Andocaeculus beatrizrosso sp. nov. (Figs. 31–59) Etymology. The specific name is a noun in apposition in honor to Beatriz Rosso, researcher of Universidad Nacional de Córdoba, Argentina, in recognition of her fundamental contributions to the knowledge of water mites. Diagnosis. Andocaeculus species of weyrauchi species group; aspidosoma with characteristic lateral constriction at level of seta Pa (Figs. 32, 34A), anterior border straight (Figs. 33, 35A) or with a medial incision in some exemplars (Figs. 32, 34A, D); palp tarsus with 3 eupathidia (Figs. 36, 47); b2 absent and d2 present (Figs. 32, 33), most setae on dorsal plates D, L and P not regressive, ellipsoidal; setae e2 and e2’ separated by ridges (Figs. 32–33). Setae of v’ series of leg I basifemur, telofemur and genu pedunculated, not pointed (Figs. 39A, 41B). Basifemur v’ (Figs. 40A, 42B), pedicelated, inserted in a long cuticular process, shorter than in other species of group, 54–60, 70–84 + long; RBaf, 0.56–0.59, RBaf + , 0.69–0.79, RTef, 0.43–0.53, RTef + , 0.61–0.78; tibia I (Figs. 40A, 43A–B) generally with 1 regressive seta in series d and tibia (Figs. 40B, 44D) II with one, solenidion φ present on tibia IV, RTi3: 3.2–4; setae l1” of tarsus II not eupathidial. Type material. Holotype, male (MACN-Ar 41112). Argentina, Provincia de Mendoza, Departamento de Las Heras, Cerro Arcos, S32.858206°, W68.938977° (+/- 500 m), elev. 1150 masl (+/- 50 m), 24.Dec.2012, A. Porta leg., Berlese extraction of debris of xeric scrub; 6 paratypes, males (MACN-Ar 41105, 41106, 41113 to 41116), same data, cleared in lactic acid and preserved in alcohol; 7 paratypes, males (MACN-Ar 41107 to 41110, 41118 to 41119, 41121), same data, 29.Mar.2011, mounted on slides; 2 paratypes, females (MACN-Ar 41121, 41122), same data, mounted on slides; 3 paratypes, females (MACN-Ar 41137 to 41139), same data, cleared in lactic acid and preserved in alcohol; 6 paratype tritonymphs (MACN-Ar 41140-41145), same data, mounted on slides; 3 deutonymph paratypes (MACN-Ar 41147 to 41149), same data, 10.May.2013, mounted on slides; 2 larvae paratypes (MACN-Ar 41111, 41117), same data, 10.May.2013, mounted on slides; 2 paratypes, adults (MACN-Ar 41103, 41153), same data, in individual vials, preserved in alcohol; 2 paratypes, adults (CAI 4336, 4337), same data, in individual vials, preserved in alcohol 7 paratypes, 5 adults and 2 larvae, on individual stubs with exemplars mounted for SEM (MACN-Ar 41104), same data; 1 male paratype (MACN-Ar 41120) from Provincia de Mendoza, Mendoza Capital, Reserva de Flora Nativa Bosque Xerófilo, S32.891690°, W68.864855° (+/- 200 m), elev. 850 masl (+/- 5 m), 25.Dic.2012, A. Porta leg., Berlese extraction of debris of xeric scrub; 1 paratype tritonymph (MACN-Ar 41146), same data, mounted on slide; 1 paratype protonymph paratype (MACN-Ar 41150), same data, 08–10.May.2013, mounted on slide; 3 paratypes, adults (MACN-Ar 41150 to 41152), same data, preserved in alcohol; 2 paratypes, adults (MACN-Ar 41124, 41125), same data, 25.Dec.2012, preserved in alcohol. Description Male (n=10). Color (Fig. 31). Gnathosoma and dorsal sclerotized plates on idiosoma dark brown, membranous integument pale cream, legs also dark brown but darker than rest of body. Gnathosoma (Figs. 32, 33, 34A, D, 35A). Covered by aspidosoma in dorsal view. Chelicerae (Fig. 36B) typical for family, 44 wide at base, 144–160 long, movable digit hook-like with 3 minute teeth; cheliceral seta ch slender (Figs. 35B–C), 6 long. Palp (Figs. 36B, 48) 4-segmented, trochanter glabrous; femorogenu with 2 clavate setae in dorsal series, d1 shorter than d; d1 and d 28–34 and 48–60 long, respectively; tibia with prominent calcar d on projecting tubercle, 26–34 long; 3 clavate setae d1, l’ and l”, 28–34, 26–36 and 32–34 long, respectively, plus one more calcar ld, 32–38 long. Palp tarsus subconical, 50–64 long, 10–16 wide at base, solenidion ω present on antiaxial face, 6–7 setae: 3 eupathidia, (u) and l ’, in addition to v, d, l” and, only in some exemplars, l1’ (Figs.36A, 48B–C). Subcapitulum (Fig. 35A) posteriorly rounded, anteriorly subconical, 156–160 long, 198–220 wide at level of palp insertion, setae m 34 long, anterior to n, 30 long, two pairs of adoral setae or 1-2 (Fig. 35B,C), 8 long. Idiosoma (Figs. 32, 33). In dorsal view subtriangular, 822–853 long, 616–671 wide at level of coxa IV. Aspidosoma (Fig. 34A). Subtriangular, 258–275 long, posterior margin 213–221 wide, anterior margin 60–66 wide, projecting horizontally over gnathosoma. Anterior portion with characteristic lateral constriction. Anterior margin straight (Figs. 22, 35A) or with medial incision in some exemplars (Figs. 34A, D). Seta Po (Fig. 34A, D) very long, its length 86–120, expanded distally, with anterior third setulate. Trichobothrial seta bo spatulate, 80–96 long, upturned in distal half and broadening at tip (Fig. 34D). Aspidosomal setation regressive, Pa (Fig. 34B) regressive, subcylindrical, 8 long, located on a transversal ridge; Pm regressive (Fig. 34C), cylindrical, 6 long, inserted on lateral border of aspidosoma, setae Pp absent. Lateral eye plates (Fig. 34A) separated from other plates; anterior pair of lateral eyes 14–22 diameter, posterior pair 22–24. Median eye 42–54 long, 44–50 wide. Hysterosoma. Dorsal view. With 5 dorsal plates D, (L), M, P (Figs. 32, 33). Dorsal plate D subtrapezoidal, 260–295 long, anterior margin 197–209 long, posterior margin 274–295 long, with one longitudinal and two transversal ridges, setae a1, b1, c1, ellipsoidal, lengths: a1: 26–30, b1: 28–30, c1: 26–28; distance between setal insertions a1-a1: 98–106, b1-b1: 54–58, c1-c1: 64–74, a1-b1: 102–106, b1-c1: 108–120. Paired lateral plates L 320–336 long, 72–80 wide; a2 regressive, 9 long; b2 absent and c2 present with p=0.5; lyrifissure ia located on external side of plate, 26–30 long; lyrifissure im located on median line of plate, 12–18 long. Median plates M fused, with d1 and d2 36–38 and 29–36 long, respectively; distance between setal insertions d1-d1: 96–120, d1-d2: 130–146, d2-d2: 367–371. Posterior plates P divided by ridges, setae e1, e2 and e2’, 46–48, 38–44, and 38–44 long, respectively; in some exemplars e2 ” present, setae e2 and e2’ inserted in different sectors of plate, separated by a ridge; distance between setal insertions e1-e1: 100–126, e1-e2: 86–96. Seta hs on posterior border of idiosoma, 40–42 long. Ventral view, podosoma and opisthosoma (Figs. 37, 38). Membranous integument striate. Coxal setation formula: 4:2–3:2–3:2, coxal setae clavate. Measurements of coxal setae, 1a: 20–26; 1b: 30–32; 1c: 32–36; 1d: 36–46; 2a: 18–22; 2b: 22–23; 2c: 16; 3a: 16–22; 3b: 16–22; 3c: 26; 4a: 20–22; 4b: 20–21; 4c: 16–22. Aggenital plates (Fig. 31B) poorly sclerotized, triangular, 72–80 long, 66–80 wide and not differentiable from integument in exemplars cleared in lactic acid. Genital opening 150–156 long, aggenital plates 34–42 wide, with 6 pairs of genital setae, 20– 24 long. Anal opening 136–146 long, adanal plates 32–34 wide, with two pairs of adanal clavate setae, 8–12 long. Pseudanal plate 164–170 long, 64–68 wide, with three pairs of setae, ps1 clavate, 34–36 long, normally developed, ps2 and ps3 regressive, ps2 10 long, ps3 8–9 long, membranous integument with 9 pairs of setae. Internal genitalia (Fig. 39) with sclerites as in other species of the group, relatively large for Caeculidae in relation to idiosoma width, 130–140 wide at level of anterior ventral arch, 90–116 long. Ventral side (Fig. 39A) with 9 pairs of setae, 3 pairs anterior and medial, 2 pairs inserted on sclerites on each side, and 2 posterior pairs, all setae simple and unbranched, relatively long for the family in relation to the size of genital sclerite (cfr. Otto, 1993, Fig. 8; Fuangarworn & Butcher, 2015, Fig. 14; Ott & Ott, 2018, Fig. 4). Two laterodorsal apophyses clearly visible from dorsal side (Fig. 39B). Legs (Figs. 40–47). Measurements see Table 9. Most setae inserted on big cuticular tubercles. Leg I (Figs. 40A, 41–43). Trochanter (Figs. 40A, 42A) with one seta in series l’ and one regressive seta in d. Basifemur (Figs. 40A, 42B) with one seta in v’, pedicelated, inserted in a long cuticular process, shorter than in other species of the group, 54–60, 70–84 + long, RBaf: 0.56–0.59, RBaf + : 0.69–0.79, setae d, l”, v” and v”1 regressive, l’ petiolate. Telofemur (Figs. 40A, 42B) with seta v’ subequal to v’ in telofemur, but more curved, 46–50, 70–76 + long, RTef: 0.43–0.53, RTef +: 0.61–0.78; setae d, l’, l” and v” regressive. Genu (Figs. 42C–D) with 3 regressive setae in series d, 1 and 3 petiolate setae in l’ and l” series, respectively, l’, 46–48, 56–70 + long, 3 petiolate setae in v’ series with v’1 medially displaced, v’, 42–44, 50–64 + long, v2’, 40–46, 56–64 + long; 2 petiolate setae in v” series. Tibia (Figs. 39A, 42D, 43A–B, E) usually with only 1 regressive seta in series d, excepting 2 exemplars asymmetrically with 2 (Fig. 40A); 4 pedunculated setae in each lateral series, 4 spinous setae with blunt tip in ventral series, (v2) medially displaced, v’, 60–64, 70–74 + long, v1’, 50–54, 60–66 + long v”, 50–52 long, solenidion φ and famulus κ” inserted laterodorsally and distally (Figs. 43B, E). Tarsus (Figs. 41, 42D, 43C–D, F–G) with each of series l’, l”, v’ and v” with 3–4, 3–4, 4, 4 setae, respectively, l1” usually eupathidial with p=11/14 (Fig. 41), with some asymmetry (Fig. 43F), solenidion ω inserted dorsally (Figs. 41, 43G) at level of l1”, simple in structure (Fig 41), famulus ε inserted in antiaxial facies (Fig. 41) at level of v1”; eupathidia: er’, (st) and, putatively, l1”, trichobothria bt 16–20 long, two claws ol’ and ol”, 10 and 28–30 long, respectively. Setal count (solenidia): 2, 6, 5, 12, 18–19(1), 19–21(1). Leg II (Fig. 40B, 44). Trochanter (Fig. 44A) with d regressive and l’ and l” petiolate. Basifemur (Fig. 44C) with d and l” regressive, l’, v”, v1” spatulate. Telofemur (Fig. 44C) with d, l’, v and l” regressive, l’ spatulate normally developed. Genu (Fig. 44B) with 3 regressive setae in series d, 3 and 2 pedunculated setae in series l’ and l”, respectively, 2 setae in series v’, seta v” 58–60 long. Tibia (Fig.44D, F) with 1 regressive seta in series d, 4 and 3 pedunculated setae in series l’ and l”, respectively, 3 spinous setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 44F). Tarsus (Fig. 44E) with 3 setae in series l’, l” and 4 in v’ and v”, solenidion ω inserted dorsally at level of l”, only er’ eupathidial; (st) absent; trichobothria bt 14–20 long. Claws ol’ and ol” length 10–12 and 30–34, respectively. Setal count (solenidia): 3, 5, 5, 11, 14(1), 17(1). Leg III (Figs.45A, 46). No eupathidia present. Trochanter (Figs. 45A, 46A) with d regressive and l” petiolate. Basifemur (Fig. 46B) with d regressive, l’, v’, v” and v1” petiolate. Telofemur (Fig. 46B) with d, l’, v” and l” regressive and v’ petiolate. Genu (Fig. 46B) with 3–4 and 1–2 regressive setae in series d and l’, respectively, and 3 pedunculated setae in series v’ and l”, 2 setae in series v”. Tibia (Fig.46D, F) with 2–4 regressive seta in series d, 4 and 3 pedunculated setae in series l’ and l”, respectively, 5 pointed setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 46G), RTi3: 3.2–4. Tarsus (Fig. 46E) with 1 and 3 setae in series l’, l”, respectively, and 3 in v’ and v”; er” present, er’ absent, trichobothria bt 100–110 long. Claws ol’ and ol”, 10 and 36–44 long, respectively. Setal count (solenidia): 2, 5, 5, 12–14, 16–18(1), 12. Leg IV (Figs.45B and 47). Trochanter (Figs. 45A, 46A) with d and l” petiolate and v’ regressive. Basifemur (Fig. 46B) with v” regressive and l’, v’ spatulate. Telofemur (Fig. 46B) with d, l’ and v” regressive and v’ and l” spatulate. Genu (Fig. 47C) with 2 regressive setae in series d, 2 pedunculated setae in each series v’ and l”, 1 seta in series v”. Tibia (Fig.47D, F–G) with 2–3 regressive seta on tibia series d, 3 pedunculated setae in series l” and 4 pointed setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 47F–G). Tarsus (Fig. 47E) with 1 and 2 setae in series l’ and l”, respectively; 3 setae in series v’ and v”, er” present, er’ absent, trichobothria bt 114–140. Claws ol’ and ol” 9–10 and 50–56 long, respectively. Setal count (solenidia): 3, 3, 5, 7, 13–14(1), 11. Female (n=3). Externally similar to male except in size. Gnathosoma. Chelicera 128–136 long. Palp 4-segmented, with chaetotaxy as in male. Subcapitulum 176–192 wide at level of palp insertion, 150–170 long, setae m anterior to n, two pairs of adoral setae or1-2. Idiosoma. 786–924 long, 572–687 wide at level of coxa IV. Aspidosoma. Subtrapezoidal, 258–274 long, posterior margin 213–266 wide, anterior margin 68–80 wide, projecting horizontally over gnathosoma. Seta Po 100–106 long, trichobothrial seta bo 88–120 long. Aspidosomal setation regressive, Pa and Pm minute, Pp absent; anterior pair of lateral eyes 16–20 diameter, posterior pair 18–22. Median eye 40 long, 48 wide. Hysterosoma. Dorsal view. Dorsal plate D subtrapezoidal, 258–320 long, anterior margin 216–287 long, posterior margin 280–307 long, a1: 22–28, b1: 28–38, c1: 30–38; distance between setal insertions, a1-a1: 100–116, b1-b1: 50–72, c1-c1: 46–76, a1-b1: 102–116, b1-c1: 96–128. Paired plates L 360 long, 60–80 wide; a2 regressive 9 long; b2 and c2 absent, ia and im 30–32 and 16–20 long, respectively. Median plates M fused, with d1, d2, 36–42 and 26–30 long, respectively, distance between setal insertions, d1-d1: 90–130, d2-d2: 347–435, d1-d2: 134–158. Posterior plates P, divided by ridges, setae e1, e2, e2’ and e2 ” , 38–44, 38–46, 36–46 and 36–40 long, respectively. Setae e2’ and e2” separated by a ridge from e2, expression of e2” variable (p=0.5), distance between setal insertions, e1-e1: 110–140, e1-e2: 60–98. Seta hs on posterior border of idiosoma, 34–36 long. Ventral view, podosoma and opisthosoma. Coxal setation formula 4:3:3:2–3. Genital opening 150–162 long, with 6 pairs of genital setae, 13–18 long. Anal opening 140–148 long, adanal plates 40–44 wide, with two pairs of adanal clavate setae, 16–18 long. Pseudanal plate 160–162 long, 56 wide, with three pairs of setae. Legs. Measurements see table 10. Ratios, RBaf: 0.5–0.58; RBaf + : 0.77–0.83; RTef: 0.43–0.51; RTef + : 0.73–0.85. Larva (n=3) (Figs 49–59) Gnathosoma. Covered by aspidosoma in dorsal view (Fig. 53). Chelicera 46 long; cheliceral seta ch present. Palp (Fig. 54) 4-segmented, trochanter glabrous, femorogenu with 1 clavate seta in dorsal series, 9–10.4 long; tibia with a calcar d, 11 long, located on a projecting tubercle, 2 clavate setae l’ and l”, and 1 calcar setae ld; 8–11 long, tarsus subconical, solenidion ω present on antiaxial face, 5 setae present, setae v, l’, and d in addition to 2 eupathidia (u). Subcapitulum (Fig. 54) 90 wide at level of palp insertion, 60 long, setae m 12–13 long, anterior to n, 10 long, two pairs of adoral setae or 1-2. Idiosoma (Figs. 49–50). 264–320 long, 184–224 wide at level of coxa III. Aspidosoma (Figs. 51–52). Subtrapezoidal, 106–108 long, posterior margin 100–106 wide, anterior margin 32–36 wide, projecting horizontally over gnathosoma. Seta Po 38 long, trichobothrial seta bo 42 long (Figs. 51B, 52), Pa and Pm minute, Pp absent. Lateral eye plates (Figs. 49, 50A, 51A) separa