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Patagonian vegetation has dramatically changed in composition and distribution over the last 23,000 years. Although the vegetation history has been inferred from individual pollen records, the regional patterns and drivers of vegetation development are poorly understood. High resolution pollen and charcoal data from eleven sites located along the eastern flanks of the Patagonian Andes (41–43°S) were examined to reconstruct the Lateglacial and Holocene vegetation and fire history of steppe/forest ecotone and separate the relative influence of climatic versus non-climatic factors in shaping the patterns of ecological change. Pollen data indicate that, as the Lateglacial climate became progressively wetter, the initial steppe vegetation was replaced by open forest of Nothofagus in the Lateglacial and early Holocene periods, and by closed forest in the late Holocene. Fire activity was lowest during the Lateglacial/early-Holocene transition and gradually increased through the Holocene. Prior to ca 5000 cal yr BP, the conifer Austrocedrus chilensis possibly persisted in isolated populations along the eastern boundary of its modern distribution. Cooler/more humid conditions after ca 5000 cal yr BP allowed the development of the modern mixed Nothofagus–Austrocedrus forest. The paleoenvironmental record points to the sensitivity of the forest/steppe ecotone in the past, not only to climate but also to complex environmental feedbacks that amplified the effects of climate change. Fil: Iglesias, Virginia. State University of Montana; Estados Unidos Fil: Whitlock, Cathy. State University of Montana; Estados Unidos Fil: Markgraf, Vera. State University of Colorado Boulder; Estados Unidos. University of Arizona; Estados Unidos Fil: Bianchi, Maria Martha. Secretaría de Cultura de la Nación. Dirección Nacional de Cultura y Museos. Instituto Nacional de Antropología y Pensamiento Latinoamericano; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Oficina de Coordinación Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales "Bernardino Rivadavia"; Argentina

The vast diversity of present vegetation and environments that occur throughout South America (12°N to 56°S) is the result of diverse processes that have been operating and interacting at different spatial and temporal scales. Global factors, such as the concentration of CO2 in the atmosphere, may have been significant for high altitude vegetation during times of lower abundance, while lower sea levels of glacial stages potentially opened areas of continental shelf for colonisation during a substantial portion of the Quaternary. Latitudinal variation in orbital forcing has operated on a regional scale. The pace of climate change in the tropics is dominated by precessional oscillations of c. 20 kyr, while the high latitudes of the south are dominated by obliquity oscillations of c. 40 kyr. In particular, seasonal insolation changes forced by precessional oscillations must have had important consequences for the distribution limits of species, with potentially different effects depending on the latitude. The availability of taxa, altitude and human impact, among other events, have locally influenced the environments. Disentangling the different forcing factors of environmental change that operate on different timescales, and understanding the underlying mechanisms leads to considerable challenges for palaeoecologists. The papers in this Special Issue present a selection of palaeoecological studies throughout South America on vegetation changes and other aspects of the environment, providing a window on the possible complexity of the nature of transitions and timings that are potentially available.

Patagonian vegetation has dramatically changed in composition and distribution over the last 16,000. yr. Although patterns of vegetation change are relatively clear, our understanding of the processes that produce them is limited. High-resolution pollen and charcoal records from two lakes located at lat 41°S provide new information on the postglacial history of vegetation and fire activity at the forest-steppe ecotone, and help clarify the relative importance of local and regional drivers of late-Holocene ecological change. Our results suggest that late-glacial parkland was colonized by shrubs at ca. 11,200. cal. yr BP and this vegetation persisted until 4900. cal. yr BP, when increased humidity allowed for the establishment of Nothofagus forest. The late Holocene is characterized by oscillations in forest dominance largely driven by changes in humidity, possibly associated with the onset or strengthening of ENSO. In the last 4900. yr, humid periods (4900-3800 and 2850-1350. cal. yr BP) have promoted Nothofagus forest, whereas drier times (3800-2850 and 1350-450. cal. yr BP) have favored Austrocedrus expansion. At intermediate moisture levels, however, the lower forest supported both taxa, and fire became an important control of community composition, with severe, infrequent fires facilitating Nothofagus regeneration and high fire frequency and intensity supporting Austrocedrus. © 2012. Fil: Iglesias, Virginia. State University of Montana; Estados Unidos Fil: Whitlock, Cathy. State University of Montana; Estados Unidos Fil: Bianchi, Maria Martha. Universidad Nacional del Comahue. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales.; Argentina Fil: Villarosa, Gustavo. Universidad Nacional del Comahue. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales.; Argentina Fil: Outes, Ana Valeria. Universidad Nacional del Comahue. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte. Instituto Patagónico de Estudios de Humanidades y Ciencias Sociales.; Argentina

Recent airborne pollen records data from Northern Patagonia (San Carlos de Bariloche, Argentina, Lat. 41.1435° S, Long. 71.375° W, 800 m elevation) suggest that pollen transport takes place from the west to the east slope of the Andes. However, the atmospheric characteristics responsible of this transport have not yet been studied. The aim of this paper is to assess potential source areas and to describe the involved atmospheric mechanisms of the trans-Andean pollen transport. Methodology relies on the analysis of backward trajectories of air masses calculated with the HYSPLIT 4.9 regional model for particular days where airborne pollen of Weinmannia trichosperma Cav. was detected east of the Andes. This pollen type was selected because it is found regularly at localities in eastern Patagonia beyond its present-day distribution. Weinmannia's substantial presence during early Holocene times would also benefit from better knowledge of its transport mechanisms. Correspondence between atmospheric trajectories and the position of sources was checked using GIS maps. Mode T, Principal Component Analysis (PCA) with Varimax rotation was used to identify the main spatial structure of geopotential height anomalies producing the calculated trajectories. Eighty-eight cases showed that the calculated directions of trajectories trended from the Northwest to Southwest passing over the Chilean region of W. trichosperma distribution. PCs results showed two patterns of negative anomalies over southern Patagonia. The prevailing circulation pattern which drives airborne transport is the presence of a trough located south of 37 to 40° S with its axis over western Patagonia. The synoptic situations for two cases highly correlated with principal component scores were described.

210Pb, 137Cs and 14C dated sediments of two late Holocene landslide lakes in the Provincial Park Lagunas de Yala (Laguna Rodeo, Laguna Comedero, 24°06′S, 65°30′W, 2100 m asl, northwestern Argentina) reveal a high-resolution multi-proxy data set of climate change and human impact for the past ca. 2000 years. Comparison of the lake sediment data set for the 20th century (sediment mass accumulation rates MARs, pollen spectra, nutrient and charcoal fluxes) with independent dendroecological data from the catchment (fire scars, tree growth) and long regional precipitation series (from 1934 onwards) show that (1) the lake sediment data set is internally highly consistent and compares well with independent data sets, (2) the chronology of the sediment is reliable, (3) large fires (1940s, 1983/1984-1989) as documented in the local fire scar frequency are recorded in the charcoal flux to the lake sediments and coincide with low wet-season precipitation rates (e.g., 1940s, 1983/1984) and/or high interannual precipitation variability (late 1940s), and (4) the regional increase in precipitation after 1970 is recorded in an increase in the MARs (L. Rodeo from 100 to 390 mg cm-2 yr-1) and in an increase in fern spores reflecting wet vegetation. The most significant change in MARs and nutrient fluxes (Corg and P) of the past 2000 years is observed with the transition from the Inca Empire to the Spanish Conquest around 1600 AD. Compared with the pre-17th century conditions, MARs increased by a factor of ca. 5 to >8 (to 800 +130, -280 mg cm-2 yr-1), PO4 fluxes increased by a factor of 7, and Corg fluxes by a factor of 10.5 for the time between 1640 and 1930 AD. 17th to 19th century MARs and nutrient fluxes also exceed 20th century values. Excess Pb deposition as indicated by a significant increase in Pb/Zr and Pb/Rb ratios in the sediments after the 1950s coincides with a rapid expansion of the regional mining industry. Excess Pb is interpreted as atmospheric deposition and direct human impact due to Pb smelting. © 2006 Elsevier Ltd and INQUA. Fil: Lupo, Liliana Concepcio. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina. Universidad Nacional de Jujuy; Argentina Fil: Bianchi, Maria Martha. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina. Universidad Nacional del Comahue; Argentina Fil: Aráoz, Ezequiel. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina. Universidad Nacional de Tucumán. Facultad de Ciencias Naturales e Instituto Miguel Lillo. Laboratorio de Investigaciones Ecológicas de las Yungas; Argentina Fil: Grau, Hector Ricardo. Universidad Nacional de Tucumán. Facultad de Ciencias Naturales e Instituto Miguel Lillo. Laboratorio de Investigaciones Ecológicas de las Yungas; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Lucas, Christoph. University of Bern; Suiza Fil: Kern, Raoul. University of Bern; Suiza Fil: Camacho, María. Universidad Nacional de Jujuy; Argentina Fil: Tanner, Willi. University of Bern; Suiza Fil: Grosjean, Martin. University of Bern; Suiza

The history of the low-elevation forest and forest-steppe ecotone on the east side of the Andes is revealed in pollen and charcoal records obtained from mid-latitude lakes. Prior to 15,000 cal yr BP, the vegetation was characterized by steppe vegetation with isolated stands of Nothofagus. The climate was generally dry, and the sparse vegetation apparently lacked sufficient fuels to burn extensively. After 15,000 cal yr BP, a mixture of Nothofagus forest and shrubland/steppe developed. Fire activity increased between 13,250 and 11,400 cal yr BP, contemporaneous with a regionally defined cold dry period (Huelmo/Mascardi Cold Reversal). The early-Holocene period was characterized by an open Nothofagus forest/shrubland mosaic, and fire frequency was high in dry sites and low in wet sites; the data suggest a sharp decrease in moisture eastward from the Andes. A shift to a surface-fire regime occurred at 7500 cal yr BP at the wet site and at 4400 cal yr BP at the dry site, preceding the expansion of Austrocedrus by 1000–1500 yr. The spread of Austrocedrus is explained by a shift towards a cooler and wetter climate in the middle and late Holocene. The change to a surface-fire regime is consistent with increased interannual climate variability and the onset or strengthening of ENSO. The present-day mixed forest dominated by Nothofagus and Austrocedrus was established in the last few millennia.

Late Pleistocene palaeolakes on the eastern side of Andes (glacial lakes) and in the Patagonian Steppe (pluvial lakes) along 41°S latitude were studied. Natural dams (glaciers, moraines) that maintained a high water level in glacial palaeolakes were melted or cut by erosion during Late Pleistocene climate amelioration. With the retreat of the glaciers from the Andean valleys during prominent regional deglaciation, outflow from the eastern side of Andes to the Pacific Ocean was formed about 13.2 ka BP. As a result, the water level in glacial palaeolakes fell rapidly in stages. These changes were often preceded by volcanic-tectonic activity, which is suggested to be the “touch of butterfly” releasing mass balance adjustment in the changing landscape. The lacustrine tephra stratigraphic pattern from the Andes, comprising more than 50 horizons since the last glaciation, is presented. The correlation of palaeoenvironmental changes between the Andes and the Patagonian Steppe is suggested on the basis of two dacite marker horizons. The deglaciation event that led to development of the new hydrological system in the Andean palaeolakeland formed several large but short-lived palaeolakes in closed basins in the Patagonian Steppe probably due to melting of local ice and permafrost accumulated during glaciation on the surrounding mountains and elevated plains.

Several high-resolution continental records have been reported recently in sites in South America, but the extent to which climatic variations were synchronous between the northern and southern hemispheres during the Late-glacial–Holocene transition, and the causes of the climatic changes, remain open questions. Previous investigations indicated that, east of the Andes, the middle and high latitudes of South America warmed uniformly and rapidly from 13 000 14C yr BP, with no indication of subsequent climate fluctuations, equivalent, for example, to the Younger Dryas cooling. Here we present a multiproxy continuous record, radiocarbon dated by accelerated mass spectroscopy, from proglacial Lake Mascardi in Argentina. The results show that unstable climatic conditions, comparable to those described from records obtained in the Northern Hemisphere, dominated the Late-glacial–Holocene transition in Argentina at this latitude. Furthermore, a significant advance of the Tronador ice-cap, which feeds Lake Mascardi, occurred during the Younger Dryas Chronozone. This instability suggests a step-wise climatic history reflecting a global, rather than regional, forcing mechanism. The Lake Mascardi record, therefore, provides strong support for the hypothesis that ocean–atmosphere interaction, rather than global ocean circulation alone, governed interhemispheric climate teleconnections during the last deglaciation. © 1997 John Wiley & Sons, Ltd.