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Victrix (Poliobrya) akbet Volynkin, Titov & Černila, sp. nov. (Figs 1–4, 17, 18, 25, 28–30) urn:lsid:zoobank.org:act: B5A1BA4F-9D55-4735-B797-B82ED29BD602 Type material. Holotype (Figs 1, 17): ♂, 12.VI.2013, NE Kazakhstan, Pavlodar Region, Bayanaul district, Bayanaul Mts., vic. of Kempirtas Mt., steppe near rocks, 420 m, 50°51’24.65” N 75°34’37.21” E, A.V.Volynkin, S.V. Titov & M. Černila leg., slide AV1256 ♂ Volynkin (Coll. NHMUK, ex coll. CAV). Paratypes: 118 specimens of both sexes, with the same data as the holotype, slides AV0940 ♂, AV1257 ♂, AV1271 ♀ Volynkin (Colls STP, CAV, MČK, MDS, ASV, NHMUK); 41 specimens of both sexes, 12–14. VI.2014, NE Kazakhstan, Pavlodar Region, Bayanaul district, Bayanaul Mts., eastern coast of Toraygyr lake, Obet Bulak gorge, N50°52.113', E75°40.286', 380 m, steppe near rocks, Volynkin A.V. & Titov S.V. leg. (Colls STP and CAV); 10 ♂, 8 ♀, 16. VI.2017, same locality, Titov S.V. leg. (Coll. STP); 8 ♂, 4 ♀, 16. VI.2018, same locality and collector (Coll. STP); 3 ♂, 27.V.2015, NE Kazakhstan, Pavlodar area, Bayanaul district, Bayanaul Mts., vic. of Nayzatas Mt., 50°51'7.81"N 75°34'1.44"E, M. Černila, S.V. Titov & M. Kučinić leg. (Coll. STP); 3 ♂, 2–3. VI.2015, Central Kazakhstan, Karaganda region, Aktogay district, Bektau Ata Mts., 47°25'57.0"N 74°48'00.0"E, S. V.Titov & M. Černila leg. (Colls STP & CAV); 1 ♂, 07. VI.2012, E Kazakhstan, Zaisan district, south of Zaisan Lake, Manrak Mts., h= 1400 m, 47°18’ N, 84°36’ E, Yakovlev R.V. leg., slide AV1305 ♂ Volynkin (Coll. CAV). Etymology. Akbet is a highest mountain in the Bayanaul mountain massif. Diagnosis. Externally, V. akbet resembles dark specimens of V. fabiani (Figs 5–8), but differs by its shorter forewing with a less elongated apex and darker forewing ground color. The male genitalia of V. akbet (Figs 17, 18) are more similar to those of V. frigidalis (Figs 9, 10, 19, 20) but differ by their somewhat slender uncus, basally broader and distally narrower valva, smaller subapical valva process and basally narrower vesica. Compared to those of V. fabiani (Figs 21, 22), the male genitalia of V. akbet have a relatively shorter uncus, a much broader basally valva, a longer and narrower setose valva apex field, and a shorter and broader valva subapical process. The male genitalia of V. akbet differ from those of V. umovii (Fig. 23) as their uncus is more strongly curved medially and they have a narrower valva apex, broader vesica and slightly larger cornutus. Compared to those of V. patula, the male genitalia of V. akbet have a slightly longer uncus, a significantly broader basally valva, a slightly more prominent valva apex, a more massive subapical process of valva and a larger cornutus in the vesica. The female genitalia, V. akbet (Fig. 25) differs from V. frigidalis (Fig. 26) by its shorter apophyses posteriores, somewhat longer lateral processes of the antrum, and broader and longer sclerotized anterior part of the ductus bursae. Compared to those of V. umovii (Fig. 27), the female genitalia, V. akbet have a significantly broader antrum and larger appendix bursae. The females of V. fabiani and V. patula are unknown. Molecular data. As the male genitalia of Poliobrya species are very similar, COI 5’ sequences of four specimens of V. akbet from two localities (Bayanaul Mts. and Bektau-Ata Mts.) were compared with COI 5’ sequences of one V. frigidalis specimen from SW Mongolia, two V. fabiani specimens from two localities in West Mongolia and six V. umovii specimens from three localities in Russia and West Mongolia (see Table 1). The distance between the specimens of V. akbet and the specimen of V. frigidalis is 2.32–2.48% (Fig. 34). The distance between the specimens of V. akbet and the specimens of V. fabiani is 2.02–2.34%. The distance between the specimens of V. akbet and the specimens of V. umovii is 1.54–1.85% (Fig. 34). COI 5' sequences of Victrix akbet can be diagnosed from those of sequenced V. frigidalis, V. fabiani and V. umovii specimens by the following unique combination of six character states: 102(G), 133(T), 241(T), 400(T), 407(T) and 553(C), while V. fabiani is characterized by a unique combination of the following six character states: 16(A), 142(C), 368(A), 389(A), 499(G) and 562(G); V. frigidalis is characterized by a unique combination of seven character states: 69(T), 169(C), 220(C), 229(G), 370(G), 474(G) and 574(T); Victrix umovii is widespread, and COI 5' sequences of its different populations have five variable states, while three character states are unique for the species in comparison with other sequenced Poliobrya species: 364(C), 379(C) and 646(C) (see Table 2). Description. Adult (Figs 1–4, 28–30). Forewing length 11–14 mm in males (12 mm in holotype) and 13–14 mm in females. Antennae ciliate. Head and thorax dark, blackish brown with admixture of olive-green scales; abdomen pale brown with a mixture of blackish brown scales. Forewing moderately broad, trigonal, with rounded apex. Forewing ground color dark greenish brown, often with light brown suffusion in the medial area. Orbicular and reniform stigmata dark, blackish-brown; subbasal line blackish, wavy; antemedial line irregularly wavy, double, whitish proximally and blackish distally; postmedial line wavy-dentate, double, blackish proximally and whitish distally; subterminal line diffuse, irregularly wavy, pale brown; terminal line blackish, present as a row of blackish dots between the veins; cilia greenish brown. Hindwing pale greyish brown, subterminal and medial lines present, both diffused, dark brownish grey. Discal spot large, semilunar, diffused, dark brownish grey. Cilia brownish grey. Male genitalia (Figs 17, 18). Uncus long, narrow, curved, apically pointed. Tegumen short, moderately broad. Juxta broad, shield-shaped. Vinculum short, V-shaped. Valva massive, lobe-like, basally broad, distally slightly narrowed, its apical part with setose field; subapical process moderately long, trigonal, apically pointed. Aedeagus long, narrow, almost straight; vesica moderately long, curved, with long plate-like cornutus on short subapical diverticulum. Female genitalia (Fig. 25). Ovipositor short, conical. Apophyses posteriores and anteriores elongated, thin. Antrum large, sclerotized, funnel-like, with short trigonal postero-lateral processes. Ductus bursae long, its posterior part short, membranous; anterior part long, sclerotized, dorso-ventrally flattened, curved posteriorly. Appendix bursae short, C-shaped; corpus bursae short, saccate. Distribution and bionomics. The new species is known from the Kazakh Upland (Bayanaul mountain massif in the Pavlodar Region and Bektau-Ata mountain massif in the Karaganda Region) and the Saur-Tarbagatai mountain massif (Manrak Ridge in the East Kazakhstan Region). In both, Bayanaul and Bektau Ata Mts., V. akbet inhabits rocky buttes at medium altitudes (Figs 28–33). Moths fly from late May until late June. Preimaginal stages are unknown. 10 16 69 102 133 142 169 220 229 241 265 274 287 364 368 370 379 389 400 407 412 433 474 499 548 553 562 574 586 622 646 Voucher No. V. akbet PT T T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986009 Bektau-Ata V. akbet PT T T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986010 Bektau-Ata V. akbet PT C T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986011 Bayanaul V. akbet PT T T C G T T T T A T G C T T G A A G T T A T A T C C A C T A T MH986012 Bayanaul V. frigidalis T T T A C T C C G A A C T T G G A G C C A T G T T T A T T G T MH 986005 SW Mongolia V. fabiani BC ZSM T A C A C C T T A A A C T T A A A A C C A T A G C T G C G G T W Mongolia Lep 90260 V. fabiani T A C A C C T T A A A C T T A A A A C C A T A G C T G C T G T MH 986004 W Mongolia V. umovii Mongolian T T C A C T T T A A A C C C G A C G C C A C A T C T A C T A C MH986006 Altai V. umovii T T C A C T T T A A A C T C G A C G C C G T A T T T A C T A C MH 986007 Russian Altai V. umovii T T C A C T T T A A A C T C G A C G C C G T A T C T A C T A C MH 986008 Russian Altai V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19106 S Ural V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19107 S Ural V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19108 S Ural

Victrix (Poliobrya) akbet Volynkin, Titov & Černila, sp. nov. (Figs 1���4, 17, 18, 25, 28���30) urn:lsid:zoobank.org:act: B5A1BA4F-9D55-4735-B797-B82ED29BD602 Type material. Holotype (Figs 1, 17): ♂, 12.VI.2013, NE Kazakhstan, Pavlodar Region, Bayanaul district, Bayanaul Mts., vic. of Kempirtas Mt., steppe near rocks, 420 m, 50��51���24.65��� N 75��34���37.21��� E, A.V.Volynkin, S.V. Titov & M. Černila leg., slide AV1256 ♂ Volynkin (Coll. NHMUK, ex coll. CAV). Paratypes: 118 specimens of both sexes, with the same data as the holotype, slides AV0940 ♂, AV1257 ♂, AV1271 ♀ Volynkin (Colls STP, CAV, MČK, MDS, ASV, NHMUK); 41 specimens of both sexes, 12���14. VI.2014, NE Kazakhstan, Pavlodar Region, Bayanaul district, Bayanaul Mts., eastern coast of Toraygyr lake, Obet Bulak gorge, N50��52.113', E75��40.286', 380 m, steppe near rocks, Volynkin A.V. & Titov S.V. leg. (Colls STP and CAV); 10 ♂, 8 ♀, 16. VI.2017, same locality, Titov S.V. leg. (Coll. STP); 8 ♂, 4 ♀, 16. VI.2018, same locality and collector (Coll. STP); 3 ♂, 27.V.2015, NE Kazakhstan, Pavlodar area, Bayanaul district, Bayanaul Mts., vic. of Nayzatas Mt., 50��51'7.81"N 75��34'1.44"E, M. Černila, S.V. Titov & M. Kučinić leg. (Coll. STP); 3 ♂, 2���3. VI.2015, Central Kazakhstan, Karaganda region, Aktogay district, Bektau Ata Mts., 47��25'57.0"N 74��48'00.0"E, S. V.Titov & M. Černila leg. (Colls STP & CAV); 1 ♂, 07. VI.2012, E Kazakhstan, Zaisan district, south of Zaisan Lake, Manrak Mts., h= 1400 m, 47��18��� N, 84��36��� E, Yakovlev R.V. leg., slide AV1305 ♂ Volynkin (Coll. CAV). Etymology. Akbet is a highest mountain in the Bayanaul mountain massif. Diagnosis. Externally, V. akbet resembles dark specimens of V. fabiani (Figs 5���8), but differs by its shorter forewing with a less elongated apex and darker forewing ground color. The male genitalia of V. akbet (Figs 17, 18) are more similar to those of V. frigidalis (Figs 9, 10, 19, 20) but differ by their somewhat slender uncus, basally broader and distally narrower valva, smaller subapical valva process and basally narrower vesica. Compared to those of V. fabiani (Figs 21, 22), the male genitalia of V. akbet have a relatively shorter uncus, a much broader basally valva, a longer and narrower setose valva apex field, and a shorter and broader valva subapical process. The male genitalia of V. akbet differ from those of V. umovii (Fig. 23) as their uncus is more strongly curved medially and they have a narrower valva apex, broader vesica and slightly larger cornutus. Compared to those of V. patula, the male genitalia of V. akbet have a slightly longer uncus, a significantly broader basally valva, a slightly more prominent valva apex, a more massive subapical process of valva and a larger cornutus in the vesica. The female genitalia, V. akbet (Fig. 25) differs from V. frigidalis (Fig. 26) by its shorter apophyses posteriores, somewhat longer lateral processes of the antrum, and broader and longer sclerotized anterior part of the ductus bursae. Compared to those of V. umovii (Fig. 27), the female genitalia, V. akbet have a significantly broader antrum and larger appendix bursae. The females of V. fabiani and V. patula are unknown. Molecular data. As the male genitalia of Poliobrya species are very similar, COI 5��� sequences of four specimens of V. akbet from two localities (Bayanaul Mts. and Bektau-Ata Mts.) were compared with COI 5��� sequences of one V. frigidalis specimen from SW Mongolia, two V. fabiani specimens from two localities in West Mongolia and six V. umovii specimens from three localities in Russia and West Mongolia (see Table 1). The distance between the specimens of V. akbet and the specimen of V. frigidalis is 2.32���2.48% (Fig. 34). The distance between the specimens of V. akbet and the specimens of V. fabiani is 2.02���2.34%. The distance between the specimens of V. akbet and the specimens of V. umovii is 1.54���1.85% (Fig. 34). COI 5' sequences of Victrix akbet can be diagnosed from those of sequenced V. frigidalis, V. fabiani and V. umovii specimens by the following unique combination of six character states: 102(G), 133(T), 241(T), 400(T), 407(T) and 553(C), while V. fabiani is characterized by a unique combination of the following six character states: 16(A), 142(C), 368(A), 389(A), 499(G) and 562(G); V. frigidalis is characterized by a unique combination of seven character states: 69(T), 169(C), 220(C), 229(G), 370(G), 474(G) and 574(T); Victrix umovii is widespread, and COI 5' sequences of its different populations have five variable states, while three character states are unique for the species in comparison with other sequenced Poliobrya species: 364(C), 379(C) and 646(C) (see Table 2). Description. Adult (Figs 1���4, 28���30). Forewing length 11���14 mm in males (12 mm in holotype) and 13���14 mm in females. Antennae ciliate. Head and thorax dark, blackish brown with admixture of olive-green scales; abdomen pale brown with a mixture of blackish brown scales. Forewing moderately broad, trigonal, with rounded apex. Forewing ground color dark greenish brown, often with light brown suffusion in the medial area. Orbicular and reniform stigmata dark, blackish-brown; subbasal line blackish, wavy; antemedial line irregularly wavy, double, whitish proximally and blackish distally; postmedial line wavy-dentate, double, blackish proximally and whitish distally; subterminal line diffuse, irregularly wavy, pale brown; terminal line blackish, present as a row of blackish dots between the veins; cilia greenish brown. Hindwing pale greyish brown, subterminal and medial lines present, both diffused, dark brownish grey. Discal spot large, semilunar, diffused, dark brownish grey. Cilia brownish grey. Male genitalia (Figs 17, 18). Uncus long, narrow, curved, apically pointed. Tegumen short, moderately broad. Juxta broad, shield-shaped. Vinculum short, V-shaped. Valva massive, lobe-like, basally broad, distally slightly narrowed, its apical part with setose field; subapical process moderately long, trigonal, apically pointed. Aedeagus long, narrow, almost straight; vesica moderately long, curved, with long plate-like cornutus on short subapical diverticulum. Female genitalia (Fig. 25). Ovipositor short, conical. Apophyses posteriores and anteriores elongated, thin. Antrum large, sclerotized, funnel-like, with short trigonal postero-lateral processes. Ductus bursae long, its posterior part short, membranous; anterior part long, sclerotized, dorso-ventrally flattened, curved posteriorly. Appendix bursae short, C-shaped; corpus bursae short, saccate. Distribution and bionomics. The new species is known from the Kazakh Upland (Bayanaul mountain massif in the Pavlodar Region and Bektau-Ata mountain massif in the Karaganda Region) and the Saur-Tarbagatai mountain massif (Manrak Ridge in the East Kazakhstan Region). In both, Bayanaul and Bektau Ata Mts., V. akbet inhabits rocky buttes at medium altitudes (Figs 28���33). Moths fly from late May until late June. Preimaginal stages are unknown. 10 16 69 102 133 142 169 220 229 241 265 274 287 364 368 370 379 389 400 407 412 433 474 499 548 553 562 574 586 622 646 Voucher No. V. akbet PT T T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986009 Bektau-Ata V. akbet PT T T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986010 Bektau-Ata V. akbet PT C T C G T T T T A T A C T T G A A G T T A T A T C C A C T A T MH986011 Bayanaul V. akbet PT T T C G T T T T A T G C T T G A A G T T A T A T C C A C T A T MH986012 Bayanaul V. frigidalis T T T A C T C C G A A C T T G G A G C C A T G T T T A T T G T MH 986005 SW Mongolia V. fabiani BC ZSM T A C A C C T T A A A C T T A A A A C C A T A G C T G C G G T W Mongolia Lep 90260 V. fabiani T A C A C C T T A A A C T T A A A A C C A T A G C T G C T G T MH 986004 W Mongolia V. umovii Mongolian T T C A C T T T A A A C C C G A C G C C A C A T C T A C T A C MH986006 Altai V. umovii T T C A C T T T A A A C T C G A C G C C G T A T T T A C T A C MH 986007 Russian Altai V. umovii T T C A C T T T A A A C T C G A C G C C G T A T C T A C T A C MH 986008 Russian Altai V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19106 S Ural V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19107 S Ural V. umovii T T C A C T T T A A A T T C G A C G C C A T A T C T A C T A C MM19108 S Ural, Published as part of Volynkin, Anton V., Titov, Sergey V., ��ernila, Matja��, Truuverk, Andro & Saldaitis, Aidas, 2019, A new species of Victrix Staudinger, 1879 from Kazakhstan (Lepidoptera, Noctuidae), pp. 325-336 in Zootaxa 4563 (2) on pages 326-328, DOI: 10.11646/zootaxa.4563.2.6, http://zenodo.org/record/2601247, {"references":["Volynkin, A. V. & Gyulai, P. (2018) A new species of Athaumasta Hampson, 1906 (Lepidoptera, Noctuidae, Bryophilinae) from the Altai Mountains of Mongolia and China. Zootaxa, 4508 (4), 594 - 600. https: // doi. org / 10.11646 / zootaxa. 4508.4.10","Volynkin, A. V. (2012) Noctuidae of the Russian Altai (Lepidoptera). Proceedings of the Tigirek State Natural Reserve, 5, 1 - 339."]}

Dasypolia (Dasypolia) volynkini Ronkay, Ronkay, Gyulai & Pekarsky, 2014 (Figs 2, 6) Dasypolia (Dasypolia) volynkini G. Ronkay, L. Ronkay, Gyulai & Pekarsky, 2014, Fibigeriana Supplement 2: 146, pl. 29, figs 5, 6, gen. fig. 7 (Type locality: " Kazakhstan, Sary-Su river, 200 m, 230 km E Kzyl-Orda"). Type material examined: Paratypes: 2 females, SE Kazakhstan, Alma-Ata area, 3 km SE Arkharly Pass, 1070 m, 44 º 13 '36.33'' N, 77 º 43 '53.64'' E, 2–3.x. 2010, leg. Taranov B.T. & Egorov P.V., slide AV0963f Volynkin (Coll. A. Volynkin, Barnaul). Additional material examined: 1 male, 12–13.iv. 2014, SE Kazakhstan, Almaty area, 10 km E of Kerbulak, Sholak Mts., h= 800 m, 43 ° 55 '7.66"N, 77 ° 47 '18.45"E, Egorov P.V. & Rakhimov R.D. leg., slide AV 1332m Volynkin (Coll. A. Volynkin, Barnaul); 1 male, 19–20.iv. 2014, SE Kazakhstan, Almaty area, Zhetyzhol Mts., 9 km NW of Ulken-Sulutor (old Krasnogorka) village, h= 1250 m, 43 ° 20 '32.70"N, 75 °06'44.40"E, Egorov P.V. & Rakhimov R.D. leg. (Coll. A. Volynkin, Barnaul). Diagnosis of male. Male wingspan 24–25 mm. The male of D. volynkini (Fig. 2) has the ground colour and the wing pattern same as in the female, but male antennae are serrate and fasciculate. The male genitalia of D. volynkini (Fig. 6) are close to D. minuta (Figs 1, 5) and Dasypolia akkeregeshena Ronkay, Ronkay, Gyulai & Pekarsky, 2014 (Figs 3, 7) but differ from D. minuta by the shorter and broader uncus, the larger dorso-medial process of juxta, the more rounded vinculum, narrower and apically more rounded harpe, and the medially broader vesica; from D. akkeregeshena differ by the somewhat longer dorso-medial process of juxta, the more rounded vinculum, the basally broader valva, the terminally narrower harpe, the differently shaped dorso-medial sclerotized plate of the distal segment of aedagus, and the medially broader vesica. From the third species of the species-complex, Dasypolia zolotuhini Ronkay, Ronkay, Gyulai & Pekarsky, 2014 (Figs 4, 8) differ clearly by the apically broader uncus, the flattened and broader harpe, the reduced costal extension, the differently shaped dorso-medial sclerotized plate of the distal segment of aedagus, and the larger and stronger sclerotised carinal plate. Description of male genitalia (Fig. 6). Uncus moderately long, broad, obtuse apically; tegumen short, penicular lobes moderately broad; vinculum short, U-shape. Juxta shield-like, with two narrow latero-apical and one large dorsomedial processes. Valva elongated, basally broad, distally much narrowed, apically rounded; sacculus broad, setose; costa broad and strongly sclerotised, costal extention broad and very short; clasper moderately broad, curved; harpe moderately broad, curved, apically club-like broadened. Aedeagus elongated, narrow; carinal plate strongly sclerotised, with several thorns; vesica membranous, tubular, curved ventrally, broadened medially. Bionomics and distribution. The species is known from southern Kazakhstan (the type-locality, Sary-Su river valley) and several localities in southeastern Kazakhstan (northeastern part of the Tien Shan mountain massif). As well as other members of the D. minuta species-complex (Ronkay et al. 2014), D. volynkini has overwintering males (the both known males were collected in mid April), which is unusual for the nominate subgenus. In southeastern Kazakhstan all specimens were collected at low altitudes (800–1250 m.). The species inhabits dry low mountains, in Arkharly pass the paratypes were collected in habitat with predominance of Spiraea, Ephedra, Atraphaxis and Prunus shrubs, and various herbs, e. g., Ferula and Poaceae; in Sholak mountains (Fig. 9) the habitat dominated by Salsola, Atraphaxis and Prunus shrubs; in Zhetyzhol mountains (Fig. 10) the habitat dominated by Rosa shrubs and various grasses (Poaceae, etc.).

Euchalcia matovi Volynkin & Titov, sp. n. (Figs 1���5, 11, 12, 17, 18, 22) Type material. Holotype: male, 09.vi. 2013, E Kazakhstan, East Kazakhstan area, Urdzhar district, Tarbagatai Ridge, 6.7 km N of Kyzymbet (Alekseevka) village, mesophilous shrubby slopes, 1300 m. 47 �� 18.365 ��� N, 81 �� 32.152 ��� E, Volynkin A.V., Titov S.V. & Černila M. leg. Slide AV0865 Volynkin (Coll. ZISP). Paratypes: 22 males, 15 females, with the same labels as holotype, slides AV0853, AV0863, AV0864, AV0870, AV0871, AV0872, AV0873, AV0874 Volynkin (males), AV0866, AV0867, AV 1177, AV 1178 Volynkin (females) (Colls ZISP, AVB, STP, MČK); 2 females, 9.vii. [19] 67, Kazakhstan, Dzhungarsky Alatau, I. Kostin [leg.] / Euchalcia inconspicua, Zolotarenko det., slides AV 1177, AV 1178 Volynkin (females) (Coll. SZMN). Diagnosis. The new species is the third and the smallest member of the E. inconspicua species-complex. Externally E. matovi (Figs 1���5) is close to E. anthea (Figs 9, 10), but differs from it by smaller size (wingspan of E. anthea 31���36 mm), somewhat paler, more brilliant forewing colouration and less contrast pattern; from E. inconspicua (Figs 7, 8) differs by smaller size (wingspan of E. inconspicua 32���37 mm), broader forewing in females, with less acutely pointed apex of forewing in both sexes, much paler forewing colouration with paler medial field and less contrast pattern. E. matovi is also externally similar to E. shugnana (Sheljuzhko, 1929) (Fig. 6), but differs from it by somewhat less acutely pointed apex of forewing in both sexes, broader forewing in females, more brilliant forewing colouration. The male genitalia of the new species (Figs 11, 12) are close to those of E. inconspicua (Figs 13, 14). The genital capsule of E. matovi differs by longer and narrower uncus, narrower and distally more angled valva with larger ventro-medial triangular lobes; E. matovi has a larger aedeagus in comparison to the genital capsule of E. inconspicua, the vesica of E. matovi is longer, and subterminal cornuti are somewhat longer and more to robust. From E. anthea (Fig. 15) the male genitalia of E. matovi differ by medially broader valva with larger ventro-medial triangular lobes, somewhat longer aedeagus in comparison with the genital capsule, shorter and broader basal tube of vesica, and longer and stronger subterminal cornuti; from E. shugnana (Fig. 16) the male genitalia differ by longer uncus, longer harpe and the vesica structure: in E. matovi the basal tube is much longer, subterminal cornuti longer and stronger. The female genitalia of E. matovi (Figs 17, 18) differ from those of E. inconspicua (Fig. 19) by longer ductus bursae with much stronger sclerotised bulbous anterior part; from E. anthea (Fig. 20) differ by shorter and not S-shaped ductus bursae and somewhat stronger sclerotised bulbous anterior part; from E. shugnana (Fig. 21) differ by much larger, more rounded anterior part of ductus bursae, larger appendix bursae, longer corpus bursae. Description. Adult (Figs 1���5, 22). Wingspan 28���31 mm, length of forewing 12���14 mm. Antennae filiform. Head, thorax and abdomen ochreous; tegulae and patagia golden ochreous; Forewing broad, with pointed apex. Ground colour of forewing ochreous with metallic golden sheen; basal line thin, brown, wavy, indistinct; antemedial and postmedial lines thin, dark golden brown; antemedial line bent at A 1, postmedial line smooth curved; suffusion of the medial area varies from golden ochreous to golden brown; submarginal line thin, smooth curved, dark golden brown, with conspicuous dark golden brown shadow at inner margin; terminal line thin, dark brown; cilia ochreous or brown; orbicular, reniform and subcellular stigma conspicuous, with thin dark brown borders. Hindwing ochreous or ochreous brown; terminal band wide, dark, fuzzy; medial band thin, slightly wavy, dark brown; discal spot thin, brown, indistinct. Male genitalia (Figs 11, 12). Uncus narrow, long, curved, apically pointed; tegumen broad, moderately long; juxta broad, shield-like, with long conical apical process; vinculum long, V-shaped; valva moderately broad, distally narrowed, angled, with well developed ventro-medial triangular lobes; sacculus relatively small, clavus small, short, triangular; harpe long, thin; aedeagus large, moderately broad; vesica tubular, consists of broad tubular basal part and sphaerical subterminal bulb with three-six large subterminal cornuti and one terminal cornutus. Female genitalia (Figs 17, 18). Ovipositor short, conical. Apophyses posteriores and anteriores long, thin. Ostium bursae membranous; ductus bursae moderately long, tubular, rugose, with strongly sclerotised bulbous proximal plate at junction to corpus bursae; appendix bursae small, elliptical, membranous; ductus bursae membranous, sack-like, moderately long. Distribution and bionomics. The new species is known from the south-west part of Tarbagatai Ridge in East Kazakhstan and the Dzhungarsky Alatau Mts. in South-East Kazakhstan. At the type locality, E. matovi inhabits mesophilous slopes with Lonicera and Rosa shrubs (Fig. 23). Etymology. The species name is dedicated to Dr. Alexey Matov (ZISP), an expert on Asian Noctuoidea., Published as part of Volynkin, Anton V., Titov, Sergey V. & Ivanova, Maria S., 2014, A new species of Euchalcia H��bner, [1821] from Kazakhstan (Lepidoptera, Noctuidae), pp. 493-497 in Zootaxa 3784 (4) on pages 493-497, DOI: 10.11646/zootaxa.3784.4.8, http://zenodo.org/record/227990

Perigrapha (Perigrapha) yasawii Volynkin, Titov & Knyazev, sp. n. (Figs 1���3, 11, 12) Type material. Holotype: male, 14.iv. 2014, S Kazakhstan, South Kazakhstan Region, 39 km WSW of Turkestan city, Syrdarya river valley, h= 180 m, N 43 �� 10.006 ���, E 67 �� 50.389 ���, at light. Volynkin A.V., Titov S.V. & Knyazev S.A. leg. Slide AV 1242 Volynkin (Coll. Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia [ZISP]). Paratypes: 26 males, with the same data as holotype (Colls A. Volynkin, Barnaul, Russia [AVB]; S. Knyazev, Omsk, Russia [SKO]; S. Titov, Pavlodar, Kazakhstan [STP]; ZISP); 1 male, Perovsk [S Kazakhstan, Kyzyl-Orda Region, vicinity of Kyzyl-Orda city], Syr-Darya reg., 2.iv. 1909, E. Miller [leg.] (Coll. ZISP). Slides AV 1230, AV 1243 Volynkin (males). Diagnosis. The new species belongs to P. circumducta complex of the P. i-cinctum ([Denis & Schifferm��ller], 1775) species-group including P. sechuana G. Ronkay, L. Ronkay & Hacker, 2010, P. circumducta circumducta (Lederer, 1855) and P. circumducta pallescens (Draudt 1934) (Hreblay 1996; Ronkay et al. 2001; Ronkay et al. 2010). P. yasawii sp. n. (Figs 1���3) differs externally from P. circumducta circumducta (Figs 4���6) by its somewhat smaller size (wingspan of P. circumducta circumducta 42���53 mm), broader pectination of male antennae, more concolorous brownish-grey ground colour of forewings (in P. c. circumducta ground colour varies from reddishbrown to blackish-brown), pale subbasal area, pale subterminal area with narrower dark field outwards the postmeidal line, somewhat smaller orbicular stigma, more rounded reniform stigma, much narrower suborbicular patch and more greyish hindwings; from P. circumducta pallescens (Figs 7, 8) by its smaller size, broader pectination of male antennae, brownish-grey ground colour of forewings, more concolorous subterminal area with narrower dark field outwards the postmeidal line, more rounded reniform stigma, much narrower suborbicular patch and more greyish hindwings; from P. sechuana (Figs 9, 10) by its smaller size, narrower forewings, concolorous brownish-grey ground colour of forewings (in P. sechuana ground colour reddish-brown), paler subbasal area, pale subterminal area with narrower dark field outwards the postmeidal line, somewhat smaller orbicular stigma, more rounded reniform stigma, much narrower suborbicular patch and less unicolorous, more greyish hindwings with paler basal area. The male genitalia of P. yasawii sp. n. (Figs 11, 12) differ from those of P. circumducta (Figs 13) by the proximally broader, almost quadrangular uncus, more quadrangular juxta, broader clasper with shorter harpe and broader proximal part of ampulla; from P. sechuana (Fig. 14) by proximally broader, almost quadrangular uncus, broader clasper with shorter harpe, less asymmetrical and apically more dilated distal parts of valvae, longer aedeagus and much longer, less dorsally recurved vesica. Description. Adult (Figs 1���3). Male. Wingspan 37���43 mm. Antennae bipectinate, with long branches. Head, thorax and abdomen brownish-grey. Ground colour of forewing pale brownish-grey, medial area dark, greyishbrown, paler near the costal margin; antemedial and postmedial lines thin, dark grey; antemedial line almost straight, slightly curved near the costal margin; postmedial line almost straight, only arcuate around the cell; subterminal line indistinct; terminal area darker than subterminal; terminal line blackish, divided into short spots; orbicular and reniform stigmata and suborbicular patch being connected with reniform stigma with merged edges, pale greyish-ochreous, bordered with blackish; cilia grey. Hindwing greyish-brown; discal spot and medial line indistinct, dark; cilia dark grey. Female unknown. Male genitalia (Figs 11, 12). Uncus short, almost quadrangular, apically slightly rounded; tegumen short, weak; penicular lobes weak, narrow; juxta long, almost quadrangular; vinculum short, V-shaped; valva long, relatively narrow; cuculli long, straight, narrow, without pollex-like extensions, asymmetrical; distal part of left cucullus longer, apically narrower than that of right cucullus; ampulla long, slender, moderately curved, apically pointed; clasper short, distally strongly broadened, harpe very short; aedeagus very long, curved; vesica long, twisted, with subconical medial diverticulum with very small field of weak and short spinules, and large terminal field of spinules on broad diverticulum. Distribution and bionomics. The new species is known from two localities in South Kazakhstan. P. y as a w i i sp. n. inhabits dry shrubby river valleys (Fig. 15). Etymology. The species name is dedicated to Khodja Ahmed Yasawi, a Turkic poet and Sufi, whose mausoleum is in Turkestan city, less than 40 km from the type locality., Published as part of Volynkin, Anton V., Titov, Sergey V. & Knyazev, Svyatoslav A., 2014, A new Perigrapha Lederer, 1857 from South Kazakhstan (Lepidoptera, Noctuidae), pp. 292-296 in Zootaxa 3856 (2) on pages 292-296, DOI: 10.11646/zootaxa.3856.2.8, http://zenodo.org/record/224978, {"references":["Ronkay, G., Ronkay, L., Gyulai, P. & Hacker, H. (2010) New Orthosiini (Lepidoptera, Noctuidae, Hadeninae) species and genera from the wide sense Himalayan region. Esperiana, 15, 127 - 221.","Hreblay, M. (1996) Neue palaarktische Taxa aus der Gattung Perigrapha Lederer, 1857 (Lepidoptera, Noctuidae). Esperiana, 4, 65 - 94.","Ronkay, L., Yela, J. L. & Hreblay, M. (2001) Hadeninae II. Noctuidae Europaeae, 5. Entomological Press, Soro, 1 - 452."]}

Orthosia (Orthosia) ronkayorum Volynkin & Titov, sp. n. (Figs 1–4, 11, 12, 17, 21, 22) Type material. Holotype: male, 08.v. 2013, NE Kazakhstan, Pavlodar area, Ekibastuz district, 12 km NW Shiderty village, coast of Shiderty Reservoir, 51 ° 47 '54.21" N, 74 ° 35 '11.57" E, Volynkin A.V. & Titov S.V. leg. (Coll. Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia [ZISP]). Slide AV0855 Volynkin. Paratypes: 1 female, with the same data as holotype (Coll. A. V. Volynkin, Barnaul, Russia [AVB]); 15 males, 1 female, 19.iv. 2012, NE Kazakhstan, Pavlodar area, Ekibastuz district, 11–12 km NW Shiderty, 51 ° 48 ' N, 74 ° 35 ' E, S.V. Titov leg. (Colls. ZISP; AVB; S. V. Titov, Pavlodar, Kazakhstan [STP]). Slides AV0845 Volynkin (male), AV0846 Volynkin (female). Diagnosis. The species belongs to the O. incerta species-group. It is the northernmost species of the O. picata lineage which includes five described species: O. picata (Bang-Haas, 1912) (Figs. 5 –7), O. faqiri Hreblay & Plante, 1994, O. feda Hreblay & Plante, 1994, O. ariuna Hreblay, 1991 (Fig. 8) and O. reshoefti Hreblay, 1994. O. ronkayorum is closely related to Central Asian O. picata and O. ariuna. It is the smallest species of the group. As well as other members of the group, it is an externally variable species, and other than its small size the remaining differences are in the genitalia; in addition, O. ronkayorum has relatively smaller eyes than the related species O. picata. The male genitalia of O. ronkayorum (Figs 11, 12) are most similar to those of O. picata (Figs 13, 14), but differ by longer and medially broader uncus, narrower juxta, longer vinculum, somewhat narrower apical part of cucullus, narrower basal part of pollex, smaller clavus, broader distal part of clasper and somewhat shorter thorn of carina; from other Central Asian species of the group – Mongolian O. ariuna (Fig. 15), the male genitalia of the new species differ by broader medial part of uncus, narrower juxta, somewhat shorter vinculum, smaller cucullus with narrower neck, narrower and longer pollex, smaller clavus, broader distal part of clasper, broader lateral bar of carina and longer thorn of carina; from Pakistanian O. faqiri and O. feda (figured by Hreblay & Plante 1994) the male genitalia of O. ronkayorum differ by broader medial part of uncus, somewhat narrower juxta, longer vinculum, narrower valva, smaller cucullus with narrower neck, narrower basal part of pollex, smaller clavus, broader distal part of clasper, larger lateral bar of carina and longer thorn of carina; from O. incerta (Hufnagel, 1766) (Figs. 9, 10, 16) which occur sympatrically in North-East Kazakhstan, the male genitalia of O. ronkayorum differ by apically pointed uncus, shorter and narrower juxta, longer vinculum, smaller and narrower cucullus, narrower pollex, smaller clavus, broader lateral bar of carina and shorter thorn of carina. In the female genitalia O. ronkayorum (Fig. 17) differs from O. picata (Fig. 18) by less sclerotised ostium bursae, somewhat shorter ductus bursae, shorter and less sclerotised appendix bursae and larger corpus bursae; from O. ariuna (Fig. 19) differs by less sclerotised ostium bursae with shorter lateral crests, larger broad posterior part of ductus bursae, shorter and stronger sclerotised appendix bursae, broader corpus bursae and longer signa; from O. faqiri differs by larger broad posterior part of ductus bursae, shorter and stronger sclerotised appendix bursae; from O. reshoefti (figured by Hreblay & Plante 1994) differs by larger broad posterior part of ductus bursae, larger corpus bursae and longer signa; from O. incerta (Fig. 20) differs by shorter and less sclerotised ostium bursae with shorter, less sclerotised and not dentate lateral crests, broader ductus bursae with much more strongly broadened posterior part, stronger sclerotised appendix bursae, somewhat broader corpus bursae. Description. Adult (Figs 1–4, 21, 22). Wingspan 28–32 mm, length of forewing 12–15 mm. Antennae biserrate on males, dentate on females. Head, thorax and abdomen brown or dark brown. Forewing elongated, with pointed apex. Ground colour of forewing monotonous brown or dark brown. Antemedial, postmedial and terminal lines indistinct. Submarginal line slightly wavy, pale, with dark borders inwards. Claviform absent. Reniform and orbicular brown, with dark borders, reniform with shadow posteriorly. Cilia brown or dark brown. Hindwing pale, brownish-grey. Discal spot semilunar, indistinct. Cilia pale brownish-grey. Male genitalia (Figs 11, 12). Uncus narrow, medially broadened, apically pointed. Tegumen moderately long, penicular lobes narrow. Juxta rectangular, with two narrow apical processes. Vinculum long, V-shaped. Valva elongated, medially curved, with S-shaped costal margin. Cucullus small, triangular, apically rounded, with narrow, apically pointed pollex. Corona absent. Sacculus large, long, distally strongly sclerotised. Clavus small, rounded, lobe-like. Ampulla long, curved, apically pointed, strongly sclerotised. Clasper moderately long, distally strongly broadened, with small, digitus-like harpe. Aedeagus long, slightly curved. Carina with elliptical lateral bar and strong, long, narrow, acute thorn. Vesica tubular, elongated, ventro-laterally recurved, with conical, apically rounded medial diverticulum. Female genitalia (Fig. 17). Ovipositor short, conical. Apophyses posteriores and anteriores long, thin. Ostium bursae broad, trapezoidal, with sclerotised lateral crests. Ductus bursae long, anteriorly curved; its posterior part heavily sclerotised, strongly broadened; posterior part rugose. Appendix bursae moderately long, broad, twisted, apically rounded, with long sclerotised plate. Corpus bursae membranous, sack-like, with four long band-like signa. Etymology. The specific name is dedicated to László and Gábor Ronkay. Distribution and bionomics. Known only from the coast of Shiderty Reservoir in North-East Kazakhstan. The species inhabits the edges of ponds with Salix, in the steppe zone (Fig. 23). Adults were collected in April, 2012 and May, 2013, at mercury vapor light.

Orthosia (Orthosia) picata (Bang-Haas, 1912) (Figs 5���7, 13, 14, 18) Taeniocampa incerta forma picata Bang-Haas, 1912, Deutsche Entomologische Zeitschrift Iris 26: 156 (Type locality: [China, Xinjiang, SE Tien Shan Mts.] ���Karagai-tau���). Taeniocampa incerta var. pallida Staudinger, 1888, nec. Haworth, 1809 (Type locality: [China, Xinjiang] Kuldja) Taeniocampa incerta var. pallidior Staudinger, 1901, nec. Staudinger, 1888 (Type locality: Fergana; Issyk-Kul). Material examined. 1 male, [SE Kazakhstan] Alma-Ata, 5.v. [19] 37, I.N. Filipjev (Coll. AVB); 2 females, SE Kazakhstan, 30 km NW Bakanas, Ili river valley, h= 380 m, 44 �� 52 '47.11"N, 75 �� 57 '41.26"E., Egorov P.V. leg. (Coll. AVB); 5 males, 5 females, 05���08.v. 2012, W Mongolia, Hovd aimak, Dzhungarian Gobi, 46 km S Altai somon, Bodonchijn-Gol river valley, h= 1270 m, 45 ˚ 46 ��� N, 92 ˚ 10 ��� E., Yakovlev R.V. leg. (Coll. AVB); 1 male, 0 7.06. 2011 SW Mongolia, Hovd aimak, 30 km S Altai somon, Bodonchijn-Gol river valley (under stream), Elkhony-Ekhen-Tal place, h= 1200 m, 45 �� 43 ��� N; 92 ��05��� E. Yakovlev R.V. leg. (Coll. AVB). Slides AV0684, AV0850, AV0854, AV0855 Volynkin (males), AV0849, AV0851, AV0852 Volynkin (females). Distribution. Tien Shan Mts., Pamir-Alai Mt. system, NW China, SW Mongolia. The species is reported here from Mongolia for the first time., Published as part of Volynkin, Anton V. & Titov, Sergey V., 2014, A new species of Orthosia Ochsenheimer, 1816 from North-East Kazakhstan (Lepidoptera, Noctuidae), pp. 494-500 in Zootaxa 3753 (5) on page 495, DOI: 10.11646/zootaxa.3753.5.7, http://zenodo.org/record/229615