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Andocaeculus Coineau, 1974 Type species Microcaeculus brundini Franz, 1962 Revised diagnosis. The original diagnosis of Coineau (1974a: 279) stated that mites of the genus Andocaeculus corresponds to caeculids with trichobothria present on all leg tarsi, with claws unequally sized, and having dorsal plates without intensive neotrichy. To these characters, we could add: gnathosoma not visible from above; only one seta Po (short or long); palpal femur with at most two setae; anal plate with two setae; femora divided into telofemur and basifemur on all legs; ω solenidion dorsally located, ε antiaxial and hidden, famulus κ” regressive; eupathidia of the typical ‘baguette de tambour’ of most of the genera of the family (Coineau 1974a) and (st) absent on tarsus II., Published as part of Porta, Andrés O., Pizarro-Araya, Jaime & Ramírez, Martín J., 2021, Revision and phylogeny of the genus Andocaeculus (Acari: Caeculidae) I: the A. weyrauchi species group, pp. 1-78 in Zootaxa 4945 (1) on page 6, DOI: 10.11646/zootaxa.4945.1.1, http://zenodo.org/record/4614242, {"references":["Franz, H. (1962) Ein neuer Microcaeculus aus Sudamerika. (Acari, Trombidiformes) Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 101 - 102, 98 - 100.","Coineau Y. (1974 a) Elements pour une monographie morphologique, ecologique et biologique des Caeculidae (Acariens). Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 81, 1 - 299, pls. 1 - 24."]}

Andocaeculus weyrauchi (Franz, 1964) (Figs 2–17) Microcaeculus weyrauchi Franz, 1964: 89–90; Taylor, Gunawardene and Kinnear, 2013: 449. A. weyrauchi, Coineau 1974a: 279. Type material. Type series from “ Puna El Infernillo in 3000 m Seehöhe am Fusse von flechtenbewachsenen Felsblöcken aus Moos, Rasen und Erde unter diesem am 13. 9. 1963 gesiebt ”, Herbert Franz collector, not examined (it should be deposited in MNHN, not found). Examined material. 4 females (MACN-Ar 41058, 41059, 41135, 41136), Argentina, Provincia de Tucumán, Abra el Infiernillo, S26.74272°, W065.77283° (+/- 50 m), alt. 3012 masl (GPS), under stones, hand collected and on litter of xeric vegetation, extracted with Berlese funnels, A. Porta leg., cleared with lactic acid, preserved in alcohol; 1 adult (MACN-Ar 41056), same data, not cleared, preserved in alcohol; 2 adults (MACN-Ar 41058), same data. preserved in alcohol; 10 adults in a vial (MACN-Ar 4134), same data, preserved in alcohol; 1 larva (MACN-Ar 41065), same data, mounted on slide; 4 protonymphs (MACN-Ar 41066 to 41069), same data, mounted on slides; 2 deutonymphs (MACN-Ar 41070 and 41071), same data, mounted on slides; 6 tritonymphs (MACN-Ar 41072 to 41077), same data, mounted on slides; 14 females (MACN-Ar 41060 to 41065, 41126 to 41033), same data, mounted on slides; 1 male (MACN-Ar 41078), same data, mounted on slide; 3 individual stubs with exemplars mounted for SEM (MACN-Ar 41057), same data; 5 individuals, (MACN-Ar 41079 to 41083), vouchers of BOLDSYSTEM data SPDAR1609-16 to 1609-20, same data, preserved in alcohol; 2 adults in a vial (FML-Ar 00286), same data, preserved in alcohol. Remark. The examined material has been assigned to A. weyrauchi on the basis of being the only species collected in the type locality and because it exhibits a morphology that corresponds exactly with that of A. weyrauchi in Franz’s description. Diagnosis. A. weyrauchi differs from other species of the group by the following combination of characters: subtrapezoidal form of the idiosoma (Figs. 3, 4); anterior margin of aspidosoma straight (Fig. 5A); palp tarsus with 4 eupathidia; opisthosoma with a relatively less regressive chaetotaxy (Figs. 3 and 4): most setae normal-sized, elongated, b2 and c2 present in some exemplars, neotrichy sometimes present on d2 seta sector (Fig. 3 and 4); with four pairs of setae on P plates, setae e2 and e2’ with e2” separated by ridges (Fig. 4); setae of series v’ on basi and telofemur I of adults, elongated, distally blunt, relatively long for the group, 54–66 and 80–96 + long in females, RBaf: 0.52–0.6, RBaf + : 0.75–0.9, RTef: 0.5–0.57, RTef + : 0.75–0.84; genua I and II with 3 and tibiae I and II with 2 regressive setae of series d (Figs. 9D and 10 D); setae of series v’ on genua and tibiae I and II pointed; setae l1’ always eupathidial on leg I and II (Figs. 9A–J); solenidion present on tibia IV (Figs. 11B, 13 B–C), RTi3: 2.6–4.3. Redescription Female (n=14). Color (Fig. 2). Gnathosoma and dorsal sclerotized plates on idiosoma brown, membranous integument pale cream, legs and hypostome dark brown. Gnathosoma (Figs. 3, 5). Covered by aspidosoma in dorsal view. Chelicera (Fig. 5A) typical for family, 54–58 wide at base, 132–170 long, movable digit hook-like with 3 minute teeth; cheliceral seta ch slender, 6 long. Palp (Figs. 14B, 15) 4-segmented, trochanter glabrous, femorogenu with 2 clavate setae in dorsal series, d1 26–40 long, d, 42–54 long; tibia with two calcars l’ and d, with d on projecting tubercle and 3 clavate setae d1, ld and l”, setal lengths, l’: 24–30, l”: 30–36, ld: 30–38, d: 26–33, d1: 26–34. Tarsus subconical, 42–54 long, 16–22 wide at base, solenidion ω present on antiaxial face, with 6–7 setae, v, l1’, l ” in addition to 4 eupathidia, (u), l’ and d. Subcapitulum (Figs. 6, 7) posteriorly rounded, anteriorly subconical, 148–230 wide at level of palp insertion, 146–170 long, setae m 22 long, anterior to n, 22 long, two pairs of adoral setae or 1–2, 12–14 long. Idiosoma. In dorsal view subtrapezoidal (Figs. 3, 4), 860–1110 long, 593–790 wide at level of coxa IV. Aspidosoma (Fig. 5). Subtrapezoidal, 284–324 long, posterior margin 221–276 wide, anterior margin 70–90 wide, projecting horizontally over gnathosoma. Anterior margin straight, with a central depression. Seta Po (Fig. 5D) 100–120 long, expanded distally, with anterior third setulate, dorsally visible from anterior margin of aspidosoma. Trichobothrial seta bo spatulate, 90–104 long, distal half upturned, broadening at tip (Fig. 5D). Aspidosomal setation regressive, Pa (Fig. 5B) regressive, subcylindrical, 10–12 long, located on a transversal ridge that is united in some exemplars to longitudinal ridge; Pm regressive (Fig. 5C), cylindrical, 10–11 long, on lateral border, setae Pp absent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 18–24 diameter, posterior pair 24–32 (Fig. 5A), median eye 40 long, 60 wide. Hysterosoma. Dorsal view. With 5 dorsal plates D, (L), M and P (Figs. 3, 4). Dorsal plate D subtrapezoidal, 279–398 long, anterior margin 184–232 long, posterior margin 280–320 long, with one longitudinal and two transversal and ridges, setae a1, b1, c1, ellipsoidal, lengths a1: 24–39, b1: 28–36, c1: 36–50; distance between setal insertions a1-a1: 106–130, b1-b1: 64–80, c1-c1: 80–98, a1-b1: 116–142, b1-c1: 108–152. Paired plates L 344–440 long, 74–100 wide; a2 regressive, 12–28 long; b2 present with p=0.36 (n=14), 32–40 long, c2 when present (p=0.43, n=14) ellipsoidal, 24–40 long; lyrifissure ia located on external side of plate, halfway between a2 and b2, 24–34 long; lyrifissure im located on median line of plate, halfway between b2 and c2, 12–18 long. Median plates M fused, with d1, d2 46–60 and 30–52 long, respectively; sometimes with neotrichy (p=0.29, n=14) on d2 sector; distance between setal insertions d1-d1: 122–178, d1-d2: 126–154, d2-d2: 308–434. Posterior plate P divided by ridges, setae e1, e2, e2’ and e2”, 50–58, 34–44, 32–44 and 30–40 long, respectively. Setae e2’ and e2” inserted on different sector of plate, which is separated by a ridge from e2; distance between setal insertions e1-e1: 110–138, e1-e2: 84–94. Seta hs on posterior border of idiosoma, 40–46 long. Ventral view, podosoma and opisthosoma (Fig. 6, 7). Membranous integument striate. Coxal setation formula 4:3:4:2–3, coxal setae clavate; their lengths, 1a: 20–28, 1b: 26–34, 1c: 32–38, 1d: 42–46; 2a: 23–26, 2b: 24–30, 2c: 26–30, 3a: 20–28, 3b: 24–28, 3c: 24, 3d: 22–24, 4a: 25–30, 4b: 24–26, 4c: 20–28, 4d: 20–30. Genital opening 140–174 long, 60–66 wide, with 6 pairs of genital setae, short, 16–22 long. Aggenital plate not observable. Anal opening 142–144 long, adanal plates 38–40 wide, each with two adanal clavate setae, 16–20 long. Pseudanal plate 80–82 long, 54–60 wide, with three pairs of setae, ps1 clavate 30–49 long, normally developed, ps2 and ps3 regressive, ps2 16–22 long; ps3 16–20 long, membranous integument with 8 pairs of setae. Legs (Figs. 8–14). Measurements see Table 2. Leg I. Trochanter (Fig. 9A) with one seta in series l’ and one regressive seta in d (Figs. 8A, 9A). Basifemur (Figs. 8A, 9B) with setae on v’ not pointed, distally blunt, but longer than other species of group, 54–64 and 80–96 + long, RBaf: 0.52–0.6, RBaf + : 0.75–0.9, 2 setae in series v”, both regressive (Fig. 9K), one pedicelated seta in series l’ and one regressive in series l” and d. Telofemur (Figs. 8A, 9B) with seta v’ subequal to v’ on basifemur, 54–66, 80–96 + long, RTef: 0.5–0.57, RTef + : 0.75–0.84, setae l’; l” and v” regressive. Genu (Figs. 8A, 9C) with 3 setae in series d; 2 setae in series l’, distal l’ elongated, 54–60, 70–78 + long; 3 pedicelated setae in series l”; series v’ with 3 pointed setae, with v1’ medially displaced; v’, 62–74, 82–100 + long, v2’, 46–62, 64–80 + long; series v” with 2 setae (Fig. 9K). Tibia (Figs. 8A, 9D–E, K) with 2 regressive setae in series d, 4 setae on both lateral series; series v’ with 4 pointed setae, 62–94, 74–106 + long, v2’ medially displaced; series v” with 4 pointed setae (Fig. 9K), v2” medially displaced, v” 52–70 long. Solenidion φ and famulus κ ” inserted dorsoantiaxially on distal portion of segment (Fig. 9D–E). Tarsus (Figs. 9G–J, 14A and C) with 3–4 setae in series l’, 4 setae in series l”, with l1” eupathidial, 3–4 setae in series v’ and 4 setae in series v”; trichobothria tb 24–34 long. Solenidion ω inserted dorsally (Figs. 9G, 14A) at level of l1”, simple in structure (Figs. 14A and C). Famulus inserted at antiaxial facies (Fig. 14C) at level of l1”. Four eupathidia: l1”, er’ and (st). Two claws ol’ and ol” lengths 8–10 and 30–42, respectively. Setal count (solenidia): 2, 6, 5, 13, 18(1), 20–22(1). Leg II (Figs. 8B and 10). Trochanter (Fig. 10A) with one seta in each lateral series and one regressive seta in series d. Basifemur (Fig. 10B) d, l” and v1” regressive, v’ and v” pedunculated. Telofemur (Figs. 8B, 10B) with d, l’, l” and v” setae regressive and v’ pedunculated. Genu (Figs. 8A, 10C) with 3–4 regressive setae in d series, 3 pedunculated setae in l’ and 4 setae in l”, v1’ displaced externally (Fig. 8B), v” long and pointed (Fig. 8B), 72–84 long. Tibia (Fig.10D) with two regressive seta in d, 4 pedunculated setae in l’ and 3–4 on l”, 3 pointed setae in v’ and v”, solenidion inserted as on tibia I (Figs. 10E, F). Tarsus (Figs. 8B and 10G–I) with 4 setae in series l’ and l”, l1” always eupathidial, 4 setae in series v’ and 3 in v”, solenidion ω inserted dorsally at level of l’, famulus antiaxial, (st) pair absent, trichobothria bt 24–34 long. Two eupathidia: l1” and er’. Claws ol’ and ol” lengths 7–10 and 28–36, respectively. Setal count: 3, 5, 5, 13–14, 15(1), 18(1). Leg III (Figs. 11A and 12). Setae on series v’ and v” spatulate, eupathidia absent. Trochanter (Fig. 12A) with anterior border rounded, with pedunculated setae: v’ and l” and 1 regressive in d. Basifemur (Fig. 12B) with l” and v’ elongated and d and v” regressive. Telofemur (Fig. 12B) with l’, l”, v” and d regressive and v’ elongated and pedunculated. Genu (Figs.11A, 12C) with 2–3, 1–2 regressive setae in series d and l’, respectively; 3 pedunculated setae in series v’ and 3 regressive in series v”. Tibia (Figs. 12D, F) with 3–4 regressive setae in series d, 3–4 setae in series l” and 4–5 in both ventral series, solenidion φ inserted on antiaxial face of segment. Tarsus (Fig. 12E) with 3–4, 2, 4–5, 3–4 setae in series l’, l”, v’ and v”, respectively, er” present and er’ absent; trichobothria bt 108–120 long, claws ol’ and ol” of length 12–16 and 26–36, respectively. Setal count: 3, 4, 5, 12–14, 15–18(1), 14–17. Leg IV (Figs. 11B and 13) Setae as on leg III. Trochanter (Fig. 13A) with 3 setae: l” pedunculated and v’ and d regressive. Basifemur (Fig. 13E) with v’ regressive and l’ and v” pedunculated. Telofemur (Fig. 11B, 13E) with l’, v’ and d regressive, with setae of series v’ and l” elongated and pedunculated. Genu (Fig. 13G) with 2 and 1 regressive setae in series d and l’, respectively; 2 elongated and pedunculated setae each in series v’ and l”. Tibia (Figs. 13B–D) with 3 regressive setae in series d, 4 setae in v series and 3 in series l”, solenidion φ present (Fig. 13 B–C). Tarsus (Fig. 13F) with 2–4, 1–2, 3, 3 setae in series l’, l”, v’ and v”, respectively, er” present and er’ absent, claws ol’ and ol” lengths 12–16 and 42–52, respectively. Trichobothria 108–150 long. Setal count: 3, 3, 5, 9, 14(1), 11–14. Male (n=1). Externally similar to female except in size. Gnathosoma. Chelicera 132–136 long. Palp 4-segmented, with chaetotaxy as in female. Subcapitulum 192 wide at level of palp insertion, 146 long, setae m anterior to n, two pairs of adoral setae. Idiosoma. 860 long, 592 wide at level of coxa IV. Aspidosoma. Subtrapezoidal, 269 long, posterior margin 221 wide, anterior margin 70 wide, projecting horizontally over gnathosoma. Seta Po 100 long, trichobothrial seta bo 90 long. Aspidosomal setation regressive, Pa and Pm minute, Pp absent; anterior pair of lateral eyes 20–22 diameter, posterior pair 26–32. Hysterosoma. Dorsal view. Dorsal plate D subtrapezoidal, 299 long, anterior margin 197 long, posterior margin 254 long, a1: 22–26, b1: 26, c1: 32; distance between setal insertions, a1-a1: 92, b1-b1: 50, c1-c1: 58, a1- b1:108–110, b1-c1: 114–118. Paired plates L 336–344 long, 70 wide; a2 regressive, 16 long; b2 and c2 absent, ia and im, 30–32 and 16 long, respectively. Median plates M fused, with d1, d2, 46–48 and 30 long, respectively, distance between setal insertions, d1-d1: 96, d2-d2: 292, d1-d2: 100–102. Posterior plates P, divided by ridges, setae e1, e2, e2’, 44–50, 34–36 and 30 long, respectively. Setae e2’ separated by ridge from e2, distance between setal insertions, e1-e1: 90, e1-e2: 68–76. Seta hs on posterior border of idiosoma, 50 long. Ventral view, podosoma and opisthosoma. Coxal setation formula 4:3:4:3. Genital opening 162 long, with 6 pairs of genital setae, 24–30 long. Aggenital plate subtriangular, poorly sclerotized. Anal opening 154 long, with two pairs of clavate adanal setae, 16–18 long. Pseudanal plate 168–180 long, 60 wide, with three pairs of setae. Genital sclerites as other species of group (Fig. 39), with nine pairs of simple, unbranched setae, 14–16 long. Legs. Measurement see Table 3. Ratios: RBaf: 0.54; RBaf + : 0.78; RTef: 0.54; RTef + : 0.76. Larva (n=1) Gnathosoma. Covered by aspidosoma in dorsal view. Chelicera 42–44 long; cheliceral seta ch present. Palp 4-segmented, trochanter glabrous, femorogenu with 1 clavate seta in dorsal series, 9–10 long; tibia with a spiniform d seta, 11 long, located on a projecting tubercle, 2 clavate setae l’ and l”, and two spiniform setae ld; 8–11 long, tarsus subconical, solenidion ω present on antiaxial face, 5 setae present: v, l’, and d in addition to 2 eupathidia, (u). Subcapitulum 90 wide at level of palp insertion, 60 long, setae m 12–13 long, anterior to n, 10 long, two pairs of adoral setae or 1-2. Idiosoma 355 long, 269 wide at level of coxa III. Aspidosoma. Subtrapezoidal, 136 long, posterior margin 132 wide, anterior margin 40 wide, projecting horizontally over gnathosoma. Seta Po 38 long, trichobothrial seta bo 42 long. Aspidosomal setation regressive, Pa and Pm minute, Pp absent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 8 diameter, posterior pair 8–10. Hysterosoma. Dorsal view (Fig. 16A). Dorsal plates poorly defined; only setae a1, b1, c1, a2, d1, e1 and hs present. Lengths of anterior hysterosomal setae, a1: 8–10, b1: 9–10, c1: 9; a2 regressive, 10 long, distance between setal insertions, a1-a1: 58, b1-b1: 38, c1-c1: 52, a1-b1: 45, b1-c1: 43–44. Lengths of posterior hysterosomal setae, d1: 22, e1: 28, distance between setal insertions d1-d1: 48, e1-e1: 38. Ventral view, podosoma and opisthosoma (Fig. 16B). Coxal setation formula 2:0:1, setae 16–18 long, Claparede’s organs present on anterior margin of coxae II. Anal opening 70 long, hs 16 long, ventral lyrifissure ih observable at level of anal plate. Legs (Fig. 17). Measurements see table 7. Femora entire on all legs, Rf: 0.65–0.73 Leg I. Tr, 0, Published as part of Porta, Andrés O., Pizarro-Araya, Jaime & Ramírez, Martín J., 2021, Revision and phylogeny of the genus Andocaeculus (Acari: Caeculidae) I: the A. weyrauchi species group, pp. 1-78 in Zootaxa 4945 (1) on pages 7-28, DOI: 10.11646/zootaxa.4945.1.1, http://zenodo.org/record/4614242, {"references":["Franz, H. (1964) Neue Caeculiden aus Afrika und Sudamerika. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 103 - 104, 82 - 93.","Taylor, C. K., Gunawardene, N. R. & Kinnear, A. (2013) A new species of Neocaeculus (Acari: Prostigmata: Caeculidae) from Barrow Island, Western Australia, with a checklist of world Caeculidae. Acarologia, 53 (4), 439 - 452. https: // doi. org / 10.1051 / acarologia / 20132105","Coineau Y. (1974 a) Elements pour une monographie morphologique, ecologique et biologique des Caeculidae (Acariens). Memoires du Museum National d'Histoire Naturelle, Serie A, Zoologie, 81, 1 - 299, pls. 1 - 24."]}

Porta, Andrés O., Pizarro-Araya, Jaime, Ramírez, Martín J. (2021): Revision and phylogeny of the genus Andocaeculus (Acari: Caeculidae) I: the A. weyrauchi species group. Zootaxa 4945 (1): 1-78, DOI: https://doi.org/10.11646/zootaxa.4945.1.1

Andocaeculus beatrizrosso sp. nov. (Figs. 31–59) Etymology. The specific name is a noun in apposition in honor to Beatriz Rosso, researcher of Universidad Nacional de Córdoba, Argentina, in recognition of her fundamental contributions to the knowledge of water mites. Diagnosis. Andocaeculus species of weyrauchi species group; aspidosoma with characteristic lateral constriction at level of seta Pa (Figs. 32, 34A), anterior border straight (Figs. 33, 35A) or with a medial incision in some exemplars (Figs. 32, 34A, D); palp tarsus with 3 eupathidia (Figs. 36, 47); b2 absent and d2 present (Figs. 32, 33), most setae on dorsal plates D, L and P not regressive, ellipsoidal; setae e2 and e2’ separated by ridges (Figs. 32–33). Setae of v’ series of leg I basifemur, telofemur and genu pedunculated, not pointed (Figs. 39A, 41B). Basifemur v’ (Figs. 40A, 42B), pedicelated, inserted in a long cuticular process, shorter than in other species of group, 54–60, 70–84 + long; RBaf, 0.56–0.59, RBaf + , 0.69–0.79, RTef, 0.43–0.53, RTef + , 0.61–0.78; tibia I (Figs. 40A, 43A–B) generally with 1 regressive seta in series d and tibia (Figs. 40B, 44D) II with one, solenidion φ present on tibia IV, RTi3: 3.2–4; setae l1” of tarsus II not eupathidial. Type material. Holotype, male (MACN-Ar 41112). Argentina, Provincia de Mendoza, Departamento de Las Heras, Cerro Arcos, S32.858206°, W68.938977° (+/- 500 m), elev. 1150 masl (+/- 50 m), 24.Dec.2012, A. Porta leg., Berlese extraction of debris of xeric scrub; 6 paratypes, males (MACN-Ar 41105, 41106, 41113 to 41116), same data, cleared in lactic acid and preserved in alcohol; 7 paratypes, males (MACN-Ar 41107 to 41110, 41118 to 41119, 41121), same data, 29.Mar.2011, mounted on slides; 2 paratypes, females (MACN-Ar 41121, 41122), same data, mounted on slides; 3 paratypes, females (MACN-Ar 41137 to 41139), same data, cleared in lactic acid and preserved in alcohol; 6 paratype tritonymphs (MACN-Ar 41140-41145), same data, mounted on slides; 3 deutonymph paratypes (MACN-Ar 41147 to 41149), same data, 10.May.2013, mounted on slides; 2 larvae paratypes (MACN-Ar 41111, 41117), same data, 10.May.2013, mounted on slides; 2 paratypes, adults (MACN-Ar 41103, 41153), same data, in individual vials, preserved in alcohol; 2 paratypes, adults (CAI 4336, 4337), same data, in individual vials, preserved in alcohol 7 paratypes, 5 adults and 2 larvae, on individual stubs with exemplars mounted for SEM (MACN-Ar 41104), same data; 1 male paratype (MACN-Ar 41120) from Provincia de Mendoza, Mendoza Capital, Reserva de Flora Nativa Bosque Xerófilo, S32.891690°, W68.864855° (+/- 200 m), elev. 850 masl (+/- 5 m), 25.Dic.2012, A. Porta leg., Berlese extraction of debris of xeric scrub; 1 paratype tritonymph (MACN-Ar 41146), same data, mounted on slide; 1 paratype protonymph paratype (MACN-Ar 41150), same data, 08–10.May.2013, mounted on slide; 3 paratypes, adults (MACN-Ar 41150 to 41152), same data, preserved in alcohol; 2 paratypes, adults (MACN-Ar 41124, 41125), same data, 25.Dec.2012, preserved in alcohol. Description Male (n=10). Color (Fig. 31). Gnathosoma and dorsal sclerotized plates on idiosoma dark brown, membranous integument pale cream, legs also dark brown but darker than rest of body. Gnathosoma (Figs. 32, 33, 34A, D, 35A). Covered by aspidosoma in dorsal view. Chelicerae (Fig. 36B) typical for family, 44 wide at base, 144–160 long, movable digit hook-like with 3 minute teeth; cheliceral seta ch slender (Figs. 35B–C), 6 long. Palp (Figs. 36B, 48) 4-segmented, trochanter glabrous; femorogenu with 2 clavate setae in dorsal series, d1 shorter than d; d1 and d 28–34 and 48–60 long, respectively; tibia with prominent calcar d on projecting tubercle, 26–34 long; 3 clavate setae d1, l’ and l”, 28–34, 26–36 and 32–34 long, respectively, plus one more calcar ld, 32–38 long. Palp tarsus subconical, 50–64 long, 10–16 wide at base, solenidion ω present on antiaxial face, 6–7 setae: 3 eupathidia, (u) and l ’, in addition to v, d, l” and, only in some exemplars, l1’ (Figs.36A, 48B–C). Subcapitulum (Fig. 35A) posteriorly rounded, anteriorly subconical, 156–160 long, 198–220 wide at level of palp insertion, setae m 34 long, anterior to n, 30 long, two pairs of adoral setae or 1-2 (Fig. 35B,C), 8 long. Idiosoma (Figs. 32, 33). In dorsal view subtriangular, 822–853 long, 616–671 wide at level of coxa IV. Aspidosoma (Fig. 34A). Subtriangular, 258–275 long, posterior margin 213–221 wide, anterior margin 60–66 wide, projecting horizontally over gnathosoma. Anterior portion with characteristic lateral constriction. Anterior margin straight (Figs. 22, 35A) or with medial incision in some exemplars (Figs. 34A, D). Seta Po (Fig. 34A, D) very long, its length 86–120, expanded distally, with anterior third setulate. Trichobothrial seta bo spatulate, 80–96 long, upturned in distal half and broadening at tip (Fig. 34D). Aspidosomal setation regressive, Pa (Fig. 34B) regressive, subcylindrical, 8 long, located on a transversal ridge; Pm regressive (Fig. 34C), cylindrical, 6 long, inserted on lateral border of aspidosoma, setae Pp absent. Lateral eye plates (Fig. 34A) separated from other plates; anterior pair of lateral eyes 14–22 diameter, posterior pair 22–24. Median eye 42–54 long, 44–50 wide. Hysterosoma. Dorsal view. With 5 dorsal plates D, (L), M, P (Figs. 32, 33). Dorsal plate D subtrapezoidal, 260–295 long, anterior margin 197–209 long, posterior margin 274–295 long, with one longitudinal and two transversal ridges, setae a1, b1, c1, ellipsoidal, lengths: a1: 26–30, b1: 28–30, c1: 26–28; distance between setal insertions a1-a1: 98–106, b1-b1: 54–58, c1-c1: 64–74, a1-b1: 102–106, b1-c1: 108–120. Paired lateral plates L 320–336 long, 72–80 wide; a2 regressive, 9 long; b2 absent and c2 present with p=0.5; lyrifissure ia located on external side of plate, 26–30 long; lyrifissure im located on median line of plate, 12–18 long. Median plates M fused, with d1 and d2 36–38 and 29–36 long, respectively; distance between setal insertions d1-d1: 96–120, d1-d2: 130–146, d2-d2: 367–371. Posterior plates P divided by ridges, setae e1, e2 and e2’, 46–48, 38–44, and 38–44 long, respectively; in some exemplars e2 ” present, setae e2 and e2’ inserted in different sectors of plate, separated by a ridge; distance between setal insertions e1-e1: 100–126, e1-e2: 86–96. Seta hs on posterior border of idiosoma, 40–42 long. Ventral view, podosoma and opisthosoma (Figs. 37, 38). Membranous integument striate. Coxal setation formula: 4:2–3:2–3:2, coxal setae clavate. Measurements of coxal setae, 1a: 20–26; 1b: 30–32; 1c: 32–36; 1d: 36–46; 2a: 18–22; 2b: 22–23; 2c: 16; 3a: 16–22; 3b: 16–22; 3c: 26; 4a: 20–22; 4b: 20–21; 4c: 16–22. Aggenital plates (Fig. 31B) poorly sclerotized, triangular, 72–80 long, 66–80 wide and not differentiable from integument in exemplars cleared in lactic acid. Genital opening 150–156 long, aggenital plates 34–42 wide, with 6 pairs of genital setae, 20– 24 long. Anal opening 136–146 long, adanal plates 32–34 wide, with two pairs of adanal clavate setae, 8–12 long. Pseudanal plate 164–170 long, 64–68 wide, with three pairs of setae, ps1 clavate, 34–36 long, normally developed, ps2 and ps3 regressive, ps2 10 long, ps3 8–9 long, membranous integument with 9 pairs of setae. Internal genitalia (Fig. 39) with sclerites as in other species of the group, relatively large for Caeculidae in relation to idiosoma width, 130–140 wide at level of anterior ventral arch, 90–116 long. Ventral side (Fig. 39A) with 9 pairs of setae, 3 pairs anterior and medial, 2 pairs inserted on sclerites on each side, and 2 posterior pairs, all setae simple and unbranched, relatively long for the family in relation to the size of genital sclerite (cfr. Otto, 1993, Fig. 8; Fuangarworn & Butcher, 2015, Fig. 14; Ott & Ott, 2018, Fig. 4). Two laterodorsal apophyses clearly visible from dorsal side (Fig. 39B). Legs (Figs. 40–47). Measurements see Table 9. Most setae inserted on big cuticular tubercles. Leg I (Figs. 40A, 41–43). Trochanter (Figs. 40A, 42A) with one seta in series l’ and one regressive seta in d. Basifemur (Figs. 40A, 42B) with one seta in v’, pedicelated, inserted in a long cuticular process, shorter than in other species of the group, 54–60, 70–84 + long, RBaf: 0.56–0.59, RBaf + : 0.69–0.79, setae d, l”, v” and v”1 regressive, l’ petiolate. Telofemur (Figs. 40A, 42B) with seta v’ subequal to v’ in telofemur, but more curved, 46–50, 70–76 + long, RTef: 0.43–0.53, RTef +: 0.61–0.78; setae d, l’, l” and v” regressive. Genu (Figs. 42C–D) with 3 regressive setae in series d, 1 and 3 petiolate setae in l’ and l” series, respectively, l’, 46–48, 56–70 + long, 3 petiolate setae in v’ series with v’1 medially displaced, v’, 42–44, 50–64 + long, v2’, 40–46, 56–64 + long; 2 petiolate setae in v” series. Tibia (Figs. 39A, 42D, 43A–B, E) usually with only 1 regressive seta in series d, excepting 2 exemplars asymmetrically with 2 (Fig. 40A); 4 pedunculated setae in each lateral series, 4 spinous setae with blunt tip in ventral series, (v2) medially displaced, v’, 60–64, 70–74 + long, v1’, 50–54, 60–66 + long v”, 50–52 long, solenidion φ and famulus κ” inserted laterodorsally and distally (Figs. 43B, E). Tarsus (Figs. 41, 42D, 43C–D, F–G) with each of series l’, l”, v’ and v” with 3–4, 3–4, 4, 4 setae, respectively, l1” usually eupathidial with p=11/14 (Fig. 41), with some asymmetry (Fig. 43F), solenidion ω inserted dorsally (Figs. 41, 43G) at level of l1”, simple in structure (Fig 41), famulus ε inserted in antiaxial facies (Fig. 41) at level of v1”; eupathidia: er’, (st) and, putatively, l1”, trichobothria bt 16–20 long, two claws ol’ and ol”, 10 and 28–30 long, respectively. Setal count (solenidia): 2, 6, 5, 12, 18–19(1), 19–21(1). Leg II (Fig. 40B, 44). Trochanter (Fig. 44A) with d regressive and l’ and l” petiolate. Basifemur (Fig. 44C) with d and l” regressive, l’, v”, v1” spatulate. Telofemur (Fig. 44C) with d, l’, v and l” regressive, l’ spatulate normally developed. Genu (Fig. 44B) with 3 regressive setae in series d, 3 and 2 pedunculated setae in series l’ and l”, respectively, 2 setae in series v’, seta v” 58–60 long. Tibia (Fig.44D, F) with 1 regressive seta in series d, 4 and 3 pedunculated setae in series l’ and l”, respectively, 3 spinous setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 44F). Tarsus (Fig. 44E) with 3 setae in series l’, l” and 4 in v’ and v”, solenidion ω inserted dorsally at level of l”, only er’ eupathidial; (st) absent; trichobothria bt 14–20 long. Claws ol’ and ol” length 10–12 and 30–34, respectively. Setal count (solenidia): 3, 5, 5, 11, 14(1), 17(1). Leg III (Figs.45A, 46). No eupathidia present. Trochanter (Figs. 45A, 46A) with d regressive and l” petiolate. Basifemur (Fig. 46B) with d regressive, l’, v’, v” and v1” petiolate. Telofemur (Fig. 46B) with d, l’, v” and l” regressive and v’ petiolate. Genu (Fig. 46B) with 3–4 and 1–2 regressive setae in series d and l’, respectively, and 3 pedunculated setae in series v’ and l”, 2 setae in series v”. Tibia (Fig.46D, F) with 2–4 regressive seta in series d, 4 and 3 pedunculated setae in series l’ and l”, respectively, 5 pointed setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 46G), RTi3: 3.2–4. Tarsus (Fig. 46E) with 1 and 3 setae in series l’, l”, respectively, and 3 in v’ and v”; er” present, er’ absent, trichobothria bt 100–110 long. Claws ol’ and ol”, 10 and 36–44 long, respectively. Setal count (solenidia): 2, 5, 5, 12–14, 16–18(1), 12. Leg IV (Figs.45B and 47). Trochanter (Figs. 45A, 46A) with d and l” petiolate and v’ regressive. Basifemur (Fig. 46B) with v” regressive and l’, v’ spatulate. Telofemur (Fig. 46B) with d, l’ and v” regressive and v’ and l” spatulate. Genu (Fig. 47C) with 2 regressive setae in series d, 2 pedunculated setae in each series v’ and l”, 1 seta in series v”. Tibia (Fig.47D, F–G) with 2–3 regressive seta on tibia series d, 3 pedunculated setae in series l” and 4 pointed setae in both ventral series, solenidion φ inserted as in tibia I (Fig. 47F–G). Tarsus (Fig. 47E) with 1 and 2 setae in series l’ and l”, respectively; 3 setae in series v’ and v”, er” present, er’ absent, trichobothria bt 114–140. Claws ol’ and ol” 9–10 and 50–56 long, respectively. Setal count (solenidia): 3, 3, 5, 7, 13–14(1), 11. Female (n=3). Externally similar to male except in size. Gnathosoma. Chelicera 128–136 long. Palp 4-segmented, with chaetotaxy as in male. Subcapitulum 176–192 wide at level of palp insertion, 150–170 long, setae m anterior to n, two pairs of adoral setae or1-2. Idiosoma. 786–924 long, 572–687 wide at level of coxa IV. Aspidosoma. Subtrapezoidal, 258–274 long, posterior margin 213–266 wide, anterior margin 68–80 wide, projecting horizontally over gnathosoma. Seta Po 100–106 long, trichobothrial seta bo 88–120 long. Aspidosomal setation regressive, Pa and Pm minute, Pp absent; anterior pair of lateral eyes 16–20 diameter, posterior pair 18–22. Median eye 40 long, 48 wide. Hysterosoma. Dorsal view. Dorsal plate D subtrapezoidal, 258–320 long, anterior margin 216–287 long, posterior margin 280–307 long, a1: 22–28, b1: 28–38, c1: 30–38; distance between setal insertions, a1-a1: 100–116, b1-b1: 50–72, c1-c1: 46–76, a1-b1: 102–116, b1-c1: 96–128. Paired plates L 360 long, 60–80 wide; a2 regressive 9 long; b2 and c2 absent, ia and im 30–32 and 16–20 long, respectively. Median plates M fused, with d1, d2, 36–42 and 26–30 long, respectively, distance between setal insertions, d1-d1: 90–130, d2-d2: 347–435, d1-d2: 134–158. Posterior plates P, divided by ridges, setae e1, e2, e2’ and e2 ” , 38–44, 38–46, 36–46 and 36–40 long, respectively. Setae e2’ and e2” separated by a ridge from e2, expression of e2” variable (p=0.5), distance between setal insertions, e1-e1: 110–140, e1-e2: 60–98. Seta hs on posterior border of idiosoma, 34–36 long. Ventral view, podosoma and opisthosoma. Coxal setation formula 4:3:3:2–3. Genital opening 150–162 long, with 6 pairs of genital setae, 13–18 long. Anal opening 140–148 long, adanal plates 40–44 wide, with two pairs of adanal clavate setae, 16–18 long. Pseudanal plate 160–162 long, 56 wide, with three pairs of setae. Legs. Measurements see table 10. Ratios, RBaf: 0.5–0.58; RBaf + : 0.77–0.83; RTef: 0.43–0.51; RTef + : 0.73–0.85. Larva (n=3) (Figs 49–59) Gnathosoma. Covered by aspidosoma in dorsal view (Fig. 53). Chelicera 46 long; cheliceral seta ch present. Palp (Fig. 54) 4-segmented, trochanter glabrous, femorogenu with 1 clavate seta in dorsal series, 9–10.4 long; tibia with a calcar d, 11 long, located on a projecting tubercle, 2 clavate setae l’ and l”, and 1 calcar setae ld; 8–11 long, tarsus subconical, solenidion ω present on antiaxial face, 5 setae present, setae v, l’, and d in addition to 2 eupathidia (u). Subcapitulum (Fig. 54) 90 wide at level of palp insertion, 60 long, setae m 12–13 long, anterior to n, 10 long, two pairs of adoral setae or 1-2. Idiosoma (Figs. 49–50). 264–320 long, 184–224 wide at level of coxa III. Aspidosoma (Figs. 51–52). Subtrapezoidal, 106–108 long, posterior margin 100–106 wide, anterior margin 32–36 wide, projecting horizontally over gnathosoma. Seta Po 38 long, trichobothrial seta bo 42 long (Figs. 51B, 52), Pa and Pm minute, Pp absent. Lateral eye plates (Figs. 49, 50A, 51A) separa

Andocaeculus burmeisteri sp. nov. (Figs 18–30) Material. Holotype, female (MACN-Ar 41087). Argentina, Santa Cruz province, Departamento Deseado, Cerro Pico Truncado, 15 km SW from town, S46.875391°, W68.069625° (+/- 100 m) elev. 164 masl, 03.Nov.2014, under rocks and from debris from xeric scrub in Cerro Pico Truncado, A. Porta leg., hand and Berlese collected, preserved in alcohol; 5 paratypes, females (MACN-Ar 41085, MACN-Ar 41090 to 41095), same data, preserved in alcohol; 2 paratypes, females (MACN-Ar 41088, 41089), same data, mounted on slides; 2 paratypes, adults (MACN-Ar 41086), same data, on 2 individual stubs with exemplars mounted for SEM; 1 paratype, adult (MACN-Ar 41096), same data, voucher of BOLDSYSTEM data SPDAR1609-46, preserved in alcohol; 1 paratype, female (MACN-Ar 41097), Argentina, Provincia de Chubut, Departamento de Sarmiento, 6 km E from Sarmiento town, S45.577516°, W69.089723° (+/- 100 m), elev. 321 masl, 02.Nov.2014, under rocks, A. Porta leg., hand collected. Etymology. The specific name is a patronymic in honor to Karl Hermann Konrad Burmeister (1807–1892), German-Argentinian first director of the MACN, who named the type locality of the species as Cerro Pico Truncado in 1889. Diagnosis. As for the A. weyrauchi species group but differing from the rest of the species by: size and general form of idiosoma, being more ovate (Figs. 19, 20); 4 eupathidia on palp tarsus; anterior portion of aspidosoma characteristic (Figs. 20, 21A), with a depression and a duck-beak appearance and with a characteristic lateral constriction at level of seta Pa; setae of opisthosomal plates smaller (Figs. 19 and 20), for example c1, 20–22 long; setae e2 and e2’ not separated by ridges; setae b2, c2 absent; d2 generally absent (Figs. 19, 20); legs III and IV (Figs. 27, 29) more slender, in particular tibia III (Figs. 27A, 28F) more than six times longer than wide, RTi3: 6.3–7.6; trochanter III (Figs. 27A, 28A arrow) with a characteristic anterior cylindrical protuberance on anterior portion of trochanter; solenidion φ absent on tibia IV. Description. Color (Fig. 18). Gnathosoma and dorsal sclerotized plates on idiosoma dark brown, membranous integument pale cream, legs also dark brown but darker that rest of body. Female (n=8) Gnathosoma. Covered by aspidosoma in dorsal view (Figs. 19–21). Cheliceral typical for family, 130–150 long, movable digit hook-like with 3 minute teeth; cheliceral seta ch slender, 6 long. Palp (Figs. 29B and 30) trochanter glabrous; femorogenu with 2 clavate setae in dorsal series, d1 shorter than d, d1 22–36 long, d 40–50 long; tibia with prominent calcar d on projecting tubercle, 28–36 long, ld and l” calcars, 34–36 and 30–34 long, respectively, l’ and d1 petiolate, 24–30 and 38–46 long, respectively; tarsus subconical, 44–56 long, width at base 20–24, ω solenidion present on antiaxial face, 7 setae, l”, v and l1’ in addition to 4 eupathidia (u), l’ and d. Subcapitulum (Figs. 22–23) posteriorly rounded, anteriorly subconical, 196–212 wide at level of palp insertion, 170–174 long, setae m, anterior to n, 15–19 long, two pairs of adoral setae or 1-2, 10 long. Lateral lips and labrum typical for family. Idiosoma. In dorsal view (Figs. 19–20) subtriangular, 1040–1100 long, 680–720 wide at level of coxa IV. Aspidosoma (Fig. 21). Subtrapezoidal, 316–332 long, posterior margin 269–300 wide, anterior margin, 111– 126 long, straight, and with an anterior depression and with a characteristic lateral constriction at level of seta Pa. Seta Po (Fig. 21C) 98–120 long, with anterior third expanded distally and setulate, dorsally visible from anterior margin of aspidosoma (Fig. 21A). Trichobothrial setae bo, spatulate, 88–100 long, upturned in half distally and broadening at tip (Fig. 21C). Aspidosomal setation regressive, Pa (Fig. 21B) regressive, subcylindrical, 5 long, located on tranversal ridge; Pm regressive, cylindrical, 8 long, setae Pp absent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 20 diameter, posterior pair 22–30 (Figs. 20, 21A). Hysterosoma. Dorsal view. With 5 dorsal plates: D, (L), M, P (Figs. 19–20). Dorsal plate D subtrapezoidal, 312–332 long, anterior margin 256–300 long, posterior margin 336–371 long with one longitudinal and two transversal ridges, setae a1, b1 and c1 very small, measurements a1: 10–14, b1: 20–26, c1: 20–22; distance between setal insertions, a1-a1: 108–124, b1-b1: 84, c1-c1: 66–70, a1-b1: 112–140, b1-c1: 130–142. Paired plates L 376–408 long, 78–84 wide; a2 regressive, 10–14 long; b2 and c2 absent, ia, 30–34 long at same level of b1, in antiaxial side of the plate; im at level of c 1 in medial line of the plate, 12–20 long. Median plate M fused, d1: 28–32, d2 generally absent, distance d1-d1: 134–154. Posterior plates P without ridges, setal measurements, e1: 26–36; e2: 18–26; e2’ : 16–22 and e2”: 20; distance between setal insertions, e1-e1: 134–146, e1-e2: 92–116. Seta hs on posterior border of idiosoma, 26–30 long. Ventral view, podosoma and opisthosoma (Figs. 22, 23). Membranous integument striate. Coxal setation formula: 4:3–4:4:3, coxal setae clavate. Measurements of coxal setae, 1a: 16–24; 1b: 18–26; 1c: 36–44; 1d: 40–46; 2a: 22; 2b: 18–26; 2c: 26–28; 3a: 14–18; 3b: 20–26; 3c: 26; 4a: 22; 4b: 16–22; 4c: 20–24. Genital opening 146–150 long, 50–60 wide, with 7 pairs of genital setae, 10–18 long. Anal opening 190–192 long, 92–112 wide with two pairs of adanal setae, clavate, 15–16 long. Pseudanal plate, 80–100 long, 42 wide, with three pairs of setae, ps1; ps3; ps3, 16–20; 18–24 and 18–24 long, respectively; membranous integument with 10–11 pairs of setae. Legs (Figs. 24–29A). Measurements see Table 8. Leg I. Trochanter (Fig. 24A, 25A) with only one seta in l’, 18–20 long, and one regressive seta in d series. Basifemur (Figs. 24A, 25B) with setae on v’ spatulate, 66–76, 100–110 + long, RBaf: 0.62–0.86, RBaf + : 0.94–1.13, setae d, l”, v” and v”1 regressive, l’ petiolate. Telofemur (Figs. 24A, 25B) with seta v’ subequal, in size and shape, to v’ on basifemur, 66–80, 100–114 + long, RTef: 0.61–0.8, RTef +: 0.92–1.18, setae l’, l”, v’ and v” regressive. Genu and tibia (Figs. 24A, 25 C-E) with 3 and 2 regressive setae in series d, respectively; setae of series v’ and v” pointed. Genu (Fig. 25C–D) with 3 petiolate setae in l” series, one very elongated spatulate setae in l’ series, 64–70, 80–94 + long, 3 setae in v’ series (Figs. 24A, 25D) with v1’ medially displaced, v’, 88–106, 106–132 + long, v1’, 52–66, v2’, 72–84, 94–114 + two setae in v” series. Tibia (Figs. 24A, 25E) with 4 spatulate setae in in both lateral series, 3 subequal pointed setae on v’ series, 88–104, 108–124 + long and 2 setae in v”, 40–59 long, solenidion φ and famulus κ” (Fig. 25G) inserted laterodorsally on distal portion of tibia. Tarsus (Figs. 25H–I, 29A) with each of series l’, l”, v’ and v” with 3–4, 3–4, 4, 4 setae, respectively, trichobothrial seta bt 28–32 long; solenidion ω inserted dorsally (Fig. 25H–I) at level of l’, simple in structure (Fig. 29A). Famulus inserted in antiaxial facies (Fig. 25F) at level of l1” (Fig. 29A) with only 4 eupathidia: l1”, er’ and (st). Two claws ol’ and ol” of lengths 8–10 and 22–30, respectively. Setal count (solenidia): 2, 5, 5 13, 16(1), 20–22(1). Leg II (Fig. 24B and 26). Setae in series l’ spatulate. Trochanter (Fig. 26A) with regressive seta in series d and with 1 seta in each lateral series. Basifemur and telofemur (Fig. 26B) each with a long and pointed seta v”, 70–82 long on basifemur; this segment with l’ and v” spatulate and one regressive seta in d, telofemur with l’ and v” spatulate, and v’1 and d regressive. Genu with 3 regressive seta in series d (Figs. 24B and 26F) and 2 in tibia d (Figs. 24B and 26C). Genu with 4 and 3 pedunculated setae in series l’ and l”, respectively, 2 pointed setae in series v’ and one very large pointed seta in series v”, 70–100 long. Tibia (Fig. 26C) with 4 setae in each lateral series and 3 on in each ventral series, solenidion inserted as on tibia I. Tarsus (Figs.26D–E, G) with 4 setae on in series l’; v’ and v”, and 3 on l”; solenidion ω inserted dorsally at a level between l1” and l’ (Figs. 26G), famulus inserted in antiaxial facies at level of l1” (Figs. 26D–E); trichobothria (Fig.26E, G) 30–38 long. Two eupathidia: l1” and er’. Claws ol’ and ol” lengths 6–8 and 20–26, respectively. Setal count (solenidia): 3, 5, 5, 13, 16(1), 17(1). Leg III (Figs.27A and 28). Setae of series v’ spatulate and elongated, eupathidia absent. Trochanter (Figs. 27A, 28A) with one characteristic cylindrical protuberance on anterior facies (Fig. 28A, arrow), v’ and d* present. Basifemur (Figs. 27A, 28B) with one seta v’ and 3 regressive l”, v” and v1”. Telofemur (Fig. 28B) with one seta in v’ and 4 regressive: l”, v”, d and l’. Genu (Figs. 27A, 28C) with 3–4 regressive setae in series d, one in l’ and 3 in v” and 3 setae in series l” and v’. Tibia (Figs. 27A, 28F) very elongated, ratio length/width at least 6 (RTi3: 6.3–7.6), with 2–3 regressive setae in series d, solenidion φ present, 4 setae in l” and 5 setae in both ventral series. Tarsus (Figs. 28D–E) with 3–4, 2, 4, 4 in series l’, l”, v’ and v”, respectively; trichobothria bt 106–124 long, er” present, er’ absent, claws ol’ and ol” of length 6–10 and 24–28, respectively. Setal count: 2, 4, 5, 13–14, 17(1), 15–16. Leg IV (Fig. 27B). Setae of series v’ as on leg III. Trochanter with one d and one l”. Basifemur with one seta each of l”, v’ and v”. Telofemur with 1 regressive seta in l’ and d, and 1 on each of v’, v” and l”. Genu with 2 setae in series l”, 2 long petiolate setae in series v’, 2 regressive in series d and 1 in series v”. Tibia with 3 regressive setae each in l” and d series, 3 petiolate in series v’ and 4 pointed in series v”, solenidion φ absent. Tarsus with 3 and 4 setae in each ventral series v ’ and v”, respectively; 1 and 1–2 petiolate setae in series l” and l’, respectively, er” present and er’ absent, trichobothria bt 114–136 long, claws ol’ and ol” lengths 10–12 and 36–46, respectively. Setal count: 2, 3, 5, 8, 13, 11.

Andocaeculus burmeisteri sp. nov. (Figs 18–30) Material. Holotype, female (MACN-Ar 41087). Argentina, Santa Cruz province, Departamento Deseado, Cerro Pico Truncado, 15 km SW from town, S46.875391°, W68.069625° (+/- 100 m) elev. 164 masl, 03.Nov.2014, under rocks and from debris from xeric scrub in Cerro Pico Truncado, A. Porta leg., hand and Berlese collected, preserved in alcohol; 5 paratypes, females (MACN-Ar 41085, MACN-Ar 41090 to 41095), same data, preserved in alcohol; 2 paratypes, females (MACN-Ar 41088, 41089), same data, mounted on slides; 2 paratypes, adults (MACN-Ar 41086), same data, on 2 individual stubs with exemplars mounted for SEM; 1 paratype, adult (MACN-Ar 41096), same data, voucher of BOLDSYSTEM data SPDAR1609-46, preserved in alcohol; 1 paratype, female (MACN-Ar 41097), Argentina, Provincia de Chubut, Departamento de Sarmiento, 6 km E from Sarmiento town, S45.577516°, W69.089723° (+/- 100 m), elev. 321 masl, 02.Nov.2014, under rocks, A. Porta leg., hand collected. Etymology. The specific name is a patronymic in honor to Karl Hermann Konrad Burmeister (1807–1892), German-Argentinian first director of the MACN, who named the type locality of the species as Cerro Pico Truncado in 1889. Diagnosis. As for the A. weyrauchi species group but differing from the rest of the species by: size and general form of idiosoma, being more ovate (Figs. 19, 20); 4 eupathidia on palp tarsus; anterior portion of aspidosoma characteristic (Figs. 20, 21A), with a depression and a duck-beak appearance and with a characteristic lateral constriction at level of seta Pa; setae of opisthosomal plates smaller (Figs. 19 and 20), for example c1, 20–22 long; setae e2 and e2’ not separated by ridges; setae b2, c2 absent; d2 generally absent (Figs. 19, 20); legs III and IV (Figs. 27, 29) more slender, in particular tibia III (Figs. 27A, 28F) more than six times longer than wide, RTi3: 6.3–7.6; trochanter III (Figs. 27A, 28A arrow) with a characteristic anterior cylindrical protuberance on anterior portion of trochanter; solenidion φ absent on tibia IV. Description. Color (Fig. 18). Gnathosoma and dorsal sclerotized plates on idiosoma dark brown, membranous integument pale cream, legs also dark brown but darker that rest of body. Female (n=8) Gnathosoma. Covered by aspidosoma in dorsal view (Figs. 19–21). Cheliceral typical for family, 130–150 long, movable digit hook-like with 3 minute teeth; cheliceral seta ch slender, 6 long. Palp (Figs. 29B and 30) trochanter glabrous; femorogenu with 2 clavate setae in dorsal series, d1 shorter than d, d1 22–36 long, d 40–50 long; tibia with prominent calcar d on projecting tubercle, 28–36 long, ld and l” calcars, 34–36 and 30–34 long, respectively, l’ and d1 petiolate, 24–30 and 38–46 long, respectively; tarsus subconical, 44–56 long, width at base 20–24, ω solenidion present on antiaxial face, 7 setae, l”, v and l1’ in addition to 4 eupathidia (u), l’ and d. Subcapitulum (Figs. 22–23) posteriorly rounded, anteriorly subconical, 196–212 wide at level of palp insertion, 170–174 long, setae m, anterior to n, 15–19 long, two pairs of adoral setae or 1-2, 10 long. Lateral lips and labrum typical for family. Idiosoma. In dorsal view (Figs. 19–20) subtriangular, 1040–1100 long, 680–720 wide at level of coxa IV. Aspidosoma (Fig. 21). Subtrapezoidal, 316–332 long, posterior margin 269–300 wide, anterior margin, 111– 126 long, straight, and with an anterior depression and with a characteristic lateral constriction at level of seta Pa. Seta Po (Fig. 21C) 98–120 long, with anterior third expanded distally and setulate, dorsally visible from anterior margin of aspidosoma (Fig. 21A). Trichobothrial setae bo, spatulate, 88–100 long, upturned in half distally and broadening at tip (Fig. 21C). Aspidosomal setation regressive, Pa (Fig. 21B) regressive, subcylindrical, 5 long, located on tranversal ridge; Pm regressive, cylindrical, 8 long, setae Pp absent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 20 diameter, posterior pair 22–30 (Figs. 20, 21A). Hysterosoma. Dorsal view. With 5 dorsal plates: D, (L), M, P (Figs. 19–20). Dorsal plate D subtrapezoidal, 312–332 long, anterior margin 256–300 long, posterior margin 336–371 long with one longitudinal and two transversal ridges, setae a1, b1 and c1 very small, measurements a1: 10–14, b1: 20–26, c1: 20–22; distance between setal insertions, a1-a1: 108–124, b1-b1: 84, c1-c1: 66–70, a1-b1: 112–140, b1-c1: 130–142. Paired plates L 376–408 long, 78–84 wide; a2 regressive, 10–14 long; b2 and c2 absent, ia, 30–34 long at same level of b1, in antiaxial side of the plate; im at level of c 1 in medial line of the plate, 12–20 long. Median plate M fused, d1: 28–32, d2 generally absent, distance d1-d1: 134–154. Posterior plates P without ridges, setal measurements, e1: 26–36; e2: 18–26; e2’ : 16–22 and e2”: 20; distance between setal insertions, e1-e1: 134–146, e1-e2: 92–116. Seta hs on posterior border of idiosoma, 26–30 long. Ventral view, podosoma and opisthosoma (Figs. 22, 23). Membranous integument striate. Coxal setation formula: 4:3–4:4:3, coxal setae clavate. Measurements of coxal setae, 1a: 16–24; 1b: 18–26; 1c: 36–44; 1d: 40–46; 2a: 22; 2b: 18–26; 2c: 26–28; 3a: 14–18; 3b: 20–26; 3c: 26; 4a: 22; 4b: 16–22; 4c: 20–24. Genital opening 146–150 long, 50–60 wide, with 7 pairs of genital setae, 10–18 long. Anal opening 190–192 long, 92–112 wide with two pairs of adanal setae, clavate, 15–16 long. Pseudanal plate, 80–100 long, 42 wide, with three pairs of setae, ps1; ps3; ps3, 16–20; 18–24 and 18–24 long, respectively; membranous integument with 10–11 pairs of setae. Legs (Figs. 24–29A). Measurements see Table 8. Leg I. Trochanter (Fig. 24A, 25A) with only one seta in l’, 18–20 long, and one regressive seta in d series. Basifemur (Figs. 24A, 25B) with setae on v’ spatulate, 66–76, 100–110 + long, RBaf: 0.62–0.86, RBaf + : 0.94–1.13, setae d, l”, v” and v”1 regressive, l’ petiolate. Telofemur (Figs. 24A, 25B) with seta v’ subequal, in size and shape, to v’ on basifemur, 66–80, 100–114 + long, RTef: 0.61–0.8, RTef +: 0.92–1.18, setae l’, l”, v’ and v” regressive. Genu and tibia (Figs. 24A, 25 C-E) with 3 and 2 regressive setae in series d, respectively; setae of series v’ and v” pointed. Genu (Fig. 25C–D) with 3 petiolate setae in l” series, one very elongated spatulate setae in l’ series, 64–70, 80–94 + long, 3 setae in v’ series (Figs. 24A, 25D) with v1’ medially displaced, v’, 88–106, 106–132 + long, v1’, 52–66, v2’, 72–84, 94–114 + two setae in v” series. Tibia (Figs. 24A, 25E) with 4 spatulate setae in in both lateral series, 3 subequal pointed setae on v’ series, 88–104, 108–124 + long and 2 setae in v”, 40–59 long, solenidion φ and famulus κ” (Fig. 25G) inserted laterodorsally on distal portion of tibia. Tarsus (Figs. 25H–I, 29A) with each of series l’, l”, v’ and v” with 3–4, 3–4, 4, 4 setae, respectively, trichobothrial seta bt 28–32 long; solenidion ω inserted dorsally (Fig. 25H–I) at level of l’, simple in structure (Fig. 29A). Famulus inserted in antiaxial facies (Fig. 25F) at level of l1” (Fig. 29A) with only 4 eupathidia: l1”, er’ and (st). Two claws ol’ and ol” of lengths 8–10 and 22–30, respectively. Setal count (solenidia): 2, 5, 5 13, 16(1), 20–22(1). Leg II (Fig. 24B and 26). Setae in series l’ spatulate. Trochanter (Fig. 26A) with regressive seta in series d and with 1 seta in each lateral series. Basifemur and telofemur (Fig. 26B) each with a long and pointed seta v”, 70–82 long on basifemur; this segment with l’ and v” spatulate and one regressive seta in d, telofemur with l’ and v” spatulate, and v’1 and d regressive. Genu with 3 regressive seta in series d (Figs. 24B and 26F) and 2 in tibia d (Figs. 24B and 26C). Genu with 4 and 3 pedunculated setae in series l’ and l”, respectively, 2 pointed setae in series v’ and one very large pointed seta in series v”, 70–100 long. Tibia (Fig. 26C) with 4 setae in each lateral series and 3 on in each ventral series, solenidion inserted as on tibia I. Tarsus (Figs.26D–E, G) with 4 setae on in series l’; v’ and v”, and 3 on l”; solenidion ω inserted dorsally at a level between l1” and l’ (Figs. 26G), famulus inserted in antiaxial facies at level of l1” (Figs. 26D–E); trichobothria (Fig.26E, G) 30–38 long. Two eupathidia: l1” and er’. Claws ol’ and ol” lengths 6–8 and 20–26, respectively. Setal count (solenidia): 3, 5, 5, 13, 16(1), 17(1). Leg III (Figs.27A and 28). Setae of series v’ spatulate and elongated, eupathidia absent. Trochanter (Figs. 27A, 28A) with one characteristic cylindrical protuberance on anterior facies (Fig. 28A, arrow), v’ and d* present. Basifemur (Figs. 27A, 28B) with one seta v’ and 3 regressive l”, v” and v1”. Telofemur (Fig. 28B) with one seta in v’ and 4 regressive: l”, v”, d and l’. Genu (Figs. 27A, 28C) with 3–4 regressive setae in series d, one in l’ and 3 in v” and 3 setae in series l” and v’. Tibia (Figs. 27A, 28F) very elongated, ratio length/width at least 6 (RTi3: 6.3–7.6), with 2–3 regressive setae in series d, solenidion φ present, 4 setae in l” and 5 setae in both ventral series. Tarsus (Figs. 28D–E) with 3–4, 2, 4, 4 in series l’, l”, v’ and v”, respectively; trichobothria bt 106–124 long, er” present, er’ absent, claws ol’ and ol” of length 6–10 and 24–28, respectively. Setal count: 2, 4, 5, 13–14, 17(1), 15–16. Leg IV (Fig. 27B). Setae of series v’ as on leg III. Trochanter with one d and one l”. Basifemur with one seta each of l”, v’ and v”. Telofemur with 1 regressive seta in l’ and d, and 1 on each of v’, v” and l”. Genu with 2 setae in series l”, 2 long petiolate setae in series v’, 2 regressive in series d and 1 in series v”. Tibia with 3 regressive setae each in l” and d series, 3 petiolate in series v’ and 4 pointed in series v”, solenidion φ absent. Tarsus with 3 and 4 setae in each ventral series v ’ and v”, respectively; 1 and 1–2 petiolate setae in series l” and l’, respectively, er” present and er’ absent, trichobothria bt 114–136 long, claws ol’ and ol” lengths 10–12 and 36–46, respectively. Setal count: 2, 3, 5, 8, 13, 11., Published as part of Porta, Andrés O., Pizarro-Araya, Jaime & Ramírez, Martín J., 2021, Revision and phylogeny of the genus Andocaeculus (Acari: Caeculidae) I: the A. weyrauchi species group, pp. 1-78 in Zootaxa 4945 (1) on pages 28-41, DOI: 10.11646/zootaxa.4945.1.1, http://zenodo.org/record/4614242