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Modified key to Palaearctic Oiceoptoma (modified from Růžička et al. 2004) The following couplets are modified to accommodate the newly described species. 5. Pronotum with diffuse median spots (usually composed of several scattered darker spots or almost indistinct, Figs. 4B, E). Scutellum more distinctly punctured medially than laterally (Fig. 6B). Elytra brown to black, subapical surface with rugosities (Figs. 2A, D, 9C). Male genital segment with ventrite 9 anteriorly distinctly longer than tergum 9 in ventral view (Fig. 12E).��� Proventrite with dense punctation laterally; medially dark brown to black, laterally only narrowly orange in ventral view (Fig. 5B). Median lobe of aedeagus subapically distinctly sinuous laterally, tapering to a widely rounded apex (Figs. 11C���D). Basal portion oval, robust, not exceeding the width of aedeagus (Figs. 11C���D). Female tergum 7 posteriorly with wide, only weakly expressed emargination (Fig. 16A). Female tergum 8 truncate posteriorly (Fig. 13C). (Central and north-eastern China, Far East of Russia, Korean Peninsula to Japan)..................................................... O. subrufum (Lewis) ��� Pronotum with four black distinct spots, arranged in a trapezoidal pattern (Figs. 4A, C). Scutellum with homogeneous punctation (Figs. 6A, C). Elytra subapically with simple surface (Figs. 1A, D, 2E, 3A, 9A���B, D). Male genital segment with ventrite 9 anteriorly as long as tergum 9 in ventral view (Figs. 12D, F).................................................. 6 6. Spots on pronotum distinctly delimited, smooth and glossy, only with sparsely distributed, minute punctures with very short setae (Figs. 4C, F). Proventrite with only minute, superficial punctation laterally; completely dark brown to black in ventral view (Fig. 5C). Elytra black (Figs. 2E, 3A, 6F, 9D). Apex of elytron short and rounded in female (Figs. 2E). Surface of scutellum and elytra with very weak, superficial punctation (Figs. 6C, F, 9D). Median lobe of aedeagus gradually tapered to a slender, sharp apex (Fig. 11E). Basal portion oval, less robust, not exceeding the width of aedeagus (Fig. 11E). Female tergum 7 posteriorly simple, without emargination (Fig. 16C). Female tergum 8 broadly truncate posteriorly (Fig. 14A). Female ventrites 5 and 6 longer medially, posteriorly exposing only a shorter area of brick-wall pattern (Figs. 14B, 15E). (Japan)............................................................................................... O. nigropunctatum (Lewis) ��� Spots on pronotum vaguely delimited, regularly covered with fine, distinct punctures, covered by long, orange setation (Figs. 4A, D). Proventrite with dense punctation laterally; medially dark brown to black, laterally widely orange in ventral view (Fig. 5A). Elytra light to dark brown (Figs. 1A, D, 6D, 9A���B). Apex of elytron elongated to blunt tip in female (Figs. 1D, 8A, 9B). Surface of scutellum and elytra with distinct, deep punctation (Figs. 6A, D, 9A���B), Median lobe of aedeagus gradually tapered to widely rounded, triangular apex (Figs. 11A���B). Basal portion widely oval, very narrow, markedly exceeding the width of aedeagus (Figs. 11A���B). Female tergum 7 posteriorly with narrow, deep emargination (Fig. 17A). Female tergum 8 regularly rounded posteriorly (Figs. 13A, 17A). Female ventrites 5 and 6 very short medially, posteriorly with long area of brick-wall pattern (Figs. 15A���B, F) (China: Zhejiang Province)..... O. tangi Sommer, Růžička, Nishikawa & Schneider, new species, Published as part of Sommer, David, R����i��ka, Jan, Nishikawa, Masaaki & Schneider, Jan, 2021, Oiceoptoma tangi (Coleoptera: Silphidae: Silphinae), a new species of carrion beetle from eastern China, pp. 445-472 in Zootaxa 4949 (3) on pages 470-471, DOI: 10.11646/zootaxa.4949.3.2, http://zenodo.org/record/4640397, {"references":["Ruzicka, J., Hava, J. & Schneider, J. (2004) Revision of Palaearctic and Oriental Oiceoptoma (Coleoptera: Silphidae). Acta Societatis Zoologicae Bohemicae, 68, 30 - 51."]}

A new species, Oiceoptoma tangi Sommer, Růžička, Nishikawa & Schneider, new species, from Zhejiang Province, China, is described and illustrated. The new species is closely related to O. subrufum (Lewis, 1888), distributed in Central and North-eastern China, the Korean Peninsula, Far East of Russia and Japan, and to O. nigropunctatum (Lewis, 1888), endemic to Japan. All three species are keyed, and distribution maps of all three species are presented.

Oiceoptoma nigropunctatum (Lewis, 1888) (Figs. 2E, 3A���C, 4C, F, 5C, F, 6C, F, 7C, 9D, 11E, 14F) Material examined. Japan, Kyushu, 1 ♂ (JRUC), Oita prefecture, Kuju mts, Mt. Kurodake [ca. 33��19���N 131��13���E], Sh��nai-machi, 19.v.1997, F. Hirokawa leg., S. Nomura det.; 1 ♂ (JRUC), Tokushima prefecture, Kisawa-son, Dosut��ge, Naka-gun, 18.vii.���11.x.2004, M. Yoshida leg., S. Nomura det.; 1 ♀ (JRUC), Oita prefecture, Kuju mts, Mt. Kurodake [ca. 33��19���N 131��13���E], 16.viii.1992, Ryo Noda leg., S. Nomura det., Published as part of Sommer, David, R����i��ka, Jan, Nishikawa, Masaaki & Schneider, Jan, 2021, Oiceoptoma tangi (Coleoptera: Silphidae: Silphinae), a new species of carrion beetle from eastern China, pp. 445-472 in Zootaxa 4949 (3) on page 446, DOI: 10.11646/zootaxa.4949.3.2, http://zenodo.org/record/4640397, {"references":["Lewis, G. (1888) Notes on the Japanese species of Silpha. The Entomologist, an Illustrated Journal of General Entomology, 21, 7 - 10."]}

Oiceoptoma tangi Sommer, Růžička, Nishikawa & Schneider, new species (Figs. 1A���F, 3D���F, 4A, D, 5A, D, 6A, D, 7A���C, 8A, C���D, 9A���B, 10A���D, 11A���B, 12A, D, 13A���B, 14 C���E, 15A���B, F, 17A���C) Oiceoptoma subrufum: Růžička et al. 2004: 47 (misidentification). Type locality. China, Zhejiang Province, Anji City, Longwangshan Mt., Pinxi Creek, 30��24���N, 119��27���E, alt. 1000��� 1100 m. Type material (30 specimens). Holotype, ♂ (SNUC), ��� China: Zhejiang Prov. | Anji City | Longwangshan Mt. | Pinxi Creek | 30��24���N, 119��27���E || alt. 1000~ 1100 m | 09-VI-2012 | HU & YIN leg. [p]���. Paratypes, 6 ♂♂ 3 ♀♀ (SNUC), 1 ♂ 1 ♀ (BMNH), 2 ♂♂ 2 ♀♀ (JRUC), same data as holotype ���; 2 ♂♂ 3 ♀♀ (SNUC), ��� China, Zhejiang Prov. | Anji County | Shenxi | 30�� 25��� N, 119�� 24��� E | alt. 400 m, 07-VI-2012 | ZHU Jian-Qing leg. [p]���; 1 ♀ (EUMJ), ��������� [= China] ��������� [= Zhejiang Prov.] Oi������ [= Lin���an-shi] | MRH��� [= Mt. West Tianmu Shan] | 6~ 11���V���1998 | ��� ����� [= Li-Zhen Li] IJDz [= leg.] [p]���; 1 ♀ (SNUC), ���16~20���V���20 [p] 06 [hw] | MRH��� [= Mt. West Tianmu Shan] ������ [= Lin���an-shi] [hw] | ������ [= Zhejiang Prov.] Bwx [hw] || Mt. w. Tianmu | 16~20. V, 2006 | Bi Wen-xuan leg. [hw]���; 1 ♂ (SNUC), ��� West Tianmushan | Lin���an County | 11~ 15���VI���2006 | alt. 300���400 m | HU & TANG leg. [p]���; 1 ♀ (SNUC), ��� China: Zejiang Prov. | Lin���an City | Mt. West Tianmu | alt. 300~ 500 m | 29���V~ 3���VI���2010 | Jian-Qing ZHU leg. [p]���; 1 ♂ (JRUC, DNA voucher JRUC-SIL-0054), ��� CHINA: Zhejiang Prov., | Lin���an County, West | Tianmushan Nat. Res., | 250 m NW of Original | Temple of Lion Sect, | 30.3431��N 119.4316��E, | J. H��jek & J. Růžička leg. || (WT10) 2.���6.vii.2017; || 1165 m, baited pitfall | traps #06 (fish meat, | ripening cheese), broad- | leaved deciduous forest | with Rhododendron [p]���; 2 ♀♀ (JRUC, DNA vouchers JRUC-SIL- 0010, JRUC-SIL-0011), ��� CHINA: Zhejiang Prov., | Lin���an County, West | Tianmushan Nat. Res., | 250 m NW of Original | Temple of Lion Sect, | 30.3431��N 119.4316��E, | Cheng-Bin Wang leg. || (WT10) 2.���6.vii.2017; || 1165 m, baited pitfall | traps #06 (fish meat; alcohol, vinegar + sugar), broad- | leaved deciduous forest | with Rhododendron [p]���; 2 ♂♂ (JSCC), ���NW. CHINA | TIENMUCHAN [without date and collector���s name, hw with a pencil] || Oiceoptoma | subrufum | (Lewis, 1888) ♂ | Jan Růžička det. 200[p]3[hw]���. Description. Male (Figs. 1A���C). Body oval, dorso-ventrally flattened. Pronotum orange with four black spots (Fig. 4A). Elytra light to dark brown. Head, antennae, legs and ventral part of body black. Flight wings fully developed, functional (Fig. 8A). Head. Black, lustrous surface with regular punctation (Fig. 3D). Covered with short, recumbent orange setation. Clypeus anteriorly widely notched (Fig. 8D). Anterior margin of clypeus with slightly irregular row of dense, long, orange setation in dorsal view. Eye kidney-shaped in lateral view (Fig. 17C). Head with short row of long erect setae behind eyes (Fig. 3D). Frons with marked dorsal tentorial pits and a transverse, elevated crest posteriorly (Figs. 3D, 17C). Maxilla with deeply punctuated cardo and base of stipes in ventral view, covered by long setae (Fig. 3E). Antennae. Medium-sized, with last four antennomeres forming a distinct club (Figs. 7A���C). Pronotum moderately transverse, widest posteriorly (Fig. 4A). Pronotal disc orange, medially with four black, vaguely delimited spots, arranged in symmetrical, trapezoidal pattern (Fig. 4A). Spots are regularly covered with fine, distinct punctures, bearing long, orange setation (Fig. 4D). Margin rimmed only in anterior half; anteriorly with wide, shallow excision, which is medially straight; anterior angles weakly elevated. Posterior margin slightly sinuous, weakly sinuate postero-laterally (Figs. 4A, 6A). Surface with dense punctation, punctures medium-sized, clearly separated, from disc to margin deeply impressed (Figs. 4A, 5D). Pronotum covered with longer, orange setation. Scutellum. Irregularly triangular, widely vaulted medially. Surface with dense, distinct, homogenous punctation; covered with longer orange setation, semierect anteriorly and distinctly recumbent medially and posteriorly (Fig. 6A). Elytra subparallel, at the widest point (along 2/3 of its length) wider than pronotum (Fig. 1A). Humerus with distinct denticle (Fig. 3F). Elytron with three distinctly elevated ridges; only first and second ridge reaching apex of elytron; external ridge shortest, reaching only two thirds of elytral length (Figs. 1A, 6D, 9A). Second ridge in last third slightly sinuous (Fig. 9A). Third ridge posteriorly terminated at apical callus (Fig. 9A). Ridges on both sides covered with a row of short, recumbent, black setae. Lateral margin of elytra strongly elevated only in anterior half, disappearing posteriorly. Apex of elytron regularly rounded (Figs. 1A, 9A). Basal part of elytra and humeri with short, recumbent, black setae (Fig. 3F). Surface dull, with isodiametric microsculpture (Figs. 6A, D); subapical surface regularly flat, without rugosities; with dense, distinct, regular punctation; punctures clearly separated by spaces 1.0���1.5 times their own diameter (Figs. 6D, 9A). Elytra with distinct, subrectangular punctures, with bluish metallic lustre covering most of the surface in ventral view (Figs. 8A, C). Elytral epipleura sparsely, finely but distinctly punctured in ventral view (Fig. 8A). Ventrum finely punctate, covered with short, recumbent, brown to black setation (Figs. 5A, 10C). Proventrite with dense punctation laterally; medially dark brown to black, laterally widely orange in ventral view (Fig. 5A). Mesocoxae widely separated (Fig. 10C). Abdominal ventrites with distinct impressions laterally in ventral view (Figs. 1B, 15A). Ventrite 6 extremely reduced in length medially, with expanded brick-wall pattern on intersegmental membrane (Figs. 15A���B, F). Legs. Protarsomeres 1���4 only slightly expanded, covered with short, dense, yellow setation (Figs. 14C���D). Pro-, / meso- and metatibia with two apical spurs of different length (Figs. 14C���D). Metatibia almost straight, only very slightly curved in ventral view (Fig. 10D). Abdominal segments. Tergite 10 relatively short in length (Fig. 12A). Ventrite 9 oval, elongate, anteriorly as long as tergite 9, constricted and deeply desclerotized medially in ventral view (Fig. 12D). Spiculum gastrale robust, elongate in ventral view (Fig. 12D). Aedeagus. Median lobe stout, robust, gradually tapered to widely rounded, triangular apex (Figs. 11A���B). Internal sac heavily sclerotized, medial portion composed of a great number of very small, regular, rounded denticles; basal part containing two strips of elongate, slender denticles (Fig. 11B). Parameres robust, almost straight, longer than median lobe, subapically constricted with rounded apex (Figs. 11A���B). Basal portion widely oval, very narrow, markedly exceeding the width of aedeagus (Figs. 11A���B). Sexual dimorphism. Female. Similar to male except for the following structures: Protarsus simple, basal protarsomere elongated (Fig. 14E). Apex of elytron elongated to a blunt tip (Figs. 1D���E, 8A, 9B). Tergum 7 with distinct, deep, narrow, U-shaped medial emargination (Fig. 17A). Tergum 8 subquadrate, regularly rounded posteriorly (Figs. 13A, 17A). Ventrite 8 widely transverse (Figs. 13B, 17B). Tergite 9 elongate, rounded; tergite 10 pentagonal, apex posteriorly regularly rounded with dense microsetae (Figs. 10A���B, 13A). Coxite laterally constricted on outer margin subapically in ventral view; stylus inserted lateroapically, its length slightly shorter than apex of coxite (Figs. 10A���B, 13A���B). Measurements. Elytra length 5.81���6.82 mm in males (mean 6.30 mm; 6.27 mm in the holotype), 6.72���7.26 mm (mean 6.96 mm) in females; pronotum length 3.41���3.82 mm in males (mean 3.63 mm; 3.55 mm in the holotype), 3.50���3.81 mm (mean 3.62 mm) in females; pronotum width 5.62���6.66 mm in males (mean 6.07 mm; 5.92 mm in the holotype), 5.63���6.42 mm (mean 6.10 mm) in females. Variability. Elytra mostly dark brown, light brown in some (probably subteneral) specimens. Spots on pronotum slightly differently delimited, being less separated from surrounding surface in some specimens. Differential diagnosis. Oiceoptoma tangi Sommer, Růžička, Nishikawa & Schneider, new species, is similar and probably closely related to O. nigropunctatum and O. subrufum. All three species share the following combination of characters: (1) orange pronotum with relatively short pubescence; (2) four dark spots on pronotal disc; (3) dark brown/black elytra with narrow epipleura; and (4) absence of patchy velvet pattern on elytral dorsal surface (see Růžička et al. (2004) for more details). The three species can be separated from each other using the character states listed in the modified key and in Table 1 below. Etymology. Patronymic, named after Liang Tang (Shanghai, China), an enthusiastic specialist of staphylinoid beetles (Agyrtidae, Staphylinidae: Steninae, Scaphidiinae, Staphylininae, Omaliinae etc.), who first recognized this species was unnamed. Collecting circumstances. The large type series was collected near Pinxi Creek on old large mammal carrion, together with numerous Necrodes littoralis (Linnaeus, 1758) (Silphidae), Creophilus maxillosus (Linnaeus, 1758) (Staphylinidae) and some clown beetles (Histeridae) (Figs. 18A���C). Three specimens were also collected on West Tianmushan, 250 m above the Original Temple of Lion Sect, using fish-baited pitfall traps in a broadleaved deciduous forest with Rhododendron (Figs. 19A���B). The specimens were collected between May and the beginning of July. ...Continued on the next page Distribution. So far known only from three localities (Longwangshan Mt., Shenxi, Tianmu Shan Mts.) in Zhejiang Province, China (Figs. 20���21)., Published as part of Sommer, David, R����i��ka, Jan, Nishikawa, Masaaki & Schneider, Jan, 2021, Oiceoptoma tangi (Coleoptera: Silphidae: Silphinae), a new species of carrion beetle from eastern China, pp. 445-472 in Zootaxa 4949 (3) on pages 466-469, DOI: 10.11646/zootaxa.4949.3.2, http://zenodo.org/record/4640397, {"references":["Ruzicka, J., Hava, J. & Schneider, J. (2004) Revision of Palaearctic and Oriental Oiceoptoma (Coleoptera: Silphidae). Acta Societatis Zoologicae Bohemicae, 68, 30 - 51.","Lewis, G. (1888) Notes on the Japanese species of Silpha. The Entomologist, an Illustrated Journal of General Entomology, 21, 7 - 10.","Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae, secundum Classes, Ordines, Genera, Species, cum Characteri- bus, Differentiis, Synonymis, Locis. Tomus I. Editio Decima, Reformata. L. Salvii, Holmiae, (4) + 823 + (1) pp. https: // doi. org / 10.5962 / bhl. title. 542"]}

Oiceoptoma subrufum (Lewis, 1888) (Figs. 2A���D, 4B, E, 5B, E, 6B, E, 7D, 8B, 9C, 11C���D, 12B, E, 13C���D, 14G, H, 15C���D) Material examined. Japan, Honshu, 1 ♂ 1 ♀ (JRUC), Yamanashi prefecture, Makioka-machi, Otometoge pass [ca. 35��16���N 138��59���E], 1450 m, 8.viii.1996, S. Nomura leg. et det. China, Gansu Province, 1 ♂ 1 ♀ (JRUC), Lazikou valley, 34��09.9���N 103��51.9���E, 2120 m, 28.vi.2005, J. H��jek, D. Kr��l & J. Růžička leg., Published as part of Sommer, David, R����i��ka, Jan, Nishikawa, Masaaki & Schneider, Jan, 2021, Oiceoptoma tangi (Coleoptera: Silphidae: Silphinae), a new species of carrion beetle from eastern China, pp. 445-472 in Zootaxa 4949 (3) on page 446, DOI: 10.11646/zootaxa.4949.3.2, http://zenodo.org/record/4640397, {"references":["Lewis, G. (1888) Notes on the Japanese species of Silpha. The Entomologist, an Illustrated Journal of General Entomology, 21, 7 - 10."]}

Oiceoptoma thoracicum (Linnaeus, 1758) (Figs 21, 40) Distribution in Turkey. Trabzon, Rize (Háva et al. 1998). Remarks. Widely distributed Transpalaearctic species, known from Europe to Japan (Růžička 2015). In central Europe, abundant in forested habitats (e.g., Růžička 1994). In Turkey, recorded from two provinces (Trabzon, Rize) of the Black Sea Region, located in the northeast of Turkey, dominated by humid climatic characteristics with high precipitation (Fig. 40). No recently collected specimens from Turkey were available., Published as part of Çiftçi, Derya, Růžička, Jan, Hasbenli, Abdullah & Şahin, Ülfet, 2018, The large carrion beetles (Coleoptera: Silphidae) of Turkey: a review with a new species record, pp. 555-591 in Zootaxa 4441 (3) on page 577, DOI: 10.11646/zootaxa.4441.3.10, http://zenodo.org/record/1301572, {"references":["Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tomus I. Laurentii Salvii, Holmiae, [4] + 824 + [1] pp. [in Latin]","Hava, J., Ruzicka, J. & Schneider, J. (1998) Faunistic records of Silphidae (Coleoptera) from Turkey. Klapalekiana, 34, 173 - 181.","Ruzicka, J. (1994) Seasonal activity and habitat associations of Silphidae and Leiodidae: Cholevinae (Coleoptera) in central Bohemia. Acta Societatis Zoologicae Bohemicae, 58, 67 - 78."]}

Oiceoptoma inaequale (Fabricius, 1781) Figs. 6, 7, 20, 33, 46 Silpha inaequalis Fabricius, 1781: 87. Diagnosis. Body length 13���16 mm, broadly oval to subparallel, entirely black. Head with short row of long, erect hairs behind each eye. Pronotum broadest at base, disc with short black hairs. Elytral apices conjointly rounded; humerus with tooth; disc tricostate, intervals distally with weak rugulose sculpturing; epipleuron wide in posterior half with upper oblique portion twice width of lower vertical portion, coloration not differing from remainder of body. Hind femur of males unmodified. Range. Quebec south to Florida, west to Ontario, South Dakota and Texas (Anderson & Peck 1985, Peck & Kaulbars 1987). Texas distribution. See Fig. 46. This species is confined to the Texan and Austroriparian biotic provinces of Texas. It occurs in the following Texas vegetational areas: pineywoods, gulf prairies and marshes, post oak savannah, blackland prairies, and cross timbers and prairies. Confirmed counties (30): Brazos, Cherokee, Cooke, Dallas, Delta, Denton, Ellis, Falls, Fannin, Grimes, Harris, Harrison, Houston, Hunt, Jackson, Lamar, Leon, Madison, McLennan, Montague, Nacogdoches, Newton, Parker, Polk, Robertson, Sabine, San Jacinto, Smith, Tyler, Walker, Wood. Collections: BRC, EGRC, JAC, PLM, SFAC, SHSU, TAMU, TORC, TTU, UTIC, WTAM. Seasonality in Texas. See Fig. 33. The adult seasonality profile of this species (based on 117 occurrence records: Appendix I) is unimodal, with a large peak in spring. Biological Notes. Anderson and Peck (1985) report that adults are mainly diurnal and appear to be restricted to deciduous forest habitats. In Kansas, this species prefers wooded areas but is often present in open fields (Lingafelter 1995). Data from examined labels. Collecting methods: hanging sugarbait trap, flight intercept trap, UV light, Lindgren funnel trap, pit-fall trap, hanging carrion trap, elevated flight intercept trap, Malaise trap, baited pit-fall trap. Habitat records: grassland [Malaise], beech���magnolia forest [pit-fall trap and Lindgren funnel trap], upland forest [flight intercept trap], bottomland forest [hanging sugarbait trap], in leaf litter, beating Quercus marylandica, and in rotting logs. Carrion records: snake/squirrel [hanging carrion trap], deer, opossum, dog, tiger [dead zoo specimen], pig, cat, shad, and cow. This species is present in the eastern forested biotic provinces of Texas, with a few records in the open prairie vegetational areas in the northeastern area of the state. Anderson (1982), Schubeck (1983), Cole (1942) and Watson and Carlton (2005) provide details on the natural history of this species., Published as part of Mullins, Patricia L., Riley, Edward G. & Oswald, John D., 2013, Identification, distribution, and adult phenology of the carrion beetles (Coleoptera: Silphidae) of Texas, pp. 221-251 in Zootaxa 3666 (2) on page 226, DOI: 10.11646/zootaxa.3666.2.7, http://zenodo.org/record/220054

Oiceoptoma rugulosum Portevin, 1903 Figs. 8, 22, 35, 48 Silpha inaequalis var. rugulosa Portevin, 1903: 333. Diagnosis. Body length 13���16 mm, broadly oval to subparallel, entirely black. Head with small eyes, short row of long erect hairs behind each eye. Pronotum broadest at base, disc with short black hairs. Elytral apices conjointly rounded; humerus with tooth; disc tricostate, intervals distally with rugulose sculpturing; epipleuron narrow in posterior half with upper horizontal portion nearly equal in width to lower vertical portion, coloration not differing from remainder of body. Hind femur of males unmodified. Range. North Carolina south to Florida, west to Texas (Peck & Kaulbars 1987). Texas distribution. See Fig. 48. This species is widespread in the Texan and Austroriparian biotic provinces of Texas, with scattered records from the Balconian province and two records from the Kansan province (Lubbock County in the Texas Panhandle). It occurs in the following Texas vegetational areas: pineywoods, gulf prairies and marshes, post oak savannah, blackland prairies, cross timbers and prairies, south Texas plains, and Edwards Plateau. Confirmed counties (29): Anderson, Bell, Bexar, Blanco, Brazos, Comal, Dallas, Denton, Galveston, Gillespie, Houston, Jackson, Jefferson, Kerr, Lamar, Lee, Limestone, Lubbock, Nacogdoches, Navarro, Polk, Refugio, Robertson, San Jacinto, Travis, Tyler, Uvalde, Victoria, Walker. Collections: EGRC, PLM, TAMU, SHSU, SFAC, SRSU, TTU, UTIC. Seasonality in Texas. See Fig. 35. Adults of this species have been collected in Texas in almost every month of the year. The adult seasonality profile of this species (based on 89 occurrence records: Appendix I) is bimodal, with large peaks in the spring and fall. Biological Notes. Peck and Kaulbars (1987) state that this species is active in the cooler months of the year and is found in both forested and open habitats. Data from examined labels. Collecting methods: UV light trap, pit-fall traps, dung beetle traps, hanging carrion trap, pit fall trap. Habitat records: sandy meadows [pit-fall trap], under bark and in dog food. Carrion records: snake [hanging carrion trap], deer, opossum, rabbit, cow, beaver, cat and snake. This species is found mostly in the eastern forested regions of Texas, with a few records from the plains regions of Texas. The Texas seasonality profile for this species shows a distinctly bimodal distribution with very few occurrence records from May to September, supporting the statement by Peck and Kaulbars (1987) that the species is active in cooler months. Life history data for this species are lacking., Published as part of Mullins, Patricia L., Riley, Edward G. & Oswald, John D., 2013, Identification, distribution, and adult phenology of the carrion beetles (Coleoptera: Silphidae) of Texas, pp. 221-251 in Zootaxa 3666 (2) on pages 228-229, DOI: 10.11646/zootaxa.3666.2.7, http://zenodo.org/record/220054

Oiceoptoma noveboracense (Forster, 1771) Figs. 21 34, 47 Silpha noveboracensis Forster, 1771: 17 (see Peck and Miller (1993) for synonymy). Diagnosis. Body length 13���16 mm, broadly oval to subparallel, brown to black, pronotum with red-orange margins. Head with small eyes, short row of long erect hairs behind each eye. Pronotum broadest at base, disc with short golden hairs. Elytral apices conjointly rounded; humerus with tooth; disc tricostate, intervals distally with weak rugulose sculpturing; epipleuron wide in posterior half with upper oblique portion twice width of lower vertical portion, coloration not differing from remainder of body. Hind femur of males unmodified. Range. Nova Scotia south to Mississippi, west to Alberta, Montana and Texas (Anderson & Peck 1985, Peck & Kaulbars 1987). Texas distribution. See Fig. 47. This species is confined to the northeastern areas of the Texan and Austroriparian biotic provinces of Texas. It occurs in the following Texas vegetational areas: pineywoods and post oak savannah. Confirmed counties (2): Lamar, Wood. Collections: SFAC. Seasonality in Texas. See Fig. 34. The adult seasonality profile of this species (based on 7 occurrence records: Appendix I) is unimodal, with a peak in early spring, but the available Texas data are limited. Biological Notes. Anderson and Peck (1985) report that adults are primarily diurnal and are most commonly found in forested habitats. Lingafelter (1995) also states that this species is mainly a woodland species. Data from examined labels. Collecting methods: Malaise trap, pit-fall trap. Habitat records: tall grass [Malaise], old growth hardwood forest [Malaise], pine and oak [pit-fall traps], flowers. There are only seven occurrence records of this species in Texas. This species is restricted to the northeastern forested areas of the state. Cole (1942), Schubeck (1969, 1983), and Anderson (1982) discussed the natural history of this species., Published as part of Mullins, Patricia L., Riley, Edward G. & Oswald, John D., 2013, Identification, distribution, and adult phenology of the carrion beetles (Coleoptera: Silphidae) of Texas, pp. 221-251 in Zootaxa 3666 (2) on pages 227-228, DOI: 10.11646/zootaxa.3666.2.7, http://zenodo.org/record/220054

A taxonomic revision of the subgenus Chrysosilpha Portevin, 1921 (of the genus Necrophila Kirby & Spence, 1828) ispresented. Three valid species are recognized: (1) N. (C.) formosa (Laporte, 1832), comb. nov. (ex Silpha Linnaeus, 1758),widely distributed from Laos and Vietnam through the Malay Peninsula, to Sumatra and Bali, with Silpha chloroptera Laporte, 1840 and Chrysosilpha chloroptera var. magnifica Portevin, 1921 as newly established junior subjectivesynonyms; (2) N. (C.) renatae (Portevin, 1920), comb. nov. (ex Silpha), endemic to Sulawesi; and (3) N. (C.) viridis (Motschulsky, 1861), comb. nov. (ex Oiceoptoma Leach, 1815), endemic to the Philippines. Lectotypes are designated for Silpha formosa Laporte, 1832; Silpha chloroptera Laporte, 1840 and Chrysosilpha chloroptera var. magnifica Portevin,1921. Georeferenced records for all three species are mapped. Parsimony analysis supports the monophyly of Chrysosilpha, with N. (C.) formosa as the sister to a clade of N. (C.) renatae and N. (C.) viridis. Geometricalmorphometrics (thin-plate spline) discriminated the three species of Chrysosilpha; the first two relative warp axesindicated 70.31% shape variation in males and 77.18% in females, which was further confirmed by MANOVA to be highlysignificant. Canonical variate analysis indicated no overlap between the three taxa and enabled a 100% correct classification of each specimen to its group mean.