1. Tiphia Fabricius 1775
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Han, Qian, Wang, He-Sheng, Chen, Bin, and Li, Ting-Jing
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Insecta ,Arthropoda ,Tiphia ,Animalia ,Biodiversity ,Tiphiidae ,Hymenoptera ,Taxonomy - Abstract
Subgenus Tiphia Fabricius, 1775 Tiphia Fabricius, 1775: 354; Allen 1975: 5; Tsuneki 1985: 3, 10; Tsuneki 1986a: 5, 9; Boni Bartalucci 2011: 342; Belokobylskij & Lelej 2017: 150. Type species Tiphia femorata Fabricius, 1775. Diagnosis. Lateral surface of propodeum dorsally with oblique wrinkles (Figs 58, 76, 92, 99); T1 anteriorly without a transverse row of short longitudinal striae, without subposterior margin, or not extending and covering subapical puncture groove in lateral view (Figs 14, 16, 40, 49, 67) (Tsuneki 1985, 1986a; Han et al. 2021). Key to the Chinese species of the subgenus Tiphia Fabricius, 1775 ♀♀ 1. Inner face of hind basitarsus medially without longitudinal groove (Figs 4, 107, 139)................................ 2 - Inner face of hind basitarsus medially with longitudinal groove (Fig. 119)....................................... 19 2. Lateral carina and medial longitudinal carina of propodeal areola with duplicate carina (Fig. 6 in Gussakovskij, 1938)............................................................................. T. (T.) duplicata Gussakovskij, 1938 - Lateral carina and medial longitudinal carina of propodeal areola without duplicate carina........................... 3 3. Propodeal dorsal face laterally with one short, transverse carina outside of areola...................................................................................................... T. (T.) brevicarinata Allen & Jaynes, 1930 - Propodeal dorsal face laterally without short, transverse carina outside of areola................................... 4 4. Tegula more than 1.5× as long as middle width.............................................................. 5 - Tegula (Figs 1, 37) atmost slightly longer than middle width................................................... 7 5. Propodeal areola with one additional longitudinal carina between medial longitudinal carina and lateral carina (Fig. 2C in Chen, 1995)........................................................................... T. (T.) mutata Chen, 1995 - Propodeal areola without one additional longitudinal carina between medial longitudinal carina and lateral carina......... 6 6. Lateral groove of S1 longer than posterior half, reaching the anterior apex...................... T. (T.) yangi Chen, 1995 - Lateral groove of S1 shorter than posterior half....................................... T. (T.) yushana Tsuneki, 1986 7. Propodeal areola with one additional longitudinal carina between medial longitudinal and lateral carinae (Plate 2, Fig. 12 in Allen & Jaynes, 1930)........................................................... T. (T.) lyrata Magretti, 1892 - Propodeal areola without one additional longitudinal carina between medial longitudinal and lateral carinae, only with medial longitudinal carina.................................................................................... 8 8. Posterior margin of tegula with transverse impressed line (Fig. 1H in Han & Kim, 2009)............................. 9 - Posterior margin of tegula without transverse impressed line (Fig. 37)........................................... 12 9. Apex of clypeus medially truncated (Fig. 159 in Tsuneki, 1985).................... T. (T.) yonaguniensis Tsuneki, 1985 - Apex of clypeus medially emarginated................................................................... 10 10. Subposterior area of T1 with 2–3 transverse rows of punctures (Fig. 40 in Tsuneki, 1986a).... T. (T.) ilanensis Tsuneki, 1986 - Subposterior area of T1 with one transverse row of punctures................................................. 11 11. Vertex, pronotum and mesoscutum with smooth interspaces between punctures.............................................................................................. T. (T.) rufomandibulata rufomandibulata Smith, 1873 - Vertex, pronotum and mesoscutum with coriaceous interspaces between punctures........ T. (T.) r. taipeiana Tsuneki, 1986 12. Mesoscutum with anterior medial groove connecting notaulus................................................. 13 - Mesoscutum with anterior medial groove separated from notaulus (Figs 37, 60)................................... 14 13. Mid and hind femora bright red; pygidium anteriorly with dense punctures................... T. (T.) scutensis Allen, 1975 - Mid and hind femora black; pygidium anteriorly with a few scattered punctures.... T. (T.) totopunctata Allen & Jaynes, 1930 14. Pronotum latero-ventrally with distinct groove in middle (Figs 106, 140)........................................ 15 - Pronotum latero-ventrally without distinct groove in middle...................... T. (T.) cilicincta Allen & Jaynes, 1930 15. Apex of clypeus medially truncated or rounded (Figs 2, 104).................................................. 16 - Apex of clypeus medially emarginated (Figs 61, 137)........................................................ 18 16. S1 anteriorly with medial longitudinal carina............................. T. (T.) sternocarinata Allen & Jaynes, 1930 - S1 (Figs 6, 109) anteriorly without medial longitudinal carina................................................. 17 17. Wings completely transparent (Fig. 1); anterior half of pygidium (Fig. 7) with sparse small punctures mixed with some big ones............................................................ T. (T.) flavobrunnea Han, Chen & Li, sp. nov. - Wings infuscate (Fig. 103); anterior half of pygidium (Fig. 110) with dense and continuous punctures................................................................................................ T. (T.) lawrencei Allen, 1975 18. Mid and hind femora (Fig. 136) bright red; propodeal areola (Fig. 142) irregularly pitted along inner and outer margins of both lateral and medial longitudinal carinae............................................ T. (T.) weismani Krombein, 1982 - Mid and hind femora (Fig. 60) black; propodeal areola (Fig. 66) normal, not pitted along inner and outer margins of both lateral and medial longitudinal carinae............................................... T. (T.) wuyiana Chen & Yang, 2003 19. Propodeal areola with one additional longitudinal carina between medial longitudinal carina and lateral carina.......... 20 - Propodeal areola without one additional longitudinal carina between medial longitudinal carina and lateral carina, only with medial longitudinal carina............................................................................. 21 20. Propodeal dorsal face laterally with one obliquely longitudinal carina outside of areola....... T. (T.) compressa Smith, 1855 - Propodeal dorsal face laterally without longitudinal carina outside of areola............... T. (T.) wushensis Tsuneki, 1986 21. Apex of clypeus medially triangular (Fig. 62 in Tsuneki, 1986a)....................... T. (T.) triangulata Tsuneki, 1986 - Apex of clypeus medially truncated or emarginated (Figs 9, 82, 127)........................................... 22 22. Propodeal dorsal face laterally (Fig. 131) with regularly strong sculpture outside of areola................................................................................................... T. (T.) toreuta Boni Bartalucci, 2010 - Propodeal dorsal face laterally without regularly strong sculpture outside of areola................................ 23 23. Tegula (Fig. 13) more than 1.5× as long as middle width..................................................... 24 - Tegula (Figs 16, 25) atmost slightly longer than middle width................................................. 27 24. Subposterior area of T1 close to middle, and (Fig. 14) medially with one transverse row of short longitudinal striae......................................................................... T. (T.) longistria Han, Chen & Li, sp. nov. - Subposterior area of T1 close to apex, and without one transverse row of short longitudinal striae..................... 25 25. Frons with medial longitudinal impunctate area............................ T. (T.) longitegulata Allen & Jaynes, 1930 - Frons without medial longitudinal impunctate area.......................................................... 26 26. Apex of clypeus medially truncated; T1 slightly longer than middle width, semicircular shape.................................................................................................... T. (T.) alishana Ishikawa, 1967 - Apex of clypeus medially slightly emarginated; T1 distinctly less than 0.5× as long as middle width, subrectangle shape (Fig. 49 in Tsuneki, 1986a)........................................................ T. (T.) tsukengensis Tsuneki, 1986 27. Pygidium impunctate, only with dense long striae reaching posterior apex....................................... 28 - Pygidium anteriorly and medially with dense or sparse punctures, posteriorly smooth or coriaceous (Figs 20, 27, 44, 88, 124).................................................................................................. 31 28. Lower frons with dense punctures....................................................................... 29 - Lower frons with sparse punctures....................................................................... 30 29. Pronotal dorsal face medially with one transverse row of big and deep punctures..... T. (T.) ordinaria ordinaria Smith, 1873 - Pronotal dorsal face medially with one transverse row of small and shallow punctures........ T. (T.) o. atayal Tsuneki, 1986 30. S6 posteriorly with medial short longitudinal carina................ T. (T.) brevilineata brevilineata Allen & Jaynes, 1930 - S6 posteriorly without medial short longitudinal carina............................ T. (T.) b. formosana Tsuneki, 1986 31. Mandible with distinct medial transverse groove (Fig. 1K in Han & Kim 2009).......... T. (T.) popilliavora Rohwer, 1924 - Mandible (Figs 17, 23, 41, 82, 118) without distinct medial transverse groove, atmost with obsolete intermittent impressed line............................................................................................... 32 32. Posterior half of pronotum medially with short, shallow and longitudinal line (Fig. 1L in Han & Kim 2009)........................................................................................... T. (T.) vernalis Rohwer, 1924 - Posterior half of pronotum medially without short, shallow and longitudinal line.................................. 33 33. Each of tibiae with two spurs apically........................................... T. (T.) unicolor (Lepeletier, 1845) - Fore tibia with one spur apically, mid tibia with one to two spurs apically, hind tibia with two spurs apically............ 34 34. Mid and hind femora bright red (Fig. 80); lateral carina of propodeal areola almost parallel posteriorly, providing rectangular shape (Fig. 86)...................................................................................... 35 - Mid and hind femora black or light brown (Figs 16, 22, 40, 117); lateral carina of propodeal areola convergent posteriorly or subparallel, providing subtrapezoidal or subrectangular shape (Figs 21, 25, 43, 121)............................... 40 35. Subposterior area T1 with one transverse row of punctures................................................... 36 - Subposterior area of T1 with irregularly scattered punctures................................................... 38 36. Frons and vertex with smooth interspaces between punctures; tegula red and semitransparent...... T. (T.) agilis Smith, 1873 - Frons and vertex with coriaceous interspaces between punctures; tegula dark red and opaque........................ 37 37. Pronotum latero-ventrally with complete groove and deep punctures in middle...... T. (T.) pigmentata Allen & Jaynes, 1930 - Pronotum latero-ventrally with interrupted groove in middle, without deep punctures....... T. (T.) taiwana Ishikawa, 1967 38. Lower frons with dense punctures (Fig. 82)........................................... T. (T.) dayi Krombein, 1982 - Lower frons with sparse punctures....................................................................... 39 39. Head, mesosoma and metasoma with coriaceous interspaces between punctures in dorsal view.................................................................................................. T. (T.) femorata (Fabricius, 1775) - Head, mesosoma and metasoma with smooth interspaces between punctures in dorsal view......................................................................................... T. (T.) femorata biseculata Allen & Jaynes, 1930 40. Posterior margin of tegula with transverse weak, impressed line............................................... 41 - Posterior margin of tegula without transverse weak, impressed line (Figs 25, 43).................................. 42 41. Mesopleuron with coriaceous interspaces between punctures............................. T. (T.) alticola Tsuneki, 1986 - Mesopleuron with smooth interspaces between punctures.............................. T. (T.) takasago Tsuneki, 1986 42. Apex of clypeus medially truncated...................................................................... 43 - Apex of clypeus medially emarginated or slightly emarginated (Figs 17, 23, 41, 118)............................... 50 43. T1 anteriorly with dense and minute punctures........................................ T. (T.) tibetana Turner, 1908 - T1 anteriorly without dense and minute punctures.......................................................... 44 44. S1 without lateral groove........................................................ T. (T.) minutostriata Ma, 1936 - S1 with lateral groove on posterior half................................................................... 45 45. Mesoscutum with anterior medial groove connecting notaulus.............................. T. (T.) frater Parker, 1935 - Mesoscutum with anterior medial groove separated from notaulus.............................................. 46 46. Pygidium posteriorly smooth...................................................... T. (T.) sternata Parker, 1935 - Pygidium (Fig. 44) posteriorly with a few longitudinal striae.................................................. 47 47. Frons with medial longitudinal narrow carina or impunctate area............................................... 48 - Frons without medial longitudinal narrow carina or impunctate area............................................ 49 48. Frons with medial longitudinal narrow carina; vertex laterally with dense punctures along inner margin of eye................................................................................ T. (T.) inconspicua Allen & Jaynes 1930 - Frons (Fig. 41) with medial longitudinal narrow impunctate area; vertex laterally (Fig. 42) with sparse punctures along inner margin of eye............................................................... T. (T.) mongolica Tsuneki, 1986 49. Anterior carina of pronotum complete and strong.......................... T. (T.) minutopunctata Allen & Jaynes 1930 - Anterior carina of pronotum incomplete, medially weak.............................. T. (T.) choui Chen & Yang, 1991 50. Second intercubital vein of forewing nearly straight (Fig. 22)................................................. 51 - Second intercubital vein of forewing strongly or slightly sinuous (Figs 16, 117)................................... 52 51. Propodeal areola with complete sharp medial longitudinal carina approaching posterior margin of areola.................................................................................... T. (T.) nervidirecta Allen & Jaynes, 1930 - Propodeal areola (Fig. 25) with interrupted medial longitudinal carina reaching posterior 1/2 of areola............................................................................................ T. (T.) chihpenchia Tsuneki, 1986 52. Surface of propodeal areola smooth (Fig. 121)............................................................. 53 - Surface of propodeal areola coriaceous (Fig. 21)............................................................ 55 53. Lower frons with sparse punctures............................................ T. (T.) pempuchiensis Tsuneki, 1986 - Lower frons (Fig. 118) with dense punctures............................................................... 54 54. Propodeal areola (Fig. 121) normal, not pitted along inner margin of lateral carina............ T. (T.) s-secunda Allen, 1975 - Propodeal areola irregularly pitted along inner margin of lateral carina.............. T. (T.) retincisura Chen & Yang, 1991 55. Pronotum latero-ventrally without distinct groove in middle............................. T. (T.) hohrai Tsuneki, 1986 - Pronotum latero-ventrally with distinct groove in middle..................................................... 56 56. Pygidium (Fig. 20) with medial longitudinal carina..................... T. (T.) mediocarinata Han, Chen & Li, sp. nov. - Pygidium without medial longitudinal carina.............................................................. 57 57. S1 anteriorly with coarse punctures............................................. T. (T.) jinana Yang & Chen, 1988 - S1 anteriorly without coarse punctures................................................................... 58 58. Anterior half of pronotum with sparse punctures...................................... T. (T.) horiana Tsuneki, 1986 - Anterior half of pronotum with dense punctures.................................... T. (T.) malayana, Published as part of Han, Qian, Wang, He-Sheng, Chen, Bin & Li, Ting-Jing, 2023, A taxonomic revision of the nominotypical subgenus Tiphia Fabricius, 1775 (Hymenoptera: Tiphiidae: Tiphiinae) from China, with three new species and a key to the Chinese species, pp. 1-43 in Zootaxa 5284 (1) on pages 3-8, DOI: 10.11646/zootaxa.5284.1.1, http://zenodo.org/record/7918890, {"references":["Fabricius, J. C. (1775) Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. 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