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A remarkable new species of pulmonate snail was recently collected in a small Atlantic Rainforest fragment near the city of Canavieiras, state of Bahia, Brazil, an area known for a high diversity of land snails. It is described herein as Leiostracus fetidus sp. nov. and can be easily identified by its color pattern of irregular brown to black axial stripes on a white to yellow background, a reddish axial band “separating” the white peristome from the rest of the shell and a broad brown spiral band surrounding the umbilical region. Other diagnostic features include a relatively small size, a proto columellar fold and two very weak folds delimiting the basal region of the aperture. This discovery is a reminder of how little this fauna is known and also an alarm for proper conservation of these forest fragments. KEYWORDS. Atlantic Rainforest, Bulimulinae, Canavieiras, Leiostracus fetidus sp. nov., Stylommatophora. RESUMO. Uma nova especie de Leiostracus da Bahia, Brasil (Gastropoda, Pulmonata, Orthalicidae)

The type status is described of 96 taxa classified within the superfamily Orthalicoidea and present in the Mollusca collection of the Museum für Naturkunde der Humboldt-Universität zu Berlin. Lectotypes are designated for the following taxa: Orthalicus elegans Rolle, 1895; Bulimus maranhonensis Albers, 1854; Orthalicus nobilis Rolle, 1895; Orthalichus tricinctus Martens, 1893. Orthalicus sphinx tresmariae is introduced as new name for Zebra sphinx turrita Strebel, 1909, not Z. quagga turrita Strebel, 1909. The following synonyms are established: Zebra crosseifischeri Strebel, 1909 = Orthalicus princeps fischeri Martens, 1893; Orthalicus isabellinus Martens, 1873 = O. bensoni (Reeve, 1849); Zebra zoniferus naesiotes Strebel, 1909 = Orthalicus undatus (Bruguière, 1789); Porphyrobaphe (Myiorthalicus) dennisoni pallida Strebel, 1909 = Hemibulimus dennisoni (Reeve, 1848); Zebra delphinus pumilio Strebel, 1909 = Orthalicus delphinus (Strebel, 1909); Orthalicus (Laeorthalicus) reginaeformis Strebel, 1909 = Corona perversa (Swainson, 1821); Bulimus (Eurytus) corticosus Sowerby III, 1895 = Plekocheilus (Eurytus) stuebeli Martens, 1885. The taxon Bulimus (Eudioptus) psidii Martens, 1877 is now placed within the family Sagdidae, tentatively in the genus Platysuccinea. Appendices are included with an index to all the types of Orthalicoidea extant (including those listed by Köhler 2007) and a partial list of letters present in the correspondence archives.

prolatus, Anctus Simone & Casati, 2013 Anctus prolatus Simone & Casati, 2013: 151���153 (figs 35���41). Gastropoda, Odontostomidae Holotype: MZSP 112450. Paratypes 1 (3 spc): MZSP 111836. Parartype 2: MZSP 112350. Paratype 3 (2 spc): MZSP 112362. Paratype 4 (3 spc): MZSP 112372. Localities: Brazil, Piau��, Serra da Capivara, Coronel Jos�� Dias Municipality, close to S��tio do Moc�� Town, Toca de Cima dos Pil��o Cave, 8��51'47.10'' S, 42��33'26.96'' W; 1) sta. P6; all collected in 2012. Collector: R. Casati. Preservation: Dry., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on page 43, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Simone L. R. L. & Casati R. 2013. New land mollusk fauna from Serra da Capivara, Piaui, Brazil, with a new genus and five new species (Gastropoda: Orthalicoidea, Streptaxidae, Subulinidae). Zootaxa 3683 (2): 145 - 158. http: // dx. doi. org / 10.11646 / zootaxa. 3683.2.4"]}

maculata, Oxychona Salvador & Cavallari, 2013 Oxychona maculata Salvador & Cavallari, 2013a: 315–316 (figs 2–11). Gastropoda, Orthalicidae Holotype: MZSP 108005. Paratype 1: MZSP 74500. Localities: Brazil, Bahia, Ilhéus Municipality, 14°47' S, 39°2' W, Dec. 1994; 1) Itapetinga Municipality, 15°15' S, 40°15' W, Jan. 2007. Collectors: A. Bodart; 1) C. Lima. Preservation: Dry.

capivara, Cyclodontina Simone & Casati, 2013 Cyclodontina capivara Simone & Casati, 2013: 149���151 (figs 16���22). Gastropoda, Odontostomidae Holotype: MZSP 112448. Paratypes 1 (19 spc): MZSP 111837. Paratypes 2 (18 spc): MZSP 111838. Paratypes 3 (9 spc): MZSP 111845. Localities: Brazil, Piau��, Serra da Capivara, Coronel Jos�� Dias Municipality, close to S��tio do Moc�� town, Toca de Cima dos Pil��o Cave, 8��51'47.10'' S, 42��33'26.96'' W; 1���2) same, sta. P1; all collected in 2012; 3) from type locality. Collector: R. Casati. Preservation: Dry. Remarks: Specimens in the lot MZSP 111835 were listed as paratypes of Rhinus gilbertus by Simone & Casati (2013, original description), but are in fact topotypes of C. capivara, which is described in the same paper. The lot MZSP 111845 contains 9 shells and not 8 as reported in the original description of C. capivara. Simone & Casati (2013) simultaneously reported the catalogue number MZSP 111845 among the paratypes of both C. capivara and Rhinus gilbertus. This catalogue number does not contain paratype specimens of R. gilbertus: this information is incorrect in the original description and should be disregarded., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on page 13, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Simone L. R. L. & Casati R. 2013. New land mollusk fauna from Serra da Capivara, Piaui, Brazil, with a new genus and five new species (Gastropoda: Orthalicoidea, Streptaxidae, Subulinidae). Zootaxa 3683 (2): 145 - 158. http: // dx. doi. org / 10.11646 / zootaxa. 3683.2.4"]}

tessa, Anostoma Simone, 2012 Anostoma tessa Simone, 2012b: 435���436 (figs 15���20). Gastropoda, Orthalicidae Holotype: MZSP 103914. Paratype: MZSP 103915. Paratypes (37 spc): MZSP 103916. Locality: Brazil, Bahia, Santa Maria da Vit��ria, ~ 13��24' S, 44��12' W, ~ 460 m altitude (type locality), Jan. 2012. Collector: J. Coltro. Preservation: Dry., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on page 49, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Simone L. R. L. 2012 b. Taxonomical study on a sample of pulmonates from Santa Maria da Vitoria, Bahia, Brazil, with description of a new genus and four new species (Mollusca: Orthalicidae and Megalobulimidae). Papeis Avulsos de Zoologia 52 (36): 431 - 439."]}

gilbertus, Rhinus Simone & Casati, 2013 Rhinus gilbertus Simone & Casati, 2013: 153���154 (figs 23���34). Gastropoda, Simpulopsidae Holotype: MZSP 112449. Paratypes 1 (27 spc): MZSP 111827. Paratypes 2 (50 spc): MZSP 111839. Paratypes 3 (9 spc): MZSP 111844. Localities: Brazil, Piau��, Serra da Capivara, Coronel Jos�� Dias Municipality, close to S��tio do Moc�� Town, Toca de Cima dos Pil��o Cave, 8��51'47.10'' S, 42��33'26.96'' W; 1���2) same, sta. P2; 3) same, sta. P1. Collector: R. Casati. Preservation: Dry. Remarks: Specimens in the lot MZSP 111835 are listed as paratypes of Rhinus gilbertus by Simone & Casati (2013, original description), but are in fact topotypes of C. capivara, which is described in the same paper. Simone & Casati (2013) simultaneously reported the catalogue number MZSP 111845 among the paratypes of Rhinus gilbertus and C. capivara. This catalogue number does not contain paratypes of R. gilbertus; this information is incorrect in the original description and should be disregarded., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on page 25, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Simone L. R. L. & Casati R. 2013. New land mollusk fauna from Serra da Capivara, Piaui, Brazil, with a new genus and five new species (Gastropoda: Orthalicoidea, Streptaxidae, Subulinidae). Zootaxa 3683 (2): 145 - 158. http: // dx. doi. org / 10.11646 / zootaxa. 3683.2.4"]}

breweri, Plekocheilus Breure & Schl��gl, 2010 Plekocheilus (Eurytus) breweri Breure & Schl��gl, 2010: 57���58 (figs 2D���2G, 3C���D, 4B���4F). Gastropoda, Orthalicidae Paratypes (2 spc): MZSP 92471. Locality: Venezuela, Bol��var, Chimant�� Massif, Cur��-tepui, 05��16'44.9" N, 62��00'56.35" W, 2405 m depth (type locality), Feb. 2009. Collector: J. Sch��gl. Preservation: Dry., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on pages 12-13, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Breure A. S. H. & Schlogl J. 2010. Additional notes on Orthalicidae from Chimanta massif, Venelzuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828 (Mollusca: Gastropoda). Zootaxa 2416: 51 - 60."]}

vlceki, Plekocheilus Breure & Schl��gl, 2010 Plekocheilus (Plekocheilus) vlceki Breure & Schl��gl, 2010: 52���54 (figs 2A���C, 3A, 3B, 4A). Gastropoda, Orthalicidae Paratype: MZSP 92470. Locality: Venezuela, Bolivar State, Chimant�� Massif, Chur��-tepui, Sima Northwest, 2100 m altitude (type locality), Feb. 2009. Collector: J. Schl��gl. Preservation: Dry., Published as part of Cavallari, Daniel C., Dornellas, Ana Paula S. & Simone, Luiz Ricardo L., 2016, Second annotated list of type specimens of molluscs deposited in the Museu de Zoologia da Universidade de S��o Paulo, Brazil, pp. 1-59 in European Journal of Taxonomy 213 on page 52, DOI: 10.5852/ejt.2016.213, http://zenodo.org/record/3840125, {"references":["Breure A. S. H. & Schlogl J. 2010. Additional notes on Orthalicidae from Chimanta massif, Venelzuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828 (Mollusca: Gastropoda). Zootaxa 2416: 51 - 60."]}

An alphabetical list of 352 type lots of molluscs housed in the Museu de Zoologia da Universidade de São Paulo is presented following the standards of the previous list by Dornellas & Simone (2011), with a few adjustments. Important items listed herein include types of species described after the previous compilation, as well as recently acquired paratypes of Asian Pomatiopsidae and Diplommatinidae (Gastropoda) taxa described by Rolf A.M. Brandt (1960s), P. Temcharoen (1970s) and W.J.M. Maassen (2000s), all of which belonged to the private collection of Jens Hemmen, Wiesbaden, Germany. Relevant items also include types of recently described species coming from the French-Brazilian Marion Dufresne MD55 expedition, and other types deposited by researchers from Brazil and the world. A list of authors and photographs of specimens are also provided.

Liguus crenatus barbouri Clench, 1929 Fig. 134 Liguus crenatus barbouri Clench, 1929: 18. – Neubert & Janssen, 2004: 236, pl. 24 fig. 295. – Breure, 2013: 12, figs 19C, 19ii. Type locality: “ Pinecrest region, central Everglades, Fla. Hammock no. 21 (Farnum number). J.N. Farnum, collector”. Label: “Hammock No. 7 / Everglades Pine / Crest region / Florida / J.N. Farnum leg.”. Dimensions: “Length 51.5 Width 27 (...) mm.”; figured specimen herein H 43.2, D 22.8, W 6.9. Type material: MHNG-INVE-64938, two paratypes (ex MCZ). Remarks: In original publication only the holotype is mentioned; MCZ 84527. The type status of this material is, however, not disputed. See also Neubert & Janssen, 2004: 236, who accept Clench’s statement “found in nearly all the hammocks of the Pinecrest region” as sufficient statement to warrant the status of paratypes. Current systematic position: Orthalicidae, Liguus faciatus (Müller, 1774)., Published as part of Breure, Abraham S. H., 2016, Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Muséum d'histoire naturelle, Geneva, pp. 57-103 in Revue suisse de Zoologie 123 (1) on page 64, DOI: 10.5281/zenodo.46290, {"references":["Clench W. J. 1929. Some new Liguus from the Florida Everglades. The Nautilus 43: 18 - 21.","Neubert E. & Janssen R. 2004. Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv fur Molluskenkunde 133: 193 - 297.","Breure A. S. H. 2013. Annotated type catalogue of Orthalicoidea (Mollusca, Gastropoda) in the Museum fur Naturkunde, Berlin. ZooKeys 279: 1 - 101."]}

Liguus fasciatus viridis Clench, 1934 Fig. 133 Liguus fasciatus viridis Clench, 1934: 105, pl. 6 fig. 11. – Johnson, 2003: 25. Type locality: “La Caoba, near Dolores, Central Soledad, Cienfuegos, Santa Clara, Cuba ”. Label: “ 2 miles E of Dolores / Soledad, Cienfuegos / Cuba ”. Dimensions: “Length 48.0 / Width 23.0 (…) mm.”; figured specimen herein H 55.8, D 26.7, W 7+. Type material: MHNG-INVE-64933, one paratype (ex Fairchild, ex MCZ). Remarks: Clench presented dimensions of the holotype and five paratypes (in the MCZ collection), but did not mention how many specimens were in the type series. However, besides the type locality, he mentioned eight localities under paratypes, “[a]ll (…) localities in the vicinity of Central Soledad”. Current systematic position: Orthalicidae, Liguus fasciatus (Müller, 1774)., Published as part of Breure, Abraham S. H., 2016, Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Muséum d'histoire naturelle, Geneva, pp. 57-103 in Revue suisse de Zoologie 123 (1) on page 86, DOI: 10.5281/zenodo.46290, {"references":["Clench W. J. 1934. New mollusks in the genus Liguus from Cuba and the Isles of Pines. West Indian mollusks, 8. Occasional Papers of the Boston Society of Natural History 8: 101 - 124.","Johnson R. I. 2003. Molluscan taxa and bibliographies of William James Clench and Ruth Dixon Turner. Bulletin of the Museum of Comparative Zoology 158: 1 - 46."]}

Liguus fasciatus archeri Clench, 1934 Fig. 139 Liguus fasciatus archeri Clench, 1934: 106, pl. 7 fig. 5. – Johnson, 2003: 4. – Neubert & Janssen, 2004: 236, pl. 24 fig. 291. – Breure, 2013: 10, figs 19B, 19i. Type locality: “ Mogote de Ramon Millo, Viñales, Pinar del Rio, Cuba ”. Label: “Viñales, Pinar / del Rio, Cuba / A. Archer leg.”. Dimensions: “Length 55.5 / Width 25.0 (…) mm.”; figured specimen herein H 54.5, D 24.5, W 6+. Type material: MHNG-INVE-64921, two paratypes (ex MCZ). Remarks: The holotype is MCZ 80901. The data of the specimens correspond to the original publication. The current systematic position is after Richardson (1993). Current systematic position: Orthalicidae, Liguus fasciatus (Müller, 1774)., Published as part of Breure, Abraham S. H., 2016, Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Muséum d'histoire naturelle, Geneva, pp. 57-103 in Revue suisse de Zoologie 123 (1) on page 63, DOI: 10.5281/zenodo.46290, {"references":["Clench W. J. 1934. New mollusks in the genus Liguus from Cuba and the Isles of Pines. West Indian mollusks, 8. Occasional Papers of the Boston Society of Natural History 8: 101 - 124.","Johnson R. I. 2003. Molluscan taxa and bibliographies of William James Clench and Ruth Dixon Turner. Bulletin of the Museum of Comparative Zoology 158: 1 - 46.","Neubert E. & Janssen R. 2004. Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv fur Molluskenkunde 133: 193 - 297.","Breure A. S. H. 2013. Annotated type catalogue of Orthalicoidea (Mollusca, Gastropoda) in the Museum fur Naturkunde, Berlin. ZooKeys 279: 1 - 101.","Richardson L. 1993. Bulimulacea: catalog of species. Amphibulimidae, Anadromidae, Grangerellidae, Odontostomidae, Orthalicidae. Tryonia 27: 1 - 164."]}

Fueron revisadas las colecciones malacológicas históricas “Cleto Sánchez Falcón” y “M. L. Jaume”, depositadas en el Museo de Historia Natural “Tomás Romay” y el Museo “Jorge Ramón Cuevas”. De ambas colecciones se copiaron los datos de etiqueta del material tipo. La validez de la información de etiqueta para cada lote fue revisada con las descripciones originales correspondientes a cada especie, revisiones taxonómicas de familias y catálogos actualizados. Se registraron 434 ejemplares, incluidos en 56 subespecies, 34 especies y seis (6) familias; estos se corresponden con 85 localidades y 16 colectores. La colección “Cleto Sánchez Falcón” posee 368 ejemplares de las familias Annulariidae, Cerionidae, Megalomastomidae, Helicinidae, Orthalicidae y Urocoptidae, siendo esta última la más representada. La colección “M. L. Jaume” tiene 66 ejemplares de 36 subespecies de Liguus fasciatus (Müller), Orthalicidae.

The type status is described of 57 taxa from the superfamily Orthalicoidea in the collection of the Brussels museum. Two new species are described: Stenostylus perturbatus sp. n., and Suniellus adriani sp. n. New lectotypes are designated for Bulimulus (Naesiotus) amastroides Ancey, 1887; Bulimulus blanfordianus Ancey, 1903; Bulimulus montivagus chacoensis Ancey, 1897; Bulimus coloratus Nyst, 1845; Plecochilus dalmasi Dautzenberg, 1900; Placostylus porphyrostomus elata Dautzenberg, 1923; Bulimulus ephippium Ancey, 1904; Bulimus fulminans Nyst, 1843; Bulimus funckii Nyst, 1843; Orphnus thompsoni lutea Cousin, 1887; Bulimus melanocheilus Nyst, 1845; Orphnus thompsoni nigricans Cousin, 1887; Orphnus thompsoni olivacea Cousin, 1887; Bulimulus pollonerae Ancey, 1897; Orphnus thompsoni zebra Cousin, 1887. New combinations are: Bostryx borellii (Ancey, 1897); Bostryx carandaitiensis (Preston, 1907); Protoglyptus mazei (Crosse, 1874); Kuschelenia (Vermiculatus) sanborni (Haas, 1947). New synonymies are established for the following nominal taxa: Orphnus thompsoni var. lutea Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Orphnus thompsoni var. nigricans Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Thaumastus nystianus var. nigricans Cousin, 1887 = Drymaeus (Drymaeus) nystianus (Pfeiffer, 1853); Orphnus thompsoni var. olivacea Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845); Orphnus thompsoni var. zebra Cousin, 1887 = Kara thompsonii (Pfeiffer, 1845).

Two new sibling species are described, Bostryx multiconspectus spec. nov and B. primigenius spec. nov., one of which is carinate. From the carinate taxa known in the Orthalicidae, 16 are occurring at localities in the Andes from northern Peru to northwestern Argentina. They belong to three genera, Bostryx, Neopetraeus and Scutalus. It is hypothesized that carination originated independently in these lineages.

A faunal overview is presented of the molluscan families Amphibulimidae, Megaspiridae, Odontostomidae, Orthalicidae, Simpulopsidae in Bolivia, Ecuador, and Peru. These Central Andean countries are known for their biodiverse malacofauna, of which the superfamily Orthalicoidea takes relatively a large share. In this paper the five families containing 103 (sub)species, for which systematic information (original publication, type locality, type depository, summarizing literature) and distributional records are presented. All species are illustrated by photographs of the type material or, if this could not be located, by a reproduction of the original figure. The following new taxon is introduced: Thaumastus (Thaumastus) sumaqwayqu sp. n. Junior subjective synonyms are established for: Plekocheilus (Sparnotion) Pilsbry, 1944 = Plekocheilus (Eudolichotis) Pilsbry, 1896; Scholvienia (Thomsenia) Strebel, 1910 = Scholvienia Strebel, 1910; Sultana (Trachyorthalicus) Strebel, 1909 = Sultana (Metorthalicus) Pilsbry, 1899; Plekocheilus (Eurytus) conspicuus Pilsbry, 1932 = Thaumastus (Thaumastus) hartwegi (Pfeiffer in Philippi, 1846); Zebra gruneri Strebel, 1909 = Orthalicus maracaibensis (Pfeiffer, 1856); Scholvienia jaspidea minor Strebel, 1910 = Scholvienia alutacea (Reeve, 1850); Bulimus bifasciatus unicolor Philippi, 1869 = Scholvienia brephoides (d'Orbigny, 1835). A new status is given to Plekocheilus mcgintyi 'Pilsbry' H.B. Baker, 1963 (subspecies of Bulinus piperitus Sowerby I, 1837); Strophocheilus superstriatus var. prodeflexus Pilsbry, 1895 (subspecies of Bulinus piperitus Sowerby I, 1837); Thaumastus (Quechua) salteri maximus Weyrauch, 1967 (subspecies of Thaumastus (Quechua) olmosensis Zilch, 1954); Pseudoglandina agitata Weyrauch, 1967 (nomen inquirendum). New combinations are: Clathrorthalicus corydon (Crosse, 1869), and Cyclodontina chuquisacana (Marshall, 1930). Lectotypes are now designated for Bulimus incisus Hupé, 1857 and Bulinus piperitus Sowerby I, 1837.

We have incorporated an additional 56 species of land snails and slugs in our ribosomal (r) RNA molecular phylogeny. The new taxa include representatives of several important groups. The molecular tree now includes 160 species of stylommatophoran land snails and slugs in 144 genera in 61 families. In the rDNA tree, the Stylommatophora are principally divided into an ‘achatinoid’ and a ‘non-achatinoid’ clade. Within these clades, several major land snail groups, including the Orthurethra, Elasmognatha, Limacoidea, and Helicoidea, are supported. Overall, the rDNA molecular phylogeny has remained stable following the incorporation of the additional taxa, with these additions having little impact on the major evolutionary patterns in the tree. Taxonomic coverage of the Orthurethra, Orthalicidae, Camaenidae, and Bradybaenidae is increased significantly. The camaenids and bradybaenids form a complex, and both appear to be paraphyletic. Several families of uncertain affinity, such as the Sagdidae and Thyrophorellidae, are included for the first time. The Sagdidae are shown to belong to the Helicoidea, and the Thyrophorellidae to the Achatinoidea. © 2006 The Linnean Society of London, Biological Journal of the Linnean Society, 2006, 87, 593–610.

Plagiodontes patagonicus (d'Orbigny, 1835) is a quite variable species endemic to southern Buenos Aires province (Argentina). Its taxonomic characterization did not include any quantitative analysis of variability for an accurate discrimination, and the extremes of its variation were described as different subspecies or species. In this paper, shell measurements and angles, and quantitative data on the terminal male genitalia were studied by Principal Component Analysis and Varimax multivariate analysis. Typical Plagiodontes patagonicus and the largest form known as P. patagonicus magnus Hylton-Scott, 1951 showed an almost continuous pattern of shell variation, which is positively correlated with the altitude gradient over their geographical range, which in turn is correlated with a rainfall gradient, i.e. they constitute a size-form cline that does not allow objective delimitation of different morphospecies. Data from the genital system were also arranged as a geographical gradient within the P. patagonicus patagonicus-P. patagonicus magnus continuum. The variability of protoconch sculpture and apertural teeth also indicate recognition of them as a single taxon. Fil: Cazzaniga, Néstor Jorge. Universidad Nacional del Sur. Departamento de Biología, Bioquímica y Farmacia; Argentina Fil: Piza, Julia. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Bahía Blanca; Argentina. Universidad Nacional del Sur. Departamento de Biología, Bioquímica y Farmacia; Argentina Fil: Ghezzi, Natalia S.. Universidad Nacional del Sur. Departamento de Biología, Bioquímica y Farmacia; Argentina

Thaumastus caetensis Pena, Salgado and Coelho, 2011 is a recently described species from Serra da Piedade (19°49’19” S, 43°40’46” W), Caeté municipality, Minas Gerais, southeastern Brazil. We present a new record of this species based on a young specimen collected in the highlands of Reserva Particular do Patrimônio Natural Santuário do Caraça, at Serra do Caraça (20°08’04” S, 43°27’09” W). This species is probably restricted to the “campos rupestres”, above 1,400 m above sea level and the new record represents the only known protected occurrence within the species’ range.

Anctus prolatus new species (Figs. 35–41) Types. Holotype MZSP 112450. Paratypes MZSP 111836, 3 shells (sta. P 6), MZSP 112350, 1 young shell, 112362, 2 shells, 112372, 3 shells; all from type locality. Type locality. BRAZIL. Piauí; Serra da Capivara; Coronel José Dias municipality, close to Sítio do Mocó town, Cave Toca de Cima dos Pilão, 8 ° 51 ’47.10”S 42 ° 33 ’26.96”W (R. Casati col, 2012). Diagnosis. Shell elongated, with width/length ratio of ~ 39 %. Axial undulations relatively strong, almost forming sculpture. Peristome elongated, rectangular, lacking teeth. Umbilicus occluded by planar surface of inner lip. Description. Shell of about 25 mm. Color cream, with sparse axial, pale brown bands. Periostracum not preserved. Outline bulimuloid; width ~ 39 % of length. Protoconch of 2 whorls, opaque, conical, suture shallow; smooth, pigmented of dark brown (Figs. 39–40); occupying ~ 3 % of shell length and ~ 12 % of shell width; border protoconch-teleoconch unclear, orthocline. Spire of ~ 7 flattened whorls, tall, being wider in last whorl (Figs. 35, 40); angle ~ 40 °; suture shallow, weakly marked. Sculpture uniform series of axial, narrow undulations, from suture to suture, ~ 40 in penultimate whorl; interspaces between undulations equivalent to their width. Peristome deflected outwards, expanding ~ 10 % width of preceding whorl; aperture orthocline (Figs. 36, 38, 41). Aperture narrow, rectangular, ~ 4 times longer than wide (Figs. 35, 40, 41), occupying ~ 46 % of shell length and ~ 35 % of width; lacking teeth except for narrow thickness in inferior half of outer lip (Figs. 40–41); outer lip mostly straight, with small rounded portions only in both ends; anterior region as narrow spatula; inner almost straight, weakly concave, superior third oblique and weakly convex, inferior 2 / 3 almost straight and narrow, as tube surrounding umbilicus. Callus thick, planar, simple (Figs. 35, 38, 40). Umbilicus occluded by planar surface of inner lip, somewhat projected ventrally (Figs. 35, 38, 41). Measurements (in mm): Holotype (Fig. 35–39): 25.2 by 10.1; paratype MZSP 111836: 20.1 by 9.3; MZSP 111836 (Figs. 40–41): 24.1 by 10.8. Distribution. Only known from type locality. Habitat. Caatinga semi-dry environment; collected inside caves. Material examined. Types. Non-type material from type locality MZSP: 112403, 10 shells, 112422, 1 shell, 112439, 1 shell. Etymology. The specific epithet is derived from Latin prolatus, meaning extended, elongated, an allusion to the elongation of the shell of compared with other congener species. Systematic remarks. The generic attribution to Anctus prolatus is based on the dark-brown protoconch, the axial sparse brown bands, the elliptical shape and the antero-posteriorly elongated peristome. The genus has only 3 species (Breure, 1974; Simone, 2006). Anctus prolatus differs from A. angiostomus (Wagner, 1827), from Bahia, in lacking a developed palatalis tooth in middle region of the outer lip, by the narrower umbilicus and by the slender outline (width/length ratio ~ 39 %, against ~ 46 % of A. angiostomus). It differs from A. laminiferus (Ancey, 1888), also from Bahia, in having a much narrower peristome, in having a wider umbilicus, and a narrower outline (width/ length ratio of A. laminiferus is ~ 47). It differs from A. surgillatus (Pfeiffer, 1857), from Bolivia, in having more elongated aperture (46 % of shell length, against ~ 37 of A. surgillatus), and much narrower peristome (width/length ratio ~ 25 % against 48 % of A. surgillatus). The elongation of the aperture of most Anctus species is for fixing and enchasing the shell to dry branches of plants (Simone, 1998). This is a behavior that can be extrapolated to A. prolatus. Anctus prolatus, which expands the occurrence of the genus northwards in Brazilian territory, and is the first occurrence outside from Bahia state., Published as part of Simone, Luiz Ricardo L. & Casati, Rafael, 2013, New land mollusk fauna from Serra da Capivara, Piauí, Brazil, with a new genus and five new species (Gastropoda: Orthalicoidea, Streptaxidae, Subulinidae), pp. 145-158 in Zootaxa 3683 (2) on pages 151-153, DOI: 10.11646/zootaxa.3683.2.4, http://zenodo.org/record/218653, {"references":["Breure, A. S. H. (1974) Catalogue of Bulimulidae (Gastropoda, Euthyneura) II. Odontostomidae. Basteria, 38, 109 - 127.","Simone, L. R. L. (2006) Land and freshwater molluscs of Brazil. EGB. Fapesp. Sao Paulo, 390 pp.","Simone, L. R. L. (1998) Anatomical description of Anctus angiostomus (Wagner, 1827) from northeastern Bahia, Brazil (Gastropoda, Pulmonata, Bulimulidae). Studies on Neotropical Fauna and Environment, 33, 170 - 177. http: // dx. doi. org / 10.1076 / snfe. 33.2.170.2160"]}

Rhinus gilbertus new species (Figs. 23–34) Types. Holotype MZSP 112449 (Figs. 24–28). Paratypes MZSP 111827, 27 shells (sta. P 2), MZSP 111835, 98 shells (sta. P 2), MZSP 111839, 50 shells (sta. P 2), MZSP 111844, 9 shells (sta. P 1) (Figs. 30–31), MZSP 111845, 9 shells (Figs. 32–33), USNM, 2 shells; MNRJ, 2 shells. All from type locality. Type locality. BRAZIL. Piauí; Serra da Capivara; Coronel José Dias municipality, close to Sítio do Mocó town, Cave Toca de Cima dos Pilão, 8 ° 51 ’47.10”S 42 ° 33 ’26.96”W (R. Casati col, 2012). Diagnosis. Shell somewhat elongated, with width/length ratio of ~ 40 %. Axial undulations relatively strong, almost forming sculpture. Peristome occupying ~ 30 % of shell length, strongly deflected. Umbilicus opened, narrow. Description. Shell of about 20 mm. Color pure white to pale brown. Periostracum not preserved. Outline somewhat turriform to bulimuloid; width ~ 40 % of length. Protoconch of 1.5 whorls, opaque, obese, suture wellmarked; sculptured by delicate, uniform, undulating striae (Figs. 23, 29); occupying ~ 5 % of shell length and ~ 25 % of shell width; border protoconch-teleoconch clear, slightly prosocline (angle ~ 10 ° with longitudinal axis) (Figs. 28, 31, 33). Spire of ~ 6 rounded whorls, tall, being wider in penultimate whorl; angle ~ 50 ° (Figs. 24–26, 29, 30– 33); suture well-marked. Sculpture uniform series of axial, narrow undulations, from suture to suture, ~ 60 in penultimate whorl; interspaces between undulations equivalent to their width. Body whorl equal (Figs. 30, 31) to ~ 5 % narrower (Figs. 24, 26, 32, 33) than preceding whorl. Peristome deflected outwards, expanding ~ 10 % width of preceding whorl; aperture orthocline (Figs. 25, 31, 33). Aperture elliptical, occupying ~ 30 % of shell length and ~ 50 % of width (Figs. 24, 30, 32); lacking teeth; outer lip rounded, with middle region slightly straight; inner lip highly concave, superior half almost straight, inferior half also almost straight and wide, as tube surrounding umbilicus; low, blunt columellar carina somewhat distinct (Fig. 25). Callus thin, planar, simple, narrowly attached to penultimate whorl (Figs. 24, 25, 30, 32). Umbilicus opened, narrow, partially occluded by inner lip (Fig. 27, 34). Measurements (in mm): Holotype MZSP 112449: 21.9 by 8.3 (Figs. 24–28); paratypes: 111844 (Figs. 30–31): 18.7 by 7.5; MZSP 111845 (Figs. 32–33): 18.7 by 8.1. Distribution. Only known from type locality. Habitat. Caatinga semi-dry environment; collected inside caves. Material examined. Types. Non-type material from type locality MZSP: 112347, 10 shells, 112356, 10 shells, 112357, 9 shells, 112367, 4 shells, 112368, 7 shells, 112373, 7 shells, 112399, 15 shells, 112415, 25 shells, 112420, 15 shells, 112432, 40 shells, 112437, 20 shells, 112442, 18 shells. Etymology. The specific epithet refers to the father of junior author, Gilberto Casati, who provided financial support and encouragement from his academic affairs. Systematic remarks. The main character of the genus Rhinus is the hairy periostracum, a character that cannot be seen in the examined material, composed of mostly long dead shells. However, the resemblance of R. gilbertus with several species of this genus suggests the generic attribute. Another character suggestive of the genus is the striated protoconch (Figs. 23, 29). Only few of the Rhinus that have elongated shell; most has a more globose shape. The elongated-shaped Rhinus (Simone, 2006), R. gilbertus resembles R. constrictus (Pfeiffer, 1841), from NE Brazil, differing in being more slender (width/length ratio of ~ 40 %, against ~ 62 % of that species); it also resembles R. evelinae Leme, 1986, from São Paulo, and R. obeliscus Haas, 1936, from Santa Catarina. It differs in having a slightly more rounded outline, as that two species are still more elongated (ratio of both species ~ 35 %), and two fewer whorls. R. gilbertus is the most elongated Rhinus from the NE Brazil, as elongated shape is more common in SE Brazilian species. A young specimen is shown in Fig. 29, displaying a different shape., Published as part of Simone, Luiz Ricardo L. & Casati, Rafael, 2013, New land mollusk fauna from Serra da Capivara, Piauí, Brazil, with a new genus and five new species (Gastropoda: Orthalicoidea, Streptaxidae, Subulinidae), pp. 145-158 in Zootaxa 3683 (2) on pages 153-154, DOI: 10.11646/zootaxa.3683.2.4, http://zenodo.org/record/218653, {"references":["Simone, L. R. L. (2006) Land and freshwater molluscs of Brazil. EGB. Fapesp. Sao Paulo, 390 pp."]}

Cyclodontina capivara new species (Figs. 16–22) Types. Holotype MZSP 112448 (Figs. 16–18). Paratypes MZSP 111837, 19 shells (sta. P 1), MZSP 111838, 18 shells (sta. P 1), MZSP 111845, 8 shells (Figs. 19–20), USNM, 2 shells; MNRJ, 2 shells. All from type locality. Type locality. BRAZIL. Piauí; Serra da Capivara; Coronel José Dias municipality, close to Sítio do Mocó town, Cave Toca de Cima dos Pilão, 8 ° 51 ’47.10”S 42 ° 33 ’26.96”W (R. Casati col, 2012). Diagnosis. Shell well sculptured axially, with ribs uniform and well-spaced; apex slightly pointed. Peristome with 6 somewhat equidistant, well-developed teeth, occupying ~ 70–76 % of shell length. Description. Shell of about 20 mm. Color pure white. Periostracum wanting. Outline slightly turriform; width ~ 30 % of length. Protoconch of 2 whorls, opaque, tall, suture well-marked; sculptured by delicate and uniform reticulation (Figs. 21, 22); occupying ~ 6 % of shell length and ~ 26 % of shell width; border protoconch-teleoconch clear, slightly prosocline. Spire of ~ 8 whorls, tall, being wider in two penultimate whorls; angle ~ 25 (Fig. 19) to 35 ° (Fig. 20); suture shallow but well-marked. Sculpture uniform series of axial, slightly prosocline, narrow ribs, from suture to suture, ~ 30 in penultimate whorl; interspaces between ribs ~ 5 times their width. Sculpture absent in inferior half of body whorl in young specimens (Fig. 21), with low carina in periphery. Body whorl varying from equal (Figs. 19, 20) to ~ 10 % narrower (Figs. 16–18) than two preceding whorls. Three spiral wide grooves corresponding to palatal tooth in ¼ whorl preceding peristome (Figs. 17, 18). Peristome entirely deflected outwards, expanding ~ 20 % width of preceding whorl; aperture slightly prosocline, ~ 10 ° related to longitudinal shell axis. Aperture oval, occupying ~ 20 % of shell length and ~ 70–76 % of width (Figs. 16, 19, 20); greatly occluded by series of blunt and tall teeth (Figs. 16, 19, 20), teeth fold-like, disposed rather perpendicularly to peristome; as follows: 2 teeth on inner lip, located somewhat equidistant from each other on inner lip, columellaris (inferior) tooth ~ 8 times longer than wide, almost straight, edge broader than base, disposed obliquely (distal end more anterior than posterior end), parietalis (superior) tooth about as long as inferior tooth, but slightly thinner and more perpendicularly organized; four palatal tooth, equidistantly located on outer lip, all them ~ 4 times longer than wide, edges slightly broader than base, palatalis superior (second tooth) with ~double height than remaining teeth; basalis tooth as smallest. Callus thick but planar, simple, widely attached to penultimate whorl (Figs. 16, 19); fecal and urinary furrows narrow and shallow, but clearly visible in both sides of callus (Fig. 16). Umbilicus wanting. Measurements (in mm): Holotype (Figs. 16–18): 19.2 by 5.9; paratypes MZSP 111845 # 1 (Fig. 19): 20.4 by 5.8; # 2 (Fig. 20): 17.0 by 5.5. Distribution. Only known from type locality. Habitat. Caatinga semi-dry environment; collected inside caves. Material examined. Types. Non-type material from type locality MZSP: 112348, 10 shells, 112352, 12 shells, 112359, 11 shells, 112336, 9 shells, 112370, 7 shells, 112374, 8 shells, 112401, 35 shells, 112414, 40 shells, 112417, 40 shells, 112430, 50 shells, 112434, 20 shells, 112441, 20 shells. Etymology. The specific epithet refers to the local of occurrence, the Serra da Capivara, being attributed in apposition. Systematic remarks. Cyclodontina capivara has resemblance to some congeneric species that possess welldeveloped axial sculpture, which are only a small part of the Cyclodontina species (Simone, 2006). It differs from C. cylindrica (Baker, 1913), from Ceará to Pernambuco, in having more developed teeth in peristome, mainly palatal ones, a more slender outline and axial ribs more spaced. It differs from C. gemellata (Ancey, 1901), from Goiás, in being much slender, in having a more pointed spire and by the aperture being much smaller. It differs from C. scabrella (Anthony & Dohn, 1882), from Paraíba, in having a slender outline, less regular axial ribs, and the aperture is narrower. Cyclodontina capivara also resembles Clessinia costata (Pfeiffer, 1848), from Minas Gerais, differing in having more developed teeth in peristome, more pointed spire and less-spaced axial ribs. There is no other species in such C. capivara can be confused. Young specimens of C. capivara are rare. They clearly show that the characteristic sculpture is restricted to the superior half of the teleoconch whorls (Fig. 21), and that the inferior half (exposed only in the body whorl) is smooth.

Uma amostra de Pulmonata coletada em Santa Maria da Vitória, interior da Bahia, Brasil, na Caatinga semiárida, é estudada taxonomicamente. Das 5 espécies, 4 revelaram-se novas, incluindo um gênero novo. Os táxons novos dão os Bulimulidae (1) Kora corallina gen. et sp. n. caracterizado pela concha alongada com apertura algo deslocada do eixo da concha e um dente oblíquo no nível médio do lábio interno; (2) Spixia coltrorum, principalmente caracterizado pelas voltas desiguais da espira, escultura delicada e perístoma com 4 dentes equidistantes; (3) Anostoma tessa, principalmente caracterizado pela espira bojuda e um canal anal bem desenvolvido; e pelo Megalobulimidae (4) Megalobulimus amandus, principalmente caracterizado pela protoconcha pontuda esculturada por uma densa quantidade de cordas axiais. Rhinys suturalis é a punica espécie previamente conhecida, mas sua distribuição geográfica é expandida para o sul até a Bahia. Uma discussão a respeito da necessidade de melhoria do estudo sobre a malacofauna do interior do Nordeste e a importância da preservação do bioma Caatinga é também fornecida. A sample of Pulmonata collected in Santa Maria da Vitória, interior of Bahia, Brazil, in Caatinga semi-arid environment, is studied taxonomically. From the five species, four are revealed as new, including a new genus. The new taxa are the Bulimulidae (1) Kora corallina gen. et sp. n. characterized by the elongated shell with aperture somewhat dislocated from the shell axis, and an oblique tooth in middle level of inner lip; (2) Spixia coltrorum, mainly characterized by an uneven spire, delicate sculpture and peristome with 4 equidistant teeth; (3) Anostoma tessa, mainly characterized by a broad spire and well-developed anal canal; and the Megalobulimidae (4) Megalobulimus amandus, mainly characterized by pointed protoconch sculptured by dense quantity of axial cords. Rhinus suturalis is the only previously known species, but its geographic distribution is expanded southwards to Bahia state. A discussion with respect to necessity for improving the study on the malacofauna from the interior region of the Brazilian Northeast and the importance for preservation of the Caatinga biome is also provided.

Plekocheilus (Eurytus) camaritagua spec. nov. (Figs 14 D, 18 A���B) Diagnosis. Shell globose, with flat whorls and hardly to not impressed suture, that descends slightly in front. Color pattern with dark brown, oblique streaks, bordered with a narrow, yellowish ���shadow.��� Sculptured with a fine granulation arranged in spirals. Description. Shell 57 mm, 1.5 times longer than wide, imperforate, sides hardly convex, rather solid. Color pale yellowish-brown with darker lines of the same color along the growth striae, crossed by reddish-brown, oblique streaks, showing some zigzags especially on the upper part of the last whorl, and bordered by a narrow, yellowish ���shadow;��� upper whorls lighter, eroded. Surface not shining, upper whorls with growth striae, on the last whorls with spiral lines of dot-like granulation, especially visible on the dorsal side of the last whorl. Protoconch eroded. Whorls 3.9, very flat, the last whorl approx. 0.9 times total height, suture not or hardly impressed, slightly descending in front. Aperture ovate, somewhat ear-shaped, 1.8 times longer than wide, 0.72 times the total length; peristome thin and hardly reflexed, pinkish (faded). Columellar margin straight, entering the aperture with a weak fold, hardly dilated and somewhat receding. Parietal wall with a thin callus. Dimensions in mm: Holotype H 57, D 37.5, HA 41, WA 23.5, LW 52, 3.9 whorls. Type locality. Colombia, Amazonas, Corregimiento La Pedrera [1 �� 20 ��S 69 �� 35 ��W, 80 m], Resguardo Ind��gena de Camaritagua, towards El Francisco. Type material. Holotype (ICNB), J. Cort��s leg./ 2004. Remarks. Although the shell is somewhat worn, its characteristics separate it from all other Plekocheilus (Eurytus) species. It may be compared to P. (E.) fusitorsus (Oberwimmer, 1931) from which it differs by (1) having a shorter spire; (2) a less impressed suture; (3) a less expanded lip. It may also be compared to P. (E.) coloratus (Nyst, 1845) which, however, (1) has the whorls more convex; (2) has a larger angle of the attachment of the lip to the body whorl; (3) has the spire more pointed. Finally, this new species may be compared to P. (E.) tepuiensis Breure, 2009, from which it differs in (1) having a different, more brownish color, with the yellowish ���shadow��� pattern; (2) having a less pointed spire; (3) having a different height/number of whorls ratio. The only known (possibly subadult) shell, the holotype, was found in SE Colombian rainforest, where snail collecting for scientific purposes has been very limited. According to the label, indigenous people use the mucus of these snails for scrubbing their skin. The color pattern appears to be similar to both the yellow trait specimens of P. coloratus from Sierra Nevada de Santa Marta and to P. fusitorsus (Oberwimmer, 1931) from Venezuelan Guayana. Etymology. This very distinctive species is named after the Camaritagua, indigenous people of the region where it was found. The epithet is used as a noun in apposition., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 37, DOI: 10.5281/zenodo.278895, {"references":["Nyst, H. (1845) Description de deux Bulimes nouveaux de la Colombie. Bulletin de l'Academie royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 12 (1), 227 - 229, pl.","Breure, A. S. H. (2009) New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World. Zootaxa, 2065, 25 - 50."]}

Dryptus jubeus (Fulton, 1908) (Figs 3 A���B) Strophocheilus (Dryptus) jubeus Fulton 1908: 86, text fig. Type locality: Venezuela, Capas. Holotype BMNH 1905.5. 3.1. Dryptus jubens [sic] (Fulton); Richardson 1995: 200 (references). Material. Venezuela, M��rida, Capas [Capaz]: USNM 603094 (paratype); mountains La Paz [misspelling for Capaz?], 2000 m, A. Gabaldon leg.: USNM 109534; Torondoy mountains, 1600 m, Gabaldon leg.: USNM 109535. Altitudinal range. The localities reported here range from 1600���2000m. Remarks. The type material of both D. jubeus and D. guerini show some differences, viz. D. jubeus has (1) a more pointed spire; (2) a broader, whitish parietal callus; and (3) a pinkish lip. We tentatively consider D. jubeus a valid taxon., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 8, DOI: 10.5281/zenodo.278895, {"references":["Fulton, H. C. (1908) Description of a new species of Strophocheilus. Proceedings of the Malacological Society of London, 8, 86 - 87.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458."]}

Plekocheilus (Eurytus) dalmasi (Dautzenberg, 1900) (Figs 9 C, 10 V ���W) Plecochilus dalmasi Dautzenberg 1900: 151, pl. 9 fig. 1. No type locality given. Lectotype RBINS /MT 668 (Breure 2011). Plekocheilus dalmsi [sic] Dautzenberg; Richardson 1995: 308 (references). Material. ?Magdalena/La Guajira, Sierra Nevada de Santa Marta, leg./ 1896: RBINS; Boyac��, km 61.5 road Soacha���La Punta, 6 km E Los Pinos, Alto de Granados, P��ramo de Pisva [sic, Pisba], 3620 m, Cleef leg./ 14.vi. 1972: RMNH. Remarks. Breure (1978) located the type material, of which one specimen is figured here. The specimen from P��ramo de Pisba is tentatively referred to this species. This locality forms part of the Cocuy area, which is known for its endemics (see also Breure 2008) and is phytogeographically related to the Sierra Nevada de Santa Marta (Cleef, pers. comm.). The single shell in the lot from P��ramo de Pisba (RMNH) lacks the upper whorls (due to fracture) and shows traces of oblique chestnut-brown color streaks on a golden-brown ground color. The white lip is characteristic in all material examined., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 29, DOI: 10.5281/zenodo.278895, {"references":["Dautzenberg, P. (1900) Croisieres du yacht Chazalie dans l'Atlantique. Mollusques. Memoires de la Societe Zoologique de France, 13, 145 - 156, pl. 9 - 10.","Breure, A. S. H. (2011) Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species. ZooKeys, 101, 1 - 50.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Breure, A. S. H. (1978) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen, 164, 1 - 255."]}

Group of Plekocheilus (Eurytus) pulicarius Remarks. The following species are treated from the study area: P. (E.) pulicarius, P. (E.) labiosus and P. (E.) piperitus., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 46, DOI: 10.5281/zenodo.278895

Plekocheilus (Eurytus) subglandiformis (Mousson, 1873) (Figs 9 C, 10 L���N) Bulimus subglandiformis Mousson 1873: 6. Type locality: Colombia, Bogot��. Lectotype ZMUZ 511633 (Breure 1976). Plekocheilus subglandiformis (Mousson); Richardson 1995: 321 (references, synonymy); Neubert and Janssen 2004: 230, pl. 1 fig. 2; Simone, 2006: 150, fig. 506. Material. Antioquia, Sons��n, Wallis leg.: SMF 21603 (ex W. Kobelt); Frontina [Frontino] (ZMUZ 233 [sic = 511632, see remarks]) (Simone, 2006);?Cundinamarca, ���Bogota���: ZMUZ 511633.?, U.S. of Colombia: USNM 316120 (ex Henderson Colln, ex Evezard Colln); Columbia: ZMUZ 511631 (not seen). Remarks. The specimen pictured by Neubert and Janssen (2004) does not show the characteristic marbled band below the suture, consisting of brown spots that fade into yellowish; these yellowish spots may continue in waving, axial bands. If the ZMUZ and SMF specimens originate from the same series, which Neubert and Janssen consider ���very probable,��� the specimens in SMF could be regarded as paralectotypes. However, it should be noted that their specimen is from ��� Antigua [sic, most probably Antioquia], Sons��n,��� while Mousson���s specimen in ZMUZ is labelled with ���Bogota.��� Also, Breure���s (1976) statement of a ���unique specimen ZMUZ 511633 ��� is in error, for two reasons: (1) we have a file of material at ZMUZ, listing three lots as P. subglandiformis (511631,��� Columbia;��� 511632, ��� Frontina ��� and 511633, ���Bogot����� [lectotype]); (2) Simone (2006: 150, fig. 506) shows two specimens from ZMUZ, the lectotype (cited in error as ZMUZ 233), and the specimen from Frontina (cited in error as ZMUZ 233). Mousson only mentions Bogot�� in his paper (Mousson 1873: 1), without giving further details on (other) localities in his description., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 29, DOI: 10.5281/zenodo.278895, {"references":["Mousson, A. (1873) Zweite Notiz uber einige von Herrn Gustav Wallis aus dem nordlichen Sud-Amerika zuruckgebrachte Mollusken. Malakozoologische Blatter, 21, 1 - 19.","Breure, A. S. H. (1976) Types of Bulimulidae (Gastropoda, Euthyneura) in the Zoologisches Museum, Universitat Zurich. In: Malacologische Opstellen. Backhuys, Rotterdam, 1 - 4, pls. 1 - 3.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Neubert, E. & Janssen, R. (2004) Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv fur Molluskenkunde, 133, 193 - 297.","Simone, L. R. L. (2006) Land and freshwater molluscs of Brazil. EGB / Fapesp, Sao Paulo, 390 pp."]}

Plekocheilus (Eurytus) piperitus (Sowerby I, 1837) (Figs 17 G���J) Bulimus piperitus Sowerby I 1837 (1832���1841): fig. 93. Type locality: [Peru, Dept. San Mart��n] Huallaga. Type material: not located. Richardson 1995: 318 (references). Material. Ecuador, Tungurahua, Rio Pastaza watershed, ex E. Doremus: DMNH 56470; ��� Ecuador:��� USNM 316099 (ex Henderson Colln, ex Evezard Colln, as P. piperitus roseolabrum), USNM 532641 (ex Fulton, as Eurytus piperitus roseolabrum). Brazil, Amazonas, Igarap�� Anib�� [Remarks. This species is known from northern Peru and northwestern Brazil (Ram��rez et al. 2003; Simone 2006), and was not mentioned under this name by Breure and Borrero (2008) for the Ecuadorian malacofauna. It may be restricted to lowlands (Huallaga is ~ 310m; ���Rio Pastaza watershed��� may refer to anywhere in a wide range of elevations). The date of Sowerby���s publication of P. piperitus was erroneously given as 1833 by Breure (1979). Recently, Petit (2009) has given a valuable overview on the main publications of the Sowerby family; from his paper the correct date (1837) can be extracted (see also Shaw 1909). Lots USNM 316099 and 532641 associate this species with Plekocheilus (E.) roseolabrum (E.A. Smith 1877), which is considered here as a distinct species from Ecuador., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 48, DOI: 10.5281/zenodo.278895, {"references":["Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Simone, L. R. L. (2006) Land and freshwater molluscs of Brazil. EGB / Fapesp, Sao Paulo, 390 pp.","Ramirez, R., Paredes, C. & Arenas, J. (2003) Moluscos del Peru. Revista de Biologia Tropical, 51 (Supplement 3), 225 - 284.","Breure, A. S. H. & Borrero, F. J. (2008) An annotated checklist of the land snail family Orthalicidae (Gastropoda: Pulmonata: Orthalicoidea) in Ecuador, with notes on the distribution of the mainland species. Zootaxa, 1768, 1 - 40.","Breure, A. S. H. (1979) Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen, 168, 1 - 215.","Petit, R. E. (2009) George Brettingham Sowerby I, II & III: their conchological publications and Molluscan taxa. Zootaxa, 2189, 1 - 218.","Shaw, H. O. N. (1909) On the dates of issue of Sowerby's \" Conchological illustrations, \" from the copy preserved in the Radcliffe Library, Oxford. Proceedings of the Malacological Society of London, 8, 333 - 340.","Smith, E. A. (1877) Descriptions of new species of South American Helicidae in the British Museum. Proceedings of the Zoological Society of London, 1877, 361 - 365."]}

Plekocheilus (Aeropictus) succineoides zilchi Breure, 1977 (Fig. 9 B) Plekocheilus (Aeropictus) zilchi Breure 1977: 260, figs 2, 21��� 22. Type locality: Dept. Boyac��, SW Labranza Grande, Quebrada Comijoque. Holotype SMF 245387. Plekocheilus zilchi Breure; Richardson 1995: 325 (references); Neubert & Janssen, 2004: 235, pl. 1 fig. 5. Material. Boyac��, SW Labranza Grande, Quebrada Comijoque (05�� 32 ' N 072�� 40 ' W, 2800 m, type locality).?, Bogot��: USNM 124663 (partim); UMMZ 242812 (Marsh Colln). Remarks. This taxon was differentiated by Breure (1977) using both conchological and anatomical characters. It occurs on the eastern escarpments of the Cordillera Oriental, ca. 90 km from the nearest record of the nominate subspecies., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 17, DOI: 10.5281/zenodo.278895, {"references":["Breure, A. S. H. (1977) Notes on Bulimulidae (Gastropoda, Euthyneura), 5. On some collections from Colombia. Archiv fur Molluskenkunde, 107, 257 - 270.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Neubert, E. & Janssen, R. (2004) Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae. Archiv fur Molluskenkunde, 133, 193 - 297."]}

Plekocheilus (Aeropictus) veranyi (Pfeiffer, 1848) Bulimus veranyi Pfeiffer 1848 a: 230. Type locality: Venezuela, Edo. M��rida, Chachopo. Lectotype BMNH 1975297 (Breure 1978). Plekocheilus (Aeropictus) veranyi (Pfeiffer); Tello 1975: 201. Plekocheilus veranyi (Pfeiffer); Richardson 1995: 323 (references, synonymy; see remarks). Plekocheilus (Orcesiellus) succinoides [sic] (Petit); Tello 1975: 201. Bulimus scytodes Pfeiffer 1853: 256. Type locality: Andes of Colombia. Type material: not located. Material. Venezuela, Barinas, Serran��a de Santo Domingo, Jackson Colln, 3000 ft [~ 1000 m, in error?]: DMNH 148721, USNM 109523 (Gabaldon leg., 3000 m); M��rida, Chachopo: BMNH; between Chachopo and Timote, C. Beets leg./ 1953: RMNH; M��rida, Jackson Colln: DMNH 160801;?M��rida, Escorial Mountains, A. Gabaldon & sons leg.: USNM 206502 (8400 ft [~ 2520 m]), USNM 206504 (9600 ft [~ 2880 m]); Glorial: DMNH 141740; ���M��rida:��� UMMZ 147737 (ex Rolle); ���Menda��� (sic): UMMZ 147736, 242815;?Miranda, Caracas, ex M.L. Jaume: CMC C 13777.?, San Domingo: UMMZ 242814 (Marsh Colln), UMMZ 147735; ��� Venezuela:��� CMC C 11004 ���11006, USNM 109526; New Granada: ANSP 66405, ANSP 66408, BMNH (ex Cuming), ZMUZ 511641 [not seen], ZMUZ 511642 [not seen]. Altitudinal range. The localities reported here range between 2520 and 3000 m (see remarks). Remarks. The date of first publication mentioned by Richardson (1995) is erroneous. He cited ��� 1847 Philippi, Abbild. Beschr. neuer Conchyl. 3: 20, pl. 8 figs 5, 9 (1851)���. However, P. veranyi is mentioned on page 36, under the heading ��� Bulimus. Tab. VIII. Januar 1849.��� Thus the first valid publication was in Pfeiffer (1848 a). Of the taxa grouped under this species by Richardson (1995: 324), we consider P. cathcartiae and P. quadricolor to be distinct species. This species is very similar to P. delicatus, but differs in being decidedly smaller. Colombian material, labeled as P. veranyi, should all be considered P. delicatus. Two reasons lead us to believe that the altitude of DMNH 148721 (3000 ft) may be in error: (1) Another lot from the same location (USNM 109523) is clearly labeled with an elevation of 3000 meters; (2) All known species of the subgenus Aeropictus, similar to P. succineoides, are known only from high elevations, mainly p��ramo (i.e., 2500 m and above); if correct, 3000 ft would be by far the lowest elevation known for any species in this group, which we consider unlikely. We have been unable to trace the localities ���Glorial��� and ���Escorial Mountains;��� the latter, may refer to a location near P��ramo El Escorial (~ 3000 m), or near the town of El Escorial (~ 3100 m), both within the Sierra Nevada de M��rida. Tello (1975) included P. succinoides [sic] in his list of Venezuelan land snails (cf. Weyrauch), apparently as a misinterpretation of a portion of Weyrauch���s (1967) description of the subgenus Orcesiellus. Contrary to Tello (1975: 201), we consider it likely that the species mentioned from M��rida by this author is P. veranyi (Pfeiffer)., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 12, DOI: 10.5281/zenodo.278895, {"references":["Pfeiffer, L. (1848 a) Descriptions of nineteen new species of Helicea, from the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London, 1847, 228 - 232.","Breure, A. S. H. (1978) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen, 164, 1 - 255.","Tello, J. (1975) Catalogo de la Fauna Venezolana. VIII. Mollusca. Published by the author, Caracas, 599 pp.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458."]}

“ Plekocheilus ” victor (Pfeiffer, 1854) Remarks. This taxon, described as Bulimus victor from Antioquia, is listed by Breure and Schouten (1985) under the subfamily Orthalicinae (= family Orthalicidae sensu Breure et al. 2010). The affiliation of this taxon to Plekocheilus as used by Pilsbry (1895 [1895 –1896]) and Richardson (1995) is uncertain. It will be treated in a forthcoming paper in this series.

Plekocheilus (Eurytus) taylorianus (Reeve, 1848) (Figs 15 C���D) Bulimus taylorianus Reeve 1849: pl. 81 fig. 602. Type locality: Ecuador, near Quito, Chimborazo. Lectotype BMNH 1874.12. 11.271 (Breure 1978). Plekocheilus taylorianus (Reeve); Richardson 1995: (references, synonymy; see remarks). Material. Ecuador. Chimborazo, Mt. Chimborazo, 21424 ft [6424 m, in error], ex Evezard Colln ex Henderson Colln: USNM 316070; Cotopaxi, Sigchos, W Latacunga: DMNH 141642; Imbabura, Ibarra: DMNH 128769; Napo, Nachiyacu: DMNH 141669; Topo: DMNH 76772, DMNH 128672, DMNH 151005); Pastaza, Mera: FMNH 202059, DMNH 141649, DMNH 183697; Puyo: DMNH 141684; Pichincha: DMNH 141681; Milpe: UF 166225 [not seen], DMNH 141658; Mindo: DMNH 141664, UF 166226 (near Rio Blanco) [not seen]; Nanegal: EPN; Pacto: EPN; Pintag: EPN; Gualea: FMNH 61138, FMNH 31173, CM 106225 (A. Cousin leg./ 1898); Quito, Stearns Colln: UMMZ 13841.?, ��� Ecuador:��� CMC C 11056, CMC C 11057, FMNH 100722, RMNH.? ��� Peru,��� F.M. Hele leg.: UMMZ 147732. Remarks. The records available suggest that P. taylorianus is confined to Ecuador, but it may extend into northernmost Peru. K��hler (2007): 127, fig. 5 mentioned a paralectotype in Berlin (ZMB 112.722). However, the picture shown is a different species of Plekocheilus. The lots available do not have actual records of altitude, but several of the localities listed are in the ~ 2200���2890 m range. However, Archidona and Puyo are lower (605 and 950 m), and a wide range of altitudes exists around Mindo (1400���4780 m). The label of lot USNM 316070 is clearly in error as the maximum elevation of Mt. Chimborazo is 6310 m; it is likely that those specimens were collected at a much lower elevation in the foothills of Chimborazo., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on pages 42-43, DOI: 10.5281/zenodo.278895, {"references":["Reeve, L. A. (1848 - 1850) Conchologia iconica: or illustrations of the shells of molluscous animals, 5. Monograph of the genus Bulimus. Reeve, Benham, and Reeve, London, 89 pls + legend., i - xi.","Breure, A. S. H. (1978) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen, 164, 1 - 255.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Kohler, F. (2007) Annotated type catalogue of the Bulimulinae (Pulmonata, Orthalicoidea, Bulimulidae) in the Museum fur Naturkunde Berlin. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 83, 125 - 159."]}

Plekocheilus (Plekocheilus) fulminans (Nyst, 1843) (Figs 7 D���E) Bulimus fulminans Nyst 1843: 261, pl. 7 fig. 1. Type locality: Venezuela, Est. Sucre. Lectotype RBINS /MT 2351 (Breure 2011). Plekocheilus fulminans (Nyst); Richardson 1995: 312 (references). Material. ?, ��� Columbia,��� Thorey leg.: ANSP 328096. Venezuela, Sucre, Cuman��: DMNH 76763, USNM 307543; Angostura: CMC C 10997. ��� Venezuela:��� CMC C 10733, CMC C 10734, UMMZ 147716 (G.A. Lathrop leg.). Guyana, Mt. Roraima: UMMZ 147717 (ex Linter, ex Tomlin). Remarks. According to Abbott (1989) and Haas (1955), this species has a wide distribution throughout Venezuela and northeastern Colombia. However, we have not seen any specimens from Colombia that can unambiguously be attributed to this taxon. The specimen from ANSP figured here matches the figure from Nyst (1843) quite well. Altitude data have been reported only for P. fulminans linterae (Sowerby III, 1890) and P. fulminans alticola (Haas, 1955) from the highlands in Venezuelan Guayana (see Breure 2009). The only other specific locality known, for the nominate form, is Cuman�� (near sea level), which might not be the actual collection locality (see also the general remark in the introduction). Breure and Schl��gl (2010) discussed the relationships between the nominate taxon and the taxa from Venezuelan Guayana., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 20, DOI: 10.5281/zenodo.278895, {"references":["Nyst, H. (1843) Notice sur deux coquilles colombiennes du genre Bulimus. Memoires du Societe royale de Sciences, Liege, 1, 261 - 262, pl. 7.","Breure, A. S. H. (2011) Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species. ZooKeys, 101, 1 - 50.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Abbott, R. T. (1989). Compendium of Landshells. American Malacologists Inc., Melbourne, Florida, 240 pp.","Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 - 387.","Breure, A. S. H. (2009) New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World. Zootaxa, 2065, 25 - 50.","Breure, A. S. H. & Schlogl, J. (2010) Additional notes on Orthalicidae from the Chimanta massif, Venezuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828. Zootaxa, 2416, 51 - 60."]}

Subgenus Plekocheilus (Eurytus) Albers, 1850 Type species by subsequent designation (Albers 1860): Helix pentadina d���Orbigny, 1835. Distribution. West Indies (St. Lucia, St. Vincent), Venezuela, Guyana, Brazil, Bolivia, Peru, Ecuador, Colombia, Panama. Remarks. In the last comprehensive revision of this genus, Pilsbry (1895 [1895 ���1896]) divided this subgenus into seven groups of species. His grouping was based solely on shell morphology and mainly used shape, surface sculpture, columellar fold, expansion of peristome and suture of last whorl to differentiate among species. Anatomical data have been presented for relatively few of the more than 60 taxa known in this subgenus (Breure 1978, 2009; Breure & Schl��gl 2010). Our grouping is thus based mainly on shell morphology, supplemented by the sparse anatomical data available for Colombian species. An additional comment on ecological settings where species of Eurytus occur is warranted. Based on ecological information then available, Breure (1979) suggested that two groups of species may be distinguished within this subgenus, namely a group that lives mainly in rain forest at altitudes of up to circa 1000 m, and a second group of species that live mainly in montane forest at altitudes of circa 1250���3500 m. We continue to adhere to this view, but have chosen to provide this information under each species when available, rather than providing a listing by elevation. Further, a number of species are still known only from the type collections, often without precise locality data., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 25, DOI: 10.5281/zenodo.278895, {"references":["Albers, J. C. (1860) Die Heliceen nach naturlicher Verwandtschaft systematisch geordnet, 2 Ausgabe (ed. E. von Martens). Engelmann, Leipzig, xviii + 359 pp.","Pilsbry, H. A. (1895 - 1896) Manual of Conchology, (2) 10. American Bulimi and Bululimuli. Strophocheilus, Plekocheilus, Auris, Bulimulus. Academy of Natural Sciences of Philadelphia, Philadelphia, iv + 213 pp, 51 pls.","Breure, A. S. H. (1978) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda). Zoologische Verhandelingen, 164, 1 - 255.","Breure, A. S. H. (2009) New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World. Zootaxa, 2065, 25 - 50.","Breure, A. S. H. & Schlogl, J. (2010) Additional notes on Orthalicidae from the Chimanta massif, Venezuelan Guayana, with descriptions of new species of Plekocheilus Guilding, 1828. Zootaxa, 2416, 51 - 60.","Breure, A. S. H. (1979) Systematics, phylogeny and zoogeography of Bulimulinae (Mollusca). Zoologische Verhandelingen, 168, 1 - 215."]}

Dryptus moritzianus (Pfeiffer, 1847) (Fig. 2 C, 4 B) Bulimus moritzianus Pfeiffer 1847: 66. Type locality: Venezuela, Caracas. Lectotype ZMB 10464 a (K��hler 2007: 128, fig. 13). Bulimus (Dryptus) moritzianus Pfeiffer; Tello 1975: 197. Dryptus moritzianus (Pfeiffer); Richardson 1995: 201 (references). Stropocheilus moritzianus wilsoni Pilsbry 1895 [1895 ���1896]: 39, pl. 21 fig. 48. Type locality: not given. Lectotype ANSP 7775 (Baker, 1963). Material. Arauca, ���Alto Arauca��� [= Upper Rio Arauca]: DMNH 128665 (R.W. Jackson Colln), ANSP 220773 (B.R. Bales Colln); Casanare, Tamara, Colombia [either T��mara, or Taman��, see remarks], A.A. Olsson leg./ 1943: ANSP 180086; Norte de Santander, ���Sta. Librada (Sarare)��� [= Santa Librada], R.P.E. Rochereau leg./ 1937: DMNH 128666; Pamplona (Pilsbry 1895 [1895 ���1896]); humid region south of the Cucuta of Tama [Tam��], Hno. Nic��foro Mar��a leg.: ANSP 168416.?, Colombia: FMNH 31040, RBINS, ZMH 7346, ZMH 7347, UF 22129 (not seen).?, ��� U.S. of Colombia:��� CM 62.4690; ���Andes, New Granada,��� Margnier leg./ 1849: ZMUZ 511563; ��� New Granada:��� ANSP 3577 ���3579, 32901, 50486, ZMUZ 511573 (Hoffmann leg./ 1851), UMMZ 144495 (Ponsonby Colln). Venezuela. M��rida, Capas [Capaz], 2000 m, J.R. Tomlin leg.: UMMZ 144492; Miranda?, Caracas: ZMB; ��� Venezuela:��� CM 62.4689 (Hartman Colln), UMMZ 144497 (as Strophocheilus moritzianus wilsoni Pilsbry; ex Tomlin, ex Ponsonby, ex Rolle), CM 62.30565 (ex Rolle). Remarks. Regarding the collection locality of DMNH 12866 [Sta. Librada (Sarare)], there are places called Santa Librada in Quind��o, Tolima, Vichada, and Cundinamarca. However, several reasons lead us to believe that the location in question is in the eastern plains of Colombia, and probably in Norte de Santander: (1) all other confirmed Colombian records for this species, and most of the species of Dryptus come from the lowlands of that region, and from Venezuela; (2) this species has not been collected in Quind��o, Tolima or Cundinamarca; (3) a region known as ���regi��n del Sarare��� in the Cordillera Oriental (Eastern Range of the Andes), and ���Sarare, Santa Librada, 1300���1600 m ��� are localities from which paratypic collections of ferns, Danae betancurii A. Rojas, 2009 have been reported (Rojas-Alvarado 2009 a). Thus, we place lots labeled Santa Librada and Santa Librada (Sarare) within Norte de Santander. We are aware that there is a Santa Librada in Vichada, and a location called El Sarare in Arauca within ~ 150 km of each other, both at less than 200 m of elevation. Of note is that if Rochereau���s actual collection localitites were in Arauca/Vichada, they would still be consistent with a distribution of this species in the eastern plains of Colombia, including equivalent elevations. Likewise, there are several locations called T��mara, or Taman��, in various departments of Colombia (Casanare, Boyac��, Tolima); we consider that it is more likely either Quebrada T��mara, or Cerro Taman��, both in Norte de Santander. In addition to the Venezuelan distribution indicated here, Tello (1975) listed Puerto Cabello (Edo. Carabobo), and Edo. Trujillo as localities for this species., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on pages 9-10, DOI: 10.5281/zenodo.278895, {"references":["Pfeiffer, L. (1847) Diagnosen neuer Heliceen. Zeitschrift fur Malakozoologie, 4, 65 - 71.","Kohler, F. (2007) Annotated type catalogue of the Bulimulinae (Pulmonata, Orthalicoidea, Bulimulidae) in the Museum fur Naturkunde Berlin. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 83, 125 - 159.","Tello, J. (1975) Catalogo de la Fauna Venezolana. VIII. Mollusca. Published by the author, Caracas, 599 pp.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Pilsbry, H. A. (1895 - 1896) Manual of Conchology, (2) 10. American Bulimi and Bululimuli. Strophocheilus, Plekocheilus, Auris, Bulimulus. Academy of Natural Sciences of Philadelphia, Philadelphia, iv + 213 pp, 51 pls.","Baker, H. B. (1963) Type land snails in the Academy of Natural Sciences of Philadelphia Part II. Land Pulmonata, exclusive of North America north of Mexico. Proceedings of the Academy of Natural Sciences of Philadelphia, 115, 191 - 259.","Rojas-Alvarado, A. F. (2009 a) Novelties in Danaea moritziana complex (Marattiaceae) from Colombia. Metodos en Ecologia y Sistematica, 4, 8 - 19."]}

Plekocheilus (Eurytus) jucundus (Pfeiffer, 1855) Bulimus jucundus Pfeiffer 1855 c: 290. Type locality: Colombia, Antioquia. Richardson 1995: 315 (references). Remarks. We have been unable to recognize this taxon in the material we have seen. The original figures suggest that it may prove to be a synonym of P. plectostylus (Pfeiffer, 1848). The location of the type is unknown., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 53, DOI: 10.5281/zenodo.278895, {"references":["Pfeiffer, L. (1855 c) Descriptions of fifty-seven new species of Helicea, from Mr. Cuming's collection. Proceedings of the Zoological Society of London, 1855, 286 - 298.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458."]}

Plekocheilus (Eurytus) tetensii (Dunker, 1875) (Figs 14 B, 16 N���R) Bulimus tetensii Dunker 1875: 29. Type locality: Colombia, Sierra Nevada de Santa Marta. Type material: not located. Plekocheilus tetensii (Dunker); Richardson 1995: 319 (references). Material. Cesar, Rio Guatapur��, Sierra Nevada de Santa Marta: USNM 599530 (8000���9000 ft [~ 2400���2700 m], Carriker leg./ 1946, as Plekocheilus coloratus lamarckianus), USNM 599524, partim (12000 ft [~ 3600 m], as P. coloratus subplicatus), USNM 599525 (as P. coloratus subplicatus); La Guajira, Sierra Nevada de Santa Marta, P��ramos de Macotama, 10000���15000 ft [~ 3000���4500 m], Carriker leg.: ANSP 151815; Cerro Pintado, Sierra de Perij��, 10300���10600 ft [~ 3090���3180 m], Carriker leg./ 1942, all as P. (E.) piperatoides: FMNH 62362; USNM 488807, USNM 488808 [Label indicates one specimen was collected at 9000 ft], MCZ 183872); Sierra Nevada de Santa Marta, ���northern slope, above Pueblo vigio [Pueblo Viejo], 1400 m:��� ZMB 39226 [considered as topotypes by K��hler 2007: 127, fig. 6]; Magdalena, Cuchilla G��inue, Cleef et al. leg./ 1977, in p��ramo vegetation (Calamagrostis sp. dominant) (Breure 1977, 1984): RMNH (Filo Alto La Cumbre, 3700m), ICNB (3000 m). Altitudinal range. 1400���4500 m. If the material has been correctly labeled, this represents the highest elevation reported here for species of Plekocheilus and Dryptus. Remarks. We consider K��hler���s decision (2007) to regard specimens from Pueblo Viejo as topotypes as somewhat arbitrary, as there is no evidence that Dunker���s specimens originate from that specific locality. The specimens from Sierra de Perij��, Cerro Pintado are tentatively referred to this species, as they differ in being smaller and having the lip and parietal callus pink., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on pages 48-49, DOI: 10.5281/zenodo.278895, {"references":["Dunker, W. (1875) Zwei neue Bulimi aus der Sierra Nevada (Columbia). Jahrbucher der deutschen malakozoologischen Gesellschaft, 2, 220 - 221, pl. 6 figs. 1 - 4.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Kohler, F. (2007) Annotated type catalogue of the Bulimulinae (Pulmonata, Orthalicoidea, Bulimulidae) in the Museum fur Naturkunde Berlin. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 83, 125 - 159.","Breure, A. S. H. (1977) Notes on Bulimulidae (Gastropoda, Euthyneura), 5. On some collections from Colombia. Archiv fur Molluskenkunde, 107, 257 - 270.","Breure, A. S. H. (1984) Survey of the land Mollusca of the Sierra Nevada de Santa Marta, Colombia. In: Hammen, T. van der & Ruiz, P. M. (eds.), Studies on Tropical Andean ecosystems, 2, La Sierra Nevada de Santa Marta (Colombia) Transecto Buritaca ─ La Cumbre. Cramer, Berlin / Stuttgart, pp. 487 - 500."]}

Group of Plekocheilus (P.) fulminans Remarks. The following species are treated from the study area: P. (P.) blainvilleanus, P. (P.) fulminans, P. (P.) loveni, and P. (P.) appuni., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 18, DOI: 10.5281/zenodo.278895

Subgenus Plekocheilus (Eudolichotis) Pilsbry 1896 Type species by original designation: Bulimus distortus Brugui��re, 1789. Distribution. West Indies (Grenada, Trinidad), Venezuela, Guyana, Suriname, Brazil, Colombia, Panama., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on page 50, DOI: 10.5281/zenodo.278895, {"references":["Bruguiere, J. C. (1789 - 1792) Histoire naturelle des Vers, Tome premier. In: Encyclopedie methodique, ou par ordre de matiere, 11. Panckoucke, Paris, 1 e Partie, pp. i - xviii + 1 - 344 (1789), 2 e Partie, pp. 345 - 757 (1992)."]}

Plekocheilus (Aeropictus) guentheri (Sowerby III, 1892) comb. nov. (Figs 5 M���P, 9 A) Bulimus (Plekocheilus) guildingi Dohrn 1875: 306, pl. 10 figs 5���6. Type locality: ���Neu-Granada.��� Syntype ZMB 101709. Not Bulimus guildingi Pfeiffer, 1842. Bulimus guentheri Sowerby III 1892: 296, pl. 23 figs 7���8. Type locality: ��� U.S. of Colombia.��� Plekocheilus guentheri (Sowerby); Richardson 1995: 315 (references). Plekocheilus (Eurytus) ameghinoi Parodiz 1962: 451. New name for B. guildingi Dohrn, 1875 not Pfeiffer, 1842. Plekocheilus ameghinoi Parodiz; Richardson 1995: 302 (references, synonymy). Material. Antioquia/C��rdoba, Rio Esmeralda, Upper Rio Sin��, Carriker leg./ 1949: USNM 601774.?, ���U S Colombia:��� ANSP 104845, USNM 532640, USNM 532644 (ex Fulton, Chamberlain Colln). Remarks. Both the Rio Esmeralda and the Rio Sin�� run from northern Antioquia to C��rdoba; the reference to ���Upper Rio Sin����� suggests that the specific location is probably in the highlands of the border area between both departments in the Parque Nacional Natural Paramillo. This is the first precise locality for this species. Pilsbry (1901: 129) was the first to notice the homonymy of this species with Bulimus guildingi Pfeiffer, 1842; therefore he suppressed Dohrn���s name in favour of Sowerby���s. Later Parodiz (1962) introduced the name Plekocheilus (Eurytus) ameghinoi as a new name for P. guildingi (Dohrn), apparently unaware of Pilsbry���s action. K��hler (2007: 127, fig. 3) found three probable syntypes of Bulimus guildingi Dohrn (type locality: ���Neu Granada ���) in the Berlin museum., Published as part of Borrero, Francisco J. & Breure, Abraham S. H., 2011, The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas, pp. 1-59 in Zootaxa 3054 on pages 17-18, DOI: 10.5281/zenodo.278895, {"references":["Dohrn, H. (1875). Ueber einige von Herrn G. Wallis in Neugranada gesammelte Landconchylien. Jahrbucher der deutschen malakozoologischen Gesellschaft, 2, 292 - 309, pl. 10.","Richardson, C. L. (1995) Bulimulidae: catalog of species. Tryonia, 28, 1 - 458.","Parodiz, J. J. (1962) New and little-known species of South and Central American land snails (Bulimulidae). Proceedings of the United States National Museum, 113 (3462), 429 - 456, pls. 1 - 2.","Pilsbry, H. A. (1901 - 1902) Manual of Conchology (2) 14. Oriental bulimoid Helicidae; Odontostominae; Cerionidae. Academy of Natural Sciences, Philadelphia, 1 - 302, pls. 1 - 62.","Kohler, F. (2007) Annotated type catalogue of the Bulimulinae (Pulmonata, Orthalicoidea, Bulimulidae) in the Museum fur Naturkunde Berlin. Mitteilungen aus dem Museum fur Naturkunde Berlin, Zoologische Reihe, 83, 125 - 159."]}