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Alpine glaciers are sensitive indicators of changes in climate, and their ubiquity in mountainous regions make them valuable proxies for terrestrial climate reconstructions worldwide. However, the timing and extent of glacier change across the South American mid-latitudes through the Holocene are still poorly constrained relative to their counterparts in the Northern Hemisphere. Here we report a new 10Be surface exposure-based chronology of moraines recording a series of progressively less-extensive glacier advances of Glaciar Torre (Argentina, 49.3°S/73.0°W) since the Last Glacial Maximum, with expansions culminating at 17,600 ± 900, 13,500 ± 500, 9700 ± 400, 6900 ± 200, 6100 ± 300, 4500 ± 200, and 530 ± 60 yr BP. The declining magnitude of Holocene glacier expansions parallels a gradual rise in local summer insolation intensity during the Holocene, while individual advances occurred during inferred negative Southern Annular Mode (SAM)-like states at centennial to millennial timescales. These observations suggest that (i) summer insolation intensity modulated antiphased trends in glacier extent in the polar hemispheres during the Holocene, and that (ii) centennial-scale ‘SAM-like’ temperature and precipitation anomalies paced glacier fluctuations throughout Patagonia. Given the persistence of the inferred ’SAM-like’ anomalies throughout the Holocene, the modern measured trend towards positive SAM index conditions could mark the onset of a fundamental shift in the climate of the Southern Hemisphere midlatitudes that warrants consideration in projections of future climate.

The history and dynamics of deciduous Nothofagus forests along the eastern slopes of the central Patagonian Andes (44°-49°S) remain insufficiently studied and understood, particularly at timescales ranging from centuries to millennia. Available fossil pollen records point to time-transgressive responses of the arboreal vegetation to climatic changes during the Last Glacial Termination (T1) and early Holocene, and spatial heterogeneity since then along north-south, east-west, and elevation transects. The degree to which these results represent biogeographic and climatic trends, varying environmental gradients, or site-specific phenomena has not been assessed in detail. Here we present a fossil pollen and macroscopic charcoal record from Lago Churrasco (45°41′S, 71°49′W), a small closed-basin lake located in the deciduous Nothofagus forest zone of the central-east Andes of Chilean Patagonia. Our results suggest that Nothofagus trees colonized newly deglaciated terrains at ∼16,000 cal yr BP and formed scrublands/woodlands several millennia earlier than reported by previous studies east of the Andes. This suggests expansion and local densification of tree populations sourced from the eastern margin of the Patagonian Ice Sheet during the Last Glacial Maximum, with the additional implication that temperature and precipitation conditions favorable for tree survival and reproduction developed early during T1. We posit that the amount of moisture delivered by the Southern Westerly Winds was not a limiting factor for arboreal expansion during T1 in this sector of the central Patagonian Andes. Closed-canopy Nothofagus forests established at ∼10,000 cal yr BP and have remained essentially invariant despite climate change and natural disturbance regimes. This resilience was challenged and exceeded by human disturbance during the 20th century through the use of fire, leading to deforestation and spread of invasive exotic species in an extraordinarily rapid event. Our record suggests a permanent influence of the Southern Westerly Winds over the last 10,000 years, with relatively modest variations at centennial and millennial timescales. Fil: Moreno, P. I.. Universidad de Chile; Chile Fil: Simi, E.. Universidad de Chile; Chile Fil: Villa Martínez, R. P.. Universidad de Magallanes; Chile Fil: Vilanova, Isabel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales "Bernardino Rivadavia"; Argentina

This paper reviews current understanding of deglaciation in North, Central and South America from the Last Glacial Maximum to the beginning of the Holocene. Together with paleoclimatic and paleoceanographic data, we compare and contrast the pace of deglaciation and the response of glaciers to major climate events. During the Global Last Glacial Maximum (GLGM, 26.5-19 ka), average temperatures decreased 4° to 8°C in the Americas, but precipitation varied strongly throughout this large region. Many glaciers in North and Central America achieved their maximum extent during the GLGM, whereas others advanced even farther during the subsequent Heinrich Stadial 1 (HS-1). Glaciers in the Andes also expanded during the GLGM, but that advance was not the largest, except on Tierra del Fuego. HS-1 (17.5-14.6 ka) was a time of general glacier thickening and advance throughout most of North and Central America, and in the tropical Andes; however, glaciers in the temperate and subpolar Andes thinned and retreated during this period. During the Bølling-Allerød interstadial (B-A, 14.6-12.9 ka), glaciers retreated throughout North and Central America and, in some cases, completely disappeared. Many glaciers advanced during the Antarctic Cold Reversal (ACR, 14.6-12.9 ka) in the tropical Andes and Patagonia. There were small advances of glaciers in North America, Central America and in northern South America (Venezuela) during the Younger Dryas (12.9-11.7 ka), but glaciers in central and southern South America retreated during this period, except on the Altiplano where advances were driven by an increase in precipitation. Taken together, we suggest that there was a climate compensation effect, or ‘seesaw’, between the hemispheres, which affected not only marine currents and atmospheric circulation, but also the behavior of glaciers. This seesaw is consistent with the opposing behavior of many glaciers in the Northern and Southern Hemispheres.

We present a high-resolution precisely dated terrestrial paleovegetation/paleoclimate record from Lago Pichilaguna, northwestern Patagonia (40°–44°S), which spans continuously from the Last Glacial Maximum (LGM) to the present. We find abundant and continuous presence of arboreal pollen (chiefly Nothofagus) during the LGM, accompanied by other trees, shrubs, and alpine herbs. These results suggest Subantarctic parkland and/or scattered woodlands under a cold and hyperhumid climate during the LGM (∼25,000–17,800 cal. yr BP) with expansion of Nothofagus under relatively warm interstadial conditions between 25,000 and 19,200 cal. yr BP. This was followed by cooling and a precipitation increase between 19,200 and 17,800 cal. yr BP, which was contemporaneous with the youngest LGM advance of Andean glaciers in the region and maximum influence of the Southern Westerly Wind (SWW). The Last Glacial Termination (T1) started at 17,800 cal. yr BP and featured the spread of thermophilous trees and ferns characteristic of North Patagonian rainforests, along with lake level lowering. These results suggest a warm pulse and southward shift of the SWW, concurrent with a rapid collapse of Andean glacier lobes. Subsequent changes led to the establishment of closed-canopy rainforests under peak interstadial warmth between ∼16,000 and 15,000 cal. yr BP. We detect a shift to cold/wet conditions during the Antarctic Cold Reversal (14,800–12,700 cal. yr BP) and a precipitation decline during Younger Dryas time, followed by maximum temperature, relatively lower lake level and minimum SWW influence between 11,300 and 7700 cal. yr BP. Precipitation then rose punctuated by centennial-scale variations since 6200 cal. yr BP. Chilean-European deforestation and spread of invasive exotic species started at ∼350 cal. yr BP aided by fire. We conclude that temperate rainforests have persisted with little interruption since T1, with major changes in floristic composition driven by climate change and fires. Rainforest composition and heterogeneity declined in response to Chilean/European disturbance during the 1600s and intensified since the 1800s. These events constitute the fastest/largest-magnitude vegetation changes of the last ∼25,000 years.

South Westland, New Zealand, and southern Chile, are two narrow continental corridors effectively confined between the Pacific Ocean in the west and high mountain ranges in the east which impart significant influence over regional climate, vegetation and soils. In both these southern mid-latitude regions, evidence for extensive and repeated glaciations during cold phases of the Quaternary is manifested by arrays of successively older glacial drift deposits with corresponding outwash plain remnants. In South Westland, these variably aged glacial landforms are mantled by layered (multisequal) soils characterised by slow loess accretion and pedogenesis in an extreme leaching and weathering environment. These cover-bed successions have undergone repeated coupled phases of topdown and upbuilding soil formation that have been related to fluctuating cycles of interglacial/warm and glacial/cold climate during the Quaternary. In this study, we recognise multisequal soils overlying glacial landforms in southern continental Chile but, unlike the spodic (podzolic) soil sequences of South Westland, these are of dominantly volcanigenic (andic) provenance and are very similar to multisequal soils of andic provenance that predominate in, and adjacent to, areas of rhyolitic to andesitic volcanism in North Island, New Zealand. Here we develop a soil-stratigraphic model to explain the observed occurrence of multisequal soils mantling dominantly glacial landforms of southern continental Chile. Based on proxy data from southern Chile, we propose that persistent vegetation cover and high precipitation on the western side of the Andes, during colder-than-present episodes tended to suppress the widespread production of glacially-derived loessial materials despite the pervasive occurrence of glacial and glacio-fluvial deposits that have frequently inundated large tracts of this landscape during the Quaternary. Given the lack of loess cover-beds that have traditionally assisted in the relative dating of glacial episodes prior to the Late Quaternary, surface exposure dating techniques could provide another chronological alternative to address this issue. However, there have been two main obstacles to successfully apply this dating technique in Patagonia. First, minimum exposure ages may be obtained on moraines older than the last glacial cycle due to erosion, although dating outwash plains is more robust. Second, on the wet western side adjacent to the Andes, persistent vegetation cover during both glacial and post-glacial times, as well as widespread inundation by volcanic mass-flows, appear preventive. We make a case that soil genesis within this region appears to be dominated by a constant flux of intermittently erupted Andean-sourced tephra which has continued to upbuild soils at the ground surface separated by intervals where topdown weathering processes are intensified. As already demonstrated by New Zealand studies, multisequal soil successions have a clear implied connection to coupled glacial and interglacial climate cycles of the Quaternary. On this basis, similar sequences in northwest Patagonia provide a relatively untapped archive to enable Quaternary glacial and environmental changes in this pervasively glaciated volcanic region to be constructed.

Iniciativa Cientifica Milenio NC120066 FONDECYT 11121280 1160488 1151469 CONICYT USA 2013-0035 NSF-BCS 1263474 Fulbright

We describe the stratigraphy, age and correlation of a prominent tephra marker, named Lepue Tephra, extensively distributed in north-western Patagonia. Lepue Tephra is well dated at c. 11 000 cal a BP from numerous lake and soil cover-bed sequences and its recognition is useful for assessing the rate and timing of deglaciation as well as associated environmental changes in this region during the last glacial termination and early Holocene. Lepue Tephra has attributes typical of a complex and compositionally zoned phreatomagmatic eruptive. While the initial rhyolitic phase can be readily distinguished from multiple eruptive products sourced from the adjacent Volcan Chaiten, the main erupted end member is of basaltic–andesitic bulk composition − similar to younger tephras sourced from Holocene monogenetic cones adjacent to the Volcan Michimahuida massif (tMim). Lepue Tephra can be correlated to an equivalent-aged pyroclastic flow deposit (Amarillo Ignimbrite) prominently distributed in the south-eastern sector of tMim. The source vent for these co-eruptive events is obscured by an extensive ice field and is currently unknown. The widespread radially symmetrical distribution of Lepue Tephra centred on tMim cannot be attributed solely to volcanological considerations. Reduced Southern Hemisphere westerly wind influence interpreted from climate proxies at the time of eruption are also implicated.

We examine the response of Nothofagus forests to climate change and disturbance regimes over the last 3200 years near Coyhaique (45°S), central–east Chilean Patagonia, using fine-resolution pollen and charcoal records from lake sediment cores. Closed-canopy deciduous Nothofagus forests have dominated the region with little variation until the arrival of Chilean–European settlers, suggesting a predominance of cool-temperate and wet conditions. Within this state we identify centennial-scale episodes of forest fragmentation, increase in littoral macrophytes and volcanic/paleofire disturbance between 2700 and 3000 cal a BP, 2200 and 2500 cal a BP and over the last ∼250 years, which we interpret as intervals with negative hydrologic balance. Natural variability caused little impact on the physiognomy and composition of the vegetation in pre-European time, in contrast to the accelerated shift that started during the late 19th century associated with deforestation, homogenization and synchronization of ecosystem changes at the landscape level, and spread of exotic plant species brought by Chilean and European settlers during a warm/dry interval. The resilience of deciduous Nothofagus forests to natural disturbance regimes and climate change was exceeded by large-scale human disturbance since the late 19th century by fire, timber exploitation and livestock grazing. These disturbances caused an ecosystem shift towards artificial meadows and scrublands with frequent high-magnitude fires.

Southern South America is the only continental landmass that intersects the core of the Southern Westerly Winds (SWW), and thus is important for studying their role as a driver/conduit for the initiation/propagation of climate signals since the last glaciation. Their interaction with the Southern Ocean (SO) affects global climate through its influence on high-latitude upwelling and biological productivity, deep-water convection sites and, consequently, ventilation of CO2 from the deep ocean. Variations in the SWW-SO coupled system have been postulated as fundamental drivers of climate change during glacial terminations and the current interglacial. Hence, deciphering the evolution of the SWW from sensitive locations in the southern middle latitudes is essential for understanding important climatic transitions during and since the Last Glacial Termination (T1). Terrestrial records from the central Patagonian Andes (CPA) (44°-49°S), however, show heterogeneities in the timing, rates, and direction of climate change during T1, impeding detailed assessment of its drivers at regional, hemispheric, and global scales. Here we present new data on glacier, vegetation, and fire-regime changes in the Coyhaique sector (45°34′S) of CPA to improve our understanding on the timing and structure of the T1, including the behavior of the SWW. Our results indicate glacial recession from the youngest Last Glacial Maximum moraines just before ∼17.9 ka and development of an ice-dammed proglacial lake during the early stages of T1. Drainage of the ice-dammed lake, triggered by renewed glacial recession, was near-synchronous with the onset of a gradual multi-millennial trend toward arboreal dominance that started at ∼16 ka east and west of the Andes at that latitude. We detect increased influence of the SWW at ∼45°S starting at ∼16.6 ka, relative to the first millennium of T1, that led to positive anomalies in precipitation between ∼16–14.4 and ∼12.8–11.5 ka, followed by negative anomalies between ∼11.5–9 ka. The synchronous spread of arboreal vegetation east and west of the CPA divide during T1, despite the trans-Andean precipitation contrasts, suggests an upward shift in the temperature-controlled Andean tree line, underscoring the role of deglacial warming as the critical driver for afforestation at regional scale. Fil: Vilanova, Isabel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales "Bernardino Rivadavia"; Argentina Fil: Moreno, P. I.. Universidad de Chile; Chile Fil: Miranda, C. G.. Universidad de Chile; Chile Fil: Villa Martínez, R. P.. Universidad de Magallanes; Chile

The deep-time dynamics of coupled socio-ecological systems at different spatial scales is viewed as a key framework to understand trends and mechanisms that have led to the Anthropocene. By integrating archeological and paleoenvironmental records, we test the hypothesis that Chilean societies progressively escalated their capacity to shape national biophysical systems as socio-cultural complexity and pressures on natural resources increased over the last three millennia. We demonstrate that Pre-Columbian societies intentionally transformed Chile’s northern and central regions by continuously adjusting socio-cultural practices and/or incorporating technologies that guaranteed resource access and social wealth. The fact that past human activities led to cumulative impacts on diverse biophysical processes, not only contradicts the notion of pristine pre-Industrial Revolution landscapes, but suggests that the Anthropocene derives from long-term processes that have operated uninterruptedly since Pre-Columbian times. Moreover, our synthesis suggests that most of present-day symptoms that describe the Anthropocene are rooted in pre-Columbian processes that scaled up in intensity over the last 3000 years, accelerating after the Spanish colonization and, more intensely, in recent decades. The most striking trend is the observed coevolution between the intensity of metallurgy and heavy-metal anthropogenic emissions. This entails that the Anthropocene cannot be viewed as a universal imprint of human actions that has arisen as an exclusive consequence of modern industrial societies. In the Chilean case, this phenomenon is intrinsically tied to historically and geographically diverse configurations in society-environment feedback relationships. Taken collectively with other case studies, the patterns revealed here could contribute to the discussion about how the Anthropocene is defined globally, in terms of chronology, stratigraphic markers and attributes. Furthermore, this deep-time narrative can potentially become a science-based instrument to shape better-informed discourses about the socio-environmental history in Chile. More importantly, however, this research provides crucial “baselines” to delineate safe operating spaces for future socio-ecological systems.

Robust glacial chronologies are needed to address the fundamental questions of when and why Ice Age climates begin and end. Well-preserved glacial deposits left by large ice sheet lobes adjacent to Estrecho de Magallanes (53°S; Chile) in southernmost South America provide a unique opportunity to reconstruct the timing and fine structure of the last two major glaciations, as well as the last termination. We present a new precise 10Be surface exposure dataset of 34 moraine boulders directly tied to a recently published, high resolution glacial geomorphic map of the deposits left by the Magallanes lobe. We find that the southern section of the Patagonian Ice Sheet was more extensive during Marine Isotope Stage 4 (MIS 4) than during MIS 2, representing the first direct dating of the MIS 4 glacier culmination in South America. Similar to the MIS 2 glacial maxima, within MIS 4 there were multiple advances that we date (n = 6 samples) to between 67.5 ± 2.1 and 62.1 ± 2.0 ka. A similarly timed MIS 4 advance was identified in New Zealand, indicating that this is a hemisphere-wide glacier-climate signal, which is further corroborated by South Atlantic and Pacific temperature proxy records. Inboard of the MIS 4 moraine complex, we date a sequence of geomorphically distinct MIS 2 moraines that represent separate major periods of glacial stability. The MIS 2 maximum extent occurred by 27.4 ± 0.8 ka (n = 4; arithmetic mean, with the standard error of the mean and 3% propagated production rate error) and was followed by at least four more full glacial culminations at 25.7 ± 0.8 (n = 3), 23.9 ± 0.8 (n = 5), 19.1 ± 0.7 (n = 3), and 18.1 ± 0.6 ka (n = 3), which represent periods when the glacier was in equilibrium with the climate for long enough to form major moraines. About 18 km inboard, this sequence is followed by smaller-scale recessional moraine crests, deposited on drumlinized terrain rather than a moraine drift, that we date to 18.0 ± 0.8 ka, indicating the glacier was in net retreat at this time. Tentative results from a 2D ice sheet model suggest that the Magallanes lobe may have reached mapped inner and outer MIS 2 moraines from a climate with approximately 4.5 °C and 5.5 °C cooler summers, respectively, assuming ∼25% less annual precipitation, relative to modern climate. We hypothesize that during the last glacial cycle, shifts in the subtropical and subantarctic fronts, and related ocean-atmosphere patterns, explain MIS 4 to 2 glacial behavior in the southern mid-latitudes.

We present results from Lago Negro, a small closed-basin lake adjacent to Lago Rosselot, to examine the vegetation and environmental history of an insufficiently studied sector of Chiloe Continental (41°30′-44°S) in northwestern Patagonia. Lake sediment cores from Lago Negro reveal 27 tephra deposited since ∼12.7 ka, including two prominent rhyodacite tephra marker beds erupted from Volcan Melimoyu, and a stratified basal clastic unit we attribute to meltwater discharge from an ice tongue that originated from Monte Queulat and covered Lago Rosselot during its expanded position, presumably Antarctic Cold Reversal in age. The pollen record shows closed-canopy North Patagonian rainforests since ∼12.7 ka, with variations in species composition and structure that suggest dynamic responses of the vegetation to past environmental changes. Vegetation responses to climate in the Lago Negro record were modulated, sometimes interrupted, by high magnitude and frequent disturbance regimes, most notably during maxima in explosive volcanic activity (∼9.5–7.2 ka and ∼3.6–1.6 ka) and heightened fire activity. Since Lago Negro is the southernmost palynological site so far investigated in the region and is located within a volcanically active sector, it provides a valuable perspective for assessing past vegetation responses along environmental gradients since the last glaciation. When compared with other sites throughout northwestern Patagonia, our record reveals a distinct north-to-south gradient in temperature and precipitation, with peak temperature and rainfall seasonality in the north, and a west-to-east gradient in disturbance regimes, with maximum frequency and magnitude of explosive volcanic events in the east. These gradients have modulated the response of rainforest vegetation to climate forcing at regional scale since ∼12.7 ka. We identify negligible differences in timing for the majority of key vegetation signals during the initial phase of the Lago Negro record, and propose that plant colonization and expansion along the ∼360 km long corridor through the Pacific slope of the northwestern Patagonian Andes was a rapid process during the Last Glacial Termination.

Climate and disturbance regimes play key roles in shaping the structure, composition and functioning of terrestrial ecosystems. Despite this importance, very few stratigraphic studies in the temperate rainforests from northwestern Patagonia have explored this relationship in detail along a time continuum through the entire Holocene. Here we present a high-resolution fossil pollen and charcoal record from Lago Fonk (median resolution: 20 years), a small closed-basin lake in the lowlands of the Chilean Lake District (41°S), where wildfires and explosive volcanism have intermittently taken place during the Holocene, along with pronounced human-induced disturbance in post-colonial time. Our results show persistence of temperate rainforest throughout the Holocene, with changes in the composition and structure of Valdivian rainforests (VRF) at millennial timescales. We detect centennial-scale alternations in dominance between the VRF tree Eucryphia/Caldcluvia and generalist trees found in VRF and North Patagonian rainforests after ∼6.5 cal ka BP. Intervals dominated by VRF coincide with enhanced fire occurrence signaling negative hydroclimate anomalies with a mean duration of ∼150 years, which alternate with positive hydroclimate anomalies lasting ∼312 years on average. Our results suggest that the magnitude and rapidity of vegetation changes detected at 10.2–9.9, 4.0–3.0, ∼1.0, and ∼0.7 cal ka BP were amplified by disturbance regimes, and led to the establishment and maintenance of Eucryphia/Caldcluvia-dominated forests in the Longitudinal Valley of the Chilean Lake District. On several occasions the higher incidence of fire disturbance during warm/dry climate intervals coincided with episodes of heightened explosive volcanic activity from multiple eruptive centers within the Southern Andean Volcanic Zone.

An important coupled ocean-atmospheric system in the mid- and high latitudes involves the Southern Westerly Winds (SWW) and the Southern Ocean (SO), which controls climate in the southernmost third of the world, deep water formation, and ventilation of CO2 from the deep ocean. Most studies have examined its role as a driver of atmospheric CO2 concentrations during glacial terminations, but very few have investigated its influence during the Holocene, i.e. the current interglacial. A fundamental problem, however, is resolving whether the SWW strength increased or declined during the early Holocene (∼11.5–7.5 ka, ka = 1000 cal yr BP) in sectors adjacent to the Drake Passage. Here we assess past changes in SWW influence over the last ∼17,000 years using terrestrial paleoclimate records from southwestern Patagonia (∼52°S). We detect a zonally symmetric Early Holocene Westerly Minimum which diminished wind stress and upwelling on the SO, contributing to a contemporary decline in atmospheric CO2 concentrations and enrichment in the stable carbon isotope ratio of atmospheric CO2 (δ13Catm). Our mid-latitude data also indicate a shift to strong SWW influence at ∼7.5 ka which correlates with a sustained increase in atmospheric CO2 and halt in the δ13Catm rise, suggesting enhancement of high-latitude ocean ventilation by an invigorated SWW-SO coupled system.

We examine hydroclimate changes at centennial/millennial timescales since 16,000 yr BP in northwestern Patagonia based on the pollen and charcoal record from Lago El Salto, a small closed-basin lake located in the Chilean Lake District (41°38′48.02″S, 73° 5′48.42″W). We observe cold/wet conditions between 14,500–16,000 yr BP, followed by further cooling with increased precipitation until 13,000 yr BP, enhanced precipitation seasonality and/or variability between 11,600–13,000 yr BP, and an extended warm-and-dry interval between 7600 and 11,300 yr BP with peak paleofire activity. Colder-and-wetter than present conditions and muted paleofire activity prevail between 5300 and 7600 yr BP, followed by alternating cold/wet and centennial-scale warm/dry phases starting at 5300 yr BP with three conspicuous megadroughts since 2500 yr BP. The most recent megadrought occurred during the Medieval Climate Anomaly. We identify a cold reversal that spans the Antarctic Cold Reversal (ACR) and the Younger Dryas (YD) chrons with stronger-than-present westerly influence during the former and enhanced variability during the latter. These results extend the northern limit of strong cooling and increase in precipitation during the ACR and the southern limit of influence of strong hydrologic variations during the YD in terrestrial environments, suggesting an overlap in the spheres of influence of processes originating from southern and northern polar latitudes. An extended warm southern westerly wind (SWW)-minimum interval is evident between 7600 and 11,300 yr BP, followed by a rapid shift to cool-moist conditions between 5300 and 7600 yr BP brought by a mid-Holocene SWW maximum. Since then we observe centennial-scale hydroclimate variability, which has driven biodiversity and fire-regime shifts of evergreen temperate rainforests.

By virtue of its location in the southern mid-latitudes, northwestern Patagonia (40°-44°S) offers the opportunity to unravel the mechanisms involved in the initiation and propagation of paleoclimate signals of hemispheric and global significance. Of particular importance is deciphering the evolution of the Southern Westerly Winds (SWW) considering their influence at continental, zonal, hemispheric, and global scale. Here I present a multi-decadal record from Lago Proschle, a small closed-basin lake located on the Pacific slope of northwestern Patagonia, to examine the timing and structure of vegetation, fire, and climate change along a continuum since the onset of the Last Glacial Termination (T1). The record shows rapid glacier recession during T1, reaching the western Andean foothills in ∼400 years or less. Nothofagus-dominated forests established between ∼17.4–17 ka, followed by closed-canopy North Patagonian Rain Forests (NPRF) with shade-tolerant thermophilous trees between ∼16.3–15.4 ka. These changes suggest an abrupt warming trend and an increase in precipitation at ∼16.3 ka, associated with a northward shift of the SWW. Subsequent increases in cold-tolerant hygrophilous trees between ∼15.4–12.8 ka imply a shift to cold, more humid conditions during the Antarctic Cold Reversal, suggesting stronger SWW influence. This was followed by stand-replacing fires brought by a sudden decline in summer precipitation at ∼12.8 ka, associated with a millennial-scale southward shift of the SWW which was contemporaneous with the onset of the Younger Dryas. Dominance of thermophilous, summer-drought tolerant Valdivian rainforest trees and high fire activity ensued between ∼10.3–7.8 ka, suggesting peak warmth and overall decline in annual precipitation associated with weakening of the SWW during the early Holocene. A multi-millennial cooling and wetting trend started at ∼7.8 ka, brought by stronger SWW influence, followed by recurrent, centennial-scale variations in temperature and precipitation starting at ∼6.4 ka. Deforestation, fire, and spread of non-native herbs by Chilean/European settlers began during the late 18th century. Abrupt vegetation changes in the Lago Proschle record were driven by rapid climate changes over the last 17,400 years amplified, in some instances, by fire disturbance.

The degree to which vegetation and site type have influenced fire regimes through the Holocene has not been investigated in detail in the temperate ecosystems of southern Patagonia. Here we present a first attempt using a paired-basin approach to study the evolution of fire regimes in sectors dominated by humid Nothofagus forests and the xeric Patagonian steppe in the Magallanes region of Chilean Patagonia (51°S). We analyzed sediment cores from two small lakes and a bog located within the same climate zone on opposite sides of the forest-steppe ecotone, ~28 km apart. The position of this biological boundary east of the Andes is controlled by the strength and position of the southern westerly winds, which constitute the sole source of precipitation throughout western Patagonia. Our results indicate that fires have occurred in the study region repeated times over the last ~13,000 years at bi- and tridecadal timescales. Sectors currently dominated by Patagonian steppe feature high frequency and low magnitude of local fires, and vice versa in humid forests. Climate-driven expansion of Nothofagus scrubland/woodland into steppe environments over the last ~4,200 years increased the magnitude and lowered the frequency of fire events, culminating with peak Nothofagus abundance, fire magnitude and frequency during the last millennium. We also detect divergences between lake-based vs. bog-based paleofire histories among paired sites located within the Patagonian steppe, ~12 km apart, which we attribute to local burning of the bog at times of lowered water table. This divergence suggests to us that bog-based vegetation and fire histories exacerbate a local, azonal, signal blurring extra-local or regional regimes, thus accounting for some discrepancies in the Quaternary paleovegetation/paleoclimate literature of southern Patagonia. Fil: Moreno, Patricio I.. Universidad de Chile; Chile Fil: Vilanova, Isabel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”; Argentina Fil: Villa Martínez, Rodrigo P.. Universidad de Magallanes; Chile Fil: Francois, Jean P.. Universidad de Playa Ancha; Chile

The Southern Westerly Winds (SWW) are the surface expression of geostrophic winds that encircle the southern mid-latitudes. In conjunction with the Southern Ocean, they establish a coupled system that not only controls climate in the southern third of the world, but is also closely connected to the position of the Intertropical Convergence Zone and CO2 degassing from the deep ocean. Paradoxically, little is known about their behavior since the last ice age and relationships with mid-latitude glacier history and tropical climate variability. Here we present a lake sediment record from Chilean Patagonia (51°S) that reveals fluctuations of the low-level SWW at mid-latitudes, including strong westerlies during the Antarctic Cold Reversal, anomalously low intensity during the early Holocene, which was unfavorable for glacier growth, and strong SWW since ∼7.5 ka. We detect nine positive Southern Annular Mode-like events at centennial timescale since ∼5.8 ka that alternate with cold/wet intervals favorable for glacier expansions (Neoglaciations) in southern Patagonia. The correspondence of key features of mid-latitude atmospheric circulation with shifts in tropical climate since ∼10 ka suggests that coherent climatic shifts in these regions have driven climate change in vast sectors of the Southern Hemisphere at centennial and millennial timescales. Fil: Moreno, P.I.. Universidad de Chile; Chile Fil: Vilanova, Isabel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”; Argentina Fil: Villa-Martínez, R.. Universidad de Magallanes; Chile Fil: Dunbar, R.B.. University of Stanford; Estados Unidos Fil: Mucciarone, D.A.. University of Stanford; Estados Unidos Fil: Kaplan, M.R.. Columbia University; Estados Unidos Fil: Garreaud, R.D.. Universidad de Chile; Chile Fil: Rojas, M.. Universidad de Chile; Chile Fil: Moy, C.M.. University of Otago; Nueva Zelanda Fil: De Pol-Holz, R.. Universidad de Magallanes; Chile Fil: Lambert, F.. Pontificia Universidad Católica de Chile; Chile

We describe the stratigraphy, age, constituent geochemistry and phenocryst thermobarometry of a closely spaced Holocene tephra couplet from Volcan Melimoyu (VMm), located in the Northern Patagonian Andes. The lower tephra unit (La Junta Tephra, Mm-1) is distinctly banded comprising a dominant lower layer of rhyodacitic (∼70% SiO2) pumiceous ash and lapilli (Mm-1p) that abruptly transitions to a subordinate upper scoriaceous layer (Mm-1s) of basaltic andesite composition (∼53% SiO2). This bimodality within Mm-1 contrasts significantly with the closely overlying Santa Ana Tephra (Mm-2) that has a homogeneous trachyte-dacite (∼63% SiO2) composition and is intermediate between the two magmatic end members of Mm-1. We propose a genetic affiliation between Mm-1 and Mm-2, and that the latter event likely represents a hybridised-remnant of those discrete magmas involved in the earlier Mm-1 eruption. To test this hypothesis we applied whole rock elemental mixing and fractional crystallisation model to reproduce the composition and crystallinity of Mm-2. Results indicate that Mm-2 can be reproduced by mixing ∼70% Mm-1p with ∼30% Mm-1s, with subsequent ∼13% fractional crystallisation of plagioclase, and minor amphibole, orthopyroxene, magnetite and biotite. Equilibrium P-T conditions calculated from Mm-1p phenocrysts point towards magma residency at moderately shallow depths (200–290 MPa, ∼7–10-km depth, 850–1000 °C), whereas Mm-1s phenocrysts indicate higher overall P-T conditions (240–480 MPa, ∼8.5–17-km depth, 1080–1150 °C). P-T conditions determined for Mm-2 (∼290 MPa, ∼10-km depth, 930–1000 °C) are similar to those of Mm-1p. There is no physical and/or geochemical evidence of mafic magma involvement in the Mm-2 eruption. Similar compositionally bimodal tephra are known from other Northern Patagonian Andean centres (i.e. Playas Blanca-Negra Tephra, Antillanca; Lepue Tephra, Michimahuida; Ho and H3 eruptions of Hudson) suggests that the intrusion of mafic magma into more silicic magma bodies is a common occurrence throughout this Andean sector. These widely dispersed, compositionally bimodal tephra not only provide key insights into pre-eruptive magmatic conditions and triggering processes, but can also be readily identified geochemically, and thereby be more fully utilised within future hazard- and paleoenvironmental-related studies.

South America lost around 52 genera of mammals during a worldwide event known as the Late Quaternary Extinction episode. More than 80% of South American mammals weighing > 44 kg succumbed. Analysis of the megafaunal extinction chronology in relation to human arrival and major climate changes have revealed slightly different extinction patterns in different eco-regions of the continent, highlighting the importance of detailed regional analysis in order to understand how the possible drivers of extinction operated. Here we present an analysis of the megafaunal extinction in the Ultima Esperanza (UE) area of southwestern Patagonia, Chile. We have compiled a comprehensive chronology of megafaunal extinctions and earliest human occupation between 18–7 cal ka BP, based on radiocarbon dates from published literature. We calculated confidence intervals using the GRIWM method to estimate the times of human arrival and megafaunal local extinctions, and then compared these events to the timing of major climate and vegetation changes, fire frequency increase, and the Reclus volcanic eruption. Our results suggest that a combination of human impacts and climate–vegetation change drove megafaunal extinctions in the UE area, with the balance of factors being taxon specific; the volcanic eruption does not seem to have exacerbated extinctions. Competition between humans and mega-carnivores seems to be the most plausible cause for the extinction of the mega-carnivores. Coexistence of humans with extinct horses, extinct camels, and mylodonts for several thousand years rules out a scenario of blitzkrieg overkill of megafauna by humans. The transition of vegetation from cold grasslands to Nothofagus forests corresponds with the disappearance of Hippidion saldiasi and Lama cf. owenii. The later full establishment of Nothofagus forests and an increasing fire frequency coincided with the disappearance of mylodonts. A climate-driven reduction of open environments plausibly reduced herbivore's populations making them susceptible to local extinction.

Diatoms can access mineral iron from dust if it is reduced, and reduced particulate iron is common in glacial dust sources., Little is known about the bioavailability of iron (Fe) in natural dusts and the impact of dust mineralogy on Fe utilization by photosynthetic organisms. Variation in the supply of bioavailable Fe to the ocean has the potential to influence the global carbon cycle by modulating primary production in the Southern Ocean. Much of the dust deposited across the Southern Ocean is sourced from South America, particularly Patagonia, where the waxing and waning of past and present glaciers generate fresh glaciogenic material that contrasts with aged and chemically weathered nonglaciogenic sediments. We show that these two potential sources of modern-day dust are mineralogically distinct, where glaciogenic dust sources contain mostly Fe(II)-rich primary silicate minerals, and nearby nonglaciogenic dust sources contain mostly Fe(III)-rich oxyhydroxide and Fe(III) silicate weathering products. In laboratory culture experiments, Phaeodactylum tricornutum, a well-studied coastal model diatom, grows more rapidly, and with higher photosynthetic efficiency, with input of glaciogenic particulates compared to that of nonglaciogenic particulates due to these differences in Fe mineralogy. Monod nutrient accessibility models fit to our data suggest that particulate Fe(II) content, rather than abiotic solubility, controls the Fe bioavailability in our Fe fertilization experiments. Thus, it is possible for this diatom to access particulate Fe in dusts by another mechanism besides uptake of unchelated Fe (Fe′) dissolved from particles into the bulk solution. If this capability is widespread in the Southern Ocean, then dusts deposited to the Southern Ocean in cold glacial periods are likely more bioavailable than those deposited in warm interglacial periods.

Concern is growing that global climate change will have widespread impact on the world’s terrestrial ecosystems, but future impacts are imperfectly constrained by ecosystem models and direct observations. All available data, including records of past ecological change, need to be utilized to assess the range of potential future outcomes. Here we show that pervasive ecosystem transformations occurred in response to warming and associated climatic changes during the last glacial-to-interglacial transition, which was of comparable magnitude to climatic change projected to occur in the next 100 to 150 years under high-emission scenarios. We used data from 596 published paleoecological records to examine compositional and structural changes in global terrestrial vegetation since the last glacial period, and to project the magnitude of ecosystem transformations under different emission scenarios in the future. Our results indicate that terrestrial ecosystems are highly sensitive to temperature change, and suggest that without major reductions in greenhouse-gas emissions to the atmosphere (i.e., in line with those targeted in the 2015 Paris Agreement), most terrestrial ecosystems worldwide are at risk of major transformation, with accompanying disruption of ecosystem services and impacts on biodiversity.

The 2008 eruption of Volcán Chaitén (VCha) in northwestern Patagonia was the first explosive rhyolitic eruption to have occurred within a century and provided an unprecedented scientific opportunity to examine all facets of the eruption ranging from magma rheology/ascent rates to ash-fall effects on biota and infrastructure. Up to very recently it was thought that the latest eruption prior to the 2008 event occurred c. 9750 cal. a BP. Although a number of researchers have recognised additional eruptive products, but their stratigraphy, age, and geochemical attributes have not been systematically described and/or recorded. In this study, we provide a detailed examination of andic cover-beds and tephra-bearing lake sequences located both proximally and distally to VCha, which record a series of hitherto unknown rhyolitic eruptive products and place all previous observations firmly within a coherent stratigraphic framework. Through major- and trace-element glass shard geochemistry we are able to confidently verify eruptive source. A total of 20 discrete tephra beds are recognised, with at least 10 having widespread areal distributions and/or depositional imprints broadly comparable to, or greater than, the 2008-tephra event. This record indicates that VCha has been continuously but intermittently active as far back as the end of the Last Glacial Maximum (c. 18,000 cal a BP) with two dominant, genetically related magma types and an intermediary ‘mixed’ type. Before this the eruptive record has been largely obscured and/or erased by widespread Andean piedmont glaciation. However, based on the tempo of VCha activity over the last c. 18,000 years, older VCha eruptives can be anticipated to occur as well as future hazardous explosive events. The new eruptive inventory will ultimately be useful for correlating equivalent-aged sequences and refining long-term eruptive tempo as well as corresponding temporal changes in magmatic evolution. Fil: Alloway, Brent V.. The University of Auckland; Nueva Zelanda. University Of Wollongong; Australia Fil: Pearce, Nick J.G.. Aberystwyth University; Reino Unido Fil: Moreno, Patricio I.. Universidad de Chile; Chile Fil: Villarosa, Gustavo. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte. Instituto Andino Patagónico de Tecnologías Biológicas y Geoambientales. Universidad Nacional del Comahue. Instituto Andino Patagónico de Tecnologías Biológicas y Geoambientales.; Argentina Fil: Jara, Ignacio. Victoria University Of Wellington; Nueva Zelanda Fil: De Pol-Holz, Ricardo. Universidad de Magallanes; Chile Fil: Outes, Valeria. Universidad de Chile; Chile

Paleoecological studies from the northern Patagonian Andes (40–44°S) have identified past changes in vegetation, fire regimes and paleoclimate since the last glaciation, including variations in strength and position of the Southern Westerly Winds (SWW). The extent to which records west and east of the Andes provide a congruent paleoclimatic history, however, has not been explored in detail in the literature. Physical and biological contrasts are evident between these regions today and are to be expected in paleoclimate reconstructions. In this context, we present pollen and charcoal records from sediment cores collected in Lago Espejo, a small closed-basin lake located in the core sector of the northern Patagonian Andes that spans uninterrupted the last ∼15,400 years. Following glacier withdrawal, the vegetation surrounding Lago Espejo features scattered Nothofagus woodlands, including relatively thermophilous rainforest trees between ∼15,400 and 14,400 cal yr BP. The disappearance of these trees and an abrupt rise in Nothofagus at ∼14,400 cal yr BP mark the establishment of closed-canopy forests during the Antarctic Cold Reversal, followed by increases in the cold-tolerant hygrophilous conifer Podocarpus nubigena during the Younger Dryas (∼12,700–11,500 cal yr BP). The Holocene vegetation consists of Nothofagus-dominated forests with modest variation in composition and structure until the present, attesting to the resilience of these forest communities to climate change and natural disturbance regimes. Rapid deforestation, anthropogenic fires and the establishment of artificial meadows with exotic herbs introduced by Europeans at ∼150 cal yr BP, triggered a rapid, large-magnitude landscape transformation unprecedented in the last 14,000 years. The timing and structure of vegetation changes revealed by the Lago Espejo record suggest that changes in the SWW were the main driver of vegetation and fire regimes in the Andes of northern Patagonia over the last 15,400 years. Comparison between multiple reconstructions from northern Patagonia reveals overall coherent vegetation and fire regime changes in the western and Andean sectors, and a spatially variable and more divergent behaviour in sites located further east. This spatial patter is akin to the present-day correlation between precipitation and SWW in this region.

Few studies have examined in detail the sequence of events during the last glacial termination (T1) in the core sector of the Patagonian Ice Sheet (PIS), the largest ice mass in the Southern Hemisphere outside of Antarctica. Here we report results from Lago Edita (47°8′ S, 72°25′ W, 570 m a.s.l.), a small closed-basin lake located in a valley overridden by eastward-flowing Andean glaciers during the Last Glacial Maximum (LGM). The Lago Edita record shows glaciolacustrine sedimentation until 19 400 yr BP, followed by organic sedimentation in a closed-basin lake and a mosaic of cold-resistant hygrophilous conifers and rainforest trees, along with alpine herbs between 19 400 and 11 000 yr BP. Our data suggest that the PIS retreated at least ∼ 90 km from its LGM limit between ∼ 21 000 and 19 400 yr BP and that scattered, low-density populations of cold-resistant hygrophilous conifers, rainforest trees, high-Andean and steppe herbs thrived east of the Andes during the LGM and T1, implying high precipitation levels and southern westerly wind (SWW) influence at 47° S. The conifer Podocarpus nubigena increased between 14 500 and 13 000 yr BP, suggesting even stronger SWW influence during the Antarctic Cold Reversal, after which it declined and persisted until 11 000 yr BP. Large increases in arboreal pollen at ∼ 13 000 and ∼ 11 000 yr BP led to the establishment of forests near Lago Edita between 10 000 and 9000 yr BP, suggesting a rise in the regional tree line along the eastern Andean slopes driven by warming pulses at ∼ 13 000 and ∼ 11 000 yr BP and a subsequent decline in SWW influence at ∼ 11 000 yr BP. We propose that the PIS imposed a regional cooling signal along its eastern, downwind margin through T1 that lasted until the separation of the northern and southern Patagonian ice fields along the Andes during the Younger Dryas period. We posit that the withdrawal of glacial and associated glaciolacustrine environments through T1 provided a route for the dispersal of hygrophilous trees and herbs from the eastern flank of the central Patagonian Andes, contributing to the afforestation of the western Andean slopes and pacific coasts of central Patagonia during T1. Fil: Henríquez, William I.. Victoria University Of Wellington; Nueva Zelanda Fil: Villa-Martinez, Rodrigo. Gaia-antártica Universidad de Magallanes; Chile Fil: Vilanova, Isabel. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales ; Argentina Fil: De Pol-Holz, Ricardo. Gaia-antártica Universidad de Magallanes; Chile Fil: Moreno, Patricio. Instituto de Ecología y Biodiversidad, Departamento de; Chile

We present detailed pollen and charcoal records from Lago Pichilafquén (~ 41°S) to decipher the effects of climate change and varying disturbance regimes on the composition and structure of the vegetation on the Andean foothills of northwestern Patagonia during the last 2600 yr. Here, temperate rainforests have dominated the landscape since 2600 cal yr BP with variations ranging from cool-temperate and wet north Patagonian rainforests to relatively warm and summer-drought-resistant Valdivian rainforests. We interpret relatively warm/dry conditions between 1900–2600, 690–750 and 320–430 cal yr BP, alternating with cold/wet conditions between 1500–1900, 750–1100 and 430–690 cal yr BP. Rapid deforestation and spread of plants introduced by Europeans occurred at 320 and 140 cal yr BP. The record includes five tephras with ages of 2130, 1460, 1310, 1210, and 340 cal yr BP, all of which precede local fire events and increases in trees favored by disturbance by less than 100 yr. We conclude that centennial-scale changes in the southern westerlies were the primary driver of vegetation shifts in northwestern Patagonia over the last 2600 yr. Within this interval, local disturbance regimes altered the structure, composition, and dynamics of the lowland rainforest vegetation during several discrete, short-lived episodes.

The biomisation method is used to reconstruct Latin American vegetation at 6000±500 and 18 000±1000 radiocarbon years before present (14C yr BP) from pollen data. Tests using modern pollen data from 381 samples derived from 287 locations broadly reproduce potential natural vegetation. The strong temperature gradient associated with the Andes is recorded by a transition from high altitude cool grass/shrubland and cool mixed forest to mid-altitude cool temperate rain forest, to tropical dry, seasonal and rain forest at low altitudes. Reconstructed biomes from a number of sites do not match the potential vegetation due to local factors such as human impact, methodological artefacts and mechanisms of pollen representivity of the parent vegetation. At 6000±500 14C yr BP 255 samples are analysed from 127 sites. Differences between the modern and the 6000±500 14C yr BP reconstruction are comparatively small; change relative to the modern reconstruction are mainly to biomes characteristic of drier climate in the north of the region with a slight more mesic shift in the south. Cool temperate rain forest remains dominant in western South America. In northwestern South America a number of sites record transitions from tropical seasonal forest to tropical dry forest and tropical rain forest to tropical seasonal forest. Sites in Central America show a change in biome assignment, but to more mesic vegetation, indicative of greater plant available moisture, e.g. on the Yucatán peninsula sites record warm evergreen forest, replacing tropical dry forest and warm mixed forest presently recorded. At 18 000±1000 14C yr BP 61 samples from 34 sites record vegetation reflecting a generally cool and dry environment. Cool grass/shrubland is prevalent in southeast Brazil whereas Amazonian sites record tropical dry forest, warm temperate rain forest and tropical seasonal forest. Southernmost South America is dominated by cool grass/shrubland, a single site retains cool temperate rain forest indicating that forest was present at some locations at the LGM. Some sites in Central Mexico and lowland Colombia remain unchanged in the biome assignments of warm mixed forest and tropical dry forest respectively, although the affinities that these sites have to different biomes do change between 18 000±1000 14C yr BP and present. The "unresponsive" nature of these sites results from their location and the impact of local edaphic influence.

A high-resolution macroscopic charcoal record from Lago Melli (42°46′S, 73°33′W) documents the occurrence of forest fires in the lowlands of Isla Grande de Chiloé, southern Chile, over the last 16,000 yr. Our data suggest that fire activity in this region was largely modulated by the position/intensity of the southern westerlies at multi-millennial time scales. Fire activity was infrequent or absent between 16,000–11,000 and 8500–7000 cal yr BP and was maximal between ∼ 11,000–8500 and 3000–0 cal yr BP. A mosaic of Valdivian/North Patagonian rainforest species started at ∼ 6000 cal yr BP, along with a moderate increase in fire activity which intensified subsequently at 3000 cal yr BP. The modern transition between these forest communities and the occurrence of fires are largely controlled by summer moisture stress and variability, suggesting the onset of high-frequency variability in summer precipitation regimes starting at ∼ 5500 cal yr BP. Because negative anomalies in summer precipitation in this region are teleconnected with modern El Niño events, we propose that the onset of El Niño-like variability at ∼ 5700–6200 cal yr BP led to a reshuffling of rainforest communities in the lowlands of Isla Grande de Chiloé and an increase in fire activity.

We report pollen and charcoal records from Vega Ñandú (∼ 51°0′S, 72°45′W), a small mire located near the modern forest-steppe ecotone in Torres del Paine National Park, southern Chile. The record shows an open landscape dominated by low shrubs and herbs between 12,600 and 10,800 cal yr BP, under cold and relatively humid conditions. Nothofagus experienced frequent, large-amplitude oscillations between 10,800 and 6800 cal yr BP, indicating recurrent transitions between shrubland/parkland environments, under warm and highly variable moisture conditions. A sustained increase in Nothofagus started at 6800 cal yr BP, punctuated by step-wise increases at 5100 and 2400 cal yr BP, implying further increases in precipitation. We interpret these results as indicative of variations in the amount of precipitation of westerly origin, with prominent increases at 6800, 5100, and 2400 cal yr BP. These pulses led to peak precipitation regimes during the last two millennia in this part of SW Patagonia. Our data suggest variations in the position and/or strength of the southern margin of the westerlies, most likely linked to variations in the extent and/or persistence of sea ice and sea-surface temperature anomalies in the Southern Ocean. Over the last two centuries the record shows a forest decline and expansion of Rumex acetosella, an exotic species indicative of European disturbance.

Holocene fire–climate–vegetation linkages are mostly understood at individual sites by comparing charcoal and pollen records with other paleoenvironmental proxy and model simulations. This scale of reconstruction often obscures detection of large-scale patterns in past fire activity that are related to changes in regional climate and vegetation. A network of 31 charcoal records from southern South America was examined to assess fire history along a transect from subtropic to subantarctic biomes. The charcoal data indicate that fire activity was greater than present at ca. 12,000 cal yr BP and increased further and was widespread at 9500 cal yr BP. Fire activity decreased and became more spatially variable by 6000 cal yr BP, and this trend continued to present. Atmospheric circulation anomalies during recent high-fire years show a southward shift in westerlies, and paleoclimate model simulations and data syntheses suggest that such conditions may have prevailed for millennia in the early Holocene when the pole-to-equator temperature gradients were weaker and annual temperatures were higher than present, in response to orbital-time-scale insolation changes.

Palynologic and stratigraphic data from Laguna Tahui (42°50′S, 73°30′W) indicate cool–temperate and humid conditions there between 14,000 and 10,000 14C yr B.P., followed by warmer and drier-than-present conditions between 10,000 and 7000 14C yr B.P., and subsequent cooling and rise in precipitation over the last 5800 14C yr. The thermophilous Valdivian trees Eucryphia cordifolia and Caldcluvia paniculata reached their maximum abundance during the early Holocene warm–dry phase (10,000–7000 14C yr B.P.), followed by a rise in lake levels and reexpansion of North Patagonian conifers starting at 7000 and 5800 14C yr B.P., respectively. Variations in the stratigraphic and geographic distribution of temperate rainforests in southern Chile suggest multimillennial trends in temperature and westerly activity, which are spatially and temporally coherent with paleoclimate records from neighboring regions. Climate variability at millennial and submillennial time scales may account for the establishment and persistence of fine-scale mosaics of Valdivian and North Patagonian rainforest species in low- to mid-elevation communities since ∼5800 14C yr B.P.

A pollen record from Lago Condorito (41°45′S, 73°07′W) shows that North Patagonian Rain Forest taxa predominated between about 13,000 and 12,200 14C yr B.P. in the lowlands of southern Chile, near the city of Puerto Montt. This was followed by the expansion and persistence of the conifer Podocarpus nubigena between 12,200 and 9900 14C yr B.P. Trees favored by disturbance expanded between 11,200 and 9900 14C yr B.P., concurrent with sharp and sustained increases of microscopic charcoal particles. Taxa of low-elevation rain forests expanded and became more diverse in pulses centered at 9900 and 9000 14C yr B.P., following the disappearance of P. nubigena. These data suggest conditions approaching modern climate between about 13,000 and 12,200 14C yr B.P. The climate cooled between 12,200 and 9900 14C yr B.P., then quickly warmed to interglacial conditions. Stand-replacing fires occurred near Lago Condorito between 11,200 and 9900 14C yr B.P., under cool-temperate, humid conditions. The proximity and reported antiquity of the Monte Verde archeological site raise the possibility that these fires were set by human activities.

The significance and cause of the decline in biomass burning across the Americas after ad 1500 is a topic of considerable debate. We synthesized charcoal records (a proxy for biomass burning) from the Americas and from the remainder of the globe over the past 2000 years, and compared these with paleoclimatic records and population reconstructions. A distinct post- ad 1500 decrease in biomass burning is evident, not only in the Americas, but also globally, and both are similar in duration and timing to ‘Little Ice Age’ climate change. There is temporal and spatial variability in the expression of the biomass-burning decline across the Americas but, at a regional–continental scale, ‘Little Ice Age’ climate change was likely more important than indigenous population collapse in driving this decline. peerReviewed

The prevailing view in the palaeoclimate literature of the last 20 years is that the Southern Westerly Winds (SWW) were intensified over southern Australia and Tasmania during the warmer-than-present early Holocene (11-8 ka). At similar latitudes on the opposite side of the southern mid-latitudes, palaeoclimate studies have suggested a poleward shift of the northern edge of the westerlies and focusing at 49 degrees S in southern South America. This zonal asymmetry contrasts with the modern day zonal symmetry displayed by the SWW and poses a formidable challenge to an understanding of the modes of climatic variability of the southern extra-tropics. This paper presents an updated synthesis of continuous, radiocarbon-dated palaeoenvironmental data from the westerlies zone of influence in all Southern Hemisphere continents. Synchronous multi-millennial trends in moisture, vegetation, fire, and hydrologic balance are remarkably consistent with the way the SWW changes impact upon the climate in Southern Hemisphere landmasses in the modern climate. Considering the modern relationships between local precipitation and zonal wind speeds, it is suggested that the SWW changed in a zonally symmetric manner at multi-millennial scale between 14 and 5 ka. Regional asymmetry develops after 5 ka across the Southern Hemisphere, with a pattern of precipitation anomalies akin to the modern functioning of El Nitio - Southern Oscillation, which started similar to 6200 years ago.

The Southern Hemisphere westerly winds are an important component of the climate system at hemispheric and global scales. Variations in their intensity and latitudinal position through an ice-age cycle have been proposed as important drivers of global climate change due to their influence on deep-ocean circulation and changes in atmospheric CO2. The position, intensity, and associated climatology of the southern westerlies during the last glacial maximum (LGM), however, is still poorly understood from empirical and modelling standpoints. Here we analyse the behaviour of the southern westerlies during the LGM using four coupled ocean-atmosphere simulations carried out by the Palaeoclimate Modelling Intercomparison Project Phase 2 (PMIP2). We analysed the atmospheric circulation by direct inspection of the winds and by using a cyclone tracking software to indicate storm tracks. The models suggest that changes were most significant during winter and over the Pacific ocean. For this season and region, three out four models indicate decreased wind intensities at the near surface as well as in the upper troposphere. Although the LGM atmosphere is colder and the equator to pole surface temperature gradient generally increases, the tropospheric temperature gradients actually decrease, explaining the weaker circulation. We evaluated the atmospheric influence on the Southern Ocean by examining the effect of wind stress on the Ekman pumping. Again, three of the models indicate decreased upwelling in a latitudinal band over the Southern Ocean. All models indicate a drier LGM than at present with a clear decrease in precipitation south of 40°S over the oceans. We identify important differences in precipitation anomalies over the land masses at regional scale, including a drier climate over New Zealand and wetter over NW Patagonia.

Climate is an important control on biomass burning, but the sensitivity of fire to changes in temperature and moisture balance has not been quantified. We analyze sedimentary charcoal records to show that the changes in fire regime over the past 21,000 yrs are predictable from changes in regional climates. Analyses of paleo- fire data show that fire increases monotonically with changes in temperature and peaks at intermediate moisture levels, and that temperature is quantitatively the most important driver of changes in biomass burning over the past 21,000 yrs. Given that a similar relationship between climate drivers and fire emerges from analyses of the interannual variability in biomass burning shown by remote-sensing observations of month-by-month burnt area between 1996 and 2008, our results signal a serious cause for concern in the face of continuing global warming. ispartof: Global Biogeochemical Cycles vol:26 issue:4 status: published