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Some well-managed organic soils are known to have higher crop yield potential than conventionally managed soils due to the greater soil quality and the ability to tolerate weed competition. However, low available soil mineral N and P in some organic systems may mask such soil quality–related benefits. We hypothesize that when plant-available N and P are not limiting, tillage-based highly diverse organic crop rotations have less yield loss (better crop tolerance) due to weed competition and higher crop yields than no-till conventional systems with low-diversity rotations. A greenhouse study was carried out in Saskatoon, Canada, using long-term (18-yr) organically managed soils (ORG) and no-till conventional soils (CONV) with three crop rotation diversities (LOW, MEDIUM, and HIGH) to compare the crop tolerance to weed competition under standard soil nutrient management conditions and under excess supply of mineral N and P. Under fertilized conditions, crop biomass increased by 50% and 69% in ORG and CONV systems, respectively. Weed biomass was similar between ORG and CONV systems under nonfertilized conditions but was 14% greater in CONV when excessive N and P were supplied. Crop biomass loss (crop tolerance) was not different among cropping systems under excess fertilizer or under standard fertilizer levels. Even with greater weed biomass under fertilized conditions, the CONV system showed crop tolerance similar to that of the ORG system. Under nonfertilized conditions, the crop biomass yield was 43% lower in ORG compared with CONV, and even after mineral N and P were applied, ORG systems showed less (17%) crop biomass than CONV. Further, differences in crop tolerance were not identified among crop rotations under both fertilizer levels. Overall, this study revealed that there were no yield benefits or better crop tolerance to weed competition in organically managed soils compared with no-till conventional soils, even under nonlimiting soil macronutrient conditions.

With the increasing resistance of wild oat (Avena fatua L.) to herbicides, there is a need to evaluate the potential of alternative cropping systems based on integrated weed management principles. A 5-yr field study at eight sites across Canada was used to evaluate the profitability of alternative cropping systems that have the potential to control wild oat using cultural practices in conjunction with herbicides. Cultural practices included twice the recommended seeding rates, fall-seeded winter crops, barley (Hordeum vulgare L.) silage, fallow, and alfalfa (Medicago sativa L.). Seven of the 14 cropping systems in this study did not include wild oat herbicide for three consecutive years, controlling wild oat entirely by cultural practices. Cropping system profitability varied by location. For many locations, combinations of barley silage and fall-seeded winter crops without wild oat herbicide application were as profitable as a system of canola (Brassica napus L.) and wheat (Triticum aestivum L.) with wild oat herbicide applied every year. Unprofitable systems generally included those with fallow, alfalfa, and fall-seeded winter crops in regions with rates of high winter kill. Wild oat control can be achieved with diverse cropping systems that are as profitable as conventional annual cropping that relies on herbicide control of wild oat.

The escalating evolution of weed species resistant to acetolactase synthase (ALS)-inhibitor herbicides makes alternative weed control strategies necessary for field crops that are dependent on this herbicide group. A fully integrated strategy that combined increased crop seeding rates (2X or 4X recommended), mechanical weed control with a minimum-tillage rotary hoe, and reduced-rate non—ALS inhibitor herbicides was compared with herbicides, rotary hoe, and seeding rates alone as a method of controlling ALS inhibitor—tolerant Indian mustard as a model weed. The full-rate herbicide treatment had the lowest weed biomass (98% reduction) and the highest yield of all treatments in 3 of 4 site-years, regardless of seeding rate. The fully integrated treatment at the 4X seeding rate had weed suppression rates equal to the full herbicide treatment at the recommended seeding rate. The fully integrated and reduced-rate herbicide treatments at the 4X seeding rate reduced weed biomass by 89% and 83%, respectively, compared with the control at the recommended seeding rate. The rotary hoe treatment alone resulted in poor weed control (≤38%), even at the highest seeding rate. Fully integrated and reduced-rate herbicide treatments at 2X and 4X seeding rates had yields equal to those of the full herbicide treatment at the recommended seeding rate. Partially or fully integrated weed control strategies that combine increased crop seeding rates and reduced-rate non—ALS inhibitor herbicides, with or without the use of a rotary hoe, can control weeds resistant to ALS-inhibitor herbicides, while maintaining crop yields similar to those achieved with full-rate herbicides. However, combining increased seeding rate, reduced-rate herbicides, and mechanical rotary hoe treatment into a fully integrated strategy maximized weed control, while reducing reliance on and selection pressure against any single weed control tactic.Nomenclature: Indian mustard, Brassica juncea (L.) Czern.

Concern over the development of herbicide-resistant weeds has led to interest in integrated weed management systems that reduce selection pressure by utilizing mechanical and cultural weed control practices in addition to herbicides. Increasing crop seeding rate increases crop competitive ability and thus can enhance herbicide efficacy. However, it is unknown how increasing the seeding rate affects an herbicide’s efficacy. The objective of this study was to examine the interaction between increasing seeding rate and herbicide dose to control weeds. To meet this objective, the herbicide fluthiacet-methyl was applied to field-grown lentil, with Indian mustard, a proxy for wild mustard, used as a model weed. The experiment was a factorial design with four lentil seeding rates and seven herbicide rates. Overall the herbicide dose response was altered by changing lentil seeding rate. Increasing lentil seeding rate decreased the weed biomass production when herbicides were not applied. In two of the four site-years, increasing lentil seeding rate lowered the herbicide ED50, the dose required to result in a 50% reduction in weed biomass. Increasing the crop seeding rate altered the dose response to provide greater weed control at lower herbicide rates compared with normal crop seeding rates. Increased seeding rates also resulted in higher and more stable crop seed yields across a wider range of herbicide dosages. These results suggest that dose–response models can be used to evaluate the efficacy of other weed management practices that can interact with herbicide performance.

Canola (Brassica napus L.) often experiences high seed losses prior to and during harvest operations. Pod drop and pod shatter are responsible for pre-harvest seed losses in canola although relatively little is known about the significance of pod drop to total pre-harvest seed losses in canola. This research describes a method to measure the predisposition for pod drop among canola genotypes that was developed and tested across different sites from 2013 to 2015. This method, referred to as pod-retention resistance, consists of the use of a force gauge to quantify the force required to break the pod pedicle at the pedicle-rachis junction. Two sets of field experiments were used, the first developed and modeled the factors that influence pod-retention resistance, pod drop and seed shatter. A second set of experiments was used to validate these findings on a broader range of genotypes and determine whether a relationship between pod retention resistance and pod drop could be determined. Results showed that the pod-retention resistance was significantly greater for pods on the proximal half of the rachis compared to the distal half of the rachis. Several factors contributed strongly to variation in pod retention resistance (rachis position, genotype and site) and pod drop (site, rachis position and genotype). The relative importance of individual factors varied between pod retention resistance and pod drop suggesting no relationship between the two. However, weight-adjusted pod retention resistance, i.e., average pod retention resistance corrected for the average weight of dropped pods, was strongly correlated with pod drop, particularly in the validation experiments. This method could be used by canola breeders to select for and develop genotypes with reduced predisposition for seed losses via pod drop.

Alternative cropping systems are gaining attention throughout the world in order to increase the sustainability of agro-ecosystems. On the Canadian prairies, the tillage-based crop-fallow system has been replaced by no-till reduced input systems or tillage-based organic systems with more diversity in crop rotations but with no external inputs. However, the long-term effects of these alternative systems on weed and yield parameters have not been assessed. A study to examine weed and crop yield dynamics under diverse cropping systems was conducted within a 18-year cropping systems study near Scott, Saskatchewan. The trial was laid out in a split-split-plot design with four replicates. The main plots were three levels of inputs; a high input system (HIGH) that used tillage and inputs to maximize yield, a reduced system (RED) that used no-till practices and minimal inputs, and a tillage-based organic system (ORG) with no external inputs. The subplots were cropping diversity (rotations); fallow-annual grains (LOW), diversified annual grains (DAG), and diversified annuals and perennial forage (DAP). The sub-sub plots were the six phases of each rotation. There was an input by rotation interaction for weed biomass but not for weed density and crop yields. ORG systems had 7× and 4× greater weed density (107 plants m −2 ), 4× higher weed biomass (154 kg ha −1 ), and 32% and 35% lower yields (1052 kg ha −1 ) than RED and HIGH systems respectively. RED and HIGH input systems had similar crop yields and lower weed density than ORG. The LOW rotation had the lowest weed density. LOW and DAG rotations had similar yields, which were higher than in DAP. All systems showed an increase in weed density and biomass over time but did not impact on crop yields which was increasing over time likely influenced by a concurrent increase in rainfall. This study concludes that eliminating tillage and reducing agrochemicals is possible but eliminating agrochemicals requires better crop rotations for weed management as well as for nutrient management.

In western Canada, more money is spent on wild oat herbicides than on any other weed species, and wild oat resistance to herbicides is the most widespread resistance issue. A direct-seeded field experiment was conducted from 2010 to 2014 at eight Canadian sites to determine crop life cycle, crop species, crop seeding rate, crop usage, and herbicide rate combination effects on wild oat management and canola yield. Combining 2× seeding rates of early-cut barley silage with 2× seeding rates of winter cereals and excluding wild oat herbicides for 3 of 5 yr (2011 to 2013) often led to similar wild oat density, aboveground wild oat biomass, wild oat seed density in the soil, and canola yield as a repeated canola–wheat rotation under a full wild oat herbicide rate regime. Wild oat was similarly well managed after 3 yr of perennial alfalfa without wild oat herbicides. Forgoing wild oat herbicides in only 2 of 5 yr from exclusively summer annual crop rotations resulted in higher wild oat density, biomass, and seed banks. Management systems that effectively combine diverse and optimal cultural practices against weeds, and limit herbicide use, reduce selection pressure for weed resistance to herbicides and prolong the utility of threatened herbicide tools.

Silene latifolia Poir. (white cockle or white campion) is an important invasive weed in North American agriculture. It exhibits dioecy, therefore, both male and female plants are required in order for seed production to occur. However, dioecious species being invasive is not common because of their limitations in pollination and subsequent seed production. The objective of this study is to determine the effect of pollination timing and distance on seed production of Silene latifolia. A series of experiments including pollination exclusion, timing and pollination distance were conducted in 2009 and 2010 at or around Saskatoon, Saskatchewan. For pollination exclusion, exclosures were built around the natural female plants for exclosure, sham-exclosure, and male and female combined treatments. Pollination timing was studied by applying exclosure, non-exclosure, night-exclosure, and day-exclosure treatments to individual female plants. Female plants were transplanted along a linear interval at six different distances from the pollen source to study the effect of pollination distance. S. latifolia was exclusively insect-pollinated and pollination occurred both day and night; however, in one year, pollination occurred mainly at night. Female plants that were in the range of 0–4 m from a compatible pollen source experienced no limitation to pollination. However, when the distance was increased further up to 128 m, pollination levels and subsequent seed production were declined. Moreover, there were differences in seed production between years suggesting that pollination was affected by the environmental conditions during pollination and the crop that white cockle was grown in. These experiments indicate that seed production in S. latifolia is limited by insect-pollination. Although there was pollination limitation for seed production at greater distances from a pollen source, the high fecundity rate (3000–18000 seeds per plant) resulted in a large seed output. Thus, we believe that a dioecious species may require characteristics that compensate the pollination limitation for a successful invasion.

Seedbank persistence in canola seeds is related to their potential to develop secondary dormancy. This can result in volunteer weed problems many years after canola production. The potential to be induced into secondary dormancy is controlled by both the canola genetics and the environment of the mother plant. However, the effect of time of harvesting on secondary dormancy potential is not known. The objective of this study was to determine the effect of harvest timing on potential to develop seed dormancy in canola. Six harvest samples were collected weekly from two canola genotypes (5440 and 5020) starting from 10 to 20% seed color change on the main stem until they were fully ripened. Freshly harvested seeds of 5440 and 5020 showed 13 and 16% primary dormancy at 32 and 33 d after flowering (DAF), respectively, but dormancy decreased with harvest timings and no dormancy was observed when seeds were fully mature (78 DAF). After dormancy induction, 10% of 5440 seeds were dormant at 32 DAF, but 94% of seeds were dormant at 78 DAF. Similarly, 70% of 5020 seeds were dormant at 33 DAF, but 90% of seeds were dormant at 68 DAF. Thus, seeds had lower potential to secondary dormancy at early development but have a high potential to secondary dormancy induction at full maturity. This study suggests that windrowing these canola genotypes at the recommended time (60% seed color change on the main stem) may reduce ability of the seed to develop secondary dormancy and thus reduce the persistence of seeds in the soil seedbank.

Cowcockle, an introduced summer annual weed of the Northern Great Plains, is being considered for domestication because of its high quality starch, cyclopeptides, and saponins. Loss of seed dormancy is one of the key desirable traits for domestication. To determine the potential for domestication of this species, an understanding of the seed dormancy and germination patterns is required. The objectives of this study were to evaluate seed dormancy in cowcockle ecotypes and determine how temperature and light affect seed dormancy. We evaluated 15 populations of cowcockle for primary dormancy by exposing them to five temperatures (5, 7.5, 10, 15, and 20 C) under two temperature regimes (constant and alternating) in both dark and light conditions. Freshly matured seeds of all the populations showed high levels of primary dormancy except ‘Mongolia’. Lower levels of dormancy at medium temperatures (10 and 15 C) and greater dormancy at low and high temperatures suggest conditional dormancy, a state at which seeds germinate over a narrower range of conditions compared to nondormant seeds. The effects of temperature regime, light, and their interaction was significant only at suboptimal (5 and 7.5 C) and supraoptimal (20 C) temperatures. Under these conditions, alternating temperatures were more effective in breaking the conditional dormancy, followed by light. The variation in optimum temperature, light, and their interactions among the cowcockle populations may be due to the plants evolving to adapt to their local environments. From a domestication perspective, the conditional dormancy in cowcockle can be observed as an evolutionary mechanism that prevents untimely germination following maturity and may not be a major obstacle for its domestication. Nomenclature: Cowcockle, Vaccaria hispanica (P. Mill.) Rauschert, VAPPY.

Weed control with herbicides is not possible in several systems including control of wild oat (Avena fatua L.) in tame oat (Avena sativa L.), red rice (Oryza sativa L. var. sylvatica) in rice (Oryza sativa L.), and in organic systems. Competitive crop cultivars can be used to manage weed competition if selective weed control by herbicides is not possible. However, most existing competitive crop cultivars often have low weed-free yield and poor grain quality. We hypothesize that the progeny of a cross between a competitive forage-type oat cultivar and a high-yielding and high grain quality grain oat cultivar will have high yield and good grain quality as well as high competitive ability (CA). The objective of this study was to evaluate progeny lines from a cross between a tall, competitive, forage-type oat cultivar with a semidwarf, high-yielding milling oat for their CA against wild oat. A field study was performed in two locations in Saskatoon, SK, Canada, in 2008 and 2009 using seven progeny lines and the two parents. Oat was seeded with and without wild oat at a target crop and weed density of 250 plant m⁻² in a randomized block design with four replicates. All the genotypes were high yielding and did not differ in grain yield or quality either in weed-free or weedy conditions. Some of the tall (CDC Baler, SA050498, and SA050479) and short (SA050040 and Ronald) genotypes had high average grain yield under both weed-free and weedy conditions compared with the other genotypes. The tall oat line SA050479 was among the most weed suppressive, which resulted in less wild oat biomass. The seedling total leaf area, crop height, and the seedling third leaf area were negatively correlated with wild oat biomass. This study demonstrates that a cross between a high-yielding and a highly competitive genotype can result in progeny with high CA, yield, and crop quality.

Effective weed management strategies are limited in organic cropping systems because herbicide use is prohibited. Enhancing crop competitive ability by integrating both cultural and mechanical weed control methods is a key strategy in such instances, but the relative efficacy of different cultural and mechanical strategies and their interactions and additive effects when combined is not well known. The objective of this study was to determine the individual and additive effects of cultural and mechanical methods on weed suppression and crop yield under organic conditions. A study was performed in two organically managed oat (Avena sativa L.) cropping systems in Saskatoon, SK, Canada, in 2008 and 2009. Three cultural practices, two crop genotypes (competitive and less competitive), high and standard crop densities (500 and 250 plants m–²), narrow and standard row spacings (11.5 and 23 cm), and two post-emergence tillage levels (harrowing and nonharrowed control) were factorially applied in a randomized block design. Increasing crop density and harrowing increased grain yield by 11 and 13%, respectively. Competitive genotype and high crop density reduced weed biomass by 22 and 52%, respectively. Combining high crop density with post-emergence harrowing increased the grain yield by 25%. The combined treatment effects on weed biomass were more profound, as the competitive genotype, increased seeding rate, and post-emergence harrowing decreased weed biomass by 71% compared with standard practices. Integrating cultural and mechanical weed management practices was superior to the use of individual practices because they additively control weeds in an organic cropping system.

The challenges associated with the adoption of conservation tillage and/or low-input cropping systems, whether organic or herbicide-free, across Canada are shaped by scale, environment, and local practices. A study in eastern Canada captured the challenges of introducing low-input cropping systems in mature (20+ yr) tillage treatments in a barley/red clover/corn/soybean rotation. Each mature tillage system came with its own weed problems, but this did not affect crops such as barley and red clover, which produced similar yields across high and low input systems. However, some form of primary tillage was needed to achieve adequate weed control and yield in organic (ORG) and herbicide-free (HF) systems. The HF and ORG systems with no-till actually failed to produce a corn crop but produced soybean yields that were half or less than that for other treatments. The successful combination of conservation tillage practices and low-input systems in eastern Canada would thus appear to be crop-specific, being easier to achieve in competitive cereal crops. In western Canadian organic agriculture, tillage is practiced with low-disturbance chisel plows instead of inversion plows. However, green manure crops (summer cover crops) are often terminated with tandem discs. Both roller crimpers and mowing can successfully kill annual green manure crops such as field pea and rye, and usually result in reduced weed growth following termination. However, the lack of tillage can result in lower crop yields in wheat following green manure terminated by roller crimping compared to tillage. Challenges for no-till organic practices include perennial weed control and soil fertility. Overall, flexible crop production programs such as the former Manitoba Pesticide Free Production program and the “Agriculture raisonnéeTM” program in Québec are more likely to promote sustained environmental, economic, and social prosperity than rigid adherence to organic or no-till practices.

Organic farmers in western Canada rely on tillage to control weeds and incorporate crop residues that could plug mechanical weed-control implements. However, tillage significantly increases the risk of soil erosion. For farmers seeking to reduce or eliminate tillage, potential alternatives include mowing or using a roller crimper for terminating green manure crops (cover crops) or using a minimum tillage (min-till) rotary hoe for mechanically controlling weeds. Although many researchers have studied organic crop production in western Canada, few have studied no-till organic production practices. Two studies were recently conducted in Saskatchewan to determine the efficacy of the following alternatives to tillage: mowing and roller crimping for weed control, and min-till rotary hoeing weed control in field pea (Pisum sativum L.). The first study compared mowing and roller crimping with tillage when terminating faba bean (Vicia faba L.) and field pea green manure crops. Early termination of annual green manure crops with roller crimping or mowing resulted in less weed regrowth compared with tillage. When compared with faba bean, field pea produced greater crop biomass, suppressed weeds better and had less regrowth. Wheat yields following pea were not affected by the method of termination. Thus, this first study indicated that roller crimping and mowing are viable alternatives to tillage to terminate field pea green manure crops. The second study evaluated the tolerance and efficacy of a min-till rotary harrow in no-till field pea production. The min-till rotary hoe was able to operate in no-till cereal residues and multiple passes did not affect the level of residue cover. Field pea exhibited excellent tolerance to the min-till rotary hoe. Good weed control occurred with multiple rotary hoe passes, and pea seed yield was 87% of the yield obtained in the herbicide-treated check. Therefore, this second study demonstrated that min-till rotary hoeing effectively controls many small seeded annual weeds in the presence of crop residue and thus can reduce the need for tillage in organic-cropping systems.

Phosphorus (P) can be low in soil under low input organic management; however, beneficial crop plant associations with arbuscular mycorrhizal fungi (AMF) are known to promote crop nutrition and increase phosphorus uptake. Thus, management strategies that promote AMF associations are particularly desirable for low-input cropping systems. The objectives of this study were to determine the impact of seeding rate on AMF colonization and the impact of AMF colonization on P concentration and uptake by organically grown field pea and lentil. Field experiments examined the impact of three seeding rates of field pea and lentil on P uptake and crop yield. Phosphorus accumulation was examined further in a controlled growth chamber experiment, in which field pea was sown at rates corresponding to those used in the field and harvested at 10-day intervals until 50 days after emergence. In the field, the level of AMF colonization of roots remained at 80% for field pea, while colonization of lentil increased with increasing seeding rates from 77% to 88%. The level of AMF colonization of field pea achieved in the growth chamber after 50 days was 80% for the two highest seeding rates and 60% for the low seeding rate. The rate at which AMF colonization occurred did not vary between treatments. Ultimately, AMF colonization level did not affect P accumulation. In contrast to several previous studies, both field and growth chamber experiments revealed that AMF colonization was not reduced at higher seeding rates. These results suggest that organic farmers may increase seeding rates without adversely affecting P nutrition.

Summary Secondary seed dormancy has been linked to seedbank persistence of volunteer oilseed rape (Brassica napus) in western Canada. It has been suggested that there is a genetic component to secondary seed dormancy expression in oilseed rape, but little is known of its importance in relation to non-genetic factors. In a series of experiments we investigated the relative importance of genotype, seed size, time of windrowing and pre- and post-harvest environment on the expression of secondary seed dormancy. We found that genotype contributed between 44 and 82% to the total variation in secondary seed dormancy. A broad range in secondary seed dormancy expression was observed among 16 genotypes examined. Nevertheless, three-quarters of the genotypes investigated exhibited relatively high potential for the expression of secondary seed dormancy (back-transformed mean 71% dormant seeds). Seed size contributed 21% to the total variation, while the influence of seed maturity (harvest regime) on secondary seed dormancy expression was negligible. Despite diverging environmental conditions during the four growing seasons spanning these experiments, the influence of pre-harvest environment on seed dormancy expression was relatively small and ranged from 0.1% to 4.5%. Secondary seed dormancy potential decreased over time during seed storage. This decrease was greatest when seeds were stored at ambient temperatures and least when seeds were stored at −70°C.