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An article published by Gressitt (1941) has been ignored by all longicornists, including Gressitt himself. However, according to ICZN, it meets all criteria as an official publication, and this status affects three taxa that were formally described therein: Bacchisa (Bacchisa) rigida (Gressitt, 1941) = Chreonoma rigida Gressitt, 1941 = Chreonoma rigida Gressitt, 1942 homonym and syn. nov.; Tetraophthalmus sikang (Gressitt, 1941) = Chreonoma sikang Gressitt, 1941 = Chreonoma sikanga Gressitt, 1942, syn. nov.; Anastathes parva hainana Gressitt, 1941 = Anastathes parva hainana Gressitt, 1942, homonym and syn. nov.

Cylindroeme Vives, 2019 8柱ḃk牛Β Cylindroeme Vives, 2019: 1. Type species: Cylindroeme vietnamica Vives, 2019, by original designation. Distribution. China, Vietnam., Published as part of Lin, Mei-Ying & Li, Kai-Qin, 2022, A new species, Cylindroeme yunnanensis sp. nov. from China (Coleoptera, Cerambycidae, Cerambycinae), pp. 294-300 in Zootaxa 5159 (2) on page 295, DOI: 10.11646/zootaxa.5159.2.7, http://zenodo.org/record/6777338, {"references":["Vives, E. (2019) Descripcion de un nuevo genero de Oemini de Vietnam (Coleoptera, Cerambycidae). (Cerambicidos nuevos o interesantes de Vietnam. Pars X) [Description of a new Oemini from Vietnam belonging to a new genus (Coleoptera, Cerambycidae). (Cerambicidae new or little known from Vietnam. Pars X)]. Faunitaxys, 7 (10), 1 - 2."]}

Cylindroeme vietnamica Vives, 2019 åfihk牛 Figures 8–10 & 13-17 Cylindroeme vietnamica Vives, 2019: 1, figs. 1–6. 1. TL: Vietnam: Ha Giang. TD: CEV. Cylindroeme vietnamica: Wang, Xie & Wang, 2022: 129, figs. 1–2, 9, 12. Specimens examined. Vietnam: 9 males and 1 female, Dong Van, Ha Giang Province, 2019.IV (CTN). China: 1 male, China, Guizhou, Guiyangshi, Huaxiqu, Guizhoudaxue, Nanxiaoqu (South campus of Guizhou University), 2007.IX, Ye Liu leg. (IZCAS). Distribution. China (new record): Guizhou. Vietnam: Ha Giang. Remarks. This species is recorded from China for the first time. Compared the only one Guizhou specimen with the Ha Giang specimens in CTN, the median struts of median lobe from Guizhou are slightly shorter than one specimen dissected from Ha Giang. However, all other characters, such as the median notch and ridges on pronotum, longitudinal ridges on elytra, are not different. Therefore, we identified it as C. vietnamica., Published as part of Lin, Mei-Ying & Li, Kai-Qin, 2022, A new species, Cylindroeme yunnanensis sp. nov. from China (Coleoptera, Cerambycidae, Cerambycinae), pp. 294-300 in Zootaxa 5159 (2) on page 297, DOI: 10.11646/zootaxa.5159.2.7, http://zenodo.org/record/6777338, {"references":["Vives, E. (2019) Descripcion de un nuevo genero de Oemini de Vietnam (Coleoptera, Cerambycidae). (Cerambicidos nuevos o interesantes de Vietnam. Pars X) [Description of a new Oemini from Vietnam belonging to a new genus (Coleoptera, Cerambycidae). (Cerambicidae new or little known from Vietnam. Pars X)]. Faunitaxys, 7 (10), 1 - 2.","Wang, Y. - Q., Zhao, W. - W. & Xie, G. - L. (2022) A revision of the genus Cylindroeme Vives (Coleoptera, Cerambycidae, Cerambycinae). ZooKeys, 1082, 127 - 134. https: // doi. org / 10.3897 / zookeys. 1082.75816"]}

Cylindroeme shii Wang, Xie & Wang, 2022 石氏åfihk牛 Cylindroeme shii Wang, Xie & Wang, 2022: 130, figs. 3–8, 10–11, 13–24. TL: China: Qinghai. TD: YU. Distribution China: Qinghai. Remarks. This species is distinguished from other two congeners by the upper eye lobes widely separated., Published as part of Lin, Mei-Ying & Li, Kai-Qin, 2022, A new species, Cylindroeme yunnanensis sp. nov. from China (Coleoptera, Cerambycidae, Cerambycinae), pp. 294-300 in Zootaxa 5159 (2) on page 300, DOI: 10.11646/zootaxa.5159.2.7, http://zenodo.org/record/6777338, {"references":["Wang, Y. - Q., Zhao, W. - W. & Xie, G. - L. (2022) A revision of the genus Cylindroeme Vives (Coleoptera, Cerambycidae, Cerambycinae). ZooKeys, 1082, 127 - 134. https: // doi. org / 10.3897 / zookeys. 1082.75816"]}

Genus Anaches Pascoe, 1865 Anaches Pascoe, 1865: 160. Type species: Sthenias dorsalis Pascoe, 1858, by original designation. Remarks. The genus can be divided into two species group by the elytral bands. The “wide banded” species group recently includes four valid species, A. dorsalis (Pascoe, 1858), A. medioalbus (Breuning, 1956), A. murzini (Lazarev, 2020), and A. wenhsini Holzschuh & Lin, 2013. They share the common characters: 1) the whitish pubescent part on elytra nearly or even more than half of elytra length in lateral view; 2) the greyish parts within the pubescent band form two bands, separated by a white pubescent polyline. In this paper, we do not try to solve this “ dorsalis ” species group except the new synonyms. Anaches cylindricus (Gressitt, 1939) neither belong to this species group, nor belong to the following species group which includes the new species. This species resembles to species of the subgenus Albosthenias Breuning, 1961 of the genus Sthenias Dejean, 1835. It won’t be discussed in this paper either. The new species belongs to the “narrow banded” species group, which includes three species, A. albaninus (Gressitt, 1942), A. m-signatus sp. nov., and A. yitingi Holzschuh & Lin, 2013. They share the common characters: 1) the whitish pubescent part on elytra much less than half of elytra length in lateral view, only nearly one third or even less than one third; 2) the greyish parts within the pubescent band form only one transverse band. In this paper, we describe one new species, and make additional descriptions of the male genitalia of the other three species. Distribution China, Vietnam, Laos, Thailand, India, Nepal, Bangladesh., Published as part of Lin, Mei-Ying & Weigel, Andreas, 2022, A study on the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae Pteropliini), with a new species and two new synonyms, pp. 123-132 in Zootaxa 5133 (1) on page 124, DOI: 10.11646/zootaxa.5133.1.6, http://zenodo.org/record/6521516, {"references":["Pascoe, F. P. (1865) Longicornia Malayana; or, a descriptive catalogue of the species of the three longicorn families Lamiidae, Cerambycidae and Prionidae, collected by Mr. A. R. Wallace in the Malay Archipelago. The Transactions of the Entomological Society of London, Series 3, (3) 2, 97 - 224, 4 pls.","Breuning, S. (1956) Die Ostasien-Cerambyciden im Museum A. Koenig, Bonn. Bonner Zoologische Beitrage, 7 (1 - 3), 229 - 236.","Lazarev, M. A. (2020) A new species of genus Sthenias Laporte, 1840 (Coleoptera: Cerambycidae) from Sichuan province of China. Entomologische Blatter und Coleoptera, 116, 057 - 059.","Holzschuh, C. & Lin, Y. - L. (2013) Beitrag zur Bockkaferfauna von Taiwan, mit Beschreibung neuer Arten (Coleoptera, Cerambycidae). In: Lin, M. - Y. & Chen, C. - C. (Eds.), In memory of Mr. Wenhsin Lin. Formosa Ecological Company, Taipei, pp. 147 - 158.","Breuning, S. (1961) Catalogue des Lamiaires du Monde (Col. Ceramb.). Verlag des Museums G. Frey, Tutzing bei Munchen, (4), 183 - 284."]}

Anaches m-signatus Lin & Weigel sp. nov. (Figs. 1–3, 9, 12–13) Description. Male (Figs. 1–2). Length 9.9–12.0 mm, width 3.0–3.6 mm. Body more than 3 times as long as wide; integument black brown to black; surface of head, pronotum, elytra, femora and ventral surface densely covered with short recumbent pubescence, mostly brown or greyish white. Scape with fine recumbent short brown or white pubescence, all antennomeres provided with long brown setae sparsely at ventral side, and sparser toward apical segments. Antennomeres III to XI provided with a whitish basal ring. Occiput, pronotum, basal third and apical fourth of elytra mostly brownish, with some blackish and whitish spots. Elytra with a distinct whitish transverse band near middle, narrowed towards elytral suture, with grayish parts in middle. The anterior margins of the whitish bands on two elytra together form “V”-shaped (Figs. 1a, 2a, 3a). The posterior margin of the whitish band on each elytron forms “m”-shaped (Fig. 3a). Legs with femora and tibiae mostly clothed with grayish pubescence with many black spots; dorsum of tarsal segments densely clothed with fine recumbent black brown or whitish pubescence. Ventral surface of thorax similar to pronotum, metasternum with more grayish pubescence, similar to ventral side of femora. Abdominal sternites (Fig. 1c) mostly clothed with grayish to brownish pubescence except the apical margin of sternite III and the sexual setal patches on sternite IV covered with golden brown long sexual setae (Figs. 1c, 9a, 9b). Head slightly narrower than pronotum. Labrum trapezoidal with rounded angles, covered with golden yellow setae at margin, maxillary palpi brown, apical palpomere conical. Inferior eye lobe slightly longer than gena below it. Antenna with last three segments surpassing elytral apex; relative lengths of segments from base to apex: 15: 4: 20: 19: 18: 18: 18: 17: 17: 16: 16. Pronotum wider than long, 1.2 times as width as long; disc with clear basal and distal transverse grooves (Figs. 1a, 2a) and a middle longitudinal groove (Fig. 2b). Prosternal process moderately constricted between procoxae, then steeply declined toward dilated apex. Procoxal cavities completely closed. Elytra ca. 2.2 times as long as wide, parallel sided before apical tenth, then roundly narrowed to truncated apices; inner angles and outer angles are obtuse. Apical margin of sternite VII curved. Apex of hind femur reaching middle of abdominal sternite V. Female (Figs. 3a–3c). Length 10.2–11.8 mm, width 3.1–3.6 mm. Female is identical to male, with antenna slightly shorter, with last two segments surpassing elytral apex and abdominal sternites quite different. Unlike the male, female abdominal sternites (Fig. 3c) covered with grayish pubescence intermixed by many black spots, without any golden brown long sexual setae. Male terminalia (Figs. 12–13). Tegmen length about 2.0 mm; lateral lobes rather straightly tapered from middle to narrowly rounded apices, each about 0.4 mm long and 0.2 mm wide; median lobe plus median struts slightly curved, subequal to tegmen in length; median struts shorter than half of whole median lobe in length; apex of ventral plate weakly projected (Fig. 13a); median foramen elongate; internal sac with two hook-shaped sclerites (Figs. 13b, 13c). Tergite VIII (Figs. 12a & 12c) trapezoidal, apex truncated with round angles, provided with medium long setae along apical and lateral sides. Diagnosis. This new species belongs to the “narrow banded” species group. It is mostly similar to A. yitingi Holzschuh & Lin, 2013, but can be easily distinguished from it by the following features: the whitish band wider; the anterior margins of the whitish bands “V”-shaped, instead of “U”-shaped; the posterior margin of the whitish band forms “m”-shaped, instead of an oblique line; the apex of tergite VIII truncated, not slightly emarginated; the apex of ventral plate of median lobe not as shaped as that of A. yitingi. Etymology. This species is named after the m-shaped macula margin (Fig. 3a, in the red rectangle) on each elytron. Distribution. China: Zhejiang Province, Hunan Province, Guangxi Province. Type specimens examined. Holotype, 1 ♂, Guangxi Province, Longsheng County, Maoershan (广OiAEüDz 儿山), 2011-VI-3, leg. Xin-Lei Huang (ëadz) (IZCAS, IOZ (E) 2224561). 9 Paratypes: 1 ♂, Guangxi Province, Jinxiu County, Dayaoshan, Gubao (广Oiŵħkä山古保), 850m, 2021-IV-09, leg. J. T. Zhao (CCCC, C21Z0046); 1 ♀, Guangxi Province, Jinxiu County, Dayaoshan, Pingbanshan (广Oiŵħkä山平Ú山), 2016-V-14, leg. Jin-Teng Zhao (ŭffiss) (CCCC, C16Z0698); 1 ♂, Hunan Province, Yanling County, Shennonggu (DZAE ẍĸ县ñƌ ĕ), alt. 640m, 2008-VII-5, leg. Zhuo Yang (杨卓) (IZCAS, IOZ (E) 2224562); 1 ♀, Hunan Province, Yanling County, Shiduxiang, env. Taoyuandongcun (DZAE ẍĸ县十δ乡fiů洞村周B), alt. 870m, 2008-VII-6, leg. Gan-Yan Yang by beating (IZCAS, IOZ (E) 2224563); 2 ♂, 3 ♀, Zhejiang Province, Gutianshan National Nature Reserve (CAW): 1♂: CSP15, 2010, 618m, 118,13°E / 29,25°N, NE2; 1♂: same data, SE1; 1♀: CSP03, 2010, 720m, 118,12°E / 29,24°N, NW2; 1♀: CSP06, 2010, 880m, 118,15°E / 29,25°N, NW2; 1♀: CSP09, 2010, 348m, 118,11°E / 29,24°N, SW3. Remarks. This new species is quite similar to Desisa pterolophioides Gressitt from Fujian, China (Gressitt, 1942a) (now in the genus Macropraonetha Breuning, 1961) by color and pubescence pattern, but can be separated from it by the elytron with one tubercle with tufted hairs before basal one fifth, the elytral apex truncated instead of narrow rounded, and the anterior and posterior margins of white band on elytra with different shapes., Published as part of Lin, Mei-Ying & Weigel, Andreas, 2022, A study on the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae Pteropliini), with a new species and two new synonyms, pp. 123-132 in Zootaxa 5133 (1) on pages 124-126, DOI: 10.11646/zootaxa.5133.1.6, http://zenodo.org/record/6521516, {"references":["Holzschuh, C. & Lin, Y. - L. (2013) Beitrag zur Bockkaferfauna von Taiwan, mit Beschreibung neuer Arten (Coleoptera, Cerambycidae). In: Lin, M. - Y. & Chen, C. - C. (Eds.), In memory of Mr. Wenhsin Lin. Formosa Ecological Company, Taipei, pp. 147 - 158.","Gressitt, J. L. (1942 a) New longicorn beetles from China: VIII (Coleoptera: Cerambycidae). Lingnan Natural History Survey and Museum, Special Publication, 2, 1 - 6.","Breuning, S. (1961) Catalogue des Lamiaires du Monde (Col. Ceramb.). Verlag des Museums G. Frey, Tutzing bei Munchen, (4), 183 - 284."]}

Anaches albaninus (Gressitt, 1942) (Figs. 6–8, 11, 16–17) Pterolophia albanina Gressitt, 1942b: 85, pl. I, fig. 5. Pterolophia albanina: Gressitt, 1951: 464; Hua, 2002: 227. Pterolophia (Hylobrotus) albanina: Breuning, 1961: 252; Löbl & Smetana, 2010: 318; Lin, 2015, 260, 2 figs.; Lin, 2017: 382, pl. 33, fig. 8; Lin & Yang, 2019: 372; Danilevsky, 2020: 455. Anaches albaninus: Lin & Lazarev, 2021: 72. Male terminalia (Figs. 16–17). Tegmen length about 2.5 mm; lateral lobes rather straightly tapered from middle to narrowly rounded apices, each about 0.5 mm long and 0.2 mm wide; median lobe plus median struts slightly curved, subequal to tegmen in length; median struts shorter than half of whole median lobe in length; apex of ventral plate pointed (Fig. 17a); median foramen elongate; internal sac with 2 hook-shaped sclerites (Figs. 17b, 17c). Tergite VIII (Figs. 16a & 16c) semicircled, apex widely rounded, provided with medium long setae along apical and lateral sides. Diagnosis. A. albaninus differs from the new species A. m-signatus sp. nov. by following features: different withish elytral band (much narrower, the anterior border not so oblique, and posterior border not “m”shaped), elytral punctuation weaker, and pro-femora in males slender. Type specimens examined. Holotype, ♀, Zhejiang, T’ienmushan, 1937-V-21, leg. O. Piel (IZCAS, IOZ (E) 217646). Other specimens examined. Shaanxi: 1 ♂, Foping, Longcaoping, alt. 1256m, 2008-VII-3, leg. Ming Bai (IZCAS); 1 ♀, Zhouzhi, Houzhenzi, alt. 1280m, 2008-V-5-6, leg. Hao Huang (IZCAS). Anhui: 1 ♀, Huangshan, 1936-VI-24 (IZCAS); Zhejiang: 1 ♂, 1 ♀, T’ienmushan, 1936-VI-25, leg. O. Piel (IZCAS); 1 ♀, Tianmushan, 1937- VII-19 (IZCAS); 1 ♂, Hangzhou, Lin’an, Qingliangfeng Ziranbaohuqu (Nat. Rev.), Shunxiwu, 450m, 118°56'19.51" E, 30°02'38.47" N, 2012-VI-24, leg. Hao Xu & Jian-Yue Qiu by malaise trap (MYNU); 1 ♂, Hangzhou, Lin’an, Xitianmushan, 2021-VII-13–15, leg. Tao Li (CSJ); 1 ♀, Anji, Longwangshan, alt. 490m, 1996-VI-12, leg. Xing-Ke Yang (IZCAS); 1 ♀, Lishui, Yujikengbaohuzhan, alt. 1371m, 27°41′41.77″N, 119°35′26.50″E, 2017-VII-10, leg. Yan-Dong Chen (IZCAS); 2 ♂, Zhejiang Province, Gutianshan National Nature Reserve (CAW): 1 ♂: CSP18, 2010, 569m, 118,12°E / 29,25°N, SE2; 1 ♂: CSP27, 2010, 665m, 118,14°E / 29,25°N, NE2. Jiangxi: 1 ♂, Ku-ling, 1935-VII-25, leg. O. Piel (IZCAS). Fujian: 1 ♂, Chong’an, Guadun, 1973-VI-11, leg. Pei-Yu Yu (IZCAS); 1 ♂, 1 ♀, Jianyang, Aotou, 1973-V-27-28, leg. Pei-Yu Yu (IZCAS). Distribution. China: Henan, Shaanxi, Gansu, Jiangsu, Anhui, Zhejiang, Hubei, Jiangxi, Hunan, Fujian, Guangxi, Sichuan. Remarks. The record from Inner Mongolia by Xu & Neng (2010) is a misidentification, we do not include it. The records from Heilongjiang, Hebei by Hua (2002), repeated by other authors (Löbl & Smetana 2010, Lin 2015, Lin 2017, Lin & Yang 2019, Lin & Lazarev 2021) are doubtful, so we do not include them., Published as part of Lin, Mei-Ying & Weigel, Andreas, 2022, A study on the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae Pteropliini), with a new species and two new synonyms, pp. 123-132 in Zootaxa 5133 (1) on pages 128-130, DOI: 10.11646/zootaxa.5133.1.6, http://zenodo.org/record/6521516, {"references":["Gressitt, J. L. (1942 b) Nouveaux longicornes de la Chine orientale. Notes d'Entomologie Chinoise, 9, 79 - 97, 2 pls.","Gressitt, J. L. (1951) Longicorn beetles of China. In: Lepesme, P. (Ed.), Longicornia, etudes et notes sur les longicornes. Vol. 2. Paul Lechevalier, Paris, pp. 1 - 667, 22 pls.","Hua, L. - Z. (2002) Cerambycidae. In: Hua, L. - Z. (Ed.), List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, pp. 189 - 237.","Breuning, S. (1961) Catalogue des Lamiaires du Monde (Col. Ceramb.). Verlag des Museums G. Frey, Tutzing bei Munchen, (4), 183 - 284.","Lobl, I. & Smetana, A. (2010) Catalogue of Palaearctic Coleoptera. Vol. 6. Chrysomeloidea. Apollo Books, Stenstrup, 924 pp. https: // doi. org / 10.1163 / 9789004260917","Lin, M. - Y. (2015) Album of Type Specimens of Longhorn Beetles Deposited in National Zoological Museum of China. Henan Science and Technology Press, Zhengzhou, 374 pp.","Lin, M. - Y. (2017) Insect Fauna of the Qinling Mountains. Vol. VI. Coleoptera II. Cerambycid-beetles. World Publishing Corporation, Xi'an, 510 pp., 37 pls.","Lin, M. - Y. & Yang, X. - K. (2019) Catalogue of Chinese Coleoptera. Vol. IX. Chrysomeloidea: Vesperidae, Disteniidae, Cerambycidae. Science Press, Beijing, 575 pp.","Danilevsky, M. L. (Ed.) (2020) Catalogue of Palaearctic Coleoptera. Vol. 6 / 1. Chrysomeloidea I (Vesperidae, Disteniidae, Cerambycidae). Updated and Revised 2 nd Edition. Brill, Leiden, 712 pp. https: // doi. org / 10.1163 / 9789004440333","Lin, M. - Y. & Lazarev, M. A. (2021) Four new combinations in the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae, Pteropliini). Humanity space International almanac, 10 (1), 70 - 76.","Holzschuh, C. & Lin, Y. - L. (2013) Beitrag zur Bockkaferfauna von Taiwan, mit Beschreibung neuer Arten (Coleoptera, Cerambycidae). In: Lin, M. - Y. & Chen, C. - C. (Eds.), In memory of Mr. Wenhsin Lin. Formosa Ecological Company, Taipei, pp. 147 - 158.","Xu, P. & Neng, N. (2010) Coloured Illustrations of Longhorned Beetles in Mongolian Plateau. Chinese Agricultural University Press, Beijing, 150 pp."]}

Anaches yitingi Holzschuh & Lin, 2013 (Figs. 4–5, 10, 14–15) Anaches yitingi Holzschuh & Lin, 2013: 154, fig. 10. Anaches yitingi: Lin, 2015, 256, 2 figs.; Lin & Yang, 2019: 362; Danilevsky, 2020: 449; Lin & Lazarev, 2021: 74. Male terminalia (Figs. 14–15). Tegmen length about 2.0 mm; lateral lobes rather straightly tapered from middle to narrowly rounded apices, each about 0.4 mm long and 0.2 mm wide; median lobe plus median struts slightly curved, slightly longer than tegmen in length; median struts shorter than half of whole median lobe in length; apex of ventral plate strongly projected (Fig. 15a); median foramen elongate; internal sac with 2 hook-shaped sclerites (Figs. 15b, 15c). Tergite VIII (Figs. 14a & 14c) trapezoidal, apex slightly emarginated with round angles, provided with medium long setae along apical and lateral sides. Diagnosis. This species is mostly similar to A. albaninus (Gressitt, 1942), but can be easily distinguished from it by the following features: the whitish band more oblique; the anterior margins of the whitish bands “U”-shaped, instead of slightly oblique line; the sexual patches on sternite IV closer to each other (Figs. 10a, 10b), instead of well separated (Figs. 11a, 11b); the apex of tergite VIII emarginated (Figs. 14a, 14c), instead of rounded (Figs. 16a, 16c); the apex of ventral plate of median lobe projected (Fig. 15a), instead of pointed (Fig. 17a). Type specimens examined. 1 ♂, 1 ♀, paratypes, Taiwan, Pingtung County, Mt. Dahan, 2007-V-26, leg. Wenhsin Lin (IZCAS, IOZ (E) 1905283–84). Distribution. China: Taiwan.

Anaches medioalbus (Breuning, 1956) (Figs. 18–21) Paramesosella medioalba Breuning, 1956: 234. Sthenias semicylindricus Hayashi, 1974: 45. syn. nov. Paramesosella medioalba; Löbl & Smetana, 2010: 316; Lin & Yang, 2019: 369; Danilevsky, 2020: 453. Anaches semicylindricus: Holzschuh & Lin, 2013: 152; Lin & Yang, 2019: 362; Danilevsky, 2020: 449; Lin & Lazarev, 2021: 74. Sthenias (s. str.) murzini Lazarev, 2020: 57, figs. 1-3. syn. nov. Anaches medioalbus: Lin & Lazarev, 2021: 73. Anaches murzini: Lin & Lazarev, 2021: 74. Remarks. According to the type pictures of Sthenias semicylindricus Hayashi, 1974 (Figs. 20-21), it is similar to A. medioalbus (Breuning, 1956) (Figs. 18-19), by the body shape, elytral apex shape, antennal segments, main pubescence maculae on elytra, with differences on irregular grayish white spots over pronotum and elytra. However, the pair of type specimens were obviously rubbed, which was the reason of lacking the white spots. Compared with a fresh specimen from Taiwan (Fig. 7 by Holzschuh & Lin 2013) and specimens from Fujian (Fig. 19), we cannot find differences between the two taxa, so we propose the synonymy. This is agreed by Carolus Holzschuh (personal communication). However, Wen-Xuan Bi proposes a more complicated opinion, that all the records of A. dorsalis from Mainland China were misidentifications, and all of them, including the Anaches medioalbus from Fujian, should be Anaches albonotatus Pic, 1932. We agreed with Wen-Xuan Bi that the specimens in IZCAS we examined are not the true A. dorsalis and we re-identify them as A. medioalbus (Breuning, 1956) in this paper. To clarity the true identity of A. albonotatus described from northern Vietnam (Tonkin) it needs more study. Holzschuh (2021) synonymized Sthenias murzini Lazarev, 2020 with Sthenias dorsalis Pascoe, 1858, which is the type species of the genus Anaches Pascoe, 1865. However, Anachus murzini (Lazarev, 2020) is not a synonym of A. dorsalis (Pascoe, 1858), because it is much closer to A. medioalbus (Breuning, 1956) than to A. dorsalis (Pascoe, 1858), which has been misunderstood for a long time. The true A. dorsalis (Pascoe, 1858) is endemic to India and nearby. We understood that the “ A. dorsalis ” in Holzschuh (2021) is infact A. medioalbus in this paper, and we agreed with him on the variations of elytral maculae and the shape of elytral apex both are infrasubspecific features, therefore we synonymized A. murzini (Lazarev, 2020) with A. medioalbus (Breuning, 1956) herein. Type specimens examined. Holotype of Paramesosella medioalba Breuning, 1956 (Figs. 18a, 18b), ♀, Kwangtseh-Fukien, 1937-VIII-19 (ZFMK); Holotype and paratype of Sthenias semicylindricus Hayashi, 1974, holotype (Fig. 21), ♂, Taiwan, Mizuho, 1972-VII-6, leg. Mizunuma (OSAKA, M. Hayashi Coll. OMNH [98-32]); paratype, 1 ♀ (Fig. 20), Taiwan, Mizuho, 1972-VII-5, leg. Mizunuma (OSAKA, M. Hayashi Coll. OMNH [98- 32]). Other specimens examined. Zhejiang: 1 ♂, Tianmushan, 350 m, 1999-VI-5, leg. Ming-Yuan Gao (IZCAS, IOZ (E) 1896988); 1 ♂, 1 ♀, Hangzhou, Lin’an, Qingliangfeng Ziranbaohuqu (Nat. Rev.), Shunxiwu, 450m, 118°56'19.51" E, 30°02'38.47" N, 2012-VI-24, leg. Hao Xu & Jian-Yue Qiu by malaise trap (MYNU). Fujian: 1 ♀, Huangkeng, 1965-VII (IZCAS); 1 ♀, Jianxi, 1981-IV-21 (IZCAS). Guangxi: 1 ♀, Jinxiu, Fenzhan, alt. 800 m, 1999-V-13, leg. Hui Xiao (IZCAS, IOZ (E) 1896989). Guizhou: 1 ♀, Guiyang, Huaxi, 2011- VI (IZCAS). Distribution China: Shaanxi, Zhejiang, Fujian, Taiwan, Hong Kong, Guangxi, Chongqing, Sichuan, Guizhou, Yunnan., Published as part of Lin, Mei-Ying & Weigel, Andreas, 2022, A study on the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae Pteropliini), with a new species and two new synonyms, pp. 123-132 in Zootaxa 5133 (1) on page 130, DOI: 10.11646/zootaxa.5133.1.6, http://zenodo.org/record/6521516, {"references":["Breuning, S. (1956) Die Ostasien-Cerambyciden im Museum A. Koenig, Bonn. Bonner Zoologische Beitrage, 7 (1 - 3), 229 - 236.","Lobl, I. & Smetana, A. (2010) Catalogue of Palaearctic Coleoptera. Vol. 6. Chrysomeloidea. Apollo Books, Stenstrup, 924 pp. https: // doi. org / 10.1163 / 9789004260917","Lin, M. - Y. & Yang, X. - K. (2019) Catalogue of Chinese Coleoptera. Vol. IX. Chrysomeloidea: Vesperidae, Disteniidae, Cerambycidae. Science Press, Beijing, 575 pp.","Danilevsky, M. L. (Ed.) (2020) Catalogue of Palaearctic Coleoptera. Vol. 6 / 1. Chrysomeloidea I (Vesperidae, Disteniidae, Cerambycidae). Updated and Revised 2 nd Edition. Brill, Leiden, 712 pp. https: // doi. org / 10.1163 / 9789004440333","Holzschuh, C. & Lin, Y. - L. (2013) Beitrag zur Bockkaferfauna von Taiwan, mit Beschreibung neuer Arten (Coleoptera, Cerambycidae). In: Lin, M. - Y. & Chen, C. - C. (Eds.), In memory of Mr. Wenhsin Lin. Formosa Ecological Company, Taipei, pp. 147 - 158.","Lin, M. - Y. & Lazarev, M. A. (2021) Four new combinations in the genus Anaches Pascoe, 1865 (Coleoptera, Cerambycidae, Lamiinae, Pteropliini). Humanity space International almanac, 10 (1), 70 - 76.","Lazarev, M. A. (2020) A new species of genus Sthenias Laporte, 1840 (Coleoptera: Cerambycidae) from Sichuan province of China. Entomologische Blatter und Coleoptera, 116, 057 - 059.","Pic, M. (1932) Nouveautes diverses. Melanges Exotico-Entomologiques, 60, 1 - 36.","Holzschuh, C. (2021) Neue Synonymien, Neumeldungen fur China und Beschreibung von zehn neuen Bockkafern aus Asien (Coleoptera, Cerambycidae). Les Cahiers Magellanes, NS, 41, 84 - 105.","Pascoe, F. P. (1865) Longicornia Malayana; or, a descriptive catalogue of the species of the three longicorn families Lamiidae, Cerambycidae and Prionidae, collected by Mr. A. R. Wallace in the Malay Archipelago. The Transactions of the Entomological Society of London, Series 3, (3) 2, 97 - 224, 4 pls."]}

Pseudomeges marmoratus (Westwood, 1848) (Figs. 3, 10, 12) Hammaticherus marmoratus Westwood, 1848: 11, pl. V, fig. 1. Type locality: Himalayan region. Merges [sic] marmoratus: Lacordaire, 1869: 311. Pseudomeges marmoratus: Breuning, 1943: 157; 1944: 300. Material examined. China (Xizang): 2 males, Xizang, Chayu, Zhuowagong, 2,500 m, 2011.VII.1, leg. Wen-Xuan Bi (CBWX); 1 male, Xizang, Motuo, 1,200 m, 2011.VIII.18, leg. Wen-Xuan Bi (CBWX); 1 male, ditto except 1,100m, 2013.VIII.19, leg. Wen-Xuan Bi (CBWX); 3 males, ditto except 2014.VII.25 (CBWX); 1 female, Xizang, Motuo, 108K, 1,400 m, 2013.VIII.16, leg. Wen-Xuan Bi (CBWX); 1 female, Xizang, Bomi County, Tongmaizhen, Tongmaidaqiao, alt. 2,025m, 30.09632˚N, 95.06610˚E, 2019.VII.19, leg. Hao-Dong Yin by light trap (IZAS). China (Yunnan): 1 female, Yunnan, Longchuan, Mangdong, 1,770 m, 2015.IX.19, leg. Y.-T. Chung (CCCC); 1 female, Yunnan, Ruili, Bangdashan, 1,432 m, 2016.IX.17, leg. Xiao-Dong Yang (CCCC); 1 male, Yunnan, Longchuan, Husa Xiang, 1,700 m, 2017.VI.24, leg. Yung-Jen Chang (CCCC). Complementary description. Male. Body length 40.5–61.3 mm, humeral width 12.3–19.3 mm. Legs with tarsi four-segmented, but tarsomeres IV and V in some individuals incompletely fused (Fig. 12). Male endophallus (Fig. 10) stout, about 2.4 times as long as the median lobe, weakly curved near apical two-fifths dorsally; APH well defined, BPH and MPH vaguely defined due to the highly reduced cs; MPH subdivided into MT, CT and PB by constriction, CT and PB moderately swollen at middle; PB with pronounced si and indistinct sf; relative lengths of APH: MPH: BPH: PB: CT: MT = 1.0: 2.9: 0.5: 0.6: 1.0: 1.3. APH, divided into ab and bb; im moderately developed; ab gradually broadened distally; bb provided with a pair of rod-like sclerites associated with the paired ejaculatory ducts, in dorsal view (Fig. 10a) protuberant behind the widely separated gn and deeply emarginated apically. Spicules mainly distributed on CT, apical part of PB and whole surface of ab. Female. Body length 57.4–66.7 mm, humeral width 19.6–22.3mm. Distribution. China (new Country record): Xizang, Yunnan; Laos, India, Myanmar, Bhutan. Remarks. This species and Pseudomeges varioti Le Moult, 1946 show allopatric distribution in Yunnan. The former is distributed in the westernmost region (Longchuan County), while the latter was reported from the southern end (Jinghong City: Weigel, Meng & Lin 2013). Pseudomeges marmoratus can be distinguished from the latter by its head smaller, not protruding at sides in front view, lower eye lobe larger, nearly as long as gena, instead of only 0.6 times, antennal scape thicker, antennomere III straight, antennomeres III to V not annulated with light pubescence at base, elytra longer compare to its head and pronotum united, scutellum slender, not transverse., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on page 247, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Westwood, J. O. (1848) The cabinet of oriental entomology; being a selection of rare and beautiful insects, natives of India and the adjacent islands. W. Smith, London ii + 88 pp., 42 pls. https: // doi. org / 10.5962 / bhl. title. 34273","Lacordaire, J. T. (1869) Histoire naturelle des insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome neuvieme. Premiere partie. Paris: Librairie encyclopedique de Roret, 409 pp.","Breuning, S. (1943) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (89 - 106), 137 - 280.","Breuning, S. (1944) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (107 - 135), 281 - 512.","Le Moult, E. (1946) Description d'un nouveau Lamiaire de l'Annam (Col., Cerambycidae). Miscellanea Entomologica 43 (8), 136.","Weigel, A., Meng, L. - Z. & Lin, M. - Y. (2013) Contribution to the Fauna of Longhorn Beetles in the Naban River Watershed National Nature Reserve. Formosa Ecological Company, Taiwan, 219 pp."]}

Meges gravidus (Pascoe, 1858) (Figs. 1, 6, 8) Monohammus gravidus Pascoe, 1858: 245. Type locality: North China. Meges gravidus: Pascoe, 1866: 272. Merges [sic] gravidus: Lacordaire, 1869: 311. Apriona multimaculata Pic, 1933: 31. Type locality: Tche-Fou (Yantai), Shandong, China. Synonymized by Breuning, 1944: 445 Type material examined. Holotype, female, “ Monochamus / gravidus / type Pasc ”, “ Meges / gravidus / Pas ”, “ Borneo? ”, “Type” (NHML). Examined by photographs taken by Xicui Du in 2013. Other material examined. China (Henan): 1 female, Henan (IZAS). China (Anhui): 1 female, Anhui, Yuexi, Yaoluoping, Menkanling, 1,100 m, 2021.VII.7, leg. Wen-Xuan Bi (CBWX); 1 female, ditto except 1,129 m, 2021. VII.6, leg. J.-T. Zhao (CCCC). China (Zhejiang): 1 male, Zhejiang, Xitianmushan, 350 m, 2005.VII.9, leg. Wen-Xuan Bi (CBWX); 1 female, ditto except 2006.VII.1-9 (CBWX); 1 female ditto except 2009.VIII.18, leg. Liang Tang (CBWX); 1 male, ditto except 2016.VII.4, leg. Wen-Xuan Bi (CBWX); 1 female, ditto except 2016.VII.5 (CBWX); 1 female, ditto except 2016.VII.24, leg. Yong-Xiang Wu (CBWX); 1 male, ditto except 1,250 m, 2016. VII.29, leg. Wen-Xuan Bi (CBWX); 1 male, ditto except 510 m, 2016.VIII.6 (CBWX); 1 female, ditto except 350 m, 2018.VII.7, leg. Xiao-Bin Song (CBWX); 1 female, Zhejiang, Hangzhou, 1978.VI (IZAS, IOZ (E) 1896993); 1 male, Zhejiang, Anji, Baofuzhen, Lidongwu, 750 m, 2017.VIII.11, leg. Wen-Xuan Bi (CBWX); 1 male, Zhejiang, Ningbo, Tiantongshan, 2009.VI, leg. Bin-Jie Ge (CBWX). China (Hunan): 1 male, Changsha, Yuelushan, 1957. VIII.2 (IZAS). China (Fujian): 1 female, Fujian, Quanzhou, Dehuaxian, Chishuizhen, leg. Jia-Hong Lin & Nan-Yi Tsai (CCCC); 1 male, Fujian, Wuyishan, Sangang, 740 m, 1997.VIII.5, leg. Jian Yao (IZAS). China (Guangxi): 1 female, Guangxi, Xingan, Jinshi, 1,230 m, 2017.VII.17, leg. Y.-Q. Lu (CCCC). Complementary description. Male (Fig. 1). Body length 37.3–40.4 mm, humeral width 11.1–12.1 mm. Male endophallus (Fig. 8) long and slender, more than 3 times as long as the median lobe, strongly curved dorsally near apical third; APH defined by the presence of af, limit between BPH and MPH indicated by a dorsal swelling; cs highly reduced, nearly unrecognizable; MPH vaguely subdivided into MT, CT and PB, without significant swelling or protuberance; PB surrounded by sf and provided with si; relative lengths of APH: MPH: BPH: PB: CT: MT = 1.0: 8.4: 1.4: 1.1: 2.8: 4.7. APH slightly curved ventrally at base with im complete and subsclerotized, clearly divided into ab and bb; ab cylindrical, moderately swollen; bb constricted near middle, provided with a pair of rodlike sclerites associated with the paired ejaculatory ducts (Fig. 5a), with gn situated near the apex. Spicules mainly distributed on dorsal surface or along a narrow ventral midline of CT, around the distal part of PB and the whole surface of ab. Female. Body length 39.9–47.2 mm, humeral width 11.8–13.6 mm. Distribution. China: Shandong, Henan, Shaanxi, Anhui, Zhejiang, Hunan, Fujian, Guangxi (new Province record). Remarks. This species was originally described from Borneo (Pascoe 1858, label of the holotype interpreted as “Borneo?”), but no additional specimen has been reported from there since. Study of series specimens from several localities of China suggest that the "North China " proposed in Pascoe (1866) is the correct type locality of this species. Thus, this species is accordingly excluded from the fauna of Borneo., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on page 244, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Pascoe F. P. (1858) On new genera and species of longicorn Coleoptera. Part III. The Transactions of the Entomological Society of London, (2) 4 (6 - 7), 236 - 266, pls XXV, XXVI. https: // doi. org / 10.1111 / j. 1365 - 2311.1858. tb 01823. x","Pascoe, F. P. (1866) Longicornia Malayana; or, a descriptive catalogue of the species of the three longicorn families Lamiidae, Cerambycidae and Prionidae, collected by Mr. A. R. Wallace in the Malay Archipelago. The Transactions of the Entomological Society of London, (3) 3, 225 - 336.","Lacordaire, J. T. (1869) Histoire naturelle des insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome neuvieme. Premiere partie. Paris: Librairie encyclopedique de Roret, 409 pp.","Pic, M. (1933) Descriptions et notes sur divers Cerambycides [Col.]. Bulletin de la Societe Entomologique de France, 38 (2), 30 - 32. https: // doi. org / 10.3406 / bsef. 1933.14517","Breuning, S. (1944) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (107 - 135), 281 - 512."]}

Pseudomeges aureus Bi, Chen & Lin sp. nov. (Figs. 4, 11) Type material. Holotype: male, “ Yunnan, Pingbian, Daweishan / 2021.IV.9 / leg. Jin Chen ” “ 2041 m / 22˚54.6524̓ N / 103˚41.8066̓ E” (IZAS). Paratypes: 1 male, same data to holotype; 2 males, “ Yunnan, Pingbian, Daweishan / 2,022 m / 2016.IV.22 / leg. Xiao-Dong Yang ” (CCCC); 1 male, ditto except 1,982 m, 2016.IV.21, leg. Chao Wu (CCCC). Description. Male (Fig. 4). Body length 52.6–58.1 mm, humeral width 15.9–18.1 mm. Integument of body, elytra and appendages blackish. Head with mandibles, frons, genae and vertex moderately covered with fine pale yellow pubescence. Antennae with scape, pedicel and the base of antennomere III to V vaguely covered with similarly colored pubescence, which gives the entire antennae a uniformly darkened appearance. Pronotum mostly covered with similarly colored pubescence as head but denser, except for the tip of lateral spine, a discal median short line and two small spots at sides of the former line which are glabrous. Scutellum covered with the same pubescence as pronotum. Elytra predominantly covered with dense pale pubescence, plus some dark golden colored pubescence forming irregular pattern over the whole elytron. Ventral surface and legs covered with moderately dense yellowish pubescence, relatively denser on posterior margin of prosternum, entire metanepisternum, posterior margin of metaventrite and anterior margin of abdominal ventrite I. Body elongate. Head slightly wider than pronotal base; moderately large, protruding toward sides near the lower eye lobe in front view, thence gradually convergent anteriorly; lower eye lobe 0.9 times as long as gena; frons ca. 1.7 times wider than high, with a fine median groove extending from anterior margin to occiput; vertex moderately concave, with prominent antennal insertions. Antennae ca. 1.9 times as long as body length, with apical 5 antennomeres surpassing elytral apices; antennomere III 1.4 times as long as scape, slightly longer than IV, antennomere IV to X gradually decreasing in length and width, antennomere X slightly longer than XI. Pronotum cylindrical, more than 0.8 times as long as the basal width; lateral spines conical with sharply pointed apex, slightly curved backwards; disk smooth, weakly convex, provided with an indistinct postmedian callus and two lateral calli slightly behind the middle; with a few setigerous punctures at both sides of the postmedian callus and behind the lateral spines. Scutellum subequal in length and width, broadly rounded posteriorly. Elytra elongate, widest across humeri, ca. 1.7 times as wide as pronotal base, 2.3 times as long as the humeral width; subparallel-sided in most of its length, then gradually convergent toward separately rounded apices; disk smooth, sparsely punctate mainly at base or near the suture, with two weak longitudinal carinae extending from the base to apical one-fifth. Abdomen with ventrite I about twice as long as each of the following ventrites. Legs moderately long and stout, metafemora extending ventrite IV in ventral view. Endophallus in everted condition (Fig. 11) similar to Pseudomeges marmoratus as described above except BPH distinctly swollen and bb of APH less protruding near middle in dorsal view. Female. Unknown. Etymology. From the Latin aureus, meaning gold-colored, referred to the appearance of the elytral maculae. Distribution. China: Yunnan (Pingbian County). Remarks. The relatively more widely distributed species Pseudomeges varioti Le Moult, 1946 has the coloration or pattern of the elytral pubescence variable and some individuals are similar to this new species. But this new species can be distinguished from males of P. varioti by the body relatively slenderer; head smaller, in front view only protruding laterally near the lower eye lobe (instead of protruding in most of the head height); lower eye lobe larger, about 0.9 times as long as gena (instead of 0.6 times); antennae indistinctly annulated; scutellum not transverse; elytra relatively narrower, about 2.3 times as long as the humeral width (instead of 1.9–2.0 times)., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on pages 248-249, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Le Moult, E. (1946) Description d'un nouveau Lamiaire de l'Annam (Col., Cerambycidae). Miscellanea Entomologica 43 (8), 136."]}

Pseudomeges varioti Le Moult, 1946 (Fig. 5) Pseudomeges varioti Le Moult, 1946: 136. Type locality: Dalat, Annam, Vietnam. Pseudomeges gigas Lepesme, 1947: 19, fig. 1. Type locality: Tranninh, Xieng-Khouang, Laos. Synonymized by Weigel, 2012: 411. Material examined. China (Guangdong): 1 female, Guangdong, Nanling, 2010, leg. Li-Zhen Li (EUMJ); China (Yunnan): 1 female, Yunnan, Menghai County, Nanbanhe Nat. Rev., Guomenshan Station, 1120 m, 22.24497°N, 100.60635°, 2011.X.12, leg. Zhi-Ming Ma (IZAS). Thailand: 1 male, Thailand, Chiang Mai, Doi Inthanon, 1989. V.24, local collector (EUMJ); 1 female, Thailand, Chiang Mai, Doi Inthanon, 1988.VIII.6 (EUMJ); 1 male, ditto except 1988.VIII.22 (EUMJ); 1 male, ditto except 1989.V.24 (EUMJ); 1 female ditto except 1989.IX.9 (EUMJ). Myanmar: 1 male, S. E. Burma, Dauna, 1991.X.5, local collector (EUMJ). Laos: 1 female, Laos, Phu Soai Dao, 1992.IV.16, local collector (EUMJ). Distribution. China: Guangdong (new Province record), Hainan, Sichuan, Yunnan; Vietnam; Laos; Myanmar (new country record). Remarks. Weigel, Meng & Lin (2013) reported P. varioti from Xizang based on some pictures from internet and personal communications with collectors, but it was due to a misidentification of P. marmoratus. Therefore P. varioti is excluded from the fauna of Xizang., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on pages 247-248, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Le Moult, E. (1946) Description d'un nouveau Lamiaire de l'Annam (Col., Cerambycidae). Miscellanea Entomologica 43 (8), 136.","Lepesme, P. (1947) Deux remarquables Cerambycides [Col.] nouveaux d'Indochine. Bulletin de la Societe Entomologique de France, Paris, 52, 19 - 21, 2 figs. https: // doi. org / 10.3406 / bsef. 1947.15935","Weigel, A., Meng, L. - Z. & Lin, M. - Y. (2013) Contribution to the Fauna of Longhorn Beetles in the Naban River Watershed National Nature Reserve. Formosa Ecological Company, Taiwan, 219 pp."]}

Meges tonkineus (Clermont, 1932) comb. nov. (Figs. 2, 7, 9) Magninia tonkinea Clermont, 1932: 215, fig. on page 215. Type locality: Tien-Yen, Tonkin, Vietnam. Type material examined. Holotype, female, “ Tonkin / Tien-yen ”, “ex. Jeanvoine / Coll. Clermont ”, “ Type ”, “ Magninia / tonkinea / Clerm. / type ”, “ tonkinea Clerm ”, “TYPE”, “ Museum Paris / Coll. M. Pic ” (MNHN). Examined by photographs taken by Gan-Yan Yang & Hong-Liang Shi in 2011. Other material examined. China (Guangxi): 1 male, Guangxi, Jinxiu, Dayaoshan, 900 m, 2018.VIII.11, local collector (CCCC). China (Hainan): 1 male, Hainan, Ledong, Jianfengling, 950 m, 2015.V.7-12, leg. Yu-Tang Wang & Nan-Yi Tsai (CCCC); 1 female, ditto except 2018.V.21, local collector (CCCC); 1 female, ditto except 2018.VI.2, local collector (CCCC). Complementary description. Male (Fig. 2). Body length 48.5–49.4 mm, humeral width 14.6–15.0 mm. General appearance similar to the female described by Clermont (1932). Body slightly slenderer. Antenna about 2.0 times of body length. Legs relatively longer and thicker. Endophallus in everted condition (Fig. 9), almost identical to Meges gravidus as described above, but relatively slenderer. Female. Body length 55.0– 57.3 mm, humeral width 16.6–17.8 mm. Distribution. China: Hainan (new Province record), Guangxi; Vietnam: “ Tonkin ”. Remarks. The resemblances of the endophallic structures between this species and Meges gravidus (Figs. 8. 9.), indicate a close relationship. This species can be easily distinguished from the latter by its larger body size, antennae longer, protibial subapical tooth sharper, elytra broader with smooth base which lacks remarkable granule, elytral humeri remarkably spined and elytral disk bearing fewer maculae of relatively lighter color. Huang et al. (2002) firstly recorded this species from Guangxi, China. We confirmed the distribution in Guangxi and added Hainan as a new locality based on new material., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on pages 245-247, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Clermont, J. (1932) Description d'un genre et d'une espece nouveaux de Cerambycidae de la sous-famille des Lamiinae [Col.]. Bulletin de la Societe Entomologique de France, 37 (15), 214 - 216. https: // doi. org / 10.3406 / bsef. 1932.14460","Pascoe F. P. (1858) On new genera and species of longicorn Coleoptera. Part III. The Transactions of the Entomological Society of London, (2) 4 (6 - 7), 236 - 266, pls XXV, XXVI. https: // doi. org / 10.1111 / j. 1365 - 2311.1858. tb 01823. x","Westwood, J. O. (1848) The cabinet of oriental entomology; being a selection of rare and beautiful insects, natives of India and the adjacent islands. W. Smith, London ii + 88 pp., 42 pls. https: // doi. org / 10.5962 / bhl. title. 34273","Huang, J. - H., Zhou, S. - Y. & Wang, S. - N. (2002) A checklist of Cerambycidae from Mao'er Mountain Natural Reserve, Guangxi (Coleoptera: Cerambycidae). Journal of Guangxi Normal University, 20 (3), 64 - 68."]}

Meges Pascoe, 1866 stat. res. Meges Pascoe, 1866: 272. Type species: Monochamus gravidus Pascoe, 1858, by monotypy. Magninia Clermont, 1932: 215. Type species: Magninia tonkinea Clermont, 1932, by monotypy. Syn. nov. Redescription. Body elongate and stout, medium to large (ca. 37–57 mm long). Head width subequal to pronotal width at base. Eyes coarsely faceted, emarginate; lower lobes vertical. Frons wider than long. Antennal tubercles moderately prominent and separated. Antennae ca. 1.7–2.0 times (in male) or 1.3–1.4 times (in female) as long as body length; scape with apical cicatrix developed; antennomere III longest, about twice as long as scape, distinctly longer than antennomere IV, antennomeres IV to X successively shorter and narrower, last antennomere slightly longer than penultimate; basal 6–7 antennomeres very sparsely fringed beneath. Both maxillary and labial terminal palpomeres fusiform. Pronotum transverse, cylindrical; each side with a long sharp lateral spine, thickened at base and curved backwards; disk weakly convex, with three indistinct calli arranged as inverted triangle; procoxal cavities closed posteriorly. Scutellum broadly rounded posteriorly. Elytra elongate, subparallel-sided in basal half, gradually convergent toward conjointly rounded apices with long sutural spine; disk provided with two more or less distinct longitudinal carinae. Mesocoxal cavities open to mesepimera. Legs moderately long, stout; protibia with a subapical tooth beneath (weak in females); mesotibia with a subapical oblique groove externally; tarsus four segmented; tarsal claws free, divaricate. Male endophallus with cs highly reduced or nearly absent; MPH long and slender, without significant swelling or protuberance; PB provided with sf and si; APH provided apically with a pair of rod-like sclerites associated with the paired ejaculatory ducts. Distribution. China, Vietnam. Remarks. The genus Meges was proposed by Pascoe (1866) for Monochamus gravidus Pascoe, 1858. The genus Magninia Clermont, 1932 was established based on M. tonkinea Clermont, 1932. Clermont (1932) differentiated this genus from Meges by “les deux sillons oculaires” (“the two eye furrows”, meaning uncertain) less oblique, the scutellum more widely rounded and frons with a pair of deep comma-shaped dimple anteriorly (=pretentorial pit). Breuning (1944) synonymized Meges Pascoe, 1866 with Monochamus Dejean, 1821 without providing any reason, but retained Magninia as valid. In his key, Breuning (1944) separated Magninia from Monochamus mainly by its elytra provided with humeral spines. However, based on the comparison of high-quality photographs of the type specimens of Monochamus gravidus and Magninia tonkinea, and the investigation of fresh material of both species, no significant difference can be found to support their generic separation, especially regarding the considerable similarities of their endophallic structures (Figs. 5, 6). Therefore, Magninia Clermont, 1932 is herein synonymized with Meges Pascoe, 1866. In addition, the genus Meges is proposed herein to be resurrected from synonymy with Monochamus Dejean, 1821 by combination of the following characters: body stout, antennae relatively short, pronotal lateral spines long and sharp, curved backwards, elytra carinated, elytral apices spined, protibia with a subapical tooth beneath, male endophallus with cs highly reduced or nearly absent, MPH without significant swelling or protuberance, PB provided with sf and si, and APH provided apically with a pair of rod-like sclerites associated with the ejaculatory ducts. While in M. sutor sutor (Linnaeus, 1758), the type species of Monochamus, body relatively slender and antennae relatively long, pronotal lateral spines less acute apically and not curved, elytra without carinae, elytral apices rounded, protibia untoothed, male endophallus with cs well developed, MPH swollen near middle and apex, PB lacking sf and si, and APH provided with a single V-shaped sclerite. Pascoe (1858) mentioned the similarities between his Monochamus gravidus and Hammaticherus marmoratus Westwood, 1848 (the type species of Pseudomeges Breuning, 1944). Based on the comparison of both type species in this study, the genus Meges can be distinguished from Pseudomeges by the antennal scape with the cicatrix more prominent and delimited by a distinct carina (“scape provided with a complete cicatrix” in Breuning 1943), elytral apices provided with long sutural spine, protibia with a subapical tooth beneath, male endophallus with the MPH relatively longer and slenderer., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on pages 243-244, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Pascoe, F. P. (1866) Longicornia Malayana; or, a descriptive catalogue of the species of the three longicorn families Lamiidae, Cerambycidae and Prionidae, collected by Mr. A. R. Wallace in the Malay Archipelago. The Transactions of the Entomological Society of London, (3) 3, 225 - 336.","Pascoe F. P. (1858) On new genera and species of longicorn Coleoptera. Part III. The Transactions of the Entomological Society of London, (2) 4 (6 - 7), 236 - 266, pls XXV, XXVI. https: // doi. org / 10.1111 / j. 1365 - 2311.1858. tb 01823. x","Clermont, J. (1932) Description d'un genre et d'une espece nouveaux de Cerambycidae de la sous-famille des Lamiinae [Col.]. Bulletin de la Societe Entomologique de France, 37 (15), 214 - 216. https: // doi. org / 10.3406 / bsef. 1932.14460","Breuning, S. (1944) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (107 - 135), 281 - 512.","Westwood, J. O. (1848) The cabinet of oriental entomology; being a selection of rare and beautiful insects, natives of India and the adjacent islands. W. Smith, London ii + 88 pp., 42 pls. https: // doi. org / 10.5962 / bhl. title. 34273","Breuning, S. (1943) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (89 - 106), 137 - 280."]}

Pseudomeges Breuning, 1944 Pseudomeges Breuning,1944: 300. Type species: Hammaticherus marmoratus Westwood, 1848, by original designation., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2022, Taxonomic studies on the genera Meges Pascoe, 1866 and Pseudomeges Breuning 1944 from China (Coleoptera, Cerambycidae, Lamiinae, Lamiini), pp. 242-250 in Zootaxa 5120 (2) on page 247, DOI: 10.11646/zootaxa.5120.2.4, http://zenodo.org/record/6389228, {"references":["Breuning, S. (1944) Etudes sur les Lamiaires (Coleop. Cerambycidae). Douzieme tribu: Agniini Thomson. Novitates Entomologicae, 3 eme supplement (107 - 135), 281 - 512.","Westwood, J. O. (1848) The cabinet of oriental entomology; being a selection of rare and beautiful insects, natives of India and the adjacent islands. W. Smith, London ii + 88 pp., 42 pls. https: // doi. org / 10.5962 / bhl. title. 34273"]}

6 Cerambycidae species are newly recorded for different provinces of China.

Tsounkranaglenea hefferni sp. nov. (Figs. 1-14) Type material. Holotype, male (Fig. 1), Malaysia, Sabah, Crocker Range, 2003-II-16, leg. local coll. (IZCAS). Paratypes: 1 male, Malaysia, Sabah, Mt. Trus Madi, 1500-2000m, 1998-III-11, leg. Affenddy (DHCO); 1 female, Malaysia, Sabah, Mt. Trus Madi, 1500-2000m, 1998-III /V, leg. local coll. (IZCAS); 1 female, Malaysia, Sabah, Crocker Range, 2010-III-18, local coll. (DHCO); 1 female, same data but 1998-III-19 (DHCO). Other material (examined only by photographs). 1 male with length 11.5 mm, no. S11.9-151. Nobuo Ohbayashi showed the first author a specimen from Malaysia with dorsal view and lateral view pictures in 2011, which was surely this species. It may be from Trus Madi, however, the owner Mr. Minoru Sawai could not be contacted during this research. Description (based on two males and three females): Male: length: 11.0– 11.5 mm, humeral width: 3.0– 3.2 mm. Female: length: 11.0–14.0 mm, humeral width: 3.4–4.2 mm. Body black. Head black, with frons covered with dense grayish white to yellow pubescence (Figs. 2e, 11d), which extends as two short vittae to occiput; antennae black. Prothorax covered with dense brownish pubescence in dorsal view except a whitish pubescent spot at base for both sexes and a whitish spot in middle of apical part for female only (Figs. 11a, 12); with a longitudinal grayish white to yellow pubescence in lateral view (Figs. 1b, 11b). Scutellum black, covered by brownish (basal part) and whitish (apical part) pubescence. Elytra black, covered by rather dense brownish pubescence, at basal fourth and along suture with some silver erect setae, basal setae much longer than others (Fig. 12), and with grayish white to yellow pubescence markings on each elytron as following: (for both sexes) a transverse vitta at base; a apical vitta; (for only female) a long line along second row of punctures (counting from lateral carina) from nearly base to apical tenth, normally with a subapical interruption; a lateral line along the puncture row near lateral carina from nearly base to middle (Fig. 11b); a short longitudinal vitta along suture just behind scutellum (Fig. 11a, sometimes missing, Fig. 12). Ventral surface black with grayish white pubescence except a long stouter line with interruptions caused by lacking pubescence (Figs. 1b, 11b). Legs reddish brown to dark brown, covered with grayish pubescence. Frons longer than broad, width less than (male) or subequal to (female) two eyes’s combined width. Antennomere ratio: male, 25:5:36:28:27:26:25:24:22:20:21; female, 25:5:36:26:25:24:23:21:19:18:19. Prothorax longer than (male) to as long as (female) broad. Elytron truncated apically, with a short tooth at inner angle and a sharp and long tooth at outer angle. Hind femur reaching base of (female) or the bent part of (male) fifth abdominal segment, first hind tarsal segment longer than the following two segments combined. The sternite VII in male (Figs. 2b, 2c, 3b–d) strongly bent at base, the apical half reduced and transformed into two slender knife-shaped teeth (Figs. 3b–d), with long brown brush hairs on the outer side. Male terminalia (Figs. 6–10): Tegmen about 3.0 mm in length; lateral lobes slender (Figs. 8a, 8c), each about 1.2 mm long and 0.3 mm wide, provided with a thin setae, strongly protruding lobe at ventral base (Fig. 8b), apical half with medium long setae; ringed part elbowed in widest portion, converging; basal piece bifurcated apically (Figs. 8c); median lobe plus median struts strongly curved (Figs. 9b, 9d), shorter than tegmen; median struts more than one third and less than half of whole median lobe in length; dorsal plate slightly shorter than ventral plate; apex of ventral plate strongly emarginated, forming two lateral apical lobes (Figs. 9a, 9b); median foramen slightly elongated; internal sac about triple length of median lobe, with two pairs of basal plate-like sclerites (located behind apex of median struts), one band of distinct supporting armature, some barely separated plate-like sclerites between them, and three rod-like sclerites at the end (Figs. 10a–10c), rod-like sclerites about 1.0 mm, much shorter than median lobe or tegmen. Tergite VIII (Figs. 6a–6c) longer than broad, strongly narrowed before apex (Fig. 6b), apex emarginated with two short ear-shaped lateral lobes, with dense and long brown setae along the narrowed part (Figs. 6a, 6b). Sternite VIII with spiculum relictum as Fig. 7b, sternite IX (spiculum gastrale) as Fig. 7a. Female genitalia (Figs. 13–14): spermathecal capsule composed of an apical orb and a long and strongly curved stalk, the stalk more than triple the length of the apical orb, curved twice at basal part (Fig. 14). Spiculum ventrale longer than abdomen. In our observation, spiculum ventrale measured 7.7 mm for an adult compared with abdomen which measured 5.7 mm in ventral view. Etymology. The species is named after Mr. Daniel Heffern (Texas, USA), who offered the interesting material for this study and provided the first author some wonderful saperdine specimens from Malaysia for research. Diagnosis. This species resembles Glenea subaurata Breuning, 1950 at first glance by the similar shape and colour. The specific differences are easily defined by the male sternite VII. Without specimens for dissection, we can not conclude whether Glenea subaurata Breuning, 1950 should be moved to the new genus Tsounkranaglenea or not. Remarks. Glenea subaurata Breuning, 1950 (Figs. 15–18) was described based on “Type un male de Sumatra: Si-Rambé, XII-1890 - III-1891, leg. E. Modigliani”, with “Longuer 10 mm, Largeur 2mm. 1/3”. The holotype male (Figs. 15–16) is deposited in MSNG and had been examined by the first author in June, 2008. Another male specimen (Figs. 17–18) is deposited in NHMB, with a label written “ PARATYPE ”. Though it has a Breuning’s handwriting label “glenea / subaurata / mihi. P. T. / Breuning det.”, it is not a paratype because the locality “Madam / Sumatra ” and the individual specimen were not mentioned in the original paper (Breuning, 1950b). Here we confirmed that the identification is correct and it is a useful additional material for the specific definition of Glenea subaurata Breuning, 1950., Published as part of Lin, Mei-Ying & Ge, Si-Qin, 2021, Tsounkranaglenea hefferni gen. et sp. nov. from Sabah, Malaysia (Coleoptera Cerambycidae, Lamiinae: Saperdini), pp. 289-297 in Zootaxa 5048 (2) on pages 291-296, DOI: 10.11646/zootaxa.5048.2.9, http://zenodo.org/record/5552118, {"references":["Breuning, S. (1950 b) Nouveaux Lamiaires du Musee de Genes (Coleoptera, Cerambycidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova, 64, 170 - 201."]}

Tsounkranaglenea gen. nov. Type species: Tsounkranaglenea hefferni sp. nov. Diagnosis. It is mostly similar to Glenea by the lateral elytral carinae and truncate elytral apex, but can be distinguished by the elongated, bended and rake-shaped sternite VII. In fact, it differs from all other saperdine genera by the peculiar sternite VII in males. It also differs from Paraglenea Bates, 1866, Heteroglenea Gahan, 1897 (as defined in Lin, Montreuil et al, 2009) and Pareutetrapha Breuning, 1952 by the male claws of fore and hind legs simple instead of appendiculate or bifid and elytral apex truncated with sharp teeth instead of rounded or slightly truncated without sharp teeth. It also differs from Eumecocera Solsky, 1871 and Stenostola Dejean, 1835 by the elytra with lateral carinae and male claws of fore and hind legs simple instead of appendiculate or bifid. The combination of the following characters makes the new genus easily separable from other saperdine genera: prothorax without lateral tubercles, elytra with distinct lateral carinae, elytral apex truncated with long spines at outer angle, male claws with appendiculated tooth only in mesotarsi and female claws all simple, male sternite VII elongated and bended into a rake-shape. Description. Small-sized (under 15 mm). Head not broader than prothorax. Eyes deeply emarginate, not divided, lower eye lobe much vertically longer than (male) to subequal to (female) gena. Antennae longer than body, in male slightly longer than female, basal segments fringed with sparse setae, scape slightly expanded, second antennomere short, third antennomere always the longest, 4 th antennomere subequal to (female) to slightly longer than (male) scape, 4 th to 10 th slightly and gradually decreasing in length except 11 th being slightly longer than 10 th. Prothorax cylindrical, without lateral tubercles, slightly narrowed around basal fifth. Elytra subparallel, truncated apically, with sharp teeth at both inner and outer angles, each with two distinct lateral carinae starting from the base and combined into apical outer tooth (Figs 1b, 11b). Procoxal cavity closed posteriorly (Fig. 11c), mesocoxal cavity open to mesepimeron, metanepisternum more than twice as wide anteriorly as posteriorly. Protarsi with first segment expanded in male (Fig. 1a), mesotibiae with an oblique groove with setae (Fig. 1b), hind femur reaching fifth abdominal segment, hind tarsi with first segment longer than the following two combined. Male claws: only anterior claws of mesotarsi appendiculate with small teeth (Figs. 4–5), posterior claws of mesotarsi without teeth, and claws of pro- and metatarsi simple. Females claws simple (Figs. 11 a-11b). Male sternite VII elongated and bent into a rake-shape (Figs. 1–3), female sternite VII with a median groove (Fig. 11c). Male terminalia. Apex of male tergite VIII emarginated (Figs. 6a–6c). Lateral lobes slender, with a strong tooth at ventral base (Fig. 8b); ringed part elbowed in the widest portion, converging; basal piece well-developed and bifurcated (Fig. 8c). Median lobe strongly curved, shorter than tegmen, dorsal plate shorter than ventral plate, apex of ventral plate emarginated (Fig. 9a). Median foramen not elongated. Endophallus with one band of supporting armature, 4 basal plate-like sclerites, and 3 rod-like sclerites. Ejaculatory duct single. Female terminalia: Setae of sternite VIII dense and long. Spermathecal capsule and gland positioned on apex of spermathecal duct. Spermathecal capsule strongly sclerotized, composed of an apical orb and a long stalk, spiculum ventrale longer than abdomen. Etymology. The generic name is a combination of a Greek word tsounkrána (τσουγκράνα) and the genus name Glenea. The Greek word “tsounkrána” refers to the shape of sternite VII in male, which looks like a rake. Gender feminine. Distribution. Malaysia. Remarks. It is very similar to Glenea (Breuning, 1956; Breuning, 1958) by the elytral lateral carinae and truncated elytral apex, and the following characters are quite common in Glenea members: endophallus with 4 basal plate-like sclerites (Lin et al., 2009; Lin, Tavakilian et al., 2009a,b; Lin & Lin, 2011; Lin & Yang, 2011a, b; Lin et al., 2018), and 3 rod-like sclerites (Lin et al., 2009; Lin, Tavakilian et al., 2009a,b; Lin & Lin, 2011; Lin & Yang, 2011a, b; Lin & Dai, 2012; Lin, 2013); spermathecal capsule strongly sclerotized, composed of an apical orb and a long stalk (Lin et al., 2009; Lin, Tavakilian et al., 2009b; Lin & Yang, 2011a, b; Lin & Dai, 2012). We separate it from Glenea based on the following reasons.: 1) Glenea is heterogeneous (Lin, Montreuil et al., 2009;), even though outer characters are very similar (Lin & Tavakilian, 2012), this peculiar species does not match with any type species of the subgenera; 2) Though most of characters can be found in the previous Glenea members, and the peculiar male sternite might not be suitable for generic level, but with only one band of supporting armature and the emarginated apex of the ventral plate of the median lobe, this convinced the authors to make a new genus. Most members of Saperdini have zero or two bands of supporting armature, rounded to the pointed apex of the ventral plate of the median lobe (Lin et al., 2009; Lin, Tavakilian et al., 2009a, b; Lin & Yang, 2011a, b; Lin & Dai, 2012; Lin, 2013; Lin et al., 2018). 3) Although the sexual dimorphism (on pubescence markings) referred to the subgenus Glenea (Acutoglenea) Breuning, 1958, and the dark integument referred especially to G. (A.) versuta basaloides Breuning 1958, G. (A.) versuta maura Pascoe, 1867, it can not be included in the subgenus Acutoglenea because of the non-simple male claws and very different male terminalia (based on the first author’s unpublished data). Besides, the type species Glenea (Acutoglenea) acuta (Fabricius, 1801) has a stouter female, with elytral length less than twice basal width, fourth antennomere much shorter than scape, which are very different from the new taxon herein described. 4) We have checked the subgenus Glenea (Lineatoglenea) Breuning, 1950, which is represented by a unique type species Glenea (Lineatoglenea) lineatopunctata Breuning, 1950 from Malaysian Borneo. There are no images available, and we did not have an opportunity to examine the type specimen which should be deposited in University of Malaysia, Sarawak (Breuning, 1950a). Based on the original description, it shares with the new taxon by antennae longer than body, similar antennomere ratio, pronotum and elytra, however it differs from the new taxon by the fifth male abdominal segment provided at the end with a short median longitudinal ridge. 5) We have checked the subgenus Glenea (Spiniglenea) Breuning, 1958, which is represented by a unique type species Glenea (Spiniglenea) spinosipennis Breuning, 1958 from Malaysian Borneo. It also has no images available, and we did not have an opportunity to examine the type specimen which should be deposited in University of Malaysia, Sarawak (Breuning, 1958b). Based on the original description, it is difficult to separated it from the new taxon on genus level, since it was based only on a female. However, it is surely not the same species. 6) We compared the new taxon with Glenea (Metaglenea) Breuning, 1956, which is represented by a species from Sumatra, and Glenea (Porphyrioglenea) Breuning, 1956, which is represented by a species from West Malaysia and Sabah, East Malaysia. They can be easily distinguished from the new taxon by the very close antennal tubercles, shorter and stouter antennae. Glenea (Pseudotanylecta) Breuning, 1956, Glenea (Subgrossoglenea) Breuning, 1956, Glenea (Tanylecta) Pascoe, 1866 also from Malaysia and Indonesia, can be separated by the close and protruding antennal tubercles. 7) Glenea (Poeciloglenea) Aurivillius, 1920, Glenea (Punctoglenea) Breuning, 1956, Glenea (Reginoglenea) Breuning, 1956, Glenea (Rubroglenea) Breuning, 1956, Glenea (Rufoglenea) Breuning, 1956, Glenea (Stiroglenea) Aurivillius, 1920, Glenea (Vanikoroglenea) Breuning, 1956, Glenea (Vittiglenea) Breuning, 1956, Glenea (Volumnia) Thomson, 1860 and all other subgenera have been studied by the first author, and none of them are suitable for the new taxon. 8) The new taxon differs from Glenea (Lobunguiglenea) Lin & Tavakilian, 2014 by the male claws with only anterior claw of mesotarsus appendiculate with small lobe in inner side, instead of all claws appenciculated in outer sides, and genitalia with median lobe strongly curved, apex of ventral plate emarginated, instead of genitalia with median lobe slightly curved, apex of ventral plate pointed., Published as part of Lin, Mei-Ying & Ge, Si-Qin, 2021, Tsounkranaglenea hefferni gen. et sp. nov. from Sabah, Malaysia (Coleoptera Cerambycidae, Lamiinae: Saperdini), pp. 289-297 in Zootaxa 5048 (2) on page 290, DOI: 10.11646/zootaxa.5048.2.9, http://zenodo.org/record/5552118, {"references":["Lin, M. Y., Montreuil, O., Tavakilian, G. & Yang, X. K. (2009) Reinstatement of the genus Heteroglenea Gahan, with four new combinations, four new synonyms and three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Zootaxa, 2137 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2137.1.1","Breuning, S. (1952) Revision einiger Gattungen aus der Gruppe der Saperdini Muls. (Col. Cerambycidae). Entomologischen Arbeiten aus dem Museum G. Frey, Tutzing bei Munchen, 3 (1), 107 - 213, 3 pls.","Breuning, S. (1956) Revision der Gattung Glenea Newman. Entomologische Arbeiten aus dem Museum G. Frey, Tutzing bei Munchen, 7 (1), 1 - 199.","Lin, M. - Y., Yang, X. - K. & Lin, Y. - L. (2009) Taxonomic Notes on Glenea pseudoscalaris (Fairmaire, 1895) (Coleoptera, Cerambycidae, Lamiinae). Longicornists, Special Bulletin of the Japanese Society of Coleopterology, No. 7, 385 - 390.","Lin, M. - Y., Tavakilian, G., Montreuil, O. & Yang, X. - K. (2009 a) Eight species of the genus Glenea Newman, 1842 from the Oriental Region, with description of three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Zootaxa, 2155 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2155.1.1","Lin, M. - Y. & Yang, X. - K. (2011 a) A new species Glenea shuteae sp. nov. from Yunnan, China: compared with Glenea decolorata Heller (Coleoptera, Cerambycidae, Lamiinae, Saperdini). Acta Zootaxonomica Sinica, 36 (1), 40 - 44.","Lin, M. - Y., Dar, M. A. & Akbar, S. A. (2018) Taxonomy and distribution of Glenea beesoni Heller, 1926 (Coleoptera: Cerambycidae: Lamiinae: Saperdini) from Indian Himalayas. Oriental Insects, 52 (3), 221 - 228. https: // doi. org / 10.1080 / 003053 16.2017. 1397565","Lin, M. - Y. & Dai, L. (2012) Description of a new species of the genus Glenea from Tibet, China (Coleoptera, Cerambycidae, Lamiinae, Saperdini). ZooKeys, 216, 5 - 11. https: // doi. org / 10.3897 / zookeys. 216.3360","Lin, M. - Y., Tavakilian, G., Montreuil, O. & Yang, X. - K. (2009 b) A study on the indiana & galathea species-group of the genus Glenea, with descriptions of four new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Annales de la Societe Entomologique de France, New Series, 45 (2), 157 - 176. https: // doi. org / 10.1080 / 00379271.2009.10697599","Lin, M. Y. & Tavakilian, G. (2012) A New Genus Bifidunguiglenea gen. nov. Is Erected for the Species Glenea gestroi Gahan (Cerambycidae: Lamiinae: Saperdini). PLoS ONE, 7 (7), e 40768, 1 - 5. https: // doi. org / 10.1371 / journal. pone. 0040768","Breuning, S. (1950 a) Nouvelles formes de Lamiaires (Cerambycidae, Coleoptera). The Sarawak Museum Journal, 5 (2), 373 - 388.","Breuning, S. (1958 b) Revision der Gattung Glenea Newm. (Col. Ceramb.) (3. Fortsetzung und Schluss). Entomologische Arbeiten aus dem Museum G. Frey, Tutzing bei Munchen, 9 (3), 804 - 907.","Pascoe, F. P. (1866) Catalogue of longicorn Coleoptera, collected in the Island of Penang by James Lamb, Esq. (Part I.). Proceedings of the Zoological Society of London, 1866, 222 - 267, pls. XXVI - XXVIII.","Aurivillius, C. (1920) Neue oder wenig bekannte Coleoptera Longicornia. 17. Arkiv for Zoologi, 13 (9), 1 - 43 (= 361 - 403), figs. 73 - 81. https: // doi. org / 10.5962 / bhl. part. 20147","Thomson, J. (1860) Essai d'une classification de la famille des cerambycides et materiaux pour servir a une monographie de cette famille. chez l'auteur [James Thomson] et au bureau du tresorier de la Societe entomologique de France, Paris, xvi + 128 pp. https: // doi. org / 10.5962 / bhl. title. 9206","Lin, M. - Y. & Tavakilian, G. (2014) A strange new species Glenea judsoni sp. nov. from Taiwan, China (Coleoptera: Cerambycidae: Lamiinae: Saperdini). In: Hua, L. - Z. (Ed.), Feelings in China. Memorial works of Dr. J. L. Gressitt of the 100 th Anniversary on his Birthday. Sun Yat-sen University, Guangzhou, pp. 161 - 167."]}

Lin, Mei-Ying, Ge, Si-Qin (2021): Tsounkranaglenea hefferni gen. et sp. nov. from Sabah, Malaysia (Coleoptera Cerambycidae, Lamiinae: Saperdini). Zootaxa 5048 (2): 289-297, DOI: https://doi.org/10.11646/zootaxa.5048.2.9

Glenea albosignatipennis Breuning, 1950 (Figures 37–38, 44) Glenea (s. str.) albosignatipennis Breuning, 1950: 260. TL: India, North Belgaum, Bombay. TD: IFRI. Glenea (Glenea) albosignatipennis: Breuning, 1956a: 61; Breuning, 1956b: 713. Type specimen examined Holotype, 1 ♀, on twig of, Tarewadi 2600 feet, N. Belgaum, 1938.V.21, J.A. Graham (IFRI). Other specimens examined India: 1 ♂, ‘ paratype’ (unmentioned in Breuning 1950 and therefore not type), Somwarpet, Coorg, S. India (NHMB, ex FREY). 1 ♀, Somwarpet, Coorg, S. India (BMNH, with a yellow label ‘ Data unreliable. See Brit. Mus. 1949–314’.) Description complementary to Breuning (1950, 1956b). The only examined male specimen (Figure 38) matches with females very well in colour and patterns of pubescent maculae. Male with antennae longer than body length, labrum medially provided with two pairs of suberect, distinctly elongate yellowish brown setae arising from respective punctures, sternite VII with a distinct umbo prior to apical opening (Figure 38 (b)), apex rounded. Both male and female with simple claws. Distribution India (Karnataka and Maharashtra). Remarks Breuning (1950: 261) described this species based on ‘Longueur 12 mm. Largeur 3 mm 1/ 3. Type une female de North Belgaum, Bombay (J.A. Graham)’, wrote something about males in the original description, and repeated the sexual difference in Breuning (1956b), also only mentioning the holotype female (Figure 37). However, one male specimen in NHMB was labelled as paratype and with a handwritten identification label by Breuning (Figure 38 (d)). Since it was not mentioned in the original literature (Breuning 1950), it cannot be treated as paratype. The second author examined another female of this species deposited at BMNH, with the same locality label as the male in Figure 38, but with a yellow label noted ‘Data unreliable. See Brit. Mus. 1949–314’. Michael Geiser (m.geiser@nhm.ac.uk, 7 January 2020) from BMNH informed us that The famous Cerambycids specialist E.F. Gilmour was caught stealing specimens from the London museum back in the 1940s. Most of his specimens were later returned to the museum, but there was a suspicion that he could have swapped around some of the labels and faked some of the data to wipe out his traces. As far as I am aware, the vast majority of the specimens still have the correct labels, but the curators at the time added this label as a cautionary measure, because in some cases the label data could have been compromised The male specimen in NHMB (Figure 38) having the exact same locality label proves that the female in BMNH has the correct labels. Therefore, the locality Somwarpet, Coorg, S. India, is correct; it is now in Karnataka state, south India, which is situated nearly 600 km away from the type locality Tarewadi village, now in Kolhapur district of Maharashtra state., Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 234-238, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Breuning S. 1950. Descriptions de nouveaux Lamiaires de l'Inde (Coleopteres). Indian For Rec (New Ser) Entomol. 7 (8 [1949]): 255 - 265.","Breuning S. 1956 a. Revision der Gattung Glenea Newman. Entomol Arb Mus G Frey. 7 (1): 1 - 199.","Breuning S. 1956 b. Revision der Gattung Glenea Newm. (1. Fortsetzung). Entomol Arb Mus G Frey. 7: 671 - 893."]}

Glenea vellayaniensis sp. nov. (Figures 1–12) Type material Holotype: ♂, (1) India: Kerala / Vellayani / 8.43333333 N, 76.98416667 E / 5 May 2019, at light / S. R. Hiremath Coll. (2) HOLOTYPE/ Glenea vellayaniensis sp. nov. /des. Hiremath and Lin, 2020 (red label). Paratypes (2 specimens, both with a white locality label as given below, besides a second pink label: ‘ PARATYPE / Glenea vellayaniensis sp. nov. /des. Hiremath & Lin, 2020’): 1♀ with label as follows: India: Kerala / Vellayani; 17. November. 2016/ 8.43333333 N, 76.98416667 E /S. R. Hiremath Coll.; 1♀ with label as follows: India: Kerala / Vellayani; 5. i. 2017 / 8.43333333 N, 76.98416667 E /S. R. Hiremath Coll. Description Male (n = 1) (Figure 1). Body length measured from vertex to elytral apex 6.29 mm; humeral width 1.92 mm. General body colour reddish brown; head, antennomeres I–II, pronotum, elytra and ventral side of body reddish brown; apical half of mandibles black; anteclypeus and antennomeres III–XI yellowish brown; maxillary and labial palpi, legs (except tarsal claws) yellowish brown; tarsal claws reddish brown. Head with frons thickly adpressed with cream yellow hairs; outer margin of upper eye lobes on each side of vertex ornamented with transverse, oblong band of adpressed yellow hairs; area surrounding lower eye lobes, gena, postclypeus thickly adpressed with white hairs. Basal half of mandibles adpressed with thick white hairs. Apical half of labrum adpressed with transverse band of pale white hairs; middle portion of labrum adorned with transversely arranged, seven golden yellow suberect setae, setae at middle smaller than remaining. Head slightly wider than pronotum, randomly covered with punctures subequal in size to those on pronotum, medially impressed with a fine dark brown longitudinal sulcus running from postclypeus to vertex. Labrum thick, protruding, longer than anteclypeus, lying over mandibles; mandibles concealed in frontal view; dorsal surface of labrum with a median transverse ridge, followed by a postmedian shallow excavation; apical margin smooth and truncated. Eyes with lower lobes 2.17 times as long as gena, connected to upper lobes by five rows of ommatidia. Antennal tubercles weakly developed, widely separated, with space between them transversely flat. Antenna with antennomeres I–VII moderately setose beneath with dark brown suberect setae; integument of antennomere I–XI uniformly covered with dark brown and yellowish brown recumbent setae. Antennae surpassing elytral apex near distal end of antennomere VIII, 1.35 times as long as body. Scape thick, cylindrical, slightly narrowed at base, integument uniformly covered with shallow punctures. Ratio of lengths of antennomeres: 1.00: 0.29: 1.35: 1.35: 1.06: 1.00: 0.94: 0.88: 0.88: 0.82: 0.82. Pronotum with pronotal disc faintly covered with yellowish brown recumbent setae, intermixed with uniformly distributed golden yellow erect setae originating from respective punctures; pronotal disc, on each side, ornamented with a broad, longitudinal, sublateral band of thick, adpressed creamy yellow hairs, turning white postmedially, inner margin of this band distinctly concave at its proximal third; lower lateral sides of pronotum, on each side, just above procoxae ornamented with a broad, longitudinal band of thick adpressed white hairs; basal middle of pronotum ornamented with a short, angulate patch of thickly adpressed hairs, white intermixed with yellow; centro-notal area with pubescence of white and yellowish brown disposed into a faint, median, narrow longitudinal band. Pronotum transverse, 1.15 times as wide as long, 0.68 times as long as humeral width; apical margin broader than basal margin; lateral margin on each side slightly swollen at middle and weakly constricted before basal margin. Pronotal disc densely covered with fine punctures; impressed ante-basally with a shallow, transverse groove; pronotal disc just before ante-basal groove, impressed at its middle with a weakly raised, short ridge. Apical margin straight, basal margin weakly sinuate. Scutellum short, tongue-shaped with distinctly arcuate apex; prominently covered with thickly adpressed yellow hairs. Elytra uniformly covered with moderately dense, yellowish grey pubescence; elytral disc uniformly intermixed with suberect golden yellow setae arising from respective punctures; outer margin and apex of elytron fringed with single row of golden yellow suberect setae. Elytral disc, on each side, ornamented with pubescent spots arranged as follows: pre-medially with two spots, medially with one and post-medially with four spots along with a short, sub-lateral pubescent band. First premedian spot located at basal quarter, between humeral prominence and scutellum, made up of thick yellow pubescence, large, broadly bean-shaped with white halo on its borders; second premedian spot located sub-laterally at basal third, small, oval, made up of thick white hairs. Median spot largest, placed between sutural and sub-lateral margins, transversely oval, made up of thick yellow hairs bordered with halo of white hairs; anterior margin angulate at its middle, posterior margin obliquely straight. Post-median spots arranged as follows: first and second spots placed obliquely side by side, just behind middle; first spot located along sutural margin, hemispherical, made up of thick yellow hairs bordered with halo of white hairs, antero-laterally joins median spot along sutural margin; second spot touches sub-lateral margin, circular, made up of thick white hairs; third spot at apical third of elytral disc, largest of post-median spots, made up of thick yellow hairs, bordered with halo of white hairs; second and third spots joined laterally along sub-lateral margin by a short, narrow, longitudinal band of white hairs; fourth spot located on elytral apex, transversely oval, subequal in size to first and second post-median spots, made up of thick yellow hairs, anteriorly bordered with narrow band of white hairs. Elytra elongate, 0.68 times as long as body, 3.30 times as long as pronotum, 2.22 times as long as humeral width; broadest at base, gradually narrowed towards apex. Humeral prominence well developed, obtusely ridged. Sublateral margin impressed with a longitudinal carina, slightly curved at basal fourth, originating from humeral prominence, extending to apical third of elytra. Elytral disc dorsally flat, coarsely punctate up to apical third, punctures becoming finer towards apex; elytral apex truncate with sutural angle dentate and marginal angle stretched into acute spine. Legs with faintly adpressed golden yellow recumbent setae, long and dense on tibiae, interspersed with randomly distributed suberect setae of similar colour. Tarsal claws appendiculate on all legs. Sternites with median portion of pro-, meso- and metasternum uniformly covered with fine, golden yellow recumbent hairs; mesepisternum, mesepimeron and metepisternum thickly adpressed with creamy white hairs; lateral sides of metasternum thickly adpressed with creamy yellow hairs. Abdomen with median longitudinal space uniformly covered with golden yellow recumbent hairs. Ventrites I–IV, laterally on each side, ornamented with a large, thickly adpressed white-haired spot. Sternite VII, laterally on either side with a post-basal spot of similar hairs as previous. Abdomen with sternite VII 0.23 times as long as abdomen, 1.33 times as long as previous segment; disc without a medio-longitudinal sulcus; apex weakly emarginate. Male genitalia (Figures 4–6). Tergite VIII (Figure 4) trapezoidal with apical margin truncated, moderately fringed with light brown elongate setae; lateral margins weakly curved in basal third, fringed with light brown elongate setae from basal third to apex, sparser proximally, denser at distal fourth on inner side; median disc of tergite VIII adorned with randomly distributed light brown, minute, adpressed setae of similar colour. Sternite VIII rectangular, proximal two-thirds with angular reddish brown patch, apex on each side of middle adorned with a transverse patch of medium to elongate, light brown, suberect setae. Spiculum gastrale about 4.43 times as long as spiculum relictum, Y-shaped with median arm slightly longer than lateral arms, distinctly curved at apex; lateral arms obliquely straight, bearing membranous dilation at their distal end. Tegmen (Figure 5 (a– c)) about 1.43 mm long, curved at middle in lateral view (Figure 5 (c)). Basal piece present, reddish brown, distally arcuate, entire surface uniformly covered with minute, suberect spinules. Roof present. Ringed part converging, constricted near widest portion; manubrium hollow, thick in appearance due to internally explanated membrane on each side, in both dorsal and ventral planes; constricted just before basal end; basal end of manubrium abridged by a rectangular membrane, truncated apically (Figure 5 (a)). Lateral lobes (Figure 5 (b)) separated, about 0.25 times as long as total length of tegmen with basal margins transversely ridged; inner margins straight, outer margins oblique, gradually arcuate towards apex; apex rounded. Integument in ventral view red-brown except for a small lighter apical area; basal margin of integument impressed with a transverse row of reddish brown, medium sized, suberect setae; remaining surface randomly covered with similar setae except for a few glabrous spaces near central area; apex concentrated with elongate light brown setae. Median lobe (Figure 6 (a–b)) subequal to length of tegmen, arcuate in lateral view; basal struts originate almost near apical fourth; ventral plate (Figure 6 (b)) dark brown at apical one-fifth, interspersed with a few randomly distributed punctures; apex obtusely pointed, lateral sides asymmetrical, with right side weakly concave near apex. Endophallus (Figures 7–9) in lateral view about 3.90 times as long as median lobe; BPH about 0.28 times as long as endophallus, with membrane transversely plicate; distal end bearing CS embedded in ventral membrane, dorsal membrane embedded with a pair of median sclerotic plates (Figure 8). MPH with MT short, slightly bulged at distal end, about 0.09 times as long as endophallus; membrane of MT randomly distributed with a few round spicules, dorsal membrane at proximal third impressed with a median patch of densely distributed angulate spicules (Figure 8). CT uniformly tubular, curved at middle, about 0.19 times as long as endophallus, proximal two-thirds of membrane uniformly covered with angulate spicules, gradually turning obtuse towards beginning of apical third; apical third dark, densely and compactly adorned with C-shaped spicules, reaching up to proximal half of PB; PB tubular in proximal half, bulged at distal half, as long as MT, uniformly but not densely adorned with C-shaped spicules, gradually turning into U-shaped spicules towards APH. APH funnel shaped, as long as MT, uniformly and closely adorned with stout, broadly U-shaped spicules; junction between PB and APH circumscribed by membranous fringe; apex of APH continued with a membranous tube, bearing RS at distal end. RS about 0.19 times as long as endophallus, uniformly curved in lateral view, composed of two closely appressed, sclerotised plates, of which ventral plate (Figure 9 (b)) dorso-ventrally flattened with lateral sides curved upwards towards dorsal plane distally, broadly encasing dorsal plate; dorsal plate (Figure 9 (a)) made up of two closely approximated inconspicuous rods, forming tubular structure; proximal ends bearing two asymmetrical, falcate tines, one tine being longer than other; apex of longer tine bearing divaricate claw-like projection. ED single, arises medially from distal end of dorsal plate of RS. Female (n = 2) (Figures 2–3). Body length 6.86–6.91 mm, humeral width 2.21–2.26 mm. Similar to male in general appearance, with the following differences. Head with lower eye lobes 1.30–1.33 times as long as genae; mandibles distinctly visible in frontal view. Antennae 1.31 times as long as body. Ratio of lengths of antennomeres: 1.00: 0.25: 1.25: 1.25: 0.95: 0.90: 0.85: 0.80: 0.75: 0.70: 0.70. Pronotum 1.20–1.21 times as wide as long, 0.62–0.65 times as long as humeral width. Elytra 0.72 times as long as body, 3.47–3.55 times as long as pronotum, 2.19–2.26 times as long as humeral width. Last abdominal ventrite robust, dorsally convex, about 0.30–0.32 times as long as length of abdomen, 2.10–2.22 times as long as previous segment; disc medially impressed with a dark brown longitudinal sulcus; apex broadly crescent shaped. Female genitalia (Figures 10–12) with ovipositor (Figure 10 (a)) about 1.10 mm long, in dorsal view thickly covered with adpressed light brown setae at apical third (except coxite lobe and stylus), immediately followed by compactly disposed cells, in scale-like arrangement throughout basal two-thirds except outer margin on each side, which is randomly distributed with a few circular punctures. Coxite lobe light brown, randomly covered with circular punctures except at apex; apex of coxite lobe in dorsal view (Figure 10 (b)) transversely impressed with a single row of closely congregated circular punctures with six setae arising from them, outer setae on either side longer and pointed, remaining setae placed between them, shorter, subequal to each other and obtusely pointed. Stylus dome-shaped, red brown, continuous with coxite lobe; basal portion of stylus impressed with a few remotely distributed punctures; apex of stylus bears a single seta arising from associated puncture. Vaginal plates flap shaped, weakly sclerotised, basally red brown. Bursa copulatrix (Figure 12) elongate, basal two-thirds tubular and bi-sinuate, apical third distinctly capitate. Spermathecal duct distinctly longer than spermatheca, enters bursa copulatrix at apical third, uniformly tubular, with its distal end slightly curved. Spermatheca (Figure 11) arise on spermathecal duct separately, before spermathecal gland, red brown with basal end obliquely curved, distinctly moulded into an elongate, U-shaped tubule up to apical third; apex curved, distinctly capitate. Spermathecal gland sac like, basally with a distinctly sclerotised ringed plate giving rise to a short sclerotised tube. Tignum longer than abdomen. In the specimen examined, tignum measured about 4.94 mm in length while abdomen measured about 3.17 mm. Differential diagnosis Glenea vellayaniensis sp. nov. is closely allied to G. pulchella Pascoe, 1858 and G. vestalis Heller, 1934 by the integument colouration, shape of elytra, and yellow hairs covering the body; however, it can be easily distinguished from them by the appendiculate claws (simple claws for G. pulchella and G. vestalis) and distinct shape of the male genitalia, viz. median lobe distinctly curved in lateral view (vs median lobe weakly curved in lateral view in G. pulchella and G. vestalis), tegmen with lateral lobes medium sized and wider (vs tegmen with lateral lobes distinctly elongate and slender in G. pulchella and G. vestalis), basal piece arcuate distally (vs basal piece bifurcated distally in G. pulchella and G. vestalis), ringed part of tegmen constricted near widest portion (vs ringed part of tegmen weakly geniculated in G. pulchella and G. vestalis), manubrium of ringed part hollow, constricted just before basal end, which is short, and abridged by rectangular membrane (vs manubrium of the ringed part closely approximated from middle, continued to distal end as a slender rod in G. pulchella and G. vestalis), median arm of spiculum gastrale slightly longer than lateral arms (vs median arm of spiculum gastrale distinctly longer than lateral arms in G. pulchella and G. vestalis) and RS composed of two closely appressed, sclerotised plates, of which ventral plate is dorso-ventrally flattened to encase dorsal plate, while dorsal plate is made up of two closely approximated inconspicuous rods, forming tubular structure; proximal ends bear two asymmetrical, falcate tines, one tine being longer than other; apex of longer tine bearing divaricate claw like projection (vs RS without any modifications as previous but made up of 4 rods in G. pulchella and 3 rods in G. vestalis). In female genitalia, spermatheca with stalk strongly moulded into U-shaped tubule (vs stalk slightly curved basally in G. pulchella and G. vestalis). It is also similar to Bifidunguiglenea gestroi (Gahan, 1894) in the integument colouration, body shape and yellow-haired maculae, but can be easily distinguished from the latter by the following characters: claws appendiculate (vs claws bifid in B. gestroi); male genitalia with median lobe distinctly curved in lateral view (vs median lobe indistinctly curved in lateral view in B. gestroi); tegmen with lateral lobes medium sized and wider (vs lateral lobes distinctly short and stout in B. gestroi) lateral lobes separated and ventral face sparsely covered with randomly distributed suberect setae (vs lateral lobes confluent except apex, ventral face densely covered with suberect setae in B. gestroi); ringed part of tegmen constricted near widest portion (vs ringed part of tegmen weakly geniculated in B. gestroi); manubrium of ringed part hollow, constricted just before basal end; basal end short, abridged by rectangular membrane (vs manubria of the ringed part joined with each other at basal third, continued to basal end as short, slender rod in B. gestroi); tergite VIII with apical margin truncated (vs apical margin of tergite VIII angulately projected at middle in B. gestroi); female genitalia with stalk of spermatheca strongly moulded into U-shaped tubule (vs spermatheca with stalk nearly straight in B. gestroi). Etymology The name refers to the type locality of the species, Vellayani campus of the Kerala Agricultural University, situated on the banks of the Vellayani lake, on the outskirts of Trivandrum, in the capital city of Kerala, India. Life history All three specimens were collected by the blue light of Actinic BL TL 8 W tubes. Distribution India (Kerala). Remarks on generic affinity Glenea vellayaniensis sp. nov. is an atypical member of the genus Glenea because of the appendiculate female claws. Claws vary greatly between the males of Glenea spp. (Gahan 1897; Lin et al. 2009b; Lin and Yang 2011), but all of them have simple claws in the females. When the female claws are not simple, they are placed in their own genus in most cases (Lin et al. 2009a; Lin and Tavakilian 2012). The genera Eumecocera Solsky, 1871 and Pareutetrapha Breuning, 1952 have appendiculate claws in both male and female. However, the new species can not be combined with any of these genera because of the presence of a lateral elytral carina, a truncated elytral apex, and distinctly different male genitalia. Meanwhile, some members currently placed in the genus Glenea also have appendiculate claws in both sexes, such as G. lineatocollis Thomson, 1860 and G. tenuilineata Thomson, 1879. However, the new species differs from them not only by morphological characters such as elytral shape, colour and haired pattern, but also by the fe, Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 208-217, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Pascoe FP. 1858. XVII. On new genera and species of Longicorn Coleoptera. Part III. Trans R Entomol Soc Lond. (2) 4 (6 - 7): 236 - 266.","Heller KM. 1934. New and Little-Known Philippine Coleoptera. Philipp J Sci. 54 (2): 279 - 307.","Gahan CJ. 1894. Viaggio di Leonardo Fea in Birmania e Regioni vicine. LVI. A list of the Longicorn Coleoptera collected by Signor Fea in Burma and the adjoining regions, with descriptions of the new Genera and species. Ann Mus Civ Stor Nat Genova. 34: 5 - 104.","Gahan CJ. 1897. Notes on the longicorn genus Glenea Newm., with descriptions of new species. Ann Mag Nat Hist. 19 (6): 473 - 493. doi: 10.1080 / 00222939708680567","Lin M-Y, Tavakilian G, Montreuil O, Yang X-K. 2009 b. A study on the indiana & galathea speciesgroup of the genus Glenea, with descriptions of four new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Ann Soc Entomol Fr. 45 (2): 157 - 176. doi: 10.1080 / 00379271.2009.10697599","Lin M-Y, Yang X-K. 2011. A new species Glenea shuteae sp. nov. from Yunnan, China, compared with Glenea decolorata Heller (Coleoptera, Cerambycidae, Lamiinae, Saperdini). Acta Zoot Sinica. 36 (1): 40 - 44.","Lin M-Y, Montreuil O, Tavakilian G, Yang X-K. 2009 a. Reinstatement of the genus Heteroglenea Gahan, with four new combinations, four new synonyms and three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Zootaxa. 2137: 1 - 22. doi: 10.11646 / zootaxa. 2137.1.1.","Lin M-Y, Tavakilian GL. 2012. A New Genus Bifidunguiglenea gen. nov. is erected for the Species Glenea gestroi Gahan (Cerambycidae: Lamiinae: Saperdini). PLoS ONE. 7 (7): e 40768. doi: 10.1371 / journal. pone. 0040768","Solsky S. 1871. Coleopteres de la Siberie orientale. Horae Soc Entomol Rossicae. 7 (1870 - 1871): 334 - 406.","Breuning S. 1952. Revision einiger Gattungen aus der Gruppe der Saperdini Mulsant. (Col., Cerambycidae). Entomol Arb Mus G Frey. 3: 107 - 211.","Thomson J. 1860. Essai d'une classification de la famille des cerambycides et materiaux pour servir a une monographie de cette famille. Paris: chez l'auteur [James Thomson] et au bureau du tresorier de la Societe entomologique de France; p. xvi + 1 - 128.","Thomson J. 1879. Typi Cerambycidarum Appendix 1 a. Revue Mag Zool Paris. 7 (3): 1 - 23."]}

Glenea pseudoalbosignatipennis sp. nov. (Figures 26–33, 43) Type material HOLOTYPE: ♀, with labels as follows: (1) India, Karnataka, Shimoga Dist., Agumbe Ghat, 2000 ft., V.2001 (CCH). (2) HOLOTYPE/ Glenea pseudoalbosignatipennis sp. nov. /des. Hiremath & Lin, 2020 (red label). Paratypes (4 specimens, with a white locality label as given below, besides a second pink label: ‘ PARATYPE / Glenea pseudoalbosignatipennis sp. nov. /des. Hiremath & Lin, 2020’): 3♂, India: Kerala: Kozhikode/ Chappanthottam: Melukavu Panchayat / 11.70055556 N, 75.81833333 E / 09. December 2018/S. R. Hiremath Coll./Ex. Syzygium jambos; 1♂, with label as follows: India: Karnataka / Kudremukha Peak / 13.13361111 N, 75.28416667 E / 13. May 2011, 1195 m/K.D. Prathapan & K. Shameem Coll. Note: the paratypes were lost in a fire accident between acceptance and publication of this manuscript. However, all specimens were thoroughly studied, and relevant measurements and all necessary illustrations including that of the male genitalia are provided. Description Male (n = 4) (Figure 26 (a–d)). Body length measured from vertex to elytral apex 10.74–11.52 mm; humeral width 3.66–3.96 mm. General body colour brick reddish brown, head and pronotum darker than elytra, faintly covered with fine, minute, recumbent, yellowish grey hairs, denser on elytra, legs and ventral side of body. Head with frons, vertex, postclypeus, labrum, base of mandibles brick reddish brown; anteclypeus yellowish brown with marginal angles ornamented with yellow spot; eyes, apical half of mandibles black; labial and maxillary palpi yellowish brown. Frons with lateral sides ornamented with thickly haired, creamy yellow, longitudinal band at base of antennal tubercles, continued between upper eye lobes, traversing vertex as slightly narrowed longitudinal bands; frons between lateral bands moderately adpressed with creamy yellow hairs. Posterior sides of eye lobes ornamented with narrow bands of similar hairs, which continue onto genae as wide, pubescent banding, anteriorly merging with lateral bands of frons on their inner side. Base of anteclypeus ornamented with creamy yellow, transverse, thickly haired band, interspersed on lateral sides with three pairs of elongate, suberect, dark brown setae arising from respective punctures. Labrum adorned on lateral sides with two pairs of punctures, each giving rise to paired, conjoint, elongate, suberect, dark brown setae (however, in one specimen these setae are asymmetrical with right side bearing one extra seta arising separately on outer side). Inner margins of eye lobes, frons, vertex and genae interspersed with randomly distributed, dark brown, suberect setae arising from respective punctures. Head slightly narrower than pronotum, moderately covered with deep punctures; frons weakly convex in lateral view, medially impressed with fine, dark brown sulcus, running from postclypeus to vertex; antennal tubercles weakly produced, divergent, widely separated at base, area between antennal tubercles flat, anteriorly inclined; eyes finely faceted, distinctly emarginated; upper eye lobes connected to lower eye lobes by 6–7 rows of ommatidia; lower eye lobes oval, 2.13–2.38 times as long as genae. Antennae surpass elytral apex at base of antennomere IX, 1.32–1.34 times as long as body. Antennomeres I–IX reddish brown, antennomeres I–III shiny, antennomeres IV–IX matt in appearance, antennomeres X and XI dark brown, matt in appearance. Antennomeres I–VI fringed beneath with elongate, dark brown, suberect setae, shorter on antennomeres V and VI. Base of scape towards outer side on dorsum, ventro-apical side of antennomeres VII and VIII bearing a single, dark brown, elongate, suberect seta. Integument of antenna covered with fine, yellowish grey, recumbent, faint hairs on scape, pedicel, base and ventral side of antennomeres III; remaining antennomeres adorned with dark brown recumbent setae, interspersed with a few randomly distributed minute, yellowish grey suberect setae. Scape cylindrical, weakly and gradually narrowed towards base; base of scape emarginated on inner side. Antennomere XI gradually and weakly thickened towards obtusely pointed apex. Ratio of lengths of antennomeres: 1.00: 0.22–- 0.24: 1.24–1.26: 1.00: 1.04: 1.04: 1.04: 0.91–0.92: 0.91–0.92: 0.80–0.83: 1.04. Prothorax dark brown, interspersed with randomly distributed red brown to dark brown suberect setae. Pronotal disc medially ornamented with a distinctly narrow, creamy yellow (creamy white in preserved specimens), longitudinal band, interrupted postmedially and reappears as oval, haired spot near basal margin; sublateral and lower lateral sides ornamented on each side with a broad, thickly haired, creamy yellow (creamy white in preserved specimens) longitudinal band, inner margins obliquely bisinuate, outer margins nearly straight; central area of these bands ornamented on each side with a pre-medial, irregularly oval, dark brown spot, remaining area interspersed with six dark brown, remotely and randomly distributed, elongate, suberect setae arising from dark-spotted base. Pronotum 1.13–1.26 times wider than long, 0.56–- 0.60 times as long as humeral width, apical margin slightly broader than basal margin; lateral sides slightly swollen at middle and weakly constricted just behind middle; pronotal disc densely covered with coarse, deep punctures; centro-notal area distinctly convex, postmedially continued as a short ridge, longitudinally impressed with a fine, dark brown sulcus; apical margin transversely straight, basal margin distinctly convex at middle. Scutellum short, tongue-shaped, apical half thickly covered with creamy white, adpressed hairs; rounded apically. Elytra reddish brown in general, postmedial disc darker than premedial disc, basal third interspersed with several dark brown, elongate, suberect setae; sparsely fringed with a few similar setae at apex. Elytral disc on each side, ornamented with one epipleural, three premedial, one medial and one postmedial, thickly haired spots; elytral epipleura ornamented with a small, creamy white, oval spot at base; premedian spots creamy white, disposed broadly in triangular fashion: first spot circular, situated at basal third, second spot smaller than previous, circular, obliquely placed, just behind basal third in close proximity to sub-lateral carina, third spot oblong, largest of premedial spots, placed behind basal third along sutural margin, and close to middle; medial spot largest, creamy yellow (creamy white in preserved specimen), irregularly oval, situated between sutural and sub-lateral margins, apical margin of this spot convex at middle, basal margin distinctly notched at middle; postmedian spot circular, creamy white, situated at apical fourth, subequal to premedian, sutural spot. Elytra elongate, about 0.68–0.69 times as long as body, 3.33–3.59 times as long as pronotum, 2.00–2.02 times as long as humeral width, wider at base, gradually narrowed towards apex; apex generally emarginated (Figures 26 (a) and 43(a)) (slightly truncate in one specimen), sutural angle dentate, marginal angle stretched into acute spine. Humeral prominence distinctly and angularly produced (Figure 43 (b)). Elytra with premedial disc covered with coarse punctures, punctures on postmedial disc gradually turning finer and sparsely distributed. Elytral disc impressed with two longitudinal rows of coarse punctures, gradually becoming finer towards apex between sublateral and lateral margin. Sublateral margin impressed with two longitudinal carinae: one begins at base of humeral prominence, other begins just below humeral prominence; carinae parallel to each other before merging with one another at apical fourth, continued as short carina, terminating along acute spine at marginal angle of elytral apex. Elytra in lateral view, weakly convex at base and apical fourth, flat at middle, slightly sloped pre-apically. Legs yellowish brown, except reddish brown tarsal claws. Outer and inner margins of fore-tibiae sparsely fringed with a few yellowish brown, suberect setae. Inner margins of mid and hind tibiae fringed with medium sized, suberect setae same colour as previous. Tarsi interspersed with a few dark brown, recumbent setae at their apex. Tarsal claws divaricate. Sternites with pro-, meso- and metasternum reddish brown. Mesepisternum ornamented on each side with a medium sized, creamy white, oval spot. Lateral sides of metasternum ornamented on each side with a large, wide, creamy yellow (creamy white in preserved specimens) haired band; sternal space between lateral bands broadly triangular, densely adpressed with golden yellow hairs; posterior margin of metasternum, on either side of discrimen, ornamented with small, transverse oval, creamy white spots. Metepisternum on anterior half ornamented with a creamy white, oval, haired spot, smaller than that on mesepisternum; posterior half ornamented with a longitudinal haired band, wider anteriorly and narrowed posteriorly. Outer face of metacoxa with a transverse, creamy white haired band. Abdomen with ventrites reddish brown. Ventrites I–IV ornamented on each side with a pair of haired bands side by side; inner band on ventrite I transverse, largest among all remaining bands; bands on ventrite IV sometimes fused with each other. Sternite VII ornamented on each side with two haired spots, obliquely placed one behind the other, posterior spot larger than anterior; Sternite VII convex pre-apically, 0.25 times as long as total length of abdomen, and 1.67–1.71 times as long as preceding segment; apical margin distinctly notched at middle. Male genitalia (Figures 27–29). Tergite VIII (Figure 27) U-shaped, with its apical margin broadly obtusely angulate; lateral sides fringed with a few small- to medium-sized, red brown setae at middle, fringed with elongate, red brown, curved setae up to apical margin; apical margin at its middle densely fringed with medium sized, light brown setae; median disc randomly covered with small, red brown, recumbent setae, arising from oval to irregularly raised structures, interspersed with circular bodies. Sternite VIII boat shaped, red-brown; basal margin ornamented on either side of middle with transversely arranged minute, light coloured spinules, sometimes interspersed with a few suberect setae, lateral sides covered with a few suberect setae; apical margin on either side of middle covered with several irregularly dispersed elongate, red brown, suberect setae, interspersed with several minute spinules in a small patch. Spiculum gastrale Y-shaped, 1.81 times as long as spiculum relictum; median arm shorter than lateral arms, separated up to preapical area, abridged medio-longitudinally by flat membrane; apex curved leftward. Spiculum relictum straight or sometimes curved near its middle. Tegmen (Figure 28 (a–c)) 2.65 mm long; in lateral view (Figure 28 (c)), distinctly concave near middle and straight on either side. Basal piece present, distal margin curved, postmedial disc impressed with a transversely curved ridge; entire surface covered with minute, angulate spinules, denser premedially. Roof present. Ringed part converging, constricted near widest portion; manubrium broadly V-shaped with arms distinctly sinuate, basal end surmounted by a short dorso-ventrally flattened areolated membrane (Figure 28 (a)). Lateral lobes (Figure 28 (b)) elongate, cylindrical, 0.33 times as long as tegmen; basal margin obliquely straight; inner and outer margins straight up to apex, apex obliquely rounded; base of inner margins slightly produced behind as thick, curved, widened rod; integument light coloured at apex, lateral margins bearing a few, small, suberect setae, impressed with two elongate setae near apical fourth, apex interspersed with a few randomly distributed, distinctly elongate setae; disc of lateral lobes randomly covered with several medium sized setae except basal margin; basal margin impressed with similar setae arranged transversely. Median lobe (Figure 29 (a–b)) subequal to tegmen, curved in lateral view (Figure 29 (a)); basal struts begin near pre-apical area; ventral plate (Figure 29 (b)) with apical margins appears grooved, preapical area on either side of middle randomly covered with a few minute punctures. Endophallus (Figures 30–32) excluding APH 2.29 times as long as median lobe. BPH 0.57 times as long as median lobe, cylindrical at apical three-fourths, basal fourth distinctly spherical; membrane transversely plicate, densely covered with semi-circular spicules. MPH with MT distinctly short, 0.21 times as long as median lobe, 0.39 times as long as CT; ventral membrane with anterior and posterior half bulged, middle portion depressed; anterior half of ventral membrane transversely plicate, covered with circular spicules. MT bears two pairs of sclerotic plates (Figure 31 (a–c)): anterior pair present adpressed to dorsal membrane at its middle, broadly rectangular, widest at middle, proximal end slightly narrowed; posterior pair placed post-medially, suspended towards ventral membrane, irregular, conical with surface wavy. CT distinctly elongate, subequal to BPH, uniformly cylindrical, with distal end bearing a weakly developed medio-dorsal swelling on ventral membrane; membrane of CT at basal third covered with minute, angulate spicules, remaining surface up to pre-apical area randomly and densely covered with minute circular spicules, lateral sides remotely interspersed with larger, hollow, circular spicules, ventral side with a medio-longitudinal patch of densely distributed angulate spicules; apical area on dorsal side covered with medium-sized, angulate spicules, along with a few larger, hollow, circular spicules on lateral side. PB vessel-shaped, 0.41 times as long as median lobe, with anterior half cylindrical, posterior half distinctly spherical; cylindrical portion with anterior third weakly reticulate, randomly covered with large, hollow, circular spicules, densely and compactly covered with setae like adpressed spicules, becoming fine and short towards bulged portion, lateral side interspersed with a few large, hollow, circular spicules; bulged portion densely and uniformly covered with minute, irregular to semi-circular spicules. RS (Figure 32 (a–b)) as long as CT, 0.55 times as long as median lobe, curved and indistinctly twisted in lateral view, composed of three rods – two lateral and one median; lateral rods abridged at pre-apical area, proximal ends weakly spatulate with transversely corrugated integument, distal ends obtusely pointed; median rod distinctly narrow, with apex giving rise to ED. ED single. Female (n = 1) (Figure 26 (e)). Body length 15.10 mm, humeral width 5.20 mm. Similar to male in general appearance with the following differences: antennae shorter compared with males; pronotum with inner margins of sublateral bands distinctly bisinuate in apical half, obliquely straight in basal half; elytra with first spot largest among premedial spots disposed in triangular fashion; third spot circular to oval, subsutural in position, slightly subequal to second spot; sternite VII medially impressed with a dark brown longitudinal sulcus. Remarks RS closely embedded in APH, concealed inside PB of endophallus (Figure 30). Attempts to inflate the APH were futile and resulted in tearing off the endophallus at PB. Hence, APH was not examined. The female description is based on the measurements and image provided by Carolus Holzschuh, which was photographed by Bruno Brudermann (Austria). Differential diagnosis The new species is similar to G. albosignatipennis Breuning, 1950 and G. signaticollis Gahan, 1889. However, it differs from G. albosignatipennis by the following characters: general body colour brick reddish brown (vs reddish brown in G. albosignatipennis); lower eye lobes of male 2.13–2.38 times as long as genae (vs lower eye lobes of male 5 times as long as genae in G. albosignatipennis); pronotum wider than long (vs pronotum as long as wide in G. albosignatipennis); scutellum tongue-shaped (vs scutellum pentagonal in G. albosignatipennis); humeral prominence (Figure 43 (b)) comparatively distinctly protruding (vs humeral prominence (Figure 44 (b)) comparatively less protruding in G. albosignatipennis); lateral and median bands on pronotum not united ante-basally (vs lateral and median bands united with each other ante-basally in G. albosignatipennis); median longitudinal band discontinuous (vs median longitudinal band continuous in G. albosignatipennis); inner margins of lateral bands obliquely sinuate (vs inner margins of lateral bands straight in G. albosignatipennis); base of elytral epipleura with a creamy white, oval, haired spot (vs elytral epipleural spot absent in G. albosignatipennis); premedian disc of elytron with three haired spots, among them one spot is located along sutural margin (vs pre-median disc of elytron with only two haired spots and sutural spot absent in G. albosignatipennis); post-median disc of elytron with a circular, pre-apical haired spot (vs post-median disc of elytron with a curved, pre-apical haired spot in G. albosignatipennis); elytral apex (43a), on each side, generally emarginated in the new species (elytral apex (44a), on each side, obliquely truncated in G. albosignatipennis); mesepisternum ornamented on each side with an oval spot (vs mesepisternum entirely covered with a large, haired spot in G. albosignatipennis); lateral sides of abdominal ventrites I–IV ornamented with two spots on each side, arranged side by side (vs abdominal ventrite II with two spots on each side, arranged side by side, and ventrites I, III and IV with a single transverse band on each side in G. albosignatipennis); sternite VII with two spots on each side, arranged one behind the other (vs abdominal sternite VII with a single, post-median spot on each side in G. albosignatipennis). The new species can be differentiated from G. signaticollis by the following characters: general body colour brick reddish brown (vs reddish brown or sometimes general colour of head, pronotum and underside of the body except sternite VII black in G. signaticollis); lower eye lobes of male 2.13–2.38 times as long as genae (vs lower eye lobes of male 3 times as long as genae in G. signaticollis); pronotum of male wider than long (vs pronotum of male as long as wide in G. signaticollis); scutellum tongue-shaped (vs scutellum semicircular in G. signaticollis); humeral prominence (Figure 43 (b)) comparatively distinctly protruding (vs humeral prominence (Figure 45 (b)) comparatively less protruding in G. signaticollis); lateral and median bands on pronotum not united ante-basally (vs lateral and median bands united with each other ante-basally in G. signaticollis); median longitudinal band on pronotal disc discontinuous (vs median longitudinal band on pronotal disc continuous or reduced to anterior and posterior spots or broadly divided into halves in G. signaticollis); inner margins of lateral bands of pronotum obliquely sinuate (vs inner margins of lateral bands of pronotum straight or slightly sinuate or indented at middle in G. signaticollis); central area of lateral bands on, Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 225-234, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Breuning S. 1950. Descriptions de nouveaux Lamiaires de l'Inde (Coleopteres). Indian For Rec (New Ser) Entomol. 7 (8 [1949]): 255 - 265.","Gahan CJ. 1889. Descriptions of new or little-known species of Glenea in the collection of the British Museum. Trans R Entomol Soc Lond. 1889 (2): 213 - 225."]}

Glenea vestalis Heller, 1934 stat. reinstated (Figures 19–25) Glenea vestalis Heller, 1934: 284, Figure 2. TL: Philippines. TD: SMTD. Glenea pulchella: Aurivillius 1926: 111 (partim). Glenea (Glenea) pulchella: Breuning 1956a: 195 (partim). Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956a: 196. [Unavailable name, infrasubspecies from Tonkin, Vietnam] Glenea (Glenea) pulchella m. preapiceconjuncta Breuning, 1956a: 197. [Unavailable name, infrasubspecies from Ceram, Indonesia] Glenea (Glenea) pulchella m. transversevittata Breuning, 1956a: 197. [Unavailable name, infrasubspecies from Fundortangabe] Glenea (Glenea) pulchella m. vestalis: Breuning 1956a: 197. Glenea (Glenea) pulchella: Hüdepohl 1996: 18. [Misidentification] Type specimen examined Holotype (Figure 19 (a–c)), ♀, Philippines, Arorey, 1923.VIII (SMTD). Other Specimens examined Philippines: 1 ♂, ‘paratype’ of Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956, Dapitan, Mindanao, Baker (MHNL, ex collection P. Lepesme); 1 ♂, ‘paratype’ of Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956, Zamboanga, Mindanao, Baker (MHNL, ex collection P. Lepesme); 1 ♀, ‘paratype’ of Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956, Surigao, Mindanao, Baker (NHMB (Frey)); 1 ♂, Philippines (BMNH); 1 ♂, 2 ♀♀, Zamboanga Mindanao, Baker (NMNH); 2 ♀♀, Island Samar, Baker (NMNH); 4 ♂♂, 2 ♀♀, Kolambugan, Mindanao, Baker (NMNH); 1 ♂, 1 ♀, Davao Mindanao, Baker (NMNH); 2 ♂♂, 2 ♀♀, Mt. Makiling, Luzon, Baker (NMNH); 2 ♂♂, 4 ♀♀, Dapitan, Mindanao, Baker (NMNH); 1 ♂, 1 ♀, Butuan Mindanao, Baker (NMNH); 1 ♀, Surigao, Mindanao (NMNH, Tippmann Coll. ’57, 213112); 2 ♀♀, Philippines, Ch Semper (MNHN); 1 ♂, Mindanao (Figure 20 (a,b), MNHN); 3 ♀♀, Mindanao, 1903–1904, J. Waterstradt (MNHN); 1 ♀, Philippines, N. Luzon, Cagayan, Sta. Ana, June 2014, local coll. (DHCO). Malaysia 1 ♀, Java (Meuwen Bay) (MNHN); 2 ♀♀, ‘paratypes’ of Glenea (Glenea) pulchella m. preapiceconjuncta Breuning, 1956, Borneo, Sandakan, Baker (NHMB (Frey)); 1 ♀, Malacca, Perak, W. Doherty (MNHN). Indonesia 1 ♀ (Figure 21 (a,b)), ‘type’ of Glenea (Glenea) pulchella m. preapiceconjuncta Breuning, 1956, Moluccas, Ceram (= Moluques, Seram) (BMNH, ex Fry Coll, 1905.100); 1 ♀, Maluku, Seram, 35 km E Pasahari, Unit O, 24–30 October 1998, leg. J. Horák (DHCO). Vietnam 1 ♀ (Figure 22 (a,b)), ‘type’ of Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956, Tonkin, Hoa-Binh, A. de Cooman (MHNL, ex Coll. Lepesme, 2002, ex Coll. J. Clermont). Description complementary to Heller (1934) and Breuning (1956a). Male genitalia (Figures 23–24) Tegmen length about 3.5 mm; lateral lobes extremely long and slender, each about 1.8 mm long and 0.05 mm wide, apex covered with short, reddish brown setae; basal piece bifurcated distally; median lobe plus median struts slightly curved, much shorter than tegmen (22:35); median struts about two-thirds of whole median lobe in length; dorsal plate slightly shorter than ventral plate; ventral edge of median orifice slightly pointed; median foramen elongated; internal sac about 3 times as long as median lobe plus median struts, with four pieces of basal armature and three rods; rods about 1.0 mm, shorter than one-third of tegmen. Tergite VIII elongate, U-shaped with its apical margin weakly notched in middle, integument with short setae. Ventrite IX subequal to ringed part of tegmen in length. Female genitalia (Figure 25): Spermatheca elongate, with its stalk curved at base and capsule oval. Spermathecal gland originating from a distinctly sclerotised ringed plate (Figure 25). Tignum much longer than abdomen. In our observation, tignum 8.5 mm for an adult with a 5.3 mm long abdomen in ventral view. Diagnosis This species is very similar to G. pulchella Pascoe at first glance, with body reddish brown and similar yellow-haired maculae. However, they differ from each other in the absence vs presence of the small spot at the centre of apical half. Male genitalia exhibit distinct differences also: lateral lobes shorter than half of tegmen and apex of tergite VIII truncated in G. pulchella (Figures 15 and 16) vs lateral lobes subequal to half of tegmen and apex of tergite VIII weakly notched at middle in G. vestalis (Figures 23 and 24). Distribution Philippines, Malaysia, Indonesia, Vietnam. Remarks Breuning (1956a) treated G. vestalis as a morph of G. pulchella, and subsequent authors considered them synonyms (Hüdepohl 1996; Tavakilian and Chevillotte 2020). However, they are two distinct species, and G. vestalis is herein reinstated from synonymy with G. pulchella. Aurivillius (1926: 111) wrote: ‘Specimens from Mindanao have a small sulphur yellow lateral dot behind the middle of elytra; this dot is wanting in specimens from Borneo and Malacca but still more developed in a specimen from Ceram’. Examination of specimens from these localities by the second author revealed that specimens referred to as having a ‘small sulphur yellow lateral dot’ behind the middle of the elytra, from Philippines and Ceram (now Indonesia, Maluku, Seram), are G. vestalis, while the specimens from Borneo and Malacca, in which ‘this dot is wanting’, are G. pulchella. Breuning (1956a) described three morphs, which are all infrasubspecific. The second author examined the ‘type’ and ‘paratypes’ of these morphs; the ‘type’ of Glenea (Glenea) pulchella m. preapiceconjuncta Breuning, 1956 (Figure 21 (a,b)) from ‘ Insel Ceram im Britischen Museum’, and the ‘type’ of Glenea (Glenea) pulchella m. postmediopunctata Breuning, 1956 (Figure 22 (a,b)) from ‘ Tonkin: Hoa-Binh in coll. Lepesme’ are both females and are both considered Glenea vestalis Heller, 1934 based on the presence of a postmedial, sulphur yellow lateral dot on the elytra. The ‘type’ of Glenea (Glenea) pulchella m. transversevittata Breuning, 1956, based on ‘ein female ohne Fundortangabe in der Sammlung Itzinger’, could not be examined; however, in all probability it belongs to G. vestalis, since Breuning (1956a: 197) compared it with m. postmediopunctata and arranged it between m. preapiceconjuncta and m. vestalis Hell., Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 221-225, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Heller KM. 1934. New and Little-Known Philippine Coleoptera. Philipp J Sci. 54 (2): 279 - 307.","Aurivillius C. 1926. Revision of the Philippine species of the Glenini (Coleoptera, Longicornia). Philipp J Sci. D 30: 89 - 115.","Breuning S. 1956 a. Revision der Gattung Glenea Newman. Entomol Arb Mus G Frey. 7 (1): 1 - 199.","Hudepohl K-E. 1996. Uber sudostasiatische Cerambyciden XIV (Coleoptera, Cerambycidae, Lamiinae). Entomofauna Z Entomolog Ansfelden. 17 (1): 1 - 24.","Tavakilian G, Chevillotte H. 2020. Titan: base de donnees internationales sur les Cerambycidae ou Longicornes. Version; [accessed 2020 Jan 8]. http: // titan. gbif. fr / index. html."]}

Glenea pulchella Pascoe, 1858 (Figures 13–18) Glenea pulchella Pascoe, 1858: 260. TL: Malacca. TD: BMNH. Glenea vesta Pascoe, 1866: 260, pl. 28, Figure 3. [Unnecessary new name for Glenea pulchella Pascoe, 1858] Glenea pulchella: Aurivillius 1926: 111 (partim). Glenea (Glenea) pulchella: Breuning 1956a: 195 (partim). Type specimen examined Holotype (Figure 13 (a–e)), ♂, Malacca (BMNH, ex Pascoe Coll. 93–60). Other specimens examined Malaysia: 1 ♀, Bornéo Occ., Pontianak, 1899 (MNHN) (Figure 14); 1 ♀, Sarawak (MNHN, Museum Paris Coll. H.W. Bates 1952, ex Musaeo, H.W. Bates 1892); 3 ♂♂ 8 ♀♀, Sandakan Borneo, Baker (NMNH); 1 ♀, Sabah, Mt. Trus-Madi, 18 March 2011, local coll. (DHCO); 1 ♀, Sabah Crocker Range, vic. Trus Madi, 13 March 2000, local coll. (DHCO); 1 ♀, Sabah Crocker Range, April 1998, local coll. (DHCO); 1 ♂, Borneo (IRSNB, ex Coll. Nonfried); 1 ♀, Borneo (IRSNB, ex Coll. F. de Moffaris); 1 ♀, Borneo, Pontanak (NHMB, ex FREY); 2 ♀♀, Borneo Occ. Pontianak, 1899 (MNHN); 1 ♂, Borneo, 1891, W. Doherty (MNHN, ex Coll. R. Oberthür, 1952); 1 ♀, Kuching, 1902 (MHNL, ex collection P. Lepesme); 1 ♀, Borneo occ., Pontianak, 1899 (MHNG); 4 ♀♀, Borneo occ., Pontianak, 1899 (MNHN). Singapore 2 ♂♂, Singapore, coll. Wallace (MNHN, ex Musaeo James Thomson); 2 ♂♂, Singapore (MNHN); 1 ♀, Singapore (BMNH); 1 ♀, Singapore (MNHN, ex Musaeo Mniszech). Description complementary to Pascoe (1858) and Breuning (1956a). Male: length: 8.8–10.3 mm, humeral width: 2.6–3.1 mm. Female: length: 11.3–13.4 mm, humeral width: 3.6–4.3 mm. Both male and female with simple claws. Male genitalia (Figures 15–16) Tegmen length about 2.6 mm; lateral lobes long and slender, each about 1.0 mm long and less than 0.1 mm wide, apex covered with short, reddish brown setae; basal piece bifurcated distally; median lobe plus median struts slightly curved, shorter than tegmen (11:13); the median struts about 2/3 of the whole median lobe in length; dorsal plate subequal to ventral plate; ventral edge of median orifice round; median foramen hardly elongated; internal sac 2 times longer than combined length of median lobe and median struts, with 2 pairs of basal armature and 4 rods; each rod about 1.0 mm, shorter than half of tegmen. Tergite VIII trapeziform, apex truncated, with short setae. Length of ventrite IX subequal to ringed part of tegmen. Female genitalia (Figures 17–18) Spermatheca rounded, with a moderately long and curved stem at its base. Spermathecal gland originating from a distinctly sclerotised ringed plate (Figure 17). Tignum much longer than abdomen. Tignum 8.8 mm for an adult with a 5.3 mm long abdomen in ventral view. Diagnosis Glenea pulchella differs from G. vellayaniensis sp. nov. by colour and haired maculae in the following body regions: (1) scape and antennomere II dark brown to black (vs scape and antennomere II reddish brown in G. vellayaniensis sp. nov.); (2) sublateral macula on pronotum made up of yellow hairs with straight inner margin (vs sublateral macula on pronotum made of yellow hairs intermixed with creamy white hairs with proximally concave inner margin in G. vellayaniensis sp. nov.); (3) basal elytral yellow-haired maculae semicircular (vs basal elytral yellow-haired maculae bean-shaped in G. vellayaniensis sp. nov.); (4) middle elytral yellow-haired maculae oval, without small spots posteriorly (vs median maculae on elytra transversely oval with small spots posteriorly in G. vellayaniensis sp. nov.); (5) elytral apex not covered by yellow hairs, but last maculae located before apex (vs elytral apex covered with yellow haired maculae in G. vellayaniensis sp. nov.). Distribution Malaysia, Singapore. Remarks Pascoe (1866: 260) wrote ‘I have altered the specific name pulchella, it having been previously used by Hope’. And Pascoe gave the species the new name ‘ Glenea vesta ’. Pascoe (1867: 370) wrote ‘ Glenea pulchella Hope, sec. J. Thomson, Ess. & c., p. 58’ from Sarawak. We checked page 58 of Thomson (1857) and found nothing related to ‘ Glenea pulchella Hope’; then we checked page 58 of Thomson (1860), and there it was written ‘ Glenea pulchella, Hope Syn.: G. conspuncta, Melly’. However, ‘Hope’s species was not described before 1860’ (Aurivillius 1926: 111). ‘ Glenea pulchella Hope’ described by Pascoe (1867) was renamed Glenea pascoei Aurivillius, 1923, while ‘ Glenea pulchella Hope’ described by Thomson (1860) from Sylhet was renamed Glenea pulchra Aurivillius, 1926. Glenea pulchella Pascoe, 1858 is the earliest name, and therefore the new name ‘ Glenea vesta ’ is not required. Breuning (1956a) treated G. vestalis Heller, 1934 as a morph and described several morphs (infrasubspecific). However, his ‘morphs’ are good species. Mukhopadhyay and Biswas (2000) reported the distribution range of G. pulchella as India: Meghalaya, Bangladesh, Burma; while Mitra et al. (2016) mentioned India: Karnataka, Manipur, Meghalaya, Sikkim, Tamil Nadu, West Bengal. These distribution records are not reliable, based on neither specimens nor trustable identifications. The first author inquired for supportive material from the Zoological Survey of India (ZSI), Kolkata; however, there are no specimens of G . pulchella in the collection. The known localities of this species are, to our knowledge, limited to Malaysia and Singapore., Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 217-221, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Pascoe FP. 1858. XVII. On new genera and species of Longicorn Coleoptera. Part III. Trans R Entomol Soc Lond. (2) 4 (6 - 7): 236 - 266.","Pascoe FP. 1866. Catalogue of Longicorn Coleoptera, collected in the Island of Penang by James Lamb, Esq. (Part I.). Proc Zool Soc Lond. 1866: 222 - 267.","Aurivillius C. 1926. Revision of the Philippine species of the Glenini (Coleoptera, Longicornia). Philipp J Sci. D 30: 89 - 115.","Breuning S. 1956 a. Revision der Gattung Glenea Newman. Entomol Arb Mus G Frey. 7 (1): 1 - 199.","Pascoe FP. 1867. Longicornia Malayana; or, a Descriptive Catalogue of the Species of the three Longicorn Families Lamiidae, Cerambycidae and Prionidae collected by Mr. A. R. Wallace in the Malay Archipelago. (Part IV). Trans R Entomol Soc Lond. 3 (3, 4): 337 - 464.","Thomson J. 1857. Essai monographique sur le groupe des Tetraophthalmites, de la famille des cerambycides (longicornes). In: Thomson J, editor. Archives Entomologiques ou recueil contenant des illustrations d'Insectes nouveaux ou rares. Tome premier. Paris: Bureau du Tresorier de la Societe Entomologique de France; p. 45 - 67, 514 + [1].","Thomson J. 1860. Essai d'une classification de la famille des cerambycides et materiaux pour servir a une monographie de cette famille. Paris: chez l'auteur [James Thomson] et au bureau du tresorier de la Societe entomologique de France; p. xvi + 1 - 128.","Aurivillius C. 1923. Cerambycidae: Lamiinae II. Pars 74. In: Schenkling S, editor. Coleopterorum catalogus. Volumen XXIII. Cerambycidae II. Berlin: W. Junk; p. 323 - 704.","Heller KM. 1934. New and Little-Known Philippine Coleoptera. Philipp J Sci. 54 (2): 279 - 307.","Mukhopadhyay P, Biswas S. 2000. Insecta: Coleoptera: Cerambycidae. Fauna Meghalaya State Fauna Ser Zool Surv India Kolkata. 4 (5): 41 - 67.","Mitra B, Das P, Mallick K, Chakraborti U, Majumder A. 2016. New records of longhorn beetles (Cerambycidae: Coleoptera) from Manipur State India with Checklist. J Zool St. 3 (1): 19 - 26."]}

Glenea signaticollis Gahan, 1889 (Figures 39–42, 45) Glenea signaticollis Gahan, 1889: 219. TL: India, Bombay. TD: BMNH. Type specimen examined India (Bombay) (BMNH). Examined based on an image. Other specimens examined 1 ♀, Madura, Coll. Dr. Itzinger (NHMB, ex FREY); 1 ♀, Nilgiri Hills, S. India (G.V. Campbell). 1 ♀, Nilgiri Hills, Coll. A.K. Weld Downing (Travancore Insect Collection, KAU). Description Body reddish brown; antenna reddish brown; head and pronotum reddish brown or black; legs yellowish brown; ventral body black, except for sternite VII reddish brown. Head with frons, genae and scutellum pale yellow in holotype (it is probably a male specimen), while often black in females; anteclypeus, labial and maxillary palpi yellowish brown; postclypeus black, apical margin thickly adpressed with creamy white hairs interspersed with four pairs of suberect, elongate, reddish brown setae; labrum reddish brown, medially interspersed with two suberect, elongate, yellowish brown setae arising from respective punctures, apical margin indisticntly emarginated; frons with two longitudinal, pale yellow-haired bands at sides, extending backwards and traversing vertex; frontal disc, between lateral bands, moderately covered with sparse pale yellow hairs; genae densely covered with pale yellow hairs; lower eye lobes 3 times as long as genae in males, 0.5 times as long as genae in females; frons as wide as one lower eye lobe in males and three-fourths wider than one lower eye lobe in females; head coarsely and densely punctured. Antenna One-quarter longer than body in male, slightly longer than body in female; scape short and thin. Ratio of lengths of antennomeres in female: 1.00: 0.13: 1.40: 1.13: 1.00: 0.93: 0.93: 0.87: 0.80: 0.73: 0.87. Pronotum as long as broad in males, slightly transverse in females, coarsely and densely punctured; prothorax with five longitudinal, pale yellow-haired bands: one located in middle of pronotum, narrow, often rubbed at middle in preserved specimens; two located at lateral sides of prothorax, each distinctly broad, indented or straight at middle of inner margins, while outer margins irregularly sinuated, top three bands narrowly united ante-basally in females, while broadly united in holotype; other two located at ventro-lateral sides of prothorax, each broad, connecting with outer margins of procoxae. Scutellum semi-circular, densely covered with white hairs. Elytra subparallel, apex distinctly truncated, sutural margin pointed apically, marginal angle stretched into elongate spine (Figures 39 (a) and 45(a)); proximal three-fourths of elytral disc coarsely and densely punctured; sublateral margin impressed with two longitudinal carinae: one begins at base of humeral prominence, terminates well before elytral apex, the other begins just below humeral prominence, terminating along apical spine of marginal angle; humeral prominence angularly projected (Figure 45 (b)); elytral disc densely covered with minute yellowish brown hairs, each elytron with seven pale yellowhaired spots: one oval haired spot at base of elytral epipleura; three situated at premedian and disposed broadly in obliquely triangular fashion: first one situated at basal third, in middle of disc between sutural and sublateral margin, second one situated obliquely to first, just behind basal third, along inner side of sublateral carina, third one larger than the first two spots and situated just before middle, situated obliquely to second one at middle of disc between sutural and sublateral margins; remainder of three situated behind middle: first one largest, oval, slightly situated behind middle, close to sutural margin and situated away from sublateral margin; second one circular, situated at apical fourth, situated along inner side of sublateral carina; third one situated apically and transverse. Legs with tarsal claws divaricate. Sternites densely covered with short, fine, grey-white hairs; mesepisternum ornamented on each side with a transverse creamy white, thickly haired spot; metasternum ornamented on each side with a broad, oblique, creamy white, thickly haired spot; metepisternum on each side ornamented postmedially with an oval, creamy white thickly haired spot, much smaller than haired spot on metasternum. Ventrite densely covered with short fine, grey-white hairs; each side of ventrite I–IV covered with two short longitudinal white-haired spots posteriorly, spots often connected to each other by a narrow white haired band; sternite VII, on each side, covered with a short longitudinal white-haired spot postemedially. In female, sternite VII convex, about 0.32 times as long as total abdominal length, about 3.00 times as long as ventrite IV; disc of sternite VII medially impressed with a fine, dark brown longitudinal sulcus, with a short, medial ridge pre-apically; apical margin broadly concave. Female genitalia (Figure 42 (a–b)) with ovipositor about 1.35 mm long, in dorsal view, up to base of coxite lobe, randomly and sparsely distributed with circular punctures, apical third densely covered with short, spiculate grey setae. Coxite lobe (Figure 42 (b)) yellowish brown, randomly and remotely distributed with a few circular punctures; apical margin adorned with about five small setae, flanked on either side with an elongate seta. Stylus (Figure 42 (b)) distinct, dome-shaped; integument yellowish brown; in dorsal view, provided with two circular punctures near base; apex of stylus bears three distinctly elongate setae arising from associated punctures. Vaginal plates sub-rectangular, weakly sclerotised, separated from one another; integument mostly translucent with faint, yellowish brown tinge; basal half linearly reddish brown on lateral sides. Bursa copulatrix elongate, bisinuate with distinct curvature at apical third; proximally narrow tube, gradually widening towards distal end; integument strongly adorned with transverse plications except basal third. Spermathecal duct slightly longer than bursa copulatrix and distinctly longer than spermatheca, enters bursa copulatrix at apical third, basally translucent tubule, followed by thick, yellowish brown, sickle-shaped tube up to middle; immediately behind middle, slightly curved and continued as a slender tube towards apex. Spermatheca and spermathecal gland opens separately on spermathecal duct. Spermatheca yellowish brown, opens on lateral side of apex of spermathecal duct, and connected to it via a distinctly slender tubule; stalk thick, elongate tubule, narrowed basally and curved towards apex; capsule distinctly capitate. Spermathecal gland strongly wrinkled tubular sac, basally without sclerotised ringed plate. In the specimen examined, tignum measured about 5.95 mm in length while abdomen measured about 5.08 mm. Distribution India (Tamil Nadu: Madurai, Nilgiri Hills; either Maharashtra or Karnataka). Remarks The description comes from Gahan (1889) and Breuning (1956b) and from studying a single female specimen along with the images of the holotype and other identified materials of G. signaticollis. The female genitalia is described and illustrated based on a single female specimen held at the Travancore Insect Collection, KAU, India. Gahan (1889) doubtfully indicated Bombay as the type locality; however, ‘Bombay’ usually referred to the erstwhile Bombay presidency in British India and not particularly to a city. Since other materials studied in the present research originate from Western Ghats, it is safe to assume that the holotype might have been collected from the presentday Western Ghats region of either Maharashtra or Karnataka, parts of which were under erstwhile Bombay presidency., Published as part of Hiremath, Sangamesh R. & Lin, Mei-Ying, 2021, Description of two new species of Glenea Newman, 1842 from southern India and reinstatement of Glenea vestalis Heller, 1934 (Coleoptera: Cerambycidae: Lamiinae: Saperdini), pp. 205-245 in Journal of Natural History 55 (3 - 4) on pages 238-243, DOI: 10.1080/00222933.2021.1900442, http://zenodo.org/record/5473739, {"references":["Gahan CJ. 1889. Descriptions of new or little-known species of Glenea in the collection of the British Museum. Trans R Entomol Soc Lond. 1889 (2): 213 - 225.","Breuning S. 1950. Descriptions de nouveaux Lamiaires de l'Inde (Coleopteres). Indian For Rec (New Ser) Entomol. 7 (8 [1949]): 255 - 265.","Breuning S. 1956 b. Revision der Gattung Glenea Newm. (1. Fortsetzung). Entomol Arb Mus G Frey. 7: 671 - 893."]}

Ithocritus similis Bi & Lin, sp. nov. (Figs. 4, 5, 10, 11, 14, 16, 17, 20) Type material. Holotype: male, ��� CHINA. Yunnan, Gongshan / Dulongjiang, Maku / 1500m, 2015.VIII.7 / leg. Xiao-Dong Yang ��� (SNUC). Paratypes: 1 female, same locality as holotype ��� 2015.VII.21 / leg. Chao Wu ��� (CCCC); 1 female, ditto ��� 2015.VIII.9 / leg. Wen-Xuan Bi ��� (CBWX); 1 male, ��� CHINA. Yunnan, Gongshan / Dulongjiangxiang / 1480m, 2017.VII.30 / local collector��� (CCCC). Description. Male (Fig. 4). Body length 30.0��� 32.5 mm, humeral width 10.0��� 10.1 mm. Integument of body, antennae and legs blackish; elytra dark-brown. Head except for anterior margin of frons, genae and antennal insertions, pronotum except for sides and posterolateral margins, scutellum and most of elytra densely covered with yellowish-white pubescence. Elytron with blackish pubescent macula involving humerus, blackish pubescent macula near scutellum, and a few small irregular blackish spots near basal and apical third. Ventral sur f ace predominantly covered with fine blackish pubescence, except for anteromedian area of prosternum, most of mesanepisternum, posterior angle of metasternum, postmedian area of metanepisternum, sides of abdominal ventrite I to V, and anterolateral area of metacoxae covered with yellowish-white pubescence forming bright maculae. Body elongate, subcylindrical, feebly narrowed posteriorly. Head slightly wider than pronotal base, occiput with several umbilicate punctures anteriorly; eyes emarginate, coarsely faceted; lower eye lobe 1.5 times as long as width, 2.2 times as long as gena. Antenna long and slender, about 2.7 times body length, finely punctate, basal 3 antennomeres sparsely fringed beneath with short setae; scape gradually thickened apically, with numerous granules near outer edge (Fig. 11); 3rd antennomere 2.8 times as long as scape, 1.2 times as long as 4th, 1.5 times as long as 5th; relative length of antennomeres as follows: 3.5: 0.7: 10: 8.5: 6.9: 6.5: 6.0: 5.9: 6.7: 6.7: 11.6. Pronotum broader than long, 0.7 times as long as basal width, width across lateral spines about 1.5 times of basal width; lateral spine stout, strongly thickened at base with acute apex; disk weakly convex, provided with several setigerous granules at sides after middle. Scutellum broadly rounded posteriorly. Prosternal process widened apically; procoxal cavities widely open posteriorly. Mesosternal process without tubercle and obliquely sloped in lateral view. Elytra elongate, ca. 2.1 times as long as humeral width, slightly convergent toward conjointly rounded apices with short sutural teeth; sparsely provided with a few small granules near humeri; disk smooth, impunctate. Legs moderately long, stout; fore femora with coarse wrinkling sculpturing on both sides; metafemora reaching apical fourth of elytra; tarsus five segmented, tarsal claws divaricate. Male genitalia. Tergite VIII, tegmen and median lobe as in Fig. 17. Endophallus in everted condition (Fig. 20) robust, about 2.3 times as long as median lobe, curved near apical third dorsally; APH defined, limit between BPH and MPH indicated by basal-lateral tubercle (blt); crescent-shaped sclerite absent; MPH subdivided into MT+CT and PB by constrictions, MT slightly longer than CT or PB; CT with lateral tubercles (ltc, Fig. 20d) developed, elongate, directed forward, PB strongly swollen laterally near base, thence, in ventral view, gradually narrowed (Fig. 20d) or expanded in lateral view (Fig. 20) toward apex; APH not subdivided, moderately swollen, weakly sclerotized dorsally from base to gonopore; apical furrow (af) with internal membrane (im) incomplete, represented by short dorsal appendix; spicules mainly distributed on ventral apical surface of MT and apical half of PB; ejaculatory ducts (ej) paired, gonopore (gn) situated near apex of dorsal side of APH. Female (Fig. 5). Body length 35.0��� 35.5 mm, humeral width 11.1���11.4 mm. Almost identical to male in general appearance. Antenna about 2.0 times body. Head relatively larger, about 1.2 times wider than pronotal base. Abdo- men with ventrite V (Fig. 16) widely protruded, apex 0.24 times basal width. Legs relatively slenderer and shorter, fore femora without wrinkling sculpturing. Etymology. From the Latin, meaning similar or resembling, referring to the similarities between the new species and Ithocritus ruber. Distribution. China: Yunnan (Gongshan County). Remarks. Ithocritus ruber also occurs in the type locality of the new species. Both species share highly morphological similarities, especially regarding the yellowish-white colored individuals (Figs. 3���5), but the new species can be distinguished by the metanepisternum and metacoxae provided with bright colored maculae (instead of absent); scutellum densely pubescent, obscuring integument (instead of with pubescence relatively sparser, not obscuring integument); elytra dark-brown (instead of reddish ochraceous); female with abdominal ventrite V widely protruded, apex 0.24 times as basal width (instead of 0.13 times); male with endophallus relatively more robust, CT with lateral tubercles developed, elongate, directed forward (instead of weakly developed), PB strongly swollen laterally near base, thence, in ventral view, gradually narrowed toward apex (instead of weakly swollen laterally and ventrally near base, thence strongly constricted toward subparallel-sided apex)., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on pages 455-457, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Hope, F. W. (1839) Descriptions of some new insects collected in Assam, by William Griffith, Esq., assistant surgeon in the Madras Medical Service. Proceedings of the Linnean Society of London, 1, 42 - 44.","Breuning, S. (1956) Revision des \" Petrognathini \". Longicornia, III, 349 - 392."]}

Genus Ithocritus Lacordaire, 1872 Ithocritus Lacordaire, 1872: 448. Type species: Monochamus ruber Hope, 1839, by original designation. Ithocritus; Gemminger, 1873: 3059; Aurivillius, 1922: 206; Breuning, 1956: 354; L��bl & Smetana, 2010: 291; Lin & Jiroux, 2011: 108; Ohbayashi & Lin, 2012: 237; Lin & Yang, 2019: 340. Remarks. Including the types species, two of the three known species of the genus have been investigated in this study. Both species with granules near outer edge of scape (Fig. 11), which is inconsistent with the tribal definition proposed by Breuning (1956) and Ohbayashi & Lin (2012). Male genitalia of the two studied species has the endophallus lacking the crescent-shaped sclerites (cs), which accordant with the structure observed in Petrognatha Leach, 1819 (type genus of the tribe), Ioesse Thomson, 1864, and Pseudapriona Breuning, 1936. The consistency of this character among these studied genera superficially stabilized their tribal treatment. However, the absence of such crescent-shaped sclerites in the male endophallus have been reported in several taxa from different tribes of subfamily Lamiinae, e.g. some species of Phytoeciini (as ���basal-lateral-ventral sclerites���, Kasatkin 2006), Mimocagosima Breuning, 1968 and Thermistis Pascoe, 1867 of Saperdini (as ���basal armature��� ��� Lin, Li & Yang 2008; Lin et al. 2012), Acanista Pascoe, 1864 of Acanthocinini, Pentacosmia Newman, 1842 and Probatodes Thomson, 1864 of Rhodopinini (both genera currently in Desmiphorini), Rosenbergia Ritsema, 1881 of Batocerini (as ���crescent-shaped sclerites���, Ślipiński & Escalona 2013), and have also been observed in Pseudomeges Breuning, 1943 and Myagrus Pascoe, 1878 of Lamiini (both genera currently in Monochamini) (Bi, unpublished). The species of Apriona Chevrolat, 1852 and Batocera Dejean, 1835 (Batocerini) from China lack the crescent-shaped sclerites, while Batocera spp. from Australia were reported by Ślipiński & Escalona (2013) having ���large crescent-shaped sclerites���, and the species of Microcriodes Breuning, 1943, also Batocerini, have such sclerites. These variations undoubtedly indicate a more complicated situation, making questionable the tribal position of Ithocritus., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on page 454, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Lacordaire, J. T. (1872) Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Famille LXVIII. Longicornes. (suite). Sous-famille III. LAMIIDES. Librairie Encyclopedique de Roret, Paris, 9 (2), 411 - 930.","Hope, F. W. (1839) Descriptions of some new insects collected in Assam, by William Griffith, Esq., assistant surgeon in the Madras Medical Service. Proceedings of the Linnean Society of London, 1, 42 - 44.","Gemminger, M. (1873) Cerambycidae (Lamiini). In: Gemminger, M. & von Harold, E. (Eds.), Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom X. Cerambycidae (Lamiini), Bruchidae. E. H. Gummi, Monachii, pp. 2989 - 3216.","Aurivillius, C. (1922) Cerambycidae: Lamiinae I. Pars 73. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. XXIII. Cerambycidae II. W. Junk, Berlin, 322 pp.","Breuning, S. (1956) Revision des \" Petrognathini \". Longicornia, III, 349 - 392.","Lobl, I. & Smetana, A. (2010) Catalogue of Palaearctic Coleoptera. Vol. 6. Apollo Books, Stenstrup, 924 pp. https: // doi. org / 10.1163 / 9789004260917 _ 004","Lin, M. - Y. & Jiroux, E. (2011) Notes on the genera Pseudapriona Breuning, 1936, Ithocritus Lacordaire, 1872 and Ioesse Thomson, 1864, of the tribe Petrognathini (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, New Series, 5, 104 - 114, 33 figs.","Ohbayashi, N. & Lin, M. - Y. (2012) A Review of the Asian Genera of the Petrognathini, with Description of a New Species and Proposal of a New Synonym (Coleoptera, Cerambycidae, Lamiinae). Japanese Journal of Systematic Entomology, 18 (2), 235 - 251.","Lin, M. - Y. & Yang, X. - K. (2019) Catalogue of Chinese Coleoptera Volume IX. Chrysomeloidea: Vesperidae, Disteniidae, Cerambycidae. Science Press, Beijing. 575 pp.","Kasatkin, D. G. (2006) The internal sac of aedeagus of longhorned beetles (Coleoptera: Cerambycidae): morphology, nomenclature of structures, taxonomic significance. Caucasian Entomological Bulletin, 2 (1), 83 - 104. https: // doi. org / 10.23885 / 1814 - 3326 - 2006 - 2 - 1 - 83 - 104","Lin, M. - Y., Li, W. - Z. & Yang, X. - K. (2008) Taxonomic review of three saperdine genera, Mandibularia Pic, Mimocagosima Breuning and Parastenostola Breuning (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Zootaxa, 1773 (1), 1 - 17. https: // doi. org / 10.11646 / zootaxa. 1773.1.1","Lin, M. - Y., Chou, W. - I., Kurihara, T. & Yang, X. - K. (2012) Revision of the genus Thermistis Pascoe 1867, with descriptions of three new species (Coleoptera: Cerambycidae: Lamiinae: Saperdini). Annales de la Societe Entomologique de France, New Series, 48 (1 - 2), 29 - 50. https: // doi. org / 10.1080 / 00379271.2012.10697749","Slipinski, S. A. & Escalona, H. E. (2013) Australian Longhorn Beetles (Coleoptera: Cerambycidae). Introduction and Subfamily Lamiinae. Vol. 1. CSIRO Publishing, Collingwood, xviii + 484 pp., 221 figs. https: // doi. org / 10.1071 / 9781486300044"]}

Falsimalmus niger Breuning, 1956 (Figs. 6, 7, 18, 21) Falsimalmus niger Breuning, 1956: 359, fig. 3. Falsimalmus niger; Rondon & Breuning, 1970: 486; Ohbayashi & Lin, 2012: 250, figs. 26-29; Lin & Yang, 2014: 312. Material examined. 1 male, China, Yunnan, Yingjiang, Nabang, 473m, 2016.V.30, leg. Xiao-Dong Yang (CCCC). Complementary description. Male (Figs. 6, 7). Body length 24.0 mm, humeral width 8.2 mm. Male genitalia. Tergite VIII, tegmen and median lobe as in Fig. 18. Endophallus in everted condition (Fig. 21) moderately long and slender, about 2.7 times as long as median lobe, curved dorsally near middle; BPH, MPH and APH well-defined; crescent-shaped sclerites (cs) present; MPH subdivided into MT, CT and PB by constrictions; MT slightly shorter than CT and PB combined; MT normally cylindrical, CT swollen near base and apex respectively, PB with anterior bulb generally rudimentary; APH strongly constricted, elongate, broader near base, with apical bulb (ab) 3.8 times longer than apical bubble (bb) (Fig. 21e); apical furrow (af) with internal membrane (im) developed, elongate and sclerotized; spicules distributed on basal half of CT, apical half of PB and with distinctly larger spicules on apical bulb of APH throughout; ejaculatory ducts paired, gonopore (gn) situated at apex of APH. Distribution. China (new country record): Yunnan (Yingjiang County); Laos (Vientiane); Myanmar (Dawns, Tenasserim); Thailand (Chiang Mai)., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on page 459, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Breuning, S. (1956) Revision des \" Petrognathini \". Longicornia, III, 349 - 392.","Rondon, J. A. & Breuning, S. (1970) Lamiines du Laos. Pacific Insects Monograph, 24, 315 - 571.","Ohbayashi, N. & Lin, M. - Y. (2012) A Review of the Asian Genera of the Petrognathini, with Description of a New Species and Proposal of a New Synonym (Coleoptera, Cerambycidae, Lamiinae). Japanese Journal of Systematic Entomology, 18 (2), 235 - 251.","Lin, M. - Y. & Yang, X. - K. (2014) One new record species, Ithocritus ruber (Hope, 1839) from China (Coleoptera: Cerambycidae: Lamiinae: Petrognathini). Zoological Systematics, 39 (2), 309 - 312. https: // doi. org / 10.11865 / zs 20140214"]}

Ithocritus ruber (Hope, 1839) (Figs. 1���3, 8, 9, 12, 13, 15, 19) Monochamus ruber Hope, 1839: 43; Hope, 1840: 441, pl. 30, fig. 5. Ithocritus ruber: Lacordaire, 1872: 448; Gemminger, 1873: 3059; Aurivillius, 1922: 206; Breuning, 1956: 354, fig. 1; Mukhopadhyay & Biswas, 2000: 59; L��bl & Smetana, 2010: 291; Lin & Jiroux, 2011: 108, figs. 13-18; Ohbayashi & Lin, 2012: 238, figs. 1-5; Lin & Yang, 2014: 309, figs. 1-11; Mitra et al., 2016: 44; 2017: 86; Lin & Yang, 2019: 340. Type material examined. Syntypes of Monochamus ruber, 2 males, India, Assam, leg. William Griffith (in OUMNH). Examined by photographs taken by James Hogan. Other materials examined. China (Yunnan): 1 male, 1 female, Yunnan, Longchuan, Husa Xiang, 24.358315��N, 97.834079��E, alt. 1353m, 2013.VIII.10, leg. Zhi-Shun Song, Qiang-Feng Zheng (IZCAS); 1 male, Yunnan, Gongshan, Maku, 1200m, 2015.VI.26, leg. Xiao-Dong Yang (CCCC); 1 male, ditto except 1500m, 2015.VIII.7 (CCCC); 1 male, ditto except 1250m, 2015.VI.27, leg. Wen-Xuan Bi (CBWX); 1 female, ditto except 1500m, 2015.VII.18 (CBWX); 1 male, ditto except 1250m, 2015.VII.20 (CBWX); 1 male, ditto except 2015.VII.2 (CBWX); 1 female, ditto except 1500m, 2015.VIII.9 (CBWX); 1 female, ditto except 1500m, 2015.VII.21, leg. Chao Wu (CBWX). China (Xizang): 2 males, 1 female, Xizang, Motuo, Baricun, 1850m, 2014.VII.26, leg. Wen-Xuan Bi (CBWX); 1 male, ditto except 2014.VII.29 (CBWX); 1 female, Xizang, Linzhi City, M��dog County, Dexingxiang, Yarang power station, 29.27022��N, 95.248532��E, alt. 706m, 2019.VIII.1, leg. Hao-Dong Yin (IZCAS). Complementary description to Breuning (1956) and Lin & Jiroux (2011). Integument of body, antennae and legs blackish; elytra reddish ochraceous. Head, pronotum, elytra predominantly covered with bright pubescence which is individually variable from brick-red to pale orange or yellowish white (Figs. 1���3). Ventral surface predominantly covered with fine blackish pubescence, except for anteromedian of prosternum, most of mesanepisternum, posterior angle of metasternum and lateral sides of abdominal ventrite I to V covered with same variably colored pubescence as in dorsal surface. Scutellum moderately pubescent, not obscuring integument, giving it darkened appearance (Figs. 8, 9). Male genitalia with endophallus, in everted condition (Fig. 19), about 2.3 times as long as median lobe, curved near apical fourth dorsally; APH defined, BPH and MPH roughly defined by basal-lateral tubercle (blt); crescent-shaped sclerite absent; MPH subdivided into MT+CT and PB by constrictions, MT ca. 1.5 times longer than CT or PB; MT with ventral swelling (vs) generally rudimentary; CT with lateral tubercles (ltc, Fig. 19d) weakly developed; PB weakly swollen laterally and ventrally near base, thence, in ventral view, strongly constricted toward subparallel-sided apex (Fig. 19d); APH not subdivided, in lateral view, moderately swollen, slightly protruding ventrally; apical furrow (af) with internal membrane (im) incomplete, represented by short dorsal appendix; spicules mainly distributed on ventral apical surface of MT and apical half of PB; ejaculatory ducts (ej) paired, gonopore (gn) situated near apex of dorsal side of APH. Female with ventrite V (Fig. 15) narrowly protruded, apex 0.13 times basal width. Distribution. China: Yunnan, Xizang (new province record); India: Assam (Mukhopadhyay & Halder 2003), Meghalaya, Sikkim and West Bengal; Bangladesh (Silhet = Sylhet); Myanmar (Kachin, Mandaley), Nepal., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on pages 454-455, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Hope, F. W. (1839) Descriptions of some new insects collected in Assam, by William Griffith, Esq., assistant surgeon in the Madras Medical Service. Proceedings of the Linnean Society of London, 1, 42 - 44.","Hope, F. W. (1840) Descriptions of some new insects, collected in Assam by William Griffith, Esq., assistant-surgeon in the Madras Medical Service, and attached to the late scientific mission to Assam. The Transactions of the Linnean Society of London, 18, 435 - 447, pls. 30 - 31. https: // doi. org / 10.1111 / j. 1095 - 8339.1838. tb 00192. x","Lacordaire, J. T. (1872) Histoire Naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Famille LXVIII. Longicornes. (suite). Sous-famille III. LAMIIDES. Librairie Encyclopedique de Roret, Paris, 9 (2), 411 - 930.","Gemminger, M. (1873) Cerambycidae (Lamiini). In: Gemminger, M. & von Harold, E. (Eds.), Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom X. Cerambycidae (Lamiini), Bruchidae. E. H. Gummi, Monachii, pp. 2989 - 3216.","Aurivillius, C. (1922) Cerambycidae: Lamiinae I. Pars 73. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. XXIII. Cerambycidae II. W. Junk, Berlin, 322 pp.","Breuning, S. (1956) Revision des \" Petrognathini \". Longicornia, III, 349 - 392.","Mukhopadhyay, P. & Biswas, S. (2000) Coleoptera: Cerambycidae. Zoological Survey of India, State Fauna Series 4. Fauna of Meghalaya, 5, 41 - 67.","Lobl, I. & Smetana, A. (2010) Catalogue of Palaearctic Coleoptera. Vol. 6. Apollo Books, Stenstrup, 924 pp. https: // doi. org / 10.1163 / 9789004260917 _ 004","Lin, M. - Y. & Jiroux, E. (2011) Notes on the genera Pseudapriona Breuning, 1936, Ithocritus Lacordaire, 1872 and Ioesse Thomson, 1864, of the tribe Petrognathini (Coleoptera, Cerambycidae, Lamiinae). Les Cahiers Magellanes, New Series, 5, 104 - 114, 33 figs.","Ohbayashi, N. & Lin, M. - Y. (2012) A Review of the Asian Genera of the Petrognathini, with Description of a New Species and Proposal of a New Synonym (Coleoptera, Cerambycidae, Lamiinae). Japanese Journal of Systematic Entomology, 18 (2), 235 - 251.","Lin, M. - Y. & Yang, X. - K. (2014) One new record species, Ithocritus ruber (Hope, 1839) from China (Coleoptera: Cerambycidae: Lamiinae: Petrognathini). Zoological Systematics, 39 (2), 309 - 312. https: // doi. org / 10.11865 / zs 20140214","Mitra, B., Das, P., Chakraborti, U., Mallick, K. & Majumder, A. (2016) Longhorn beetles (Cerambycidae: Coleoptera) of Meghalaya with eight new records. The Journal of Zoology Studies, 3 (4), 39 - 47.","Mitra, B., Chakraborti, U., Mallick, K., Bhaumik, S. & Das, P. (2017) An updated list of cerambycid beetles (Coleoptera: Cerambycidae) of Assam, India. Records of the Zoological Survey of India, 117 (1), 78 - 90. https: // doi. org / 10.26515 / rzsi / v 117 / i 1 / 2017 / 117286","Lin, M. - Y. & Yang, X. - K. (2019) Catalogue of Chinese Coleoptera Volume IX. Chrysomeloidea: Vesperidae, Disteniidae, Cerambycidae. Science Press, Beijing. 575 pp."]}

Genus Falsimalmus Breuning, 1956 (Figs. 6, 7, 18, 21) Falsimalmus Breuning, 1956: 358. Type species: Falsimalmus niger Breuning, 1956, by original designation. Falsimalmus; Rondon & Breuning, 1970: 486; Ohbayashi & Lin, 2012: 249. Remarks. This genus consists of a single species and has been placed in the Petrognathini since Breuning (1956) described it. Ohbayashi & Lin (2012) mentioned that the ���head with frons is not fully vertical, but more or less inclined anteriad.��� According to our observation, the head of this genus is normally vertical (Fig. 7). In addition, the endophallic structure of the only species of this genus has been investigated for the first time and described herein. As shown in the following section, the endophallus of this genus has normally developed crescent-shaped sclerites (absent in Petrognatha gigas (Fabricius, 1793) and Ithocritus spp.); a developed, complete, elongate and sclerotized internal membrane (im) of apical furrow (incomplete and represented by a short dorsal appendix in Ithocritus spp.); a strongly constricted and elongate APH, which is associated with larger spicules (APH swollen and lacking spicules in Ithocritus spp.). These features are obviously different from the other members of this tribe studied by the first author, but partially (especially the constricted and spiculate APH with reduced apical bubble and the developed internal membrane of apical furrow) resemble some Lamiini genera, e.g. Echinovelleda Breuning, 1936 (Bi 2018: figs. 25, 26), indicating an uncertain tribal position of this genus, which is consistent with the suggestion presented by Ohbayashi & Lin (2012). A further examination based on molecular methods may confirm its systematic position., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on pages 457-459, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Breuning, S. (1956) Revision des \" Petrognathini \". Longicornia, III, 349 - 392.","Rondon, J. A. & Breuning, S. (1970) Lamiines du Laos. Pacific Insects Monograph, 24, 315 - 571.","Ohbayashi, N. & Lin, M. - Y. (2012) A Review of the Asian Genera of the Petrognathini, with Description of a New Species and Proposal of a New Synonym (Coleoptera, Cerambycidae, Lamiinae). Japanese Journal of Systematic Entomology, 18 (2), 235 - 251.","Hope, F. W. (1839) Descriptions of some new insects collected in Assam, by William Griffith, Esq., assistant surgeon in the Madras Medical Service. Proceedings of the Linnean Society of London, 1, 42 - 44.","Bi, W. - X. (2018) Studies on the Flightless Lamiinae (Coleoptera: Cerambycidae) from China: II. Echinovelleda Breuning, 1936, Paroriaethus Breuning, 1936 and Lonyarbon gen. nov. Japanese Journal of Systematic Entomology, 24 (2), 267 - 276."]}

Tribe Petrognathini Blanchard, 1845 Diagnosis (modified from Ohbayashi & Lin 2012). Antennal scape without cicatrix or granules at apex; mandible neither elongate nor longitudinally grooved; eyes emarginate; pronotum provided with lateral spines; mesocoxal cavity open to epimeron; metepisternum not distinctly widened; legs stout; mesotibiae with an external sinus; tarsal claws simple and divaricate., Published as part of Bi, Wen-Xuan, Chen, Chang-Chin & Lin, Mei-Ying, 2020, Notes on the tribe Petrognathini Blanchard, 1845 from China, with description of a new species from Yunnan (Coleoptera, Cerambycidae, Lamiinae), pp. 453-460 in Zootaxa 4732 (3) on page 454, DOI: 10.11646/zootaxa.4732.3.7, http://zenodo.org/record/3667282, {"references":["Ohbayashi, N. & Lin, M. - Y. (2012) A Review of the Asian Genera of the Petrognathini, with Description of a New Species and Proposal of a New Synonym (Coleoptera, Cerambycidae, Lamiinae). Japanese Journal of Systematic Entomology, 18 (2), 235 - 251."]}