ALE–PLE 0, PME–PME 0.16, PME–PLE 0.10. Leg measurements: leg I 7.81 (2.34, 2.51, 1.72, 1.24); II 6.53 (2.10, 2.13, 1.21, 1.09); III 6.41 (2.03, 1.88, 1.48, 1.02); IV 8.15 (2.51, 2.51, 1.88, 1.25). Epigyne: subequal to 2 times wider than long with 2 weak teeth, pointed at end, medially situated and widely separated, atrium small and close to the posterior margin of the epigyne, posterior margin of atrium 4 times wider than long; spermathecae 2 times longer than wide, spermathecal heads originating medially from the spermathecae; fertilization ducts 2 times longer than wide (Fig. 6 A–B). Variation. Total length of females 5.87, 6.33 (n=2). Distribution. Known only from Tibet, China (Fig. 8).

Genus Draconarius Ovtchinnikov, 1999 Type species. Draconarius venustus Ovtchinnikov, 1999, by original designation. Diagnosis and description. Male palp with cymbial furrow about 1/2 cymbial length (Fig. 3B) or more than 1/2 cymbial length (Fig. 5B); conductor short and pointed at the end (Figs 3B, 5B); lamella of conductor large, the left side higher than the right in ventral view (Figs 3B, 5B); median apophysis spoon-shaped. The epigyne with short teeth, medially situated and widely separated (Figs 1A, 2A, 4A, 6A) or anteriorly situated and close together (Fig. 7A); copulatory ducts short; spermathecae large and broad; spermathecal heads located anteriorly (Figs 2B, 7B) or medially (Figs 1B, 4B, 6B). For other diagnostic characters, see Ovtchinnikov (1999) and Wang (2003)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, Four new species of the genus Draconarius Ovtchinnikov, 1999 (Araneae, Agelenidae) from the Tibetan Plateau, China, pp. 141-154 in Zootaxa 4648 (1) on page 142, DOI: 10.11646/zootaxa.4648.1.7, http://zenodo.org/record/3353959, {"references":["Ovtchinnikov, S. V. (1999) On the supraspecific systematics of the subfamily Coelotinae (Araneae, Amaurobiidae) in the former USSR fauna. Tethys Entomological Research, 1, 63 - 80.","Wang, X. (2003) Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences, 54, 499 - 662."]}

Draconarius budanlaensis Z. Zhao & S. Li, sp. n. Figs 2, 8 Type material. Holotype ♀ (IZCAS-Ar39617): Budanla Mountain (N29.02890º, E92.35338º, 4930 m), Nyalam County, Shigatse, Tibet, China, 10. VIII. 2014, Y.C. Li leg. Etymology. The specific name refers to the type locality, Budanla Mountain; adjective. Diagnosis. Draconarius budanlaensis sp. n. is most similar to D. yadongensis (Hu & Li, 1987). The epigyne of both species has broad teeth that are widely separated; atrium flat and wide; spermathecae close together medially. The two species differ by the locations of the epigynal teeth (laterally in D. budanlaensis sp. n. in Fig. 2A and centrally in D. yadongensis in fig. 20.1 in Hu & Li 1987); the shape of the epigynal teeth (broad and short in D. budanlaensis sp. n. in Fig. 2A and as wide as long in D. yadongensis in fig. 20.1 in Hu & Li 1987), and the length of the posterior margin of the atrium (1/3 as long as wide in D. budanlaensis sp. n. in Fig. 2A and 5 times wider than long in D. yadongensis in fig. 20.1 in Hu & Li 1987). Description. Female (IZCAS-Ar39617). Carapace and legs brown. Clypeus, endites, labium and sternum black-brown. Abdomen grey-brown. The colour of spinnerets equals the carapace. Abdomen with five chevronshaped patterns. Total length 14.10. Carapace 6.41 long, 4.16 wide. Abdomen 7.69 long, 3.95 wide. Eye sizes and interdistances: AME 0.16, ALE 0.20, PME 0.18, PLE 0.19; AME–AME 0.21, AME–ALE 0.17, AME–PME 0.22, ALE–PLE 0, PME–PME 0.29, PME–PLE 0.28. Leg measurements: I 16.79 (4.49, 5.77, 3.92, 2.61); II 15.69 (4.16, 5.13, 3.80, 2.60); III 15.54 (4.01, 4.81, 4.16, 2.56); IV 19.48 (4.80, 6.09, 5.38, 3.21). Epigyne: about 1.5 times wider than long, with 2 broad teeth that are about 3 times wider than long, situated anteriorly and widely separated, hoods distinct, atrium flat and wide, about 1.7 times wider than long, posterior margin of atrium 3 times wider than long; spermathecae longer than wide, spermathecal heads originating anteriorly from the spermathecae; fertilization ducts 3 times wider than long (Fig. 2 A–B). Male. Unknown. Distribution. Known only from Tibet, China (Fig. 8).

Draconarius yingbinensis Z. Zhao & S. Li, sp. n. Figs 7–8 Type material. Holotype ♀ (IZCAS-Ar29625): Yingbin Road (N27.98375º, E85.98287º, 2450 m), Zhangmu Township, Nyalam County, Shigatse, Tibet, China, 29. VII. 2014, Y.C. Li leg. Paratype: 1♀ (IZCAS-Ar39626): same data as holotype. Etymology. The specific name refers to the type locality, Yingbin Road; adjective. Diagnosis. Draconarius yingbinensis sp. n. is most similar to D. nanyuensis (Peng & Yin, 1998). The epigynal teeth of both species are located anteriorly, close together, bent at the end, and have the same length to width ratio; atrium small, close to the posterior margin of the epigyne; spermathecae spherical, spermathecal heads originating anteriorly from the spermathecae. The two species differ by the shape of the epigynal teeth at the end (blunt in D. yingbinensis sp. n. in Fig. 7A and pointed in D. nanyuensis in fig. 8 in Peng & Yin 1998); the shape of the atrium (rectangular in D. yingbinensis sp. n. in Fig. 7A and with a triangular septum with atrium on both sides of septum in D. nanyuensis in fig. 8 in Peng & Yin 1998). Description. Female (IZCAS-Ar39625). Carapace, sternum, legs, abdomen and spinnerets brown; clypeus, endites and labium darker than carapace.Abdomen with five chevron-shaped patterns and covered by blueish-green hairs. Total length 6.34. Carapace 3.48 long, 2.23 wide. Abdomen 2.86 long, 2.01 wide. Eye sizes and interdistances: AME 0.11, ALE 0.20, PME 0.11, PLE 0.16; AME–AME 0.08, AME–ALE 0.06, AME–PME 0.14, ALE–PLE 0, PME–PME 0.12, PME–PLE 0.16. Leg measurements: I 8.03 (2.34, 2.66, 1.72, 1.31); II 7.11 (2.13, 2.25, 1.56, 1.17); III 6.72 (1.88, 2.03, 1.72, 1.09); IV 8.72 (2.38, 2.66, 2.38, 1.30). Epigyne: more than 1.5 times wider than long, epigynal teeth about 3 times wider than long, located anteriorly and close together, hoods distinct, atrium small, close to the posterior margin of the epigynue, about 2.5 times wider than long, epigyne 3 times wider than atrium, copulatory openings near the center of the atrium, posterior margin of atrium 3.5 times wider than long; spermathecae spherical, as long as wide, spermathecal heads originating anteriorly from the spermathecae, copulatory ducts wide; fertilization ducts 4 times longer than wide (Fig. 7 A–B). Male. Unknown. Variation. Total length of females 6.34, 6.53 (n=2). Distribution. Known only from Tibet, China (Fig. 8).

Lineacoelotes tiantaiensis Z. Zhao & S. Li, sp. n. Figs 3, 8 Type material. Holotype ♀ (IZCAS-Ar39285): Tiantai National Forest Park (N33.249180°, E107.054280°, 1074 m), Wuxiang Township, Hantai District, Hanzhong City, Shaanxi Province, China, 25. X. 2013, Y. Li and J. Liu leg. Paratypes: 2♀ (IZCAS-Ar39286 - Ar39287): same data as holotype. Etymology. The specific name referring to the type locality: Tiantai National Forest Park. Diagnosis. Females of L. tiantaiensis sp. n. are similar to those of L. funiushanensis. They can be distinguished from the latter species by the atrium thin and long, and epigynal teeth long (Fig. 3A) vs. atrium wide and short, and epigynal teeth short in L. funiushanensis (fig. 5 in Xu et al., 2008). Description. Female (holotype). Carapace and clypeus brown-yellow, endites and labium dark brown, sternum brown-yellow, the colour of legs lighter than that of carapace, with a grey ring of hair, femur surrounded by two ring hair, patella surrounded by one ring of hair, tibia surrounded by one ring of hair. Abdomen blue-brown, dorsal view of abdomen with five chevron-shaped patterns; the colour of spinnerets and abdomen are the same. Total length 7.04. Carapace 3.40 long, 2.66 wide. Abdomen 3.64 long, 2.38 wide. Eye sizes and interdistances: AME 0.12, ALE 0.19, PME 0.16, PLE 0.18; AME-AME 0.06, AME-ALE 0.07, AME-PME 0.09, ALE-PLE 0, PME-PME 0.11, PME-PLE 0.15. Leg measurements: I 11.62 (3.37, 3.84, 2.74, 1.67); II 10.19 (3.01, 3.43, 2.12, 1.63); III 9.76 (3.07, 3.08, 2.44, 1.17); IV 12.54 (3.37, 3.92, 3.43, 1.82). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 1.3 times wider than high; epigynal teeth long and wide, bent at tips; originating from anterior margin of the atrium and extending over its posterior margin, about 1/2 the height of plate, with 2.8 times longer than wide; the distance between the base of the tooth is shorter than the distance between the tip of the tooth (2.5 times of their width); epigynal atrium situated at the central part of the plate, longer than wide, with their posterior part more sclerotized than other parts of the plate; copulatory ducts large; spermathecae oval, separated from each other; spermathecal heads long, originating from the middle part of the spermathecae; fertilization duct pointing to the front, hook-shaped at the tips (Fig. 3 A–B). Male. Unknown. Variation. Total length of females 6.09–8.33. Carapace length 2.36–2.82. Distribution. Known only from Shaanxi Province, China (Fig. 8).

Lineacoelotes zhongbaensis Z. Zhao & S. Li, sp. n. Figs 4���5, 8 Type material. Holotype ♂ (IZCAS-Ar39288): Zhongba Village (N32.912870��, E108.135530��, 489 m), Houliu Township, Shiquan County, Ankang City, Shaanxi Province, China, 27. IX. 2013, Y. Li and J. Liu leg. Paratypes: 1♂ 1♀ (IZCAS-Ar39289-Ar39290): same data as holotype. Etymology. The specific name referring to the type locality: Zhongba Village. Diagnosis. Males of L. zhongbaensis sp. n. are most similar to those of L. funiushanensis. They can be distinguished from the latter by the patellar apophysis extending over the anterior edge of the patella, while the tip of the distal apophysis of the conductor (C2) is pointed in the new species (Fig. 4 B���C) vs. patellar apophysis less than the anterior edge of patella and C2 blunt in L. funiushanensis (figs 8���10 in Xu et al., 2008). Females of L. zhongbaensis sp. n. are most similar to those of other Lineacoelotes species. They can be distinguished by the spermathecal heads that are bifurcate at the tip (Fig. 5B), and the epigynal teeth that are merged at base and located on the anterior edge of epigynal plate (Fig. 5A). Description. Male (holotype). Carapace, clypeus, endites and labium brown, sternum lighter than carapace, legs darker than carapace. Abdomen grey-brown, dorsal view of abdomen with five chevron-shaped patterns, spinnerets and abdominal similar in colour and slightly yellowish-brown. Total length 6.61. Carapace 3.12 long, 2.23 wide. Abdomen 3.49 long, 1.98 wide. Eye sizes and interdistances: AME 0.13, ALE 0.17, PME 0.15, PLE 0.19; AME-AME 0.05, AME-ALE 0.05, AME-PME 0.08, ALE-PLE 0, PME-PME 0.08, PME-PLE 0.07. Leg measurements: I 12.53 (3.45, 4.18, 2.82, 2.08); II 10.55 (2.92, 3.24, 2.58, 1.81); III 10.24 (2.93, 3.01, 2.69, 1.61); IV 13.88 (3.70, 4.05, 3.92, 2.21). Chelicera with 3 promarginal and 2 retromarginal teeth. Palp: femur 2.5 times longer than wide; patella 2.4 times longer than wide; tibia 3.1 times longer than wide; patellar seta shorter than patellar; patella with 2 apophyses, lateral and dorsal; lateral one patellar apophysis wide and long, width about 1/2 the width of the patella, about 1.5 times longer than patella, dorsal one long, length about 1/2 the length of the patella; retrolateral tibial apophysis extending to the anterior margin of tibia, narrower than patellar apophysis and with a pointed tip; cymbial furrow about 1/2 the width of the cymbium and less than 1/2 of the cymbial length; embolus beginning at 7:30 o���clock position, long and distorted; conductor with two apophyses; the distal apophysis (C2) larger than the ventral apophysis (C3); median apophysis spoon-shaped (Fig. 4 A���C). Female (IZCAS -Ar39290). Spinnerets slightly cyan-blue, otherwise same as male. Total length 7.05. Carapace 3.38 long, 2.26 wide. Abdomen 3.67 long, 2.32 wide. Eye sizes and interdistances: AME 0.11, ALE 0.16, PME 0.14, PLE 0.15; AME-AME 0.05, AME-ALE 0.05, AME-PME 0.09, ALE-PLE 0, PME-PME 0.07, PME-PLE 0.09. Leg measurements: I 9.41 (2.56, 3.47, 1.93, 1.45); II 8.46 (2.53, 2.69, 1.92, 1.32); III 8.18 (2.36, 2.43, 2.08, 1.31); IV 10.12 (2.59, 3.47, 2.69, 1.37). Leg measurements: leg I missing; II 11.60 (3.53, 3.59, 2.56, 1.92); III 10.97 (3.21, 3.27, 2.82, 1.67); IV 14.56 (4.17, 4.49, 3.85, 2.05). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 1.7 times wider than high; epigynal teeth long, length of teeth about 4/5 the height of plate, with merged bases while pointed and bent at tips; originating from the anterior margin of epigyne, extending over posterior margin of atrium, 4 times longer than wide; the distance between tooth about 1.5 times of its width; epigynal atrium situated at the central part of epigyne, wider than long; copulatory ducts originating from the posterior margin of the atrium and large; spermathecae oval, next to each other; spermathecal heads long and originating from the anterior part of the spermathecae, bifurcated at the tip; fertilization duct points to the left, hook-shaped at the tips (Fig. 5 A���B). Variation. Total length of males 6.61���7.11. Carapace length 3.12���3.38. Distribution. Known only from Shaanxi, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, New species of the coelotine spider genus Lineacoelotes (Araneae: Agelenidae) from China, pp. 351-363 in Zootaxa 4623 (2) on pages 354-359, DOI: 10.11646/zootaxa.4623.2.9, http://zenodo.org/record/3255507, {"references":["Xu, X., Li, S. & Wang, X. (2008) Lineacoelotes, a new genus of Coelotinae from China (Araneae: Amaurobiidae). Zootaxa, 1700, 1 - 20."]}

Lineacoelotes lifengyuanae Z. Zhao & S. Li, sp. n. Figs 1–2, 8 Type material. Holotype ♂ (IZCAS-Ar39282): Wuma Village (N30.040500°, E106.572630°, 799 m), Jindaoxia Township, Beibei District, Chongqing Municipality, China, 30.IV. 2014, Y. Lin, H. Zhao, Y. Li, J. Wu and F. Li leg. Paratypes: 1♂ 1♀ (IZCAS-Ar39283-Ar39284): same data as holotype. Etymology. The specific name is a paronym for Fengyuan Li, one of the collectors of the new species, to commemorate her contribution to the new species. Diagnosis. Males of L. lifengyuanae sp. n. are most similar to those of L. ziboensis sp. n. They can be distinguished from the latter by the anterior distal apophysis of the conductor dorsal edge enlarged while the posterior distal apophysis of the conductor dorsal edge is small (Fig. 1B) vs. the anterior distal apophysis of the conductor dorsal edge small while the posterior distal apophysis of the conductor dorsal edge is enlarged in L. ziboensis sp. n. (Fig. 6B). Females of L. lifengyuanae sp. n. are most similar to those of other Lineacoelotes species. They can be distinguished from congeners by having long fertilization ducts (Fig. 2B) and tips of the epigynal teeth that bend abruptly (Fig. 2A). Description. Male (holotype). Carapace and clypeus brown, endites and labium dark brown, colour of sternum the same as carapace, legs brown but lighter than carapace. Abdomen blue-gray, dorsal view of abdomen with four chevron-shaped patterns, colour of spinnerets same as abdominal color. Total length 7.04. Carapace 3.40 long, 2.66 wide. Abdomen 3.64 long, 2.38 wide. Eye sizes and interdistances: AME 0.12, ALE 0.19, PME 0.22, PLE 0.21; AME-AME 0.05, AME-ALE 0.09, AME-PME 0.09, ALE-PLE 0, PME-PME 0.10, PME-PLE 0.11. Leg measurements: I 15.71 (4.17, 5.06, 3.85, 2.63); II 15.07 (3.97, 4.81, 3.85, 2.44); III 13.85 (3.60, 4.16, 3.85, 2.24); IV 17.31 (4.36, 5.13, 5.13, 2.69). Chelicera with 3 promarginal and 2 retromarginal teeth. Palp: femur 3.2 times longer than wide; patella 2.2 times longer than wide; tibia 3.1 times longer than wide; patella seta the same length as patella; patellar apophysis bent and long, extending to the anterior margin of the tibia, with pointed tip; retrolateral tibial apophysis wider than patellar apophysis, extending over tibia and with a blunt tip; cymbial furrow about 1/2 the length of the cymbium; embolus distorted, originating at 6 o’clock position; conductor with two apophyses (C2 and C3), one broad groove and one broad lamella; median apophysis spoon-shaped in ventral view; fertilization duct point to the left, colour lighter than spermathecal colour (Fig. 1 A–C). Female (IZCAS -Ar39284). Abdomen grey-brown, otherwise same as in male. Total length 9.94. Carapace 4.17 long, 2.56 wide.Abdomen 5.77 long, 3.85 wide. Eye sizes and interdistances:AME 0.11, ALE 0.16, PME 0.15, PLE 0.17; AME-AME 0.10, AME-ALE 0.08, AME-PME 0.10, ALE-PLE 0, PME-PME 0.05, PME-PLE 0.11. Leg measurements: leg I missing; II 11.60 (3.53, 3.59, 2.56, 1.92); III 10.97 (3.21, 3.27, 2.82, 1.67); IV 14.56 (4.17, 4.49, 3.85, 2.05). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 2.2 times wider than high, with distinct notch posteriorly; epigynal teeth long, length of teeth about 3/4 the height of plate, pointed and bent at tips, extending close to the posterior margin of the epigyne and about 4 times longer than wide; the distance between tooth about 2.5 times of their width; epigynal atrium more than 2 times wider than high; copulatory ducts large; spermathecae spherical and close together about 1/6 of their width; spermathecal heads originating from the anterior part of the spermathecae, through the ventral side of the copulatory duct (Fig. 2 A–B). Variation. Total length of males 6.41–7.04. Carapace length 3.40–4.17. Distribution. Known only from Chongqing Municipality, China (Fig. 8).

Lineacoelotes zhongbaensis Z. Zhao & S. Li, sp. n. Figs 4–5, 8 Type material. Holotype ♂ (IZCAS-Ar39288): Zhongba Village (N32.912870°, E108.135530°, 489 m), Houliu Township, Shiquan County, Ankang City, Shaanxi Province, China, 27. IX. 2013, Y. Li and J. Liu leg. Paratypes: 1♂ 1♀ (IZCAS-Ar39289-Ar39290): same data as holotype. Etymology. The specific name referring to the type locality: Zhongba Village. Diagnosis. Males of L. zhongbaensis sp. n. are most similar to those of L. funiushanensis. They can be distinguished from the latter by the patellar apophysis extending over the anterior edge of the patella, while the tip of the distal apophysis of the conductor (C2) is pointed in the new species (Fig. 4 B–C) vs. patellar apophysis less than the anterior edge of patella and C2 blunt in L. funiushanensis (figs 8–10 in Xu et al., 2008). Females of L. zhongbaensis sp. n. are most similar to those of other Lineacoelotes species. They can be distinguished by the spermathecal heads that are bifurcate at the tip (Fig. 5B), and the epigynal teeth that are merged at base and located on the anterior edge of epigynal plate (Fig. 5A). Description. Male (holotype). Carapace, clypeus, endites and labium brown, sternum lighter than carapace, legs darker than carapace. Abdomen grey-brown, dorsal view of abdomen with five chevron-shaped patterns, spinnerets and abdominal similar in colour and slightly yellowish-brown. Total length 6.61. Carapace 3.12 long, 2.23 wide. Abdomen 3.49 long, 1.98 wide. Eye sizes and interdistances: AME 0.13, ALE 0.17, PME 0.15, PLE 0.19; AME-AME 0.05, AME-ALE 0.05, AME-PME 0.08, ALE-PLE 0, PME-PME 0.08, PME-PLE 0.07. Leg measurements: I 12.53 (3.45, 4.18, 2.82, 2.08); II 10.55 (2.92, 3.24, 2.58, 1.81); III 10.24 (2.93, 3.01, 2.69, 1.61); IV 13.88 (3.70, 4.05, 3.92, 2.21). Chelicera with 3 promarginal and 2 retromarginal teeth. Palp: femur 2.5 times longer than wide; patella 2.4 times longer than wide; tibia 3.1 times longer than wide; patellar seta shorter than patellar; patella with 2 apophyses, lateral and dorsal; lateral one patellar apophysis wide and long, width about 1/2 the width of the patella, about 1.5 times longer than patella, dorsal one long, length about 1/2 the length of the patella; retrolateral tibial apophysis extending to the anterior margin of tibia, narrower than patellar apophysis and with a pointed tip; cymbial furrow about 1/2 the width of the cymbium and less than 1/2 of the cymbial length; embolus beginning at 7:30 o’clock position, long and distorted; conductor with two apophyses; the distal apophysis (C2) larger than the ventral apophysis (C3); median apophysis spoon-shaped (Fig. 4 A–C). Female (IZCAS -Ar39290). Spinnerets slightly cyan-blue, otherwise same as male. Total length 7.05. Carapace 3.38 long, 2.26 wide. Abdomen 3.67 long, 2.32 wide. Eye sizes and interdistances: AME 0.11, ALE 0.16, PME 0.14, PLE 0.15; AME-AME 0.05, AME-ALE 0.05, AME-PME 0.09, ALE-PLE 0, PME-PME 0.07, PME-PLE 0.09. Leg measurements: I 9.41 (2.56, 3.47, 1.93, 1.45); II 8.46 (2.53, 2.69, 1.92, 1.32); III 8.18 (2.36, 2.43, 2.08, 1.31); IV 10.12 (2.59, 3.47, 2.69, 1.37). Leg measurements: leg I missing; II 11.60 (3.53, 3.59, 2.56, 1.92); III 10.97 (3.21, 3.27, 2.82, 1.67); IV 14.56 (4.17, 4.49, 3.85, 2.05). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 1.7 times wider than high; epigynal teeth long, length of teeth about 4/5 the height of plate, with merged bases while pointed and bent at tips; originating from the anterior margin of epigyne, extending over posterior margin of atrium, 4 times longer than wide; the distance between tooth about 1.5 times of its width; epigynal atrium situated at the central part of epigyne, wider than long; copulatory ducts originating from the posterior margin of the atrium and large; spermathecae oval, next to each other; spermathecal heads long and originating from the anterior part of the spermathecae, bifurcated at the tip; fertilization duct points to the left, hook-shaped at the tips (Fig. 5 A–B). Variation. Total length of males 6.61–7.11. Carapace length 3.12–3.38. Distribution. Known only from Shaanxi, China (Fig. 8).

Lineacoelotes lifengyuanae Z. Zhao & S. Li, sp. n. Figs 1���2, 8 Type material. Holotype ♂ (IZCAS-Ar39282): Wuma Village (N30.040500��, E106.572630��, 799 m), Jindaoxia Township, Beibei District, Chongqing Municipality, China, 30.IV. 2014, Y. Lin, H. Zhao, Y. Li, J. Wu and F. Li leg. Paratypes: 1♂ 1♀ (IZCAS-Ar39283-Ar39284): same data as holotype. Etymology. The specific name is a paronym for Fengyuan Li, one of the collectors of the new species, to commemorate her contribution to the new species. Diagnosis. Males of L. lifengyuanae sp. n. are most similar to those of L. ziboensis sp. n. They can be distinguished from the latter by the anterior distal apophysis of the conductor dorsal edge enlarged while the posterior distal apophysis of the conductor dorsal edge is small (Fig. 1B) vs. the anterior distal apophysis of the conductor dorsal edge small while the posterior distal apophysis of the conductor dorsal edge is enlarged in L. ziboensis sp. n. (Fig. 6B). Females of L. lifengyuanae sp. n. are most similar to those of other Lineacoelotes species. They can be distinguished from congeners by having long fertilization ducts (Fig. 2B) and tips of the epigynal teeth that bend abruptly (Fig. 2A). Description. Male (holotype). Carapace and clypeus brown, endites and labium dark brown, colour of sternum the same as carapace, legs brown but lighter than carapace. Abdomen blue-gray, dorsal view of abdomen with four chevron-shaped patterns, colour of spinnerets same as abdominal color. Total length 7.04. Carapace 3.40 long, 2.66 wide. Abdomen 3.64 long, 2.38 wide. Eye sizes and interdistances: AME 0.12, ALE 0.19, PME 0.22, PLE 0.21; AME-AME 0.05, AME-ALE 0.09, AME-PME 0.09, ALE-PLE 0, PME-PME 0.10, PME-PLE 0.11. Leg measurements: I 15.71 (4.17, 5.06, 3.85, 2.63); II 15.07 (3.97, 4.81, 3.85, 2.44); III 13.85 (3.60, 4.16, 3.85, 2.24); IV 17.31 (4.36, 5.13, 5.13, 2.69). Chelicera with 3 promarginal and 2 retromarginal teeth. Palp: femur 3.2 times longer than wide; patella 2.2 times longer than wide; tibia 3.1 times longer than wide; patella seta the same length as patella; patellar apophysis bent and long, extending to the anterior margin of the tibia, with pointed tip; retrolateral tibial apophysis wider than patellar apophysis, extending over tibia and with a blunt tip; cymbial furrow about 1/2 the length of the cymbium; embolus distorted, originating at 6 o���clock position; conductor with two apophyses (C2 and C3), one broad groove and one broad lamella; median apophysis spoon-shaped in ventral view; fertilization duct point to the left, colour lighter than spermathecal colour (Fig. 1 A���C). Female (IZCAS -Ar39284). Abdomen grey-brown, otherwise same as in male. Total length 9.94. Carapace 4.17 long, 2.56 wide.Abdomen 5.77 long, 3.85 wide. Eye sizes and interdistances:AME 0.11, ALE 0.16, PME 0.15, PLE 0.17; AME-AME 0.10, AME-ALE 0.08, AME-PME 0.10, ALE-PLE 0, PME-PME 0.05, PME-PLE 0.11. Leg measurements: leg I missing; II 11.60 (3.53, 3.59, 2.56, 1.92); III 10.97 (3.21, 3.27, 2.82, 1.67); IV 14.56 (4.17, 4.49, 3.85, 2.05). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 2.2 times wider than high, with distinct notch posteriorly; epigynal teeth long, length of teeth about 3/4 the height of plate, pointed and bent at tips, extending close to the posterior margin of the epigyne and about 4 times longer than wide; the distance between tooth about 2.5 times of their width; epigynal atrium more than 2 times wider than high; copulatory ducts large; spermathecae spherical and close together about 1/6 of their width; spermathecal heads originating from the anterior part of the spermathecae, through the ventral side of the copulatory duct (Fig. 2 A���B). Variation. Total length of males 6.41���7.04. Carapace length 3.40���4.17. Distribution. Known only from Chongqing Municipality, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, New species of the coelotine spider genus Lineacoelotes (Araneae: Agelenidae) from China, pp. 351-363 in Zootaxa 4623 (2) on pages 352-354, DOI: 10.11646/zootaxa.4623.2.9, http://zenodo.org/record/3255507

Genus Lineacoelotes Xu, Li & Wang, 2008 Lineacoelotes Xu, Li & Wang, 2008: 4. Type species. Lineacoelotes longicephalus Xu, Li & Wang, 2008, from Sichuan Province. Diagnosis and description. Patellae of male palp (lateral view) with a strong seta that is located on a swollen and convex base. For more detail diagnosis and description, see Xu, Li & Wang, 2008. Comments. Lineacoelotes was originally described for five species: L. bicultratus (Chen, Zhao & Wang, 1991), L. funiushanensis (Hu, Wang & Wang, 1991), L. nitidus (Li & Zhang, 2002), L. longicephalus Xu, Li & Wang, 2008, and L. strenuus Xu, Li & Wang, 2008. Based on these data and the new species described below, the genus is distributed in central (Henan, Hubei), southwestern (Chongqing, Guizhou, Sichuan) China., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, New species of the coelotine spider genus Lineacoelotes (Araneae: Agelenidae) from China, pp. 351-363 in Zootaxa 4623 (2) on page 352, DOI: 10.11646/zootaxa.4623.2.9, http://zenodo.org/record/3255507, {"references":["Wang, X., Xu, X. & Li, S. (2008) Notiocoelotes, a new genus of the spider subfamily Coelotinae from Southeast Asia (Araneae, Amaurobiidae). Zootaxa, 1853, 1 - 17.","Chen, J., Zhao, J. & Wang, J. (1991) Two new species of spider of the genus Coelotes from Wudang Mountain, China (Araneae: Agelenidae). Journal of Hubei University, Natural Science Edition, 13, 9 - 12. https: // doi. org / 10.1007 / s 11859 - 008 - 0103 - 7","Hu, J., Wang, Z. & Wang, Z. (1991) Notes on nine species of spiders from natural conservation of Baotianman in Henan Province, China (Arachnoidea: Araneida). Henan Science, 9, 37 - 52.","Li, S. & Zhang, B. (2002) A new species of the genus Coelotes from Tenglongdong Cave, Hubei, China (Araneae: Amaurobiidae). Acta Zootaxonomica Sinica, 27, 466 - 468."]}

Lineacoelotes tiantaiensis Z. Zhao & S. Li, sp. n. Figs 3, 8 Type material. Holotype ♀ (IZCAS-Ar39285): Tiantai National Forest Park (N33.249180��, E107.054280��, 1074 m), Wuxiang Township, Hantai District, Hanzhong City, Shaanxi Province, China, 25. X. 2013, Y. Li and J. Liu leg. Paratypes: 2♀ (IZCAS-Ar39286 - Ar39287): same data as holotype. Etymology. The specific name referring to the type locality: Tiantai National Forest Park. Diagnosis. Females of L. tiantaiensis sp. n. are similar to those of L. funiushanensis. They can be distinguished from the latter species by the atrium thin and long, and epigynal teeth long (Fig. 3A) vs. atrium wide and short, and epigynal teeth short in L. funiushanensis (fig. 5 in Xu et al., 2008). Description. Female (holotype). Carapace and clypeus brown-yellow, endites and labium dark brown, sternum brown-yellow, the colour of legs lighter than that of carapace, with a grey ring of hair, femur surrounded by two ring hair, patella surrounded by one ring of hair, tibia surrounded by one ring of hair. Abdomen blue-brown, dorsal view of abdomen with five chevron-shaped patterns; the colour of spinnerets and abdomen are the same. Total length 7.04. Carapace 3.40 long, 2.66 wide. Abdomen 3.64 long, 2.38 wide. Eye sizes and interdistances: AME 0.12, ALE 0.19, PME 0.16, PLE 0.18; AME-AME 0.06, AME-ALE 0.07, AME-PME 0.09, ALE-PLE 0, PME-PME 0.11, PME-PLE 0.15. Leg measurements: I 11.62 (3.37, 3.84, 2.74, 1.67); II 10.19 (3.01, 3.43, 2.12, 1.63); III 9.76 (3.07, 3.08, 2.44, 1.17); IV 12.54 (3.37, 3.92, 3.43, 1.82). Chelicera with 3 promarginal and 2 retromarginal teeth. Epigyne: plate 1.3 times wider than high; epigynal teeth long and wide, bent at tips; originating from anterior margin of the atrium and extending over its posterior margin, about 1/2 the height of plate, with 2.8 times longer than wide; the distance between the base of the tooth is shorter than the distance between the tip of the tooth (2.5 times of their width); epigynal atrium situated at the central part of the plate, longer than wide, with their posterior part more sclerotized than other parts of the plate; copulatory ducts large; spermathecae oval, separated from each other; spermathecal heads long, originating from the middle part of the spermathecae; fertilization duct pointing to the front, hook-shaped at the tips (Fig. 3 A���B). Male. Unknown. Variation. Total length of females 6.09���8.33. Carapace length 2.36���2.82. Distribution. Known only from Shaanxi Province, China (Fig. 8)., Published as part of Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan & Li, Shuqiang, 2019, New species of the coelotine spider genus Lineacoelotes (Araneae: Agelenidae) from China, pp. 351-363 in Zootaxa 4623 (2) on page 354, DOI: 10.11646/zootaxa.4623.2.9, http://zenodo.org/record/3255507, {"references":["Xu, X., Li, S. & Wang, X. (2008) Lineacoelotes, a new genus of Coelotinae from China (Araneae: Amaurobiidae). Zootaxa, 1700, 1 - 20."]}

Li, Bing, Zhao, Zhe, Chen, Haifeng, Wu, Zhiyan, Li, Shuqiang (2019): New species of the coelotine spider genus Lineacoelotes (Araneae: Agelenidae) from China. Zootaxa 4623 (2): 351-363, DOI: https://doi.org/10.11646/zootaxa.4623.2.9

Troglocoelotes nongchiensis Z. Zhao & S. Li sp. n. Figs 7, 14 Type material. Holotype ♀ (IZCAS-Ar34089): Er Cave (N24°05.711’, E108°04.954’, 175 m), Nongchi Village, Du’an County, Hechi City, Guangxi Zhuang Autonomous Region, China, 27. XII. 2013, Y. Luo and local collectors leg. Paratype: 1♀ (IZCAS-Ar34090) same cave, 9. IV. 2018, Z. Chen and J. Liu leg. Etymology. The specific name refers to the type locality, Nongchi Village; adjective. Diagnosis. The female can be distinguished from T. yumiganensis sp. n. by the epigynal teeth located medially vs. anteriorly, and the posterior margin of epigynal atrium triangular vs. belt-shaped. The female is also similar to T. banmenensis sp. n. but can be distinguished from it by length of epigynal teeth longer than width and the posterior margin of epigynal atrium triangular (Fig. 7 A–B). Description. Female (holotype, IZCAS-Ar34089): Total length 5.63. Carapace 2.84 long, 2.01 wide. Abdomen 2.79 long, 1.99 wide. All eyes absent. Leg yellowish. Leg measurements: I: 9.85 (2.81, 3.14, 2.34, 1.56); II: 9.31 (2.59, 2.97, 2.19, 1.56); III: 9.22 (2.50, 2.81, 2.50, 1.41); IV: 11.79 (3.13, 3.59, 3.44, 1.63). Abdomen grey. Epigyne: atrium occupying less than 1/4 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium triangular; teeth 1/3 the atrial width, 3 times longer than width, located medio-laterally, next to atrial lateral margins, spaced of each other by 3 lengths; copulatory ducts coiled one coil around spermathecal anterior parts (Fig. 7 A–B). Male: Unknown. Distribution. China (Guangxi) (Fig. 14).

Troglocoelotes yumiganensis Z. Zhao & S. Li sp. n. Figs 12–14 Type material. Holotype ♂ (IZCAS-Ar34096): Yumigan Cave (N24°38.965’, E105°09.754’, 1549 m), Tianyakou Village, Longlin County, Baise City, Guangxi Zhuang Autonomous Region, China, 19. IV. 2018, Z. Chen and J. Liu leg. Paratypes: 1♀ (IZCAS-Ar34097): same data as in holotype; 1♀ (IZCAS-Ar34098): same cave, 14–15. XII. 2012, Z. Zhao and Z. Chen leg.; 3♀♀ (IZCAS-Ar34099-Ar34101): same cave, 5. XII. 2017, Z. Chen leg. Etymology. The specific name refers to the type locality, Yumigan Cave; adjective. Diagnosis. The male can be distinguished from T. qixianensis sp. n. by having a large lamella on conductor, small RTA, and a black spine next to RTA. From the similar T. tortus by the blunt tip of the patellar apophysis and a short cymbial furrow (1/2 the length of the cymbium vs. 2/3 the length of the cymbium). From the similar T. liangensis sp. n. by the long patellar apophysis and a narrow conductor (Fig. 12 B–C). Females can be distinguished from T. nongchiensis sp. n. by the posterior margin of epigynal atrium belt-shaped vs. triangular (Fig. 13 A–B). Description. Male (holotype, IZCAS-Ar34096): Total length 5.51. Carapace 2.69 long, 1.97 wide. Abdomen 2.82 long, 1.72 wide. Cervical groove and radial furrows pronounced. Eyes reduced, with 6 spots; anterior lateral spots and posterior lateral spots large than other spots. Legs pale. Leg measurements: I: 11.09 (2.97, 3.59, 2.81, 1.72); II: 10.63 (2.81, 3.44, 2.66, 1.72); III: 9.49 (2.51, 2.66, 2.65, 1.67); IV: 13.15 (3.13, 4.22, 3.91, 1.89). Abdomen milky white. Spinnerets milky white. Palp: patellar apophysis shorter than length of patella and with a blunt tip; VTA wide; RTA about 1/3 the length of the cymbium; distal part with black spine next to RTA; cymbial furrow 1/2 the length of the cymbium; embolus beginning at 7 o’clock position; conductor pointed to obliquely up with a large lamella (Fig. 12 A–C). Female (paratype, IZCAS-Ar34097): Total length 6.01. Carapace 3.25 long, 1.76 wide. Abdomen 2.76 long, 2.01 wide. Leg measurements: I: 10.41 (2.96, 3.41, 2.40, 1.64); II: 9.99 (2.81, 3.13, 2.49, 1.56); III: 9.31 (2.49, 2.84, 2.48, 1.50); IV: 12.40 (3.21, 3.79, 3.61, 1.79). Epigyne: atrium located posteriorly, occupying about 1/4 of the epigynal plate; epigynal teeth short, located anteriorly, with the width as its length, spaced of each other by 12 lengths; copulatory duct like 2 folded sacs and copulatory ducts overlapped of each other at their tips; spermathecae brown (Fig. 13 A–B). Distribution. China (Guangxi) (Fig. 14).

Genus Troglocoelotes Z. Zhao & S. Li gen. n. (Figs 1���15) Type species. Troglocoelotes yumiganen sis Z. Zhao & S. Li gen. n. Etymology. The generic name refers to the cave habitat and coelotine spiders. The gender is masculine. Diagnosis. Troglocoelotes is closely related to Guilotes Z. Zhao & S. Li 2018 based on the copulatory organs morphology but can be distinguished by having 3 promarginal and 2 retromarginal teeth (Fig. 1D) vs. with 5���6 promarginal and 5 retromarginal teeth in Guilotes, eyes reduced (Fig. 1A) or absent (Fig. 1B) vs. normal in Guilotes (Fig. 1C); male palp: median apophysis slice-shaped and with a bent tip (Figs 5 B���C, 8B���C, 10B���C, 12B��� C) vs. spoon-shaped, cymbium not suddenly narrowing (Figs 5C, 8C, 10C, 12C), conductor without outgrowth, tegulum and bulbus not intumescent as in Guilotes (Figs 5A, 8A, 10A, 12A); epigyne: epigynal teeth located anteriorly (Fig. 1 E���F) or medially (Figs 7A, 9A) vs. posteriorly, spermathecae ellipsoidal, anterior parts specialized, coiled by copulatory ducts, slender and long (Fig. 1 G���H) vs. spermathecae long and heads fused with copulatory ducts. Description. Small to medium sized with a total length between 3.83���9.28. Coloration: carapace and chelicerae brown; abdomen and spinnerets pale; legs yellowish or pale. Prosoma, posterior radial furrows and both sides of fovea, and chelicerae covered with hairs. Chelicerae with 3 promarginal and 2 retromarginal teeth. Eyes reduced or absent. Legs without black annular striations, leg formula 4123. Abdomen covered with hairs, with cardiac pattern in the most species, without any other marks. Spinnerets: the proximal segment is as long as the basal segment. Male palp: patellar apophysis long in the most species (Figs 8 B���C, 12B���C), but short in T. liangensis (Fig. 5 B���C), wide at base in T. tortus (Fig. 10 B���C); VTA wide and short (the tip just over patellar tibia) (Fig. 5 B���C); RTA about 1/2 the length of VTA (Figs 5C, 8C, 10C, 12C); cymbial furrow in most species long (Figs 5C, 8C, 10C), about 1/2 the length of cymbium in T. yumiganensis (Fig. 12C); embolus filiform, beginning at 6 to 7 o���clock position (Figs 5 A���C, 8A���C, 10A���C, 12A���C); conductor simple but in most species with a blunt dorsal apophysis (Figs 8 B���C, 10B���C) and a lamella (Figs 5B, 10B, 12B); median apophysis slice-shaped with bent tip (Figs 5 B���C, 8B���C, 10B���C, 12B���C). Epigyne: atrium small (Figs 3A, 4A, 6A, 7A, 9A, 11A, 13A); epigynal teeth vary in shape but point medially in the most species (Figs 3A, 4A, 6A, 7A, 9A, 13A); hoods absent; copulatory ducts long and semi-transparent with light color, spermathecae ellipsoidal with dark color, anterior parts specialized, coiled by copulatory ducts, slender and long (Figs 3B, 4B, 6B, 7B, 9B, 11B, 13B). Natural history. All species of Troglocoelotes gen. n. were collected from the deep zone of moist and dark caves (Fig. 15). Characteristics like reduced eyes and pale body coloration indicate they are true troglobionts. Distribution. Guizhou and Guangxi, China (Fig. 14). Comments. The relationships of coelotine spiders were recently analyzed using 8 genes from 286 species in 19 genera (Zhao & Li 2017). The molecular topologies supported Troglocoelotes as a new genus and suggested that Guilotes is the sister-group (Fig. 2)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 220-222, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528, {"references":["Zhao, Z. & Li, S. (2017) Extinction vs. rapid radiation: the juxtaposed evolutionary histories of coelotine spiders support the Eocene-Oligocene orogenesis of the Tibetan Plateau. Systematic Biology, 66 (6), 988 - 1006. https: // doi. org / 10.1093 / sysbio / syx 042"]}

Troglocoelotes yosiianus (Nishikawa, 1999) comb. n. Coelotes yosiianus Nishikawa 1999: 23, figs 1–5 (♀). Draconarius yosiianus (Nishikawa): Wang 2002: 69 (♀); Wang 2003: 556 (♀); Zhu et al. 2017: 415, fig.277 (♀). Diagnosis. Females are similar to T. nongchiensis sp. n. by the epigynal teeth pointed medially and with a tip bent upwards, but can be distinguished from by the copulatory ducts having more coils. Females are also similar to T. tortus by the rectangular posterior margin of epigynal and long copulatory ducts, but can be distinguished by the epigynal teeth pointed medially and the middle parts of copulatory ducts widest. Description. See Nishikawa 1999. Type material. Holotype ♀ (NSMT, not examined): Jiabao Cave (280 m), Xingren Village, Bama County, Guangxi, China, 21. IV. 1998, Y. Nishikawa leg. Paratype: 1♀ (NSMT): same cave, 6. IV. 1997, Y. Nishikawa leg. Material examined. None. Distribution. China (Guangxi) (Fig. 14).

Troglocoelotes qixianensis Z. Zhao & S. Li sp. n. Figs 8���9, 14 Type material. Holotype 6 (IZCAS-Ar34091): Qixian Cave (N25��12.225���, E106��07.085���, 934 m), Wangmo County, Southwest Guizhou Autonomous Prefecture, Guizhou, China, 14. IV. 2018, Z. Chen and J. Liu leg. Paratype: 1♀ (IZCAS-Ar34092): same data as holotype. Etymology. The specific name refers to the type locality, Qixian Cave; adjective. Diagnosis. The male can be distinguished from the similar T. yumiganensis sp. n. by the lack of any lamella on conductor vs. having a lamella and without spine next to RTA. From the similar T. liangensis sp. n. by the patellar apophysis longer than patellar length vs. less than 1/2 the patellar length, tip of the conductor pointed to anteriorly vs. posteriorly. From the similar T. tortus by the tip of the patellar apophysis pointed vs. blunt, and cymbial furrow length 1/2 the length of the cymbium vs. 2/3 the length of the cymbium (Fig. 8 B���C). Females can be distinguished from the similar T. liangensis sp. n. by the posterior margin of epigynal atrium rectangular vs. oval, epigynal teeth located next to atrium lateral margins while in other congeners located above atrium (Fig. 9 A���B). Description. Male (holotype, IZCAS-Ar34091): Total length 3.83. Carapace 1.95 long, 1.13 wide. Abdomen 1.88 long, 1.26 wide. All eyes absent. Leg pale. Leg measurements: I: 8.14 (2.26, 2.74, 1.88, 1.26); II: 7.35 (2.19, 2.34, 1.56, 1.26); III: 7.11 (2.03, 2.03, 1.79, 1.26); IV: 9.09 (2.51, 2.81, 2.51, 1.26). Abdomen and spinnerets brown. Palp: patellar apophysis longer than patellar length, blunt tip; VTA extending beyond tibia, with blunt tip; length of RTA about 1.5 times shorter than VTA; cymbial furrow about 1/2 the length of the cymbium; embolus beginning at 6 o���clock position; conductor ducts 3 times wider than spermathecal width (Fig. 8 A���C). Female (paratype, IZCAS-Ar34092): Total length 5.15. Carapace 2.34 long, 1.41 wide. Abdomen 2.81 long, 1.88 wide. Leg measurements: I: 8.46 (2.59, 2.81, 1.72, 1.34); II: 7.83 (2.50, 2.34, 1.71, 1.28); III: 7.42 (2.19, 2.42, 1.56, 1.25); IV: 9.85 (2.97, 2.97, 2.50, 1.41). Epigyne: atrium occupying less than 1/5 of the epigynal plate, situated at the posteriorly; the posterior margin of epigynal atrium rectangular; teeth 6 times shorter than atrial width, 2 times longer than width, located medio-laterally, spaced of each other by about 5 lengths; the widest parts of copulatory ducts 3 times wider than spermathecal width (Fig. 9 A���B). Distribution. China (Guizhou) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 229-230, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Troglocoelotes yumiganensis Z. Zhao & S. Li sp. n. Figs 12���14 Type material. Holotype ♂ (IZCAS-Ar34096): Yumigan Cave (N24��38.965���, E105��09.754���, 1549 m), Tianyakou Village, Longlin County, Baise City, Guangxi Zhuang Autonomous Region, China, 19. IV. 2018, Z. Chen and J. Liu leg. Paratypes: 1♀ (IZCAS-Ar34097): same data as in holotype; 1♀ (IZCAS-Ar34098): same cave, 14���15. XII. 2012, Z. Zhao and Z. Chen leg.; 3♀♀ (IZCAS-Ar34099-Ar34101): same cave, 5. XII. 2017, Z. Chen leg. Etymology. The specific name refers to the type locality, Yumigan Cave; adjective. Diagnosis. The male can be distinguished from T. qixianensis sp. n. by having a large lamella on conductor, small RTA, and a black spine next to RTA. From the similar T. tortus by the blunt tip of the patellar apophysis and a short cymbial furrow (1/2 the length of the cymbium vs. 2/3 the length of the cymbium). From the similar T. liangensis sp. n. by the long patellar apophysis and a narrow conductor (Fig. 12 B���C). Females can be distinguished from T. nongchiensis sp. n. by the posterior margin of epigynal atrium belt-shaped vs. triangular (Fig. 13 A���B). Description. Male (holotype, IZCAS-Ar34096): Total length 5.51. Carapace 2.69 long, 1.97 wide. Abdomen 2.82 long, 1.72 wide. Cervical groove and radial furrows pronounced. Eyes reduced, with 6 spots; anterior lateral spots and posterior lateral spots large than other spots. Legs pale. Leg measurements: I: 11.09 (2.97, 3.59, 2.81, 1.72); II: 10.63 (2.81, 3.44, 2.66, 1.72); III: 9.49 (2.51, 2.66, 2.65, 1.67); IV: 13.15 (3.13, 4.22, 3.91, 1.89). Abdomen milky white. Spinnerets milky white. Palp: patellar apophysis shorter than length of patella and with a blunt tip; VTA wide; RTA about 1/3 the length of the cymbium; distal part with black spine next to RTA; cymbial furrow 1/2 the length of the cymbium; embolus beginning at 7 o���clock position; conductor pointed to obliquely up with a large lamella (Fig. 12 A���C). Female (paratype, IZCAS-Ar34097): Total length 6.01. Carapace 3.25 long, 1.76 wide. Abdomen 2.76 long, 2.01 wide. Leg measurements: I: 10.41 (2.96, 3.41, 2.40, 1.64); II: 9.99 (2.81, 3.13, 2.49, 1.56); III: 9.31 (2.49, 2.84, 2.48, 1.50); IV: 12.40 (3.21, 3.79, 3.61, 1.79). Epigyne: atrium located posteriorly, occupying about 1/4 of the epigynal plate; epigynal teeth short, located anteriorly, with the width as its length, spaced of each other by 12 lengths; copulatory duct like 2 folded sacs and copulatory ducts overlapped of each other at their tips; spermathecae brown (Fig. 13 A���B). Distribution. China (Guangxi) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 234-237, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Troglocoelotes yosiianus (Nishikawa, 1999) comb. n. Coelotes yosiianus Nishikawa 1999: 23, figs 1���5 (♀). Draconarius yosiianus (Nishikawa): Wang 2002: 69 (♀); Wang 2003: 556 (♀); Zhu et al. 2017: 415, fig.277 (♀). Diagnosis. Females are similar to T. nongchiensis sp. n. by the epigynal teeth pointed medially and with a tip bent upwards, but can be distinguished from by the copulatory ducts having more coils. Females are also similar to T. tortus by the rectangular posterior margin of epigynal and long copulatory ducts, but can be distinguished by the epigynal teeth pointed medially and the middle parts of copulatory ducts widest. Description. See Nishikawa 1999. Type material. Holotype ♀ (NSMT, not examined): Jiabao Cave (280 m), Xingren Village, Bama County, Guangxi, China, 21. IV. 1998, Y. Nishikawa leg. Paratype: 1♀ (NSMT): same cave, 6. IV. 1997, Y. Nishikawa leg. Material examined. None. Distribution. China (Guangxi) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on page 232, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528, {"references":["Nishikawa, Y. (1999) A new eyeless agelenid spider from a limestone cave in Guangxi, south China. Journal of the Speleological Society of Japan, 24, 23 - 26.","Wang, X. P. (2002) A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae). Bulletin of the American Museum of Natural History, 269, 1 - 150. https: // doi. org / 10.1206 / 0003 - 0090 (2002) 269 2.0. CO; 2","Wang, X. P. (2003) Species revision of the coelotine spider genera Bifidocoelotes, Coronilla, Draconarius, Femoracoelotes, Leptocoelotes, Longicoelotes, Platocoelotes, Spiricoelotes, Tegecoelotes, and Tonsilla (Araneae: Amaurobiidae). Proceedings of the California Academy of Sciences, 54, 499 - 662.","Zhu, M. S., Wang, X. P. & Zhang, Z. S. (2017) Fauna Sinica: Invertebrata Uol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp."]}

Troglocoelotes banmenensis Z. Zhao & S. Li sp. n. Figs 4, 14 Type material. Holotype ♀ (IZCAS-Ar34081): San Cave (N25��43.002���, E107��55.796���, 505 m), Banmen Village, Sandu County, Qianna City, Guizhou, China, 20. III. 2013, H. Zhao and J. Liu leg. Paratype: 1♀ (IZCAS- Ar34082): same cave, 11. IV. 2018, Z. Chen and J. Liu leg. Etymology. The specific name refers to the type locality, Banmen Village; adjective. Diagnosis. The specimens of this species can be distinguished from the similar species T. yumiganensis sp. n. and T. nongchiensis sp. n. by the epigynal atrium occupying about 1/8 epigynal plate vs. 1/4 of the epigynal plate, the posterior margin of epigynal atrium trapezoidal while triangular in T. yumiganensis and belt-shaped in T. nongchiensis, epigynal teeth circular vs. finger-shaped in T. yumiganensis and T. nongchiensis (Fig. 4 A���B). Description. Female (holotype, IZCAS-Ar34081): Total length 6.67. Carapace 3.46 long, 1.93 wide. Abdomen 3.21 long, 1.28 wide. Eyes reduced, with 6 white spots. Leg pale. Leg measurements: I: 13.21 (3.21, 4.81, 3.14, 2.05); II: 12.32 (3.53, 3.85, 2.89, 2.05); III: 11.87 (3.21, 3.53, 3.21, 1.92); IV: 15.39 (3.85, 4.81, 4.81, 1.92). Abdomen with cardiac pattern. Epigyne: atrium oval, occupying less than 1/8 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium rectangular; teeth oval, not much longer than width, spaced of each other by 2 lengths; copulatory ducts wide, coiled anterior parts spermathecae 3 coils (Fig. 4 A���B). Male: Unknown. Distribution. China (Guizhou) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on page 223, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Troglocoelotes tortus (Chen, Zhu & Kim, 2008) comb. n. Figs 10���11, 14 Draconarius tortus Chen, Zhu & Kim 2008: 86, figs 2���5 (♀); Jiang et al. 2017: figs 1���15 (�� ♀); Zhu et al, 2017: 393, fig. 257 (♀). Diagnosis. Males can be distinguished from the similar T. qixianensis sp. n. by the thin RTA, conductor pointed posteriorly with a lamella and a strong dorsal apophysis. From the similar T. liangensis sp. n. by the patellar apophysis with a pointed tip, narrow embolus, and the long lamella of the conductor. From the similar T. yumiganensis sp. n. by the lack of a spine next to the RTA, patellar apophysis with a pointed tip, and the cymbial furrow over 2/3 the length of the cymbium (Fig. 10 B���C). Females can be distinguished from the similar T. yosiianus sp. n. by the wide and truncated epigynal teeth which are pointed anteriorly and spermathecae overlapped at their tip. From the similar T. liangensis sp. n. by copulatory ducts thin vs. wide (Fig. 11 A���B). Description. Cervical groove and radial furrows dark. All eyes absent. Leg yellowish. Abdomen grey. Male palp: patellar apophysis with pointed tip; VTA triangular; length of RTA about 1/3 the length of VTA; cymbial furrow about 2/3 the length of the cymbium; embolus beginning at 5 o���clock position; conductor with a blunt dorsal apophysis and a large lamella (Fig. 10 A���C). Epigyne: atrium 2 times wider than long, located at posteriorly; teeth short, 4 times wider than length, spaced of each other by 3 widths; copulatory ducts convoluted with 7 coils; spermathecae spaced by 2 widths (Fig. 11 A���B). Type material. Holotype ♀ (MHBU, not examined): Feilong Cave (1335 m), Shanjiao county, Xingyi City, Guizhou, China, 25 May 2004, H. Chen leg. Material examined. 1♀ (IZCAS-Ar34093): same cave (N24��55.000���, E104��53.000���, 1335 m), 8. III. 2011, C. Wang and Y. Lin leg.; 1 6 5♀♀ (IZCAS-Ar34094): same cave (N24��58.231���, E104��52.900���, 1382 �� 3 m), 15. IV. 2018, Z. Chen and J. Liu leg.; 2♀♀ (IZCAS-Ar34095): same cave (N24��58.231���, E104��52.900���, 1382 �� 3 m), 1. XII. 2017, Z. Chen leg. Distribution. China (Guizhou) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 230-232, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528, {"references":["Chen, H. M., Zhu, M. S. & Kim, J. P. (2008) Three new eyeless Draconarius spiders (Araneae, Amaurobiidae) from limestone caves in Guizhou, southwestern China. Korean Arachnology, 24, 85 - 95.","Jiang, X., Guo, X., Yu, Z. & Chen, H. (2017) First description of the male of the eyeless spider Draconarius tortus Chen, Zhu & Kim 2008 from China (Araneae: Agelenidae: Coelotinae). Zootaxa, 4294 (1), 141 - 144. https: // doi. org / 10.11646 / zootaxa. 4294.1.11","Zhu, M. S., Wang, X. P. & Zhang, Z. S. (2017) Fauna Sinica: Invertebrata Uol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp."]}

Troglocoelotes liangensis Z. Zhao & S. Li sp. n. Figs 5���6, 14 Type material. Holotype 6 (IZCAS-Ar34083): Liang Cave (N23��40.011���, E106��33.111���, 464 m), Tianyang County, Baise City, Guangxi Zhuang Autonomous Region, China, 12. XII. 2015, X. Zhang and Z. Chen leg. Paratypes: 2♀♀ (IZCAS-Ar34084-Ar34085): same cave (N23��40.118���, E106��33.957���, 464 m), 18. XII. 2012, Z. Zhao and Z. Chen leg; 3♀♀ (IZCAS-Ar34086-Ar34088): same cave (N23��39.954���, E106��34.166���, 473 �� 4 m), 7. XII. 2017, Z. Chen leg. Etymology. The specific name refers to the type locality, Liang Cave; adjective. Diagnosis. The male can be distinguished from T. qixianensis sp. n. by conductor with lamella vs. no lamella. From the similar T. tortus by the blunt tip of the patellar apophysis, wider embolus vs. filariform, and smaller median apophysis. From the similar T. yumiganensis sp. n. by the long cymbial furrow and no spine next to the RTA (Fig. 5C). Females from the similar T. tortus by epigynal atrium lip-shaped, epigynal teeth pointed medially vs. anteriorly, and copulatory ducts 2 times wider than in T. tortus (Fig. 6 A���B). Description. Male (holotype, IZCAS-Ar34083): Total length 4.63. Carapace 2.42 long, 1.56 wide. Abdomen 2.21 long, 1.53 wide. Eyes reduced, with 2 spots. Leg pale. Leg measurements: I: 10.00 (2.81, 3.13, 2.34, 1.72); II: 9.12 (2.56, 2.81, 2.19, 1.56); III: 8.45 (2.34, 2.51, 2.19, 1.41); IV: 11.25 (2.97, 3.36, 3.20, 1.72). Abdomen grey. Palp: patellar apophysis shorter than width of patella, blunt tip; VTA wide; RTA small, width about 1/3 the width of VTA; cymbial furrow more than 2/3 the length of the cymbium; embolus beginning at 6 o���clock position; conductor simple with a wide dorsal apophysis and a small lamella (Fig. 5 A���C). Female (paratype, IZCAS-Ar34084): Total length 4.29. Carapace 2.08 long, 1.41 wide. Abdomen 2.21 long, 1.39 wide. Chelicerae white. Leg measurements: I: 8.02 (2.34, 2.51, 2.03, 1.14); II: 7.90 (2.19, 2.51, 1.88, 1.32); III: 7.35 (2.03, 2.19, 1.88, 1.25); IV: 10.15 (2.56, 3.06, 2.97, 1.56). Epigyne: atrium occupying less than 1/4 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium oval; teeth small, width 2 times wider than length, located anteriorly, spaced of each other by more than 7 widths; copulatory ducts wider than spermathecal width (Fig. 6 A���B). Distribution. China (Guangxi) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 223-226, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Troglocoelotes proximus (Chen, Zhu & Kim, 2008) comb. n. Draconarius proximus Chen, Zhu & Kim 2008: 87, figs 6���9 (♀); Zhu et al. 2017: 348, fig. 216 (♀). Diagnosis. This species can be distinguished from the similar T. yosiianus by the atrium occupying 1/10 of the epigynal plate and the spermathecal anterior parts close together vs. overlapped each other. From similar T. bailongensis sp. n. by epigynal teeth pointed posteriorly vs. medially. Description. See Chen et al. 2008. Type material. Holotype ♀ (MHBU, not examined): Dongge Cave (N25��19.000���, E108��01.000���, 680 m), Laoyang Village, Libo County, Qiannan City, Guizhou, China, 16. X. 2000, H. Chen leg. Paratype: 1♀ (MHBU), Shuangshi Cave (716 m), Bale Village, Libo County, Qiannan City, Guizhou, China, 8. X. 2001, H. Chen leg. Material examined. None. Distribution. China (Guizhou) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on page 229, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528, {"references":["Chen, H. M., Zhu, M. S. & Kim, J. P. (2008) Three new eyeless Draconarius spiders (Araneae, Amaurobiidae) from limestone caves in Guizhou, southwestern China. Korean Arachnology, 24, 85 - 95.","Zhu, M. S., Wang, X. P. & Zhang, Z. S. (2017) Fauna Sinica: Invertebrata Uol. 59: Arachnida: Araneae: Agelenidae and Amaurobiidae. Science Press, Beijing, 727 pp."]}

Troglocoelotes tortus (Chen, Zhu & Kim, 2008) comb. n. Figs 10–11, 14 Draconarius tortus Chen, Zhu & Kim 2008: 86, figs 2–5 (♀); Jiang et al. 2017: figs 1–15 (Ƌ ♀); Zhu et al, 2017: 393, fig. 257 (♀). Diagnosis. Males can be distinguished from the similar T. qixianensis sp. n. by the thin RTA, conductor pointed posteriorly with a lamella and a strong dorsal apophysis. From the similar T. liangensis sp. n. by the patellar apophysis with a pointed tip, narrow embolus, and the long lamella of the conductor. From the similar T. yumiganensis sp. n. by the lack of a spine next to the RTA, patellar apophysis with a pointed tip, and the cymbial furrow over 2/3 the length of the cymbium (Fig. 10 B–C). Females can be distinguished from the similar T. yosiianus sp. n. by the wide and truncated epigynal teeth which are pointed anteriorly and spermathecae overlapped at their tip. From the similar T. liangensis sp. n. by copulatory ducts thin vs. wide (Fig. 11 A–B). Description. Cervical groove and radial furrows dark. All eyes absent. Leg yellowish. Abdomen grey. Male palp: patellar apophysis with pointed tip; VTA triangular; length of RTA about 1/3 the length of VTA; cymbial furrow about 2/3 the length of the cymbium; embolus beginning at 5 o’clock position; conductor with a blunt dorsal apophysis and a large lamella (Fig. 10 A–C). Epigyne: atrium 2 times wider than long, located at posteriorly; teeth short, 4 times wider than length, spaced of each other by 3 widths; copulatory ducts convoluted with 7 coils; spermathecae spaced by 2 widths (Fig. 11 A–B). Type material. Holotype ♀ (MHBU, not examined): Feilong Cave (1335 m), Shanjiao county, Xingyi City, Guizhou, China, 25 May 2004, H. Chen leg. Material examined. 1♀ (IZCAS-Ar34093): same cave (N24°55.000’, E104°53.000’, 1335 m), 8. III. 2011, C. Wang and Y. Lin leg.; 1 6 5♀♀ (IZCAS-Ar34094): same cave (N24°58.231’, E104°52.900’, 1382 ± 3 m), 15. IV. 2018, Z. Chen and J. Liu leg.; 2♀♀ (IZCAS-Ar34095): same cave (N24°58.231’, E104°52.900’, 1382 ± 3 m), 1. XII. 2017, Z. Chen leg. Distribution. China (Guizhou) (Fig. 14).

Troglocoelotes banmenensis Z. Zhao & S. Li sp. n. Figs 4, 14 Type material. Holotype ♀ (IZCAS-Ar34081): San Cave (N25°43.002’, E107°55.796’, 505 m), Banmen Village, Sandu County, Qianna City, Guizhou, China, 20. III. 2013, H. Zhao and J. Liu leg. Paratype: 1♀ (IZCAS- Ar34082): same cave, 11. IV. 2018, Z. Chen and J. Liu leg. Etymology. The specific name refers to the type locality, Banmen Village; adjective. Diagnosis. The specimens of this species can be distinguished from the similar species T. yumiganensis sp. n. and T. nongchiensis sp. n. by the epigynal atrium occupying about 1/8 epigynal plate vs. 1/4 of the epigynal plate, the posterior margin of epigynal atrium trapezoidal while triangular in T. yumiganensis and belt-shaped in T. nongchiensis, epigynal teeth circular vs. finger-shaped in T. yumiganensis and T. nongchiensis (Fig. 4 A–B). Description. Female (holotype, IZCAS-Ar34081): Total length 6.67. Carapace 3.46 long, 1.93 wide. Abdomen 3.21 long, 1.28 wide. Eyes reduced, with 6 white spots. Leg pale. Leg measurements: I: 13.21 (3.21, 4.81, 3.14, 2.05); II: 12.32 (3.53, 3.85, 2.89, 2.05); III: 11.87 (3.21, 3.53, 3.21, 1.92); IV: 15.39 (3.85, 4.81, 4.81, 1.92). Abdomen with cardiac pattern. Epigyne: atrium oval, occupying less than 1/8 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium rectangular; teeth oval, not much longer than width, spaced of each other by 2 lengths; copulatory ducts wide, coiled anterior parts spermathecae 3 coils (Fig. 4 A–B). Male: Unknown. Distribution. China (Guizhou) (Fig. 14).

Troglocoelotes liangensis Z. Zhao & S. Li sp. n. Figs 5–6, 14 Type material. Holotype 6 (IZCAS-Ar34083): Liang Cave (N23°40.011’, E106°33.111’, 464 m), Tianyang County, Baise City, Guangxi Zhuang Autonomous Region, China, 12. XII. 2015, X. Zhang and Z. Chen leg. Paratypes: 2♀♀ (IZCAS-Ar34084-Ar34085): same cave (N23°40.118’, E106°33.957’, 464 m), 18. XII. 2012, Z. Zhao and Z. Chen leg; 3♀♀ (IZCAS-Ar34086-Ar34088): same cave (N23°39.954’, E106°34.166’, 473 ± 4 m), 7. XII. 2017, Z. Chen leg. Etymology. The specific name refers to the type locality, Liang Cave; adjective. Diagnosis. The male can be distinguished from T. qixianensis sp. n. by conductor with lamella vs. no lamella. From the similar T. tortus by the blunt tip of the patellar apophysis, wider embolus vs. filariform, and smaller median apophysis. From the similar T. yumiganensis sp. n. by the long cymbial furrow and no spine next to the RTA (Fig. 5C). Females from the similar T. tortus by epigynal atrium lip-shaped, epigynal teeth pointed medially vs. anteriorly, and copulatory ducts 2 times wider than in T. tortus (Fig. 6 A–B). Description. Male (holotype, IZCAS-Ar34083): Total length 4.63. Carapace 2.42 long, 1.56 wide. Abdomen 2.21 long, 1.53 wide. Eyes reduced, with 2 spots. Leg pale. Leg measurements: I: 10.00 (2.81, 3.13, 2.34, 1.72); II: 9.12 (2.56, 2.81, 2.19, 1.56); III: 8.45 (2.34, 2.51, 2.19, 1.41); IV: 11.25 (2.97, 3.36, 3.20, 1.72). Abdomen grey. Palp: patellar apophysis shorter than width of patella, blunt tip; VTA wide; RTA small, width about 1/3 the width of VTA; cymbial furrow more than 2/3 the length of the cymbium; embolus beginning at 6 o’clock position; conductor simple with a wide dorsal apophysis and a small lamella (Fig. 5 A–C). Female (paratype, IZCAS-Ar34084): Total length 4.29. Carapace 2.08 long, 1.41 wide. Abdomen 2.21 long, 1.39 wide. Chelicerae white. Leg measurements: I: 8.02 (2.34, 2.51, 2.03, 1.14); II: 7.90 (2.19, 2.51, 1.88, 1.32); III: 7.35 (2.03, 2.19, 1.88, 1.25); IV: 10.15 (2.56, 3.06, 2.97, 1.56). Epigyne: atrium occupying less than 1/4 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium oval; teeth small, width 2 times wider than length, located anteriorly, spaced of each other by more than 7 widths; copulatory ducts wider than spermathecal width (Fig. 6 A–B). Distribution. China (Guangxi) (Fig. 14).

Troglocoelotes bailongensis Z. Zhao & S. Li sp. n. Figs 3, 14 Type material. Holotype ♀ (IZCAS-Ar34080): Bailong Cave (N24��30.322���, E108��39.888���, 221 m), Yizhou County, Hechi City, Guangxi Zhuang Autonomous Region, China, 13. X. 2010, C. Wang and Y. Lin leg. Etymology. The specific name refers to the type locality, Bailong Cave; adjective. Diagnosis. The holotype can be distinguished from T. proximus by epigynal teeth pointed medially vs. posteriorly, and the posterior margin of epigynal atrium rectangular vs. lip-shaped (Fig. 3 A���B). Description. Female (holotype, IZCAS-Ar34080): Total length 6.56. Carapace 2.97 long, 2.01 wide. Abdomen 3.59 long, 2.40 wide. Eyes reduced, with 2 small spots. Leg yellowish without annular striations. Legs yellowish. Leg measurements: I: 11.60 (3.21, 3.65, 2.82, 1.92); II: 10.77 (2.95, 3.53, 2.56, 1.73); III: 9.93 (2.88, 2.63, 2.82, 1.60); IV: 13.59 (3.53, 4.17, 3.97, 1.92). Epigyne: atrium occupying about 1/6 of the epigynal plate and situated the central part of epigyne; the rectangular posterior margin of epigynal atrium; epigynal teeth about 1/4 the plate length, 4 times longer than width, spaced of each other by more than 2 lengths, finger-shaped and located anteriorly (Fig. 3 A���B). Male: Unknown. Distribution. China (Guangxi) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 222-223, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Troglocoelotes proximus (Chen, Zhu & Kim, 2008) comb. n. Draconarius proximus Chen, Zhu & Kim 2008: 87, figs 6–9 (♀); Zhu et al. 2017: 348, fig. 216 (♀). Diagnosis. This species can be distinguished from the similar T. yosiianus by the atrium occupying 1/10 of the epigynal plate and the spermathecal anterior parts close together vs. overlapped each other. From similar T. bailongensis sp. n. by epigynal teeth pointed posteriorly vs. medially. Description. See Chen et al. 2008. Type material. Holotype ♀ (MHBU, not examined): Dongge Cave (N25°19.000’, E108°01.000’, 680 m), Laoyang Village, Libo County, Qiannan City, Guizhou, China, 16. X. 2000, H. Chen leg. Paratype: 1♀ (MHBU), Shuangshi Cave (716 m), Bale Village, Libo County, Qiannan City, Guizhou, China, 8. X. 2001, H. Chen leg. Material examined. None. Distribution. China (Guizhou) (Fig. 14).

Troglocoelotes qixianensis Z. Zhao & S. Li sp. n. Figs 8–9, 14 Type material. Holotype 6 (IZCAS-Ar34091): Qixian Cave (N25°12.225’, E106°07.085’, 934 m), Wangmo County, Southwest Guizhou Autonomous Prefecture, Guizhou, China, 14. IV. 2018, Z. Chen and J. Liu leg. Paratype: 1♀ (IZCAS-Ar34092): same data as holotype. Etymology. The specific name refers to the type locality, Qixian Cave; adjective. Diagnosis. The male can be distinguished from the similar T. yumiganensis sp. n. by the lack of any lamella on conductor vs. having a lamella and without spine next to RTA. From the similar T. liangensis sp. n. by the patellar apophysis longer than patellar length vs. less than 1/2 the patellar length, tip of the conductor pointed to anteriorly vs. posteriorly. From the similar T. tortus by the tip of the patellar apophysis pointed vs. blunt, and cymbial furrow length 1/2 the length of the cymbium vs. 2/3 the length of the cymbium (Fig. 8 B–C). Females can be distinguished from the similar T. liangensis sp. n. by the posterior margin of epigynal atrium rectangular vs. oval, epigynal teeth located next to atrium lateral margins while in other congeners located above atrium (Fig. 9 A–B). Description. Male (holotype, IZCAS-Ar34091): Total length 3.83. Carapace 1.95 long, 1.13 wide. Abdomen 1.88 long, 1.26 wide. All eyes absent. Leg pale. Leg measurements: I: 8.14 (2.26, 2.74, 1.88, 1.26); II: 7.35 (2.19, 2.34, 1.56, 1.26); III: 7.11 (2.03, 2.03, 1.79, 1.26); IV: 9.09 (2.51, 2.81, 2.51, 1.26). Abdomen and spinnerets brown. Palp: patellar apophysis longer than patellar length, blunt tip; VTA extending beyond tibia, with blunt tip; length of RTA about 1.5 times shorter than VTA; cymbial furrow about 1/2 the length of the cymbium; embolus beginning at 6 o’clock position; conductor ducts 3 times wider than spermathecal width (Fig. 8 A–C). Female (paratype, IZCAS-Ar34092): Total length 5.15. Carapace 2.34 long, 1.41 wide. Abdomen 2.81 long, 1.88 wide. Leg measurements: I: 8.46 (2.59, 2.81, 1.72, 1.34); II: 7.83 (2.50, 2.34, 1.71, 1.28); III: 7.42 (2.19, 2.42, 1.56, 1.25); IV: 9.85 (2.97, 2.97, 2.50, 1.41). Epigyne: atrium occupying less than 1/5 of the epigynal plate, situated at the posteriorly; the posterior margin of epigynal atrium rectangular; teeth 6 times shorter than atrial width, 2 times longer than width, located medio-laterally, spaced of each other by about 5 lengths; the widest parts of copulatory ducts 3 times wider than spermathecal width (Fig. 9 A–B). Distribution. China (Guizhou) (Fig. 14).

Troglocoelotes bailongensis Z. Zhao & S. Li sp. n. Figs 3, 14 Type material. Holotype ♀ (IZCAS-Ar34080): Bailong Cave (N24°30.322’, E108°39.888’, 221 m), Yizhou County, Hechi City, Guangxi Zhuang Autonomous Region, China, 13. X. 2010, C. Wang and Y. Lin leg. Etymology. The specific name refers to the type locality, Bailong Cave; adjective. Diagnosis. The holotype can be distinguished from T. proximus by epigynal teeth pointed medially vs. posteriorly, and the posterior margin of epigynal atrium rectangular vs. lip-shaped (Fig. 3 A–B). Description. Female (holotype, IZCAS-Ar34080): Total length 6.56. Carapace 2.97 long, 2.01 wide. Abdomen 3.59 long, 2.40 wide. Eyes reduced, with 2 small spots. Leg yellowish without annular striations. Legs yellowish. Leg measurements: I: 11.60 (3.21, 3.65, 2.82, 1.92); II: 10.77 (2.95, 3.53, 2.56, 1.73); III: 9.93 (2.88, 2.63, 2.82, 1.60); IV: 13.59 (3.53, 4.17, 3.97, 1.92). Epigyne: atrium occupying about 1/6 of the epigynal plate and situated the central part of epigyne; the rectangular posterior margin of epigynal atrium; epigynal teeth about 1/4 the plate length, 4 times longer than width, spaced of each other by more than 2 lengths, finger-shaped and located anteriorly (Fig. 3 A–B). Male: Unknown. Distribution. China (Guangxi) (Fig. 14).

Troglocoelotes nongchiensis Z. Zhao & S. Li sp. n. Figs 7, 14 Type material. Holotype ♀ (IZCAS-Ar34089): Er Cave (N24��05.711���, E108��04.954���, 175 m), Nongchi Village, Du���an County, Hechi City, Guangxi Zhuang Autonomous Region, China, 27. XII. 2013, Y. Luo and local collectors leg. Paratype: 1♀ (IZCAS-Ar34090) same cave, 9. IV. 2018, Z. Chen and J. Liu leg. Etymology. The specific name refers to the type locality, Nongchi Village; adjective. Diagnosis. The female can be distinguished from T. yumiganensis sp. n. by the epigynal teeth located medially vs. anteriorly, and the posterior margin of epigynal atrium triangular vs. belt-shaped. The female is also similar to T. banmenensis sp. n. but can be distinguished from it by length of epigynal teeth longer than width and the posterior margin of epigynal atrium triangular (Fig. 7 A���B). Description. Female (holotype, IZCAS-Ar34089): Total length 5.63. Carapace 2.84 long, 2.01 wide. Abdomen 2.79 long, 1.99 wide. All eyes absent. Leg yellowish. Leg measurements: I: 9.85 (2.81, 3.14, 2.34, 1.56); II: 9.31 (2.59, 2.97, 2.19, 1.56); III: 9.22 (2.50, 2.81, 2.50, 1.41); IV: 11.79 (3.13, 3.59, 3.44, 1.63). Abdomen grey. Epigyne: atrium occupying less than 1/4 of the epigynal plate, situated at the central part of epigyne; the posterior margin of epigynal atrium triangular; teeth 1/3 the atrial width, 3 times longer than width, located medio-laterally, next to atrial lateral margins, spaced of each other by 3 lengths; copulatory ducts coiled one coil around spermathecal anterior parts (Fig. 7 A���B). Male: Unknown. Distribution. China (Guangxi) (Fig. 14)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2019, Troglocoelotes gen. n., a new genus of Coelotinae spiders (Araneae, Agelenidae) from caves in South China, pp. 219-238 in Zootaxa 4554 (1) on pages 226-229, DOI: 10.11646/zootaxa.4554.1.7, http://zenodo.org/record/2623528

Nuconarius pseudocapitulatus (Wang, 2003) comb. n. FIGS 5–7 Draconarius pseudocapitulatus WaNG 2003: 545, fIGS 53A–B; WaNG et al. 2010: 89, fIGS 407–427; Zhu et al. 2017: 352, fIGS 219A–D. Types. Holotype: ♀ (HNNU), DANZHU He DRAINAGe, (27.631000°N, 98.621000°E, eLeV. 2700 M), 13.5 AIR KM SSW OF GONGSHAN, GONGSHAN CO., NUjIANG, Yunnan, China, 30 JUNe–5 JULY 2000, D. KAVANAUGH, C.E. GRISWOLD, H.B. LIANG, D. UBICK, H.M. YAN, D.Z. DONG LeG. Paratypes: 1♀ (HNNU), same data as holotype; 1♀ (CAS), Nujiang STATe NATURe ReSeRVe, NO. 12 BRIDGe CAMP AReA (27.715000°N, 98.502000°E, eLeV. 2775M) 16.3 AIR KM W OF GONGSHAN, NUjIANG PReFeCTURe, GAOLIGONG SHAN, Yunnan, China, 15–19 JULY 2000, H.M. YAN, D. KAVANAUGH, C.E. GRISWOLD, H.B. LIANG, D. UBICK, D.Z. DONG LeG. Other records: 37♀♀ 9♂♂ (HNNU AND CAS), GAOLIGONG SHAN, Yunnan, China. Material examined. 1♀ 1♂ (IZCAS-AR 33971), DABADI (27.786267°N, 98.510283°E, eLeV. 3200 M), 41 KM OF DeRUNG-NU AUTONOMOUS COUNTY GONGSHAN, NUjIANG OF THe LISU AUTONOMOUS PReFeCTURe, Yunnan, China, 11 DeCeMBeR 2013, Y. LI & J. LIU LeG. Diagnosis. MALeS OF THIS SPeCIeS CAN Be DISTINGUISHeD FROM N. brevipatellatus Z. ZhaO & S. LI sp. n. BY THe LONG PATeLLAR APOPHYSIS, SUBeqUAL TO THe LeNGTH OF PATeLLA, WHILe IT IS OBVIOUS SHORT IN N. brevipatellatus Z. ZhaO & S. LI sp. n.; and cymbial furrow short, less than ½ of the length of cymbium, while it is about ⅓ in N. brevipatellatus Z. ZhaO & S. LI sp. n. (FIGS 1C, 5C), AND MeDIAN APOPHYSIS ABSeNT (FIGS 1B–C, 5B–C). FeMALeS OF THe SPeCIeS CAN Be DISTINGUISHeD FROM N. brevipatellatus Z. ZhaO & S. LI sp. n. BY THe SPeRMATHeCAL HeADS WHOSe WIDTH eqUAL TO THe LeNGTH WHILe IN N. brevipatellatus Z. ZhaO & S. LI sp. n., THe WIDTH OF SPeRMATHeCAL HeADS IS LONGeR THAN THe LeNGTH (FIGS 2B, 6B); THe POSTeRIOR ePIGYNAL SCLeRITe SeMICIRCULAR AND UNCOVeReD WHILe IN N. brevipatellatus Z. ZhaO & S. LI sp. n., THe POSTeRIOR ePIGYNAL SCLeRITe COVeReD BY THe exTeNDeD POSTeRIOR eDGe OF ATRIUM (FIGS 2A, 6A). Description. Male: See WANG et al. (2010). PHOTOS OF BOTH HABITUS AND PALP ARe PROVIDeD HeRe (FIGS 5–6). Female: See WANG (2003). PHOTOS OF BOTH HABITUS AND ePIGYNe ARe PROVIDeD HeRe (FIG. 6). Variation: TOTAL LeNGTH VARIeS FROM 8.70 TO 10.80. Distribution. YUNNAN, CHINA (FIG. 7).

Hengconarius longpuensis Z. Zhao & S. Li sp. n. FIGS 17–18, 20 Type material. Holotype: ♂ (IZCAS-AR33980), LONGPU VILLAGe (28.53412°N, 98.451050°E, eLeV. 2873 M), CAWARONG TOWNSHIP, ZAYü, Tibet, China, 14 SePTeMBeR 2014, J. LIU LeG. Paratypes: 1♀ 1♂ (IZCAS-AR33981, 33982), SAMe DATA AS HOLOTYPe. Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe PINYIN "LONGPU" AND ReFeRS TO ITS TYPe LOCALITY, LONGPU VILLAGe; ADjeCTIVe. Diagnosis. THe MALeS ARe SIMILAR TO H. exilis BUT CAN eASILY Be DISTINGUISHeD FROM H. exilis BY THe PALP SHORTeR THAN THe TARSUS; THe TARSUS AS LONG AS THe TIBIA (TARSUS AS TWICe LONG AS THe TIBIA IN H. exilis), TWO BRANCHeS OF CONDUCTOR THAT DIFFeR IN LeNGTH BUT IN H. exilis, TWO BRANCHeS BeING OF THe SAMe LeNGTH (FIGS 9B, 17B); MeDIAN APOPHYSIS LONGeR THAN IT IN H. exilis (FIGS 9C, 17C). THe FeMALeS CAN Be eASILY DISTINGUISHeD FROM H. exilis BY POSTeRIOR ePIGYNAL SCLeRITe BROAD, THe HeIGHT OF IT IS ABOUT ½ THe HeIGHT OF ePIGYNe BUT IS TWICe AS ITS IN H. exilis (FIGS 10A, 18A); THe DISTANCe BeTWeeN SPeRMATHeCAe IS TWICe AS LONG AS THe DIAMeTeR OF SPeRMATHeCA WHILe IN H. exilis, THe DISTANCe IS SUBeqUAL TO THe DIAMeTeR (FIGS 10B, 18B). Description. Male (holotype): TOTAL LeNGTH 8.34. CARAPACe 3.85 LONG, 2.81 WIDe. ABDOMeN 4.49 LONG, 2.97 WIDe. EYe SIZeS AND INTeRDISTANCeS: AME 0.12, ALE 0.22, PME 0.14, PLE 0.16; AME–AME 0.08, AME–ALE 0.06, AME–PME 0.12, ALE–PLE 0, PME–PME 0.12, PME–PLE 0.13. LeG MeASUReMeNTS: I 13.54 (3.53, 4.38, 3.21, 2.42); II 12.37 (3.21, 3.91, 3.06, 2.19); III 10.97 (3.02, 3.38, 3.01, 1.56); IV 15.09 (3.85, 4.25, 4.49, 2.50). Palp: patellar apophysis absent, RTA square slice-shaped; LTA short, approximately ⅓ the length of RTA; cymbial FURROW SHORT, LeSS THAN ½ THe LeNGTH OF CYMBIUM; eMBOLIC BASe BeGINNING AT 8 O’CLOCK POSITION; CONDUCTOR SHORT AND BIFURCATe, TWO BRANCHeS SLICe-SHAPeD AND POINTeD; MeDIAN APOPHYSIS THIN AND STRAIGHT, NOT SPOON-SHAPeD (FIGS 17A–C). Female (paratype): TOTAL LeNGTH 9.94. CARAPACe 4.81 LONG, 3.25 WIDe. ABDOMeN 5.13 LONG, 3.25 WIDe. EYe SIZeS AND INTeRDISTANCeS: AME 0.14, ALE 0.19, PME 0.16, PLE 0.17; AME–AME 0.15, AME–ALE 0.10, AME– PME 0.23, ALE–PLE 0, PME–PME 0.20, PME–PLE 0.24. LeG MeASUReMeNTS: I 11.79 (3.21, 4.24, 2.59, 1.75); II 11.47 (3.48, 3.75, 2.75, 1.49); III 10.16 (3.13, 3.39, 2.13, 1.51); IV 14.44 (3.85, 4.81, 3.75, 2.03). EPIGYNe: MIDDLe SePTUM LONG BUT NOT ReACHING THe POSTeRIOR eDGe OF ePIGYNe; ANTeRIOR ePIGYNAL SCLeRITe STRONG, WITH OBVIOUS WRINKLeS; THe HeIGHT OF IT IS LeSS THAN ½ OF THe ePIGYNAL HeIGHT; THe HeIGHT OF POSTeRIOR ePIGYNAL SCLeRITe ABOUT ½ THe ePIGYNAL HeIGHT WHILe IN THe MIDDLe OF POSTeRIOR ePIGYNAL SCLeRITe NARROW, ONLY ABOUT ¼ THe ePIGYNAL HeIGHT; ATRIUM slit-shaped; spermathecae separated, the width about ¾ the length, the distance between spermathecae 2½ the DIAMeTeR OF SPeRMATHeCA AT THe ANTeRIOR PART AND TWICe AS ITS AT THe POSTeRIOR PART; SPeRMATHeCAL HeADS SITUATeD ANTeROLATeRALLY; THe WIDTH ABOUT ¼ THe LeNGTH OF FeRTILIZATION DUCT (FIGS 18A–B). Variation: TOTAL LeNGTH VARIeS FROM 8.34 TO 9.94. Distribution. TIBeT, CHINA (FIG. 20).

Hengconarius pseudobrunneus (Wang, 2003) comb. n. FIGS 19–20 Draconarius pseudobrunneus WaNG 2003: 544, fIGS 52A–B; WaNG et al. 2010: 85, fIGS 361–406, 548; Zhu et al. 2017: 350, fIGS 218A–D. Types. Holotype: ♀ (HNNU), DANZHU He DRAINAGe, (27.631000°N, 98.621000°E, eLeV. 2700 M), 13.5 AIR KM SSW OF GONGSHAN, GONGSHAN CO., NUjIANG, Yunnan, China, 30 JUNe 2000 – 5 JULY 2000, D. KAVANAUGH, C.E. GRISWOLD, H.B. LIANG, D. UBICK, H M. YAN, D.Z. DONG LeG. Paratypes: 3♀♀ (HNNU AND CAS), SAMe DATA AS HOLOTYPe. Other records: 21♀♀ 36♂♂ (HNNU AND CAS), GAOLIGONG SHAN, Yunnan, China. Material examined. 1♀ (IZCAS-AR 33983), 6 KM E OF PIANMA TOWNSHIP (26.002433°N, 98.659067°E, eLeV. 2422 M), LUSHUI COUNTY, NUjIANG OF THe LISU AUTONOMOUS PReFeCTURe, Yunnan, China, 7 DeCeMBeR 2013, Y. LI & J. Liu leg.; 1♀ (IZCAS-Ar33984), Luoshuidong (24.96838°N, 98.81840°E, elev. 1881 m), Jiangdong Village, GUDONG TOWNSHIP, TeNGCHONG COUNTY, BAOSHAN CITY, Yunnan, China, 26 NOVeMBeR 2013, Y. LI & J. LIU LeG. Diagnosis. H. pseudobrunneus IS SIMILAR TO H. incertus AND H. latusincertus IN ePIGYNe SHAPe BUT CAN Be DISTINGUISHeD FROM H. incertus BY THe MeDIALLY SITUATeD SPeRMATHeCAL HeADS, AND FROM H. latusincertus BY THe NARROW ATRIUM, CLeAR COPULATORY DUCT AND NARROW SPeRMATHeCAe (FIGS 19A–B). Description. See WANG (2003). PHOTOS OF BOTH HABITUS AND ePIGYNe ARe PROVIDeD IN THIS STUDY (FIG. 19). Variation: TOTAL LeNGTH VARIeS FROM 6.28 TO 11.23. Distribution. YUNNAN, CHINA (FIG. 20).

Genus Hengconarius Z. Zhao & S. Li gen. n. Type species. Draconarius exilis ZhaNG, Zhu & WaNG, 2005 Etymology. THe GeNeRIC NAMe IS DeRIVeD FROM THe PINYIN "HeNG", ReFeRRING TO THe HeNGDUAN MOUNTAINS WHeRe THe GeNUS IS DISTRIBUTeD AND THe "- conarius " FROM Draconarius. THe GeNDeR IS MASCULINe. Diagnosis. Hengconarius Z. ZhaO & S. LI gen. n. IS SIMILAR TO Sinodraconarius, Nuconarius Z. ZhaO & S. LI gen. n. AND Draconarius. BUT THeY CAN Be DISTINGUISHeD IN DeTAILS AS FOLLOWS: THe NeW GeNUS CAN Be DISTINGUISHeD FROM Sinodraconarius BY A PATeLLAR APOPHYSIS THAT IS NeVeR BIFURCATe AND A CONDUCTOR WHICH IS BIFURCATe; FROM Nuconarius Z. ZhaO & S. LI gen. n. BY THe MeDIAN APOPHYSIS NOT POINTeD, THe CONDUCTOR WITH BASAL LAMeLLA AND THe ePIGYNe WITH MIDDLe SePTUM, THe ANTeRIOR eDGe OF ePIGYNe WRINKLeD AND SCLeROTIZeD, THe POSTeRIOR eDGe OF ePIGYNe SCLeROTIZeD; FROM Draconarius BY THe CYMBIAL FURROW LeSS THAN ½ THe LeNGTH OF CYMBIUM, THe MeDIAN APOPHYSIS SLICe-SHAPeD, ePIGYNAL TeeTH ABSeNT, AND THe SPeRMATHeCAe SIMPLe AND NOT CONVOLUTeD. Description. SMALL TO MeDIUM-LARGe SIZeD, WITH TOTAL LeNGTHS RANGING FROM 6.28 TO 12.91. CARAPACe YeLLOW TO TAN; RADIAL ReGION WITH GReY-GReeN SHORT SeTAe; CHeLICeRAe WITH 3 PROMARGINAL AND 2 ReTROMARGINAL TeeTH. LeG FORMULA 4> 1> 2> 3. MALe PALP: PATeLLAR APOPHYSIS PReSeNT OR ReDUCeD; TWO TIBIAL APOPHYSeS: ReTROLATeRAL TIBIAL APOPHYSIS BROAD AND slice-shaped; lateral tibial apophysis small, whose length and width about ⅓ of the length and WIDTH OF ReTROLATeRAL TIBIAL APOPHYSIS, ReSPeCTIVeLY; CYMBIAL FURROW SHORT, LeSS THAN ½ THe LeNGTH OF CYMBIUM; eMBOLIC BASe BeGINNING AT ABOUT 8 O’CLOCK POSITION; CONDUCTOR BeFURCATe, WITH A SMALL BASAL LAMeLLA AND A CONDUCTOR’S DORSAL APOPHYSIS; MeDIAN APOPHYSIS THIN, USUALLY SLICe-SHAPeD. EPIGYNe: ePIGYNAL TeeTH ABSeNT; THe ANTeRIOR eDGe OF ePIGYNe WRINKLeD AND SCLeROTIZeD; THe POSTeRIOR eDGe OF ePIGYNe SCLeROTIZeD; THe SHAPe OF ATRIUM VARIABLe; ePIGYNAL HOODS LOCATeD ANTeROLATeRALLY; COPULATORY OPeNINGS LOCATeD CeNTRALLY; COPULATORY DUCT exTeNDS HORIZONTALLY; SPeRMATHeCAe SIMPLe AND SePARATeD, THe DISTANCe BeTWeeN SPeRMATHeCAe SUBeqUAL TO THe DIAMeTeR OF SPeRMATHeCA; SPeRMATHeCAL HeADS LOCATeD AT ANTeROLATeRALLY OR IN THe MIDDLe. Natural history. SPeCIeS OF Hengconarius Z. ZhaO & S. LI gen. n. WeRe ALL COLLeCTeD BY HAND OR TRAPPING MeTHODS. THeY WeRe FOUND IN MOIST AND GLOOMY PLACeS IN THe HIGHLANDS (1881 –4089 M), SUCH AS UNDeR STONeS, ROCKS AND IN LeAF-LITTeR. Composition. SO FAR eIGHT SPeCIeS ARe CONSIDeReD IN THe NeW GeNUS: H. dedaensis Z. ZHAO & S. LI sp. n., H. exilis (ZHANG, ZHU & WANG, 2005) comb. n., H. falcatus (XU & LI, 2006) comb. n., H. incertus (WANG, 2003) comb. n., H. latusincertus (WANG, GRISWOLD & MILLeR, 2010) comb. n., H. longipalpus Z. ZHAO & S. LI sp. n., H. longpuensis Z. ZHAO & S. LI sp. n. AND H. pseudobrunneus (WANG, 2003) comb. n. Distribution. SICHUAN, TIBeT AND YUNNAN, CHINA (FIG. 20). Comments. FOR THe DeTAILS OF THe ReLATIONSHIPS IN Hengconarius Z. ZhaO & S. LI gen. n., PLeASe See ZZ630, ZZ913, ZZ929, ZZ 987, SD023 AND SD041 (SOUTHeRN COeLOTeS GROUPS) IN FIGURe 3 AND SUPPLeMeNTARY FIGUReS S4– S 6 OF Z. ZHAO & S. LI (2017).

Hengconarius longipalpus Z. Zhao & S. Li sp. n. FIGS 15���16, 20 Type material. Holotype: ♂ (IZCAS-AR33978), FeNGxUe YAKOU (25.972667��N, 98.683933��E, eLeV. 3133 M), PIANMA TOWNSHIP, LUSHUI COUNTY, NUjIANG OF THe LISU AUTONOMOUS PReFeCTURe, Yunnan, China, 25 JUNe 2013, Z. ZHAO & J.C. LIU LeG. Paratype: 1♀ (IZCAS-AR33979), FeNGxUe YAKOU (25.972667��N, 98.683933��E, eLeV. 3133 M), PIANMA TOWNSHIP, LUSHUI COUNTY, NUjIANG OF THe LISU AUTONOMOUS PReFeCTURe, Yunnan, China, 8 DeCeMBeR 2013, Y. LI & J. LIU LeG. Etymology. THe SPeCIFIC NAMe IS DeRIVeD FROM THe LONG MALe PALP (LATIN PReFIx "LONGUS-" (LONG), NOUN " PALP " AND AFFIx "- US "); ADjeCTIVe. Diagnosis. THe MALeS ARe SIMILAR TO H. exilis BUT DIFFeR BY THe SHORT AND HORIZONTAL CONDUCTOR WHILe IN H. exilis, THe eND OF CONDUCTOR SLIGHTLY UPWARD (FIGS 9B, 15B); THe eND OF MeDIAN APOPHYSIS BLACK-BROWN AND OPAqUe BUT TRANSLUCeNT IN H. exilis (FIGS 9C, 15C). THe FeMALeS CAN Be eASILY DISTINGUISHeD FROM H. exilis BY THe LARGe POSTeRIOR epigynal sclerite; the height of posterior epigynal sclerite subequal to ⅓ the height of epigyne, while it is twice in H. exilis (FIGS 10A, 16A); THe BROAD SPeRMATHeCAL HeADS, WHOSe WIDTH eqUALS TO THe LeNGTH BUT IN H. exilis, THe LeNGTH OF SPeRMATHeCAL HeADS IS LONGeR THAN THe WIDTH AND FeRTILIZATION DUCT THIN (FIGS 10B, 16B). Description. Male (holotype): TOTAL LeNGTH 10.76. CARAPACe 5.74 LONG, 4.25 WIDe. ABDOMeN 5.02 LONG, 3.51 WIDe. EYe SIZeS AND INTeRDISTANCeS: AME 0.13, ALE 0.25, PME 0.23, PLE 0.24; AME���AME 0.08, AME���ALE 0.05, AME���PME 0.15, ALE���PLE 0, PME���PME 0.15, PME���PLE 0.15. LeG MeASUReMeNTS: I 29.07 (8.42, 8.15, 7.91, 4.59); II 27.04 (7.91, 7.65, 7.14, 4.34); III 25.50 (7.65, 6.88, 6.89, 4.08); IV 29.57 (8.93, 7.65, 8.41, 4.58). Palp: patellar apophysis absent; RTA square slice-shaped; LTA short, approximately ⅓ the length of RTA; cymbial FURROW SHORT, LeSS THAN �� THe LeNGTH OF CYMBIUM; eMBOLUS BeGINNING AT 8 O���CLOCK POSITION; CONDUCTOR SHORT, BIFURCATe AND HORIZONTAL; MeDIAN APOPHYSIS THIN AND SLICe-SHAPeD (FIGS 15A���C). Female (paratype): TOTAL LeNGTH 10.69. CARAPACe 5.61 LONG, 3.31 WIDe. ABDOMeN 5.08 LONG, 3.29 WIDe. EYe SIZeS AND INTeRDISTANCeS: AME 0.13, ALE 0.24, PME 0.21, PLE 0.25; AME���AME 0.10, AME���ALE 0.08, AME��� PME 0.14, ALE���PLE 0, PME���PME 0.13, PME���PLE 0.14. LeG MeASUReMeNTS: I 17.07 (5.62, 5.09, 3.82, 2.54); II 16.29 (5.61, 4.58, 3.82, 2.28); III 14.52 (4.85, 4.07, 3.56, 2.04); IV 18.34 (6.12, 5.09, 4.84, 2.29). EPIGYNe: MIDDLe SePTUM exTeNDS TO THe POSTeRIOR eDGe OF ePIGYNe; ANTeRIOR ePIGYNe WeAKLY SCLeRITeD, WITH LIGHTLY WRINKLeS, THe HeIGHT of it about �� the epigynal height; the height of posterior epigynal sclerotized is more than ⅓ the epigynal height; atrium trajectile, occupying about �� of epigynal plate; spermathecae simple and separated, the distance between the anterior spermathecae is twice the spermathecal diameters; the width of spermathecae about �� the length; SPeRMATHeCAL HeADS SITUATeD ANTeROLATeRALLY (FIGS 16A���B). Variation: TOTAL LeNGTH VARIeS FROM 10.69 TO 12.76. Distribution. YUNNAN, CHINA (FIG. 20)., Published as part of Li, Bing, Zhao, Zhe, Zhang, Chuntian & Li, Shuqiang, 2018, Nuconarius gen. n. and Hengconarius gen. n., two new genera of Coelotinae (Araneae, Agelenidae) spiders from Southwest China, pp. 237-263 in Zootaxa 4457 (2) on pages 254-256, DOI: 10.11646/zootaxa.4457.2.2, http://zenodo.org/record/1342559