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Genus Pteronemobius Jacobson, 1904 Type species: Nemobius tartarus (= Pteronemobius heydenii tartarus) Saussure, 1874, Published as part of He, Zhixin, Wang, Ning, Huang, Huateng & Ma, Libin, 2022, Two new species of ground crickets (Orthoptera: Trigonidiidae; Nemobiinae) from China, pp. 473-479 in Zootaxa 5092 (4) on page 476, DOI: 10.11646/zootaxa.5092.4.6, http://zenodo.org/record/5886518, {"references":["Saussure, H. (1874) Etudes sur les insectes orthopteres, famille des gryllides. Mission scientifique au Mexique et dans l'Amerique centrale, 6, 296 - 515."]}

Homonemobius brevipennis He & Ma, sp. nov. 短D同n'ḋ Figs. 1, 2, 3A–B, 4A–C Holotype. China: Male, Yunnan, Er’yuan, Niuguidan Village, 5.VII.2021, Libin Ma & Wei Yuan coll. Paratypes. 4 males and 4 females, the same data as the holotype; 2 males & 1 female, Yunnan, Lijiang, Gucheng County, Sheshan Park, 5.VIII.2021, Zhixin He & Ning Wang coll. (SNNU). Description. Male (Figs. 2A and 3A). Body size small. Head small and pubescent, slightly wider than the anterior margin of pronotum. Frontal rostrum twice broader than antennal scape. Eyes large. Apical three joints of maxillary palpi distinctly elongated and the 5th joint apically truncated and shaped as horseshoe shaped. Pronotum transverse, both anterior and posterior margins straight and the posterior wider than the anterior. Tegmen short; dorsal field bearing four longitudinal veins and lateral field with four Sc branches. Tympana absent. Hind tibia bears three dorsal external spurs and four internal spurs, and apical spurs comprising two outside and three inside. Genitalia (Figs. 3A–C). Epiphallus relatively short (confined to the basal half of the genitalia complex) and ectoparamere very large (almost similar to epiphallus). Median part of the epiphallus shaped as a bridge and very thin. Ectoparameres clip-shaped (viewed laterally) with an upper branch (large, curved, proximally expanded and apically acute) and lower branch (short and nearly straight). Female (Figs. 2B, 3B). Resemble male. Body size slightly larger than male. Ovipositor needle-like and slightly shorter than cercus. Coloration. Body blackish brown. Four brown longitudinal stripes at the dorsal area of the head. Legs dark brown. Measurements. Male (n=7). BL 4.22±1.29, PL 1.08±0.24, TL 1.27±0.83, HFL 2.79±0.62; Female (n=5). BL 4.45±1.10, PL 1.03±0.14, TL 1.03±0.56, OL 2.42±0.43. Etymology. The name refers to the species’ short tegmen. Distribution (Fig. 1). China (Yunnan). Remarks. This species is similar to Homonemobius curifolis Zhang et al., 2020 in color,but differin the shape of epiphallus. Of the known species, H. curifolis has a short and thick clip-like ectoparamere, which this structure is long and thin on H. brevipennis sp. nov. This new species also resembles Homonemobius amare He & Ma, 2021. The differences are: shorter tegmen, and more acute lower branch of the ectoparameres. It differs from Homonemobius nigrus Li et al., 2010 in ectoparameres (bifurcated vs. not bifurcated), and differs from H. monomorphus in coloration of body (dark brown vs. yellowish brown) and tegmen’s length (reached the 3 rd vs. 6 th segment of abdomen).

Pteronemobius osuviridis He & Ma, sp. nov. fl������n'��� Figs. 1, 3C���D, 4D���F, 5 Holotype. China: Male, Yunnan, Lincang, Linxiang County, Datian Village, 16.VIII.2021, Zhixin He, Ning Wang & Wei Yuan coll. Paratypes. 4 males and 4 females, the same data as holotype (SNNU). Description. Male (Figs 2C, 4A). Body size medium. Head round, slightly wider than pronotum. Rostrum approximately equal to antennal scape in width. Ocelli oval-like, median ocellus smallest. The apical joint of maxillary palpi longer than the third one, and it depressed, widened and with rounded apex; apical joint of labial palpi rod-like, longer than remaining sections, depressed and widened. Pronotum transverse, bearing bristles. Anterior and posterior margin straight, posterior margin wider than the anterior, both lateral edges slightly concaved in the middle. Tegmen reaching 7th abdominal segment, bearing one oblique vein and three chord veins. Dividing vein longitudinally separates the mirror as two cells. Apical region degraded. The Fore tibia only possesses a long oval-like external tympanum. Hind tibia bearing four dorsal spurs and three apical spurs on each side. Genitalia (Figs. 4D���F). Epiphallus large and dorsally broad, with apical margin broadly and medially concaved and armed with short lateral lobes. Ectoparamere clip-like (laterally viewed) and conspicuously shorter than epiphallus. In lateral view, ectoparamere aremedwith similar upper and lower branches and the upper one significantly and proximally expanded. Female (Figs 3D, 5B). Resembles male. Body size slightly smaller than male. Tegmen short, bearing five regular veins on the dorsal field. Ovipositor is sword-like and slightly shorter than cercus. Coloration. Body yellowish-brown. Four dark longitudinal stripes at the dorsal area of head. Anterior half of the abdomen colored dark brown. The apex of the hind femur bears small dark spots on two sides. The basal part of cercus greenish. Measurements. Male (n=5). BL 6.43��1.43, PL 1.42��0.46, TL 3.75��0.97, HFL 4.69��0.99; Female (n=4). BL 6.03��1.45, PL 1.49��0.31, TL 2.93��0.76, OL 4.10��0.66. Etymology. The name refers to the greenish color of the male's cercus. Distribution (Fig. 1). China (Yunnan). Remarks. This species is very similar to Pteronemobius nitidus (Bol��var, 1901) in appearance and coloration but different in body size and shape of ectoparamere. P. nitidus is larger than the new species (BL 8 mm vs. 6 mm). The male genitalia of the known species is armed with an upper branch nearly two times longer than the lower one, while the branches of the new species are similar in length., Published as part of He, Zhixin, Wang, Ning, Huang, Huateng & Ma, Libin, 2022, Two new species of ground crickets (Orthoptera: Trigonidiidae; Nemobiinae) from China, pp. 473-479 in Zootaxa 5092 (4) on pages 476-477, DOI: 10.11646/zootaxa.5092.4.6, http://zenodo.org/record/5886518, {"references":["Bolivar, I. (1901) Orthopteres, In E Zichy. Dritte Asiatischen Forschungsreise der Grafen Eugen Zichy (Zool. Ergebn). Budapest- Leipzig, 2, 223 - 243."]}

Homonemobius brevipennis He & Ma, sp. nov. ���D���n'��� Figs. 1, 2, 3A���B, 4A���C Holotype. China: Male, Yunnan, Er���yuan, Niuguidan Village, 5.VII.2021, Libin Ma & Wei Yuan coll. Paratypes. 4 males and 4 females, the same data as the holotype; 2 males & 1 female, Yunnan, Lijiang, Gucheng County, Sheshan Park, 5.VIII.2021, Zhixin He & Ning Wang coll. (SNNU). Description. Male (Figs. 2A and 3A). Body size small. Head small and pubescent, slightly wider than the anterior margin of pronotum. Frontal rostrum twice broader than antennal scape. Eyes large. Apical three joints of maxillary palpi distinctly elongated and the 5th joint apically truncated and shaped as horseshoe shaped. Pronotum transverse, both anterior and posterior margins straight and the posterior wider than the anterior. Tegmen short; dorsal field bearing four longitudinal veins and lateral field with four Sc branches. Tympana absent. Hind tibia bears three dorsal external spurs and four internal spurs, and apical spurs comprising two outside and three inside. Genitalia (Figs. 3A���C). Epiphallus relatively short (confined to the basal half of the genitalia complex) and ectoparamere very large (almost similar to epiphallus). Median part of the epiphallus shaped as a bridge and very thin. Ectoparameres clip-shaped (viewed laterally) with an upper branch (large, curved, proximally expanded and apically acute) and lower branch (short and nearly straight). Female (Figs. 2B, 3B). Resemble male. Body size slightly larger than male. Ovipositor needle-like and slightly shorter than cercus. Coloration. Body blackish brown. Four brown longitudinal stripes at the dorsal area of the head. Legs dark brown. Measurements. Male (n=7). BL 4.22��1.29, PL 1.08��0.24, TL 1.27��0.83, HFL 2.79��0.62; Female (n=5). BL 4.45��1.10, PL 1.03��0.14, TL 1.03��0.56, OL 2.42��0.43. Etymology. The name refers to the species��� short tegmen. Distribution (Fig. 1). China (Yunnan). Remarks. This species is similar to Homonemobius curifolis Zhang et al., 2020 in color,but differin the shape of epiphallus. Of the known species, H. curifolis has a short and thick clip-like ectoparamere, which this structure is long and thin on H. brevipennis sp. nov. This new species also resembles Homonemobius amare He & Ma, 2021. The differences are: shorter tegmen, and more acute lower branch of the ectoparameres. It differs from Homonemobius nigrus Li et al., 2010 in ectoparameres (bifurcated vs. not bifurcated), and differs from H. monomorphus in coloration of body (dark brown vs. yellowish brown) and tegmen���s length (reached the 3 rd vs. 6 th segment of abdomen)., Published as part of He, Zhixin, Wang, Ning, Huang, Huateng & Ma, Libin, 2022, Two new species of ground crickets (Orthoptera: Trigonidiidae; Nemobiinae) from China, pp. 473-479 in Zootaxa 5092 (4) on pages 474-475, DOI: 10.11646/zootaxa.5092.4.6, http://zenodo.org/record/5886518, {"references":["Zhang, D. X., Wang, P. & Liu, H. Y. (2020) A contribution to taxonomy of the superfamily Grylloidea Laicharting from Yunnan, China (Orthoptera: Grylloidea). Zootaxa, 4729 (4), 575 - 581. https: // doi. org / 10.11646 / zootaxa. 4729.4.8","He, Z. X., Zhang, T. & Ma, L. B. (2021) Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species. Zootaxa, 5011 (1), 1 - 70. https: // doi. org / 10.11646 / zootaxa. 5011.1.1","Li, K., He, Z. Q. & Liu, X. W. (2010) Four new species of Nemobiinae from China (Orthoptera, Gryllidae, Nemobiinae). Zootaxa, 2540 (1), 57 - 64. https: // doi. org / 10.11646 / zootaxa. 2572.1.4"]}

Pteronemobius osuviridis He & Ma, sp. nov. fl尾异n'ḋ Figs. 1, 3C–D, 4D–F, 5 Holotype. China: Male, Yunnan, Lincang, Linxiang County, Datian Village, 16.VIII.2021, Zhixin He, Ning Wang & Wei Yuan coll. Paratypes. 4 males and 4 females, the same data as holotype (SNNU). Description. Male (Figs 2C, 4A). Body size medium. Head round, slightly wider than pronotum. Rostrum approximately equal to antennal scape in width. Ocelli oval-like, median ocellus smallest. The apical joint of maxillary palpi longer than the third one, and it depressed, widened and with rounded apex; apical joint of labial palpi rod-like, longer than remaining sections, depressed and widened. Pronotum transverse, bearing bristles. Anterior and posterior margin straight, posterior margin wider than the anterior, both lateral edges slightly concaved in the middle. Tegmen reaching 7th abdominal segment, bearing one oblique vein and three chord veins. Dividing vein longitudinally separates the mirror as two cells. Apical region degraded. The Fore tibia only possesses a long oval-like external tympanum. Hind tibia bearing four dorsal spurs and three apical spurs on each side. Genitalia (Figs. 4D–F). Epiphallus large and dorsally broad, with apical margin broadly and medially concaved and armed with short lateral lobes. Ectoparamere clip-like (laterally viewed) and conspicuously shorter than epiphallus. In lateral view, ectoparamere aremedwith similar upper and lower branches and the upper one significantly and proximally expanded. Female (Figs 3D, 5B). Resembles male. Body size slightly smaller than male. Tegmen short, bearing five regular veins on the dorsal field. Ovipositor is sword-like and slightly shorter than cercus. Coloration. Body yellowish-brown. Four dark longitudinal stripes at the dorsal area of head. Anterior half of the abdomen colored dark brown. The apex of the hind femur bears small dark spots on two sides. The basal part of cercus greenish. Measurements. Male (n=5). BL 6.43±1.43, PL 1.42±0.46, TL 3.75±0.97, HFL 4.69±0.99; Female (n=4). BL 6.03±1.45, PL 1.49±0.31, TL 2.93±0.76, OL 4.10±0.66. Etymology. The name refers to the greenish color of the male's cercus. Distribution (Fig. 1). China (Yunnan). Remarks. This species is very similar to Pteronemobius nitidus (Bolívar, 1901) in appearance and coloration but different in body size and shape of ectoparamere. P. nitidus is larger than the new species (BL 8 mm vs. 6 mm). The male genitalia of the known species is armed with an upper branch nearly two times longer than the lower one, while the branches of the new species are similar in length.

Genus Homonemobius Chopard, 1935 Type species: Nemobius monomorphus (= Homonemobius monomorphus) Bol��var, 1900, Published as part of He, Zhixin, Wang, Ning, Huang, Huateng & Ma, Libin, 2022, Two new species of ground crickets (Orthoptera: Trigonidiidae; Nemobiinae) from China, pp. 473-479 in Zootaxa 5092 (4) on page 473, DOI: 10.11646/zootaxa.5092.4.6, http://zenodo.org/record/5886518, {"references":["Chopard, L. (1935) On a collection of Gryllidae from India made by Mr. B. M. Fletcher. Annals and Magazine of Natural History, London, 10 (16), 284 - 296. https: // doi. org / 10.1080 / 00222933508655047","Bolivar, I. (1900) Les Orthopteres de St-Joseph's College a Trichinopoly (Sud de l'Inde). Annales de la Societe Entomologique de France, 68, 761 - 812."]}

Pteronemobius (Pteronemobius) choui He & Ma, sp. n. (Figs. 45���46) Type material. Holotype. China: Male, Zhejiang, Tianmu Mountain National Natural Reserve, vii-21-1963, Yao Zhou coll. (NWAFU). Description. Male. Body size medium. Head round, slightly wider than pronotum. Rostrum approximately equal to antennal scape in width. Ocelli oval-like, median ocellus smallest of the three. Postclypeus transverse wide and its middle portion convex, anteclypeus concave medially and half-round. Apical joint of maxillary palpi longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transverse wide, bearing bristles; anterior and posterior margin straight, posterior margin wider than the anterior, middle groove distinctly, middle of both lateral edges slightly concave. Tegmen reaching 6th segment of abdomen, bearing an oblique vein and three chord veins, the third one and diagonal vein converge proximally. Dividing vein longitudinal and separating mirror as left and right cells. Apical region degraded, lateral region bearing two Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing four dorsal spurs on two sides respectively, of the inside spurs, first one stubby and far away from others, and the basal of fourth one expanded; apical spurs three inside and outside respectively. Genitalia. In dorsal view, epiphallic upper and lower margin concave medially; lateral lobe somewhat widened at basal part and longer than ectoparamere, curved inward and arc-shaped, distal portion somewhat acute. In ventral view, apical endoparamere funnel-shaped; ectoparamere short and truncated; the basal of guiding rod shaped as triangular. Female. Unknown. Coloration. Body reddish brown. Head brown. Pronotum brown, a small portion of posterior margin reddish brown. Legs reddish brown. Tegmen reddish brown. Abdomen dark brown. Measurements (mm). Male. BL 6.86, HW 1.78, PL 1.29, TL 3.29, HFL 3.77. Etymology. The name derived from family name of Yao Zhou in his honor. Remarks. This species is slightly similar to P. (P.) heydenii concolour in male genitalia, but different in length and shape of ectoparamere. Its coloration of body is similar to P. (P.) yezoensis, but different in characteristic of male genitalia., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on pages 43-44, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792

Dianemobius zhengi Chen, 1994 Distribution: Sichuan, Yunnan., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Chen, J. (1994) A new species of the genus Dianemobius Vickery from Sichuan Province, China (Orthoptera: Gryllidae). Entomological Research, 1, 66 - 68."]}

Polionemobius chayuensis He & Ma, sp. n. (Figs. 33���35) Type material. Holotype. China: Male, Xizang, Chayu, Xiachayu, vi-7-2019, Libin Ma coll. (SNNU). Paratypes. 3 males & 2 females, same information as holotype. Description. Male. Body size small. Head small, slightly wider than the anterior of pronotum. Rostrum slightly wider than antennal scape. Median ocellus half-round and smallest, lateral ocelli round. Eyes rather large. Postclypeus transverse wide and anteclypeus concave medially. Apical joint of maxillary palpi slightly longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transverse wide, trapezoidal; anterior margin straight and the posterior somewhat convex, posterior margin wider than the anterior, middle of both lateral edges slightly concave. Tegmen bearing an oblique vein and three chord veins. Dividing vein longitudinal and separating mirror as left and right cells. Apical region degraded, lateral region bearing four Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing four dorsal spurs inside and three outside, first of the inside stubby and far away from others; apical spurs three on both sides respectively, first of the inside longest. Genitalia. Epiphallus extremely narrowed, lateral lobe rather long. Ectoparamere shaped as triangular sheet, inside protruded. Female. Similar to male but tegmen rather short, bearing few transverse veins and primary veins almost paralleled. Ovipositor blade shape. Coloration. Body colored dark brow. Legs colored yellowish brown. Measurements (mm). Male (n=4). BL 4.68��0.53, HW 1.38��0.15, PL 1.41��0.12, TL 2.89��0.36, HFL 3.41��0.45. Female (n=2). BL 4.46��0.31, HW 1.47��0.26, PL 1.08��0.20, TL 2.01��0.33, HFL 3.53��0.42, OL 2.13��0.36. Etymology. The name refers to the type locality of Chayu County. Remarks. This species is very similar to Polionemobius taprobanensis (Walker, 1869) and Polionembius yunnanus Liu & Shi, 2014 in ectoparamere of male genitalia, but its epiphallic median lobe not obvious. It is similar to P. yunnanus in coloration of body, but differs in coloration of fourth joint of maxillary palpi (the new species colored dark, P. yunnanus colored light)., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 33, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum, 1, 1 - 224.","Liu, H. Y. & Shi, F. M. (2014) Two new species of the genus Pteronemobius Jacobson (Orthoptera: Gryllidae) from China. Far Eastern Entomologist, 284, 19 - 23."]}

Homonemobius amare He & Ma, sp. n. Distribution: Yunnan.

Dianemobius furumagiensis (Ohmachi & Furukawa, 1929) Distribution: Guangxi, Hainan, Hebei, Inner Mongolia, Shandong, Sichuan, Taiwan, Zhejiang., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Ohmachi, F. & Furukawa, H. (1929) Nemobius furumagiensis, a new species of Grylloidea. Proceedings of the Imperial Academy of Japan, 5 (8), 374 - 376. https: // doi. org / 10.2183 / pjab 1912.5.374"]}

Speonemobius minor He & Ma, sp. n. (Figs. 75���77) Type material. Holotype. China: Male, Xizang, Motuo, Beibeng, ix-11-2010, Libin Ma coll. Paratype. 1 male, Xizang, Motuo, Beibeng, v-31-2019, Libin Ma coll. (SNNU). Description. Male. Body size small for the genus. Head small, slightly wider than the anterior of pronotum. Rostrum distinctly wider than antennal scape. Ocelli round, median ocellus largest of the three. Eyes rather large. Postclypeus transversely widened, anteclypeus concave medially. Apical joint of maxillary palpi distinctly longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transversely widened, trapezoid; anterior margin straight and posterior one somewhat convex, posterior margin wider than the anterior, middle of both lateral edges slightly concave. Tegmen bearing an oblique vein and two chord veins. Apical region degraded, lateral region bearing three Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing two apical spurs inside and three outside; and hind tibia bearing three dorsal spurs on both sides respectively. Genitalia. Shaped as columnar sheet. Epiphallus widened medially and lateral lobe blunt, with gap between them shallowly concave. Ectoparamere finger-like. Female. Unknown. Coloration. Head light yellow, ornamented with grey stripes. Pronotum light yellow and bearing two pair of light patterns. Tegmen dark brown, ornamented with yellow stripes. Measurements (mm). Male (n=2). BL 4.26��0.04, HW 1.45��0.02, PL 1.03��0.02, TL 1.68��0.06, HFL 2.84��0.11. Etymology. The name refers to its small size. Remarks. This species is very similar to Speonemobius litoreus Vannini & Chelazzi, 1978 and Speonemobius alaesignatus Ingrisch, 1987 in body shape and head coloration, but it differs from them in the tegmen���s coloration and in the male genitalia. In addition, it is different from Speonemobius bifasciatus He et al., 2016 in body size and coloration, and different from Speonemobius fulvus He et al., 2016 in male genitalia, body coloration and wing length., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on pages 66-68, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Vannini, M. & Chelazzi, G. (1978) Description of a new species of intertidal cricket, Speonemobius litoreus, with notes on its ecology and behaviour. Monitore zoologico italiano, Supplement, New Series, 11 (1), 1 - 13. https: // doi. org / 10.1080 / 03749444.1978.10736574","Ingrisch, S. (1987) Neue Grillen von Borneo und aus Thailand (Insecta: Saltatoria: Grylloidea). Senckenbergiana biologica, 68 (1 - 3), 163 - 185.","He, Z. Q., Lu, H., Wang, X. Y. & Li, K. (2016) Two new species of Speonemobius from Southwest China (Orthoptera: Gryllidae: Nemobiinae). Zootaxa, 4205 (5), 454 - 458. https: // doi. org / 10.11646 / zootaxa. 4205.5.3"]}

Fibunemobius tamquam He & Ma, sp. n. Distribution: Xizang. Giganemobius jianfenglingensis (Liu & Shi, 2016) Distribution: Hainan.

Impetunemobius brunneis He & Ma, sp. n. Distribution: Guangxi. Parapteronemobius dibrachiatus (Ma & Zhang, 2015) Distribution: Guangdong., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 5, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Ma, L. B. & Zhang, T. (2015) First record of cricket genus Caconemobius (Grylloidea: Nemobiinae) from China with description of a new species. Zootaxa, 3914 (5), 585 - 590. https: // doi. org / 10.11646 / zootaxa. 3914.5.7"]}

Dianemobius csikii (Bol��var, 1901) Distribution: Hainan, Hebei, Inner Mongolia, Sichuan, Taiwan, Yunnan, Ningxia., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Bolivar, I. (1901) In Zichy. Zoologische Ergebnisse der dritten Asiatischen Forschungsreise des Grafen Eugen Zichy, 2, 223 - 243."]}

Dianemobius fascipes fascipes (Walker, 1869) Distribution: Fujian, Guangdong, Hainan, Hebei, Hubei, Hunan, Inner Mongolia, Jiangxi, Jilin, Shandong, Shaanxi, Taiwan, Yunnan, Zhejiang., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum, 1, 1 - 224."]}

Claranemobius yaoquensis He & Ma, sp. n. Distribution: Yunnan.

Impetunemobius brunneis He & Ma, sp. n. Distribution: Guangxi. Parapteronemobius dibrachiatus (Ma & Zhang, 2015) Distribution: Guangdong.

Homonemobius amare He & Ma, sp. n. Distribution: Yunnan., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 5, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792

Claranemobius yaoquensis He & Ma, sp. n. Distribution: Yunnan., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792

Erexitonemobius bellus He & Ma, sp. n. Distribution: Xizang., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on page 4, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792

Fibunemobius tamquam He & Ma, sp. n. Distribution: Xizang. Giganemobius jianfenglingensis (Liu & Shi, 2016) Distribution: Hainan., Published as part of He, Zhixin, Zhang, Tao & Ma, Libin, 2021, Crickets of subfamily Nemobiinae Saussure, 1877 (Orthoptera: Grylloidea; Trigonidiidae) from China with descriptions of new genera and new species, pp. 1-70 in Zootaxa 5011 (1) on pages 4-5, DOI: 10.11646/zootaxa.5011.1.1, http://zenodo.org/record/5157792, {"references":["Liu, H. Y. & Shi, F. M. (2016) Checklist of Nemobiinae from China (Orthoptera: Trigonidiidae). International Journal of Fauna and Biological Studies, 3 (4), 103 - 108"]}

Pteronemobius (Pteronemobius) choui He & Ma, sp. n. (Figs. 45–46) Type material. Holotype. China: Male, Zhejiang, Tianmu Mountain National Natural Reserve, vii-21-1963, Yao Zhou coll. (NWAFU). Description. Male. Body size medium. Head round, slightly wider than pronotum. Rostrum approximately equal to antennal scape in width. Ocelli oval-like, median ocellus smallest of the three. Postclypeus transverse wide and its middle portion convex, anteclypeus concave medially and half-round. Apical joint of maxillary palpi longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transverse wide, bearing bristles; anterior and posterior margin straight, posterior margin wider than the anterior, middle groove distinctly, middle of both lateral edges slightly concave. Tegmen reaching 6th segment of abdomen, bearing an oblique vein and three chord veins, the third one and diagonal vein converge proximally. Dividing vein longitudinal and separating mirror as left and right cells. Apical region degraded, lateral region bearing two Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing four dorsal spurs on two sides respectively, of the inside spurs, first one stubby and far away from others, and the basal of fourth one expanded; apical spurs three inside and outside respectively. Genitalia. In dorsal view, epiphallic upper and lower margin concave medially; lateral lobe somewhat widened at basal part and longer than ectoparamere, curved inward and arc-shaped, distal portion somewhat acute. In ventral view, apical endoparamere funnel-shaped; ectoparamere short and truncated; the basal of guiding rod shaped as triangular. Female. Unknown. Coloration. Body reddish brown. Head brown. Pronotum brown, a small portion of posterior margin reddish brown. Legs reddish brown. Tegmen reddish brown. Abdomen dark brown. Measurements (mm). Male. BL 6.86, HW 1.78, PL 1.29, TL 3.29, HFL 3.77. Etymology. The name derived from family name of Yao Zhou in his honor. Remarks. This species is slightly similar to P. (P.) heydenii concolour in male genitalia, but different in length and shape of ectoparamere. Its coloration of body is similar to P. (P.) yezoensis, but different in characteristic of male genitalia.

Speonemobius minor He & Ma, sp. n. (Figs. 75–77) Type material. Holotype. China: Male, Xizang, Motuo, Beibeng, ix-11-2010, Libin Ma coll. Paratype. 1 male, Xizang, Motuo, Beibeng, v-31-2019, Libin Ma coll. (SNNU). Description. Male. Body size small for the genus. Head small, slightly wider than the anterior of pronotum. Rostrum distinctly wider than antennal scape. Ocelli round, median ocellus largest of the three. Eyes rather large. Postclypeus transversely widened, anteclypeus concave medially. Apical joint of maxillary palpi distinctly longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transversely widened, trapezoid; anterior margin straight and posterior one somewhat convex, posterior margin wider than the anterior, middle of both lateral edges slightly concave. Tegmen bearing an oblique vein and two chord veins. Apical region degraded, lateral region bearing three Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing two apical spurs inside and three outside; and hind tibia bearing three dorsal spurs on both sides respectively. Genitalia. Shaped as columnar sheet. Epiphallus widened medially and lateral lobe blunt, with gap between them shallowly concave. Ectoparamere finger-like. Female. Unknown. Coloration. Head light yellow, ornamented with grey stripes. Pronotum light yellow and bearing two pair of light patterns. Tegmen dark brown, ornamented with yellow stripes. Measurements (mm). Male (n=2). BL 4.26±0.04, HW 1.45±0.02, PL 1.03±0.02, TL 1.68±0.06, HFL 2.84±0.11. Etymology. The name refers to its small size. Remarks. This species is very similar to Speonemobius litoreus Vannini & Chelazzi, 1978 and Speonemobius alaesignatus Ingrisch, 1987 in body shape and head coloration, but it differs from them in the tegmen’s coloration and in the male genitalia. In addition, it is different from Speonemobius bifasciatus He et al., 2016 in body size and coloration, and different from Speonemobius fulvus He et al., 2016 in male genitalia, body coloration and wing length.

Polionemobius chayuensis He & Ma, sp. n. (Figs. 33–35) Type material. Holotype. China: Male, Xizang, Chayu, Xiachayu, vi-7-2019, Libin Ma coll. (SNNU). Paratypes. 3 males & 2 females, same information as holotype. Description. Male. Body size small. Head small, slightly wider than the anterior of pronotum. Rostrum slightly wider than antennal scape. Median ocellus half-round and smallest, lateral ocelli round. Eyes rather large. Postclypeus transverse wide and anteclypeus concave medially. Apical joint of maxillary palpi slightly longer than the third, depressed and widened, with rounded apex; apical joint of labial palpi rod-like, longer than remainder sections, depressed and widened. Pronotum transverse wide, trapezoidal; anterior margin straight and the posterior somewhat convex, posterior margin wider than the anterior, middle of both lateral edges slightly concave. Tegmen bearing an oblique vein and three chord veins. Dividing vein longitudinal and separating mirror as left and right cells. Apical region degraded, lateral region bearing four Sc branches. Fore tibia only possessing external tympanum, elongate oval-like. Hind tibia bearing four dorsal spurs inside and three outside, first of the inside stubby and far away from others; apical spurs three on both sides respectively, first of the inside longest. Genitalia. Epiphallus extremely narrowed, lateral lobe rather long. Ectoparamere shaped as triangular sheet, inside protruded. Female. Similar to male but tegmen rather short, bearing few transverse veins and primary veins almost paralleled. Ovipositor blade shape. Coloration. Body colored dark brow. Legs colored yellowish brown. Measurements (mm). Male (n=4). BL 4.68±0.53, HW 1.38±0.15, PL 1.41±0.12, TL 2.89±0.36, HFL 3.41±0.45. Female (n=2). BL 4.46±0.31, HW 1.47±0.26, PL 1.08±0.20, TL 2.01±0.33, HFL 3.53±0.42, OL 2.13±0.36. Etymology. The name refers to the type locality of Chayu County. Remarks. This species is very similar to Polionemobius taprobanensis (Walker, 1869) and Polionembius yunnanus Liu & Shi, 2014 in ectoparamere of male genitalia, but its epiphallic median lobe not obvious. It is similar to P. yunnanus in coloration of body, but differs in coloration of fourth joint of maxillary palpi (the new species colored dark, P. yunnanus colored light).

Aphonoides aspidoid sp.­nov. (Figs. 12A; 16A–C) Examined­materials. Holotype.­ 1 male. China: Guangdong, Nanling, Pingdingshan, Aug. 6, 2019, He, Zhixin and Zhang, Tao coll. (SNNU); Paratypes.­ 2 males. China: Hunan, Langzhou, Mangshan, Oct. 6, 2004, Xie, Lingde coll. (SNNU). Measurements­ (mm).­ Male­ (n=3): ­ BL 13.92±2.74; HL 2.64±0.23; HW 3.65±0.57; PL 2.84±0.92; PW 3.74±0.53; FWL ­ 8.73±0.38; HFL 7.38±0.07; HTL 7.13±1.01 Etymology.­ The species name refers to the epiphallic lateral lobes blunt apically. Description.­Male: Head almost as wide as pronotum. Vertex broad and flattened, armed with densely pubescent. Frontal rostrum slightly convex in front view, inclined dorsally and ventrally. Median ocellus smallest of the three, ovoid; lateral ocellus transversely ovoid. End section of maxillary palpi almost equal to the length of the third; end section of labial palpi almost equal to the total length of remainder sections. Pronotum broad and flattened, anterior margin straight, posterior convex and middle of posterior margin angularly convex; median groove distinct. Longitudinal veins of tegmina paralleled. Hindwings longer than tegmina, uncovered portion about half length of hind femur. Inner tympanum shaped as elongate oval; outer tympanum absent. Hind tibiae evenly distributed with spines, armed with spurs distally (numbered 5:4); outer apical spurs two (equal in length) and the inner three (the dorsal one longest, two time longer than longest; middle one equal to length of longest; ventral one shortest). FIGURE­16.­ Genitalia of A. aspidoid sp.­nov. A. dorsal view; B. lateral view; C. ventral view. Genitalia.­ Lateral lobes of epiphallus trapezoidal. Median lobes of epiphallus small, apically acute. Ectoparamere proximally rod-like, not straight; apically expanded, shaped as baseball glove, with anterior side curved outward. Ectoparamere band-shaped in lateral view, included angle between ectoparamere and lateral lobes of epiphallus form an acute angle. Guiding rods shovel-like, apically acute. Coloration.­ Body brown. Clypeus and labrum yellowish. Tegmina brown. Female unknown. Remarks. This species resembles Aphonodies sabahi Gorochov, 2007 in features of body and genitalia, but are distinct in the epiphallus and ectoparamere. A. aspidoid sp.­nov. differs from A. sabahi in shape of ectoparamere, which of the new proximally not straight and what of the other is straight; ectoparamere of the new armed with outside margin apically concave and the other with inner margin concave. Besides, epiphallic lateral lobes apically inward truncated and what of the other outward truncated.

Zheng, Yanna, Xin, Zhixia, Xie, Lingde, Ma, Libin (2021): -Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure,-1878- (Orthoptera:-Gryllidae)-from-China. Zootaxa 4995 (3): 401-424, DOI: https://doi.org/10.11646/zootaxa.4995.3.1

Progoniogryllus rotundus sp. nov. Type materials. Holotype. Male. China: Xizang, Motuo, Beibeng, vi-1-2019, Ma, Libin coll. (SNNU). Paratype. China: 2 males, Xizang, Motuo, Beibeng, vi-1-2019, Ma, Libin coll. (SNNU); 1 male, Xizang, Chayu, Cibagou, vi-7-2019, Lv, Lin coll. (SNNU); 1 female, Xizang, Motuo, Beibeng, v-31-2019, Ma, Libin coll. (SNNU); 1 male, Yunnan, Lushui, Pianma, vi-20-2019, Ma, Libin coll. (SNNU); 1 female, Yunnan, Lushui, Pianma, vi-19-2019, Ma, Libin coll. (SNNU). Measurements. Male (n=5): BL 10.09��0.61, HL 1.76��0.09, HW 3.05��0.08, PL 2.06��0.08, PW 3.29��0.17, FWL 5.02��0.49, FWW 3.5��0.18, MTL 2.69��0.28, HFL 6.88��0.21; female (n=5): BL 10.84��0.59, HL 1.8��0.23, HW 3.14��0.39, PL 2.35��0.20, PW 3.33��0.25, MTL 2.33��0.36, HFL 7.50��0.25, OL 7.80��0.66 Description. Male. Body size small for Gryllinae species (Fig. 5A). In frontal view, head higher dorsoventrally, vertex rounded and convex (Fig. 9D). Dorsally viewed, occiput short, round and convex; vertex inclined. Frons convex and narrow (almost equal to the width of antennal scape), and arched in lateral view. A shallow groove present between bottom of frons and top of clypeus (along with epistomal suture). Labrum shield-like with lateral margins acutely convex and apically pointed round. End section of maxillary palpus conspicuously longer than the third; end section of labial palpus obviously longer than the total length of anterior two. Pronotum long and obviously longer than head. In dorsal view, pronotum somewhat trapezoid-like, anteriorly narrower than the posterior. Anterior margin of pronotum slightly concave and posteriorly straight. Disc flatten and possessing with a middle groove, and laterally fold down. Lateral lobes depressed posteriorly. Elytra broad and flatten, conspicuously wider than hind margin of pronotum. Basal field somewhat wide but short and almost as half length of pronotum. Oblique veins two. Diagonal vein arched. Mirror transverse and obviously wider than length. Dividing vein separating hind part oval-like. Apical field distally rounded, very short and somewhat a half length of mirror, and possessing two rows of irregular cells. Hind wings absent. Cercus long and almost as long as hind tibia. Outer tympana large and long oval-like; inner tympana absent and corresponding part pit-like. Hind femur slightly longer than tibia. Subapical spurs five pairs and inner ones obviously shorter than the outer proximally. Body pubescent, brown and ornamented with yellowish-brown and dark-brown patterns. Lateral areas of head and pronotum possessing continuous yellowish-brown stripes (Fig. 9D). Lateral lobes of pronotum almost dark brown with bottom margin yellowish-brown. Maxillary palpus colored dark and labial palpus colored light. Elytral disc yellowish-brown and most of lateral lobes colored dark brown. Fore and middle legs yellowish-brown and ornamented with brown patterns, and on femur and tibia ornamented with dark brown spots dorsally and laterally, and, especially, dark brown spots almost forming a ring on middle tibia. Hind legs and cercus colored brown and ornamented with dull yellowish brown. Abdomen dark brown dorsally and laterally, and ventrally colored yellowish-brown and ornamented with brown spots. Subgenital plate colored brown. Epiphallus almost without median lobe and slightly extended outward at the middle (Fig. 6A���C). Lateral lobes of epiphallus short and obtuse, the gap between them broad and somewhat rectangular. In lateral view, included angle between median and lateral lobe nearly arched. Ectoparamere short and slightly upward curved. In ventral view, ectoparamere comprise of two branches proximally, combined medially and curved upward. Female. Body size and coloration similar to male (Figs. 5B; 9C). Brachypterous, portions after mesonotum uncovered. Ovipositor long and slightly longer than cercus. Some samples macroperous and armed with normal elytra (Fig. 4B, D). Etymology. The name refers to its elytra armed with rounded margin apically, and its somewhat oval-like body. Remarks. Holotype and most specimens light brown and there are also some specimens that are darker. Compared with light ones, patterns or stripes of those dark samples are not clear or absent, and the coloration of legs and abdomen are darker. In addition, this species possesses rounded apical end of the tegmina., Published as part of Ma, Libin, Jing, Xuan & Zheng, Yanna, 2021, Integrative taxonomy base on morphology and molecular phylogeny with description of a new genus, Progoniogryllus gen. nov. and two new species (Orthoptera: Grylloidea: Gryllidae; Gryllinae), pp. 523-536 in Zootaxa 4995 (3) on pages 531-532, DOI: 10.11646/zootaxa.4995.3.7, http://zenodo.org/record/5057465

Genus­ Zvenella ­Gorochov,­1988 Zvenella acutangulata Xia, Liu & Yin, 1991 Zvenella scalpratus Ma & Zhang, 2012, syn.­nov. Distribution: Yunnan (Ruili, Cangyuan, Jinghong), Hainan (Yinggeling), Guangxi (Nonggang). Zvenella yunnana (Gorochov, 1985) Zvenella aequalis Ma & Zhang, 2012, syn.­nov. Distribution: Yunnan (Ruili, Mengla, Cangyuan), Guangxi (Damingshan). Zvenella geniculata (Chopard, 1931) Zvenella decussatus Ma & Zhang, 2012, syn.­nov. Distribution: Yunnan (Mengla)., Published as part of Zheng, Yanna, Xin, Zhixia, Xie, Lingde & Ma, Libin, 2021, - Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure, - 1878 - (Orthoptera: - Gryllidae) - from-China, pp. 401-424 in Zootaxa 4995 (3) on page 402, DOI: 10.11646/zootaxa.4995.3.1, http://zenodo.org/record/5056938, {"references":["Xia, K. L., Liu, X. W. & Yin, H. S. (1991) Two new species of the Chinese crickets (Orthoptera: Grylloidea). Contributions from the Shanghai Institute of Entomology, 10, 121 - 123.","Ma, L. B. & Zhang, Y. L. (2012) The crickets of genus Zvenella Gorochov, 1988 (Gryllidae: Podoscirtinae) from China. Zootaxa, 3597, 57 - 67. https: // doi. org / 10.11646 / zootaxa. 3597.1.8","Gorochov, A. V. (1985) On the Orthoptera subfamilies Itarinae, Podoscirtinae and Nemobiinae (Gryllidae) from eastern Indochina. In: Medvedev, L. N. (Ed.), Fauna i ekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam], 1985, pp. 17 - 25.","Chopard, L. (1931) On the Gryllidae from the Malay Peninsula. Bulletin of the Raffles Museum, 6, 124 - 149."]}

Progoniogryllus directus sp. nov. Type materials. Holotype. Male. China: Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU). Paratype. China: 2 males, Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU); 1 male, Xizang, Chayu, Xiachayu, vi-6-2019, Ma, Libin coll. (SNNU); 2 males, Xizang, Chayu, vi-13-2019, Yang, qichen coll. (SNNU); 2 females, Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU); 3 females, Xizang, Chayu, Xiachayu, vi-6-2019, Ma, Libin coll. (SNNU). Measurements. Male (n=5): BL 8.8��0.42, HL 1.37��0.08, HW 2.24��0.22, PL 1.83��0.07, PW 2.76��0.06, FWL 4.92��0.50, FWW 3.35��0.10, MTL 2.32��0.13, HFL 5.98��0.23; female (n=2): BL 8.21��0.34, HL 1.29��0.15, HW 2.19��0.04, PL 1.67��0.52, PW 2.69��0.07, MTL 2.61��0.15, HFL 6.22��0.17, OL 6.15��0.22 Description. Male. Body smaller than medium species of Gryllinae (Fig. 7A, B). In frontal view, head laterally widened and dorsoventrally short, and vertex broad and flat (Fig. 9B). In dorsal view, head longer than pronotum; occiput widened and slightly convex; vertex inclined; and frons somewhat convex, wide and slightly wider than antennal scape. Antennal scape transverse, shield-like. Epistomal suture shallow, sinuate and medially upward convex. Lower portion of clypeus and labrum depressed. Labrum broad with lateral margins rounded convex and apically arched. End section of maxillary palpus obviously longer than the third; end section of labial palpus conspicuously longer than the total length of remainder two. Pronotum transverse and slightly longer than the half width. Disc of pronotum broad and flattened, anteriorly concave and posteriorly straight, and lateral margin straight and inclined down. Lateral lobes of pronotum broad and flattened with ventral surface slightly inwardly concave. Tegmina very short and not reaching abdominal tip. Tegminal disc broad and flattened, almost rectangular-shaped, slightly arched laterally and pointed rounded apically. Basal field short and slightly longer than 1/3 length of pronotum. Oblique veins two; diagonal vein weakly arched and smoothly jointing with outside margin of mirror. Mirror transverse, somewhat shaped as rhombus with inner and bottom corner rounded. Dividing vein separating a small oval at mirror bottom. Apical field distally straight, very short and shorter than 1/3 length of mirror. Cercus thin and tapering. Outer tympana large and long oval-like; inner tympana very small and oval-like. Hind femur slightly longer than tibia. Subapical spurs six pairs and proximally possessing an additional small one outside. Hind tarsus armed with two rows of spines and numbered more outside. Body colored dark brown dorsally and yellowish-brown ventrally. Occiput ornamented with six yellowishbrown stripes (Fig. 9B). Of them, the outermost ones inclined outward and jointing with yellowish-brown stripes above eyes. Yellowish-brown ornamented above eyes or around ocelli. Occiput, vertex, top portions of frons and inner margin of antennal socket dark brown. Eyes and portions between antennal socket and labrum, including mandible, maxillary and labial palpi yellowish-brown. Pronotum dark brown dorsally and only ornamented with few of yellowish-brown spot laterally. Lateral lobes of pronotum colored yellowish-brown at half lower portions. Elytra yellowish brown with lateral lobes colored dark.Abdomen colored dark brown dorsally and laterally, and yellowishbrown ventrally. Subgenital plate brown. Legs yellowish-brown and ornamented with few of brown spots. Epiphallus transverse (Fig. 8A���C), median lobe weakly convex and truncate apically; lateral lobes short and obtuse, in dorsal view, gap between them large and rectangular-like. Ectoparamere prismatoidal, proximally branched with outside branch thick and inner one thin, and combined as one apically. Female. Body form and coloration similar with male (Figs. 7C, D; 9A). Brachypterous individual armed with elytra bud-like (Fig. 7C). Ovipositor (Fig. 4A, C) yellowish-brown. Etymology. The name refers to its elytra armed with straight margin apically. Remarks. Inner tympana varied, sometimes pit-like and without tympanic membrane and sometimes with a very small tympanic membrane. This species possesses straight apical end of elytra., Published as part of Ma, Libin, Jing, Xuan & Zheng, Yanna, 2021, Integrative taxonomy base on morphology and molecular phylogeny with description of a new genus, Progoniogryllus gen. nov. and two new species (Orthoptera: Grylloidea: Gryllidae; Gryllinae), pp. 523-536 in Zootaxa 4995 (3) on pages 534-535, DOI: 10.11646/zootaxa.4995.3.7, http://zenodo.org/record/5057465

Progoniogryllus directus sp. nov. Type materials. Holotype. Male. China: Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU). Paratype. China: 2 males, Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU); 1 male, Xizang, Chayu, Xiachayu, vi-6-2019, Ma, Libin coll. (SNNU); 2 males, Xizang, Chayu, vi-13-2019, Yang, qichen coll. (SNNU); 2 females, Xizang, Chayu, Zhala village, vi-5-2019, Ma, Libin coll. (SNNU); 3 females, Xizang, Chayu, Xiachayu, vi-6-2019, Ma, Libin coll. (SNNU). Measurements. Male (n=5): BL 8.8±0.42, HL 1.37±0.08, HW 2.24±0.22, PL 1.83±0.07, PW 2.76±0.06, FWL 4.92±0.50, FWW 3.35±0.10, MTL 2.32±0.13, HFL 5.98±0.23; female (n=2): BL 8.21±0.34, HL 1.29±0.15, HW 2.19±0.04, PL 1.67±0.52, PW 2.69±0.07, MTL 2.61±0.15, HFL 6.22±0.17, OL 6.15±0.22 Description. Male. Body smaller than medium species of Gryllinae (Fig. 7A, B). In frontal view, head laterally widened and dorsoventrally short, and vertex broad and flat (Fig. 9B). In dorsal view, head longer than pronotum; occiput widened and slightly convex; vertex inclined; and frons somewhat convex, wide and slightly wider than antennal scape. Antennal scape transverse, shield-like. Epistomal suture shallow, sinuate and medially upward convex. Lower portion of clypeus and labrum depressed. Labrum broad with lateral margins rounded convex and apically arched. End section of maxillary palpus obviously longer than the third; end section of labial palpus conspicuously longer than the total length of remainder two. Pronotum transverse and slightly longer than the half width. Disc of pronotum broad and flattened, anteriorly concave and posteriorly straight, and lateral margin straight and inclined down. Lateral lobes of pronotum broad and flattened with ventral surface slightly inwardly concave. Tegmina very short and not reaching abdominal tip. Tegminal disc broad and flattened, almost rectangular-shaped, slightly arched laterally and pointed rounded apically. Basal field short and slightly longer than 1/3 length of pronotum. Oblique veins two; diagonal vein weakly arched and smoothly jointing with outside margin of mirror. Mirror transverse, somewhat shaped as rhombus with inner and bottom corner rounded. Dividing vein separating a small oval at mirror bottom. Apical field distally straight, very short and shorter than 1/3 length of mirror. Cercus thin and tapering. Outer tympana large and long oval-like; inner tympana very small and oval-like. Hind femur slightly longer than tibia. Subapical spurs six pairs and proximally possessing an additional small one outside. Hind tarsus armed with two rows of spines and numbered more outside. Body colored dark brown dorsally and yellowish-brown ventrally. Occiput ornamented with six yellowishbrown stripes (Fig. 9B). Of them, the outermost ones inclined outward and jointing with yellowish-brown stripes above eyes. Yellowish-brown ornamented above eyes or around ocelli. Occiput, vertex, top portions of frons and inner margin of antennal socket dark brown. Eyes and portions between antennal socket and labrum, including mandible, maxillary and labial palpi yellowish-brown. Pronotum dark brown dorsally and only ornamented with few of yellowish-brown spot laterally. Lateral lobes of pronotum colored yellowish-brown at half lower portions. Elytra yellowish brown with lateral lobes colored dark.Abdomen colored dark brown dorsally and laterally, and yellowishbrown ventrally. Subgenital plate brown. Legs yellowish-brown and ornamented with few of brown spots. Epiphallus transverse (Fig. 8A–C), median lobe weakly convex and truncate apically; lateral lobes short and obtuse, in dorsal view, gap between them large and rectangular-like. Ectoparamere prismatoidal, proximally branched with outside branch thick and inner one thin, and combined as one apically. Female. Body form and coloration similar with male (Figs. 7C, D; 9A). Brachypterous individual armed with elytra bud-like (Fig. 7C). Ovipositor (Fig. 4A, C) yellowish-brown. Etymology. The name refers to its elytra armed with straight margin apically. Remarks. Inner tympana varied, sometimes pit-like and without tympanic membrane and sometimes with a very small tympanic membrane. This species possesses straight apical end of elytra.

Genus Progoniogryllus gen. nov. Type species: Progoniogryllus rotundus sp. nov., here designated. Diagnosis. Size small for Gryllinae species. Colored dark brown and ornamented with patches and stripes in light color, especially, above eyes there are backward forked stripes. Pronotum pubescent, armed with straight fore and hind margins and the former slightly narrower than the hind. Elytra short, far from supra-anal plate; basal and apical area of elytra rather short and almost similar in length; possessing two oblique veins and developed mirror; and media fold (fan) between apical part of lateral and dorsal area developed and expanded. Fore tibiae armed with outer tympanum and the inner one absent. Epiphallus almost without middle lobe and gap between lateral lobes rather broad. Ectoparamere simple and shaped as wide blade. Particularly, testis of the new unusually developed and occupies a large space in the abdominal cavity. Molecular phylogeny. To infer the taxonomic status of the new genus, we constructed phylogenetic relationship of two species of the new genus, 32 other Gryllinae species and three species of Gryllotalpidae (Fig. 2). The tree presented that the new taxa is included in the lineage of Modicogryllini and relevant species. Within the species we concluded, the new ones are much closer to Goniogryllus. Although they are distinct, it is significant that both the new and Goniogryllus bear similar coloration. Courtship behavior. If the male and female are placed in a box, the male will make a courtship call and approach female. The male will point the end of his abdomen at the female, and it can be observed that there is a spermatophore hanging at the end of the abdomen, but the female seems to avoid the male. Usually, the female's avoidance will not cause the male to give up, he will continue to get close to the female, constantly chase and point his tail up at the female in different positions. After courtship for about 7–15 minutes, the male keeps falling back and takes the opportunity to get under the female's body. After the external genitalia are connected, the copulation commences. During the mating, it is found that the female and the male twist at a certain angle (Fig. 3). The whole process lasts about 2 minutes, then the female leaves and the copulation ends. Etymology. The name refers to the position of the new in the phylogenic tree (fig. 2) in the article. Included species: Progoniogryllus rotundus sp. nov. and Progoniogryllus directus sp. nov. Remarks. External appearance of the new genus slightly resembles the genera Parasongella Otte, 1987, Callogryllus Sjöstedt, 1910 and Goniogryllus Chopard, 1936 since they both possess short elytra (or not) and bear yellowish stripes above eyes, but the tegmina of the new taxa are somewhat longer and possess an intact mirror, and the genitalia of the new taxa is distinct from the above. Parasongella includes three species, Parasongella ornaticeps (Chopard, 1969) (type locality: Thailand), Parasongella japonica Ichikawa, 2001 (type locality: Japan) and Parasongella nigriceps Otte, 1987 (type locality: Tanzania). Although considered in the same genus, their male genitalia are rather different from each other. The type species, P. nigriceps, bears genitalia that are very similar to Songella species and the clypeus is strongly upwards bent. The genitalic complex of P. japonica is very similar to the species of Goniogryllus, but that of P. ornaticeps is different from the former two. Thus, we do not think it warranted to combine those three species into Parasongella (Ichikawa, 2001). The male genitalia of the new genus is similar to the genus Gryllodes Saussure, 1874, and, their elytra is rather similar to each other. But the body of Gryllodes species is somewhat dorsoventrally compressed and that of the new genus is normal. Based on the phylogenic relationship which was established based on gene sequences, these two genera are distantly related.

Zvenella acutangulata ­Xia,­Liu­&­Yin­1991 (Figs. 2A; 3A, D; 4A–G) Zvenella acutangulata Xia, Liu & Yin, 1991: 121–123; Yin & Liu, 1995: 109; Gorochov, 2002: 336; Ma & Zhang, 2012: 60 Zvenella scalpratus Ma & Zhang, 2012: 60, syn.­nov. Examined­materials.­ 3 males. China: Yunnan, Ruili, Mengxiu, Jun. 12, 2013, Ma, Libin coll. (SNNU); 3 males. China: Yunnan, Jinghong, Jul. 11, 2018, Peng, Zhong coll. (SNNU); 2 males. China: Yunnan, Mengla, Yaoqu, May 24, 2013, Ma, Libin coll. (SNNU); 3 males. China: Yunnan, Mengla, Menglun, May 12, 2009, Ma, Libin coll. (SNNU); 2 males. China: Yunnan, Cangyuan, Banlao, Jun. 4, 2013, Ma, Libin coll. (SNNU); 3 males. China: Hainan, Baisha, Yinggeling, Aug. 21, 2019, He, Zhixin and Zhang, Tao coll. (SNNU); 1 male: Guangxi, Longzhou, Zhubu, Nonggang, Sep. 9, 2020, He, Zhixin coll. (SNNU); 1 male: Guangxi, Longzhou, Zhubu, Nonggang, May 6, 2019, Ma, Libin and Zhang, Tao coll. (SNNU). Measurements­ (mm).­ Male­ (n=18): ­ BL 17.30±2.12; HL 5.48±0.53; HW 3.10±0.57; PL 2.52±0.24; PW 4.00±0.33; FWL ­ 13.50±0.45; HFL 6.12±0.02; HTL 10.66±0.08 Description.­Male:­ Vertex broad and flattened, slightly inclined. Occiput slightly convex, almost as wide as anterior margin of pronotum. Frontal rostrum slightly narrower than antennal scape; slightly convex in front view, inclined dorsally and ventrally. Antennal scape shield-like. Median ocellus smallest and transversely ovoid; lateral ocelli rounded. Eyes slightly convex, about 1/3 length of head. Postclypeus shaped as narrow band; anteclypeus shaped as broad shield. Labrum shaped as rhombus, lateral margins angle-like and the apex concave medially. End section of maxillary palpi almost equal to the length of the third; end section of labial palpi depressed and widened, about two times longer than the second. Pronotum broad and flattened, anterior margin concave and the posterior convex; posterior margin slightly wider than the anterior. Basal margin of metanotal gland straight, laterally convex at the middle and apically sinuated; keels U-shaped and apically assigned at the middle of gland; elongate setae distributed at the middle of gland and this area almost rounded. Tegmina with oblique veins four. Diagonal vein proximally bifurcated and both branches connected to CuA vein. Chord veins three, connecting to the proximal of mirror by two transverse veins. Mirror shield-like, longer than width; dividing vein straight. Apical field armed with regular reticulate veins. Hindwings longer than tegmina, uncovered portion about two times longer than hind femur. FIGURE­2.­ Bodies of Z. acutangulata and Z. yunnana. A. Z. acutangulata; B. Z. yunnana. Scale bar: 10 mm. Inner tympanum larger than the outer; the outer one oval-like and the inner shaped as elongate oval. Hind tibiae evenly equipped with spines and armed with spurs distally (numbered 5:5); inner apical spurs three (the dorsal one longest, the ventral one shortest and about 1/3 length of the longest, the middle one about 1/2 length of the longest) and outer apical spurs two (equal in length) and about 1/2 length of the longest. Supra-anal plate trapezoidal, the anterior concave and posterior margin straight; the anterior about two times wider than of the posterior margin; concave at middle of super-anal plate. Genitalia.­ Middle lobes of epiphallus long, erect, shaped as rod and acute apically. Lateral lobes of epiphallus short, included angle between middle lobe and lateral lobe form a right angle. Epiphallus ventrally possessing two lateral projections, anterior one rod-like (apically rounded, about 1/3 length of middle lobe of epiphallus) and caudal one sheet-like (ventrally concave and armed with dense and sharp denticles, and dorsal margin flat). Ectoparamere apically bifurcated with upper branch apically rounded (about 1/2 length of the lower) and lower branch apically acute. Guiding rod shovel-like, proximally widened and apically slightly acute. Coloration.­ Body colored brown. Occiput light brown; ocelli yellow; eyes brown; antennal scape yellowish brown; labrum light brown. Pronotum dark brown, ornamented with axe-shaped pattern which yellowish brown. Tegmina brown. Hind tibiae dark brown. Female unknown. FIGURE­3.­ Metanotal glands (A–C) and supra-anal plates (D–F) of Zvenella species. A and D. Z. acutangulata; B and E. Z. yunnana; C and F. Z. geniculata. FIGURE­4.­ Genitalia of Z. acutangulata. A. dorsal view; B. lateral view; C. ventral view; D. caudal viewed; E. inside of male genitalia, lateral view; F. guiding rod, ventral view; G. guiding rod, lateral view. Remarks. Ma and Zhang (2012) identified some Hainan specimens as Z. scalpratus, which resemble Z. acutangulata and it was thought there might be some differences between them. The authors argued that they were different in ventral projections and lateral lobes of epiphallus as well as characters of supra-anal plate. After examining about twenty specimens from Hainan, Guangxi and Yunnan, we found these differences are intraspecific variation. In addition, based on our collections and literature records, Z. acutangulata should possess a broad distributional range. Thus, we consider that Z. scalpratus should be junior synonym of Z. acutangulata., Published as part of Zheng, Yanna, Xin, Zhixia, Xie, Lingde & Ma, Libin, 2021, - Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure, - 1878 - (Orthoptera: - Gryllidae) - from-China, pp. 401-424 in Zootaxa 4995 (3) on pages 404-407, DOI: 10.11646/zootaxa.4995.3.1, http://zenodo.org/record/5056938, {"references":["Xia, K. L., Liu, X. W. & Yin, H. S. (1991) Two new species of the Chinese crickets (Orthoptera: Grylloidea). Contributions from the Shanghai Institute of Entomology, 10, 121 - 123.","Yin, H. S. & Liu, X. W. (1995) Synopsis on the classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific & Technological Literature Publishing House, Shanhai, 239 pp.","Gorochov, A. V. (2002) Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 1. The male genitalia and Indo-Malayan Podoscirtini. Zoosystematica Rossica, 10 (12), 303 - 350.","Ma, L. B. & Zhang, Y. L. (2012) The crickets of genus Zvenella Gorochov, 1988 (Gryllidae: Podoscirtinae) from China. Zootaxa, 3597, 57 - 67. https: // doi. org / 10.11646 / zootaxa. 3597.1.8"]}

Phyllotrella planidorsalis ��Gorochov,��1988 (Figs. 6B; 8A���B; 9A���E) Phyllotrella planidorsalis Gorochov, 1988: 18, 2002: 334; Yin & Liu, 1995: 113; Kim & Hong, 2014: 63; Sun & Liu, 2019: 444 Phyllotrella fumingi Sun & Liu, 2019: 445, syn.��nov Phyllotrella hainanensis Sun & Liu, 2019: 443, syn.��nov Phyllotrella transversa Sun & Liu, 2019: 444, syn.��nov Examined��materials. 3 males. China: �� Guangdong, Nanling, Sep. 1���7, 2020, He, Zhixin coll. (SNNU); 3 males. China: �� Guangdong, Nanling, Aug. 1, 2020, Zhang, Tao coll. (SNNU); 3 males. China: �� Guangxi, Shiwandashan National Forest Park, May 8, 2019, Ma, Libin and Zhang, Tao coll. (SNNU). �� Measurements��(mm).��Male: �� BL 18.11��2.04; HL 4.48��0.53; HW 3.10��0.57; PL 3.64��0.24; PW 3.02��0.33; FWL �� 26.43��0.45; HFL 10.54��0.02; HTL 9.56��0.08 FIGURE��8.�� Metanotal gland and super-anal plate of P. planidorsalis. A. metanotal gland; B. supra-anal plate. Description.��Male:�� Head wider than the anterior of pronotum. Occiput narrowed and flattened. Vertex slightly inclined, armed with central groove and densely pubescent. Frontal rostrum extremely convex, inclined dorsally and ventrally; extremely narrowed dorsally, almost as wide as antennal scape. Antennal scape shield-like. Ocelli inconspicuous, transversely ovoid. Eye ovoid, about 1/3 length of head. Postclypeus flattened; anteclypeus concave. Labrum convex and shaped as triangular. End section of maxillary palpi slightly narrower than the third; labial palpi short and the end section shaped as knife. Pronotum trapezoid, anterior margin straight, posterior margin rounded and convex, posterior margin wider than the anterior. Pronotal disc broad. Metanotal gland densely pubescent; short setae distributed along proximal margin; distally armed with upward protrude centrally. Anterior margin of metanotal gland straight; lateral margin medially concave; posterior margin arc-shaped and convex (Fig. 8). Oblique veins six, last one proximally linked to the basal of diagonal vein by a transverse vein. Chord veins three, two external ones joined proximally and linked to diagonal vein and then converging with CuA vein, the third chord vein linking with mirror by two transverse veins. Mirror large, oval-like; dividing vein straight. Apical field shaped as triangular, about 1/3 length of tegmina, armed with irregular wing chambers. Hindwings slightly longer than the tegmina, and uncovered portion about 1/2 length of hind femur. Inner tympanum slit-like and the outer tympanum quadrate. Hind tibiae bearing spines proximally and spurs distally (numbered 5:4). Supra-anal plate symmetrically divided each side as triangular and apically pubescent. Genitalia. Lateral lobes of epiphallus band-shaped, apically bifurcated as right angle, both branches apically acute; upper branch about two times longer than the lower. Ectoparamere forked into three parts, one branch forward and head pointed; other two branches curved and band-shaped with acute apex, and included angles between each two of branches varied in individuals. FIGURE��9.�� Genitalia of P. planidorsalis. A. dorsal view; B. lateral view; C. ventral view; D. inside of male genitalia, lateral view; E. inside of male genitalia, apically enlarged view. Coloration.�� Body colored yellowish green. Eyes brown; clypeus and labrum yellowish. Pronotum yellowish green, a pair of eyelike spots of pronotum disc dark brown. Tegmina green, mirror and oblique veins brown. Female unknown. Remarks.�� Coloration of this species varied from light to dark. The specimens we describe here are colored yellowish and those specimens in dark color possessing similar type of pattern. �� P. planidorsalis was established by Gorochov (1988) and type locality is Vietnam. This species was first described by Yin & Liu (1995) in Hainan, China. Then, Sun & Liu (1995) proposed three species, P. fumingi, P. hainanensis and P. transversa, based on characters of supra-anal plate and ectoparamere. We examined specimens and determined that these differences represent intraspecific variation, espeically, the movable ectoparamere always presenting various appearance in specimens. Therefore, we consider that P. fumingi, P. hainanensis and P. transversa are junior synonyms of P. planidorsalis., Published as part of Zheng, Yanna, Xin, Zhixia, Xie, Lingde & Ma, Libin, 2021, - Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure, - 1878 - (Orthoptera: - Gryllidae) - from-China, pp. 401-424 in Zootaxa 4995 (3) on pages 411-412, DOI: 10.11646/zootaxa.4995.3.1, http://zenodo.org/record/5056938, {"references":["Gorochov, A. V. (1988) New and little-known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and certain other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), Fauna i ekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam], pp. 5 - 21.","Gorochov, A. V. (2002) Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 1. The male genitalia and Indo-Malayan Podoscirtini. Zoosystematica Rossica, 10 (12), 303 - 350.","Yin, H. S. & Liu, X. W. (1995) Synopsis on the classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific & Technological Literature Publishing House, Shanhai, 239 pp.","Sun, K. & Liu, H. Y. (2019) Review of the genus Phyllotrella Gorochov, with descriptions of three new species from China (Orthoptera: Gryllidae: Podoscirtinae: Podoscirtini). Zootaxa, 4629 (3), 441 - 447. https: // doi. org / 10.11646 / zootaxa. 4629.3.11"]}

Progoniogryllus rotundus sp. nov. Type materials. Holotype. Male. China: Xizang, Motuo, Beibeng, vi-1-2019, Ma, Libin coll. (SNNU). Paratype. China: 2 males, Xizang, Motuo, Beibeng, vi-1-2019, Ma, Libin coll. (SNNU); 1 male, Xizang, Chayu, Cibagou, vi-7-2019, Lv, Lin coll. (SNNU); 1 female, Xizang, Motuo, Beibeng, v-31-2019, Ma, Libin coll. (SNNU); 1 male, Yunnan, Lushui, Pianma, vi-20-2019, Ma, Libin coll. (SNNU); 1 female, Yunnan, Lushui, Pianma, vi-19-2019, Ma, Libin coll. (SNNU). Measurements. Male (n=5): BL 10.09±0.61, HL 1.76±0.09, HW 3.05±0.08, PL 2.06±0.08, PW 3.29±0.17, FWL 5.02±0.49, FWW 3.5±0.18, MTL 2.69±0.28, HFL 6.88±0.21; female (n=5): BL 10.84±0.59, HL 1.8±0.23, HW 3.14±0.39, PL 2.35±0.20, PW 3.33±0.25, MTL 2.33±0.36, HFL 7.50±0.25, OL 7.80±0.66 Description. Male. Body size small for Gryllinae species (Fig. 5A). In frontal view, head higher dorsoventrally, vertex rounded and convex (Fig. 9D). Dorsally viewed, occiput short, round and convex; vertex inclined. Frons convex and narrow (almost equal to the width of antennal scape), and arched in lateral view. A shallow groove present between bottom of frons and top of clypeus (along with epistomal suture). Labrum shield-like with lateral margins acutely convex and apically pointed round. End section of maxillary palpus conspicuously longer than the third; end section of labial palpus obviously longer than the total length of anterior two. Pronotum long and obviously longer than head. In dorsal view, pronotum somewhat trapezoid-like, anteriorly narrower than the posterior. Anterior margin of pronotum slightly concave and posteriorly straight. Disc flatten and possessing with a middle groove, and laterally fold down. Lateral lobes depressed posteriorly. Elytra broad and flatten, conspicuously wider than hind margin of pronotum. Basal field somewhat wide but short and almost as half length of pronotum. Oblique veins two. Diagonal vein arched. Mirror transverse and obviously wider than length. Dividing vein separating hind part oval-like. Apical field distally rounded, very short and somewhat a half length of mirror, and possessing two rows of irregular cells. Hind wings absent. Cercus long and almost as long as hind tibia. Outer tympana large and long oval-like; inner tympana absent and corresponding part pit-like. Hind femur slightly longer than tibia. Subapical spurs five pairs and inner ones obviously shorter than the outer proximally. Body pubescent, brown and ornamented with yellowish-brown and dark-brown patterns. Lateral areas of head and pronotum possessing continuous yellowish-brown stripes (Fig. 9D). Lateral lobes of pronotum almost dark brown with bottom margin yellowish-brown. Maxillary palpus colored dark and labial palpus colored light. Elytral disc yellowish-brown and most of lateral lobes colored dark brown. Fore and middle legs yellowish-brown and ornamented with brown patterns, and on femur and tibia ornamented with dark brown spots dorsally and laterally, and, especially, dark brown spots almost forming a ring on middle tibia. Hind legs and cercus colored brown and ornamented with dull yellowish brown. Abdomen dark brown dorsally and laterally, and ventrally colored yellowish-brown and ornamented with brown spots. Subgenital plate colored brown. Epiphallus almost without median lobe and slightly extended outward at the middle (Fig. 6A–C). Lateral lobes of epiphallus short and obtuse, the gap between them broad and somewhat rectangular. In lateral view, included angle between median and lateral lobe nearly arched. Ectoparamere short and slightly upward curved. In ventral view, ectoparamere comprise of two branches proximally, combined medially and curved upward. Female. Body size and coloration similar to male (Figs. 5B; 9C). Brachypterous, portions after mesonotum uncovered. Ovipositor long and slightly longer than cercus. Some samples macroperous and armed with normal elytra (Fig. 4B, D). Etymology. The name refers to its elytra armed with rounded margin apically, and its somewhat oval-like body. Remarks. Holotype and most specimens light brown and there are also some specimens that are darker. Compared with light ones, patterns or stripes of those dark samples are not clear or absent, and the coloration of legs and abdomen are darker. In addition, this species possesses rounded apical end of the tegmina.

Aphonoides aspidoid sp.­nov. (Figs. 12A; 16A–C) Examined­materials. Holotype.­ 1 male. China: Guangdong, Nanling, Pingdingshan, Aug. 6, 2019, He, Zhixin and Zhang, Tao coll. (SNNU); Paratypes.­ 2 males. China: Hunan, Langzhou, Mangshan, Oct. 6, 2004, Xie, Lingde coll. (SNNU). Measurements­ (mm).­ Male­ (n=3): ­ BL 13.92±2.74; HL 2.64±0.23; HW 3.65±0.57; PL 2.84±0.92; PW 3.74±0.53; FWL ­ 8.73±0.38; HFL 7.38±0.07; HTL 7.13±1.01 Etymology.­ The species name refers to the epiphallic lateral lobes blunt apically. Description.­Male: Head almost as wide as pronotum. Vertex broad and flattened, armed with densely pubescent. Frontal rostrum slightly convex in front view, inclined dorsally and ventrally. Median ocellus smallest of the three, ovoid; lateral ocellus transversely ovoid. End section of maxillary palpi almost equal to the length of the third; end section of labial palpi almost equal to the total length of remainder sections. Pronotum broad and flattened, anterior margin straight, posterior convex and middle of posterior margin angularly convex; median groove distinct. Longitudinal veins of tegmina paralleled. Hindwings longer than tegmina, uncovered portion about half length of hind femur. Inner tympanum shaped as elongate oval; outer tympanum absent. Hind tibiae evenly distributed with spines, armed with spurs distally (numbered 5:4); outer apical spurs two (equal in length) and the inner three (the dorsal one longest, two time longer than longest; middle one equal to length of longest; ventral one shortest). FIGURE­16.­ Genitalia of A. aspidoid sp.­nov. A. dorsal view; B. lateral view; C. ventral view. Genitalia.­ Lateral lobes of epiphallus trapezoidal. Median lobes of epiphallus small, apically acute. Ectoparamere proximally rod-like, not straight; apically expanded, shaped as baseball glove, with anterior side curved outward. Ectoparamere band-shaped in lateral view, included angle between ectoparamere and lateral lobes of epiphallus form an acute angle. Guiding rods shovel-like, apically acute. Coloration.­ Body brown. Clypeus and labrum yellowish. Tegmina brown. Female unknown. Remarks. This species resembles Aphonodies sabahi Gorochov, 2007 in features of body and genitalia, but are distinct in the epiphallus and ectoparamere. A. aspidoid sp.­nov. differs from A. sabahi in shape of ectoparamere, which of the new proximally not straight and what of the other is straight; ectoparamere of the new armed with outside margin apically concave and the other with inner margin concave. Besides, epiphallic lateral lobes apically inward truncated and what of the other outward truncated., Published as part of Zheng, Yanna, Xin, Zhixia, Xie, Lingde & Ma, Libin, 2021, - Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure, - 1878 - (Orthoptera: - Gryllidae) - from-China, pp. 401-424 in Zootaxa 4995 (3) on page 418, DOI: 10.11646/zootaxa.4995.3.1, http://zenodo.org/record/5056938, {"references":["Gorochov, A. V. (2007) Taxonomy of Podoscirtinae (Orthoptera: Gryllidae). Part 6. Indo-Malayan Aphonoidini. Zoosystematica Rossica, 15 (12), 237 - 289."]}

Genus Phyllotrella ­Gorochov,­1988 Phyllotrella planidorsalis Gorochov, 1988 Phyllotrella fumingi Sun & Liu, 2019, syn.­nov. Phyllotrella hainanensis Sun & Liu, 2019, syn.­nov. Phyllotrella transversa Sun & Liu, 2019, syn.­nov. Distribution: Hainan, Guangxi (Shiwandashan Forest), Guangdong (Nanling)., Published as part of Zheng, Yanna, Xin, Zhixia, Xie, Lingde & Ma, Libin, 2021, - Revision-of-the-cricket-subfamily-Podoscirtinae-Saussure, - 1878 - (Orthoptera: - Gryllidae) - from-China, pp. 401-424 in Zootaxa 4995 (3) on page 402, DOI: 10.11646/zootaxa.4995.3.1, http://zenodo.org/record/5056938, {"references":["Gorochov, A. V. (1988) New and little-known crickets of the subfamilies Landrevinae and Podoscirtinae (Orthoptera, Gryllidae) from Vietnam and certain other territories. In: Medvedev, L. N. & Striganova, B. R. (Eds.), Fauna i ekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam], pp. 5 - 21.","Sun, K. & Liu, H. Y. (2019) Review of the genus Phyllotrella Gorochov, with descriptions of three new species from China (Orthoptera: Gryllidae: Podoscirtinae: Podoscirtini). Zootaxa, 4629 (3), 441 - 447. https: // doi. org / 10.11646 / zootaxa. 4629.3.11"]}

Genus Progoniogryllus gen. nov. Type species: Progoniogryllus rotundus sp. nov., here designated. Diagnosis. Size small for Gryllinae species. Colored dark brown and ornamented with patches and stripes in light color, especially, above eyes there are backward forked stripes. Pronotum pubescent, armed with straight fore and hind margins and the former slightly narrower than the hind. Elytra short, far from supra-anal plate; basal and apical area of elytra rather short and almost similar in length; possessing two oblique veins and developed mirror; and media fold (fan) between apical part of lateral and dorsal area developed and expanded. Fore tibiae armed with outer tympanum and the inner one absent. Epiphallus almost without middle lobe and gap between lateral lobes rather broad. Ectoparamere simple and shaped as wide blade. Particularly, testis of the new unusually developed and occupies a large space in the abdominal cavity. Molecular phylogeny. To infer the taxonomic status of the new genus, we constructed phylogenetic relationship of two species of the new genus, 32 other Gryllinae species and three species of Gryllotalpidae (Fig. 2). The tree presented that the new taxa is included in the lineage of Modicogryllini and relevant species. Within the species we concluded, the new ones are much closer to Goniogryllus. Although they are distinct, it is significant that both the new and Goniogryllus bear similar coloration. Courtship behavior. If the male and female are placed in a box, the male will make a courtship call and approach female. The male will point the end of his abdomen at the female, and it can be observed that there is a spermatophore hanging at the end of the abdomen, but the female seems to avoid the male. Usually, the female's avoidance will not cause the male to give up, he will continue to get close to the female, constantly chase and point his tail up at the female in different positions. After courtship for about 7���15 minutes, the male keeps falling back and takes the opportunity to get under the female's body. After the external genitalia are connected, the copulation commences. During the mating, it is found that the female and the male twist at a certain angle (Fig. 3). The whole process lasts about 2 minutes, then the female leaves and the copulation ends. Etymology. The name refers to the position of the new in the phylogenic tree (fig. 2) in the article. Included species: Progoniogryllus rotundus sp. nov. and Progoniogryllus directus sp. nov. Remarks. External appearance of the new genus slightly resembles the genera Parasongella Otte, 1987, Callogryllus Sj��stedt, 1910 and Goniogryllus Chopard, 1936 since they both possess short elytra (or not) and bear yellowish stripes above eyes, but the tegmina of the new taxa are somewhat longer and possess an intact mirror, and the genitalia of the new taxa is distinct from the above. Parasongella includes three species, Parasongella ornaticeps (Chopard, 1969) (type locality: Thailand), Parasongella japonica Ichikawa, 2001 (type locality: Japan) and Parasongella nigriceps Otte, 1987 (type locality: Tanzania). Although considered in the same genus, their male genitalia are rather different from each other. The type species, P. nigriceps, bears genitalia that are very similar to Songella species and the clypeus is strongly upwards bent. The genitalic complex of P. japonica is very similar to the species of Goniogryllus, but that of P. ornaticeps is different from the former two. Thus, we do not think it warranted to combine those three species into Parasongella (Ichikawa, 2001). The male genitalia of the new genus is similar to the genus Gryllodes Saussure, 1874, and, their elytra is rather similar to each other. But the body of Gryllodes species is somewhat dorsoventrally compressed and that of the new genus is normal. Based on the phylogenic relationship which was established based on gene sequences, these two genera are distantly related., Published as part of Ma, Libin, Jing, Xuan & Zheng, Yanna, 2021, Integrative taxonomy base on morphology and molecular phylogeny with description of a new genus, Progoniogryllus gen. nov. and two new species (Orthoptera: Grylloidea: Gryllidae; Gryllinae), pp. 523-536 in Zootaxa 4995 (3) on pages 529-530, DOI: 10.11646/zootaxa.4995.3.7, http://zenodo.org/record/5057465, {"references":["Otte, D. (1987) African Crickets (Gryllidae). 9. New Genera and Species of Brachytrupinae and Gryllinae. Proceedings of the Academy of Natural Sciences of Philadelphia, 139, 315 - 374.","Sjostedt, Y. (1910) 17. Orthoptera. 5. Gryllodea. In: Sjostedt, Y. (Ed.), Wissenschaftliche ergebnisse der Schwedischen zoologischen expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906, unter leitung von prof. dr. Yngve Sjostedt. Tryckt hos P. Palmquists aktiebolag, Stockholm, pp. 91 - 124. https: // doi. org / 10.5962 / bhl. title. 1805","Chopard, L. (1936) The Tridactylidae and Gryllidae of Ceylon. Ceylon Journal of Science (Biological Science), 20, 9 - 87.","Chopard, L. (1969) Orthoptera. Vol. 2. Grylloidea. In: Sewell, R. B. S. (Ed.), The Fauna of India and the Adjacent Countries. Baptist Mission Press, Calcutta, pp. 1 - 421.","Ichikawa, A. (2001) New species of Japanese crickets (Orthoptera: Grylloidea) with notes on certain taxa. Tettigonia, 3, 45 - 58.","Saussure, H. d. (1874) Etudes sur les insectes Orthopteres, famille des Gryllides. Mission scientifique au Mexique et dans l'Amerique centrale, 6, 296 - 515."]}

Svercacheta semiobscurus (Chopard, 1961) comb. nov. (Figs. 12A, B; 13A–C) Gryllus semiobscurus Chopard, 1961: 273; Chopard, 1969: 67 Modicogryllus semiobscurus, Chopard, 1967: 81; Bhowmik, 1985: 186; Ingrisch, 1998: 339 Holotype information. Type locality: India, Assam, Ledo. Deposited at the Academy of Natural Sciences, Philadelphia, US (not examined). Measurements. Male (n=5): BL 12.73±0.52, HL 2.09±0.12, HW 3.58±0.19, PL 2.32±0.13, PW 3.97±0.36, FWL 7.48±0.38, FWW 4.18±0.25, MTL 2.83±0.27, HFL 7.35±0.28 Description. Male. Body sized medium in Gryllinae species. Head almost as long as pronotum. Occiput broad and flatten, vertex weakly inclined. Frons rounded and convex, and slightly wider than antennal scape. Antennal scape transverse and somewhat rounded. Postclypeus (top portion of clypeus) band-like with anteclypeus (lower portions) depressed. Labrum shield-like, laterally convex and apically rounded. End section of maxillary palpus longer than the third; end section of labial palpus conspicuously longer than the total length of proximal two. Pronotum transverse, broad and flattened dorsally, slightly longer than the half of width, and laterally inclined and folding down; lateral lobes somewhat flattened with lower margins inward depressed. Elytra short and not reaching abdominal tip. Elytral surface tapering with lateral margins almost arched. Basal field small and short, which nearly as half-length of pronotum. Diagonal vein straight and almost fusing a line with the outer margin of mirror. Mirror longer than width and slightly shaped as rectangular, with bottom pointed round. Apical field very short and shorter than mirror. Hind wings present and long, but not longer than cercus. Cercus thin and long, and almost equal to hind femur. Outer tympana large and longitudinal ovoid; inner tympanum small and ovoid. Hind femur conspicuously longer than tibia. Subapical spurs six pairs, short and small, and the inner proximal one always absent but the corresponding one in outside present. Hind tarsus armed with two rows of dorsal spines and outside ones more than the inner ones. Coloration. Body colored yellowish-brown and ornamented with dark brown patterns. Occiput yellowishbrown and ornamented with seven large brown spots.Area between eyes ornamented with a broad dark brown band, and a yellowish-brown line between lateral ocelli. Frons almost dark brown and only median ocellus and around portions yellowish-brown. The middle of postclypeus (dorsal part of clypeus) brown and gradually changing to yellowish-brown; anteclypeus (lower portion of clypeus), including labrum colored yellowish-brown. Mandible yellowish-brown and colored dark apically. Pronotum yellowish brown and ornamented with dark brown spots; pronotal lateral lobes dark brown and about 1/3 lower portions yellowish-brown. Elytra yellowish-brown and ornamented with brown stripes laterally. Abdomen dark brown dorsally and laterally, and yellowish brown ventrally. Subgenital plate colored brown and cercus yellowish brown. Legs yellowish brown and ornamented with few of dark spots, and hind femur ornamented with dark spots and stripes dorsally and outward. Genitalia. Epiphallus without median lobe, and lateral lobes far away from each other and the gap between them somewhat shaped as rectangular. In lateral view, epiphallic lateral lobe obviously thick and short, and armed with a short and obtuse papillary. Ectoparamere sheet-like and curved upward along with epiphallic lateral lobes. Female unknown. Materials examined. 1 male, Guangdong, Nanling, viii-23-2010, Wang, Manqiang coll. (SNNU); 5 males, Hainan, Wuzhishan, 690m, viii-2-2009, Ma, Libin coll. (SNNU); 1 male, Yunnan, Mengla, Yaoqu, 794m, vi-11- 2009, Ma, Libin coll. (SNNU); 1 male, Yunnan, Mengla, Yaoqu, 794m, v-31-2009, Ma, Libin coll. (SNNU); 1 male, Yunnan, Mengla, Yaoqu, 794m, v-29-2009, Ma, Libin coll. (SNNU); 3 males, Yunnan, Mengla, Shangyong, Longmen, 870m, v-18-2013, Ma, Libin coll. (SNNU); 1 male, Yunnan, Mengla, Shangyong, Longmen, 870m, v- 16-2013, Ma, Libin coll. (SNNU); 1 male, Yunnan, Mengla, Yaoqu, 794m, v-22-2013, Ma, Libin coll. (SNNU); 1 male, Yunnan, Jinping, Mengla, Xinmeng, 450m, v-3-2013, Ma, Libin coll. (SNNU). Remarks. Coloration of this species varied from light to dark. The specimen we describe here is colored light and those darker possess similar patterns with but distinct in ground color. Type locality of M. semiobscurus is India. Chopard (1969) described the holotype and Ingrisch (1998) identified specimens from Thailand as M. semiobscurus and noted differences between specimens from India and Thailand. These differences are mainly in the angle between epiphallic proximal part and lateral lobe, and features of basal part of epiphallus and lateral lobes of epiphallus. According to description and illustration from Chopard (1969) and Ingrisch (1998), we compared male’s genitalia of our specimens with them, epiphallic lateral lobes of the ours are somewhat shorter and thicker (lateral viewed), and the angle between the basal of epiphallus and lateral lobe as well as degree of bending of epiphallic lateral lobes are slightly different., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on pages 242-244, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Chopard, L. (1961) Les divisions du Genre Gryllus basees sur l'etude de l'appareil copulateur (Orthoptera: Gryllidae). \" EOS', Revista espanola de entomologia Tomo 37 cuaderno 3, Institute espanol de entomologia Madrid, 267 - 287.","Chopard, L. (1969) Orthoptera. Vol. 2. Grylloidea. Baptist Mission Press, Calcutta, 421 pp.","Chopard, L. (1967) Gryllides. Fam. Gryllidae; Subfam. Gryllinae (Trib. Grymnogryllini, Gryllini, Gryllomorphini, Nemobiini). Uitgeverij Dr. W. Junk, Gravenhage, 213 pp.","Bhowmik, H. K. (1985) Contribution to the Gryllid fauna of the Western Himalayas. Records of the Zoological Survey of India, 73, 1 - 84.","Ingrisch, S. (1998) The genera Velarifictoms, Modicogryllus and Mitius in Thailand (Ensifera: Gryllidae, Gryllinae). Entomologica Scandinavica, 29, 315 - 359. https: // doi. org / 10.1163 / 187631298 X 00122"]}

Ma, Libin, Zheng, Yanna, Qiao, Min (2021): Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae). Zootaxa 4990 (2): 227-252, DOI: https://doi.org/10.11646/zootaxa.4990.2.2

Eumodicogryllus vicinus (Chopard, 1968) comb. nov. Tartarogryllus vicinus Chopard, 1968: 275, illustr. 1 Modicogryllus (Eumodicogryllus) vicinus, Gorochov, 1986: 3 Holotype information. Type locality: Afghanistan, Herat, Bala Murghab. Deposited at the Moravské zemské Museum, Brno, Czech (not examined). Remarks: When proposed this species, Chopard (1968: 276) indicated that it is difficult to distinguish it from E. bordigalensis (= burdigalensis) unless using male’s genitalia to identify them. When established the subgenus Eumodicogryllus, Gorochov (1986) arranged this species in the taxon, but it is not included in the elevated genus Eumodicogryllus now (Cigliano, et al., 2021). According to features of its epiphallus (referring to Chopard, 1968: 275, Fig. 1), this species should be placed in Eumodicogryllus as done by Gorochov (1986) because it does not possess the characters of Tartarogryllus.

Modicogryllus (Promodicogryllus) nigericus (Chopard, 1961) comb. nov. Tartarogryllus nigericus Chopard, 1961: 278, illustr. 5 Holotype information. Type locality: Nigeria, Ibadan. Deposited at the Natural History Museum (BMNH), London, U.K. (not examined). Remarks. Chopard (1961) compared this species with Eumodicogryllus bordigalensis (= Tartarogryllus bordigalensis) and M. conspersus, and he indicated the genitalia of it is more similar to E. bordigalensis than M. conspersus, but this species is more like the species of the subgenus Promodicogryllus than the species E. bordigalensis. Thus, we transfer it here., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on page 234, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Chopard, L. (1961) Les divisions du Genre Gryllus basees sur l'etude de l'appareil copulateur (Orthoptera: Gryllidae). \" EOS', Revista espanola de entomologia Tomo 37 cuaderno 3, Institute espanol de entomologia Madrid, 267 - 287."]}

Svercacheta siamensis (Chopard, 1961) (Figs. 1 C, G; 2A, B; 10A, B; 11A–F) Modicogryllus siamensis Chopard, 1961: 280, plate 7; Ingrisch, 1998: 338 Modicogryllus confirmatus, Bhowmik, 1977: 230, 233, misidentification; Vasanth, 1993: 63, 172, misidentification; Ingrisch, 1998: 336 Tartarogryllus minusculus, Yin & Liu, 1995: 66, misidentification; He, 2018: 529 Modicogryllus (Lepidogryllus) siamensis, Ichikawa, Murai & Honda, 2000: 262 Lepidogryllus siamensis, Kim, 2013: 301 Svercacheta siamensis, Gorochov, 2017: 28; He, 2018: 520, 529 Modicogryllus nigrivertex Kaltenbach, 1979 (= Svercacheta nigrivertex, Gorochov, 1993: 87), synonymized by Gorochov, 2017: 28 Modicogryllus pacificus Otte, 1994, synonymized by Ingrisch, 1998: 339 Holotype information. Type locality: Sri Lanka. Deposited at the Natural History Museum (BMNH), London, U.K. (not examined). Measurements. Male (n=3): BL 12.32±0.70, HL 1.87±0.10, HW 3.33±0.08, PL 2.19±0.37, PW 3.56±0.16, FWL 6.99±0.20, FWW 3.82±0.13, MTL 2.81±0.06, HFL 6.71±0.09 Description. Male. Body sized medium in Gryllinae species. Head longer than pronotum. Occiput somewhat broad and flatten, vertex slightly inclined. Frons somewhat convex and about 1.5 times of antennal scape. Antennal scape transverse wide and almost shaped as rectangular. Clypeus depressed and conspicuously lower than frons; Postclypeus (top portion of clypeus) longitudinally narrow and band-shaped. Mandible strong. End section of maxillary palpus equal to the length of the third; end section of labial palpus almost equal to the total length of remainder two. Pronotum transverse and almost two times wider than length.Anterior margin of pronotum slightly concave and posteriorly straight, with lateral margins slightly inclined and fold down. Lateral lobes of pronotum almost flatten with anterior and posterior slightly concave. Basal field of elytra very small and as 2/3 length of pronotum. Oblique veins two. Diagonal vein nearly straight. Mirror obviously longer than width, somewhat rectangular-like but bottom margin pointed round; dividing vein separating hind portion almost triangular-like. Apical field nearly equal to the length of mirror. Hind wings present and uncovered portion obviously longer than cercus. Cercus long and longer than hind tibia. Outer tympana large and longitudinal oval-like; inner tympana rather small and oval-like. Hind tibia almost equal to the length of hind femur. Subapical spurs six pairs and inner proximal one always absent. Coloration. Body colored brown and ornamented with dark brown on head and pronotum. Occiput light brown and ornamented with six fine longitudinal stripes colored yellowish-brown. Area between eyes dark brown and a thin yellowish-brown line between lateral ocelli. Areas between frons and clypeus dark brown, but median ocellus yellowish-brown and a longitudinal fine stripe under median ocellus which colored yellowish-brown. Pronotum dark brown and distributed yellowish-brown spots on the middle and posterior; lateral lobe of pronotum dark brown with bottom margin yellowish-brown. Elytra yellowish-brown and hind wings almost white. Abdomen dorsally and laterally dark brown and ventrally yellowish-brown, but subgenital plate and cercus colored brown. Fore legs almost yellowish-brown and hind legs colored darker, hind femur ornamented with oblique stripes outside. Genitalia. Epiphallus without media lobe, lateral lobes far from each other and jointed by a transverse bridge. Inner margin of epiphallic lateral lobe straight. In lateral view, lateral lobes upward pointed and acute apically. Ectoparamere developed and, in dorsal view, the tips forked. In lateral view, ectoparamere curved dorsally and always touching epiphallic lateral lobes. Females Very similar with male. Apex of ovipositor slightly smooth (Figs. 1C, G). Materials examined. male, Hainan, Yinggeling, viii-4-2010, Jiang, Chaozhong coll. (SNNU); male, Guangdong, Fengkai, Heishiding, x-4-2014, Zhang, Tao coll. (SNNU); male, Sichuan, Chuanshan, x-7-2015, Liu, Yang coll. (SNNU). Remarks. This species was used to identified as Tartarogryllus minusculus Walker, 1869 in China for a long time (He, 2018; Yin & Liu, 1995). Gorochov (2017) considered this species is a synanthropic species and widely distributed in many tropical regions, such as Thailand, the Andaman Islands, Hawaii, Vietnam, Cambodia, South China, continental India, Sri Lanka, South Iran and Oman., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on pages 240-242, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Chopard, L. (1961) Les divisions du Genre Gryllus basees sur l'etude de l'appareil copulateur (Orthoptera: Gryllidae). \" EOS', Revista espanola de entomologia Tomo 37 cuaderno 3, Institute espanol de entomologia Madrid, 267 - 287.","Ingrisch, S. (1998) The genera Velarifictoms, Modicogryllus and Mitius in Thailand (Ensifera: Gryllidae, Gryllinae). Entomologica Scandinavica, 29, 315 - 359. https: // doi. org / 10.1163 / 187631298 X 00122","Bhowmik, H. K. (1977) Study on some Indian crickets with new distributional records of the subfamily Gryllinae (Gryllidae, Orthoptera). Records of the Zoological Survey of India, 73, 229 - 238.","Vasanth, M. (1993) Studies on crickets (Insecta: Orthoptera: Gryllidae) of Northeast India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 132, 1 - 178.","Yin, H. S. & Liu, X. W. (1995) Synopsis of the Classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific and Technological Literature Publishing House Press, Shanghai, 237 pp.","He, Z. Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4","Ichikawa, A., Murai, T. & Honda, E. (2000) Monograph of Japanese crickets (Orthoptera: Grylloidea). Bulletin Hoshizaki Green Found, 4, 257 - 332.","Kim, T. W. (2013) A taxonomic study on the burrowing cricket genus Velarifictorus with morphologically resembled genus Lepidogryllus (Orthoptera: Gryllidae: Gryllinae) in Korea. Animal Systematics, Evolution and Diversity, 29, 294 - 307. https: // doi. org / 10.5635 / ASED. 2013.29.4.294","Gorochov, A. V. (2017) Order Orthoptera, superfamily Grylloidea. Arthropod fauna of the UAE, 6, 21 - 35.","Kaltenbach, A. P. (1979) Ergebnisse der osterreichisch-indischen Andamen-Expedition 1976. Saltatoria (Insecta-Orthopteroida). Annalen des Naturhistorischen Museums in Wien Serie B Botanik und Zoologie, 82, 605 - 614.","Gorochov, A. V. (1993) Grylloidea (Orthoptera) of Saudi Arabia and adjacent countries. Fauna of Saudi Arabia, 13, 79 - 97.","Otte, D. (1994) OSF 1: Additions and Corrections-October. Journal of Orthoptera Research, 3, 86. https: // doi. org / 10.2307 / 3503412","Walker, F. (1869) Catalogue of the specimens of Dermaptera Saltatoria and supplement to the Blattariae in the collection of the British Museum. Printed for the trustees of the British Museum, London, 224 pp."]}

Modicogryllus (Modicogryllus) latefasciatus (Chopard, 1933) Gryllus latefasciatus Chopard, 1933: 2 Modicogryllus latefasciatus, Chopard, 1961: 275; Gorochov, 1996: 109 Velarifictorus latefasciatus, Yin & Liu, 1995: 61, misidentification Velarifictorus (Velarifictorus) micado, He, 2018: 530, misapplied synonymy Holotype information. Type locality: China, Sichuan Province (Kipa. N.O., Szechuan. Sven Hedins Exp. Ctr. Asien. Dr. Hummel). Deposited at the Swedish Royal Museum of Natural History (Naturhistoriska Riksmuseet, NHRM) Stockholm, Sweden (not examined). Remarks. Chopard (1933) described this species under the genus of Gryllus and moved it into the genus of Modicogryllus (Chopard, 1961), however, in his studies, he did not produce any graphic or describe the features of male genitalia. Gorochov (1996) examined the holotype and he described and illustrated this species. He indicated that the head of this species slightly resembles to the species of the genus Velarifictorus. It might since the similar reason, Yin and Liu (1995) arranged some Velarifictorus specimens (Yin & Liu, 1995: illustr. 182–184) under this name and moved this species from Modicogryllus to the genus Velarifictorus. According to the work of Yin and Liu, He (2018) considered the species Velarifictorus latefasciatus (Chopard, 1933) was junior synonym of Velarifictorus micado (Saussure, 1877). This is a mistake made over and over again and M. latefasciatus is a Modicogryllus species but not of Velarifictorus. This species has not been rediscovered by Chinese for so many years and it is also difficult for us to find it since the type locality is only recorded with the name of Province and without any more information., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on page 230, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Chopard, L. (1933) Swedish-Chinese scientific expedition to the northwestern provinces of China, under the leadership of Dr. Sven Hedin and Prof. Su Ping-chang. Insects collected by the Swedish expedition doctor Dr. David Hummel 1927 - 1930. 4. Orthoptera. 3. Gryllidae. Arkiv for Zoologi, 25 B, 1 - 4.","Chopard, L. (1961) Les divisions du Genre Gryllus basees sur l'etude de l'appareil copulateur (Orthoptera: Gryllidae). \" EOS', Revista espanola de entomologia Tomo 37 cuaderno 3, Institute espanol de entomologia Madrid, 267 - 287.","Gorochov, A. V. (1996) New and little known crickets from the collection of the Humboldt University and some other collections (Orthoptera: Grylloidea). Part 1. Zoosystematica Rossica, 4, 81 - 114.","Yin, H. S. & Liu, X. W. (1995) Synopsis of the Classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific and Technological Literature Publishing House Press, Shanghai, 237 pp.","He, Z. Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4","Saussure, H. de (1877) Melanges orthopterologiques V. fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 25, 169 - 504. https: // doi. org / 10.5962 / bhl. title. 8541"]}

Modicogryllus (Promodicogryllus) confirmatus (Walker, 1859) Acheta confirmata Walker, 1859: 221 Gryllus confirmatus, Walker, 1869: 37; Chopard, 1936: 25; Chopard, 1969: 41 Modicogryllus confirmatus, Chopard, 1961: 275; Gorochov, 1986: 3; Yin & Liu, 1995: 65; Ingrisch, 1998: 335 Svercacheta siamensis, Bhowmik, 1977: 230, 233, misidentification; Vasanth, 1993: 63, 172, misidentification. Holotype information. Type locality: Sri Lanka. Deposited at the Natural History Museum (BMNH), London, U.K. (not examined). Remarks. Ingrisch (1998) examined the holotype and suggested that those records of M. confirmatus from outside of holotype locality should be confirmed by reexamination, for example, Indian species identified by Bhowmik (1977) and Vasanth (1993) are actually Svercacheta siamensis (Chopard, 1961). This species was recorded in China by Yin and Liu (1995) and we believed that most Chinese species were identified using the references of Chopard (e.g. Chopard, 1969), however, he regarded M. consobrinus as junior synonym of M. confimatus by mistake (Chopard, 1961; Chopard, 1969; Ingrisch, 1998). According to illustration of holotype’s genitalia, Chinese species should be under the name M. consobrinus and, therefore, this species is not found in China now., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on page 232, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Walker, F. (1859) Characters of some apparently undescribed Ceylon Insects. Annals and Magazine of Natural History, London, 3 - 4, 217 - 224. https: // doi. org / 10.1080 / 00222935908697111","Walker, F. (1869) Catalogue of the specimens of Dermaptera Saltatoria and supplement to the Blattariae in the collection of the British Museum. Printed for the trustees of the British Museum, London, 224 pp.","Chopard, L. (1936) The Tridactylidae and Gryllidae of Ceylon. Ceylon Journal of Science, Biological Science, 20, 9 - 87.","Chopard, L. (1969) Orthoptera. Vol. 2. Grylloidea. Baptist Mission Press, Calcutta, 421 pp.","Chopard, L. (1961) Les divisions du Genre Gryllus basees sur l'etude de l'appareil copulateur (Orthoptera: Gryllidae). \" EOS', Revista espanola de entomologia Tomo 37 cuaderno 3, Institute espanol de entomologia Madrid, 267 - 287.","Gorochov, A. V. (1986) New and little known Grylloidea (Orthoptera) from Middle Asia and adjacent territories. Proceedings of Zoological Institute of Academy of Sciences of USSR, 140, 3 - 15.","Yin, H. S. & Liu, X. W. (1995) Synopsis of the Classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific and Technological Literature Publishing House Press, Shanghai, 237 pp.","Ingrisch, S. (1998) The genera Velarifictoms, Modicogryllus and Mitius in Thailand (Ensifera: Gryllidae, Gryllinae). Entomologica Scandinavica, 29, 315 - 359. https: // doi. org / 10.1163 / 187631298 X 00122","Bhowmik, H. K. (1977) Study on some Indian crickets with new distributional records of the subfamily Gryllinae (Gryllidae, Orthoptera). Records of the Zoological Survey of India, 73, 229 - 238.","Vasanth, M. (1993) Studies on crickets (Insecta: Orthoptera: Gryllidae) of Northeast India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 132, 1 - 178."]}

Genus Turanogryllus Tarbinsky, 1940 Type species: Gryllus lateralis Fieber, 1853, Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on page 244, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Tarbinsky, S. P. (1940) The Saltatorian Orthopterous Insects of the Azerbaidzhan S. S. R. Academy of Sciences in the Azyerbaidjankoi S. S. R., Moscow, Leningrad, 245 pp.","Fieber, F. (1853) Synopsis der europaischen Orthoptera mit besonderer Rucksicht auf die in Bohmen vorkommenden Arten. Lotos, 3, 90 - 104 + 115 - 129 + 138 - 154 + 168 - 176 + 184 - 188 + 201 - 207 + 232 - 238 + 252 - 261."]}

Genus Gryllodinus Bolívar, 1927 Gryllodes (Gryllodinus) Bolívar, 1927: 102 Eugryllodes, synonymized by Chopard, 1967: 111 Gryllodinus, Gorochov, 1979: 92 Type species: Gryllodes kerkennensis Finot, 1893 Included species: Gryllodinus kerkennensis (Finot, 1893), Gryllodinus odicus (Uvarov, 1911), Gryllodinus abditus Gorochov, 1979 and G. jijilacus sp. nov. Remarks. The distinct feature of Gryllodinus species is the clypeus is very large and strongly convex. Three species of the genus were recorded worldwide (Gorochov, 1979). Although they possess similar appearance with each other (Gorochov, 1979; Gorochov, 1986; Uvarov, 1911), the male genitalia are distinct (Gorochov, 1979). Besides overall similarity in this genus, the species also resemble to some species of Modicogryllus and Turanogryllus, from arid areas. G. odicus for example, Uvarov (1911) indicated that it is similar with G. kerkennensis and Turanogryllus lateralis (Fieber, 1853) in coloration and pattern of head and pronotum; and Gorochov (1979) indicated that it similar to M. pallipalpis, M. latefasciatus, M. pseudocyprius, M. nitidus and M. algirius, and very similar to M. pseudocyprius and M. algirius. In this study, the specimens of the genus found in China rather resemble to the species T. lateralis. And we used to confuse these two species so much that when we went to Xinjiang to collect our samples, we mistakenly assumed that the specimens of T. lateralis was the species of Gryllodinus we were looking for. Here is the genus Gryllodinus is initially introduced to China., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on pages 234-235, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Bolivar, I. (1927) Datos complementarios sobre los Ortopteros de la Peninsula Iberica. II. Grilidos. Boletin de la Real Sociedad Espanola de Historia Natural, 27, 96 - 110.","Chopard, L. (1967) Gryllides. Fam. Gryllidae; Subfam. Gryllinae (Trib. Grymnogryllini, Gryllini, Gryllomorphini, Nemobiini). Uitgeverij Dr. W. Junk, Gravenhage, 213 pp.","Gorochov, A. V. (1979) To the knowledge of crickets (Orthoptera, Gryllidae) from the fauna of Middle Asia. Entomologicheskoe Obozrenie, 58, 92 - 98.","Finot, A. (1893) Descriptions abregees de huit especes nouvelles d'Orthopteres habitant l'Algerie et la Tunisie. Bulletin de la Societe Entomologique de France, 62, 250 - 253.","Uvarov, B. P. (1911) Contribution a la faune des Orthopteres de la Steppe des Kirghises. Russkoe entomologicheskoe obozrenie, 11, 425 - 429.","Gorochov, A. V. (1986) New and little known Grylloidea (Orthoptera) from Middle Asia and adjacent territories. Proceedings of Zoological Institute of Academy of Sciences of USSR, 140, 3 - 15.","Fieber, F. (1853) Synopsis der europaischen Orthoptera mit besonderer Rucksicht auf die in Bohmen vorkommenden Arten. Lotos, 3, 90 - 104 + 115 - 129 + 138 - 154 + 168 - 176 + 184 - 188 + 201 - 207 + 232 - 238 + 252 - 261."]}

Gryllodinus jijilacus sp. nov. (Figs. 5A–C; 6A–C) Type materials. Holotype. China. Male, Xinjiang, Jijihu, vi-6-2005, Han, Huilin coll. (SNNU). Paratype, male, data similar with holotype. Measurement. Male (n=2): BL 11.92±0.5, HL1.72±0.24, HW 2.81±0.12, PL1.55±0.04, PW 3.23±0.14, FWL 8.11±0.17, FWW 4.15±0.3, MTL 2.42±0.19, HFL 5.94±0.19 Description. Male. Body sized medium in Gryllinae species. Head and pronotum short, almost as long as apical field of elytra. Occiput short and slightly convex. Vertex inclined. Clypeus very convex proximally and shaped as hemisphere in lateral view; and truncate apically, which oblique inward. Labrum shaped normally and with middle notch apically. Eyes large, its hind and lower margin straight. Rostrum very wide and nearly four times wider than antennal scape. Antennal scape transverse, rectangle-like. Apical three sections of maxillary palpus in equal length; end section of labial palpus slightly longer than the total length of proximal two. Pronotum transverse, two times wider than length; its anterior margin inward concave and almost shaped as obtuse angle, posterior margin straight and lateral margin curved to lateral lobes; lateral lobes of pronotum somewhat flat and conspicuously inward curved posteriorly. Elytral long and longer than three times of the total length of head and pronotum. Lateral margins of elytral almost straight. CuA vein long and the transvers portion broadly arched. Harp area large and harp veins two. Diagonal vein somewhat arc-like. Mirror large and slightly oval shaped. Dividing vein arched and dividing anterior portion larger than the posterior. Apical field rather short, almost as long as the half of mirror and possessing with few of cell. Hindwings present and long, with uncovered portion as long as forewings. Fore tibia only armed with outer tympana, large and shaped as longitudinal oval; the part of inner tympana should present depressed and somewhat crescent. Tarsus very short and as half-length of tibia. Hind tibia short, as half-length of hind femur; subapical spurs five or six, those presenting apically or proximally varied in size. Hind proximal tarsus armed with two rows of spines, numbered five or six respectively and right-left asymmetry. Cercus short and nearly as long as hind tibia. Coloration. Body colored yellowish brown and ornamented with brown spots or stripes. Vertex and occiput possessing brown and transvers stripes, and several irregular transvers stripes presenting between them. Frons armed with a brown and oval-shaped patch in the middle, which surrounded by ocelli and eyes. Eyes dark brown. Clypeus almost yellowish brown and just armed with few of brown patch under low margin of antennal scape. Pronotum yellowish brown and only colored brown in blade-shaped patterns and portions around them. Fore and middle legs yellowish brown; most parts of hind legs yellowish brown, but dorsum of hind femur ornamented with brown spots medially and apically, and spurs on tibia, spines on tarsus as well as claw colored brown apically. Abdominal tergites and thoracic pleura dark brown, and ventral portions yellowish brown. Genitalia. Genitalia very similar to Modicogryllus species. Epiphallus widened, with thin lateral lobes and without middle lobe. Epiphallic lateral lobe proximally broad, narrowed at middle and tapering and with acute end. The middle of epiphallus very wide and somewhat longitudinal long, with inner (proximal) part shaped as obtuse angle. Ectoparamere proximally stick-like, middle portions widened and armed with lateral lobe, and apical portions depressed laterally, inwards curved and with acute tip. Female unknown. Etymology. The name refers to the locality where discovered it. Remarks. This species possesses genitalia which is very similar to the species of Modicogryllus, but in Modicogryllus the epiphallus and ectoparamere are shaped simply, and those of the new are varied in thickness and appendant. Of the Modicogryllus, the new one bears endoparamere resembling Modicogryllus pseudocyprius Gorochov, 1996. Then, there are three species of the genus Gryllodinus Bolívar, 1927 worldwide and male’s genitalia of all of these species are rather distinct, and that of the new species is similar to Gryllodinus odicus (Uvarov, 1911) (Gorochov, 1979: figs 7–9). Comparing with G. odicus, epiphallic lateral lobes of the new rather outwardly expanded proximally and the constricting portion in the middle papillary, and these of G. odicus are simple. In addition, the new species owns similar coloration with Turanogryllus lateralis (Fieber, 1853), it might be the result of the similarity of habitat environment.

Turanogryllus rufoniger (Chopard, 1925) (Figs. 1A, E; 2D; 17A–C; 18A–D) Gryllodes rufoniger Chopard, 1925: 292 Paragryllopsis rufoniger, Chopard, 1963: 169 Turanogryllus rufoniger, Chopard, 1967: 36; Chopard, 1969: 73; Yin and Liu, 1995: 64; Ma, 2011: 68 Holotype information. Type locality: Laos, Vientiani. Deposited at the Muséum d’histoire naturelle de Genève (MNHN), Paris, France (not examined). Materials examinzed. 1 male, Yunnan, Xishuangbanna, Menglun, 1000m, iv-3-2009, Liu, Xianwei et al. coll. (SEM); 1 male, Hainan, Maoyang, Maoxing, 325m, viii-5-2009, Ma, Libin coll. (NWAFU); 2 males, Yunnan, Banna botanical garden, x-19-2014, Zhang, Tao coll. (SNNU); male, Yunnan, Jinping, Mengla, 385m, v-1-2013, Ma, Libin coll. (SNNU); 3 males, Yunnan, Mengla, Menglun, x-12–23, Zhang, Tao coll. (SNNU); male, Yunnan, Mengla, Shangyong, Longmen, 996m, v-13-2013, Ma, Libin coll. (SNNU); male, Yunnan, Cangyuan, Banlao, 1134m, vi-5- 2013, Ma, Libin coll. (SNNU); 2 males, Yunnan, Cangyuan, Banlao, 1170m, vi-5-2013, Ma, Libin coll. (SNNU); female, Yunnan, Banna botanical garden, x-19-2014, Zhang, Tao coll. (SNNU); female, Yunnan, Mengla, Menglun, 690m, v-23-2009, Ma, Libin coll. (SNNU); female, Yunnan, Mengla, Menglun, 690m, v-28-2009, Ma, Libin coll. (SNNU); female, Yunnan, Mengla, Menglun, x-12–23, Zhang, Tao coll. (SNNU); female, Yunnan, Jinping, Mengla, 385m, iv-30-2013, Ma, Libin coll. (SNNU)., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on pages 247-248, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Chopard, L. (1925) Descriptions de Gryllides nouveau (Orthopteres). Annales De La Societe Entomologique De France, 94, 291 - 332.","Chopard, L. (1963) Notes sur la faune des Orthopteroides d'Israel et de quelques regions limitophes (Blattodea, Gryllacridoidea, Gryllodea). Bulletin Research Council of Israel. B 11, 161 - 176.","Chopard, L. (1967) Gryllides. Fam. Gryllidae; Subfam. Gryllinae (Trib. Grymnogryllini, Gryllini, Gryllomorphini, Nemobiini). Uitgeverij Dr. W. Junk, Gravenhage, 213 pp.","Chopard, L. (1969) Orthoptera. Vol. 2. Grylloidea. Baptist Mission Press, Calcutta, 421 pp.","Yin, H. S. & Liu, X. W. (1995) Synopsis of the Classification of Grylloidea and Gryllotalpoidea from China. Shanghai Scientific and Technological Literature Publishing House Press, Shanghai, 237 pp.","Ma, L. B. (2011) Systematic study on the Gryllidae from China (Orthoptera: Grylloidea). In: College of Plant Protection. Vol. Doctor. Northwest A & F University, Yangling, pp. 263."]}

Turanogryllus koshunensis (Shiraki, 1911) Gryllodes koshunensis Shiraki, 1911: 61 Gryllopsis koshunensis, Chopard, 1967: 33 Velarifictorus koshunensis, Kim, 2012: 144 Turanogryllus koshunensis, He, 2018: 529 Holotype information. Type locality: China, Taiwan, Koshun. Deposited at Taipei University, Taiwan, China (not examined). Remarks. This species seems to be based on the fact that Shiraki compared this species with Velarifictorus asperses (= Gryllodes berthellus Saussure, 1877) (Walker, 1869) when he established it. Dr. Kim, Tae-Woo and Dr. Yang, Jeng-Tze considered it a species of Velarifictorus in their personal communication (Kim, 2012). Orthoptera species file online (Cigliano, et al., 2021) presents a series of photos of the holotype deposited in Taipei University. According to those pictures and morphological characters, both He (2018) and I think it should be in Turanogryllus. When note that T. eous, was mentioned by Bey-Bienko (1956) and indicated the species under the name Gryllodes koshunensis from Taiwan Island is outwardly similar to T. eous., Published as part of Ma, Libin, Zheng, Yanna & Qiao, Min, 2021, Revision of Chinese crickets of the tribe Modicogryllini Otte & Alexander, 1983 with notes on relevant taxa (Orthoptera: Gryllidae; Gryllinae), pp. 227-252 in Zootaxa 4990 (2) on page 245, DOI: 10.11646/zootaxa.4990.2.2, http://zenodo.org/record/5026190, {"references":["Shiraki, T. (1911) Monographie der Grylliden von Formosamosa, mit der Uebersicht der Japanischen Arten. Generalgouverne- ment von Formosa, Taihoku (Taipei), 129 pp.","Chopard, L. (1967) Gryllides. Fam. Gryllidae; Subfam. Gryllinae (Trib. Grymnogryllini, Gryllini, Gryllomorphini, Nemobiini). Uitgeverij Dr. W. Junk, Gravenhage, 213 pp.","Kim, T. W. (2012) First Record of the Field-Cricket Turanogryllus eous (Orthoptera: Gryllidae: Gryllinae) from Korea. Animal Systematics, Evolution and Diversity, 28, 140 - 144. https: // doi. org / 10.5635 / ASED. 2012.28.2.140","He, Z. Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4","Saussure, H. de (1877) Melanges orthopterologiques V. fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 25, 169 - 504. https: // doi. org / 10.5962 / bhl. title. 8541","Walker, F. (1869) Catalogue of the specimens of Dermaptera Saltatoria and supplement to the Blattariae in the collection of the British Museum. Printed for the trustees of the British Museum, London, 224 pp.","Bey-Bienko, G. Y. (1956) Notes on fauna and taxonomy of Gryllidae (Orthoptera) from China. Zoologicheskii Zhurnal, 35, 219 - 237."]}