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The following M. hastatus samples from Martinique are kept in the Paris Museum collection: Material examined. 1 juv.: Les Anses d’Arlet, under stones 0.5 km N of Anses d’Arlet, 11 Nov 1978; 3 juv.: Grand’Rivière, near beach, Anse Morne Rouge, dead tree, 20 Feb 1981, all leg. J.-M. Thibaud; 3 females, 7 juv.: Gros-Morne, 6 km NE of Gros-Morne, forest between Morne Bellevue and Morne de l’Etang, 600–690 m a.s.l., dead leaves and wood, 11 Feb 1981; 1 juv.: Trinité, near île Caravelle, under dry log between l’Oratoire and meteo station, 80–115 m a.s.l., 12 Feb 1981; 29 juv.: Morne Rouge, Trace des Jésuites, right bank of Lorrain River, 400–500 m a.s.l., dead wood and bamboo, 13 Feb 1981; 8 juv.: Rivière-Pilote, Anse Figuier, under bark of Cocos palm lying on sand, 24 Feb 1981; 1 juv.: Grand’Rivière, path to Morne Macouba (massif Montagne Pelée), S of savanna Anatole, forest, 550–600 m a.s.l., dead wood and leaves, 26 Feb 1981; 4 females, 2 juv.: Grand’Rivière, Morne aux Gueules (path to Montagne Pelée), cultivated land, 26.II.1981; 3 females: Fonds-Saint-Denis, Pitons du Carbet, between sides at 1100 and 1120 m a.s.l., SSE of Piton Boucher, bush savanna, mosses and dead wood, 27 Feb 1981, all leg. J.-P. Mauriès; 2 males, 9 juv.: Saint-Pierre, rocks, sugar cane, 80 m a.s.l., 02 Mar 2015, leg. M. Coulis. Remarks. As seen from the above records, M. hastatus is remarkably widespread on Martinique, ranging from nearly sea-level to mountain tops above 1100 m in elevation. Although there are no direct observations on the labels of its association with ants or termites, dead wood in the tropics is known to be widely populated and destroyed by termites. Both available males were studied in detail to reveal the species identity. The results are presented in Figs 1–20. There can be no doubt now that the Martinique samples belong to M. hastatus. The spinulate flagellum (fl) arising at the base of a simple, more or less sigmoid solenomere (sl) is especially characteristic (Figs 14–17 & 20). SEM iconography demonstrates all features typical of a small cryptic pyrgodesmid, including a flabellate collum that fully covers the head from above, the differentiated tuberculations and lobulations on the metaterga, the porosteles, the well-developed cerategument on the metazonae, etc. However, only gonopod characters are decisive for a proper species determination. This provides the first evidence of the presence of M. hastatus in the Caribbean., Published as part of GOLOVATCH, Sergei I., GEOFFROY, Jean-Jacques, MAURIÈS, Jean-Paul & VANDENSPIEGEL, Didier, 2016, Detailed iconography of the widespread Neotropical millipede, Myrmecodesmus hastatus (Schubart, 1945), and the first record of the species from the Caribbean area (Diplopoda, Polydesmida, Pyrgodesmidae), pp. 1-6 in Fragmenta Faunistica 59 (1) on pages 2-3, DOI: 10.3161/00159301FF2016.59.1.001, http://zenodo.org/record/6251868

Tuoba poseidonis (Verhoeff, 1901) Geophilus poseidonis Verhoeff, 1901: 421. LOCALITÉ TYPE. — Grèce, île Egine. RÉPARTITION ET HABITAT. — Tuoba poseidonis est connu sur le littoral circumméditerranéen de la France à la Grèce. En Italie, les données sont surtout situées sur le littoral ouest, la Sicile et la Sardaigne. Il a aussi été recensé au bord de la Mer Rouge et de la Mer Morte (Zapparoli & Iorio 2012).Dans les Alpes-Maritimes, il a été signalé par Caziot (1925) et Brölemann (1930) à l’île Saint-Honorat, dans les cordons de varech. Cette espèce méditerranéenne ne vit que le milieu étroit constitué par la zone intertidale et la zone de mouillage par les vagues, et n’est jamais trouvé au-dessus de cet ensemble. Ses micro-habitats sont, entre autres, les cordons littoraux de varechs et le dessous des posidonies échouées (Attems 1929; Brölemann 1930; Minelli & Iovane 1987; Geoffroy & Iorio 2009; Zapparoli & Iorio 2012; Iorio 2014). REMARQUES Espèce non retrouvée malgré nos prospections sur le littoral d’Èze-Bord-de-Mer à Saint-Raphaël., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 232, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["VERHOEFF K. W. 1901. - Beitrage zur Kenntnis palaarktischer Myriopoden. XVI. Aufsatz: zur vergleichenden Morphologie, Systematik und Geographie der Chilopoden. Nova Acta Abhandlungen der Kaiserlichen Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher 77: 369 - 465.","ZAPPAROLI M. & IORIO E., 2012. - The centipedes (Chilopoda) of Corsica: catalogue of species with faunistic, zoogeographical and ecological remarks. International Journal of Myriapodology 7: 15 - 68.","CAZIOT C. 1925. - Les Myriapodes du departement des Alpes- Maritimes et de la principaute de Monaco (Avec la contribution de H. W. Brolemann). Riviera scientifique 12 (1): 3 - 8.","BROLEMANN H. W. 1930. - Elements d'une faune des myriapodes de France. Chilopodes. Faune de France, 25. P. Lechevalier, Paris, 405 p.","ATTEMS C. 1929. - Myriapoda 1: Geophilomorpha, in SCHULZE F. E. & KUKENTHAL W. (eds), Das Tierreich, vol. 52. W. de Gruyter & C °, Berlin & Leipzig: 1 - 388.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34.","GEOFFROY J. - J. & IORIO E. 2009. - The French centipede fauna (Chilopoda): updated checklist and distribution in mainland France, Corsica and Monaco. Soil Organisms 81 (3): 671 - 694.","IORIO E. 2014. - Catalogue biogeographique des chilopodes (Chilopoda) de France metropolitaine. Memoires de la Societe linneenne de Bordeaux 15: 1 - 372."]}

Cryptops trisulcatus Brölemann, 1902 Cryptops biscarensis trisulcatus Brölemann, 1902: 73. LOCALITÉ TYPE. — Pyrénées-Orientales, Port-Vendres. MATÉRIEL EXAMINÉ. — Guillaumes (06), PNM, Le Coulet, conifères divers, 1470-1500 m, 8.VI.2008, réc.dét. EI: 1 ex.; Valbonne (06), Sophia-Antipolis, pins, chênes verts et chênes pubescents, 236 m, 43,625116N, 7,031673E, 11.IV.2012, réc./dét. EI: 1 ex.; Sospel (06), grotte de Sues, IV.2009, réc. JR & JML/dét. EI: 1 ex. Spécimens conservés en éthanol à 70° dans la collection personnelle de l’auteur (EI). RÉPARTITION ET HABITAT. — La répartition de cette espèce est quasiment limitée aux régions méditerranéennes d’Europe, jusqu’en Turquie; elle existe également en Algérie et en Tunisie (Iorio & Geoffroy 2008; Akkari et al. 2008). En France, C. trisulcatus n’est connu que des départements circumméditerranéens et en Corse (Iorio & Geoffroy 2008; Zapparoli & Iorio, 2012), où il est assez peu fréquent. Peu d’informations sont disponibles sur les habitats correspondant aux quelques stations connues dans le département des Alpes-Maritimes. Toutefois, les localisations des spécimens du côté français et les données écologiques italiennes (Minelli & Iovane 1987; Zapparoli 2006; Zapparoli & Iorio, 2012), confirment qu’il s’agit ici comme ailleurs d’une espèce thermophile surtout présente dans les formations arborées typiquement méditerranéennes., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 228, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["BROLEMANN H. W. 1902. - Materiaux pour servir a une faune des myriapodes de France. Feuille des Jeunes Naturalistes, 32: 73.","IORIO E. & GEOFFROY J. - J. 2008. - Les scolopendromorphes de France (Chilopoda, Scolopendromorpha): identification et distribution geographique des especes. Riviera scientifique 91 (2007): 73 - 90.","AKKARI N., STOEV P. & LEWIS J. G. E. 2008. - The scolopendromorph centipedes (Chilopoda, Scolopendromorpha) of Tunisia: taxonomy, distribution and habitats. ZooKeys 3: 77 - 102.","ZAPPAROLI M. & IORIO E., 2012. - The centipedes (Chilopoda) of Corsica: catalogue of species with faunistic, zoogeographical and ecological remarks. International Journal of Myriapodology 7: 15 - 68.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34."]}

Cryptops umbricus umbricus Verhoeff, 1931 Cryptops umbricus Verhoeff, 1931: 283. LOCALITÉ TYPE. — Italie, Umbria, Sud d’Arezzo; près de Pegli; Savona; Santaurio, Stressa au pied du Mont Mottarone. MATÉRIEL EXAMINÉ. — La Trinité (06), grotte du Plateau Tercier, VI.2008, réc. JR & JML/dét. EI: 10 ex.; Roquebrune-Cap-Martin (06), grotte de Saint-Roman, 22.VII.2010, réc. JML/dét. EI: 1 ex.; Monaco, grotte du Trocadéro, 2012, réc. JML/dét. EI: 1 ex.; Monaco, dans les galeries souterraines est, réc. JML/dét. EI: 2 ex.; Peille (06), grotte de Morgelle (dite de « Saint-Jean »), 2012, réc. JML/dét. EI: 2 ex. Spécimens conservés en éthanol à 70° dans la collection personnelle de l’auteur (EI). RÉPARTITION ET HABITAT. — C. u. umbricus n’est présent que dans les Alpes-Maritimes et les Alpes-de-Haute-Provence en France, et assez largement réparti en Italie sans y être commun (Zapparoli & Minelli 2006; Iorio & Geoffroy 2008; Iorio 2010a; présent travail). On ne le trouve quasi-exclusivement que dans les grottes, y compris dans de petits avens peu profonds (Minelli 1985; Iorio & Minelli 2005). Nous le qualifions ici d’eutroglophile en raison du fait que beaucoup de populations y sont établies en permanence, mais pour certaines, le milieu souterrain est très superficiel, voire artificiel. L’espèce présente des adaptations morphologiques nettement troglomorphes (allongement des articles antennaires, des antennes, et des pattes par rapport aux espèces épigées du même genre). REMARQUES Les spécimens examinés permettent de compléter la connaissance de certains caractères morphologiques: deux spécimens atteignaient 55 mm de long, deux autres 51 mm et 50,5 mm; cette espèce peut donc excéder les 50 mm auparavant observés. La longueur des sillons céphaliques est toujours inférieure à 40 % de celle de l’écusson céphalique chez les exemplaires étudiés (longueur = 26 à 38%), montrant la constance de ce caractère en cumulant avec tous les individus détaillés antérieurement (Iorio & Minelli 2005; Iorio 2010a). Les scies dentées des P. 21 sont dotées de: 14 + 11, 14 + 17, 15 + 14, 15 + 15, 17 + 15, 14 + x, 12 + x (immature) dents tibiales; 4 + x, 4 + 4, 4 + 5, 5 + 5, 5 + 6, 3 + x (immature) dents tarsales. Les amplitudes pour ces scies deviennent chez C. u. umbricus: 11 à 18 dents tibiales (neuf à dix chez les immatures, voire 12) et quatre à cinq dents tarsales (rarement trois = immatures)., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 228, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["VERHOEFF K. W. 1931. - Uber europaische Cryptops - Arten. Zoologischer Jahrbucher (Systematik) 62 (3): 263 - 288.","ZAPPAROLI M. & MINELLI A. 2006. - Chilopoda, in RUFFO S. & STOCH F. (eds), Checklist and distribution of the Italian fauna. Memorie del Museo civico di Storia naturale di Verona 2. Serie: 123 - 125 + CD-ROM.","IORIO E. & GEOFFROY J. - J. 2008. - Les scolopendromorphes de France (Chilopoda, Scolopendromorpha): identification et distribution geographique des especes. Riviera scientifique 91 (2007): 73 - 90.","IORIO E. 2010 a. - Description d'une nouvelle sous-espece de Cryptops umbricus Verhoeff, 1931 (Chilopoda, Scolopendromorpha, Cryptopidae). Bulletin de la Societe linneenne de Bordeaux tome 144 (N. S.) 37 (4) (2009): 471 - 481.","IORIO E. & MINELLI A. 2005. - Un Chilopode confirme pour la faune de France: Cryptops umbricus Verhoeff, 1931 (Scolopendromorpha, Cryptopidae). Bulletin mensuel de la Societe linneenne de Lyon 74 (4): 150 - 157."]}

Scutigera coleoptrata (Linnaeus, 1758) Scolopendra coleoptrata Linnaeus, 1758: 638. LOCALITÉ TYPE. — Espagne. MATÉRIEL EXAMINÉ. — Mandelieu (06), garrigue, en lisière arborée (chênes), 16 m, 43,528558N, 6,914716E, 7.V.2010, réc./dét. EI: 1 ex.; Valbonne (06), Sophia-Antipolis, pins, chênes verts et chênes pubescents, clairières pierreuses, 218 m, 43,625211N, 7,03265E, 11.IV.2012, réc./dét. EI: 2 ex.; Roquebrune-Cap-Martin (06), hôtel Vista Palace, jardins en restanques et pinèdes, 208 m, 43,756402N, 7,442974E, 18.IV.2012 et 26.VI.2012, réc./dét. EI: 3 ex.; Beausoleil (06), Giram, chênes verts, 499 m, 43,758534N, 7,416478E, 23.IV.2012, réc./dét. EI: 4 ex.; Nice (06), Pointe Madame, lisières de friches/pinèdes, 40-50 m, 3.X.2012, 43,692375N, 7,307853E, réc./dét. EI: 2 ex.; idem, 13.V.2013: 1 ex.; Menton (06), à proximité de la Gardieura, sentier GR 52, pinèdes et lisières de pinèdes, 350- 400 m, 3.V.2013, réc./dét. EI: 2 ex.; Roquebrune-Cap-Martin (06), Mont Gros, pinède claire, 600-650 m, 12.VI.2013, réc./dét. EI: 2 ex.; Menton (06), à proximité de Moniéri, mur le long de la petite route et de la lisière arbustive/arborée adjacente et près de quelques habitations en contrebas, 250-300 m, 9.X.2013 et 23.X.2013, réc./dét. EI: 9 ex.; Menton (06), Chemin des Granges de Saint-Paul, dans une lisière pierreuse de vignes, dans une friche, une prairie et une pinède, 400- 450 m, 24.X.2013, réc./dét. EI: 4 ex. Les spécimens ont été identifiés à vue pour cette espèce, très aisée à reconnaître sans prélèvement. RÉPARTITION ET HABITAT. — Cette espèce largement répartie en France et en Europe mais quasi-exclusivement synanthrope en dehors des régions circumméditerranéennes, dont la distribution s’est sans aucun doute élargie en raison de ses moeurs anthropophiles, est ici dans son territoire d’autochtonie (Iorio & Geoffroy 2007b). Dans les Alpes-Maritimes, on la trouve aussi bien en milieu naturel ou anthropisé qu’en milieu urbain et sub-urbain (Iorio 2008; présent travail), de 0 à 650 mètres d’altitude., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 216, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["LINNAEUS C. 1758. - Systema naturae. Editio Decima. Holmiae 1: 824 p.","IORIO E. & GEOFFROY J. - J. 2007 b. - Etude comparative de quatre especes du genre Cryptops Leach, 1815 (Chilopoda, Scolopendromorpha, Cryptopidae) en France. Le Bulletin d'Arthropoda 31: 29 - 35."]}

Cryptops hortensis (Donovan, 1810) Scolopendra hortensis Donovan, 1810: 23. LOCALITÉ TYPE. — Angleterre, Devon, Exeter, jardins. MATÉRIEL EXAMINÉ. — Saint-Martin-Vésubie (06), PNM, vallon Madone de Fenestre, 1500-1600 m, 44°05’17’’N- 7°18’24’’E, 22.V.2010, réc./dét. MZ: 12 ex.; Nice (06), Parc Phénix, 8.I.2013, réc. JML/dét. EI: 1 ex. Spécimens conservés en éthanol à 70° dans la collection personnelle des auteurs (EI, MZ). RÉPARTITION ET HABITAT. — C. hortensis est très largement réparti, présent de l’Asie centrale à l’Europe, et il a même été introduit dans d’autres contrées, comme en Amérique du Nord (Zapparoli 2006; Iorio & Geoffroy 2008). Il est connu dans de nombreux départements français, mais il est plus commun dans l’Ouest et le Centre du pays que dans l’Est (Iorio & Geoffroy 2008; Geoffroy & Iorio 2009). Dans les Alpes-Maritimes, il semble assez peu fréquent, à l’instar de ce qui s’observe ailleurs en France orientale et, même si peu de données écologiques sont disponibles, il a été trouvé aussi bien en milieu forestier qu’en milieu anthropisé (parcs urbains) (Iorio 2008; présent travail)., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on pages 226-227, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["DONOVAN E., 1810 - The Natural History of British Insects. London. 1810: 90 p.","IORIO E. & GEOFFROY J. - J. 2008. - Les scolopendromorphes de France (Chilopoda, Scolopendromorpha): identification et distribution geographique des especes. Riviera scientifique 91 (2007): 73 - 90.","GEOFFROY J. - J. & IORIO E. 2009. - The French centipede fauna (Chilopoda): updated checklist and distribution in mainland France, Corsica and Monaco. Soil Organisms 81 (3): 671 - 694."]}

Schendyla carniolensis Verhoeff, 1902 Schendyla carniolensis Verhoeff, 1902: 91. LOCALITÉ TYPE. — Adelsberg (Slovénie). MATÉRIEL EXAMINÉ. — Lucéram (06), Peira-Cava, La Cabanette, hêtres-conifères, 1350-1400 m, 1.X.2008, réc./dét. EI: 1♂ immature à 41pp; Menton (06), à proximité de la Gardieura, sentier GR 52, pinèdes et lisières de pinèdes, 350-400 m, 3.V.2013, réc./dét. EI: 1♀ à 49pp. Spécimens conservés en éthanol à 70° dans la collection personnelle de l’auteur (EI). RÉPARTITION ET HABITAT. — Schendyla carniolensis possède une répartition limitée au sein de l’Europe: Roumanie, Serbie, Slovénie, Autriche, Nord et Centre de l’Italie (très sporadique dans le sud du pays), France (Zapparoli 2006; Zapparoli & Minelli 2006; Geoffroy & Iorio 2009). En France, ce géophilomorphe est présent dans les Alpes-Maritimes et le Var, et de l’Aude aux Pyrénées-Atlantiques, tout en étant très sporadique à l’est des Pyrénées-Orientales (Iorio 2008, 2012, 2014). Dans les Alpes-Maritimes, cette espèce n’est citée que dans deux pinèdes (et/ou leurs lisières) et dans une forêt mixte de hêtres et de conifères, de 0 à 1400 mètres d’altitude (Iorio 2008; présent travail). Les données concernant le Nord et le Centre de l’Italie font apparaître que l’espèce est clairement sylvicole, privilégiant les forêts thermophiles et méso-thermophiles (Minelli & Iovane 1987; Zapparoli 2006). Dans le Var, elle est également citée dans une forêt à dominante de hêtres (Iorio 2012). Autrefois citée « de Cannes à Menton » par Caziot (1925), il est possible qu’elle ait régressé sur cette partie du littoral en raison de l’importante croissance de l’urbanisation depuis les années 1920, et du fait qu’elle n’y ait pas été retrouvée ailleurs qu’à Menton., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 230, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["VERHOEFF K. W. 1902. - Uber Chilopoden von Sudsteiermark Krain und Kroatien. Sitzungs-Berichte der Gesellschaft naturforschender Freunde zu Berlin: 90 - 93.","ZAPPAROLI M. & MINELLI A. 2006. - Chilopoda, in RUFFO S. & STOCH F. (eds), Checklist and distribution of the Italian fauna. Memorie del Museo civico di Storia naturale di Verona 2. Serie: 123 - 125 + CD-ROM.","GEOFFROY J. - J. & IORIO E. 2009. - The French centipede fauna (Chilopoda): updated checklist and distribution in mainland France, Corsica and Monaco. Soil Organisms 81 (3): 671 - 694.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34.","IORIO E. 2012. - Premiere contribution a la connaissance des chilopodes (Chilopoda) de la hetraie de la Sainte-Baume (Var). Bulletin de la Societe linneenne de Bordeaux tome 147 (N. S.), 40 (2): 235 - 241.","CAZIOT C. 1925. - Les Myriapodes du departement des Alpes- Maritimes et de la principaute de Monaco (Avec la contribution de H. W. Brolemann). Riviera scientifique 12 (1): 3 - 8."]}

Geophilus flavus (De Geer, 1778) Scolopendra flava De Geer, 1778: 561. LOCALITÉ TYPE. — Inconnue. MATÉRIEL EXAMINÉ. — Allos (04), PNM, 11-17.VI.1899, réc. HWB/dét. EI: 2♀ à 51pp; Allos (04), PNM, bois de Villard, 16.VI.1899, réc. HWB/dét. EI: 1♀ à 51pp; Beuil (06), PNM, Les Launes, conifères et lisière pelouse, 1600 m, 7.VI.2008, réc./ dét. EI: 2♂ à 49pp; Colmars-les-Alpes (04), PNM, Ratery, forêt domaniale de la Haute-Vallée du Verdon, mélèzaie, sol et bois mort, 1700 m, 22.VIII.2009, réc./dét. JJG: 1♀; Saint-Martin-Vésubie (06), PNM, Forêt de Boréon, parking 5, forêt mixte de conifères, 1500 m, 44°06’59’’N- 7°16’44’’E, 21.V.2010, réc./dét. MZ: 3♀ (1♀ à 51pp, 2♀ à 53pp); Monaco, aven, XI-XII.2012, réc. JML/dét. EI: 1♂ immature à 49pp; idem, I-III.2013: 1♂ immature à 51pp. Spécimens conservés en éthanol à 70° dans la collection personnelle des auteurs (EI, MZ) ou dans celle du MNHN (Myriapoda, Geophilomorpha) (JJG). RÉPARTITION ET HABITAT. — Ce géophilomorphe est largement réparti en Europe, également connu en Tunisie, et introduit dans diverses contrées, par exemple en Amérique du Nord; en France tout comme en Italie, il est connu dans de nombreux secteurs, et apparaît comme une des espèces les plus euryèces de cet ordre de chilopodes même s’il est plus souvent trouvé en forêt qu’ailleurs (Brölemann 1930; Minelli & Iovane 1987; Zapparoli 2006; Geoffroy & Iorio 2009; Iorio 2014). 20 des stations connues dans les Alpes-Maritimes informent sur les habitats: quatre forêts indéterminées, deux pessières, une pinède, une aulnaie, une hêtraie-conifères, une châtaigneraie et une forêt de feuillus dominée par le châtaignier, une mélèzaie, deux forêts de conifères, une rhodoraie/prairie limitrophe, une prairie subalpine, une prairie montagnarde et une lisière de pelouse montagnarde, un milieu méditerranéen sur une île, et dans deux avens. G. flavus est présent de 0 à 2450 mètres dans ce département., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 231, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["GEER C. DE 1778. - Memoires pour servir a l'histoire des insectes. Tome VII. P. Hesselberg, Stockholm, 1778: 950 p.","BROLEMANN H. W. 1930. - Elements d'une faune des myriapodes de France. Chilopodes. Faune de France, 25. P. Lechevalier, Paris, 405 p.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34.","GEOFFROY J. - J. & IORIO E. 2009. - The French centipede fauna (Chilopoda): updated checklist and distribution in mainland France, Corsica and Monaco. Soil Organisms 81 (3): 671 - 694.","IORIO E. 2014. - Catalogue biogeographique des chilopodes (Chilopoda) de France metropolitaine. Memoires de la Societe linneenne de Bordeaux 15: 1 - 372."]}

Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe, Geoffroy, Jean-Jacques (2015): Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s.s., synthèse des connaissances et espèces menacées. Zoosystema 37 (1): 211-238, DOI: 10.5252/z2015n1a11, URL: http://dx.doi.org/10.5252/z2015n1a11

Strigamia acuminata (Leach, 1815) Geophilus acuminatus Leach, 1815: 386. LOCALITÉ TYPE. — Angleterre, RoboroughDown près de Plymouth; champs de Battersea. MATÉRIEL EXAMINÉ. — Saint-Martin-Vésubie (06), PNM, vallon Madone de Fenestre, lariçaie, 1830 m, 44°05’36’’N- 7°21’24’’E, 22.V.2010, réc./dét. MZ: 1♂ à 39pp. Spécimens conservés en éthanol à 70° dans la collection personnelle de l’auteur (MZ). RÉPARTITION ET HABITAT. — S. acuminata est largement réparti en Europe, mais est absent des contrées les plus nordiques (Norvège, Suède), sa limite septentrionale de répartition étant le sud du Danemark (Andersson et al. 2005; Zapparoli 2006; Geoffroy & Iorio 2009; Bonato et al. 2012). Il est présent dans de nombreux départements français. Dans les Alpes-Maritimes, ce géophilomorphe est peu commun, connu en forêt et seulement dans les étages montagnards et sub-alpins (1350 à 2000 mètres d’altitude): deux lariçaies, une pessière, une forêt de nature indéfinie. Notons que le caractère strictement sylvicole de celui-ci est déjà bien établi dans le reste de sa répartition, et qu’il s’agit d’une espèce essentiellement inféodée aux forêts caducifoliées, hêtraies notamment (Iorio 2014). Toutefois, à l’instar de L. (L.) dentatus évoqué plus haut, il semble aussi plus montagnard ici et peut parfois être trouvé dans d’autres types de boisements comme l’illustrent nos données et celles de Minelli & Iovane (1987), Zapparoli (1993, 2006) et Spelda (1999), surtout dans le sud de sa répartition. Mais même dans cette dernière, les boisements caducifoliés et le hêtre en particulier restent notablement privilégiés; par exemple, sur les 73 stations où l’habitat est connu dans les Apennins centrales, 65 sont des forêts caducifoliées en tout ou partie (15 forêts mixtes), dont 43 hêtraies, et sept autres sont des forêts de conifères (sapinières, pinèdes) (Zapparoli 2006)., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 232, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["LEACH W. E. 1815. - A tabular view of the external characters of four classes of animals, wich Linne arranged under Insecta; with the distribution of the genera composing three of these classes into orders, etc. and descriptions of several new genera and species. Transactions of the Linnean Society 11: 307 - 400.","ANDERSSON G., BJARNE M., SCHELLER U., DJURSVOLL P., BUDD G. & GARDENFORS U. 2005. - Mangfotingar. Myriapoda. Nationalnyckeln till Sveriges Flora och Fauna, Upsalla 351 p.","GEOFFROY J. - J. & IORIO E. 2009. - The French centipede fauna (Chilopoda): updated checklist and distribution in mainland France, Corsica and Monaco. Soil Organisms 81 (3): 671 - 694.","BONATO L., DANYI L., SOCCI A. A. & MINELLI A. 2012. - Species diversity of Strigamia Gray, 1843 (Chilopoda: Linotaeniidae): a preliminary synthesis. Zootaxa 3593: 1 - 39.","IORIO E. 2014. - Catalogue biogeographique des chilopodes (Chilopoda) de France metropolitaine. Memoires de la Societe linneenne de Bordeaux 15: 1 - 372.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34.","ZAPPAROLI M. 1993. - Chilopodi dell'Italia Nord-Occidentale. Rivista Piemontese di Storia Naturale 14: 97 - 127.","SPELDA J. 1999. - Verbreitungsmuster und Taxonomie der Chilopoda und Diplopoda Sudwestdeutschlands. Diskriminanzanalytische Verfahren zur Trennung von Arten und Unterarten am Beispiel des Gattung Rhymogona Cook, 1896 (Diplopoda, Chordeumatida, Craspedosomatidae). Ph. D. Thesis, University of Ulm. Part I: 217 p. Part II: 324 p."]}

Pachymerium ferrugineum (C. L. Koch, 1835) Geophilus ferrugineus C. L. Koch, 1835: 2. LOCALITÉ TYPE. — Allemagne, Regensburg, Arklee. RÉPARTITION ET HABITAT. — Toute l’Europe, de la Scandinavie à l’Espagne et duRoyaume-Uni jusqu’enRussie occidentale; Proche- et Moyen-Orient; Asie occidentale; Afrique du Nord; Alaska. Introduit en Amérique du Nord et centrale, également au Chili (Zapparoli, 2009b). Dans les Alpes-Maritimes, signalé par Brölemann (1904) à Monaco; par Caziot (1925) sur tout le littoral de ce département, jusque dans les cordons de varech; par Brölemann (1930) à La Bocca (Cannes); par Iorio (2008) à Cannes la Bocca, plage, d’après un spécimen datant de 1905. Globalement, ce géophilomorphe est connu comme ayant une tendance halophile, parfois nette dans certains endroits (Brölemann 1930; Palmen & Rantala 1954), sans être strictement inféodé aux habitats particuliers du bord de mer (Minelli & Iovane 1987; Iorio 2014). REMARQUES Espèce non retrouvée malgré nos prospections sur le littoral d’Èze-Bord-de-Mer à Saint-Raphaël, y compris à Cannes la Bocca., Published as part of Iorio, Étienne, Zapparoli, Marzio, Ponel, Philippe & Geoffroy, Jean-Jacques, 2015, Les myriapodes chilopodes (Chilopoda) du Parc national du Mercantour, du département des Alpes-Maritimes et de leurs environs: description d'une nouvelle espèce du genre Lithobius Leach, 1814 s. s., synthèse des connaissances et espèces menacées, pp. 211-238 in Zoosystema 37 (1) on page 231, DOI: 10.5252/z2015n1a11, http://zenodo.org/record/5155397, {"references":["KOCH C. L. 1835. - Deutschlands Crustaceen Myriapoden und Arachniden, in (PANZER G. W. F.) HERRICH- SCHAFFER G. A. W. Deutschlands Insecten. Pustet: 136.","ZAPPAROLI M. 2009 b. - An annotated catalogue of the epigeic and cave centipedes (Chilopoda) of Sardinia, in CER- RETTI P., MASON F., MINELLI A., NARDI G. & WHITMORE D. (eds), Research on the terrestrial arthropods of Sardinia (Italy). Zootaxa 2318: 56 - 168.","BROLEMANN H. W. 1904. - Chilopodes monegasques. Bulletin du Musee oceanographique de Monaco 15: 1 - 15.","CAZIOT C. 1925. - Les Myriapodes du departement des Alpes- Maritimes et de la principaute de Monaco (Avec la contribution de H. W. Brolemann). Riviera scientifique 12 (1): 3 - 8.","BROLEMANN H. W. 1930. - Elements d'une faune des myriapodes de France. Chilopodes. Faune de France, 25. P. Lechevalier, Paris, 405 p.","PALMEN E. & RANTALA M. 1954. - On the life-history and ecology of Pachymerium ferrugineum (C. L. Koch) (Chilopoda, Geophilidae). Annales zoologici Societatis Zoologicae Botanicae Fennicae - Vanamo' 16 (3): 1 - 44.","MINELLI A. & IOVANE E. 1987. - Habitat preferences and taxocenoses of Italian centipedes. Bollettino del Museo civico di Storia naturale di Venezia 37: 7 - 34.","IORIO E. 2014. - Catalogue biogeographique des chilopodes (Chilopoda) de France metropolitaine. Memoires de la Societe linneenne de Bordeaux 15: 1 - 372."]}

Key to Hyleoglomeris species currently known to occur in China 1. Dorsum dark, usually with a vivid colour pattern............................................................. 2 ��� Body uniformly pallid to light, only slightly pigmented at best.................................................. 7 2. Body rather uniformly orange-brown, dorsum devoid of a colour pattern. Chongqing........................ H. aschnae ��� Colour pattern vivid................................................................................... 3 3. Background coloration pallid to yellow, spots on tergites brown to dark brown. Collum, second tergite and tergite 11 entirely pale. Body with a distinctive pattern of 1 + 1 large, transverse-oval, evidently separated, paramedian spots on tergites 3���10 and on anal shield, as well as of 1 + 1 increasingly small, roundish, sublateral spots on tergites 3 ���(8) 9. Yunnan....... H. maculata ��� Background coloration of dorsum dark, markings lighter...................................................... 4 4. Coloration of dorsum nearly entirely dark brown to blackish, only front third of second tergite contrasting light. Sichuan................................................................................................ H. sinensis ��� Coloration of dorsum variegated, with evident spots............................................................ 5. Tergites 3���11 with a light yellow axial row of spots flanked by two rows of rather vague, marbled, olive-yellowish spots. Hong Kong....................................................................................... H. bicolor ��� Tergites 3���11 with a light axial row of spots flanked by only one row of large, marbled yellowish spots................. 6 6. Caudal half of head and nearly entire collum dark brown to blackish. Telopod syncoxital horns usual, not hypertrophied. Jiangsu................................................................................... H. emarginata ��� Head and nearly entire collum light (Fig. 9 A, B). Telopod syncoxital horns greatly hypertrophied (Fig. 10 C). Guangxi................................................................................................. H. qiyi sp. n. 7. An evident transverse sulcus above schism on second tergite. Collum with one (front) transverse stria................... 8 ��� Surface above schism of second tergite smooth. Collum rarely with one, usually with two transverse striae, but 2 nd one some- times obliterated mid-dorsally............................................................................ 9 8. Second tergite with 8���11 striae, of which usually seven cross the dorsum. Tips of syncoxital horns with a minute setoid fila- ment (Fig. 4 E). Guizhou....................................................................... H. eusulcata ��� Second tergite with six striae, of which only 3 or 4 cross the dorsum. Tips of syncoxital horns devoid of a setoid filament (Fig. 16 E). Guangxi........................................................................ H. curtisulcata sp. n. 9. Collum with one (front) transverse stria................................................................... 10 ��� Collum with two transverse striae, of which 2 nd can be obliterated mid-dorsally.................................... 11 10. Central lobe of telopod syncoxite rather low and roundly subtriangular (Fig. 15 C). Mulun Karst, Guangxi................................................................................................... H. mashanorum sp. n. ��� Central lobe of telopod syncoxite rather high and subquadrate (Fig. 8 C). Guizhou........................ H. yinshi sp. n. 11. Collum with two transverse striae, of which 2 nd obliterated mid-dorsally. Mulun Karst.............................. 12 ��� Collum as usual, with two complete transverse striae........................................................ 13 12. Tip of telopod syncoxite horns devoid of a setoid filament (Fig. 17 D)................................ H. kunnan sp. n. ��� Tip of telopod syncoxite horns with a minute setoid filament (Fig. 14 D)............................ H. mulunensis sp. n. 13. Brownish pigment usually retained at least in some parts of head.............................................. 14 ��� Body completely unpigmented (sometimes except for 2 or 3 caudalmost ocelli).................................. 18 14. Head, antennae and collum usually with traces of brownish coloration, sometimes body either uniformly slightly infuscate (Fig. 9 D���F) or even with a faint pattern of paramedian light brown spots on following tergites........................ 15 ��� Only head with traces of pigment........................................................................ 16 15. Antennomere 6 ca 2.5 times as long as high. Second tergite with 6 or 7 transverse striae, of which (1) 2���3 crossing the dorsum. Yunnan...................................................................................... H. reducta ��� Antennomere 6 ca 2.0 times as long as high. Second tergite with 5���7 transverse striae, of which 4 or 5 crossing the dorsum. Guangxi..................................................................................... H. lii sp. n. 16. Head (especially near ocelli and T��m��sv��ry���s organ) and antennae often slightly infuscate, very light brownish. Central lobe of telopod syncoxite broadly subtrapeziform (Fig. 3 C). Hunan........................................ H. youhao sp. n. ��� Only either ocelli or antennae infuscate. Central lobe of telopod syncoxite considerably narrower..................... 17 17. Only ocelli brownish to dark brown. Larger: width 3.8 ���4.0 mm. Tips of telopod syncoxital horns with a minute lobule devoid of a setoid filament (Fig. 13 C). Guangxi......................................................... H. xueju sp. n. ��� Only antennae sometimes very faintly brownish. Smaller: width 2.5 ���3.0 mm. Tips of telopod syncoxital horns with a setoid fil- ament (Fig. 2 E). Hunan........................................................................ H. xia sp. n. 18. Caudomedial process of telopod femur with a distinct distolateral lobe (Fig. 2 C, lo)................................ 19 ��� Caudomedial process of telopod femur devoid of a distinct distolateral lobe...................................... 21 19. Tip of telopod syncoxite horn with a minute setoid filament. Yunnan.................................. H. albicorporis ��� Tip of telopod syncoxite horn devoid of a setoid filament..................................................... 20 20. Central lobe of telopod syncoxite nearly as high as lateral horns (Fig. 7 C). Second tergite with six transverse striae, of which three crossing the dorsum. Guizhou............................................................. H. gudu sp. n. ��� Central lobe of telopod syncoxite evidently lower, roundly subtriangular (Fig. 12 C). Second tergite with at least eight trans- verse striae. Guangxi...................................................................... H. heshang sp. n. 21. Telopod tibial process absent (Fig. 6 C, D). Central lobe of telopod syncoxite rather low and semi-circular (Fig. 6 C)................................................................................................... H. nigu sp. n. ��� Telopod tibial process present. Central lobe of telopod syncoxite higher.......................................... 22 22. Second tergite with 6 or 7 transverse striae, of which only two crossing the dorsum. Central lobe of telopod syncoxite high, subtrapeziform, slightly concave at ventral margin (Fig. 1 D). Hunan................................... H. tiani sp. n. ��� Second tergite with 9 or 10 transverse striae, of which seven crossing the dorsum. Central lobe of telopod syncoxite high, nar- row, linguiform (Fig. 5 C). Guizhou.............................................................. H. wuse sp. n., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 24-25, DOI: 10.5281/zenodo.214383

Hyleoglomeris qiyi sp. n. Figs 9 A���C & 10. Material examined: Holotype male (IZAS), China, Guangxi Prov., Huanjiang County, Mulun Karst, Min Li Forest, 25.09935 ��N, 107.96647 ��E, litter, Berlese extraction after sifting, 14.03. 2005, leg. Chen (CHIgx05��� 109). Paratypes. 1 female, 1 juv. (IZAS), 1 female, 1 juv. (MNHN CC 175), same locality, together with holotype. Name: To emphasize the highly peculiar syncoxite of the telopods, i.e. hypertrophied horns and a wamting central lobe, ��� qiyi ��� in Chinese meaning ���bizarre���; a noun in apposition. Diagnosis: Differs from congeners except H. venustula (Silvestri, 1917) in the hypertrophied horns and a strongly reduced central lobe of the telopod syncoxite (Silvestri 1917), from H. venustula by its far more strongly developed horns and a wanting central lobe. See also Key below. Description: Length ca 5.5 (holotype) or 6.0 mm (paratypes), width 2.5 mm (holotype) or 2.6 mm (paratypes). Coloration motley, dark brown and yellowish. Ocelli blackish, 7 + 1, translucid. T��m��sv��ry���s organ transverse-oval, only slightly wider than long. Antennomere 6 ca 1.7 times as long as high. Colour pattern mostly very clear (Fig. 9 A���C). Head, ventral side of body and legs mostly marbled light yellow-brown to red-brown; ocelli black; antennae red-brown basally, growing increasingly dark red-brown distad, pallid apically (Fig. 9 C). Collum rather broadly marbled brown along margins and narrowly brownish along both transverse striae, with a lighter, marbled, vague, yellowish, central spot (holotype), or infuscate, marbled dark brown only laterally (paratypes, Fig. 9 A, B). Second tergite with a distinct, dark brown, central, transverse, crossshaped marking extending laterad into rather wide bands along both front and caudal margins; front band somewhat lighter and nearly reaching down the schism (holotype) or almost so (paratypes, Figs 9 A, B); caudal band darker, shorter and supplied with a considerable arm stretched frontodorsally; large, paramedian, marbled yellowish, comma-shaped spots remaining laterally between both darker bands; a small, subtriangular, axial, yellowish spot lying also at caudal margin. Tergites 3���11 each with an elongate, transverse, marbled yellowish spot lying laterally between a rather broad, marbled brown interval situated above a broadly whitish lateral end of tergites, on the one hand, and a light, marbled yellowish, axial spot, on the other. Light axial spots on tergites 3���5 larger, irregularly subquadrate, each with a small brown spot at caudal margin; light axial spots on following tergites increasingly small and vague, subtriangular, most of them also showing a small brown spot at caudal margin (Fig. 9 A, B). Anal shield mostly light yellowish, with a large, dark brown, round (holotype) or vertically oblong-oval (paratypes) spot reaching neither ventral nor dorsal margin (Fig. 9 A, C). Collum with two transverse striae. Second tergite with a rather broad hyposchism reaching behind the caudal tergal margin; seven transverse striae, three starting below, one level to, the remaining three striae above schism, only one stria crossing the dorsum. Male anal shield regularly rounded at caudal margin. Male leg 17 (Fig. 10 A) with a low and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with one subapical spine. Male leg 18 (Fig. 10 B) with a broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 10 C, D) with an extremely strongly hypertrophied syncoxite, the latter being totally devoid of a central lobe, but supplied with conspicuous (nearly as long as telopods), thick, densely setose horns crowned with a strong apical uncus and a subapical bifid filament on front face. Prefemur sparsely micropapillate laterally. Caudomedial femoral process prominent, apically with a small membranous sac. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, rounded apically. Remarks: This species is remarkable due to its conspicuously enlarged and complex telopod syncoxite. A somewhat similar structure has only been known in H. venustula, from the Himalaya of Assam, India (Silvestri 1917), but the telopod syncoxital horns are much less strongly incrassate and elongate while a vestigial central lobe is still discernible. The slightly unsteady shape of the dark central spot on the anal shield is most probably only individual variation not linked to sex., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 13-15, DOI: 10.5281/zenodo.214383, {"references":["Silvestri, F. (1917) Contributions to a knowledge of the Oriental Diplopoda Oniscomorpha. 1. The family Glomeridae. Records of the Indian Museum, 13, 103 - 151."]}

Hyleoglomeris lii sp. n. Figs 9 D���F & 11. Material examined: Holotype male (IZAS), China, Guangxi Prov., Fuchuan County, Qishan Village, Cave Bai Fu Dong, 24.964284 ��N, 111.2752257 ��E, 27.01. 12, leg. Li Youbang, Li Youting & Li Yousu (CHIgx 12 ���LYB08). Paratypes. 1 male, 1 female (MNHN CC 176), 1 female (IZAS), 1 male, 1 female (GNUG), same locality, together with holotype. Name: Honours Li Youbang, the collector. Diagnosis: Differs from all congeners in its residual coloration, as well as the bisunuate, subtrapeziform, central lobe of the telopod syncoxite, the absence of a distinct chitinized lobe from the telopod caudomedial femoral process, and of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0 mm, width 2.0��� 2.9 mm in both sexes. Holotype 2.9 mm wide. Coloration usually uniformly light grey-yellow (except for the usual flavous margins) (Fig. 9 D���F), only a very faint pattern of 1 + 1 rounded darker, light grey-yellow, paramedian spots sometimes traceable on collum and several following tergites; front half of body coloured slightly more intensely. Head and collum the darkest, marbled light red-brown, collum with a light central spot. Ocelli sometimes very light brownish, at least 6 + 1, translucid, barely visible. Antennomere 6 ca 2.0 times as long as high. Second tergite with 5���7 transverse striae, two starting below, one level to, the remaining striae above schism, 4 or 5 striae (various, sometimes neither 1 st nor last from below) crossing the dorsum; last crossing stria always a little deeper than others. Male leg 17 (Fig. 11 A) with a very low and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with a ventral spine in distal third. Male leg 18 (Fig. 11 B) with an arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 11 C, D) with a rather low, linguiform, apically evidently concave, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule. Prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, narrowly rounded apically. Remark: Because this species has retained some pigment in its tegument, it would seem to be hardly more than a troglophile., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 15-16, DOI: 10.5281/zenodo.214383

Hyleoglomeris xia sp. n. Fig. 2. Material examined: Holotype male (IZAS), China, Hunan Prov., Linwu County, Sanhe Town, Tianhe Village, Cave 1, 19.06. 2009, leg. Tian Mingyi & Xue Zhihong (CHIhn09���LWX02). Paratype. 1 female (SCAU), same locality, together with holotype. Name: To emphasize the animal���s ocelli being unpigmented, ��� xia ��� in Chinese meaning ���blind���; a noun in apposition. Diagnosis: Differs from all congeners in the residual coloration of the antennae, the presence of a lobe on the caudomedial process of the telopod femur, of a setoid filament at the tip of the telopod syncoxital horns, of only a residual telopod tibial process, coupled with a roundly subtrapeziform, slightly concave, central lobe of the telopod syncoxite, as well as a semi-circular notch of male syncoxite 18, and a 3 -segmented male telopodite 17. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.0 (holotype) or 6.0 mm (paratype), width 2.5 (holotype) or 3.0 mm (paratype). Coloration nearly entirely pallid, only antennae in paratype very faintly infuscate, brownish. Ocelli at least 4 + 1, translucid, barely visible. Antennomere 6 ca 2.1���2.2 times as long as high. Second tergite with 7 or 8 transverse striae, 3 or 4 starting below, one level to, the remaining striae above schism, 3 or 4 frontmost ones crossing the dorsum. Male leg 17 (Fig. 2 A) with a very low and rather irregularly shaped outer coxal lobe; telopodite 3 -segmented, tarsus with only one subapical spine. Male leg 18 (Fig. 2 B) with a rather broadly rounded syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 2 C-E) with a high, apically slightly concave, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute, elongate lobule supplied with an adjacent mesal setoid filament (Fig. 2 E). Prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a rounded lobe (lo) on caudal face. Caudomedial process of tibia very obscure, low, membranous; tibial tubercle on caudal face evident, papillate. Tarsus strongly sigmoid, subacuminate apically. Remarks: Since this species shows such troglomorphic traits as entirely unpigmented tegument and ocelli, it can well prove to be a troglobite. Residual coloration of the antennae in one of the two specimens is noteworthy., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 4-5, DOI: 10.5281/zenodo.214383

Hyleoglomeris mashanorum sp. n. Fig. 15. Material examined: Holotype male (IZAS), China, Guangxi Prov., Huanjiang County, Mulun Karst, Cave Mashan Dong, ca 340 m a.s.l., 25.15855 ��N, 108.05865 ��E, 21.05. 2007, leg. F. Br��hier (CHIgx 07��� 21 ���04). Name. To emphasize the animal���s provenance from Cave Mashan Dong in Mulun Karst. Diagnosis: Differs from congeners except H. eusulcata, H. curtisulcata sp. n. and H. yinshi sp. n. in the presence of only one complete transverse stria on the collum, from both H. eusulcata and H. curticulcata sp. n. by the absence of a transverse sulcus on the second tergite, from H. yinshi sp. n. by the shape of the central lobe of the telopod syncoxite and of the notch in male syncoxite 18. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 7.5 mm, width 3.7 mm. Coloration entirely pallid. Ocelli at least 3 + 1, translucid, barely visible. Antennomere 6 ca 2.2 times as long as high. Collum with one complete transverse stria. Second tergite with eight transverse striae, three starting below, one level to, the remaining striae above schism, four striae (not last two ones from below) crossing the dorsum. Leg 17 (Fig. 15 A) with a low and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus devoid of subapical spines. Leg 18 (Fig. 15 B) with a narrowly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 15 C, D) with a rather low, linguiform, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a highly inconspicuous lobule (Fig. 15 D). Both prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus evidently sigmoid, subacuminate apically. Remark: Due to several troglomorphic traits, this species seems to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on page 21, DOI: 10.5281/zenodo.214383

Hyleoglomeris youhao sp. n. Fig. 3. Material examined: Holotype male (IZAS), China, Hunan Prov., Linwu County, Sanhe Town, near Changshali Village, Cave 2, 19.06. 2009, leg. Tian Mingyi & Xue Zhihong (8,12♀ CHIhn09���LWX03). Paratypes. 1 male, 2 females (IZAS), 2 males, 2 females (SCAU), 2 males, 4 females (MNHN CC 170), 1 male, 2 females (ZMUM), 1 male, 2 females (GNUG), same locality, together with holotype. Name: To emphasize the type series being large enough to share with all of the relevant repositories, ��� youhao ��� in Chinese meaning ���friendly���; a noun in apposition. Diagnosis: Differs by the residual coloration on the head, by the presence of a lobe on the caudomedial process of the telopod femur, coupled with a subquadrate, slightly concave, central lobe of the telopod syncoxite. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0���8.0 mm, width 2.8���3.7 mm in both sexes. Holotype 7.0 mm long and 3.2 mm wide. Coloration nearly entirely pallid, but parts of head (especially near ocelli and T��m��sv��ry���s organ) and antennae often slightly infuscate, very light brownish. Ocelli at least 5 + 1, translucid, mostly rather convex. Antennomere 6 ca 2.0��� 2.1 times as long as high. Second tergite with 7���9 transverse striae (usually 8), 3 or 4 starting below, one level to, the remaining striae above schism, usually 3 or 4 (2 nd to 4 th or 5 th from below) crossing the dorsum. Male leg 17 (Fig. 3 A) with a rather high and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 3 B) with a rather broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 3 C, D) with a rather high, apically very faintly concave, subquadrate, sparsely setose, central syncoxital lobe flanked by two setose horns with simple apices. Prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a rounded lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, narrowly rounded apically. Remarks: This species shows such troglomorphic traits as a nearly unpigmented body and entirely unpigmented ocelli, so it can well prove to be a troglobite. Residual coloration of the antennae and head is noteworthy., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on page 5, DOI: 10.5281/zenodo.214383

Hyleoglomeris yinshi sp. n. Fig. 8. Material examined: Holotype male (MNHN CC 174), China, Guizhou Prov., Kaiyang County, Cave Xian Yan Dong, 19.02. 2004, leg. S. Prevorčnik & B. Sket. Paratypes. 1 male, 1 female (MNHN CC 174), 1 male, 1 female (IZAS), 1 male (ZMUM), same locality, together with holotype. Name: To emphasize the animal coming from a cave, “ yinshi ” in Chinese meaning a “hermit”; a noun in apposition. Diagnosis: Differs from congeners except H. eusulcata, H. mulunensis sp. n. and H. mashanorum sp. n. in the presence of only one complete transverse stria on the collum, from H. eusulcata by the absence of a transverse sulcus on the second tergite, from both H. mulunensis sp. n. and H. mashanorum sp. n. by the subquadrate central lobe of the telopod syncoxite and the shape of the notch in male syncoxite 18. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.0–6.0 mm (male) or 6.0–7.0 mm (female), width 2.2–2.9 mm (male) or 2.7 –3.0 mm (female). Holotype 5.0 mm long, 2.3 mm wide. Coloration entirely pallid. Ocelli at least 4 + 1, translucid, barely visible. Antennomere 6 ca 2.0 times as long as high. Collum with one transverse stria. Second tergite with 8–11 transverse striae, two starting below, 1 or 2 level to, the remaining striae above schism, 3–5 striae (various, but never 1 st and/or last from below) crossing the dorsum. Male leg 17 (Fig. 8 A) with a high and highly irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 8 B) with a broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 8 C, D) with a high, linguiform, apically barely concave, sparsely setose, central syncoxital lobe flanked by two setose horns, each horn being crowned by a short minute lobule. Prefemur micropapillate laterally. Caudomedial femoral process prominent, subapically with a small lobe. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, narrowly rounded apically. Remark: Due to its troglomorphic traits (unpigmented tegument and ocelli), this species seems to be a troglobite.

Key to Hyleoglomeris species currently known to occur in China 1. Dorsum dark, usually with a vivid colour pattern............................................................. 2 – Body uniformly pallid to light, only slightly pigmented at best.................................................. 7 2. Body rather uniformly orange-brown, dorsum devoid of a colour pattern. Chongqing........................ H. aschnae – Colour pattern vivid................................................................................... 3 3. Background coloration pallid to yellow, spots on tergites brown to dark brown. Collum, second tergite and tergite 11 entirely pale. Body with a distinctive pattern of 1 + 1 large, transverse-oval, evidently separated, paramedian spots on tergites 3–10 and on anal shield, as well as of 1 + 1 increasingly small, roundish, sublateral spots on tergites 3 –(8) 9. Yunnan....... H. maculata – Background coloration of dorsum dark, markings lighter...................................................... 4 4. Coloration of dorsum nearly entirely dark brown to blackish, only front third of second tergite contrasting light. Sichuan................................................................................................ H. sinensis – Coloration of dorsum variegated, with evident spots............................................................ 5. Tergites 3–11 with a light yellow axial row of spots flanked by two rows of rather vague, marbled, olive-yellowish spots. Hong Kong....................................................................................... H. bicolor – Tergites 3–11 with a light axial row of spots flanked by only one row of large, marbled yellowish spots................. 6 6. Caudal half of head and nearly entire collum dark brown to blackish. Telopod syncoxital horns usual, not hypertrophied. Jiangsu................................................................................... H. emarginata – Head and nearly entire collum light (Fig. 9 A, B). Telopod syncoxital horns greatly hypertrophied (Fig. 10 C). Guangxi................................................................................................. H. qiyi sp. n. 7. An evident transverse sulcus above schism on second tergite. Collum with one (front) transverse stria................... 8 – Surface above schism of second tergite smooth. Collum rarely with one, usually with two transverse striae, but 2 nd one some- times obliterated mid-dorsally............................................................................ 9 8. Second tergite with 8–11 striae, of which usually seven cross the dorsum. Tips of syncoxital horns with a minute setoid fila- ment (Fig. 4 E). Guizhou....................................................................... H. eusulcata – Second tergite with six striae, of which only 3 or 4 cross the dorsum. Tips of syncoxital horns devoid of a setoid filament (Fig. 16 E). Guangxi........................................................................ H. curtisulcata sp. n. 9. Collum with one (front) transverse stria................................................................... 10 – Collum with two transverse striae, of which 2 nd can be obliterated mid-dorsally.................................... 11 10. Central lobe of telopod syncoxite rather low and roundly subtriangular (Fig. 15 C). Mulun Karst, Guangxi................................................................................................... H. mashanorum sp. n. – Central lobe of telopod syncoxite rather high and subquadrate (Fig. 8 C). Guizhou........................ H. yinshi sp. n. 11. Collum with two transverse striae, of which 2 nd obliterated mid-dorsally. Mulun Karst.............................. 12 – Collum as usual, with two complete transverse striae........................................................ 13 12. Tip of telopod syncoxite horns devoid of a setoid filament (Fig. 17 D)................................ H. kunnan sp. n. – Tip of telopod syncoxite horns with a minute setoid filament (Fig. 14 D)............................ H. mulunensis sp. n. 13. Brownish pigment usually retained at least in some parts of head.............................................. 14 – Body completely unpigmented (sometimes except for 2 or 3 caudalmost ocelli).................................. 18 14. Head, antennae and collum usually with traces of brownish coloration, sometimes body either uniformly slightly infuscate (Fig. 9 D–F) or even with a faint pattern of paramedian light brown spots on following tergites........................ 15 – Only head with traces of pigment........................................................................ 16 15. Antennomere 6 ca 2.5 times as long as high. Second tergite with 6 or 7 transverse striae, of which (1) 2–3 crossing the dorsum. Yunnan...................................................................................... H. reducta – Antennomere 6 ca 2.0 times as long as high. Second tergite with 5–7 transverse striae, of which 4 or 5 crossing the dorsum. Guangxi..................................................................................... H. lii sp. n. 16. Head (especially near ocelli and Tömösváry’s organ) and antennae often slightly infuscate, very light brownish. Central lobe of telopod syncoxite broadly subtrapeziform (Fig. 3 C). Hunan........................................ H. youhao sp. n. – Only either ocelli or antennae infuscate. Central lobe of telopod syncoxite considerably narrower..................... 17 17. Only ocelli brownish to dark brown. Larger: width 3.8 –4.0 mm. Tips of telopod syncoxital horns with a minute lobule devoid of a setoid filament (Fig. 13 C). Guangxi......................................................... H. xueju sp. n. – Only antennae sometimes very faintly brownish. Smaller: width 2.5 –3.0 mm. Tips of telopod syncoxital horns with a setoid fil- ament (Fig. 2 E). Hunan........................................................................ H. xia sp. n. 18. Caudomedial process of telopod femur with a distinct distolateral lobe (Fig. 2 C, lo)................................ 19 – Caudomedial process of telopod femur devoid of a distinct distolateral lobe...................................... 21 19. Tip of telopod syncoxite horn with a minute setoid filament. Yunnan.................................. H. albicorporis – Tip of telopod syncoxite horn devoid of a setoid filament..................................................... 20 20. Central lobe of telopod syncoxite nearly as high as lateral horns (Fig. 7 C). Second tergite with six transverse striae, of which three crossing the dorsum. Guizhou............................................................. H. gudu sp. n. – Central lobe of telopod syncoxite evidently lower, roundly subtriangular (Fig. 12 C). Second tergite with at least eight trans- verse striae. Guangxi...................................................................... H. heshang sp. n. 21. Telopod tibial process absent (Fig. 6 C, D). Central lobe of telopod syncoxite rather low and semi-circular (Fig. 6 C)................................................................................................... H. nigu sp. n. – Telopod tibial process present. Central lobe of telopod syncoxite higher.......................................... 22 22. Second tergite with 6 or 7 transverse striae, of which only two crossing the dorsum. Central lobe of telopod syncoxite high, subtrapeziform, slightly concave at ventral margin (Fig. 1 D). Hunan................................... H. tiani sp. n. – Second tergite with 9 or 10 transverse striae, of which seven crossing the dorsum. Central lobe of telopod syncoxite high, nar- row, linguiform (Fig. 5 C). Guizhou.............................................................. H. wuse sp. n.

Hyleoglomeris heshang sp. n. Fig. 12. Material examined: Holotype male (IZAS), China, Guangxi Prov., Xilin County, Zhou Bang Village, Cave Zhou Bang, Cave 2, ca 890 m a.s.l., 24.55353 ��N, 105.11073 ��E, 18.04. 2010, leg. L. Deharveng et al. (CHIgx 10���29). Paratypes. 1 female (IZAS), 1 male, 1 female (MNHN CC 177), 1 male, 1 female (SCAU), 1 male (ZMUM), same locality, together with holotype. Name: To emphasize the animal coming on a lonely cave, ��� heshang ��� in Chinese meaning a ���monk���; a noun in apposition. Diagnosis: Differs in its residual coloration of a few ocelli, as well as by the roundly subtriangular central lobe of the telopod syncoxite, the presence of a distinct chitinized lobe from the telopod caudomedial femoral process, and the absence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.5 ���6.0 mm, width 2.5 ���3.0 mm in both sexes. Holotype 6.0 mm long, 3.0 mm wide. Coloration entirely pallid to sometimes very faintly yellowish, 2 or 3 uppermost ocelli sometimes faintly brownish. Ocelli 5 + 1 or 6 + 1, translucid, often barely visible. Antennomere 6 ca 1.7���1.8 times as long as high. Second tergite with 8 or 9 transverse striae, three starting below, 1 or 2 level to, the remaining striae above schism, 5 or 6 striae (neither 1 st nor last from below) crossing the dorsum. Male leg 17 (Fig. 12 A) with a high and rather regularly rounded outer coxal lobe; telopodite 4 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 12 B) with a narrowly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 12 C���E) with a rather low, roundly subtriangular, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule (Fig. 12 E). Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face. A caudomedial process of tibia absent; tibial tubercle on caudal face evident, papillate. Tarsus rather strongly sigmoid, subacuminate apically. Remark: Due to its troglomorphic traits, this species might well prove to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 16-18, DOI: 10.5281/zenodo.214383

Hyleoglomeris kunnan sp. n. Fig. 17. Material examined: Holotype male (IZAS), China, Guangxi Prov., Huanjiang County, core region of Mulun Karst, 650 m a.s.l., Cave Ganxiao Dong, 25.182633 ºN, 108.031839 ºE, 0 7.11. 2009, leg. Tian Mingyi et al. (CHIgx09–087). Paratypes. 1 female (SCAU), same locality, together with holotype. Name. To emphasize the difficulties in reaching the type locality, “ kunnan ” in Chinese meaning “difficult”. Diagnosis: Differs from congeners except H. mulunensis sp. n. by the 2 nd transverse stria on the collum being obliterated mid-dorsally, from H. mulunensis sp. n. by the absence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 7.5 (holotype) or 9.5 mm (paratype); width 4.0 (holotype) or 4.6 mm (paratype). Coloration uniformly yellow brown (paratype) to entirely pallid (holotype). Ocelli at least 4 + 1, translucid, barely visible. Antennomere 6 ca 2.1 times as long as high. Collum with two transverse striae, but 2 nd obliterated and interrupted mid-dorsally. Second tergite with 10 or 11 transverse striae, four starting below, one level to, the remaining striae above schism, four striae (neither 1 st nor last two ones from below) crossing the dorsum. Male leg 17 (Fig. 17 A) with a high and regularly rounded outer coxal lobe; telopodite 4 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 17 B) with a broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 17 C–E) with a roundly subtrapeziform, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule (Fig. 17 E). Both prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face (Fig. 17 D). Caudomedial process of tibia rather evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, subacuminate apically. Remark: Due to its troglomorphic traits, this species seems to be a troglobite. 1: H. emarginata; 2: H. sinensis; 3: H. aschnae; 4: H. nigu sp. n.; 5: H. yinshi sp. n.; 6: H. eusulcata; 7: H. xia sp. n.; 8: H. youhao sp. n.; 9. H. tiani sp. n.; 10: H. wuse sp. n.; 11: H. mashanorum sp. n.; 12: H. curtisulcata sp. n.; 13: H. kunnan sp. n.; 14: H. mulunensis sp. n.; 15: H. qiyi sp. n.; 16: H. lii sp. n.; 17: H. gudu sp. n.; 18: H. albicorporis; 19: H. heshang sp. n.; 20: H. xueju sp. n.; 21: H. reducta; 22: H. maculata; 23: H. bicolor

Hyleoglomeris wuse sp. n. Fig. 5. Material examined: Holotype male (IZAS), China, Guizhou Prov., Libo County, Maolan Town, Cave Dong Ge Dong, 26.07. 2009, leg. Tian Mingyi & Xue Zhihong (CHIgz09���LWX 13). Paratypes. 1 female (IZAS), 1 male, 1 female (SCAU), 1 male, 2 females (MNHN CC 171), 1 male (ZMUM), 1 female (GNUG), same locality, together with holotype. Name: To emphasize the animal being completely unpigmented, ��� wuse ��� in Chinese meaning ���uncoloured���; a noun in apposition. Diagnosis: Differs from all congeners in the absence of a lobe on the caudomedial process of the telopod femur, of a setoid filament at the tip of the telopod syncoxital horns, coupled with a narrow, rather high, linguiform, central lobe of the telopod syncoxite, as well as a 3 -segmented male telopodite 17. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0��� 7.5 mm (male) or 6.5 ���8.0 mm (female), width 3.2 ���4.0 mm in both sexes. Holotype 6.0 mm long and 3.6 mm wide. Coloration entirely pallid. Ocelli at least 4 + 1, translucid, mostly barely visible. Antennomere 6 ca 2.2 times as long as high. Second tergite with 9 or 10 (usually 10) transverse striae, four starting below, one level to, the remaining striae above schism, normally seven striae (usually 1 st to 3 rd and 5 th to 8 th) from below crossing the dorsum. Male leg 17 (Fig. 5 A) with a rather high and regularly rounded outer coxal lobe; telopodite 3 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 5 B) with a rather narrowly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 5 C, D) with a high, linguiform, rather narrow, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule (Fig. 5 D). Prefemur and femur sparsely micropapillate laterally. Caudomedial femoral process prominent. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather moderately sigmoid, rounded apically. Remark: Only one species of Hyleoglomeris, H. eusulcata, has hitherto been known from Guizhou Province, also a presumed troglobite from Libo County (Golovatch et al. 2006)., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 8-9, DOI: 10.5281/zenodo.214383, {"references":["Golovatch, S. I., Geoffroy, J. - J. & Mauries, J. - P. (2006) Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema, 28 (4), 887 - 915."]}

Hyleoglomeris nigu sp. n. Fig. 6. Material examined: Holotype male (MNHN CC 172), China, Guizhou Prov., Qianxi County, Honglin Town, Cave Luo Sai Dong, 22.11. 2003, leg. L. Latella & D. Avesani. Paratypes. 1 female (MNHN CC 172), 1 female (IZAS), same locality, together with holotype. Name: To emphasize the animal being isolated in its cave, ��� nigu ��� in Chinese meaning a ���nun���; a noun in apposition. Diagnosis: Differs from all congeners in the absence of a lobe on the caudomedial process of the telopod femur, of a setoid filament at the tip of the telopod syncoxital horns, of a telopod tibial process, coupled with a low, roundly subtriangular, central lobe of the telopod syncoxite. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.5 mm (holotype) or 7.0���8.0 mm (paratypes), width 2.8 mm (holotype) or 3.0 mm (paratypes). Coloration entirely pallid. Ocelli at least 5 + 1, translucid, mostly barely visible. Antennomere 6 ca 1.8 times as long as high. Second tergite with seven transverse striae, three starting below, one level to, the remaining striae above schism, 4 (holotype) or 3 (paratypes) striae (various, but not uppermost one) from below crossing the dorsum. Male leg 17 (Fig. 6 A) with a high and regularly rounded outer coxal lobe; telopodite 4 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 6 B) with a V-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 6 C, D) with a rather low, roundly subtriangular, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule. Neither prefemur nor femur micropapillate laterally. Caudomedial femoral process prominent. Caudomedial process of tibia absent; tibial tubercle on caudal face evident, papillate and pilose. Tarsus evidently sigmoid, subacuminate apically. Remark: Due to a number of troglomorphic traits (unpigmented tegument and ocelli), this species seems to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 9-11, DOI: 10.5281/zenodo.214383

Hyleoglomeris eusulcata Golovatch, Geoffroy & Mauri��s, 2006 Fig. 4. Material examined: 1 male, 1 female (IZAS), 1 male, 2 females (MNHN CC 156), 1 male, 1 female (SCAU), 2 females (GNUG), 1 male, 1 female (ZMUM), China, Guizhou Prov., Libo County, Shuipa, Cave Shui Jiang Dong (= Shuipu Da Dong), 28.02. 1995, leg. P. Trontelj. Descriptive notes: Length ca 5.0���9.0 mm, width 2.8���4.2 mm in both sexes. Coloration nearly entirely pallid. Ocelli at least 3 + 1, translucid, barely visible. Collum with one transverse stria. Second tergite with a peculiar transverse sulcus above schism, also with 8���11 transverse striae (versus 8 or 9), three starting below, two level to, the remaining striae above schism, seven from below (versus 5 or 7, as given in Golovatch et al. 2006) crossing the dorsum (never the uppermost one). Male leg 17 (Fig. 4 A) with a rather high (versus low) and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with only one subapical spine. Male leg 18 (Fig. 4 B) with a rather narrowly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 4 C���E) with a high (versus rather low), linguiform (versus subquadrate), sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned with a minute lobule supplied with a subapical (versus apical) setoid filament (Fig. 4 E). Prefemur and, to a lesser extent, femur (versus prefemur alone) micropapillate laterally. Caudomedial femoral process prominent, subapically with a rounded lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather moderately sigmoid, narrowly rounded (versus subacuminate) apically. Remarks: This species has been the only Hyleoglomeris known from Guizhou Province, presumably being a troglobite found in Cave La Tai Dong, Libo County (Golovatch et al. 2006). The above new samples derive from a nearby cave, being in an almost complete agreement with the original description (e.g., the presence of a single transverse stria on the collum, of a strong sulcus on the second tergite, of 4 -segmented male telopodites 17, of a narrow syncoxital notch in male leg 18, of a caudal lobe on the caudomedial femoral process of the telopods, of a setoid filament at the apex of the telopod syncoxital horns). Minor variations concern only the shape of the central lobe of the telopod syncoxite (subquadrate to linguiform), the position of the apical filament of the telopod syncoxital horns (truly apical to subapical), the presence of papillae also in the laterobasal parts of the telopod femur, and the shape of the telopod tarsus (narrowly rounded to subacuminate) and a few other traits (see above)., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 6-7, DOI: 10.5281/zenodo.214383, {"references":["Golovatch, S. I., Geoffroy, J. - J. & Mauries, J. - P. (2006) Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema, 28 (4), 887 - 915."]}

Hyleoglomeris youhao sp. n. Fig. 3. Material examined: Holotype male (IZAS), China, Hunan Prov., Linwu County, Sanhe Town, near Changshali Village, Cave 2, 19.06. 2009, leg. Tian Mingyi & Xue Zhihong (8,12♀ CHIhn09–LWX03). Paratypes. 1 male, 2 females (IZAS), 2 males, 2 females (SCAU), 2 males, 4 females (MNHN CC 170), 1 male, 2 females (ZMUM), 1 male, 2 females (GNUG), same locality, together with holotype. Name: To emphasize the type series being large enough to share with all of the relevant repositories, “ youhao ” in Chinese meaning “friendly”; a noun in apposition. Diagnosis: Differs by the residual coloration on the head, by the presence of a lobe on the caudomedial process of the telopod femur, coupled with a subquadrate, slightly concave, central lobe of the telopod syncoxite. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0–8.0 mm, width 2.8–3.7 mm in both sexes. Holotype 7.0 mm long and 3.2 mm wide. Coloration nearly entirely pallid, but parts of head (especially near ocelli and Tömösváry’s organ) and antennae often slightly infuscate, very light brownish. Ocelli at least 5 + 1, translucid, mostly rather convex. Antennomere 6 ca 2.0– 2.1 times as long as high. Second tergite with 7–9 transverse striae (usually 8), 3 or 4 starting below, one level to, the remaining striae above schism, usually 3 or 4 (2 nd to 4 th or 5 th from below) crossing the dorsum. Male leg 17 (Fig. 3 A) with a rather high and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 3 B) with a rather broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 3 C, D) with a rather high, apically very faintly concave, subquadrate, sparsely setose, central syncoxital lobe flanked by two setose horns with simple apices. Prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a rounded lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, narrowly rounded apically. Remarks: This species shows such troglomorphic traits as a nearly unpigmented body and entirely unpigmented ocelli, so it can well prove to be a troglobite. Residual coloration of the antennae and head is noteworthy.

Hyleoglomeris mulunensis sp. n. Fig. 14. Material examined: Holotype male (IZAS), China, Guangxi Prov., Huanjiang County, Mulun Karst, Cave Xia Dong, ca 650 m a.s.l., 25 �� 10 ��� 13 ������N, 108 ��00��� 17 ������E, 11.03. 2009, leg. Yu Zhuangji & Wei Longtao (CHIgx09���074). Paratypes. 1 male, 1 female (MNHN CC 179), 1 male, 1 female (SCAU), same locality, together with holotype. Name. To emphasize the animal���s provenance from a cave in Mulun Karst. Diagnosis: Differs from congeners except H. kunnan sp. n. in the 2 nd transverse stria on the collum being obliterated mid-dorsally, from H. kunnan sp. n. by the presence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0��� 7.5 mm, width 3.2���3.6 mm in both sexes. Holotype 6.0 mm long, 3.5 mm wide. Coloration entirely pallid. Ocelli 3 + 1 or 4 + 1, translucid, barely visible. Antennomere 6 ca 1.9 ���2.0 times as long as high. Collum with two transverse striae, but 2 nd one obliterated and interrupted mid-dorsally. Second tergite with 8���10 transverse striae, three starting below, one level to, the remaining striae above schism, 3���5 striae (never last three ones from below) crossing the dorsum. Male leg 17 (Fig. 14 A) with a rather low and regularly rounded outer coxal lobe; telopodite 3 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 14 B) with a broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 14 C, D) with a rather low, roundly subtriangular, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by an inconspicuous lobule with a short setoid filament at base (Fig. 14 D). Both prefemur and femur only faintly papillate laterally. Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather slightly sigmoid, subacuminate apically. Remark: Due to showing several troglomorphic traits, this species might well prove to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 19-20, DOI: 10.5281/zenodo.214383

Hyleoglomeris yinshi sp. n. Fig. 8. Material examined: Holotype male (MNHN CC 174), China, Guizhou Prov., Kaiyang County, Cave Xian Yan Dong, 19.02. 2004, leg. S. Prevorčnik & B. Sket. Paratypes. 1 male, 1 female (MNHN CC 174), 1 male, 1 female (IZAS), 1 male (ZMUM), same locality, together with holotype. Name: To emphasize the animal coming from a cave, ��� yinshi ��� in Chinese meaning a ���hermit���; a noun in apposition. Diagnosis: Differs from congeners except H. eusulcata, H. mulunensis sp. n. and H. mashanorum sp. n. in the presence of only one complete transverse stria on the collum, from H. eusulcata by the absence of a transverse sulcus on the second tergite, from both H. mulunensis sp. n. and H. mashanorum sp. n. by the subquadrate central lobe of the telopod syncoxite and the shape of the notch in male syncoxite 18. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.0���6.0 mm (male) or 6.0���7.0 mm (female), width 2.2���2.9 mm (male) or 2.7 ���3.0 mm (female). Holotype 5.0 mm long, 2.3 mm wide. Coloration entirely pallid. Ocelli at least 4 + 1, translucid, barely visible. Antennomere 6 ca 2.0 times as long as high. Collum with one transverse stria. Second tergite with 8���11 transverse striae, two starting below, 1 or 2 level to, the remaining striae above schism, 3���5 striae (various, but never 1 st and/or last from below) crossing the dorsum. Male leg 17 (Fig. 8 A) with a high and highly irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus devoid of subapical spines. Male leg 18 (Fig. 8 B) with a broadly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 8 C, D) with a high, linguiform, apically barely concave, sparsely setose, central syncoxital lobe flanked by two setose horns, each horn being crowned by a short minute lobule. Prefemur micropapillate laterally. Caudomedial femoral process prominent, subapically with a small lobe. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather strongly sigmoid, narrowly rounded apically. Remark: Due to its troglomorphic traits (unpigmented tegument and ocelli), this species seems to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 11-12, DOI: 10.5281/zenodo.214383

Hyleoglomeris xueju sp. n. Fig. 13. Material examined: Holotype male (IZAS), China, Guangxi Prov., Du'an County, Xia'ao Town, Yao Nan Village, Cave Yao Nan Dong, Cave 1, ca 350 m a.s.l., 24.244233 ��N, 107.940118 ��E, debris, Berlese extraction, 26.04. 2010, leg. L. Deharveng et al. (CHIgx 10���65). Paratypes. 1 female (IZAS), 1 male, 1 female (MNHN CC 178), 1 male (SCAU), 1 male (ZMUM), same locality, together with holotype. Name. To emphasize the animal���s provenance from a cave, ��� xueju ��� in Chinese meaning a ���cave-dweller���; a noun in apposition. Diagnosis: Differs in its residual coloration of the ocelli, as well as by the high, linguiform, central lobe of the telopod syncoxite, the presence of a distinct chitinized lobe from the telopod caudomedial femoral process, and the absence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0��� 7.5 mm, width 3.8 ���4.0 mm in both sexes. Holotype 7.0 mm long, 4.0 mm wide. Coloration almost entirely pallid, but ocelli brownish to dark brown. Ocelli 4 + 1 or 5 + 1, translucid, well visible. Antennomere 6 ca 1.9 ���2.0 times as long as high. Second tergite with 7���9 transverse striae, 1���3 starting below, 1 or 2 level to, the remaining striae above schism, 4 or 5 striae (various, but not last from below) crossing the dorsum. Male leg 17 (Fig. 13 A) with a high and irregularly shaped outer coxal lobe; telopodite 4 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 13 B) with an arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 13 C, D) with a high, linguiform, central syncoxital lobe flanked by two setose horns, each latter crowned by a particularly inconspicuous lobule. Both prefemur and, to a lesser degree, femur finely papillate laterally. Caudomedial femoral process (Fig. 13 D) prominent, subapically with a distinct lateral lobe on caudal face. Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, papillate. Tarsus rather evidently sigmoid, narrowly rounded apically. Remark: Due to some of its traits being troglomorphic, this species might well prove to be a troglobite, but pigmented ocelli might rather be evidence of troglophily., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 18-19, DOI: 10.5281/zenodo.214383

Hyleoglomeris curtisulcata sp. n. Fig. 16. Material examined: Holotype male (IZAS), China, Guangxi Prov., Huanjiang County, Mulun Karst, Cave Gang Lai Dong, 25.11278 ��N, 107.98309 ��E, 14.03. 2005, leg. L. Deharveng & A. Bedos (CHIgx05��� 111). Paratypes. 1 female (IZAS), 1 male, 1 female (MNHN CC 180), same locality, together with holotype. Name: To emphasize the presence of a rather short transverse sulcus on the second tergite. Diagnosis: Differs from congeners except H. eusulcata and H. speophila Golovatch, Geoffroy & Mauri��s, 2006, from Laos (Golovatch et al. 2006), by the presence of a transverse sulcus above the schism on the second tergite (yet shorter, reaching only ca 1 / 5 height of the second tergite), from H. speophila by the presence of only one complete stria on the collum and of a lobe on the caudomedial process of the telopod femur, from H. eusulcata by the absence of a setoid filament at the tip of the syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 6.0 (paratype) to 7.5 mm (holotype) in male, 7.5 ���8.0 mm in female; width 3.7 (holotype) or 3.8 mm (paratype) in male, 3.9 or 4.5 mm in female. Coloration entirely pallid. Ocelli at least 3 + 1, translucid, barely visible. Antennomere 6 ca 2.1 times as long as high. Collum with one transverse stria. Second tergite with six transverse striae, two starting below, one level to, the remaining striae above schism, 3 or 4 striae (not last two ones from below, both forming a rather short, but evident transverse sulcus above schism) crossing the dorsum. Male leg 17 (Fig. 16 A) with a rather low and irregularly shaped outer coxal lobe; telopodite 3 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 16 B) with an arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 16 C���E) with a linguiform, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by an inconspicuous lobule (Fig. 16 E). Both prefemur and, to a lesser extent, femur micropapillate laterally. Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face (Fig. 16 D). Caudomedial process of tibia evident, membranous; tibial tubercle on caudal face distinct, non-papillate. Tarsus rather evidently sigmoid, narrowly rounded apically. Remark: Due to its troglomorphic traits, this species is likely to represent a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 21-22, DOI: 10.5281/zenodo.214383, {"references":["Golovatch, S. I., Geoffroy, J. - J. & Mauries, J. - P. (2006) Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia. Zoosystema, 28 (4), 887 - 915."]}

Hyleoglomeris tiani sp. n. Fig. 1. Material examined: Holotype male (IZAS), China, Hunan Prov., Linwu County, Huatang Town, Shimen Village, Cave Long Dong, 19.06. 2009, leg. Tian Mingyi & Xue Zhihong (CHIhn09���LWX01). Paratype. 1 male (SCAU), same locality, together with holotype. Name: Honours Tian Mingyi, one of the collectors. Diagnosis: Differs from all congeners in the absence of a lobe on the caudomedial process of the telopod femur, of a setoid filament at the tip of the telopod syncoxital horns, coupled with a rather short, subtrapeziform, faintly concave central lobe of the telopod syncoxite, a widely arch-shaped notch of male syncoxite 18, and a 3 - segmented male telopodite 17. See also Key below. Description: Length ca 6.0���7.0 mm, width 2.8 (paratype) or 3.0 mm (holotype). Coloration entirely pallid. Ocelli at least 5 + 1, translucid, mostly rather convex; T��m��sv��ry���s organ transverse-oval, only slightly wider than long. Antennomere 6 ca 2.0 times as long as high. Collum with two transverse striae. Second tergite with a narrow hyposchism not reaching behind the caudal tergal margin; 6 or 7 superficial transverse striae, 3 or 4 starting below schism, one level to, all others above schism while two (2 nd and 3 rd from below) crossing the dorsum. Male anal shield regularly rounded at caudal margin. Leg 1 (Fig. 1 A) with a conspicuous, distomesal, finger-shaped, spinigerous, coxal process (k). Leg 17 (Fig. 1 B) with a rather low and regularly rounded outer coxal lobe (cxl); telopodite 3 -segmented, tarsus with two subapical spines. Leg 18 (Fig. 1 C) with a broadly arch-shaped syncoxital notch (scxn); telopodite 4 -segmented. Telopods (Fig. 1 D, E) with a high, apically slightly concave, sparsely setose, central syncoxital lobe (scxl) flanked by two setose horns (scxh), each latter crowned with a minute, elongate lobule devoid of adjacent structures (Fig. 1 E). Prefemur and, to a lesser extent, femur micropapillate laterally. Prefemur with a well-developed frontomedial trichostele (= flagelliferous digit) (fpft). Femur with a smaller frontomedial trichostele (fft). Caudomedial femoral process (cfp) prominent, directed distomedially at ca 100 �� to femur, mostly strongly chitinized, apically with a small membranous sac (s). Caudomedial process of tibia (tip) evident, sac-shaped, membranous, with an evident, papillate tubercle at base. Tarsus subacuminate apically. Remark: Since this species shows such troglomorphic traits as entirely unpigmented tegument and ocelli, it can well prove to be a troglobite., Published as part of Golovatch, Sergei I., Liu, Weixin & Geoffroy, Jean-Jacques, 2012, Review of the millipede genus Hyleoglomeris Verhoeff, 1910 in China, with descriptions of new species (Diplopoda, Glomerida, Glomeridae), pp. 1-27 in Zootaxa 3358 on pages 2-4, DOI: 10.5281/zenodo.214383

Hyleoglomeris heshang sp. n. Fig. 12. Material examined: Holotype male (IZAS), China, Guangxi Prov., Xilin County, Zhou Bang Village, Cave Zhou Bang, Cave 2, ca 890 m a.s.l., 24.55353 °N, 105.11073 °E, 18.04. 2010, leg. L. Deharveng et al. (CHIgx 10–29). Paratypes. 1 female (IZAS), 1 male, 1 female (MNHN CC 177), 1 male, 1 female (SCAU), 1 male (ZMUM), same locality, together with holotype. Name: To emphasize the animal coming on a lonely cave, “ heshang ” in Chinese meaning a “monk”; a noun in apposition. Diagnosis: Differs in its residual coloration of a few ocelli, as well as by the roundly subtriangular central lobe of the telopod syncoxite, the presence of a distinct chitinized lobe from the telopod caudomedial femoral process, and the absence of a setoid filament at the tip of the telopod syncoxital horns. See also Key below. Description: All characters like in H. tiani sp. n., except as follows. Length ca 5.5 –6.0 mm, width 2.5 –3.0 mm in both sexes. Holotype 6.0 mm long, 3.0 mm wide. Coloration entirely pallid to sometimes very faintly yellowish, 2 or 3 uppermost ocelli sometimes faintly brownish. Ocelli 5 + 1 or 6 + 1, translucid, often barely visible. Antennomere 6 ca 1.7–1.8 times as long as high. Second tergite with 8 or 9 transverse striae, three starting below, 1 or 2 level to, the remaining striae above schism, 5 or 6 striae (neither 1 st nor last from below) crossing the dorsum. Male leg 17 (Fig. 12 A) with a high and rather regularly rounded outer coxal lobe; telopodite 4 -segmented, tarsus with two subapical spines. Male leg 18 (Fig. 12 B) with a narrowly arch-shaped syncoxital notch; telopodite 4 -segmented. Telopods (Fig. 12 C–E) with a rather low, roundly subtriangular, sparsely setose, central syncoxital lobe flanked by two setose horns, each latter crowned by a minute lobule (Fig. 12 E). Caudomedial femoral process prominent, subapically with a distinct lateral lobe on caudal face. A caudomedial process of tibia absent; tibial tubercle on caudal face evident, papillate. Tarsus rather strongly sigmoid, subacuminate apically. Remark: Due to its troglomorphic traits, this species might well prove to be a troglobite.

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul, Vandenspiegel, Didier (2011): The millipede genus Plusioglyphiulus Silvestri, 1923 in Thailand (Diplopoda, Spirostreptida, Cambalopsidae) 2940. Zootaxa 2940 (1): 1-63, DOI: 10.11646/zootaxa.2940.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2940.1.1

Glomeris mohamedanica Attems, 1900 Figs 24, 25 Glomeris europaea striata var. mohamedanica Attems, 1900: 303 (D). Glomeris mohamedanica ��� Schubart 1953: 218 (L). Type material: Tunisia, B��ja Gov., Medjez El Bab (now Mejez El Bab), date?, leg.?, ��� lectotype (here designated) (NHMW 3913), paralectotypes: 3 ���, 2 ♀ (NHMW 3913), 1 ��� (MNHN CC 153). The designation of a lectotype (from 350 syntypes, see Attems 1900) seems advisable in order to ensure that the name-bearing type shows the diagnostically important characteristics of the male sex. Other material: Tunisia, B��ja Gov., Nefza (30 km of B��ja), 2003, leg. N. Akkari, 1 ���, 1 ♀ (MNHN CC 153). Same locality, 36��57���61���N, 8��56���51���E, open area with scattered vegetation, under stones, 27.II.2004, leg. N. Akkari, 3 ���, 5 ♀, 3 juv. (MNHN CC 153). Same locality, 2.X.2005, leg. N. Akkari, 1 ���, 3 ♀ (NMNH). Tunisia, Jen- douba Gov., Beni Mtir (8 km S of A��n Draham), mixed oak forest dominated by Quercus suber, slope, under stones, 19.II.2007, leg. N. Akkari, 10 ��� (MNHN CC 153), 3 ��� (ZMUM), 2 ��� (NMNH), 2 # (FMNH). Same locality, alt. approx. 500 m, 19.II.2007, leg. N. Akkari, 2 ���, 2 ♀ (ZMUC 200113). Tunisia, Jendouba Gov., Tabarka, 10.XII.2003, leg. N. Akkari, 1 ♀ (FMNH). Tunisia, Jendouba Gov., 40 km of Jendouba, near frontier, 17.V.1988, leg. Z. M. Cop. Exp., 2 M (ZMUC 200108). Short description: Length up to 12 mm, width up to 6.0 mm. Coloration dark brown to blackish, with contrasting yellow bands, pattern as in Figs 24A, B. ��� legs 17, 18 and 19 (telopods) as in Figs 25 A-D. Remarks: This species is unusual among the presumably native North African Glomeris in belonging to the ��� Eurypleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 75-76, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Attems C (1900) Uber die Farbung von Glomeris und Beschreibung neuer oder wenig gekannter palaearktischer Myriopoden. Archiv fur Naturgeschichte 66 (1, 3): 297 - 320.","Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952)."]}

Glomeris klugii Brandt, 1833 Figs 10-14 Glomeris klugii Brandt, 1833: 195 (D). Glomeris marmorata Brandt, 1833: 196 (D), syn. n.! Glomeris fuscomarmorata Lucas, 1846: 284 (D), syn. n.! Glomeris pustulata Latr., var. marmorata ��� Brandt 1840b: 42 (R); 1841b: 148 (R); Gervais 1847: 73 (R). Glomeris fusco-marmorata (sic!) ��� Gervais 1847: 74 (R); Lucas 1849: 327, plate 1, fig. 4 (R); Pocock 1892: 27 (F); Brolemann 1921: 101 (L). Glomeris conspersa forma genuina (sic!) ��� Attems 1908: 105 (F). Glomeris fuscomarmorata ��� Schubart 1953: 218 (L); Abrous-Kherbouche & Mauri��s 1996: 586 (L). Glomeris maculosa Verhoeff, 1921: 27, fig. 3 (D), syn. n.! Glomeris maculosa ��� Schubart 1953: 218 (L). Glomeris conspersa ��� Abrous-Kherbouche & Mauri��s 1996: 572, 586 (F, L). Type material examined: Germany, ���Hercynia���, leg. Zimmermann, 2 ♀? syntypes of Glomeris marmorata Brandt, 1833 (ZMUB 39). No lectotype designation has been made here, particularly as we did not directly examine the material. Algeria, Philippeville (now Skikda), date?, leg. et det. H. Lucas, 1 ♀ syntype (���type���) of Glomeris fuscomarmorata (MNHN CC 043). Algeria, Gorges de la Chiffa, date?, leg. H. Ribaut, ��� ���type��� of Glomeris maculosa Verhoeff, 1921 (ZSM A20070924 + micropreparation A 20032085). The ��� represents only part of the syntype series, which was said to consist of 2 ��� and 1 ♀ (Verhoeff 1921). Other material (nearly all previously identified as Glomeris fuscomarmorata): Algeria, ���Bona��� (now Annaba), leg. F. Meinert, 5 ♀ (ZMUC). Algeria, le Ruisseau (environs of Algiers), 18.XII.1892, leg. P. Lesne, 2 ♀ (MNHN CC 168, entry 6-97). Algeria, Ravin de la Femme Sauvage (environs of Algiers), XII.1892, leg. P. Lesne, 2 ���, 2 ♀ (MNHN CC 168, entry 6-97). Algeria, loc.?, 1898, leg. Noucelhier, 1 ���, 3 ♀ (MNHN CC 168). Algeria, Massif de l���Edough, VII-VIII.1918, leg. Ed. Ch. (?), 1 ��� juv., 4 ♀ (MNHN CC 168). Algeria, near Algiers, ruisseau des Singes, date?, leg. et det. H. Ribaut, 1 ���, 1 ♀, 1♀ juv. (MNHN CC 043). Algeria, Algiers, 3.IV.1907, leg. et det. H. Ribaut, 2 ♀ (MNHN CC 043). Tunisia (Khroumirie), Jendouba Gov., A��n Draham, V-VI.1906, leg. H. Gadeau de Kerville, 1 ��� (MNHN CC 043). Same locality, A��n Draham, Col des Ruines, 36��47���N, 8��41���E, 19.XI.2003, leg N. Akkari, 1 ♀ (NMNH). Same locality, 5.II.2004, leg N. Akkari, 3 ♀ (ZMUM). Same locality, 19.XI.2003, leg N. Akkari, 1 ♀ (NMNH). Same locality, 18.IV.2004, leg. N. Akkari, 1 juv. (ZMUM). Same locality, alt. approx. 735 m, 9.III.2009, leg. N. Akkari and H. Enghoff, 1 ♀ (ZMUC). Same locality, alt. approx. 710 m, 11.III.2009, leg. N. Akkari and H. Enghoff, 3 ���, 1 ��� juv., 11 ♀ (ZMUC). Same locality, A��n Draham area, 5-18.V.1988, leg. Z. M. Cop. Exp., 3 ���, 2 ♀ (ZMUC 200107). Same locality, 7 km S of A��n Draham, Les Ch��nes, 22.03.1986, leg. Z. M. Cop. Exp., 2 ���, 6 ♀ (ZMUC 200109). Same locality, Fernana, 36��43���59���N, 8��40���43���E, alt. approx. 750 m, Quercus-Erica forest, 9.III.2009, leg. N. Akkari and H. Enghoff, 2 ��� juv., 3 ♀ (ZMUC). Same locality, A��n Draham, Beni M���Tir, 36��43���51���N, 8��42���19���E, alt. approx. 590 m, Quercus-Erica forest, 10.III.2009, leg. N. Akkari and H. Enghoff, 2 ���, 1 ��� juv., 2 ♀ (ZMUC). Brief redescription: Length up to 13.5 mm, width up to 6.25 mm. Coloration mostly vivid, very distinctly marbled, pattern as in Figs 10-13. Syncoxite of telopods as in Figs 14A, B, with only minor variations in shape of central lobe and in delicately bifid coxal horns. Remarks: The presumed type series of G. marmorata, even though represented by two dry, pinned specimens, has still preserved its colour pattern sufficiently well (Fig. 11) to compare it with the beautiful illustrations by Koch (1863) and hence to unequivocally synonymize this species here with G. klugii, the holotype of which has been revised elsewhere (Golovatch 2003). This formalizes Brolemann���s (1921) informal synonymy ��� G. conspersa C. Koch, 1847 (= marmorata)��� proposed in his checklist. The syntype of G. fuscomarmorata examined here has faded completely, apparently due to the long preservation in alcohol. Fortunately, most of the other samples from North Africa, especially fresh ones, have retained their coloration (Figs 10, 12), which matches quite closely the pattern depicted by Lucas (1849), based on a then recently collected syntype (Fig. 13). We are certain, however, that this species must also have been printed somewhat too dark and red compared to its natural coloration in Lucas��� (1849) Plate 1, likely because Lucas used dry material (cf Figs 11 and 13). On the other hand, G. klugii in Europe is known to exist in two colour morphs, the dark ��� undulata ��� and the light ��� conspersa ��� (Hoess 2000), of which ��� conspersa ��� is much more widespread and occupies peripheral parts of the species��� distribution area, including North Africa. Could material of ��� undulata ��� have served, at least in part, for Lucas��� (1846, 1849) descriptions and illustrations? Hoess (2000) marked as questionable populations of ��� conspersa ��� from a few small, outlying areas in the Balkans, near Algiers and Tunis, but we can confirm the presence of G. klugii in North Africa. We suggest that it could well have arrived there, particularly at the largest sea ports, through commercial activities, which have been going on since prehistoric times throughout the Mediterranean. The synonymy of G. klugii and G. fuscomarmorata proposed here therefore appears fully justified. Furthermore, we must give due tribute to Brandt (1840b, 1841a, b) who, already at the very beginning of diplopodological explorations in North Africa, wrote that his G. marmorata from Germany and Algeria were identical. Although he failed to recognize that his own G. klugii and G. marmorata actually represented the same speciesapparently because the holotype of G. klugii (surprisingly) did not show any striae on the thoracic shield, retained (in alcohol) its generally light coloration and was thought to have come from Egypt or Syria���he was essentially correct in thinking that the same species could exist on both continents. Likewise correct have been the very few subsequent records of ��� conspersa ��� in Algeria and Tunisia (Attems 1908; Abrous-Kherbouche and Mauri��s 1996), whereas most other authors believed that the North African fauna, including that of Glomeris, is fully endemic. The syntype of G. maculosa is a ��� conspersa ��� specimen of G. klugii, with the colour pattern still well traceable. Hence the synonymy of these names is also proposed here. The fact that G. klugii belongs to the ��� Eurypleuromeris ��� type provides an additional indication of its probable introduction to North Africa from Europe. Moreover, even though G. conspersa is the type-species of Eurypleuromeris, Verhoeff (1921) mistakenly attributed his G. maculosa to the ��� Stenopleuromeris ��� type. Indeed, the anterior part of tergum 3 in the syntype of G. maculosa that we have examined is probably a little narrower than is usual for European or other North African ��� conspersa ��� specimens of G. klugii, but this variation seems too modest to be considered a reliable distinction, perhaps even reflecting individual rather than geographical variation. This is another good reason to abandon the subgeneric division of Glomeris., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 62-67, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Brandt JF (1833) Tentaminum quorundam monographicorum Insecta Myriapoda Chilognatha Latreilli spectantium prodromus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 6: 194 - 209.","Lucas H (1846) Notes sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du nord de l'Afrique. Revue de Zoologie pour la Societe Cuvierienne 9: 283 - 289.","Brandt JF (1840 b) Remarques critiques sur les especes qui composent le genre Glomeris, suivies de quelques observations sur leur distribution geographique des especes en general. Bulletin scientifique publie par l'Academie Imperiale des Sciences de St. Petersbourg 7 (4 / 5): 37 - 44.","Gervais P (1847) Myriapodes. In: Walckenaer CA, Gervais P (Eds) Histoire naturelle des Insectes Apteres. Roret, Paris, 623 p.","Lucas H (1849) Histoire naturelle des animaux articules. Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842. Sciences Physiques, Zoologie 1: Crustacees, Arachnides, Myriapodes. Atlas. Imprimerie Imperiale, Paris, 322 - 390.","Pocock RI (1892) On the Myriopoda and Arachnida collected by Dr. Anderson in Algeria and Tunisia. Proceedings of the Zoological Society of London 1892: 24 - 28.","Brolemann HW (1921) Liste des Myriapodes signales dans le nord de l'Afrique. Bulletin de la Societe des Sciences Naturelles du Maroc 1 (3 - 6): 99 - 110.","Attems C (1908) Note sur les Myriapodes recueillis par M. Henri Gadeau de Kerville en Khroumirie et description de deux especes et d'une variete nouvelles provenant de cette region de la Tunisie. In: Gadeau de Kerville H (Ed) Voyage zoologique en Khroumirie (Tunisie). Rouen 1906, 103 - 116.","Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952).","Abrous-Kherbouche O, Mauries J-P (1996) Decouverte du genre Archipolydesmus en Algerie: description de trois especes nouvelles (Diplopoda, Polydesmida, Polydesmidae). Bulletin du Museum national d'Histoire naturelle, 4 e serie, 18 A (Zool.) (3 / 4): 571 - 587.","Verhoeff KW (1921) Chilognathen-Studien (91. Diplopoden-Aufsatz). Archiv fur Naturgeschichte 86 A (12): 23 - 80 (for 1920).","Koch CL (1863) Die Myriapoden. Getreu nach den Natur abgebildet und beschrieben 1: 1 - 134. H. W. Schmidt, Halle.","Golovatch SI (2003) Two new species of Glomeridellidae (Diplopoda: Glomerida) from the Middle East. Arthropoda Selecta 11 (4): 255 - 258 (for 2002).","Hoess R (2000) Bestimmungsschlussel fur die Glomeris - Arten Mitteleuropas und angrenzender Gebiete (Diplopoda: Glomeridae). Jahrbuch des Naturhistorischen Museums Bern 13: 3 - 20.","Brandt JF (1841 a) Uber die in der Regentschaft Algier vom Herrn Dr. Wagner beobachteten Myriapoden. In: Wagner M (Ed) Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838. Leipzig 3, 282 - 289."]}

Key to the known Glomeris species of North Africa 1 Tegument entirely pallid, rarely only ocelli dark. Cavernicoles................... 2 ��� Head and terga distinctly pigmented, colour pattern evident, coloration mostly vivid. Epigean................................................................................. 3 2 Collum with a single transverse stria. Syncoxital horns of telopod clearly bifid (Fig. 2D, E). Libya..................................................... G. monostriata sp. n. ��� Collum with the usual two transverse striae. Syncoxital horns of telopod surmounted with a setoid filament (Fig. 1F). Kabylia, Algeria....................................................................................................... G. troglokabyliana sp. n. 3 Colour pattern of terga 2-11 without evident, light, paramedian spots, but with alternating light and dark transverse bands (Figs 10, 12, 24).............. 4 ��� Colour pattern of terga 2-(6)11 with 1+1 or 2+2 more or less evident light spots (Figs 3, 4, 7, 15-17, 20-22, 26, 27)................................................... 5 4 Coloration of caudal halves of terga distinctly marbled throughout (Figs 10, 12)........................................................................................... G. klugii ��� Transverse bands on terga at most only slightly marbled in contact zones (Fig. 24).......................................................................... G. mohamedanica 5 Unusually large yellowish to orange spots on tergum 6, following terga 7-11 uniformly blackish (Figs 20-22)...................................... G. carthaginiensis ��� Spots on tergum 6 of same size as on adjacent terga................................... 6 6 Median spots on tergum 2 much smaller than lateral ones..... G. anisosticta ��� Either both pairs of spots on tergum 2 comparable in size or lateral ones (if present) smaller than median ones............................................................. 7 7 Coloration uniformly blackish, lighter spots (1+1 or 2+2) very vague, marbled (Figs 15-18)................................................................... G. sublimbata ��� Background coloration brown to blackish, lighter spots more distinct........ 8 8 Tergum 2 with a single central light spot against a black-brown background. Pygidium uniformly black-brown. Morocco......................... G. brolemanni ��� Tergum 2 with 2+2 light spots................................................................... 9 9 Background coloration light to dark brown, drab, pattern as in Figs 26, 27.................................................................................................... G. punica ��� Background coloration dark brown to blackish, pattern vivid.................. 10 10 Light axial stripe absent (Figs 6-8).................................... G. flavomaculata ��� Light axial stripe present (Figs 3, 4).................................. G. colorata sp. n., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on page 83, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461

Glomeris numidia Verhoeff, 1921: 28 (D), syn. n.! Glomeris numidia ��� Schubart 1953: 218 (L). Glomeris punica ��� Schubart 1953: 218 (L). Type material: Tunisia, Bou-Kournine (now Bou Kornine), date?, leg.?, ��� lectotype of G. punica Attems, 1900 (here designated) (NHMW 3910), paralectotypes: 5 ���, 1 ♀ (NHMW 3910). The designation of a lectotype (of 60 syntypes, see Attems 1900) seems advisable in order to ensure that the name-bearing type shows the diagnostically important characteristics of the male sex. Algeria, Gorges de la Chiffa, date?, leg. H. Ribaut, 2 ��� syntypes of Glomeris numidia Verhoeff, 1921 (��� ���type���, ZSM A20070944 + micropreparation A 20032086 and ��� ���?type���, A 20091438). Both ������ represent only part of the type series stated to have consisted of 2 ��� and 2 ♀ (Verhoeff 1921). Other material: Algeria, La Chiffa, date?, leg. Ch. Alluaud (219), 2 ���, 5 ♀ (MNHN CC 018, collection Brolemann CCXIX). Algeria, le Ruisseau (environs of Algiers), 18.XII.1892, leg. P. Lesne, 1 ��� (MNHN CC 018, entry 6-97). Algeria, Yakouren (Kabylia), 10.VI.1893, leg. P. Lesne, 1 ��� (MNHN CC 018, entry 6-97). Tunisia (Kroumirie), Jendouba Gov., A��n Draham, V-VI.1906, leg. H. Gadeau de Kerville, 6 ���, 19 ♀ (MNHN CC 018). Tunisia, B��ja Gov., Jebel El Jouza, close to Amdoun Village, under stones, 10.III.2005, leg. N. Akkari, 2 juv. (MNHN CC 018). Same locality, under stones and barks, 25.V.2005, leg. N. Akkari, 2 ���, 1 ♀ (NMNH). Tunisia, Jendouba Gov., A��n Draham, Col des Ruines, mixed oak forest with Quercus faginea and Q. suber, 36��47���N, 8��41���E, under stone, 19.XI.2003, leg. N. Akkari, 1 juv. (MNHN CC 018). Tunisia, Le Kef Gov., Sakiet Sidi Youssef, coniferous forest dominated by Pinus halepensis, under stones, 24.X.2003, leg. N. Akkari, 4 ���, 5 ♀ (ZMUM). Tunisia, Zaghouan Gov., Jebel Zaghouan, alt. 350-390 m, 25.II.2007, leg. N. Akkari, 2 ���, 2 ♀ (ZMUC 200114). Tunisia, Jendouba Gov., 15 km E of Tabarka, 25.III.1986, leg. Z. M. Cop. Exp., 1 ��� (ZMUC 200111). Tunisia, Thala Gov., 12 km S of Thala, 10.III.1986, leg. Z. M. Cop. Exp., 1 ��� (ZMUC 200110). Tunisia, Zaghouan Gov., 3 km SE of Zaghouan, NE side of Djebel Zaghouan, 1.XII.1974, leg. H. Wald��n, 1 ♀ (Museum G��teborg). Tunisia, Tunis Gov., Bou Kornine National Park, 18 km SE of Tunis, close to Hammam Lif Town, 36��42���53���N, 10��20���68���E, alt. 105-150 m, Thuya ��� Eucalyptus grove, dry river bed, under stones and logs, 4.III.2008, leg. P. Stoev and N. Akkari, 2 ���, 9 ♀ (FMNH), 6 ���, 4 ♀, 1 juv. (NMNH). Tunisia, Zaghouan Gov., Jebel Zaghouan, surroundings of ���Temple des Eaux���, 36��23���40���N, 10��08���09��� E, alt. approx. 300 m, forest close to road, under stones and in leaf litter, 19.III.2008, leg. N. Akkari and P. Stoev, 2 ���, 4 ♀ (FMNH), 4 ���, 2 ♀, 1 juv. (NMNH). Tunisia, Jendouba Gov., 15 km of Tabarka, 36��49���97��� N, 8��42���34���E, alt. approx. 230 m, Quercus suber forest, slope, under stones and barks, 21.III.2008, leg. P. Stoev and N. Akkari, 1 ���, 2 ♀, 1 ♀ juv. (FMNH), 1 ♀ (NMNH). Tunisia, B��ja Gov., 13 km of Nefza (road Tabarka-Nefza), 36��57���61���N, 8��56���51��� E, alt. approx. 150 m, Pinus forest, under stones, 23.III.2008, leg. P. Stoev and N. Akkari, 2 ���, 1 ♀ (FMNH). Tunisia, Jendouba Gov., close to A��n Draham, Fernana, 9.III.2009, leg. N. Akkari and H. Enghoff, 1 ��� juv., 3 ♀ (ZMUC). Tunisia, Zaghouan Gov., Jebel Zaghouan, 13.III.2009, leg. N. Akkari and H. Enghoff, 4 ���, 2 ♀, 1 ��� juv. (ZMUC). Tunisia, Jendouba Gov., 9 km of Hammam Bourguiba (W of A��n Draham), 36��48���05���N, 8��39���54���E, alt. approx. 380 m, humid Pinus forest, close to river, under stones, logs and in litter, 22.III.2008, leg. P. Stoev and N. Akkari, 1 ���, 1 juv. (NMNH). Short description: Length up to 12 mm, width up to 6.0 mm. Coloration light to dark brown, terga 2-11 each with 2+2, rather vague, paramedian spots; pattern as in Fig. 27. ��� legs 17, 18 and 19 (telopods) as in Figs 28 A-F. An always high lateral lobe of ��� coxite 17 is remarkable (Fig. 28A). Remarks: This species belongs to the ��� Stenopleuromeris ��� type. Schubart (1963) mistakenly attributed it to the ��� Eurypleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 77-79, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Verhoeff KW (1921) Chilognathen-Studien (91. Diplopoden-Aufsatz). Archiv fur Naturgeschichte 86 A (12): 23 - 80 (for 1920).","Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952).","Attems C (1900) Uber die Farbung von Glomeris und Beschreibung neuer oder wenig gekannter palaearktischer Myriopoden. Archiv fur Naturgeschichte 66 (1, 3): 297 - 320.","Schubart O (1963) Uber einige Diplopoden aus Algier. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc 43 (2): 79 - 94."]}

Eutrichodesmus reductus Golovatch, Geoffroy, Mauri��s & VandenSpiegel, sp. n. urn:lsid:zoobank.org:act: 1726A727-8FC2-4BF8-807D-8DFC1C54380C Figs 31-33 Type material: Indonesia, Sulawesi Selatan, kab. Maros: Samanggi, Gua Saripa Cave, hand collected, 18.VIII.1990, leg. A. Bedos and L. Deharveng (SULS-214), holotype ��� (MZB), paratypes: 1 ��� (MNHN JC 325), 1 ♀ (SEM). Name: To emphasize the strongly underdeveloped paraterga 2 and lack of metatergal tuberculation. Diagnosis: Differs from congeners except E. communicans Golovatch, Geoffroy, Mauri��s & VandenSpiegel, 2009 by the strongly underdeveloped paraterga 2, coupled with 19 body segments and the absence of metatergal tuberculation; from E. communicans by a very short tergal trichome, from it and other congeners in a few minor details of gonopod structure (in particular, the shape of the telopodite). Description: Length of adults of both sexes ca 4.0- 4.2 mm, width 0.45-0.5 mm, body broadest at midbody segments. Holotype ca 4.0 mm long and 0.45 mm wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 31). Adults with 19 segments, body subcylindrical (Figs 31, 32A), volvation apparently incomplete due to insufficiently wide and low paraterga. Head (Fig. 33A) slightly transverse, with a poorly separated pair of very low, paramedian tubercles above antennal sockets; antennae relatively short and clavate, antennomere 6 longest; tegument (Figs 33 C-E) and many other characters (Figs 32F, 33B) much as in E. distinctus sp. n.; collum and following metaterga devoid of tuberculation, beset with numerous, irregularly arranged, extremely short setae (Fig. 32 B-F, 33C-E). Paraterga directed ventrolaterad, slightly interrupting contour of convex dorsum, short, not reaching level of venter (Figs 32 A-D); paraterga 2 (Fig. 32B) only a little enlarged compared to following ones, indistinctly lobulate both anterolaterally and laterally, with only a single noticeable lobe forming a schism ledge, both schism and hyposchism very small; paraterga 3 and 4 not narrower than others, overlap typical already from paraterga 4, not paraterga 5 as in preceding congeners. Paraterga broadly rounded caudally, very indistinctly lobulate laterally and with a single evident lobulation caudolaterally (Figs 32 C-E, 33C). Limbus distinctly denticulate, almost hidden by nearby abundant microvilli (Fig. 33D, E). Pore formula normal, ozopores located dorsally near base of caudal corner of paraterga (Fig. 33C). Legs relatively long and slender, evidently surpassing edge of paraterga (Fig. 33B). Gonopods (Fig. 33F, G) relatively complex. Coxae abundantly micropapillate, but only with a few macrosetae. Telopodite elongate, only slightly arcuate, with a large, papillate, peculiar, distofemoral outgrowth (dp) in distal one-third and a slender but short solenomere bearing a few small setae (but no pad!) subapically at base of a very complex tip represented by two erect teeth, a longer uncus and a group of minute outgrowths. Remarks: This small-bodied, pallid, non-volvatory (= ���haplodesmid���, see Golovatch et al. 2009) species seems to be especially close, morphologically as well as geographically, to E. communicans from Vanuatu, Melanesia, southwestern Pacific (Golovatch et al. 2009), possibly representing still one more troglobite., Published as part of Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauri��s, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species, pp. 1-46 in ZooKeys 12 (12) on pages 37-40, DOI: 10.3897/zookeys.12.167, http://zenodo.org/record/576460, {"references":["Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2009) Review of the millipede family Haplodesmidae, with descriptions of some new or poorly-known species (Diplopoda: Polydesmida). In: Golovatch SI, Mesibov R (Eds) Advances in the Systematics of Diplopoda I. ZooKeys 7: 1 - 53."]}

Glomeris brolemanni Schubart, 1960 Glomeris brolemanni Schubart, 1960: 164, figs 1-3 (D). Remarks: Unfortunately, no new material of this species could be obtained for study. It is stated to differ from congeners in having a black-brown ground coloration, with the thoracic shield showing only a single, large, pale spot in the anterior part, each of the subsequent terga showing a pair of light, lateral, transverse-oval spots, and the pygidium completely dark (except for the usual pale margin, which is slightly wider laterally) (Schubart 1960). It belongs to the ��� Stenopleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on page 59, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Schubart O (1960) Ein weiterer Beitrag zur Diplopoden-Fauna Marokkos. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc 40 (3): 159 - 232."]}

Eutrichodesmus filisetiger Golovatch, Geoffroy, Mauri��s & VandenSpiegel, sp. n. urn:lsid:zoobank.org:act: CC34CE29-FD03-4796-B19E-8EFD94686705 Figs 12-15 Type material: Vietnam, Thanh Hoa Prov., Thanh Son: Lang Kho Muong, Hang Doi Cave, hand collected, 13.XII.2003, leg. L. Deharveng and team (Vn0312-27), holotype ��� (MNHN JC 320), paratypes: 1 ���, 2 subad. (MNHN JC 320), 1 ��� (SEM). Vietnam, Thanh Hoa Prov., Lang Hang: Hang Bo Muoi Cave, hand collected, 14.XII.2003, leg. L. Deharveng and team (Vn0312-33), paratype ♀ (ZMUM). Name: To emphasize the peculiar, filiform but rather short tergal setae. Diagnosis: Differs from congeners by the perfect volvation, coupled with the filiform tergal setae, evident metatergal lobulations, a regular pattern of tergal tuberculation and a few minor details of gonopod structure (in particular, the shape of the telopodite and a rudimentary distofemoral process). Description: Length of adults of both sexes ca 12-13 mm, width 2.5���2.6 mm, body broadest at segment 3 or 4. Holotype ca 12 mm long and 2.5 mm wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 12). Adults with 20 segments (Figs 12, 13A), pattern of conglobation typical of ���doratodesmids���. Head, antennae (Fig. 14A), tegument (Figs 14E, F), sterna (Fig. 14C), gonopod aperture (Fig. 14G) and many other characters (Figs 14C, D) much as in E. distinctus sp. n.; collum rather irregularly tuberculate and slightly flattened medially at anterior edge (Figs 13E, 14B). Metaterga behind collum with three transverse, rather regular, mixostictic rows of rounded, flat, undifferentiated tubercles (Figs 13A-F), each crowned with a short, 2-segmented, filiform seta (often broken off) (Figs 14E, F). Only last two metaterga a little elevated compared to preceding ones (Figs 12, 13A, D). Paraterga directed ventrolaterad, nearly continuing general outline of convex dorsum, rather broad, slightly surpassing level of venter, most of them evidently lobulate (Figs 13A-F, 14C); paraterga 2 strongly enlarged, evidently lobulate only anterolaterally but not laterally (Fig. 13B). Following paraterga broadly rounded, evidently 4-lobate laterally and 3-lobate caudolaterally (Figs 13A-D, 14G). Limbus very distinctly spiculate, almost hidden by nearby abundant microvilli (Fig. 14E). Pore formula normal, ozopores very poorly visible, located dorsally near base of ventrocaudal lobulation (Figs 13C, D). Epiproct short, with regular tuberculations (Fig. 13D). Legs relatively long and slender, slightly reaching edge of paraterga (Figs 14C, G). Gonopods (Figs 14G, H, 15A-F) relatively complex. Coxae abundantly micropapillate and setose, with two conspicuous lobes: one normal, apicolateral, the other unusual, subtriangular, median, both apparently to control telopodite movements. Telopodite elongate, slender, slightly arcuate, with only a rudimentary distofemoral process at about midway (barely seen in lateral view as a short prong in a kind of pocket) and an evident hairpad at distal one-third of acropodite (= solenomere), also marking a branching point of two long, slender, apical pieces, one lateral, bifid, a little shorter and well-sclerotized, the other lamellar, pointed and dentate. Remarks: This rather large, pallid species shows filiform tergal setae and only poorly differentiated metatergal tubercles. It is also a typical ���doratodesmid���, possibly still another troglobite., Published as part of Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauri��s, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede genus Eutrichodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae), with descriptions of new species, pp. 1-46 in ZooKeys 12 (12) on pages 16-21, DOI: 10.3897/zookeys.12.167, http://zenodo.org/record/576460

Eutrichodesmus reductus Golovatch, Geoffroy, Mauriès & VandenSpiegel, sp. n. urn:lsid:zoobank.org:act: 1726A727-8FC2-4BF8-807D-8DFC1C54380C Figs 31-33 Type material: Indonesia, Sulawesi Selatan, kab. Maros: Samanggi, Gua Saripa Cave, hand collected, 18.VIII.1990, leg. A. Bedos and L. Deharveng (SULS-214), holotype ♁ (MZB), paratypes: 1 ♁ (MNHN JC 325), 1 ♀ (SEM). Name: To emphasize the strongly underdeveloped paraterga 2 and lack of metatergal tuberculation. Diagnosis: Differs from congeners except E. communicans Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2009 by the strongly underdeveloped paraterga 2, coupled with 19 body segments and the absence of metatergal tuberculation; from E. communicans by a very short tergal trichome, from it and other congeners in a few minor details of gonopod structure (in particular, the shape of the telopodite). Description: Length of adults of both sexes ca 4.0- 4.2 mm, width 0.45-0.5 mm, body broadest at midbody segments. Holotype ca 4.0 mm long and 0.45 mm wide. Coloration uniformly pallid, shown pinkish because of a photographic artifact (Fig. 31). Adults with 19 segments, body subcylindrical (Figs 31, 32A), volvation apparently incomplete due to insufficiently wide and low paraterga. Head (Fig. 33A) slightly transverse, with a poorly separated pair of very low, paramedian tubercles above antennal sockets; antennae relatively short and clavate, antennomere 6 longest; tegument (Figs 33 C-E) and many other characters (Figs 32F, 33B) much as in E. distinctus sp. n.; collum and following metaterga devoid of tuberculation, beset with numerous, irregularly arranged, extremely short setae (Fig. 32 B-F, 33C-E). Paraterga directed ventrolaterad, slightly interrupting contour of convex dorsum, short, not reaching level of venter (Figs 32 A-D); paraterga 2 (Fig. 32B) only a little enlarged compared to following ones, indistinctly lobulate both anterolaterally and laterally, with only a single noticeable lobe forming a schism ledge, both schism and hyposchism very small; paraterga 3 and 4 not narrower than others, overlap typical already from paraterga 4, not paraterga 5 as in preceding congeners. Paraterga broadly rounded caudally, very indistinctly lobulate laterally and with a single evident lobulation caudolaterally (Figs 32 C-E, 33C). Limbus distinctly denticulate, almost hidden by nearby abundant microvilli (Fig. 33D, E). Pore formula normal, ozopores located dorsally near base of caudal corner of paraterga (Fig. 33C). Legs relatively long and slender, evidently surpassing edge of paraterga (Fig. 33B). Gonopods (Fig. 33F, G) relatively complex. Coxae abundantly micropapillate, but only with a few macrosetae. Telopodite elongate, only slightly arcuate, with a large, papillate, peculiar, distofemoral outgrowth (dp) in distal one-third and a slender but short solenomere bearing a few small setae (but no pad!) subapically at base of a very complex tip represented by two erect teeth, a longer uncus and a group of minute outgrowths. Remarks: This small-bodied, pallid, non-volvatory (= “haplodesmid”, see Golovatch et al. 2009) species seems to be especially close, morphologically as well as geographically, to E. communicans from Vanuatu, Melanesia, southwestern Pacific (Golovatch et al. 2009), possibly representing still one more troglobite.

Key to the known Glomeris species of North Africa 1 Tegument entirely pallid, rarely only ocelli dark. Cavernicoles................... 2 – Head and terga distinctly pigmented, colour pattern evident, coloration mostly vivid. Epigean................................................................................. 3 2 Collum with a single transverse stria. Syncoxital horns of telopod clearly bifid (Fig. 2D, E). Libya..................................................... G. monostriata sp. n. – Collum with the usual two transverse striae. Syncoxital horns of telopod surmounted with a setoid filament (Fig. 1F). Kabylia, Algeria....................................................................................................... G. troglokabyliana sp. n. 3 Colour pattern of terga 2-11 without evident, light, paramedian spots, but with alternating light and dark transverse bands (Figs 10, 12, 24).............. 4 – Colour pattern of terga 2-(6)11 with 1+1 or 2+2 more or less evident light spots (Figs 3, 4, 7, 15-17, 20-22, 26, 27)................................................... 5 4 Coloration of caudal halves of terga distinctly marbled throughout (Figs 10, 12)........................................................................................... G. klugii – Transverse bands on terga at most only slightly marbled in contact zones (Fig. 24).......................................................................... G. mohamedanica 5 Unusually large yellowish to orange spots on tergum 6, following terga 7-11 uniformly blackish (Figs 20-22)...................................... G. carthaginiensis – Spots on tergum 6 of same size as on adjacent terga................................... 6 6 Median spots on tergum 2 much smaller than lateral ones..... G. anisosticta – Either both pairs of spots on tergum 2 comparable in size or lateral ones (if present) smaller than median ones............................................................. 7 7 Coloration uniformly blackish, lighter spots (1+1 or 2+2) very vague, marbled (Figs 15-18)................................................................... G. sublimbata – Background coloration brown to blackish, lighter spots more distinct........ 8 8 Tergum 2 with a single central light spot against a black-brown background. Pygidium uniformly black-brown. Morocco......................... G. brolemanni – Tergum 2 with 2+2 light spots................................................................... 9 9 Background coloration light to dark brown, drab, pattern as in Figs 26, 27.................................................................................................... G. punica – Background coloration dark brown to blackish, pattern vivid.................. 10 10 Light axial stripe absent (Figs 6-8).................................... G. flavomaculata – Light axial stripe present (Figs 3, 4).................................. G. colorata sp. n.

Glomeris anisosticta Brandt, 1841 Glomeris pustulata Latr., var. microstemma Brandt, 1840b: 42 (D). Glomeris pustulata Latr., var. microstemma ��� Brandt 1841b: 148 (R); Gervais 1847: 72 (R). Glomeris pustulata Latr., var. anisosticta Brandt, 1841a: 284 (D). Glomeris pustulata genuina (= pustulata), var. microstemma ��� Verhoeff 1906: 180 (R). Glomeris anisosticta ��� Brolemann 1921: 100 (L); Schubart 1953: 218 (L). Glomeris pustulata anisosticta ��� Abrous-Kherbouche & Mauri��s 1996: 586 (L). Remarks: Unfortunately, no new material of this species could be obtained for study. It is stated to be close to G. pustulata, but differs in the paramedian spots on the thoracic shield being much smaller than each lateral spot (Brandt 1840b, 1841a, b). For the time being, we treat this taxon as a full species, but its separation from G. pustulata remains to be confirmed., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on page 59, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Brandt JF (1841 a) Uber die in der Regentschaft Algier vom Herrn Dr. Wagner beobachteten Myriapoden. In: Wagner M (Ed) Reisen in der Regentschaft Algier in den Jahren 1836, 1837 und 1838. Leipzig 3, 282 - 289.","Brandt JF (1840 b) Remarques critiques sur les especes qui composent le genre Glomeris, suivies de quelques observations sur leur distribution geographique des especes en general. Bulletin scientifique publie par l'Academie Imperiale des Sciences de St. Petersbourg 7 (4 / 5): 37 - 44.","Brandt JF (1841 b) Recueil de memoires relatifs a l'ordre des Insectes Myriapodes et lus a l'Academie Imperiale des Sciences de St. - Petersbourg. St. - Petersbourg & Leipsik, 189 pp.","Gervais P (1847) Myriapodes. In: Walckenaer CA, Gervais P (Eds) Histoire naturelle des Insectes Apteres. Roret, Paris, 623 p.","Verhoeff KW (1906) Uber Diplopoden. 4. (24.) Aufsatz: Zur Kenntnis der Glomeriden (zugleich Vorlaufer eine Glomeris - Monographie). (Beitrage zur Systematik, Geographie, Entwickelung, vergleichenden Morphologie und Biologie). Archiv fur Naturgeschichte 72 (1, 2): 107 - 226.","Brolemann HW (1921) Liste des Myriapodes signales dans le nord de l'Afrique. Bulletin de la Societe des Sciences Naturelles du Maroc 1 (3 - 6): 99 - 110.","Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952).","Abrous-Kherbouche O, Mauries J-P (1996) Decouverte du genre Archipolydesmus en Algerie: description de trois especes nouvelles (Diplopoda, Polydesmida, Polydesmidae). Bulletin du Museum national d'Histoire naturelle, 4 e serie, 18 A (Zool.) (3 / 4): 571 - 587."]}

Glomeris flavomaculata Lucas, 1846 Figs 6-9 Glomeris flavomaculata Lucas, 1846: 284 (D). Glomeris flavo-maculata (sic!) ��� Gervais 1847: 74 (R); Lucas 1849: 326, plate 1, fig. 5 (R); Pocock 1892: 27 (F); Brolemann 1921: 101 (L); Schubart 1963: 80 (R). Glomeris connexa C. Koch, v. flavo-maculata (sic!) ��� Silvestri 1896: 157 (F). Glomeris flavomaculata ��� Schubart 1953: 218 (L); Abrous-Kherbouche & Mauri��s 1996: 572, 586 (F, L). Type material examined: Algeria, environs of Algiers, date?, leg. et det. H. Lucas, ��� lectotype (here designated) (MNHN CC 042), paralectotypes: 14 ���, 19 ♀, 4 ♀ juv. (MNHN CC 042). Algeria, environs of Oran, 1846, leg.?, det. H. Lucas, paralectotypes: 1 ���, 2 ♀ (MNHN CC 042). The designation of a lectotype seems advisable in order to fix the type locality and to ensure that the name-bearing type shows the diag- nostically important characteristics of the male sex, particularly as Lucas (1846) spoke about this species being common throughout Algeria. Other material: Algeria, 1850, leg. H. Lucas,? paralectotype ��� (MNHN CC 042, entry 67-96). Algeria, Vall��e des Singes, leg. & det. H. Ribaut, 2 ���, 3 ♀ (MNHN CC 042). Brief redescription: Length up to 15 mm, width up to 6.25 mm. Coloration vivid, pattern as in Figs 6-8. ��� legs 17, 18 and 19 (telopods) as in Figs 9 A-D. Remarks: Nearly the entire type series of G. flavomaculata has faded completely, probably due to the long preservation in alcohol. Fortunately, however, the paralectotypes from near Oran and the? paralectotype ��� from Lucas��� collection (1850) have retained their coloration (Figs 6, 7), which matches quite closely the pattern well depicted by Lucas (1849), based on a then rather freshly collected syntype (Fig. 8). We are certain, however, that all diplopods printed in colour by Lucas (1849) in Plate 1 are too dark and red compared to their natural coloration, not only because pertinent alcohol material shows this, but also in view of the specific name itself, flavomaculata, clearly indicating the presence of light spots on the body. The most likely explanation is that Lucas used dry material of G. flavomaculata which had become somewhat darkened (see also below). This species belongs to the ��� Stenopleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 59-61, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Lucas H (1846) Notes sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du nord de l'Afrique. Revue de Zoologie pour la Societe Cuvierienne 9: 283 - 289.","Gervais P (1847) Myriapodes. In: Walckenaer CA, Gervais P (Eds) Histoire naturelle des Insectes Apteres. Roret, Paris, 623 p.","Lucas H (1849) Histoire naturelle des animaux articules. Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842. Sciences Physiques, Zoologie 1: Crustacees, Arachnides, Myriapodes. Atlas. Imprimerie Imperiale, Paris, 322 - 390.","Pocock RI (1892) On the Myriopoda and Arachnida collected by Dr. Anderson in Algeria and Tunisia. Proceedings of the Zoological Society of London 1892: 24 - 28.","Brolemann HW (1921) Liste des Myriapodes signales dans le nord de l'Afrique. Bulletin de la Societe des Sciences Naturelles du Maroc 1 (3 - 6): 99 - 110.","Schubart O (1963) Uber einige Diplopoden aus Algier. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc 43 (2): 79 - 94.","Silvestri F (1896) Una escursione in Tunisia (Symphyla, Chilopoda, Diplopoda). Il Naturalista Siciliano, Nuova Serie 1 (8 - 12): 143 - 161.","Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952).","Abrous-Kherbouche O, Mauries J-P (1996) Decouverte du genre Archipolydesmus en Algerie: description de trois especes nouvelles (Diplopoda, Polydesmida, Polydesmidae). Bulletin du Museum national d'Histoire naturelle, 4 e serie, 18 A (Zool.) (3 / 4): 571 - 587."]}

Glomeris punica Attems, 1900 Figs 26-28 Glomeris europaea striata var. punica Attems, 1900: 302 (D). Glomeris connexa var. punica ��� Attems 1908: 105 (F). Glomeris connexa punica ��� Brolemann 1921: 100 (L)., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on page 76, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Attems C (1900) Uber die Farbung von Glomeris und Beschreibung neuer oder wenig gekannter palaearktischer Myriopoden. Archiv fur Naturgeschichte 66 (1, 3): 297 - 320.","Attems C (1908) Note sur les Myriapodes recueillis par M. Henri Gadeau de Kerville en Khroumirie et description de deux especes et d'une variete nouvelles provenant de cette region de la Tunisie. In: Gadeau de Kerville H (Ed) Voyage zoologique en Khroumirie (Tunisie). Rouen 1906, 103 - 116.","Brolemann HW (1921) Liste des Myriapodes signales dans le nord de l'Afrique. Bulletin de la Societe des Sciences Naturelles du Maroc 1 (3 - 6): 99 - 110."]}

Glomeris colorata Golovatch, Mauri��s, Akkari & Stoev, sp. n. urn:lsid:zoobank.org:act: 672AFB57-3269-4D2C-89FB-BCDAA16D3DFC Figs 3-5 Type material. Tunisia, Siliana Gov., Jebel Sidi Aouidet (47 km from Siliana, road El Fahs-Siliana), 36��15���81���N, 9��46.46���E, alt. approx. 440 m, slope overgrown by conifers close to road, under stones, leaf litter, 28.III.2008, leg. P. Stoev and N. Akkari, holotype ��� (MNHN CC 167), paratypes: 3 ♀ (MNHN CC 167), 2 ♀ (ZMUM), 1 ♀ (NMNH), 1 ♀ (ZMUC). Tunisia, Zaghouan Gov., Jebel Mansour, near Sidi Aouidet Village, 36��12���31���N, 9��45���59���E, alt. approx. 510 m. Pinus forest with Rosmarinus officinalis bushes, under stones and leaf litter, 28.III.2008, leg. N. Akkari and P. Stoev, paratypes: 6 ♀ (FMNH), 4 ���, 1 ♀ (NMNH). Name: To emphasize the colourful tergal pattern. Diagnosis: Especially similar to G. punica and G. flavomaculata, based on the characteristic colour pattern (2+2 rows of spots on terga 2-11), but differs in the presence of a light axial line, of medially coalesced 1+1 spots on the pygidium and the lighter background coloration. Description: Length of adults of both sexes (extended, alcohol material) ranging between 9.5 (���) and 10.0- 13.5 mm (♀), width between 4.7 (���) and 5.0- 6.7 mm (♀); body broadest at thoracic shield. Holotype ca 9.5 mm long and 4.7 mm wide. Coloration (Figs 3, 4) variegated, mostly rather vivid, background usually dark brown, but sometimes mostly yellow due to expanded light spots. Head mostly light brown, evidently marbled near ocelli, with 1+1 and 2+2 small pallid spots against a light brown background just above antennal sockets; labrum pale yellowish; antennae largely dark brown, only tip pallid. Collum with a large, mostly marbled, yellowish central spot (Figs 3, 4). Thoracic shield with 2+2 large light spots, sometimes interconnected with a transversely oval marbled area in-between, but separated by a more or less evident, often incomplete, light axial line. Subsequent terga, except pygidium, with 1+1 more or less wide, light paramedian spots, usually arranged into clear stripes, a mostly interrupted and sometimes vague, light, axial line, and a pair of marbled, transversely oval, lateral areas. Pygidium with 1+1 light, paramedian, mostly coalesced spots (Fig. 4), less frequently nearly entirely light grey-brown. Venter and legs contrastingly light yellow. Head usual, transverse; T��m��sv��ry���s organ transversely oval, slightly shorter than in G. troglokabyliana sp. n.; antennae long, antennomere 6 longest, 2.4-2.5 times long- er than wide; ocelli (4)5 black + 1 transparent, all convex. Collum with two transverse striae. Thoracic shield with a narrow hyposchism almost reaching caudal tergal contour (Fig. 5A); four transverse striae, of which 1-2 anteriormost starting well in front of schism and crossing entire dorsum, the other two always abbreviated (Fig. 5A); anteriormost stria only rarely slightly interrupted dorsally. Tergum 3 relatively narrowly (Fig. 5A) and tergum 4 broadly rounded laterally. Tergal pilosity and mid-dorsal sinuosity absent. Pygidium usually with a completely regularly rounded caudal margin, only rarely extremely faintly sinuated medially at margin. ��� leg 17 (Fig. 5B) with a low outer coxal lobe; telopodite 4-segmented. ��� leg 18 (Fig. 5C) with a broadly subtriangular syncoxital notch; telopodite 4-segmented. Telopods (Figs 5D) with a high, rather regularly rounded, bare, central syncoxital lobe flanked by two setose horns, latter each crowned with a minute bulb and a short setoid filament. Tarsus quite narrowly rounded apically. Remarks: Based on the narrow tergum 3, G. colorata sp. n. can be attributed to the ��� Stenopleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 56-58, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461

Glomeris carthaginiensis Schubart, 1953 Figs 20-23 Glomeris pustulata trisulcata Brolemann, 1925: 65 (D), nom. praeocc. Glomeris pustulata carthaginiensis Schubart, 1953: 222 (N), nom. n. Type material: Tunisia, Bizerte Gov., Djebel Ichkheul (near Mateur), 29.IX.1924, leg. L. Seurat, ��� lectotype of Glomeris pustulata trisulcata Brolemann, 1925 (here designated) (MNHN CC 098, Brolemann n�� 2629), paralectotypes: 1 ���, 3 ♀ (MNHN CC 098, Brolemann n�� 2629), det. H. Brolemann. The designation of a lectotype seems advisable in order to ensure that the name-bearing type shows the diagnostically important characteristics of the male sex. Other material: Tunisia, Bizerte Gov., Ichkeul National Park (plain of Mateur), inside the park, 37��08���30���N, 9��41���03���E, under stones, 7.I.2005, leg. N. Akkari, 3 ♀, 1 juv. (ZMUC). Same locality, under stones, 24.IX.2006, leg. N. Akkari, 1 ���, 1 ♀ (NMNH). Same locality, alt. approx. 50 m, under stones, 23.III.2008, leg. N. Akkari and P. Stoev, 1 ���, 10 ♀, 4 juv. (MNHN CC 098), 1 ���, 4 ♀, 2 juv. (ZMUC), 3 ♀, 2 juv. (NHMW 7780). Same locality, under stones, 8.III.2004, leg. N. Akkari, 1 ���, 7 ♀, 1 juv. (ZMUM). Same locality, under stones, 3.XII.2006, leg. N. Akkari, 3 ♀ (ZMUM). Same locality, 12.III.2009, leg. N. Akkari and H. Enghoff, 3 ���, 8 ♀ (ZMUC). Same locality, 27.III.2007, leg. F. Vilisics, 1 ��� (HNHM). Short description: Length up to 17 mm, width up to 7.5 mm. Coloration blackish, with a characteristic pattern of 2+2 light spots on thoracic shield and, occasionally, also on terga 3 and 4, but always with a pair 1+1 light and especially large spots both on tergum 6 and pygidium; no light markings on terga 7-11 (Figs 20-22). ��� legs 17, 18 and 19 (telopods) as in Figs 23 A-C. Remarks: This taxon is here considered to be a full species. The type series, said to have originally consisted of 13 specimens of both sexes (Brolemann 1925), is now incomplete (only 5 specimens left), and strongly faded, but still with a detectable colour pattern. Variation in coloration and pattern modest, the most characteristic feature being the presence of a paramedian pair of especially large, light spots on tergum 6 and a complete absence of lighter markings thereafter until the pygidium (Figs 20-22). This species belongs to the ��� Stenopleuromeris ��� type., Published as part of Golovatch, Sergei, Mauri��s, Jean-Paul, Akkari, Nesrine, Stoev, Pavel & Geoffroy, Jean-Jacques, 2009, The millipede genus Glomeris Latreille, 1802 (Diplopoda, Glomerida, Glomeridae) in North Africa, pp. 47-86 in ZooKeys 12 (12) on pages 72-75, DOI: 10.3897/zookeys.12.179, http://zenodo.org/record/576461, {"references":["Schubart O (1953) Diplopoden aus Marokko, gesammelt vom Institut Scientifique Cherifien. Bulletin de la Societe des Sciences Naturelles du Maroc 32 (1): 199 - 225 (for 1952).","Brolemann HW (1925) Deux formes nouvelles de Diplopodes tunisiens. Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord 16: 62 - 66."]}

Genus Doratodesmus Cook in Cook & Collins, 1895 Type species: Doratonotus armatus Pocock, 1894. Proposed to replace the preoccupied Doratonotus Pocock, 1894 (Cook in Cook & Collins 1895) (type locality: Java, Indonesia). = Hoplitesmus Chamberlin, 1945. Type species: Hoplitesmus enoplus Chamberlin, 1945. Synonymized by Jeekel (1955). = Pauroplus Chamberlin, 1945. Type species: Pauroplus analdes Chamberlin, 1945, n. syn. = Eucondylodesmus Miyosi, 1956. Type species: Eucondylodesmus elegans Miyosi, 1956, n. syn. = Scolopopyge Hoffman, 1978. Type species: Scolopopyge pholeter Hoffman, 1978, n. syn. = Selminarchus Hoffman, 1978. Type species: Selminarchus hispidus Hoffman, 1978, n. syn. = Crenatidorsus Zhang in Zhang & Wang, 1993. Type species: Crenatidorsus grandifoliatus Zhang in Zhang & Wang, 1993, n. syn. Diagnosis. Body ���doratodesmid���, with or without mid-dorsal projections; conglobation complete. Gonopod coxae usually abundantly setose ventrolaterally; telopodite usually stout and strongly enlarged laterally towards end of femorite, with or without a short solenomere branch thereafter; acropodite variable, from absent to well-developed., Published as part of Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauri��s, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida), pp. 1-53 in ZooKeys 7 (7) on page 45, DOI: 10.3897/zookeys.7.117, http://zenodo.org/record/576436, {"references":["Pocock RI (1894) Chilopoda, Symphyla and Diplopoda from the Malay Archipelago. In: Weber M (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien 3: 307 - 404.","Chamberlin RV (1945) On some diplopods from the Indo-Australian Archipelago. American Museum Novitates 1282: 1 - 43.","Jeekel CAW (1955) Milliped miscellany II. Entomologische Berichten 15: 412 - 417.","Miyosi Y (1956) Beitrage zur Kenntnis japanischer Myriopoden. 17. Aufsatz: Uber eine neue Gattung von Oniscodesmidae und eine neue Art von Monotarsobius. Zoological Magazine 65 (8): 311 - 314.","Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220."]}

Doratodesmus grandifoliatus (Zhang in Zhang & Wang, 1993) n. comb. Figs 27-30. Material. China, Yunnan Prov., Mengzi County, pothole no. 2 (Ma Fa Tiao Dong), 6.I.1989, leg. P. Beron, 2 ♁♁, 1 ♀ (MNHN JC 315), 1 ♁, 3 ♀♀, 9 juv. (NMNHS), 1 ♁ (ZMUM), 1 ♀ (SEM); Yunnan Prov., Mengzi County, Longbaopo Dong Cave, 27.XI.1995, leg. I. Kos, B. Sket & F. Velkovrh, 1 ♁ & 1 ♀ in copula (OBBFUL), 1 ♁ (ZMUC), 1 ♁ (SEM). Diagnosis. Differs from congeners by the simultaneous lack of mid-dorsal outgrowths on metaterga and of lateral lobulations on paraterga 2, the presence of deep crenulations at the caudal edge of most metaterga, coupled with a peculiar shape of the gonopod telopodite supplied with a large, multidenticulate, lateral lobe and a long, slightly curved, rather simple acropodite. Short redescription. Length of adults of both sexes ca 10-11 mm, width 1.8-2.1 mm, body broadest at segments 3 and 4. Coloration rather uniformly whitish to light yellow, anterior body parts often a little infuscate, light yellowish-brown. Adults with 20 segments, conglobation pattern typical of “ Doratodesmidae ”, complete (Golovatch 2003). Superficially, strongly resembling Eutrichodesmus incisus sp. n. All characters fully agreeing with the original description (Zhang and Wang 1993), but augmented here by illustrations showing more details: head (Fig. 28 C), antennae (Fig. 28 C, D), collum (Fig. 27 D), meta- and paraterga (Figs 27 A-C, E, F; 28 B), tegument and tergal setae (Figs 28 F; 29 A, B), ozopores (Fig. 28 E), telson (Fig. 27 F; 28 A), legs (Figs 29 C, D; 30 A). Sterna usually with a rather deep, narrow depression between coxae (Fig. 28 B), only those between coxae 6, 7 and 9 much wider (Fig. 30 A). Gonopod aperture transverseoval, relatively small, far from reaching lateral sides of segment 7. Legs rather long; femoral and tarsal segments equal and longest; several basal segments microtuberculate; claw simple, slightly curved ventrad; some setae sparsely microdentate (Figs 29 C, D; 30 A). Gonopods (Figs 30 B, C) relatively complex. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally. Telopodite considerably longer than coxite, subunciform, basal half slender and setose, at about midway conspicuously enlarged, with a multidenticulate lateral lobe (lo) and a very short solenomere (sl) with a hairpad on top; acropodite relatively long and simple. Remarks. This species has hitherto been described or reported from three caves in Yunnan, China (Wang and Zhang 1993; Golovatch 2003). One of the new samples (from Longbaopo Cave) is strictly topotypic, while Ma Fa Tiao Dong Cave is known to support another millipede, Glyphiulus subgranulatus Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2007 (see Golovatch et al. 2007a).

Eutrichodesmus armatocaudatus Golovatch, Geoffroy, Mauri��s & VandenSpiegel, sp. n. urn:lsid:zoobank.org:act: B2718568-A85A-4FDE-80E9-574F67AB90E9 Figs 5-8. Type material. Vietnam, Thanh Hoa Prov., Pu Luong, Lung Cao, Hang Lang Lua Cave, 11.XII.2003, leg. L. Deharveng & team (Vn 0312-05), holotype ��� (MNHN JC 310), paratypes: 1 ���, 1 juv. (MNHN JC 310), 1 ��� (ZMUM); Ha Nam Ninh Prov., Cuc Phuong National Park, Cave of Prehistoric Man, 10.X.1998, leg. L. Deharveng (VIET-537), 1 ��� (MNHN JC 310); same locality, Water Fairy Cave, 11.X.1998, leg. L. Deharveng (VIET-541), 1 ���, 2 juv. (MNHN JC 310); same locality, cave 4 (Son Cung Cave?), 21.VI.2003, leg. P. Trontelj & B. Sket, 1 ���, 1 ♀, 3 juv. (MNHN JC 310), 1 ���, 1 juv. (ZMUC), 1 ���, 1 juv. (NMNHS), 1 ��� (MCSNV), 1 ���, 1 juv. (OBB- FUL), 1 ♀ (SEM); Ha Nam Ninh Prov., outside Cuc Phuong National Park, cave 3, 20.VI.2003, leg. P. Trontelj & B. Sket, 1 ���, 1 ♀ (ZMUM). Name. To emphasize the obvious dorsal projections on body segments 17-19. Diagnosis. Most similar to Parapauroplus monodentus Zhang in Zhang & Wang, 1993, which also shows evident mid-dorsal projections only on body segments (16)17- 19, but differs by the presence of porosteles, the slightly lower metatergal tubercles devoid of microsetae near each of the main setae, and the slightly stouter gonopod telopodite showing a shorter midway process. Description. Length of adults of both sexes ca 10-14 mm, width 2.3-2.6 mm, body broadest at segment 3 or 4; ������ usually a little smaller than ♀♀. Holotype ca 11 mm long and 2.5 mm wide. Coloration uniformly pallid or light yellowish. Adults with 20 segments, pattern of conglobation typical of ��� Doratodesmidae ��� (Figs 5 A). Head (Fig. 6 A) and tegument (Figs 5 B, C; 6 F, G) as in preceding species, but antennae somewhat longer (Fig. 6 A), collum slightly flattened mid-dorsally, not covering the head from above (Fig. 6 A). Metaterga behind collum with three transverse, rather irregular rows of tubercles (Figs 5 A-C), most of which still retain short, bisegmented setae (Fig. 6 G), metaterga 17-19 each with an evident dorsal outgrowth (Fig. 5 C). Paraterga mostly directed ventrolaterad, rather broad, slightly surpassing level of venter, clearly trisinuate caudolaterally at base, due to two lobulations (Figs 5 B, D; 6 B); paraterga 2 strongly enlarged, subvertical, margin nearly entire, with only very faint traces of a series of small lobulations anterolaterally (Fig. 6 A), both schism and hyposchism very small; paraterga 3 and 4 slightly shorter than others, overlap of following paraterga typical, latter broadly rounded and slightly 3- or 4-lobate. Limbus spiculate (Fig. 6 F). Pore formula normal, ozopores located on top of a small porostele, slightly above ventrocaudal lobulation (Figs 6 C-E). Pleurotergal carinae wanting. Epiproct elongated, also with tubercles, directed ventrocaudad, with the usual four cones just below tip; para- and hypoprocts as in Figs 5E, F. Sterna usually with a deep, narrow depression between coxae (Figs 5 D; 6 B), only sterna between coxae 6, 7 and 9 much wider (Fig. 7 A). Gonopod aperture transverseoval, relatively small, far from reaching lateral sides of segment 7. Legs relatively long; femoral and tarsal segments longest and equal; claw simple, very slightly curved ventrad; some setae very sparsely microdenticulate (Figs 5 D; 6 B). Gonopods (Figs 7; 8) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous rounded lobe frontolaterally. Telopodite considerably longer than coxite, slightly stouter, setose in its basal half, with an inconspicuous, digitiform, simple, lateral, distofemoral process (dp) at about midway, more distally with a slightly folded and enlarged shaft (= solenomere) with dense pilosity and a hairpad; seminal groove terminating subapically, acropodite short. Remarks. This species seems to be close to some of the ���doratodesmids��� showing evident mid-dorsal projections on some of the metaterga, e.g. Ascetophacus macclurei Hoffman, 1977, Cerastelachys cavernicola (Sinclair, 1901), Pocillidorsus dorsiangulatus Zhang in Zhang & Wang, 1993 and Parapauroplus monodentus Zhang in Zhang & Wang, 1993, etc. (Hoffman 1977a, 1977b; Zhang and Wang 1993). Cave 4 also supports another millipede, Glyphiulus mediobliteratus Golovatch, Geoffroy, Mauri��s & VandenSpiegel, 2007 (Golovatch et al. 2007b)., Published as part of Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauri��s, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida), pp. 1-53 in ZooKeys 7 (7) on pages 13-16, DOI: 10.3897/zookeys.7.117, http://zenodo.org/record/576436, {"references":["Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220.","Sinclair FG (1901) On the myriapods collected during the \" Skeat Expedition \" to the Malay Peninsula, 1899 - 1900. Proceedings of the Zoological Society of London 2: 505 - 533.","Hoffman RL (1977 a) The systematic position of the diplopod family Doratodesmidae, and description of a new genus from Malaya (Polydesmida). Pacific Insects 17 (2 - 3): 699 - 719.","Hoffman RL (1977 b) Diplopoda from Malayan caves collected by M. Pierre Strinati. Revue suisse de Zoologie 84 (3): 699 - 719.","Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2007 b) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda: Spirostreptida: Cambalopsidae). Part 2. The javanicus - group. Zoosystema 29 (3): 417 - 456."]}

Doratodesmus grandifoliatus (Zhang in Zhang & Wang, 1993) n. comb. Figs 27-30. Material. China, Yunnan Prov., Mengzi County, pothole no. 2 (Ma Fa Tiao Dong), 6.I.1989, leg. P. Beron, 2 ������, 1 ♀ (MNHN JC 315), 1 ���, 3 ♀♀, 9 juv. (NMNHS), 1 ��� (ZMUM), 1 ♀ (SEM); Yunnan Prov., Mengzi County, Longbaopo Dong Cave, 27.XI.1995, leg. I. Kos, B. Sket & F. Velkovrh, 1 ��� & 1 ♀ in copula (OBBFUL), 1 ��� (ZMUC), 1 ��� (SEM). Diagnosis. Differs from congeners by the simultaneous lack of mid-dorsal outgrowths on metaterga and of lateral lobulations on paraterga 2, the presence of deep crenulations at the caudal edge of most metaterga, coupled with a peculiar shape of the gonopod telopodite supplied with a large, multidenticulate, lateral lobe and a long, slightly curved, rather simple acropodite. Short redescription. Length of adults of both sexes ca 10-11 mm, width 1.8-2.1 mm, body broadest at segments 3 and 4. Coloration rather uniformly whitish to light yellow, anterior body parts often a little infuscate, light yellowish-brown. Adults with 20 segments, conglobation pattern typical of ��� Doratodesmidae ���, complete (Golovatch 2003). Superficially, strongly resembling Eutrichodesmus incisus sp. n. All characters fully agreeing with the original description (Zhang and Wang 1993), but augmented here by illustrations showing more details: head (Fig. 28 C), antennae (Fig. 28 C, D), collum (Fig. 27 D), meta- and paraterga (Figs 27 A-C, E, F; 28 B), tegument and tergal setae (Figs 28 F; 29 A, B), ozopores (Fig. 28 E), telson (Fig. 27 F; 28 A), legs (Figs 29 C, D; 30 A). Sterna usually with a rather deep, narrow depression between coxae (Fig. 28 B), only those between coxae 6, 7 and 9 much wider (Fig. 30 A). Gonopod aperture transverseoval, relatively small, far from reaching lateral sides of segment 7. Legs rather long; femoral and tarsal segments equal and longest; several basal segments microtuberculate; claw simple, slightly curved ventrad; some setae sparsely microdentate (Figs 29 C, D; 30 A). Gonopods (Figs 30 B, C) relatively complex. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally. Telopodite considerably longer than coxite, subunciform, basal half slender and setose, at about midway conspicuously enlarged, with a multidenticulate lateral lobe (lo) and a very short solenomere (sl) with a hairpad on top; acropodite relatively long and simple. Remarks. This species has hitherto been described or reported from three caves in Yunnan, China (Wang and Zhang 1993; Golovatch 2003). One of the new samples (from Longbaopo Cave) is strictly topotypic, while Ma Fa Tiao Dong Cave is known to support another millipede, Glyphiulus subgranulatus Golovatch, Geoffroy, Mauri��s & VandenSpiegel, 2007 (see Golovatch et al. 2007a)., Published as part of Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauri��s, Jean-Paul & VandenSpiegel, Didier, 2009, Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida), pp. 1-53 in ZooKeys 7 (7) on pages 36-37, DOI: 10.3897/zookeys.7.117, http://zenodo.org/record/576436, {"references":["Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220.","Golovatch SI (2003) A review of the volvatory Polydesmida, with special reference to the patterns of volvation (Diplopoda). African Invertebrates 44 (1): 39 - 60.","Golovatch SI, Geoffroy J-J, Mauries J-P, VandenSpiegel D (2007 b) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda: Spirostreptida: Cambalopsidae). Part 2. The javanicus - group. Zoosystema 29 (3): 417 - 456.","Golovatch SI, Geoffroy J-J, Mauries, J-P, VandenSpiegel D (2007 a) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda: Spirostreptida: Cambalopsidae). Part 1. The granulatus - group. Zoosystema 29 (1): 7 - 49."]}