Search

Setostylus triumphus Qi, Huang et Wang sp. n. (Figs 3–4) Diagnosis. This new species resembles the type species of the genus, S. singularis (Lane) in the male adult appearance, but it can be easily distinguished from the latter by the suboval and brown maxillary palpus of the male, and the gonocoxites having raised posteromesal part in the male terminalia. In S. singularis, the maxillary palpus is globose and black, and the gonocoxites have emarginated posteromesal part. Type material. Holotype. Male, CHINA: Zhejiang, Mt. Longwang (30.24° N, 119.26° E), 980 m, 27.VII.2018, coll. Caixia Liu, slide no. LWS-5-59. Paratype. Male, CHINA: Guangdong, Mt. Dinghu (23.10° N, 112.32° E), 235 m, 12.VI.2020, coll. Lei Qi, slide no. DHS-8-67. DNA barcode. The DNA barcode sequences from holotype and paratype are deposited in BOLD and GenBank. The BIN number is AEJ 6195 in BOLD. The GenBank accession numbers for holotype and paratype are ON190027 and ON190028, respectively. Description Male (Fig. 3). Body length (without antennae) 5.0 mm. wing length 1.4 mm, width 3.0 mm. Head (Fig. 4a) pale brown, with width 1.2 times of its height. Vertex with numerous dark setae, frons and face bare. Three ocelli in approximately linear arrangement, medial ocellus about 1/3 diameter of lateral one. Compound eyes almost hemispheric, covered with pointed setae. Clypeus pale brown with 12–16 dark brown setae. Mouthparts reduced, maxillary palpus with two segments, segment 2 erecting forward, suboval, with 8–10 setae laterally. Antenna (Fig. 4c) brown and strongly compressed; scape and pedicel nearly funnel-shaped, bearing several subapical and apical setae; flagellum pectinate and 14-segmented, each flagellomere bearing 6–10 setae on dorsal edge, segments 1–8 with a few microtrichia ventrally, subtriangular apically. Thorax (Figs 4d–e) pale brown to brown. Scutum brown, bearing dense setae, with a V-shaped streak. Scutellum brown, with a row of marginal setae. Mediotergite pale brown, bare. Anepisternum dark brown, with several setae on top corner. Laterotergite pale brown and subrectangular, bare. Halter length 0.3 mm, stem translucent; knob dark brown, bearing sparse setae. Wing (Fig. 4b) brown, without pattern, bearing dense microtrichia on surface. Wing margin with short brown setae. R 1 and R 4+5 with dark setae arranged along the veins. Vein C stops at nearly middle of R 4+5 to M 1. Sc ends at basal 1/3 of C, about half length of R 1. Vein m-cu and R+M fusion subequal in length; Rs about twice of m-cu in length. CuA curved. CuP free, with end approaching to outer margin. Legs (Fig. 3) pale brown to dark brown. Fore coxa brown. Mid coxa with basal 2/3 pale brown and bare, distal 1/3 dark brown and setose. Hind coxa dorsally covered with sparse setae. Femur, tibia and tarsus brown, covered with numerous setae. Hind tibia with two combs apically, in linear arrangement. Abdomen (Fig. 3) pale brown to dark brown, bearing dense dark setae. Tergite 1 brown, tergites 2–4 somewhat transparent, tergites 5–8 dark brown. Terminalia (Figs 4f–h) brown. Tergite 9 subtrapezoidal, with abundant setae. Gonostylus curved, narrowed gradually from base to end; basal 1/2 bearing abundant dark brown setae on surface, 8–10 black spines on inner margin medially, and having a triangular projection distal-laterally; distal 1/2 hyaline, bare, with a dark brown apical bristle; hyaline part about twice length of bristle. Gonocoxites brown, bearing dense setae; posterior margin raised and divided medially, with 8–10 black strong spines posteriorly. Cercus thinly digitate, bearing setae. Female. Unknown. Etymology. The name of this new species is derived from the Latin word triumphus (victory), referring to the V-shaped streak on the scutum., Published as part of Qi, Lei, Huang, Junhao, Wu, Hong & Wang, Qingyun, 2022, Two new species of Setostylus Matile, 1990 (Diptera: Keroplatidae) from China, pp. 443-450 in Zootaxa 5165 (3) on pages 446-448, DOI: 10.11646/zootaxa.5165.3.9, http://zenodo.org/record/6838620

Setostylus tridigitus Qi, Huang et Wang sp. n. (Figs 1–2) Diagnosis. This new species resembles S. abdominalis (Sasakawa & Tamu) in wing pattern and general appearance of the male terminalia, but it differs from the latter in the subtriangular terminal flagellomere in the male antennae, and gonocoxites having 18–20 black strong spines on each posteromesal lobiform part in the male terminalia. In S. abdominalis, the terminal flagellomere is suboval in the male antennae, and the gonocoxites have 23–25 short spines on each posteromesal lobiform part in the male terminalia. Type material. Holotype. Male, CHINA: Guangdong, Mt. Dinghu (23.92° N, 112.32° E), 72 m, 27.VIII.2020, coll. Lei Qi, slide no. DHS-8-85. Paratypes. Male, CHINA: Guizhou, Mt. Leigong (26.36° N, 108.16° E), 3.X.2019, coll. Changlie Yang, slide no. LGS-10-42; male, Guangdong, Chebaling National Nature Reserve (24.42° N, 114.15° E), 378 m, 12.VI.2020, coll. Lei Qi, slide no. CBL-8-39; male, Guangxi, Dayaoshan National Nature Reserve (24.14° N, 110.19° E), 1,520 m, 8.VI.2020, coll. Tao Li, slide no. DYS-10-70. DNA barcode. The DNA barcode sequence from holotype and paratypes are deposited in BOLD (https://boldsystems.org/) and GenBank (https://www.ncbi.nlm.nih.gov/), with the BIN number of AEI9998 and the GenBank accession number of ON190026. Description. Male (Fig. 1). Body length (without antennae) 4.5 mm. Wing length 1.3 mm, width 2.8 mm. Head (Fig. 2a) dark brown, with width 1.3 times of its height. Vertex with abundant dark setae, frons and face bare. Three ocelli arranged nearly in a line, lateral ocelli about three times of median one in diameter. Compound eyes almost hemispheric, covered with pointed setae. Clypeus brown with about 25–30 dark brown setae. Mouthparts reduced, maxillary palpus with two segments; segment 2 erecting forward, oval, with 6–8 setae laterally. Antenna (Fig. 2c) brown and strongly compressed; scape and pedicel funnel-shaped, covered with several subapical and apical setae; flagellum pectinate and 14-segmented, each flagellomere bearing 6–10 setae on dorsal edge, segments 1–8 with a few microtrichia ventrally, subtriangular apically. Thorax (Figs 2d–e) brown. Scutum brown, covered with bushy setae, three dark brown longitudinal stripes. Scutellum brown, with several setae. Mediotergite brown, bare. Anepisternum dark brown, subtriangular, with thick setae on top corner. Laterotergite pale brown and subelliptical, bare. Halter length 0.8 mm, stem translucent, bare; knob dark brown, bearing setae. Wing (Fig. 2b) brown, without pattern, veins distinct. Wing membrane covered with abundant brown microsetae. Wing margin with short brown setae. R 1 and R 4+5 with setae arranged along the veins. Vein C terminates at 1/2 of R 4+5 to M 1. Vein m-cu and R+M fusion subequal in length; Rs about twice length of m-cu. Vein M 1 and M 2 weaker than other veins. CuA slightly curved; CuP free terminally, approximates outer margin. Legs (Fig. 1) brown to dark brown. Fore coxa brown, covered with dense setae. Mid and hind coxae pale brown and bare on basal 2/3, dark brown and setose on distal 1/3. Femur, tibia and tarsus brown, covered with numerous setae. Hind tibia with two separated combs apically. Abdomen (Fig. 1) brown to dark brown, with tergite 1 brown, tergites 2–4 pale and 5-8 dark brown, covered with dense dark setae. Terminalia (Figs 2f–h) brown. Tergite 9 subpentagonal, with abundant setae. Gonostylus curved, narrowed gradually towards end; basal 2/3 covered with setae, 10–12 black spines on inner margin, and having a tiny projection distal-laterally; apical process bare and subtransparent, slightly raised at half of inner margin, with a dark brown bristle terminally, hyaline part about 1.5 times length of bristle. Gonocoxites brown, with narrow incision at middle posteriorly, bearing 18–20 black strong spines on each posteromesal lobiform part. Cercus pale brown, digitate, with numerous setae. Female. Unknown. Etymology. The name of this new species comes from the Latin tri (three) and digitus (finger), which refers to the three-toed streak on the scutum., Published as part of Qi, Lei, Huang, Junhao, Wu, Hong & Wang, Qingyun, 2022, Two new species of Setostylus Matile, 1990 (Diptera: Keroplatidae) from China, pp. 443-450 in Zootaxa 5165 (3) on pages 444-446, DOI: 10.11646/zootaxa.5165.3.9, http://zenodo.org/record/6838620

Key to the species of Setostylus (based on male adults) 1. Two ocelli.......................................................................................... 2 - Three ocelli......................................................................................... 3 2. Flagellomere 14 with an antennal apicule; gonocoxites with anterior margin not emarginated (Matile 1990: Fig. 919)............................................................................... S. singularis (Lane) [Neotropical] - Flagellomere 14 without an antennal apicule; gonocoxites with anterior margin distinctly emarginated (Matile 1990: Fig. 922)............................................................................ S. bifidus Matile [Neotropical] 3. Wing without pattern.................................................................................. 4 - Wing with distinct patterns............................................................................. 9 4. Scutellum without bristles.............................................................................. 5 - Scutellum covered with bristles.......................................................................... 6 5. Tergite 9 longer than wide (Matile 1990: Fig. 936); tarsus I 1.5 times longer than the tibia. S. rufobrunneus Matile [Oriental] - Tergite 9 wider than long (Matile 1990: Fig. 937); tarsus I 2.5 times longer than the tibia..... S. bispinosus Matile [Oriental] 6. Scutum with pattern................................................................................... 7 - Scutum without pattern; flagellomere 14 cordiform apically (Papp et al. 2006)................ S. alienus Papp [Oriental] 7. Scutum with two stripes............................................................................... 8 - Scutum with three stripes; tergite 9 subpentagonal (Fig. 2).............................. S. tridigitus sp. n. [Oriental] 8. Scape and pedicel dark brown; palpus brown; tergites 2–4 somewhat transparent (Fig. 4)...... S. triumphus sp. n [Oriental] - Scape and pedicel reddish brown; palpus reddish brown; tergites 2–4 with a large yellow basal spot on both sides (Matile 1990)......................................................................... S. stubbsi Matile [Oriental] 9. Posterior margin of tergite 9 emarginated................................................................. 10 - Posterior margin of tergite 9 rounded.................................................................... 11 10. R +M fusion longer than the stem of median fork; gonostylus dorsomedially with a small, semi-transparent, posterodorsallyprojecting tooth (Huerta & Fitzgerald, 2020: Figs 2, 3D)........................ S. xoxo Huerta & Fitzgerald [Nearctic] - R +M fusion equal to half the stem of median fork; gonostylus dorsomedially without tooth (Matile 1990: Fig. 921)................................................................................ S. pictipennis Matile [Neotropical] 11. Goncoxites with posterior margin emarginated............................................................. 12 - Goncoxites with posterior margin not emarginated, with two submedian tubercles (Ševčík et al. 2021: Fig. 14A)..................................................................... S. fangshuoi Ševčík, Hippa & Burdíková [Oriental] 12. Wing with light brown fumose along costal edge near apex (Huerta & Fitzgerald, 2020)..................................................................................................... S. bellulus (Williston) [Neotropical] - Wing with light brown fumose along costal edge near anterior apical portion of wing to distinctly marked with brown more broadly............................................................................................ 13 13. Palpus yellow....................................................................................... 14 - Palpus reddish brown; halteres with yellowish stem and black knob (Edwards 1931)...... S. innotatus (Edwards) [Oriental] 14. Costa reaching slightly beyond one-third distance from vein R 4+5 to M 1;gonostylus with a minute projection on dorsoapical part, arcuate projection ventrally (Sasakawa & Tamu 1961: Figs 1–2).......... S. abdominalis (Sasakawa & Tamu) [Palaearctic] - Costa reaching almost midway between R 4+5 to M1; gonostylus without projection (Xu et al. 2007: Figs 5 –6).......................................................................... S. chinensis Cao, Evenhuis & Zhou [Palaearctic], Published as part of Qi, Lei, Huang, Junhao, Wu, Hong & Wang, Qingyun, 2022, Two new species of Setostylus Matile, 1990 (Diptera: Keroplatidae) from China, pp. 443-450 in Zootaxa 5165 (3) on pages 448-449, DOI: 10.11646/zootaxa.5165.3.9, http://zenodo.org/record/6838620, {"references":["Matile, L. (1990) Recherches sur la systematique et l'evolution des Keroplatidae (Diptera, Mycetophilidae). Memoires du Museum National d'Histoire Naturelle, Series A, 148, 1 - 682.","Papp, L., Merz, B. & Foldvari, M. (2006) Diptera of Thailand. A summary of the families and genera with references to the species representations. Acta Zoologica Academiae Scientiarum Hungaricae, 52 (2), 97 - 269.","Huerta, H. & Fitzgerald, S. J. (2020) A new species of Setostylus Matile and new records of Setostylus bellulus (Williston) (Diptera: Keroplatidae). Zootaxa, 4722 (1), 84 - 90. https: // doi. org / 10.11646 / zootaxa. 4722.1.8","Sevcik, J., Hippa, H. & Burdikova, N. (2021) Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera. Insects, 13 (1), 19. https: // doi. org / 10.3390 / insects 13010019","Edwards, F. W. (1931) Fauna Sumatrensis. Mycetophilidae (Diptera). Tijdshrift voor Entomologie, 74, 262 - 278.","Sasakawa, M. & Tamu, N. (1961) Japanese Fungivoridae (Diptera) III: New or Little-known fungus gnats from the Tsushima Islands (Agriculture). The scientific reports of Kyoto Prefectural University. Agriculture, 13, 68 - 69.","Xu, H. C., Cao, J., Zhou, Z. J., Wu, H. & Huang, Z. W. (2007) First record of the tribe Keroplatini from China, with descriptions of two new species (Diptera: Keroplatidae). Zootaxa, 1497 (1), 35 - 40. https: // doi. org / 10.11646 / zootaxa. 1497.1.3"]}

Heteropterna (Heteropterna s. str.) fanjingshana Wang et Huang sp. n. (Figs 3���4) Diagnosis. This new species is similar to H. septemtrionalis (Okada) by having similar antennae with the last four flagellomeres yellowish white. It can be recognized by the wing having medial hyaline spot oblique outward, and the internal lobe of gonostylus apically straight in ventral view (Figs 4g, i���k). In H. septemtrionalis, the wing has a medial hyaline spot oblique inward, and the internal lobe of gonostylus apically hooked in ventral view (Matile, 1990: Figs 586���587). Type material. Holotype. Male, CHINA: Guizhou Province, Mt. Fanjing (27.95�� N, 108.77�� E), 840 m, 5.VI.2020, coll. Zhenghai Yang, slide no. FJS-10-28. Description. Male (Fig. 3). Body length (without antennae) 5.3 mm. Wing length 3.4 mm. Length of terminalia 0.5 mm. Head (Figs 4a���b) dark brown. Compound eyes (Fig. 4a) hemispheric, making up most area of head, covered with dense pubescence superficially. Three ocelli (Fig. 4b) lying between upper edges of eyes, arranged in an inverted triangle, with median ocellus at a slightly lower position, lateral ocelli about three times as median one in diameter. Mouthparts (Fig. 4a) reduced, palpus two-segmented and setose, labrum ligulate, labellum has bushy setae laterally.Antenna (Fig. 4c) pectinate and laterally compressed, about 1.5 times length of head width; scape and pedicel subtrapezoid, with setae dorsally and ventrally; flagellum 14-segmented and comb-like, covered with serried microtrichia laterally, with countable setae dorsally and ventrally; flagellomeres 1���10 brown (except 7th segment yellowish white laterally), flagellomeres 11���14 yellowish white, with last segment pale brown terminally. Thorax (Figs 4d���f) brown. Anepisternum dark brown and subtriangular, bare. Mesonotum brown; medial stripe dark brown, narrowly bandlike, reaching to posterior 2/5. Scutellum dark brown, subrectangular. Mediotergite yellowish brown and subtriangular, medial membrane inverted triangular. Laterotergite brown and subelliptical. Halter length 0.5 mm; stem translucent, with distal half bearing setae anteriorly; knob dark brown, oval. Wing (Fig. 4g) pale brown, covered with thick microtrichia superficially, outer and posterior margin with short cilia. Fuscous stripe along costa, approaching to 2/7 width of wing, stretching from base to end of costa. Medial hyaline spot irregular extending from end of Sc outward to middle of R, apical hyaline spot extending from end of R 2+3 obliquely inward to bottom margin of stripe. Veins brown to dark brown, with two rows of microsetae on C, and a row of microsetae on R 1, R 2+3 and R 4+5. Vein C terminate at 2/5 of R 4+5 -M 1. M 1 and M 2 weaker than other veins, with M 1 ending at outer margin, and M 2 ending far from outer margin. CuA curved backward at midpoint. False vein next to CuA, with free end. Vein r-m degenerated. Legs (Fig. 3) yellowish white to dark brown. Hind leg dark brown, except femora yellowish white interiorly, with thick setae. Fore and mid legs damaged, only with fuscous coxae left. Abdomen (Fig. 4h) brown, with dense setae dorso-ventrally. Tergites pale brown. Sternites dark brown, with paired suboval hyaline spots from sternites 2 to 4 on each segment. Terminalia (Figs 4i���k) brown. Tergite 9 subtrapezoidal, shallowly concave at posterior margin. Cercus subtriangular, setose dorsally. Gonocoxites with posterior 1/3 weakly sclerotized along median line; posterior protuberance horn-shaped, covered with intensive setae ventrally. In ventral view external lobe of gonostylus finely digitate, obtuse terminally, slightly shorter than width of gonocoxite, with dense bristles outside; internal lobe of gonostylus horn-like, about 2/3 length of external lobe, with setae inside and long bristles outside. In lateral view dorsal lobe of gonostylus enlarged dorsad at middle, with thick setae superficially. Female. Unknown. Etymology. This species is named after Mt. Fanjing, the collection site of type specimen; noun in genitive case., Published as part of Wang, Qingyun, Qi, Lei, Wu, Hong & Huang, Junhao, 2021, Two new species of Heteropterna Skuse, 1888 (Diptera: Keroplatidae) from China, pp. 277-284 in Zootaxa 4985 (2) on pages 281-283, DOI: 10.11646/zootaxa.4985.2.11, http://zenodo.org/record/5055074, {"references":["Matile, L. (1990) Recherches sur la systematique et l'evolution des Keroplatidae (Diptera, Mycetophilidae). Memoires du Museum National d'Histoire Naturelle, (A), 148, 1 - 682."]}

Heteropterna (Heteropterna s. str.) cuneata Wang et Huang sp. n. (Figs 1���2) Diagnosis. This new species resembles H. annulipes (Colless) in appearance by having similar wing pattern, but can be recognized by the cercus tapering to the apex, the external lobe of gonostylus apically rounded in ventral view, and the gonocoxites with posterior 1/5 split into halves in the terminalia (Figs 2i���k). In H. annulipes, the cercus is reniform, the external lobe of gonostylus has a pointed end in ventral view, and the gonocoxites are fused posteriorly in the terminalia (Colless 1966: Figs 1a���c). Type material. Holotype. Male, CHINA: Guizhou Province, Mt. Fanjing (27.8�� N, 108.62�� E), 750 m, 5.VI��� 4.VII.2020, coll. Fanliang Liu, slide no. FJS-11-20. Description. Male (Fig. 1). Body length (without antennae) 5.5 mm. Wing length 3.2 mm. Length of terminalia 0.56 mm. Head dark brown. Compound eyes (Fig. 2a) hemispheric, occupying most area of head, bearing dense pubescence on surface. Three ocelli (Fig. 2b) lying between upper edges of eyes, arranged in an inverted triangle, with lateral ones about three times of median one in diameter. Mouthparts (Fig. 2a) reduced, palpus two-segmented and porrect, labrum subtriangular, labellum has bushy setae outside. Antenna (Fig. 2c) pectinate and laterally compressed, slightly longer than head width; scape and pedicel subtrapezoid, covered with dense microtrichia laterally, having setae in different length dorsally and ventrally; flagellum 14-segmented and comb-like, bearing dense microtrichia laterally, with a few setae dorsally and ventrally; flagellomeres 1���11 brown (except 7th and 11th segment partly yellowish white), flagellomeres 12���14 yellowish white, with last one somewhat dark apically. Thorax (Figs 2d���f) brown. Prothoracic spiracle membranous, elliptical. Anepisternum dark brown and subtriangular, with sparse setae. Mesonotum brown, middle streak dark brown and cuneiform, with yellowish brown stripes along lateral sides. Scutellum fuscous, subrectangular. Mediotergite yellowish brown and subtriangular, with medial membranous area inverted triangular. Laterotergite fuscous brown, suboval. Halter length 0.38 mm; stem translucent, with setae anteriorly; knob dark brown, with sparse setae anteriorly. Wing (Fig. 2g) pale brown, having dense microtrichia on surface, outer and posterior margin with short cilia. Fuscous stripe along costal margin, about 1/4 width of wing, extending from base to end of costa. Hyaline spots irregular, with a smaller medial spot from end of Sc obliquely outward to above middle of R, and a larger apical spot from end of R 2+3 obliquely outward to above bottom margin of stripe. Veins brown to dark brown, with two rows of microsetae on C, and a row of microsetae on R 1 and R 4+5. Vein C stops at 5/12 of R 4+5 -M 1. M 1 and M 2 weaker than other veins, with M 1 ending at outer margin and M 2 ending far from outer margin. CuA curved backward at middle. False vein close to CuA, with free end. Vein r-m degenerated and weakly sclerotized. Legs (Fig. 1) yellowish white to dark brown. Coxae dark brown, covered with setae densely. Femora yellowish brown, scattered with setae. Tibiae and tarsi light brown, bearing thick microtrichia. Abdomen (Fig. 2h) brown, covered with dense setae. Tergites light brown. Sternites dark brown, with paired subelliptical hyaline spots on sternites 2 to 4. Terminalia (Figs 2i���k) brown. Tergite 9 subtrapezoidal, concave both anteriorly and posteriorly, covered with setae densely. Cercus setose dorsally, wide basally, tapering to posterior end, with distal half slightly curved inward. Gonocoxites ventrally with anterior 4/5 fused, posterior 1/5 spilt into halves; posterior protuberance subtriangular, with stout setae ventrally. In ventral view external lobe of gonostylus thinly digitate, rounded terminally, slightly shorter than width of gonocoxite, having dense long bristles laterally; internal lobe of gonostylus corniform, about 2/3 length of external lobe, with short setae inside and long bristles outside. In lateral view dorsal lobe of gonostylus subtriangular, covered with dense setae. Female. Unknown. Etymology. The name of this species is from the Latin cuneatus (cuneiform), in reference to the cuneiform streak at middle of mesonotum; adjective in genitive case., Published as part of Wang, Qingyun, Qi, Lei, Wu, Hong & Huang, Junhao, 2021, Two new species of Heteropterna Skuse, 1888 (Diptera: Keroplatidae) from China, pp. 277-284 in Zootaxa 4985 (2) on pages 278-279, DOI: 10.11646/zootaxa.4985.2.11, http://zenodo.org/record/5055074, {"references":["Colless, D. H. (1966) Diptera: Mycetophilidae. Insects of Micronesia, 12 (6), 637 - 667."]}

Two new species of Heteroptera (Diptera: Keroplatidae: Keroplatinae), H. cuneata sp. n. and H. fanjingshana sp. n. are described from Guizhou, China. Key to adult males of Heteropterna from China is given. Habitus images of adults and morphological structures are provided.

Heteropterna Skuse, 1888 Heteroptera Skuse, 1888. Linn. Soc. New South Wales, Proc. 3: 166; Edwards, 1929. Linn. Soc. New South Wales, Proc. 54: 174. Type species: Heteropterna macleayi Skuse, 1888, by monotypy. Key to the species of Heteropterna in China (based on male adults) 1 Antenna with flagellomeres 11���14 yellowish white.......................................................... 2 - Antenna with flagellomere 11 mainly brown, flagellomeres 12���14 yellowish white, mesonotum with a cuneiform medial streak................................................................................. H. cuneata sp. n. 2 Medial hyaline wing spot with posterior side curving toward wing tip, mesonotum with a band-like medial stripe, internal lobe of gonostylus apically straight in ventral view............................................. H. fanjingshana sp. n. - Medial hyaline wing spot with posterior end curving toward base of wing, mesonotum with a cuneiform medial stripe, internal lobe of gonostylus apically hooked in ventral view..................................... H. septemtrionalis (Okada), Published as part of Wang, Qingyun, Qi, Lei, Wu, Hong & Huang, Junhao, 2021, Two new species of Heteropterna Skuse, 1888 (Diptera: Keroplatidae) from China, pp. 277-284 in Zootaxa 4985 (2) on page 278, DOI: 10.11646/zootaxa.4985.2.11, http://zenodo.org/record/5055074, {"references":["Skuse, R. A. A. (1888) Diptera of Australia. Part III. The Mycetophilidae. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 1123 - 1222.","Edwards, F. W. (1929) Notes on the Ceroplatinae, with descriptions of new Australian species (Diptera, Mycetophilidae). Proceedings of the Linnean Society of New South Wales, 54 (3), 162 - 175."]}

Pseudoaerumnosa tkoci Vilkamaa, Halenius & Ševčík, 2019 (Figs. 5, 6C) New material. CHINA: Zhejiang province, Lin’an, 3 male, Tianmu Mountain, Malaise trap, 29.VII.2018, Zuluan Chen (ZAFU) [XWCZ-15, XWCZ-24, XWCZ-26]; 1 male, same data, but 27.III.2018 [XWCZ-38]. Redescription. Male. Brown. Head. Eye bridge 2–3 rows of facets. Prefrons with 10 setae. Clypeus with no setae. Palpus (Fig. 5B) one-segmented, with 3 setae and a large dorsal patch of sensillae. Antennal flagellomeres (Fig. 5A) asymmetrical and the frontal part larger. Length/width of 4th flagellomere: 2.20–2.39. Neck length/width: 0.50–0.54. Thorax. Anterior pronotum with 5–6 setae. Proepisternum with 2–3 setae. Wings (Fig. 6C). Length 2.74–2.84 mm. Width/length: 0.47–0.49. R1/R: 1.68. c/w: 0.66. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia unmodified without specialized setae. Foretibia spur longer than the tibial width. Hypopygium (Fig. 5C). Gonocoxite slightly longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe- or finger-like projections, without setose area. Gonostylus elongated with a medial lobe, which has 3 megasetae. Apical tooth with a megaseta attached to its ventral side, with 2 subapical megasetae near the tooth, all these megasetae slender and slightly curved. Tegmen wider than long, nearly straight apically, narrowed at middle. Aedeagus with distinct apical fork and basal sclerotization. Distribution. China: Taiwan, Zhejiang. Remarks. The new specimens examined show a slight difference from the type specimen by the apical part of the apical tooth in the Chinese specimens being shorter than in the holotype.

Pseudoaerumnosa regularis Chen & Huang sp. n. (Figs. 2, 3A���B, 6A) Specimens examined. Holotype: male. CHINA: Zhejiang province, Lin���an, Tianmu Mountain, Malaise trap, 29.VIII.2018, Zuluan Chen (ZAFU) [XWCZ-10]. Paratypes: CHINA: 12 males, same data as holotype (ZAFU) [XWCZ-2, XWCZ-5, XWCZ-6, XWCZ-8, XWCZ-9, XWCZ-11, XWCZ-25, XWCZ-27, XWCZ-28, XWCZ-29, XWCZ-30, XWCZ-41]. Description. Male. Brown. Head. Eye bridge 2���3 rows of facets. Prefrons with 10 setae. Clypeus with no setae. Palpus one-segmented, with 3 setae and a large dorsal patch of sensillae. Antennal flagellomeres (Fig. 2A) asymmetrical with the frontal part larger. Length/width of 4th flagellomere: 1.50���1.60. Neck length/width: 0.58��� 0.59. Thorax. Anterior pronotum with 5���6 setae. Proepisternum with 3���12 setae. Wings (Fig. 6A). Length 2.47 mm. Width/length: 0.55���0.60. R1/R: 2.26. c/w: 0.55���0.85. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia unmodified without specialized setae. Foretibia spur longer than the tibial width (Fig. 2B). Hypopygium (Figs. 2C, 3A���B). Gonocoxite longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe-like or finger-like production, without setose area. Gonostylus elongated with a medial lobe, which has 3 megasetae. Apical tooth with a long megaseta attached to its ventral side, with 2 subapical megasetae near the tooth, all these megasetae slender and slightly curved. Tegmen wider than long, nearly straight apically, basolaterally sclerotized. Aedeagus with distinct apical fork and basal sclerotization. Distribution. China: Zhejiang. Remarks. Pseudoaerumnosa regularis sp. n. is similar to P. annae in having an elongate gonostylus and the tegmen straight laterally, but differs in having a narrow aedeagus and 3 megasetae on the lobe of gonostylus, while having a broad aedeagus and 4 megasetae on the lobe of gonostylus in P. annae. Etymology. The species is named after its regularly-shaped hypopygium, from the Latin, regularis, meaning regularity., Published as part of Chen, Zuluan, Yang, Xue, Menzel, Frank, Wu, Hong & Huang, Junhao, 2021, Three Oriental species of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae) from China, pp. 551-562 in Zootaxa 4969 (3) on pages 556-558, DOI: 10.11646/zootaxa.4969.3.7, http://zenodo.org/record/4751483

Chen, Zuluan, Yang, Xue, Menzel, Frank, Wu, Hong, Huang, Junhao (2021): Three Oriental species of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae) from China. Zootaxa 4969 (3): 551-562, DOI: https://doi.org/10.11646/zootaxa.4969.3.7

Pseudoaerumnosa tianmushana Chen & Huang sp. n. (Figs. 3C–F, 4, 6B) Specimens examined. Holotype: male. CHINA: Zhejiang province, Lin’an, Tianmu Mountain, Malaise trap, 29.VII.2018, Zuluan Chen (ZAFU) [XWCZ-21]. Paratypes, 15 males, same data as holotype (ZAFU) [XWCZ-3, XWCZ-12, XWCZ-13, XWCZ-14, XWCZ-16, XWCZ-17, XWCZ-18, XWCZ-19, XWCZ-20, XWCZ-23, XWCZ- 31, XWCZ-32, XWCZ-33, XWCZ-37, XWCZ-42]. Description. Male. Brown. Head. Eye bridge with 2–3 rows of facets. Prefrons with 13 setae. Clypeus with no setae. Palpus one-segmented, with 2 setae and a large dorsal patch of sensilla. Antennal flagellomeres (Fig. 4A) asymmetrical with the frontal part larger. Length/width of 4th flagellomere: 1.88–2.09. Neck length/width: 0.93–0.94. Thorax. Anterior pronotum with 4–5 setae. Prothoracic episternum with 2–3 setae. Wings (Fig. 6B). Length 2.36–2.47 mm. Width/length: 0.46–0.48. R1/R: 1.64. c/w: 0.79–0.80. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia (Fig. 4B) unmodified, without specialized setae. Foretibia spur longer than the tibial width. Hypopygium (Figs. 3C–F, 4C). Gonocoxite longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe- or finger-like projections, without setose area. Gonostylus elongated with a medial lobe, which have 3 megasetae.Apical tooth with a long megaseta attached to its ventral side and 2 subapical megasetae near the tooth, the megasetae slender and slightly curved. Tegmen broader than long, smoothly rounded apically and basolaterally sclerotized. Aedeagus with an indistinct apical fork and a basocentral sclerotized central strip. Distribution. China: Zhejiang. Remarks. Pseudoaerumnosa tianmushana sp. n. resembles P. formosa in its elongated gonostylus and tegmen narrowed above the middle, but differs by having an aedeagus with indistinct apical fork and gonostylus medial lobe reaching to apical 1 / 3 of gonostylus, while having an aedeagus with distinct apical fork and gonostylus with medial lobe reaching to apical 1 / 4 of gonostylus in P. formosa. Etymology. This species is named after its type locality, Tianmu Mountain (Chinese: Tianmu-shan) in Zhejiang province, China.

Pseudoaerumnosa tkoci Vilkamaa, Halenius & ��ev�k, 2019 (Figs. 5, 6C) New material. CHINA: Zhejiang province, Lin���an, 3 male, Tianmu Mountain, Malaise trap, 29.VII.2018, Zuluan Chen (ZAFU) [XWCZ-15, XWCZ-24, XWCZ-26]; 1 male, same data, but 27.III.2018 [XWCZ-38]. Redescription. Male. Brown. Head. Eye bridge 2���3 rows of facets. Prefrons with 10 setae. Clypeus with no setae. Palpus (Fig. 5B) one-segmented, with 3 setae and a large dorsal patch of sensillae. Antennal flagellomeres (Fig. 5A) asymmetrical and the frontal part larger. Length/width of 4th flagellomere: 2.20���2.39. Neck length/width: 0.50���0.54. Thorax. Anterior pronotum with 5���6 setae. Proepisternum with 2���3 setae. Wings (Fig. 6C). Length 2.74���2.84 mm. Width/length: 0.47���0.49. R1/R: 1.68. c/w: 0.66. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia unmodified without specialized setae. Foretibia spur longer than the tibial width. Hypopygium (Fig. 5C). Gonocoxite slightly longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe- or finger-like projections, without setose area. Gonostylus elongated with a medial lobe, which has 3 megasetae. Apical tooth with a megaseta attached to its ventral side, with 2 subapical megasetae near the tooth, all these megasetae slender and slightly curved. Tegmen wider than long, nearly straight apically, narrowed at middle. Aedeagus with distinct apical fork and basal sclerotization. Distribution. China: Taiwan, Zhejiang. Remarks. The new specimens examined show a slight difference from the type specimen by the apical part of the apical tooth in the Chinese specimens being shorter than in the holotype., Published as part of Chen, Zuluan, Yang, Xue, Menzel, Frank, Wu, Hong & Huang, Junhao, 2021, Three Oriental species of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae) from China, pp. 551-562 in Zootaxa 4969 (3) on pages 558-561, DOI: 10.11646/zootaxa.4969.3.7, http://zenodo.org/record/4751483, {"references":["Vilkamaa, P., Halenius, P. & Sevcik, J. (2019) Review of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae), with the description of twenty-four new species. Zootaxa, 4656 (1), 1 - 42. https: // doi. org / 10.11646 / zootaxa. 4656.1.1"]}

Pseudoaerumnosa tianmushana Chen & Huang sp. n. (Figs. 3C���F, 4, 6B) Specimens examined. Holotype: male. CHINA: Zhejiang province, Lin���an, Tianmu Mountain, Malaise trap, 29.VII.2018, Zuluan Chen (ZAFU) [XWCZ-21]. Paratypes, 15 males, same data as holotype (ZAFU) [XWCZ-3, XWCZ-12, XWCZ-13, XWCZ-14, XWCZ-16, XWCZ-17, XWCZ-18, XWCZ-19, XWCZ-20, XWCZ-23, XWCZ- 31, XWCZ-32, XWCZ-33, XWCZ-37, XWCZ-42]. Description. Male. Brown. Head. Eye bridge with 2���3 rows of facets. Prefrons with 13 setae. Clypeus with no setae. Palpus one-segmented, with 2 setae and a large dorsal patch of sensilla. Antennal flagellomeres (Fig. 4A) asymmetrical with the frontal part larger. Length/width of 4th flagellomere: 1.88���2.09. Neck length/width: 0.93���0.94. Thorax. Anterior pronotum with 4���5 setae. Prothoracic episternum with 2���3 setae. Wings (Fig. 6B). Length 2.36���2.47 mm. Width/length: 0.46���0.48. R1/R: 1.64. c/w: 0.79���0.80. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia (Fig. 4B) unmodified, without specialized setae. Foretibia spur longer than the tibial width. Hypopygium (Figs. 3C���F, 4C). Gonocoxite longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe- or finger-like projections, without setose area. Gonostylus elongated with a medial lobe, which have 3 megasetae.Apical tooth with a long megaseta attached to its ventral side and 2 subapical megasetae near the tooth, the megasetae slender and slightly curved. Tegmen broader than long, smoothly rounded apically and basolaterally sclerotized. Aedeagus with an indistinct apical fork and a basocentral sclerotized central strip. Distribution. China: Zhejiang. Remarks. Pseudoaerumnosa tianmushana sp. n. resembles P. formosa in its elongated gonostylus and tegmen narrowed above the middle, but differs by having an aedeagus with indistinct apical fork and gonostylus medial lobe reaching to apical 1 / 3 of gonostylus, while having an aedeagus with distinct apical fork and gonostylus with medial lobe reaching to apical 1 / 4 of gonostylus in P. formosa. Etymology. This species is named after its type locality, Tianmu Mountain (Chinese: Tianmu-shan) in Zhejiang province, China., Published as part of Chen, Zuluan, Yang, Xue, Menzel, Frank, Wu, Hong & Huang, Junhao, 2021, Three Oriental species of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae) from China, pp. 551-562 in Zootaxa 4969 (3) on page 558, DOI: 10.11646/zootaxa.4969.3.7, http://zenodo.org/record/4751483

Pseudoaerumnosa regularis Chen & Huang sp. n. (Figs. 2, 3A–B, 6A) Specimens examined. Holotype: male. CHINA: Zhejiang province, Lin’an, Tianmu Mountain, Malaise trap, 29.VIII.2018, Zuluan Chen (ZAFU) [XWCZ-10]. Paratypes: CHINA: 12 males, same data as holotype (ZAFU) [XWCZ-2, XWCZ-5, XWCZ-6, XWCZ-8, XWCZ-9, XWCZ-11, XWCZ-25, XWCZ-27, XWCZ-28, XWCZ-29, XWCZ-30, XWCZ-41]. Description. Male. Brown. Head. Eye bridge 2–3 rows of facets. Prefrons with 10 setae. Clypeus with no setae. Palpus one-segmented, with 3 setae and a large dorsal patch of sensillae. Antennal flagellomeres (Fig. 2A) asymmetrical with the frontal part larger. Length/width of 4th flagellomere: 1.50–1.60. Neck length/width: 0.58– 0.59. Thorax. Anterior pronotum with 5–6 setae. Proepisternum with 3–12 setae. Wings (Fig. 6A). Length 2.47 mm. Width/length: 0.55–0.60. R1/R: 2.26. c/w: 0.55–0.85. R1 with dorsal and ventral setae, bM, r-m and stCuA bare. Legs. Foretibia unmodified without specialized setae. Foretibia spur longer than the tibial width (Fig. 2B). Hypopygium (Figs. 2C, 3A–B). Gonocoxite longer than gonostylus. Gonocoxites fused, ventromedial margin distinct with short to moderate setae, ventrolateral surface with long and dense setae, apicomesial corner with 1 or 2 elongated setae. Intergonocoxal area without lobe-like or finger-like production, without setose area. Gonostylus elongated with a medial lobe, which has 3 megasetae. Apical tooth with a long megaseta attached to its ventral side, with 2 subapical megasetae near the tooth, all these megasetae slender and slightly curved. Tegmen wider than long, nearly straight apically, basolaterally sclerotized. Aedeagus with distinct apical fork and basal sclerotization. Distribution. China: Zhejiang. Remarks. Pseudoaerumnosa regularis sp. n. is similar to P. annae in having an elongate gonostylus and the tegmen straight laterally, but differs in having a narrow aedeagus and 3 megasetae on the lobe of gonostylus, while having a broad aedeagus and 4 megasetae on the lobe of gonostylus in P. annae. Etymology. The species is named after its regularly-shaped hypopygium, from the Latin, regularis, meaning regularity.

Bradysia minorlobus Yang, Shi & Huang sp. n. (Fig. 3 A���E) Specimens examined. Holotype: male. CHINA: Zhejiang Province, Linan, Jixiancun, sweep-net, 15.XI.2011, leg. Kai Shi [SM01801] (ZAFU). Paratypes: 1 male, Zhejiang, Linan, Tianmu Mountain, sweep-net, 05.V.2017, leg. Kai Heller [SCILA343-17]; 2 males, Zhejiang, Linan, Taihuyuan, Jinyou Village, sweep-net, 28.IV.2016, leg. Kai Heller [SCILA155-16, SCILA157-16] (ZFMK). Description. Male. Colouration. Head dark brown; antenna, thorax, abdomen and hypopygium brown; palpus (Fig. 3C) and legs yellowish-brown; wing fumose. Head. Eye bridge with 3 rows of facets. Prefrons with 16���19 setae. Clypeus with 1���2 setae. Basal segment of palpus with 2���3 setae; 2 nd segment with 7���8 setae; 3 rd segment with 6���8 setae. Length/width of 4 th flagellomere: 2.34���2.56. Neck length/width: 0.68���0.71 (Fig. 3D). Thorax. Anterior pronotum with 3���6 setae, episternum 1 with 3���4 setae. Wings (Fig. 3C). Length 1.65���2.66 mm, width/length: 0.34��� 0.40. c/w: 0.52���0.62. R1/R: 0.58���1.07. M, Cu, r-m and stM bare. Legs. Foretibia (Fig. 3E) with a comb of 9���10 setae. Length of spur/width of foretibia 1.54���1.83. Length of femur/length of metatarsus: foreleg 1.23���1.31. Length of metatarsus/length of tibia: foreleg 0.56���0.61, hind leg 0.42���0.46. Length of hind tibia/length of thorax 1.63���1.64. Foretibia with 0 dorsal, 0 ventral, 2 prolateral and 2 retrolateral spinose setae. Midtibia without dorsal megasetae. Hypopygium (Fig. 3B). Hypopygium 0.49 times as long as wide. Gonocoxite same length as gonostylus; with one ventral elongated seta at apicomesial corner. Intergonocoxal area with small, slightly bifid, densely setose lobe. Gonostylus (Fig. 3A) elongated, densely setose apically, the apical tooth lacking; 1.97 times as long as wide; with a small subapical dorsal lobe with 5 short megasetae, which form a close group. Tegmen 0.71 times as long as wide, roundish apically; with some aedeagal teeth. Sternite 10 with 1 seta on each half. Female. Unknown. BIN. BOLD:ADD2898. Distribution. China: Zhejiang. Remarks. The morphological characters of the new species and B. chenjinae are compared in Results and Discussion. The new species resembles B. scabricornis Tuomikoski, 1960 and B. atracornea Mohrig and Menzel, 1992, but the intercoxal area of B. minorlobus Yang, Shi & Huang sp. n. has a small slightly bifid integonocoxal lobe, which is lacking in many other species. Etymology. This species is named according to its small intergonocoxal lobe of male genitalia and the specific name is combined from the Latin words ���minor��� and ��� lobus ���, meaning ���small��� and ���lobe���, respectively., Published as part of Yang, Xue, Shi, Kai, Heller, Kai, Menzel, Frank, Huang, Junhao & Wu, Hong, 2019, Morphology and DNA barcodes of two species of Bradysia Winnertz from China (Diptera, Sciaridae), with the description of Bradysia minorlobus Yang, Shi & Huang sp. n., pp. 85-94 in Zootaxa 4612 (1) on pages 90-93, DOI: 10.11646/zootaxa.4612.1.5, http://zenodo.org/record/3233638, {"references":["Tuomikoski, R. (1960) Zur Kenntnis der Sciariden (Dipt.) Finnlands. Annales Zoologici Societatis Zoologicae Botanicae Fennicae ' Vanamo', 21 (4), 1 - 164.","Mohrig, W. & Menzel, F. (1992) Neue Arten europaischer Trauermucken (Diptera, Sciaridae). An International Journal of Dipterological Research, 3, 1 - 16."]}

Bradysia minorlobus Yang, Shi & Huang sp. n. (Fig. 3 A–E) Specimens examined. Holotype: male. CHINA: Zhejiang Province, Linan, Jixiancun, sweep-net, 15.XI.2011, leg. Kai Shi [SM01801] (ZAFU). Paratypes: 1 male, Zhejiang, Linan, Tianmu Mountain, sweep-net, 05.V.2017, leg. Kai Heller [SCILA343-17]; 2 males, Zhejiang, Linan, Taihuyuan, Jinyou Village, sweep-net, 28.IV.2016, leg. Kai Heller [SCILA155-16, SCILA157-16] (ZFMK). Description. Male. Colouration. Head dark brown; antenna, thorax, abdomen and hypopygium brown; palpus (Fig. 3C) and legs yellowish-brown; wing fumose. Head. Eye bridge with 3 rows of facets. Prefrons with 16–19 setae. Clypeus with 1–2 setae. Basal segment of palpus with 2–3 setae; 2 nd segment with 7–8 setae; 3 rd segment with 6–8 setae. Length/width of 4 th flagellomere: 2.34–2.56. Neck length/width: 0.68–0.71 (Fig. 3D). Thorax. Anterior pronotum with 3–6 setae, episternum 1 with 3–4 setae. Wings (Fig. 3C). Length 1.65–2.66 mm, width/length: 0.34– 0.40. c/w: 0.52–0.62. R1/R: 0.58–1.07. M, Cu, r-m and stM bare. Legs. Foretibia (Fig. 3E) with a comb of 9–10 setae. Length of spur/width of foretibia 1.54–1.83. Length of femur/length of metatarsus: foreleg 1.23–1.31. Length of metatarsus/length of tibia: foreleg 0.56–0.61, hind leg 0.42–0.46. Length of hind tibia/length of thorax 1.63–1.64. Foretibia with 0 dorsal, 0 ventral, 2 prolateral and 2 retrolateral spinose setae. Midtibia without dorsal megasetae. Hypopygium (Fig. 3B). Hypopygium 0.49 times as long as wide. Gonocoxite same length as gonostylus; with one ventral elongated seta at apicomesial corner. Intergonocoxal area with small, slightly bifid, densely setose lobe. Gonostylus (Fig. 3A) elongated, densely setose apically, the apical tooth lacking; 1.97 times as long as wide; with a small subapical dorsal lobe with 5 short megasetae, which form a close group. Tegmen 0.71 times as long as wide, roundish apically; with some aedeagal teeth. Sternite 10 with 1 seta on each half. Female. Unknown. BIN. BOLD:ADD2898. Distribution. China: Zhejiang. Remarks. The morphological characters of the new species and B. chenjinae are compared in Results and Discussion. The new species resembles B. scabricornis Tuomikoski, 1960 and B. atracornea Mohrig and Menzel, 1992, but the intercoxal area of B. minorlobus Yang, Shi & Huang sp. n. has a small slightly bifid integonocoxal lobe, which is lacking in many other species. Etymology. This species is named according to its small intergonocoxal lobe of male genitalia and the specific name is combined from the Latin words ‘minor’ and ‘ lobus ’, meaning ‘small’ and ‘lobe’, respectively.

Bradysia chenjinae Yang, Zhang & Yang, 1993 (Fig. 2 A���C) Specimens examined. Holotype: male. CHINA: Guizhou Province, Guiyang, Institute of Biological Research, Guizhou Academy of Sciences, sweep-net, 1.IX.1987, leg. Chikun Yang (ZAFU). Paratypes: CHINA: 3 males, same data as holotype, but 3.IX.1987; 1 male, Guizhou Institute of Forestry Science, sweep-net, 7.VIII.1987, leg. Xuemin Zhang (ZAFU). (Some errors were found in the original description of the type series after checking the labels. The specimen information of the holotype in the original description was exactly the same as here provided, and the information of all the paratypes provided here were different from those in Yang et al. (1993)). Additional material: 6 males, Zhejiang Province, Longwang Mountain, VII.1995; 3 males, same data, but light-trap 20.VIII.1995; 1 male, same data, but IV.1996, leg. Hong Wu; 1 male, same data, but 12.V.1996, leg. Hong Wu; 2 males, Zhejiang Province, Linan, Jixian Village, sweep-net, 15.XII.2011, leg. Kai Shi [SM01799���1800] (The drawings are based on SM01799) (ZAFU); 1 male, Zhejiang, Linan, Taihuyuan, Jinyou Village, sweep-net, 28.IV.2016, leg. Kai Heller [SCILA154-16]; 5 males, Zhejiang, Linan, Banqiao, Mengwuli Village, Malaise trap, 28.V.2016, leg. Kai Heller [SCILA 147-16, SCILA 168-16, SCILA 174-16, SCILA 202-17, SCILA 203-17]; 1 male, Zhejiang, Changxing, Yangwan Village, Malaise trap, 18.V.2016, leg. Kai Shi [SCILA252-17]; 1 male, Zhejiang, Tianmu Mountain, sweep-net, 18.V.2016, leg. Kai Heller [SCILA346-17]; 2 males, Zhejiang, Tianmu Mountain, sweep-net, 05.V.2017, leg. Kai Heller [SCILA347-17, SCILA363-17]; 1 male, Zhejiang, Changxing, Yangwan Village, Malaise trap, 10.XII.2016, leg. Kai Shi [SCILA370-17] (ZFMK). Redescription. Male. Colouration. Head dark brown; antenna, thorax, abdomen, and hypopygium brown; palpus and legs yellowish-brown; wing fumose. Head. Eye bridge with three rows of facets. Prefrons with 19 setae. Clypeus with one seta. Basal segment of palpus with 2 setae; 2 nd segment with 6 setae; 3 rd segment with 6 setae. Length/width of 4 th flagellomere: 2.49. Neck length/width: 0.82���1.26. Thorax. Anterior pronotum with 2 setae, episternum 1 with 4 setae. Wings (Fig. 2C). Length 1.82���1.84 mm, width/length: 0.35���0.37. c/w: 0.63��� 0.71. R1/R: 0.57���0.58. M, Cu, r-m and stM bare. Legs. Foretibia with a comb of 8 setae. Length of spur/width of foretibia 1.62. Length of femur/length of metatarsus: foreleg 1.33. Length of metatarsus/length of tibia: foreleg 0.58, hind leg 0.44. Length of hind tibia/length of thorax 1.62. Foretibia with 0 dorsal, 0 ventral, 2 prolateral and 2 retrolateral spinose setae. Midtibia without dorsal spinose setae. Hypopygium (Fig. 2A). Hypopygium 1.2 times as long as wide. Gonocoxite length close to gonostylus length; with one ventral elongate seta at apicomesial corner; with densely setose ventrally. Intergonocoxal area with large, deeply bifid, densely setose lobe. Gonostylus (Fig. 2B) elongated, 2.08 times as long as wide, the apical tooth lacking. A small subapical dorsal lobe with 5 short megasetae, forming a close group. Tegmen roundish apically, 0.63 times as long as wide; with some aedeagal teeth. Sternite 10 with 1 seta on each half. Female. Unknown. BIN. BOLD:ACO5209. Distribution. China: Guizhou, Zhejiang. Remarks. In the original description of the species, the authors stated that the diagnosis of B. chenjinae was comparable to that of the similar species Bradysia chunmeiae Yang, Zhang & Yang, 1993, without specifying the characters of the two species in detail, while the latter species was characterized by its light colour and antennal characters. After comparing the type specimens of both species, we found that the two species are quite different, in that B. chunmeiae has no distinct subapical dorsal lobe on the gonostyli and a rather short neck of the antenna (neck length/width: 0.22���0.41), whereas B. chenjinae has a distinct subapical dorsal lobe on the gonostyli and a longer neck of the antenna (neck length/width: 0.82���1.26). Etymology. This species was named after Prof. Chenjin Yang for supporting Prof. Chi-kun Yang���s insect investigation in Guizhou Province in 1987., Published as part of Yang, Xue, Shi, Kai, Heller, Kai, Menzel, Frank, Huang, Junhao & Wu, Hong, 2019, Morphology and DNA barcodes of two species of Bradysia Winnertz from China (Diptera, Sciaridae), with the description of Bradysia minorlobus Yang, Shi & Huang sp. n., pp. 85-94 in Zootaxa 4612 (1) on pages 89-90, DOI: 10.11646/zootaxa.4612.1.5, http://zenodo.org/record/3233638, {"references":["Yang, C. K., Zhang, X. M. & Yang, C. Q. (1993) Taxonomy of the Sciaridae from Guizhou province. Entomotaxonomia, 15 (4), 283 - 311."]}

Dichopygina perfecta (Pettey, 1918) (Figs 2–6) Neosciara perfecta Pettey, 1918: 325, 341, figs 30, 61. Type locality: USA, Maryland, Montgomery Co., Plummers Island. Bradysia perfecta (Pettey, 1918): Stone & Laffoon, 1965: 234. Corynoptera perfecta (Pettey, 1918): Steffan, 1966: 49, 54. Dichopygina perfecta (Pettey, 1918): Mohrig et al., 2013: 200–201, fig. 28. Material examined. Canada. 1♂, New Brunswick, Fundy National Park, Devil`s Halfacre Road (45°35'22″N, 64°57' 20″W; elev. 61 m), 21 May 2013, leg. Shirley Butland (CNFDK107-14) (BIOG); 1♂, New Brunswick, Kouchibouguac National Park, Near Park Compound, behind Research House (46°46'15″N, 65°00'23″W; elev. 61m), malaise trap, 26 August 2013, leg. Bernard Martin (CNKOS809-14) (BIOG). Finland. 2♂, Lapland, Rovaniemi, Sorvanulkki, herb-rich, old-growth boreal forest, malaise trap, 28 July 2014, leg. Jukka Salmela (SCILA010-16, SCILA009-16) (BIOG). Diagnosis. The morphological differences between D. perfecta and D. bernhardi are hereby confirmed. The gonostylus of D. perfecta has almost equal 7–9 thin and straight subapical megasetae in one group, whereas D. bernhardi has 10–13 megasetae at the apical half of the gonostylus and the basalmost megasetae longer than others. In addition, the tegmen of D. perfecta is equally rounded, while it is flatter in D. bernhardi. Redescription. Colour. Thorax bright brown; abdomen, hypopygium brown; legs yellow; wing hyaline or slightly darkened. Antenna unicolour and yellowish brown. Head. Eye bridge 2–3 facets wide. Antennal setae fine, dense, shorter than segment width. Length/width of flagellomere 4 of antenna 1.40–1.80; transition of basal part to neck pronounced (Fig. 3). Neck length/segment width 0.30–0.40. Maxillary palpus bright and 3-segmented; basal segment with 1 bristles; 2nd segment short, oval; 3rd segment as long as basal segment; sensillae present. Thorax. Notum brown. Thoracic setae weak, white. Posterior pronotum bare. Mesothoracic sclerites bare. Wings (Fig. 4). Length 1.6–1.9 mm. bM, r-m bare; R 1/ R 0.60–0.80; c/w 0.63–0.72. Membrane without macrotrichia; venation weak, with faint stM; M-fork of normal shape; R 1 ending clearly before base of M-fork. Halter bright. Legs. Foretibia with dense patch of setae and curved margin; claws untoothed. Hind coxa of same colour as femora. Hind tibia 0.80–0.90 mm; Tibial spurs of equal length. Abdomen. Abdominal setae weak, sparse. Tergal setae white; sternal setae white. Hypopygium (Fig. 5). Hypopygium 0.5–0.7 times as long as wide. Base of gonocoxites with normal, weak hairs; gonocoxites narrowly separated; inner margin of gonocoxites U-shaped; inner membrane of hypopygium bare; gonostylus elongate, narrowed and curved (Fig. 2); 1.1–1.5 times longer than wide; inner margin straight, or concave in ventral view; apical part of gonostylus tapered. Apical tooth without internal structure, shorter than subapical megasetae; 1.1–1.7 times longer than broad. Awl-like setae absent. Innerside of gonostylus with 7–9 thin and straight subapical megasetae in one group. Position of basalmost megaseta 55–67% from apex. Whiplash-hair absent. Tegmen 0.6–0.7 times as long as broad; equally rounded; without special structures; central process absent (Fig. 6). Aeadeagal apical structure and teeth absent. Remarks. The species is firstly recorded in Finland and Canada. Distribution. Finland, Canada, USA (Mohrig et al., 2013)., Published as part of Leng, Ruixin, Heller, Kai, Huang, Junhao, Ye, Lan & Wu, Hong, 2018, DNA barcoding of the genus Dichopygina, with a new species from China (Diptera: Sciaridae), pp. 18-26 in Zoological Systematics 43 (1) on pages 23-24, DOI: 10.11865/zs.201802, http://zenodo.org/record/5364735, {"references":["Pettey, F. W. 1918. A revision of the genus Sciara of the family Mycetophilidae (Diptera). Annals of the Entomologcial Society of America, 11: 319 - 341.","Mohrig, W, Heller, K., Hippa, H., Vilkamaa, P., Menzel, F. 2013. Revision of the black fungus gnats (Diptera: Sciaridae) of North America. Studia Dipterologica, 19: 141 - 286."]}

Dichopygina tibetana Leng, Heller & Huang, sp. nov. (Figs. 7–12) Material examined. Holotype, 1♂. China. Tibet, Bomi, Ganjing Guesthouse, light trap, 18 July 2014, leg. Jun Xu / Mei Qin (SM02765) (SCILA001-16) (ZAFU). Paratypes. 5♂, the same data as holotype (SM02766–67, SM02818–20) (SCILA002-16, SCILA003-16, SCILA004-16, SCILA005-16, SCILA006-16) (ZAFU]. Diagnosis. The new species and D. bernhardi are similar to each other in having a rather straight gonostylus with all megasetae at the apical half of the gonostylus and the flatter top of tegmen. However, D. tibetana differs in having larger wings (wing length 1.6–1.8 mm versus 1.4–1.7 mm) and fewer gonostylar megasetae (6–8 versus 10–13). By its number of gonostylar megasetae and basalmost megasetae longer than the others, D. tibetana also resembles D. duplics. However, the new species can be distinguished by its larger wings (wing length 1.6–1.8 mm versus 1.2–1.4 mm) and broader eye bridge (facets wide 4 versus 3). Furthermore, D. tibetana resembles D. perfecta by sharing the very similar arrangement of the gonostylar megasetae, while the later species may easily be recognized by its equally rounded tegmen. Description. Colour. Thorax, abdomen, antennae and hypopygium brown; legs, palpus and wing veins yellowish brown; wings fumose. Antenna unicolour and yellowish brown. Head. Eye bridge 4 facets wide. Antennal setae fine, dense, shorter than segment width (Fig. 8). Length/width of flagellomere 4 of antenna 1.95–2.33; transition of basal part to neck pronounced; neck lengh/segment width 0.30–0.50, unicolour. Maxillary palpus 3-segmented, basal segment with 1 seta; 2nd segment with 6–8 setae; 3rd segment with 5–7 setae; sensillae present (Fig. 10). Thorax. Notum brown. Thoracic setae weak, white. Anterior pronotum with 4–6 setae, episternum 1 with 4–8 setae. Wings (Fig. 7). Wing length 1.60–1.80 mm, width/length 0.43–0.46, R 1/ R 0.64–0.85, c/w 0.49–0.57. Membrane without macrotrichia. Venation weak, with faint stM; M-fork of normal shape; R 1 ending clearly before base of M-fork; bM, r-m bare. Halter bright, short. Legs. Foreleg: foretibia (Fig. 10) with dense patch of setae and curved margin; claws untoothed; length of basitarsomere/length of foretibia 0.55–0.60; length of femur/length of metatarsus 0.56–0.86. Length of metatarsus/length of tibia: foreleg 1.17–1.48, hind leg 0.85–0.97. Length of hind tibia/length of thorax 0.95–1.16. Abdomen. Abdominal setae weak, sparse. Tergal setae white. Sternal setae white. Hypopygium (Figs 9, 12). Hypopygium 0.6–0.8 times as long as wide. Base of gonocoxites with normal, weak hairs; gonocoxites wide and strong, narrowly separated; inner margin of gonocoxites U-shaped; inner membrane of hypopygium bare. Gonostylus elongated, narrowed and curved; 1.0–1.3 times longer than wide; inner margin straight, or concave in ventral view; apex tapered. Apical tooth without internal structure, shorter than subapical megasetae; 1.0–1.3 times longer than broad. Awl-like setae absent. With 6–8 megasetae in the apical half of the gonostylus and the basalmost megasetae longer than the others. Position of basalmost megaseta 42–44% from apex. Whiplash-hair absent. Tegmen 0.70–0.75 times as long as broad, variable in shape, from subtrapezoidal to subtriangular, without sclerotized borders, with central process. Aeadeagal with apical structure and teeth. Distribution. China (Tibet). Etymology. This species is named after the Chinese province of its type locality, Tibet. Funding The work is financially supported by the National Natural Science Foundation of China (31372244), the Science and Technology Department of Zhejiang Province (2013 C32082) and Norwegian Taxonomy Initiative (NTI) (Artsprosjektet, 70184228). Acknowledgements We are grateful to Ms. Valérie Lévesque-Beaudin (Guelph, Canada) for the loan of Canadian Dichopygina specimens and to Mr. Björn Rulik (Bonn, Germany) for making available the sequences of Finnish specimens on BOLD and assistance with the treatment of the Chinese sequences., Published as part of Leng, Ruixin, Heller, Kai, Huang, Junhao, Ye, Lan & Wu, Hong, 2018, DNA barcoding of the genus Dichopygina, with a new species from China (Diptera: Sciaridae), pp. 18-26 in Zoological Systematics 43 (1) on pages 24-26, DOI: 10.11865/zs.201802, http://zenodo.org/record/5364735, {"references":["Pettey, F. W. 1918. A revision of the genus Sciara of the family Mycetophilidae (Diptera). Annals of the Entomologcial Society of America, 11: 319 - 341."]}

Leng, Ruixin, Heller, Kai, Huang, Junhao, Ye, Lan, Wu, Hong (2018): DNA barcoding of the genus Dichopygina, with a new species from China (Diptera: Sciaridae). Zoological Systematics 43 (1): 18-26, DOI: 10.11865/zs.201802

Mohrigia structura Rudzinski, 2006 (Fig. 16) Mohrigia structura Rudzinski, 2006: 458 ���460. Diagnosis. The species is characterized by its long-ovate gonostylus, densely setose on the apicoventral part, with an apical tooth on the dorsal lobe, and only one flagellate seta under the tooth; gonocoxite with a mastoid intercoxal lobe bearing 7 setae, setose on the ventral inner side on the apical third; tegmen pyramidal, parameral apodeme continuously sclerotized forming a central strip on dorsal side and with a finger-like process at the apex; maxillary palps 2-segmented, basal segment with 1 long seta (Rudzinski, 2006). Distribution. China (Taiwan, Fig. 16) (Rudzinski, 2006)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 90-91, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia truncatula Shi & Huang, sp. nov. (Figs 13, 15 C, 16) Material examined. Holotype, male. CHINA. Fujian, Wuyishan, Xingcunzhen, Tongmucun, Longdu, sweep-net, 27-IV-2012, leg. Kai Shi [SM01601]. Paratype, Zhejiang. 1 male, Lishui, Jingning, Yujikeng, sweep-net, 29-VII- 2010, leg. Lu-Jing Yang [SM01566]. Diagnosis. The species is characterized by its ovate gonostylus with a slightly curved apical tooth arising from a well-developed lobe, two to three flagellate setae on the ventral mesial margin of gonostylus; hypopygium with a small conical setose intercoxal lobe; tegmen apically truncate, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Head dark brown; palps, antenna, thorax, abdomen and hypopygium brown; legs yellowish-brown; wings fumose. Head (Figs 13 C, D). Eye bridge with 3 rows of facets. Prefrons with 7 setae. Clypeus with 1 seta. Maxillary palps 3-segmented, basal segment with 1–2 setae. Length/width index of 4th flagellomere: 2.19–2.64. Thorax. Anterior pronotum with 2–4 setae, episternum 1 with 1–2 setae. Wings (Fig. 15 C). Wing length 1.63– 2.03 mm, width/length: 0.47–0.49. c/w: 0.71–0.72. R1/R: 0.62–0.70. M, CuA, stM and r-m nonsetose. Legs (Fig. 13 E). Foretibia with bordered posteroventral patch of modified setae. Length of spur/width of foretibia 1.03–1.11. Length of femur/length of metatarsus: foreleg 1.23–1.29. Length of metatarsus/length of tibia: foreleg 0.58–0.59, hind leg 0.48. Length of hind tibia/length of thorax 1.18–1.44. Claws toothed. Hypopygium (Figs 13 A, B). The gonocoxite as long as the gonostylus. The gonostylus ovate, with densely setose on the apicoventral part, with a slightly curved apical tooth arising from a well-developed lobe, two to three flagellate setae on the ventral mesial margin of gonostylus present. Hypopygium with a small conical setose intercoxal lobe. Apex of Tegmen truncate, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Zhejiang, Fujian, Fig. 16). Remarks. This species is close to M. megalocornuta (Mohrig & Menzel, 1992) in sharing a well-developed dorsal lobe on an ovate gonostylus, and apically truncate tegmen with parameral apodeme continuously sclerotized forming a central strip on dorsal side. The new species differs in having a dorsal lobe arising from the subapical part, while it is arising from the apex in M. megalocronuta. Etymology. This species is named after its apically truncate tegmen, from the Latin adjective truncatulus, meaning truncate.

Mohrigia notolobos Xu & Huang, sp. nov. (Figs 7, 14 D, 16) Material examined. Holotype, male. China, Yunnan. Baoshan, Longyang, Lujiang, Nankang, 24°48.376'N, 98°47.342'E, 1847 m, 9-V-2009, leg. Su-Jiong Zhang [SM00938]. Diagnosis. This species is unique by an apicodorsal lobe bearing three long setae on gonocoxite, its clavate gonostylus, and relatively longer aedeagus. Description (male). Color. Head and thorax dark brown; antennae, palps, abdomen and hypopygium yellowish brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 7 E, F). Eye bridge narrow, with 3 rows of facets, ommatidia reduced medially. Maxillary palps 2- segmented, basal segment with 2 setae; 2nd segment with 5 setae. Length/width index of 4th flagellomere: 2.08. Thorax. Anterior pronotum with 1 seta, episternum 1 with 1 seta. Wings (Fig. 14 D). Wing length 1.85 mm, width/length: 0.39. c/w: 0.84. R1/R: 0.53. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 7 G). Length of spur/width of foretibia 0.98. Length of femur/length of metatarsus: foreleg 1.51. Length of metatarsus/length of tibia: foreleg 0.51, hind leg 0.46. Length of hind tibia/length of thorax 1.22. Claws toothed. Hypopygium (Figs 7 A–D). Gonocoxite as long as gonostylus. Gonostylus slender, densely setose on apicoventral part, with an apical tooth arising from a subapical dorsal lobe. Hypopygium with a conical setose intercoxal lobe, three long setae arising from apicodorsal lobe of gonocoxite. Tegmen slightly and smoothly curved, parameral apodeme laterally sclerotized on tegmen. Aedeagus relatively longer than other Mohrigia species. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Etymology. This species is named after the dorsal lobe on the gonocoxite, a Latinized Greek compound word from the nouns nōton (back) and lobos (lobe). It is an invariable noun in apposition.

Mohrigia insolenta Rudzinski, 2006 (Fig. 16) Mohrigia insolenta Rudzinski, 2006: 461 ���463. Diagnosis. The species is unique in the genus by its unusual split gonostylus and setae on wing veins M and CuA on the dorsal side. However, it resembles the type species M. hippai in having the extremely long and dense flagellate setae on the intercoxal lobe of the hypopygium. By contrast, M. hippai has an apically broadly rounded gonostylus without a lobe on the inner side; and wing veins are bare. Furthermore, M. insolenta has 2-segmented maxillary palps, while it is 3-segmented in M. hippai (Rudzinski, 2006). Distribution. China (Taiwan, Fig. 16) (Rudzinski, 2006)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 81, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia angusta Xu & Huang, sp. nov. (Figs 1, 14 C, 16) Material examined. Holotype, male. China, Shaanxi. Ningshan, Niujiliang Nature Reserve, Gangou, 33��49.5532'N, 108��47.2403'E, 1778 m, 10-VIII-2013, leg. Jun Xu [SM01896]. Diagnosis. The species is characterized by its slender gonostylus a strong tooth arising from a subapical dorsal lobe, with two long flagellate setae on the inner side of the gonostylus; hypopygium without intercoxal lobe; tegmen pyramidal, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a fingerlike process at the apex. Description (male). Color. Head and thorax dark brown; antennae, abdomen, palps and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 1 D, E). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 8 setae; 3rd segment with 6 setae. Length/width index of 4th flagellomere: 3.24���3.52. Thorax. Anterior pronotum with 3 setae, episternum 1 with 5 setae. Wings (Fig. 14 C). Wing length 1.47 mm, width/length: 0.39. c/w: 0.71. R1/R: 0.58. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 1 F). Length of spur/width of foretibia 0.98. Length of femur/length of metatarsus: foreleg 1.51. Length of metatarsus/length of tibia: foreleg 0.51, hind leg 0.46. Length of hind tibia/length of thorax 1.22. Claws toothed. Hypopygium (Figs 1 A���C). The gonocoxite as long as the gonostylus. The gonostylus slender, densely setose on the apicoventral part, with a strong tooth arising from a subapical dorsal lobe, two long flagellate setae on the inner side of the gonostylus. Hypopygium without intercoxal lobe. Tegmen pyramidal, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a finger-like process at the apex. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Shaanxi, Fig. 16). Remarks. This new species is very similar to M. rusticana, Rudzinski, 2006 and M. structura Rudzinski, 2006 in sharing a slender gonostylus with the subapical tooth arising from a dorsal lobe. A distinct difference is observed on the intercoxal lobe, since it is absent in the new species, while the other two species both have a small intercoxal lobe. Additionally, the tegmen is pyramidal in the new species and M. structura, while trapezoidal in M. rusticana. The new species and M. rusticana have 3-segmented palps, while it is 2-segmented in M. structura. Further differences could also be found on the sclerotized parameral apodeme. Etymology. This species is named after its slender gonostylus, from the Latin adjective angustus, meaning long and narrow., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 73, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia scrobiculata Xu & Huang, sp. nov. (Figs 10, 11, 14 E, 16) Material examined. Holotype, male. China, Yunnan. Yingjiang, Tongbiguan, Mulianhuatang, 24°36.004'N, 97°39.139'E, 1341 m, 20-V-2009, leg. Su-Jiong Zhang [SM00711]. Paratype: 1 male, the same data as holotype [SM00678]. Diagnosis. This species is unique in its significantly concaved inner side of the slender gonostylus and the long and strong apical tooth arising from a lobe on the dorsal margin. The species is also characterized by two long flagellate setae on the inner side of gonostylus; tegmen trapezoid, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head and thorax dark brown; antennae, palps, abdomen and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 11 A, B). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 5 setae; 3rd segment with 4–5 setae. Length/width index of 4th flagellomere: 2.22–2.57. Thorax. Anterior pronotum with 3 setae, episternum 1 with 3 setae. Wings (Fig. 14 E). Wing length 1.40–1.44 mm, width/length: 0.38. c/w: 0.63–0.68. R1/R: 0.42–0.50. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 11 C). Length of spur/width of foretibia 0.82–1.03. Length of femur/length of metatarsus: foreleg 1.84. Length of metatarsus/length of tibia: foreleg 0.49–0.51, hind leg 0.42–0.45. Length of hind tibia/length of thorax 0.79– 0.97. Claws toothed. Hypopygium (Figs 10 A–D). The gonocoxite as long as the gonostylus. The gonostylus slender and inner side significantly concaved, densely setose on the apicoventral part, with a long and strong apical tooth arising from a lobe on the dorsal margin of gonostylus, two flagellate setae on the inner side of gonostylus present. Hypopygium without an intercoxal lobe. Tegmen trapezoid, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Remarks. The paratype shares most of the diagnostic characteristics with the holotype, except it has an intercoxal lobe on the ventral base of hypopygium, while the holotype lacks the structure. The intercoxal lobe is usually a very important character in Mohrigia. However, this species is unique in having its significantly concaved inner side of the gonostylus and the long and strong apical tooth arising from a lobe on the dorsal margin, shared by both the type and paratype specimens, that were collected from the same site. Therefore, we treat the two specimens as the same species in this study. If more similar materials examined in the future, the character will have to be evaluated again. Etymology. This species is named after its significantly concaved inner side of gonostylus, from the Latin adjective scrobiculatus, meaning concave.

Mohrigia scrobiculata Xu & Huang, sp. nov. (Figs 10, 11, 14 E, 16) Material examined. Holotype, male. China, Yunnan. Yingjiang, Tongbiguan, Mulianhuatang, 24��36.004'N, 97��39.139'E, 1341 m, 20-V-2009, leg. Su-Jiong Zhang [SM00711]. Paratype: 1 male, the same data as holotype [SM00678]. Diagnosis. This species is unique in its significantly concaved inner side of the slender gonostylus and the long and strong apical tooth arising from a lobe on the dorsal margin. The species is also characterized by two long flagellate setae on the inner side of gonostylus; tegmen trapezoid, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head and thorax dark brown; antennae, palps, abdomen and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 11 A, B). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 5 setae; 3rd segment with 4���5 setae. Length/width index of 4th flagellomere: 2.22���2.57. Thorax. Anterior pronotum with 3 setae, episternum 1 with 3 setae. Wings (Fig. 14 E). Wing length 1.40���1.44 mm, width/length: 0.38. c/w: 0.63���0.68. R1/R: 0.42���0.50. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 11 C). Length of spur/width of foretibia 0.82���1.03. Length of femur/length of metatarsus: foreleg 1.84. Length of metatarsus/length of tibia: foreleg 0.49���0.51, hind leg 0.42���0.45. Length of hind tibia/length of thorax 0.79��� 0.97. Claws toothed. Hypopygium (Figs 10 A���D). The gonocoxite as long as the gonostylus. The gonostylus slender and inner side significantly concaved, densely setose on the apicoventral part, with a long and strong apical tooth arising from a lobe on the dorsal margin of gonostylus, two flagellate setae on the inner side of gonostylus present. Hypopygium without an intercoxal lobe. Tegmen trapezoid, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Remarks. The paratype shares most of the diagnostic characteristics with the holotype, except it has an intercoxal lobe on the ventral base of hypopygium, while the holotype lacks the structure. The intercoxal lobe is usually a very important character in Mohrigia. However, this species is unique in having its significantly concaved inner side of the gonostylus and the long and strong apical tooth arising from a lobe on the dorsal margin, shared by both the type and paratype specimens, that were collected from the same site. Therefore, we treat the two specimens as the same species in this study. If more similar materials examined in the future, the character will have to be evaluated again. Etymology. This species is named after its significantly concaved inner side of gonostylus, from the Latin adjective scrobiculatus, meaning concave., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 90, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157

Mohrigia inflata Shi & Huang, sp. nov. (Figs 4, 15 B, 16) Lycoriella longirostris Yang, Zhang & Yang, 1995: 210 and 211 (misidentification, in part). Material examined. Holotype, male. CHINA. Hubei, Mts. Shennongjia, Yinyuhe, sweep-net, 16-V-2012, leg. Kai Shi [SM01637]. Paratypes, Zhejiang. Kaihua, Mt. Gutianshan: 2 males, 17-VII-1992, leg. Xue-Xin Chen [SM10009, SM10011]; 1 male, yellow pan trap, 21-VII-1992, leg. Hong Wu [SM10010] (all males labeled as paratypes of Lycoriella longirostris, Yang, Zhang & Yang, 1995). 1 male, Anji, Mt. Longwangshan, sweep-net, 1- V-2012, leg. Kai Shi [SM01605]. Diagnosis. T he species is characterized by its ovate gonostylus with a strong apical tooth arising from a strong dorsal lobe, three flagellate setae on the ventral mesial margin of gonostylus; hypopygium with a small conical setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side and laterally sclerotized. Description (male). Head dark brown; antenna, thorax, abdomen and hypopygium brown; palps pale brown; legs yellowish-brown; wings fumose. Head (Figs 4 C, D). Eye bridge with 3 rows of facets. Prefrons with 15 setae. Clypeus with 1���3 setae. Maxillary palps 3-segmented, basal segment with 2 setae. Length/width index of 4th flagellomere: 2.60���3.54. Thorax. Anterior pronotum with 3���6 setae, episternum 1 with 5���7 setae. Wings (Fig. 15 B). Wing length 2.12��� 2.32 mm, width/length: 0.42���0.47. c/w: 0.61���0.72. R1/R: 0.70���0.80. M, stM and CuA nonsetose. r-m with 0���4 setae. Legs (Fig. 4 E). Length of spur/width of foretibia 0.87���1.10. Length of femur/length of metatarsus: foreleg 1.17���1.48. Length of metatarsus/length of tibia: foreleg 0.56���0.64, hind leg 0.48���0.54. Length of hind tibia/length of thorax 1.14���1.39. Claws toothed. Hypopygium (Figs 4 A, B). The gonocoxite as long as the gonostylus. The gonostylus ovate, densely setose on the apicoventral part, with a strong apical tooth arising from a strong dorsal lobe, three flagellate setae on the ventral mesial margin of gonostylus present. Hypopygium with a small conical setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side and laterally sclerotized. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Zhejiang, Hubei, Fig. 16). Remarks. Three paratypes of Lycoriella longirostris, are here treated as M. inflata sp. nov. The argument for the misidentification is discussed above in the remarks of M. megalocornuta (Mohrig & Menzel, 1992). The new species is similar to M. globulosa (Figs 3 A���C) in both having a much inflated gonostylus, but the gonostylus is relatively ovate in this specis, and nearly spherical in M. globulosa. Difference is also seen in the shape of the tegmen, which is smoothly curved in M. inflata, while triangular in M. globulosa. Etymology. This species is named after its inflated gonostylus, from the Latin adjective inflatus, meaning inflated., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 78-81, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Mohrig, W., Menzel, F. & Kozanek, M. (1992) Neue Trauermucken (Diptera, Sciaridae) aus Nord-Korea und Japan. Dipterological Research, 3, 17 - 32."]}

Mohrigia retarda Rudzinski, 2006 (Fig. 16) Mohrigia retarda Rudzinski, 2006: 452 ���453. Diagnosis. The species is characterized by its swollen gonostylus with the strong apical tooth located on a dorsal lobe, 2���3 flagellate setae on inner side of the gonostylus; gonocoxite with a mastoid intercoxal lobe that bears 2���4 setae; tegmen trapezoidal and apically truncate, parameral apodeme strongly sclerotized on lateral sides to apical margin, and also forming a central strip; maxillary palps 3-segmented, basal segment with 1 seta (Rudzinski, 2006). Distribution. China (Taiwan, Fig. 16) (Rudzinski, 2006)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 87, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia composivera Rudzinski, 2006 (Fig. 16) Mohrigia composivera Rudzinski, 2006: 457 ���458. Material examined. China, Yunnan. 1 male, Lijiang, Yulong Jokul Nature Reserve, Yunshanping, 27��09'54"N, 100��15'59"E; 3002.7 m, 12-VII-2014, leg. Jun Xu and Mei Qin [SM02342]. Tibet. 2 males, Motuo, Zhamo road, K18, 29��44'19''N, 95��40'41''E; 3286 m, 21-VII-2014, leg. Jun Xu [SM02523, SM02526]; 2 males, Linzhi, Bomi, G318, Modai, 29��42'33"N, 96��09'48"E, 3122 m, 18-VII-2014, leg. Jun Xu and Mei Qin [SM02422-02423]. Shaanxi. 2 males, Huxian, Laoyu, Baliping Zhuque Nature Park, sweep-net, 13-VII-2012, leg. Junhao Huang [SM01659���01660]. Fujian. 1 male, Wuyishan, Xingcun, Tongmu, Longdu, 27��44'55"N, 117��40'37"E, 558���598 m, 3-V-2014, leg. Jun Xu [SM02137]. Diagnosis. The species is characterized by its thick-ovate gonostylus with an apical tooth arising from an indistinct dorsal lobe, two to three flagellate setae on the inner side of the gonostylus; gonocoxite with conical intercoxal lobe bearing about 10 setae; tegmen pyramidal, parameral apodeme continuously sclerotized, forming a central strip on dorsal side and a finger-like process at the apex; ventral inner side of gonocoxite with long setae; maxillary palps 3-segmented, basal segment with 1 long seta (Rudzinski, 2006). Distribution. China (Fujian, Yunnan, Tibet, Shaanxi, Taiwan; Fig. 16) (Rudzinski, 2006). Remarks. This species was first described from Taiwan and found distributed widely in south China. All the continental specimens match the diagnostic characteristics but slight difference is found in the body size: wing length = 2.10 mm in the type specimen, whereas 1.78 mm in the male from Fujian, and 2.93 mm in the specimen from Yunnan., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 74, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia truncatula Shi & Huang, sp. nov. (Figs 13, 15 C, 16) Material examined. Holotype, male. CHINA. Fujian, Wuyishan, Xingcunzhen, Tongmucun, Longdu, sweep-net, 27-IV-2012, leg. Kai Shi [SM01601]. Paratype, Zhejiang. 1 male, Lishui, Jingning, Yujikeng, sweep-net, 29-VII- 2010, leg. Lu-Jing Yang [SM01566]. Diagnosis. The species is characterized by its ovate gonostylus with a slightly curved apical tooth arising from a well-developed lobe, two to three flagellate setae on the ventral mesial margin of gonostylus; hypopygium with a small conical setose intercoxal lobe; tegmen apically truncate, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Head dark brown; palps, antenna, thorax, abdomen and hypopygium brown; legs yellowish-brown; wings fumose. Head (Figs 13 C, D). Eye bridge with 3 rows of facets. Prefrons with 7 setae. Clypeus with 1 seta. Maxillary palps 3-segmented, basal segment with 1���2 setae. Length/width index of 4th flagellomere: 2.19���2.64. Thorax. Anterior pronotum with 2���4 setae, episternum 1 with 1���2 setae. Wings (Fig. 15 C). Wing length 1.63��� 2.03 mm, width/length: 0.47���0.49. c/w: 0.71���0.72. R1/R: 0.62���0.70. M, CuA, stM and r-m nonsetose. Legs (Fig. 13 E). Foretibia with bordered posteroventral patch of modified setae. Length of spur/width of foretibia 1.03���1.11. Length of femur/length of metatarsus: foreleg 1.23���1.29. Length of metatarsus/length of tibia: foreleg 0.58���0.59, hind leg 0.48. Length of hind tibia/length of thorax 1.18���1.44. Claws toothed. Hypopygium (Figs 13 A, B). The gonocoxite as long as the gonostylus. The gonostylus ovate, with densely setose on the apicoventral part, with a slightly curved apical tooth arising from a well-developed lobe, two to three flagellate setae on the ventral mesial margin of gonostylus present. Hypopygium with a small conical setose intercoxal lobe. Apex of Tegmen truncate, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Zhejiang, Fujian, Fig. 16). Remarks. This species is close to M. megalocornuta (Mohrig & Menzel, 1992) in sharing a well-developed dorsal lobe on an ovate gonostylus, and apically truncate tegmen with parameral apodeme continuously sclerotized forming a central strip on dorsal side. The new species differs in having a dorsal lobe arising from the subapical part, while it is arising from the apex in M. megalocronuta. Etymology. This species is named after its apically truncate tegmen, from the Latin adjective truncatulus, meaning truncate., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 95-96, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Mohrig, W., Menzel, F. & Kozanek, M. (1992) Neue Trauermucken (Diptera, Sciaridae) aus Nord-Korea und Japan. Dipterological Research, 3, 17 - 32."]}

Mohrigia subrhynchophysa Xu & Huang, sp. nov. (Figs 12, 16) Material examined. Holotype, male. China, Yunnan. Baoshan, Nankang, Gaoligong Nature Reserve, 24°49.729'N, 98°46.074'E, 2181 m, 11-V-2009, leg. Man-Man Wang [SM00887]. Diagnosis. The species is characterized by having a large coracoid dorsal lobe on gonostylus, dorsal mesial margin expanded, and two long flagellate setae on the inner side; gonocoxite with a small setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side; maxillary palps 3-segmented, basal segment with 1 seta. Description (male). Color. Head, thorax and abdomen dark brown; antennae and hypopygium yellowishbrown; mesonotum brown, palps and legs pale brown; wings fumose. Head (Figs 12 C, D). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 8 setae; 3rd segment with 6 setae. Length/width index of 4th flagellomere: 3.73. Thorax. Anterior pronotum with 4 setae, episternum 1 with 5 setae. Wings (partly broken). R1/R: 0.86. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 12 E). Length of spur/width of foretibia 0.97. Length of femur/length of metatarsus: foreleg 1.30. Length of metatarsus/length of tibia: foreleg 0.62, hind leg 0.51. Length of hind tibia/length of thorax 1.26. Claws toothed. Hypopygium (Figs 12 A, B). The gonocoxite as long as the gonostylus. Gonostylus densely setose on the apicoventral part, with the apical tooth arising from a large coracoid dorsal lobe, dorsal mesial margin expanded, two long flagellate setae on the inner side. Hypopygium with a small setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Remarks. The species is similar to M. rhynchophysa in sharing a large coracoid dorsal lobe on the gonostylus. But it differs in having a small setose intercoxal lobe and 3-segmented maxillary palps, while M. rhynchophysa bearing an oblong intercoxal lobe and 2-segmented maxillary palps. In addition, the parameral apodeme is continuously sclerotized forming a central strip on the dorsal side in the new species, while it is laterally sclerotized on the tegmen in M. rhynchophysa. Etymology. This species is named after its similarity to M. rhynchophysa (Yang, Zhang & Yang, 1993).

Mohrigia orthacantha (Yang, Zhang & Yang, 1993), new combination (Figs 8, 14 F, 16) Lycoriella orthacantha Yang, Zhang & Yang, 1993: 661 (type locality: Mt. Longqishan, Fujian). Material examined. Neotype, male (here designated). China, Fujian, Wuyishan, Xingcun, Pikenkou-Shaxizhou, 27��44'55"N, 117��40'37"E; 597 m, 1-V-2014, leg. Mei Qin [SM02297]. 1 male, Inner Mongolia, Chifeng, Saihanwula Nature Reserve, Zhenggou, 44��13.198'N, 118��44.177'E, 1231 m, 23-VII-2013, leg. Mei Qin [SM01963]. Diagnosis. This species is characterized by its kidney-shaped gonostylus with two long flagellate setae on the ventral side; gonocoxite with a small setose intercoxal lobe; tegmen rounded, parameral apodeme completely sclerotized forming a central strip in dorsal view. Description (male). Color. Head, antennae and thorax dark brown; palps, abdomen and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 8 C, D). Eye bridge with 3 rows of facets. Prefrons with 16���17 setae. Maxillary palps 3-segmented, basal segment with 2 setae; 2nd segment with 5���6 setae; 3rd segment with 4���5 setae. Length/width index of 4th flagellomere: 3.08. Thorax. Anterior pronotum with 1���2 setae, episternum 1 with 2���7 setae. Wings (Fig. 14 F). Wing length 1.94 mm, width/length: 0.41. c/w: 0.68. R1/R: 0.67���0.79. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 8 E). Length of spur/width of foretibia 0.89���0.98. Length of femur/length of metatarsus: foreleg 1.22���1.38. Length of metatarsus/length of tibia: foreleg 0.60���0.64, hind leg 0.48���0.68. Length of hind tibia/length of thorax 1.10��� 1.31. Claws toothed. Hypopygium (Figs 8 A, B). The gonocoxite as long as the gonostylus. The gonostylus kidney-shaped, with ventral and dorsal margin, basally expanded and forming an angulate corner, densely setose on the apicoventral part, a subapical tooth arising from the dorsal margin, two long flagellate setae on the ventral side of gonostylus. Hypopygium with a small setose intercoxal lobe. Tegmen rounded, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Sternite 10 with 1 seta on each half. Distribution. China (Fujian, Inner Mongolia, Fig. 16). Remarks. The species was described in the genus Lycoriella, with illustrations of the 4th flagellar segment of antenna, maxillary palp, apex of foretibia and terminalia. The authors provided the detailed measurements of the body and its different parts, without special attention to its terminalia structures, which indicate its affiliation with Mohrigia. It is unique in the genus in having a kidney-shaped gonostylus with a ventral and dorsal margin basally expanded, and its rounded tegmen. However, the holotype and paratype of this species could be not found in the ZAFU collection, in which the type should have been deposited. After careful investigation, we hereby confirm that the type material is lost. The type locality is Mt. Longqishan, which is in the southeastern part of the Wuyishan mountainous area in the Fujian province of China. During a recent field survey about 200 kilometers north of this mountainous area, a male was collected in Xingcun that matches all the descriptions and illustrations provided by Yang et al. (1993). Here we designate this specimen as the neotype of the species. The diagnosis, redescription and new illustrations (Figs 8 and 14 F) are given based on the neotype., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 86, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Yang, C. - K., Zhang, X. - M. & Yang, C. - Q. (1993) Diptera: Sciaridae. In: Huang, C. (Ed.), Animals of Longqi Mountain, Beijing, China Forestry Publishing press, pp. 654 - 664."]}

Mohrigia notolobos Xu & Huang, sp. nov. (Figs 7, 14 D, 16) Material examined. Holotype, male. China, Yunnan. Baoshan, Longyang, Lujiang, Nankang, 24��48.376'N, 98��47.342'E, 1847 m, 9-V-2009, leg. Su-Jiong Zhang [SM00938]. Diagnosis. This species is unique by an apicodorsal lobe bearing three long setae on gonocoxite, its clavate gonostylus, and relatively longer aedeagus. Description (male). Color. Head and thorax dark brown; antennae, palps, abdomen and hypopygium yellowish brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 7 E, F). Eye bridge narrow, with 3 rows of facets, ommatidia reduced medially. Maxillary palps 2- segmented, basal segment with 2 setae; 2nd segment with 5 setae. Length/width index of 4th flagellomere: 2.08. Thorax. Anterior pronotum with 1 seta, episternum 1 with 1 seta. Wings (Fig. 14 D). Wing length 1.85 mm, width/length: 0.39. c/w: 0.84. R1/R: 0.53. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 7 G). Length of spur/width of foretibia 0.98. Length of femur/length of metatarsus: foreleg 1.51. Length of metatarsus/length of tibia: foreleg 0.51, hind leg 0.46. Length of hind tibia/length of thorax 1.22. Claws toothed. Hypopygium (Figs 7 A���D). Gonocoxite as long as gonostylus. Gonostylus slender, densely setose on apicoventral part, with an apical tooth arising from a subapical dorsal lobe. Hypopygium with a conical setose intercoxal lobe, three long setae arising from apicodorsal lobe of gonocoxite. Tegmen slightly and smoothly curved, parameral apodeme laterally sclerotized on tegmen. Aedeagus relatively longer than other Mohrigia species. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Etymology. This species is named after the dorsal lobe on the gonocoxite, a Latinized Greek compound word from the nouns nōton (back) and lobos (lobe). It is an invariable noun in apposition., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 84, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157

Mohrigia infernosa Rudzinski, 2006 (Fig. 16) Mohrigia infernosa Rudzinski, 2006: 450 ���452. Material examined. China, Fujian. 1 male, Wuyishan, Xingcun, Xianfengling, 27��43'30"N, 117��39'36"E, 2-V- 2014, leg. Mei Qin [SM02454]. Diagnosis. The species is characterized by its oblong-ovate gonostylus with the inner side significantly concaved, with the apical tooth located on a dorsal lobe, and one flagellate seta under the apical tooth; gonocoxite with a big long-tongue-shaped intercoxal lobe that is apically setose, ventral inner side of gonocoxite bearing long setae; tegmen oblong-pyramidal, parameral apodeme laterally sclerotized on dorsal side; maxillary palps 2- segmented, basal segment with 3���4 setae (Rudzinski, 2006). Distribution. China (Fujian, Taiwan; Fig. 16) (Rudzinski, 2006). Remarks. This species was first described from Taiwan, China. The Fujian specimen differs by having the tooth arise from the apical fourth of gonostylus, not from the apex as in the type specimen illustrated by Rudzinski (2006). Furthermore, the new material bears very dense setae on the lateral side of the intercoxal lobe, while it is nearly bare in the type. In addition, slight differences are found in the measurements: length/width index of 4th flagellomere = 2.5, wing length = 1.93 mm and R1/R = 0.75 in the type specimen; whereas length/width index of 4th flagellomere = 3.21, wing length = 2.83 mm and R1/R = 1.27 in the Fujian specimen. Despite the above differences, we do not consider the new material as a new species due to their common and unique prolonged intercoxal lobe and laterally sclerotized parameral apodeme., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 78, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia cirricoxalis Rudzinski, 2006 (Fig. 16) Mohrigia cirricoxalis Rudzinski, 2006: 453 ���455. Diagnosis. The species is characterized by its long and dense flagellate setae on the ventral inner side of the gonocoxite that lacks of an intercoxal lobe; tegmen trapezoidal and maxillary palps 2-segmented. The species is closely related to M. retarda Rudzinski, 2006 and M. megalocornuta (Mohrig & Menzel, 1992) by having an ovate gonostylus with a strong apical tooth arising from dorsal lobe, and its trapezoid tegmen. The species differs in having long and dense flagellate setae on the ventral inner side of the gonocoxite, 2-segmented maxillary palps and lacks of an intercoxal lobe. Distribution. China (Taiwan, Fig. 16) (Rudzinski, 2006)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 73-74, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476.","Mohrig, W., Menzel, F. & Kozanek, M. (1992) Neue Trauermucken (Diptera, Sciaridae) aus Nord-Korea und Japan. Dipterological Research, 3, 17 - 32."]}

Mohrigia cylindrata Xu & Huang, sp. nov. (Figs 2, 14 A, 16) Material examined. Holotype, male. China, Tibet. Motuo, Lage, 29��31'2"N, 94��52'52"E, 3193 m, 23-VII-2014, leg. Jun Xu [SM02406]. Paratypes: 1 male, the same data as holotype [SM02407]; 1 male, Motuo, Zhamo road, K18, 29��44'19''N, 95��40'41''E, 3286 m, 21-VII-2014, leg. Jun Xu [SM02527]. Yunnan. 1 male, Baoshan, Nankang, Gaoligong Nature Reserve, 24��49.729'N, 98��46.074'E, 2181 m, 11-V-2009, leg. Su-jiong Zhang [SM00899]. Diagnosis. The species is characterized by its thick-ovate gonostylus with the dorsal mesial margin expanded, with an apical tooth arising from a subcylindrical dorsal lobe, two long flagellate setae on the inner side of the gonostylus; hypopygium with a small setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head and antennae dark brown; abdomen, hypopygium and thorax yellowishbrown; mesonotum brown; palps and legs pale brown; wings fumose. Head (Figs 2 C, D). Eye bridge with 2 rows of facets. Prefrons with 15 setae. Maxillary palps 3-segmented, basal segment with 2 setae; 2nd segment with 6���8 setae; 3rd segment with 5���8 setae. Length/width index of 4th flagellomere: 3.67���3.93. Thorax. Anterior pronotum with 3���5 setae, episternum 1 with 5���7 setae. Wings (Fig. 1 4 A). Wing length 2.24���2.64 mm, width/length: 0.35���0.37. c/w: 0.72���0.77. R1/R: 0.69���0.92. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 2 E). Length of spur/width of foretibia 0.96���1.10. Length of femur/length of metatarsus: foreleg 1.42���1.63. Length of metatarsus/length of tibia: foreleg 0.55���0.58, hind leg 0.47���0.53. Length of hind tibia/ length of thorax 1.20���1.34. Claws toothed. Hypopygium (Figs 2 A, B). The gonocoxite as long as the gonostylus. The gonostylus thick-ovate, densely setose on the apicoventral part, dorsal mesial margin expanded, with an apical tooth arising from a subcylindrical dorsal lobe, two long flagellate setae on the inner side of the gonostylus. Hypopygium with a small setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Tibet, Fig. 16). Remarks. The species is close to M. composivera Rudzinski, 2006 as they share an oval gonostylus and a small setose intercoxal lobe. But the new species differs in having a subcylindrical dorsal lobe, whereas the dorsal lobe is indistinct in M. composivera. In addition, M. composivera has a finger-like process at the apex of the central strip, while the new species does not have this structure. Etymology. This species is named after the subcylindrical dorsal lobe of the gonostylus, from the Latin adjective cylindratus, meaning cylindrical., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 74-77, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia cylindrata Xu & Huang, sp. nov. (Figs 2, 14 A, 16) Material examined. Holotype, male. China, Tibet. Motuo, Lage, 29°31'2"N, 94°52'52"E, 3193 m, 23-VII-2014, leg. Jun Xu [SM02406]. Paratypes: 1 male, the same data as holotype [SM02407]; 1 male, Motuo, Zhamo road, K18, 29°44'19''N, 95°40'41''E, 3286 m, 21-VII-2014, leg. Jun Xu [SM02527]. Yunnan. 1 male, Baoshan, Nankang, Gaoligong Nature Reserve, 24°49.729'N, 98°46.074'E, 2181 m, 11-V-2009, leg. Su-jiong Zhang [SM00899]. Diagnosis. The species is characterized by its thick-ovate gonostylus with the dorsal mesial margin expanded, with an apical tooth arising from a subcylindrical dorsal lobe, two long flagellate setae on the inner side of the gonostylus; hypopygium with a small setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head and antennae dark brown; abdomen, hypopygium and thorax yellowishbrown; mesonotum brown; palps and legs pale brown; wings fumose. Head (Figs 2 C, D). Eye bridge with 2 rows of facets. Prefrons with 15 setae. Maxillary palps 3-segmented, basal segment with 2 setae; 2nd segment with 6–8 setae; 3rd segment with 5–8 setae. Length/width index of 4th flagellomere: 3.67–3.93. Thorax. Anterior pronotum with 3–5 setae, episternum 1 with 5–7 setae. Wings (Fig. 1 4 A). Wing length 2.24–2.64 mm, width/length: 0.35–0.37. c/w: 0.72–0.77. R1/R: 0.69–0.92. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 2 E). Length of spur/width of foretibia 0.96–1.10. Length of femur/length of metatarsus: foreleg 1.42–1.63. Length of metatarsus/length of tibia: foreleg 0.55–0.58, hind leg 0.47–0.53. Length of hind tibia/ length of thorax 1.20–1.34. Claws toothed. Hypopygium (Figs 2 A, B). The gonocoxite as long as the gonostylus. The gonostylus thick-ovate, densely setose on the apicoventral part, dorsal mesial margin expanded, with an apical tooth arising from a subcylindrical dorsal lobe, two long flagellate setae on the inner side of the gonostylus. Hypopygium with a small setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Tibet, Fig. 16). Remarks. The species is close to M. composivera Rudzinski, 2006 as they share an oval gonostylus and a small setose intercoxal lobe. But the new species differs in having a subcylindrical dorsal lobe, whereas the dorsal lobe is indistinct in M. composivera. In addition, M. composivera has a finger-like process at the apex of the central strip, while the new species does not have this structure. Etymology. This species is named after the subcylindrical dorsal lobe of the gonostylus, from the Latin adjective cylindratus, meaning cylindrical.

Mohrigia megalocornuta (Mohrig & Menzel, 1992) (Figs 5, 6, 15D, 16) Lycoriella megalocornuta Mohrig & Menzel, 1992: 24 ���25. Lycoriella longirostris Yang, Zhang & Yang, 1995: 210 and 211 (syn. nov.). Mohrigia megalocornuta (Mohrig & Menzel, 1992), Menzel & Mohrig,2000: 414 ���420; Rudzinski, 2006: 460 ���461. Material examined. Zhejiang. Linan, Mt. Tianmushan: 2 males, sweep-net, 3-VII-2010, leg. Kai Shi [SM01570��� 01571]; 1 male, Kaishanlaodian-Xianrending, sweep-net, 3-VII-2010, leg. Lu-Jing Yang [SM01580]. Linan, Qingliangfeng: 1 male, sweep-net, 6���8-VIII-2008, leg. Guang-Yue Wang [SM00232]; 1 male, Mt. Longtangshan, malaise trap, 5-V-2012 [SM01756]. 1 male, Anji, Mt. Longwangshan, sweep-net, 1-V-2012, leg. Kai Shi [SM01606]. 1 male, Lishui, Jingning, Yujikeng, sweep-net, 29-VII-2010, leg. Lu-Jing Yang [SM01567]. Fujian. Wuyishan, Xingcunzhen, Tongmucun, Longdu: 1 male, sweep-net, 27-IV-2012, leg. Kai Shi [SM01603]; 1 male, sweep-net, 27-IV-2012, leg. Lu-Jing Yang [SM01597]; 1 male, sweep-net, 3-V-2014, leg. Mei Qin [SM02132]; Wuyishan, Xingcunzhen, Tongmucun, Xianfengling, Liaowangtai: 2 males, sweep-net, 2-V-2014, leg. Jun Xu [SM02169���2170]; Jianyang, Huangkeng, Lijiapo: 1 male, sweep-net, 6-V-2014, leg. Lan Ye [SM02148]; Jiangle, Mt. Longqishan: 1 male, Shaxizi, sweep-net, 13-VIII-2012, leg. Kai Shi [SM01759]; 1 male, Longtan, sweep-net, 14-VIII-2 0 12, leg. Kai Shi [SM01745]; 1 male, Kengwei, sweep-net, 15-VIII-2 0 12, leg. Jun Xu [SM01787]. Taiwan. 1 male, Xuanlan, Fushan, Botanical Garden, 670 m, sweep-net, 10-VI-2010, leg. Xiao-Yan Liu [SM01586]. The holotype and a portion of the paratypes of Lycoriella longirostris Yang, Zhang & Yang, 1995, here treated as a junior synonym of M. megalocornuta: 1 male, Zhejiang, Kaihua, Mt. Gutianshan, yellow pan trap, 21-VII- 1992, leg. Hong Wu [SM10001] (holotype of Lycoriella longirostris). 3 males, Zhejiang, Kaihua, Mt. Gutianshan, yellow pan trap, 21-VII-1992, leg. Hong Wu [SM10003, SM10006���10007]; 1 male, 50 m, yellow pan trap, 21-VII- 1992, leg. Hong Wu [SM10002]; 1 male, yellow pan trap, 17-VII-1992, leg. Hong Wu [SM10004]; 2 males, 400 m, yellow pan trap, 25.VII.1992, Hong Wu [SM10005, SM10008] (7 paratypes of Lycoriella longirostris). Diagnosis. The species is characterized by its ovate gonostylus with a strong apical tooth arising from a welldeveloped dorsal lobe, two to three flagellate setae on the ventral mesial margin of gonostylus; hypopygium with a small conical setose intercoxal lobe; tegmen nearly rectangular and apically truncate. Description (male). Head dark brown; antenna, thorax and hypopygium brown; palps, legs and abdomen pale brown; wings fumose. Head (Figs 5D, 6A). Eye bridge with 2���3 rows of facets. Prefrons with 10���15 setae. Clypeus with 1���2 setae. Maxillary palps 3-segmented, basal segment with 1 seta. Length/width index of 4th flagellomere: 2.26���2.93. Thorax. Anterior pronotum with 3���7 setae, episternum 1 with 2���7 setae. Wings (Fig. 15 D). Wing length 1.59��� 2.24 mm, width/length: 0.43���0.48. c/w: 0.62���0.77. R1/R: 0.53���0.80. M, CuA and stM nonsetose. r-m with 0���3 setae. Legs (Fig. 5B). Length of spur/width of foretibia 0.86���1.17. Length of femur/length of metatarsus: foreleg 1.18���1.38. Length of metatarsus/length of tibia: foreleg 0.57���0.63, hind leg 0.46���0.66. Length of hind tibia/length of thorax 1.17���1.42. Claws toothed. Hypopygium (Figs 5A���C). The gonocoxite as long as the gonostylus. The gonostylus ovate, densely setose on the apicoventral part, with a strong apical tooth arising from a well-developed dorsal lobe, two to three flagellate setae on the ventral mesial margin of gonostylus. Hypopygium with a small conical setose intercoxal lobe. Tegmen nearly rectangular and apically truncate. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Zhejiang, Fujian, Taiwan, Fig. 16); Japan (Mohrig & Menzel, 1992). Remarks. The nearly rectangular and apically truncate tegmen is an important characteristic of M. megalocornuta. However, Mohrig et al. (1992) and Menzel & Mohrig (2000) gave neither detailed description nor an illustration for the sclerotized tegmen. Rudzinski (2006) recorded the species from Taiwan, providing detailed description of the sclerotized parameral apodeme in tegmen forming in a rim, as well as clear illustration. In this study, a long series of materials are recorded from various localities in continental China. Yang et al. (1995) described Lycoriella longirostris, which was characterized by the maxillary palps 3- segmented, the foretibia bordered with a posteroventral patch of modified setae, and a long apical tooth on the gonostylus, based on the holotype and 15 paratypes from Mt. Gutianshan, Zhejiang, China. Among this material, the holotype and ten paratypes were now found deposited in the ZAFU collection, and the rest of paratypes were not located. After further observation of this material, the parameral apodeme of tegmen are found continuously sclerotized and forming a central strip on the dorsal side, which is a diagnostic character of the genus Mohrigia Menzel. Moreover, Lycoriella is characterized by having one flagellate seta on the medial side of the gonostylus, but two to six flagellate setae are found in all of these type specimens. Therefore, we determined that this material should belong to the genus Mohrigia. However, after careful examination of the above eleven specimens, we confirmed the holotype and seven paratypes belong to M. megalocornuta (Mohrig & Menzel, 1992) (= Lycoriella longirostris, Yang, Zhang & Yang, 1995, new synonym) (Figs 5A���C), in having the diagnostic characteristics that gonostylus ovate and with the strong apical tooth arising from a well-developed dorsal lobe, and the tegmen is nearly rectangular and apically truncate. But, the remaining three paratypes [SM10009���10011] are quite different in having an inflated gonostylus deeply bilobed and a much larger intercoxal lobe on the ventral base of hypopygium. During our study on Chinese Sciaridae, another two specimens from Hubei and Zhejiang provinces were found to be similar to the three specimens in their external morphology and structures of hypopygium. Based on the common differential characteristics, we described them below as M. inflata sp. nov. (Figs 4 A, B)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 81-82, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Mohrig, W., Menzel, F. & Kozanek, M. (1992) Neue Trauermucken (Diptera, Sciaridae) aus Nord-Korea und Japan. Dipterological Research, 3, 17 - 32.","Menzel, F. & Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement 6 (1999), 1 - 761.","Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Mohrigia subrhynchophysa Xu & Huang, sp. nov. (Figs 12, 16) Material examined. Holotype, male. China, Yunnan. Baoshan, Nankang, Gaoligong Nature Reserve, 24��49.729'N, 98��46.074'E, 2181 m, 11-V-2009, leg. Man-Man Wang [SM00887]. Diagnosis. The species is characterized by having a large coracoid dorsal lobe on gonostylus, dorsal mesial margin expanded, and two long flagellate setae on the inner side; gonocoxite with a small setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side; maxillary palps 3-segmented, basal segment with 1 seta. Description (male). Color. Head, thorax and abdomen dark brown; antennae and hypopygium yellowishbrown; mesonotum brown, palps and legs pale brown; wings fumose. Head (Figs 12 C, D). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 8 setae; 3rd segment with 6 setae. Length/width index of 4th flagellomere: 3.73. Thorax. Anterior pronotum with 4 setae, episternum 1 with 5 setae. Wings (partly broken). R1/R: 0.86. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 12 E). Length of spur/width of foretibia 0.97. Length of femur/length of metatarsus: foreleg 1.30. Length of metatarsus/length of tibia: foreleg 0.62, hind leg 0.51. Length of hind tibia/length of thorax 1.26. Claws toothed. Hypopygium (Figs 12 A, B). The gonocoxite as long as the gonostylus. Gonostylus densely setose on the apicoventral part, with the apical tooth arising from a large coracoid dorsal lobe, dorsal mesial margin expanded, two long flagellate setae on the inner side. Hypopygium with a small setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Yunnan, Fig. 16). Remarks. The species is similar to M. rhynchophysa in sharing a large coracoid dorsal lobe on the gonostylus. But it differs in having a small setose intercoxal lobe and 3-segmented maxillary palps, while M. rhynchophysa bearing an oblong intercoxal lobe and 2-segmented maxillary palps. In addition, the parameral apodeme is continuously sclerotized forming a central strip on the dorsal side in the new species, while it is laterally sclerotized on the tegmen in M. rhynchophysa. Etymology. This species is named after its similarity to M. rhynchophysa (Yang, Zhang & Yang, 1993)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 92, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Yang, C. - K., Zhang, X. - M. & Yang, C. - Q. (1993) Diptera: Sciaridae. In: Huang, C. (Ed.), Animals of Longqi Mountain, Beijing, China Forestry Publishing press, pp. 654 - 664."]}

Mohrigia globulosa Xu & Huang, sp. nov. (Figs 3, 14 B, 16) Material examined. Holotype, male. China, Tibet. Motuo, Lage, 29��31'2"N, 94��52'52"E, 3193.5 m, 23-VII-2014, leg. Jun Xu [SM02405]. Paratype: 1 male, same data as holotype [SM02404]. Diagnosis. The species is characterized by its approximately spherical gonostylus with the dorsal mesial margin weakly expanded, an apical tooth arising from a small dorsal lobe; gonocoxite with long setae on the ventral side; hypopygium with a conical setose intercoxal lobe; tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a finger-like process at the apex. Description (male). Color. Head, antennae, thorax and abdomen dark brown; palps and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 3 D, E). Eye bridge with 3 rows of facets. Prefrons with 12���18 setae. Maxillary palps 3-segmented, basal segment with 1���2 setae; 2nd segment with 7���8 setae; 3rd segment with 6���7 setae. Length/width index of 4th flagellomere: 3.79���3.90. Thorax. Anterior pronotum with 4���5 setae, episternum 1 with 5���8 setae. Wings (Fig. 14 B). Wing length 2.83��� 2.95 mm, width/length: 0.37. c/w: 0.78���0.80. R1/R: 0.69���0.81. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 3 F). Length of spur/width of foretibia 0.99���1.02. Length of femur/length of metatarsus: foreleg 1.39��� 1.42. Length of metatarsus/length of tibia: foreleg 0.55���0.57, hind leg 0.46���0.47. Length of hind tibia/length of thorax 1.26���1.31. Claws toothed. Hypopygium (Figs 3 A���C). The gonostylus as long as the gonocoxite. The gonostylus approximately spherical, densely setose on the apicoventral part, dorsal mesial margin weakly expanded, with an apical tooth arising from a small dorsal lobe, three long flagellate setae on the inner side of the gonostylus present. Ventral side of the gonocoxite with long setae. Hypopygium with a conical intercoxal lobe. Tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a finger-like process at the apex. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Tibet, Fig. 16). Remarks. Based on the greatly inflated gonostylus, the new species is similar to M. cylindrata (Figs 2 A, B). However, the new species can be diagnosed in having its gonostylus more inflated and nearly spherical, and the dorsal medial margin of gonostylus less expanded, while M. cylindrata has the gonostylus ovate and the dorsal medial margin strongly expanded. Furthermore, the tegmen of the new species is triangular and a finger-like process exists at the apex of the central strip, while it is evenly curved towards the top and without the finger-like structure in M. cylindrata. Etymology. This species is named after its nearly spherical gonostylus, from the Latin adjective globulosus, meaning spherical., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 77, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157

Mohrigia ovoidea Xu & Huang, sp. nov. (Figs 9, 15 A, 16) Material examined. Holotype, male. China, Fujian. Wuyishan, Xingcun, Xianfengling, 27��43'30"N, 117��39'36"E, 2-V-2014, leg. Mei Qin [SM02450]. Diagnosis. The species is characterized by its ovate gonostylus with a strong apical tooth arising from a dorsal lobe, with two flagellate setae on the ventral mesial margin of gonostylus; hypopygium without an intercoxal lobe, only bearing three slightly long setae on the intercoxal area; tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head, antennae and thorax dark brown; palps, abdomen and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 9 D, E). Eye bridge with 3 rows of facets. Prefrons with 6 setae. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 6 setae; 3rd segment with 5 setae. Length/width index of 4th flagellomere: 3.74. Thorax. Anterior pronotum with 2 setae, episternum 1 with 3 setae. Wings (Fig. 15 A). Wing length 1.92 mm, width/length: 0.38. c/w: 0.71. R1/R: 0.48. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 9 F). Length of spur/width of foretibia 1.00. Length of femur/length of metatarsus: foreleg 1.57. Length of metatarsus/length of tibia: foreleg 0.52, hind leg 0.35. Length of hind tibia/length of thorax 1.01. Claws toothed. Hypopygium (Figs 9 A���C). The gonocoxite as long as the gonostylus. The gonostylus ovate, densely setose on the apicoventral part, with a strong apical tooth arising from a dorsal lobe, two flagellate setae on the ventral mesial margin of gonostylus present. Intercoxal lobe of hypopygium absent, only bearing three slightly long setae on the intercoxal area. Tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Fujian, Fig. 16). Remarks. The new species resembles M. angusta (Figs 1 A, B) in having an ovate gonostylus with the subapical the tooth arising from a dorsal lobe, and the intercoxal lobe absent. The difference is mainly in the form of their ovate gonostylus, which is twice as long as wide in the new species, while M. angusta has a slender gonostylus that is about three times as long as wide. Etymology. This species is named after its ovate gonostylus, from the Latin adjective ovoideus, meaning ovate., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 86-87, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157

Mohrigia inflata Shi & Huang, sp. nov. (Figs 4, 15 B, 16) Lycoriella longirostris Yang, Zhang & Yang, 1995: 210 and 211 (misidentification, in part). Material examined. Holotype, male. CHINA. Hubei, Mts. Shennongjia, Yinyuhe, sweep-net, 16-V-2012, leg. Kai Shi [SM01637]. Paratypes, Zhejiang. Kaihua, Mt. Gutianshan: 2 males, 17-VII-1992, leg. Xue-Xin Chen [SM10009, SM10011]; 1 male, yellow pan trap, 21-VII-1992, leg. Hong Wu [SM10010] (all males labeled as paratypes of Lycoriella longirostris, Yang, Zhang & Yang, 1995). 1 male, Anji, Mt. Longwangshan, sweep-net, 1- V-2012, leg. Kai Shi [SM01605]. Diagnosis. T he species is characterized by its ovate gonostylus with a strong apical tooth arising from a strong dorsal lobe, three flagellate setae on the ventral mesial margin of gonostylus; hypopygium with a small conical setose intercoxal lobe; tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side and laterally sclerotized. Description (male). Head dark brown; antenna, thorax, abdomen and hypopygium brown; palps pale brown; legs yellowish-brown; wings fumose. Head (Figs 4 C, D). Eye bridge with 3 rows of facets. Prefrons with 15 setae. Clypeus with 1–3 setae. Maxillary palps 3-segmented, basal segment with 2 setae. Length/width index of 4th flagellomere: 2.60–3.54. Thorax. Anterior pronotum with 3–6 setae, episternum 1 with 5–7 setae. Wings (Fig. 15 B). Wing length 2.12– 2.32 mm, width/length: 0.42–0.47. c/w: 0.61–0.72. R1/R: 0.70–0.80. M, stM and CuA nonsetose. r-m with 0–4 setae. Legs (Fig. 4 E). Length of spur/width of foretibia 0.87–1.10. Length of femur/length of metatarsus: foreleg 1.17–1.48. Length of metatarsus/length of tibia: foreleg 0.56–0.64, hind leg 0.48–0.54. Length of hind tibia/length of thorax 1.14–1.39. Claws toothed. Hypopygium (Figs 4 A, B). The gonocoxite as long as the gonostylus. The gonostylus ovate, densely setose on the apicoventral part, with a strong apical tooth arising from a strong dorsal lobe, three flagellate setae on the ventral mesial margin of gonostylus present. Hypopygium with a small conical setose intercoxal lobe. Tegmen slightly and smoothly curved, parameral apodeme continuously sclerotized forming a central strip on dorsal side and laterally sclerotized. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Zhejiang, Hubei, Fig. 16). Remarks. Three paratypes of Lycoriella longirostris, are here treated as M. inflata sp. nov. The argument for the misidentification is discussed above in the remarks of M. megalocornuta (Mohrig & Menzel, 1992). The new species is similar to M. globulosa (Figs 3 A–C) in both having a much inflated gonostylus, but the gonostylus is relatively ovate in this specis, and nearly spherical in M. globulosa. Difference is also seen in the shape of the tegmen, which is smoothly curved in M. inflata, while triangular in M. globulosa. Etymology. This species is named after its inflated gonostylus, from the Latin adjective inflatus, meaning inflated.

Mohrigia hippai Menzel, 1995 (Fig. 16) Mohrigia hippai Menzel, 1995: 102 ���104; Menzel & Mohrig, 2000: 414 ���420. Material examined. China, Tibet. 1 male, Nielamu, Zhangmu, G 318, K5368, 27��57'27''N, 85��58'18''E, 2845 m, 6- VIII-2014, leg. Jun Xu and Mei Qin [SM02488]. Diagnosis. The species is characterized by its broadly clavate gonostylus, with an apical tooth on the dorsal side, and two to three flagellate setae on the apical half; gonocoxite with a wide intercoxal lobe bearing long and dense setae on the ventral side; tegmen longer than wide and apically rounded, parameral apodeme continuously sclerotized forming a central strip on dorsal side; ventral inner side of gonocoxite with long setae; R1 significantly longer than R and extended to the base of the M-fork; eye bridge narrow with perpendicular frontal process and the ommatidia lost at the center (Menzel & Mohrig, 2000). Distribution. China (Tibet��� new record; Fig. 16); Nepal (Simbhanjang), Burma (Kambaiti) (Menzel & Martens, 1995; Menzel & Mohrig, 2000). Remarks. This species was first recorded from Nepal and is new to China. The Chinese specimen shares the diagnostic characteristics, but slight differences were found (a broader gonocoxite and a narrower tegmen)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 77-78, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Menzel, F. & Martens, J. (1995) Die Sciaridae (Diptera, Nematocera) des Nepal-Himalaya. Teil I. Die blutenbesuchenden Trauermucken an Aronstabgewachsen der Gattung Arisaema (Araceae JUSS.). Studia dipterologica, 2 (1), 97 - 129.","Menzel, F. & Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement 6 (1999), 1 - 761."]}

Mohrigia angusta Xu & Huang, sp. nov. (Figs 1, 14 C, 16) Material examined. Holotype, male. China, Shaanxi. Ningshan, Niujiliang Nature Reserve, Gangou, 33°49.5532'N, 108°47.2403'E, 1778 m, 10-VIII-2013, leg. Jun Xu [SM01896]. Diagnosis. The species is characterized by its slender gonostylus a strong tooth arising from a subapical dorsal lobe, with two long flagellate setae on the inner side of the gonostylus; hypopygium without intercoxal lobe; tegmen pyramidal, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a fingerlike process at the apex. Description (male). Color. Head and thorax dark brown; antennae, abdomen, palps and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 1 D, E). Eye bridge with 3 rows of facets. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 8 setae; 3rd segment with 6 setae. Length/width index of 4th flagellomere: 3.24–3.52. Thorax. Anterior pronotum with 3 setae, episternum 1 with 5 setae. Wings (Fig. 14 C). Wing length 1.47 mm, width/length: 0.39. c/w: 0.71. R1/R: 0.58. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 1 F). Length of spur/width of foretibia 0.98. Length of femur/length of metatarsus: foreleg 1.51. Length of metatarsus/length of tibia: foreleg 0.51, hind leg 0.46. Length of hind tibia/length of thorax 1.22. Claws toothed. Hypopygium (Figs 1 A–C). The gonocoxite as long as the gonostylus. The gonostylus slender, densely setose on the apicoventral part, with a strong tooth arising from a subapical dorsal lobe, two long flagellate setae on the inner side of the gonostylus. Hypopygium without intercoxal lobe. Tegmen pyramidal, parameral apodeme continuously sclerotized forming a central strip on dorsal side and a finger-like process at the apex. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Shaanxi, Fig. 16). Remarks. This new species is very similar to M. rusticana, Rudzinski, 2006 and M. structura Rudzinski, 2006 in sharing a slender gonostylus with the subapical tooth arising from a dorsal lobe. A distinct difference is observed on the intercoxal lobe, since it is absent in the new species, while the other two species both have a small intercoxal lobe. Additionally, the tegmen is pyramidal in the new species and M. structura, while trapezoidal in M. rusticana. The new species and M. rusticana have 3-segmented palps, while it is 2-segmented in M. structura. Further differences could also be found on the sclerotized parameral apodeme. Etymology. This species is named after its slender gonostylus, from the Latin adjective angustus, meaning long and narrow.

Mohrigia rhynchophysa (Yang, Zhang & Yang, 1993), new combination Plastosciara rhynchophysa Yang, Zhang & Yang, 1993: 662 (type locality: Mt. Longqishan, Fujian); Zhang et al., 2003: 123. Diagnosis. The species is characterized by the apical tooth arising from a large coracoid dorsal lobe on the gonostylus, densely setose on the apicoventral part of gonostylus, inner side of gonostylus slightly concaved, bearing several flagellate setae; parameral apodeme laterally sclerotized on tegmen; gonocoxite with an oblong intercoxal lobe; maxillary palps 2-segmented, basal segment with 2 setae. Distribution. China (Fujian, Fig. 16) (Yang et al, 1993). Remarks. The species was described as Plastosciara rhynchophysa by Yang et al. (1993), although the authors considered it a ���strange��� member of the genus due to its unique structures on the hypopygium, slender antennal 4th flagellomere, and an arcuate patch of dense straight setae on subapical foretibia. The diagnosis above is given based on the original description and illustration, since we could not locate the type specimen in ZAFU collection, where the type should have been deposited. Yang et al. (1993) gave no detailed description, but the illustration shows the laterally sclerotized parameral apodeme, and several flagellate setae on the inner side of the gonostylus. Based on the generic characteristics of Mohrigia, P. rhynchophysa Yang, Zhang & Yang, 1993 should be transferred to the genus., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 87, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Yang, C. - K., Zhang, X. - M. & Yang, C. - Q. (1993) Diptera: Sciaridae. In: Huang, C. (Ed.), Animals of Longqi Mountain, Beijing, China Forestry Publishing press, pp. 654 - 664."]}

Mohrigia rusticana Rudzinski, 2006 (Fig. 16) Mohrigia rusticana Rudzinski, 2006: 455 ���456. Diagnosis. The species is characterized by its long-ovate gonostylus, densely setose on the apicoventral part, with an apical tooth on the dorsal lobe, 1���2 flagellate setae on the apical half of the gonostylus; gonocoxite with a conical intercoxal lobe that is densely setose on apical half; tegmen large and trapezoidal, parameral apodeme strongly sclerotized in two lateral sides and the apical margin; maxillary palps 3-segmented, basal segment with 1 long seta (Rudzinski, 2006). Distribution. China (Taiwan, Fig. 16) (Rudzinski, 2006)., Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on page 90, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476."]}

Key to the species of Mohrigia (males) 1. Maxillary palps 2-segmented (Fig. 7 E).....................................................................2 - Maxillary palps 3-segmented (Figs 8 D, 12C)................................................................7 2. Hypopygium with an unusually split gonostylus, with a wide intercoxal lobe; wing veins M and CuA with macrotrichia dor- sally............................................................. M. insolenta Rudzinski, 2006: Taiwan (ORI) - Hypopygium with an ovate or elongated oval gonostylus, with intercoxal lobe narrow or absent; wing veins M and CuA with- out macrotrichia (Fig. 14 D)..............................................................................3 3. Intercoxal lobe absent............................................. M. cirricoxalis Rudzinski, 2006: Taiwan (ORI) - Intercoxal lobe present (Fig. 7 D).........................................................................4 4. Gonostylus with a small intercoxal lobe (Fig. 7 D)............................................................5 - Gonostylus with an oblong intercoxal lobe..................................................................6 5. Gonocoxite with an apicodorsal lobe bearing three long setae, tegmen slightly and smoothly curved, parameral apodeme later- ally sclerotized on tegmen (Figs 7 C���D)....................................... M. notolobos sp. nov.: Yunnan (PAL) - Gonocoxite without apicodorsal lobe, tegmen pyramidal-tiara shape, parameral apodeme continuously sclerotized forming a central strip on dorsal side............................................ M. structura Rudzinski, 2006: Taiwan (ORI) 6. Dorsal lobe of gonostylus large and coracoid, arising from strongly inflated dorsal margin of gonostylus...................................................................... M. rhynchophysa (Yang, Zhang & Yang, 1993): Fujian (ORI) - Dorsal lobe of gonostylus relatively small, and gonostylus not inflated in dorsal margin.......................................................................................... M. infernosa Rudzinski, 2006: Fujian, Taiwan (ORI) 7. Intercoxal lobe on the ventral side of genitalia broad, and tegmen hidden by the long and dense setae on the intercoxal lobe............................................................ M. hippai Menzel, 1995: Tibet; Nepal, Burma (ORI) - Intercoxal lobe on the ventral side of genitalia narrow or absent, and tegmen never hidden by setae on the intercoxal lobe... 8 8. Hypopygium with a rounded tegmen, the gonostylus kidney-shaped (Figs 8 A���B)....................................................................... M. orthacantha (Yang, Zhang & Yang, 1993): Fujian, Inner Mongolia (PAL, ORI) - Tegmen of hypopygium not rounded, the gonostylus not kidney-shaped (Figs 10 B, 12B).............................9 9. Dorsal lobe of gonostylus large and coracoid, arising from strongly inflated dorsal margin of gonostylus (Fig. 12 A).......................................................................... M. subrhynchophysa sp. nov.: Yunnan (ORI) - Dorsal lobe of gonostylus relatively small, and gonostylus not inflated on dorsal margin (Figs 9 A, 10A)................ 10 10. Inner side of gonostylus significantly concaved with a long and strong apical tooth arising from its dorsal medial margin (Figs 10 A, D).............................................................. M. scrobiculata sp. nov.: Yunnan (ORI) - Inner side of gonostylus not significantly concaved (Figs 1 A, 9A).............................................. 11 11. Inner side of gonostylus inflated or slightly inflated (Figs 2 A, 9A)..............................................12 - Inner side of gonostylus not inflated, slender (Fig. 1 A).......................................................19 12. Intercoxal lobe absent (Fig. 9 C)................................................ M. ovoidea sp. nov.: Fujian (ORI) - Intercoxal lobe present (Figs 2 B, 3C, 4A, 5A, 13A).........................................................13 13. Gonostylus with an apical tooth arising from a well-developed dorsal lobe (Figs 2 A, 5A���B).........................14 - Gonostylus with an apical tooth arising from a relatively small dorsal lobe (Figs 3 C, 4A)............................16 14. Tegmen slightly and smoothly curved, dorsal lobe of gonostylus subcylindrical (Figs 2 A���B)........................................................................................... M. cylindrata sp. nov.: Yunnan, Tibet (ORI *) - Tegmen nearly rectangular and apically truncate, dorsal lobe of gonostylus large (Figs 5A���B, 13A���B).................15 15. Dorsal lobe arising just medially of the apex of gonostylus (Figs 5A���C)............................................................................ M. megalocornuta (Mohrig & Menzel, 1992): Zhejiang, Fujian, Taiwan; Japan (ORI) - Dorsal lobe arising from the subapical part of gonostylus (Figs 13 A���B)...... M. truncatula sp. nov.: Zhejiang, Fujian (ORI) 16. Gonostylus greatly inflated, approximately spherical (Figs 3 A���B, 4A���B)....................................... 17 - Gonostylus slightly inflated, elongate oval (Fig. 1 A).........................................................19 17. Gonostylus with two dorsal lobe, tegmen trapezoid................... M. hylotoma Menzel & Mohrig, 1999: Nepal (ORI) - Gonostylus only with one dorsal lobe, tegmen not trapezoid...................................................18 18. Tegmen triangular (Figs 3 A���C).............................................. M. globulosa sp. nov.: Tibet (ORI *) - Tegmen evenly curved apically (Figs 4 A, B).............................. M. inflata sp. nov.: Zhejiang, Hubei (PAL*) 19. Tegmen pyramidal................. M. composivera Rudzinski, 2006: Fujian, Yunnan, Tibet, Shaanxi, Taiwan (PAL, ORI) - Tegmen trapezoid.................................................... M. retarda Rudzinski, 2006: Taiwan (ORI) 20. Intercoxal lobe on ventral side of genitalia absent, tegmen pyramidal (Fig. 1 C)......... M. angusta sp. nov.: Shaanxi (PAL) - Intercoxal lobe on ventral side of genitalia present, tegmen large and trapezoidal..................................................................................................... M. rusticana Rudzinski, 2006: Taiwan (ORI), Published as part of Xu, Jun, Shi, Kai, Huang, Junhao & Wu, Hong, 2017, Review of the genus Mohrigia Menzel (Diptera, Sciaridae) from China, pp. 71-98 in Zootaxa 4300 (1) on pages 72-73, DOI: 10.11646/zootaxa.4300.1.4, http://zenodo.org/record/837157, {"references":["Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 29 (23), 449 - 476.","Yang, C. - K., Zhang, X. - M. & Yang, C. - Q. (1993) Diptera: Sciaridae. In: Huang, C. (Ed.), Animals of Longqi Mountain, Beijing, China Forestry Publishing press, pp. 654 - 664.","Menzel, F. & Martens, J. (1995) Die Sciaridae (Diptera, Nematocera) des Nepal-Himalaya. Teil I. Die blutenbesuchenden Trauermucken an Aronstabgewachsen der Gattung Arisaema (Araceae JUSS.). Studia dipterologica, 2 (1), 97 - 129.","Mohrig, W., Menzel, F. & Kozanek, M. (1992) Neue Trauermucken (Diptera, Sciaridae) aus Nord-Korea und Japan. Dipterological Research, 3, 17 - 32."]}

Mohrigia ovoidea Xu & Huang, sp. nov. (Figs 9, 15 A, 16) Material examined. Holotype, male. China, Fujian. Wuyishan, Xingcun, Xianfengling, 27°43'30"N, 117°39'36"E, 2-V-2014, leg. Mei Qin [SM02450]. Diagnosis. The species is characterized by its ovate gonostylus with a strong apical tooth arising from a dorsal lobe, with two flagellate setae on the ventral mesial margin of gonostylus; hypopygium without an intercoxal lobe, only bearing three slightly long setae on the intercoxal area; tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Description (male). Color. Head, antennae and thorax dark brown; palps, abdomen and hypopygium yellowish-brown; mesonotum brown; legs pale brown; wings fumose. Head (Figs 9 D, E). Eye bridge with 3 rows of facets. Prefrons with 6 setae. Maxillary palps 3-segmented, basal segment with 1 seta; 2nd segment with 6 setae; 3rd segment with 5 setae. Length/width index of 4th flagellomere: 3.74. Thorax. Anterior pronotum with 2 setae, episternum 1 with 3 setae. Wings (Fig. 15 A). Wing length 1.92 mm, width/length: 0.38. c/w: 0.71. R1/R: 0.48. r-m, stM, M1 and M2 bare, CuA1 and CuA2 bare. Legs (Fig. 9 F). Length of spur/width of foretibia 1.00. Length of femur/length of metatarsus: foreleg 1.57. Length of metatarsus/length of tibia: foreleg 0.52, hind leg 0.35. Length of hind tibia/length of thorax 1.01. Claws toothed. Hypopygium (Figs 9 A–C). The gonocoxite as long as the gonostylus. The gonostylus ovate, densely setose on the apicoventral part, with a strong apical tooth arising from a dorsal lobe, two flagellate setae on the ventral mesial margin of gonostylus present. Intercoxal lobe of hypopygium absent, only bearing three slightly long setae on the intercoxal area. Tegmen triangular, parameral apodeme continuously sclerotized forming a central strip on dorsal side. Aedeagus relatively short. Sternite 10 with 1 seta on each half. Distribution. China (Fujian, Fig. 16). Remarks. The new species resembles M. angusta (Figs 1 A, B) in having an ovate gonostylus with the subapical the tooth arising from a dorsal lobe, and the intercoxal lobe absent. The difference is mainly in the form of their ovate gonostylus, which is twice as long as wide in the new species, while M. angusta has a slender gonostylus that is about three times as long as wide. Etymology. This species is named after its ovate gonostylus, from the Latin adjective ovoideus, meaning ovate.

Proscleropterus rudis Colonnelli, Qin & Huang, sp. nov. (Figs 27 ���40, 44) Type Specimens. Holotype. ♂ (IZCAS), " China: Guizhou, Zunyi, Suiyang-xian, Kuankuoshui, Xiangguangshan-cun, 4 -VI- 2010, leg. Z. Wang, IOZ(E) 1803073 ". Paratypes. 1 ♀, same data as holotype but "IOZ(E) 1803619 " (IZCAS); Guizhou, Zunyi, Suiyang-xian, Kuankuoshui, Jinzi-cun, 1 ♀, 3 -VI- 2010, leg. Z. Wang, IOZ(E) 1802495 (IZCAS); Guizhou, Zunyi, Suiyang-xian, Kuankuoshui, Gongtong-gou, 1 ♀, 3 -VI- 2010, leg. Z. Wang, IOZ(E) 1803089 (IZCAS). Description. Male (holotype). LB 2.36 mm; LR 1.03 mm; WP 0.87 mm; LP 0.77 mm; WE 1.44 mm; LE 1.54 mm. Habitus as shown in Figures 27���29. Vestiture. Ventrites (Fig. 30) I and II clothed with short and apically pinnate white scales on disc, and with oval to lanceolate white scales mingled with sparse brown clavate scales on sides; ventrites III and IV with a row of sparse oval scales; ventrite V bearing dense hair-like brown scales on median concavity, and white oval and clavate scales on sides. Head (Figs 9���10). Rostrum slender, 1.38 times as long as pronotum, 4.78 times as long as wide at apex, evenly curved; subparallel-sided from base to antennal insertion, then moderately and gradually widened towards apex, where it is 1.15 times as wide as base. Antennae inserted at 0.44 length of rostrum from apex; scape moderate in length, evidently clavate, round and fringed with 2���3 short setae at apex; scape 1.15 times as long as the funicle, length ratio of funicular segments I: II: III: IV: V: VI = 2.48: 0.79: 0.73: 1.04: 1.00: 1.25 and width ratio = 1.37: 1.08: 1.00: 1.12: 1.37: 1.67. Pronotum (Figs 27, 29) 1.13 times as wide as long, 0.50 times as long and 0.61 times as wide as elytra. Pronotum with lateral sides subparallel on basal half, strongly convergent at the level of the median tubercles and then subparallel on apical 2 / 5. Elytra (Figs 27���29) oval, 1.07 times as long as wide, 2.00 times as long as and 1.65 times as wide as pronotum, with strongly prominent humeri, with large sharp granules at base of interval VIII; widest at basal fifth, then gradually convergent towards apex. Striae with punctures separated by a distance about 2���3 times longer than their diameter. Interval I with a row of sparse granules on apical 3 / 4; interval II with 2���3 sparse granules on basal fifth and with some weakly convex granules on middle; interval III with 3 weakly convex granules near base and two moderately large oblong or round tubercles composed of 3���4 merged granules basad and apicad of middle; interval IV narrow, without tubercles, only bearing a few sharp granules along its entire length; interval V with rather large short 5 -capitate tubercle near base, another small tubercle composed of 3 merged granules and a convex granule basad and apicad of middle on one elytron, the other bearing only a small tubercle composed of 2 merged granules, and a round 7 -capitate tubercle on apical fourth; interval VI with sculpture similar to that of IV; interval VII with a few large sharp granules scattered along its basal half, and a weak smooth elongate tubercle composed of 2���3 merged smaller granules apicad of middle; interval VIII with large tubercle at base, then a weak smooth elongate one composed of 2���3 merged smaller granules apicad of middle, rest of interval VIII with simple, irregular granules; interval IX with a tubercle composed of 4 merged granules on basal third, and a weak smooth elongate one composed of 2���4 merged smaller granules apicad of middle, rest of interval IX and lateral intervals with simple, irregular granules. Underside. Ventrites (Fig. 30) with coarse and moderately dense punctures; length ratio of ventrites I: II: III: IV: V = 5.03: 2.53: 1.04: 1.00: 2.67 and width ratio = 1.88: 1.55: 1.25: 1.16: 1.00. Pygidium (Fig. 31) moderately transverse, about 1.24 times as wide as long, coarsely and moderately punctured and with fine median carina. Terminalia and genitalia. Penis (Figs 32���33) broad, relatively thick in lateral profile; sides weakly widening from base to apical sixth, then strongly convergent apically; apical projection (Fig. 34) blunt, rounded at apex. Endophallus (Fig. 32) with a pair of plate-like sclerites in the apical part, numerous rounded spicules in the median part, and a pair of tubular structures at base. Sternite VIII broken during dissection; spiculum gastrale (Fig. 36) robust, bent leftward, and far exceeding in length the aedeagal body or its apodeme. Tegmen (Fig. 35) with apodeme slender, nearly as long as the diameter of the tegminal ring, weakly widened at apex. Otherwise as in P. shennongjianus Qin & Huang, sp. nov. Female (n = 3). LB 2.46���2.74 mm (mean, 2.63 mm). LR 1.20���1.28 mm (mean, 1.25 mm). WP 0.87���0.91 mm (mean, 0.89 mm). LP 0.81���0.89 mm (mean, 0.84 mm). WE 1.56���1.68 mm (mean, 1.60 mm). LE 1.46���1.69 mm (mean, 1.59 mm). Rostrum (Figs 11���12) slightly slenderer than that of male, 1.45���1.57 times as long as pronotum, about 1.21 times as long as in males. Antennae inserted just basad of middle of rostrum. Pronotum 0.98���1.11 times as wide as long. Elytra 0.94���1.03 times as long as wide. Tibiae simple, not mucronate. Ventrites I and II moderately inflated, sparsely punctured. Ventrite V with shallow median concavity. Pygidium smaller than that of male, 1.22 times wider than long. Terminalia and genitalia. Tergite VIII (Fig. 37) with pair of combs of dense, long setae along apical margin. Sternite VIII (Fig. 38) with a pair of patches of several minute setae near apex; arms slender and apically fuse, nearly 0.4 times as long as apodeme, nearly half as long as coxite and stylus combined. Coxites (Fig. 39) robust, subdivided into two pieces, nearly 5.5 times as long as styli; styli apicolaterally inserted, moderate in length, nearly 2.2 times as long as wide. Spermatheca (Fig. 40) C-shaped, collum slightly convex; ramus indistinct, outline almost uniformly continuous; cornu slender, strongly curved and attenuate. Otherwise as in male. Etymology. The species name is a Latin word ��� rudis ��� meaning ���coarse���, to indicate its rougly sculptured elytra. Distribution. China (Guizhou; Fig. 44). Biological notes. Plant association unknown. Remarks. The new species differs from P. shennongjianus Qin & Huang, sp. nov. in having plumper body, with wider pronotum subparallel-sided on apical 2 / 5, oval elytra with sides gradually convergent towards apex, whereas P. shennongjianus Qin & Huang, sp. nov. has more elongate body, pronotum subparallel-sided on apical half, subcordate elytra with sides evenly convergent towards apex. Moreover, the two species have different distribution of tubercles on elytral intervals., Published as part of Mei Qin, Junhao Huang, Enzo Colonnelli, Runzhi Zhang & Hong Wu, 2016, Two new species and a key to species of the genus Proscleropterus Korotyaev (Coleoptera: Curculionidae) from China, pp. 173-185 in Zoological Systematics 41 (2) on pages 180-185, DOI: 10.11865/zs.201616, http://zenodo.org/record/270328