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Understanding the severity and extent of near surface critical zone (CZ) disturbances and their ecosystem response is a pressing concern in the face of increasing human and natural disturbances. Predicting disturbance severity and recovery in a changing climate requires comprehensive understanding of ecosystem feedbacks among vegetation and the surrounding environment, including climate, hydrology, geomorphology, and biogeochemistry. Field surveys and satellite remote sensing have limited ability to effectively capture the spatial and temporal variability of disturbance and CZ properties. Technological advances in remote sensing using new sensors and new platforms have improved observations of changes in vegetation canopy structure and productivity; however, integrating measures of forest disturbance from various sensing platforms is complex. By connecting the potential for remote sensing technologies to observe different CZ disturbance vectors, we show that lower severity disturbance and slower vegetation recovery are more difficult to quantify. Case studies in montane forests from the western United States highlight new opportunities, including evaluating post‐disturbance forest recovery at multiple scales, shedding light on understory vegetation regrowth, detecting specific physiological responses, and refining ecohydrological modeling. Learning from regional CZ disturbance case studies, we propose future directions to synthesize fragmented findings with (a) new data analysis using new or existing sensors, (b) data fusion across multiple sensors and platforms, (c) increasing the value of ground‐based observations, (d) disturbance modeling, and (e) synthesis to improve understanding of disturbance. Key Points: Increasing remote sensing capabilities improve observations of disturbance severity and recovery in varying critical zone settingsCase studies present how new approaches from multiple platforms and sensors better quantify post‐disturbance recovery of montane forestsWe suggest integrating the new approaches to refine remote sensing capabilities and resolve critical zone processes at multiple scales [ABSTRACT FROM AUTHOR]

Understanding patterns of species coexistence is a fundamental challenge in ecology. The physical environment is believed to play an important role, influencing patterns of dispersal and biotic interactions across space and time. Floodplain forest species are presumed to interact strongly with their environment, as evidenced by pronounced spatial variation in forest composition associated with flood‐driven abiotic constraints. Questions of how, and to what degree, fine‐scale heterogeneity interacts with broad‐scale hydrogeomorphology to influence patterns of functional composition remain unresolved. We examined how functional diversity (i.e. richness and range of functional traits) and ecological strategies (i.e. functional trait combinations) of floodplain forest communities varied across regional and local gradients of flooding in a northern temperate region of the United States of America. We found functional diversity of woody overstory species varied across hydrogeomorphic settings and that different settings altered associations between functional diversity and both the relative elevation above and proximity to rivers. Ecological strategies shifted over local gradients of relative elevation and distance to channel with different magnitudes and directions depending on the broader hydrogeomorphic context. We found evidence that interactions among flood regimes and landform position impose different levels of functional constraint. These results indicate patterns of community assembly are not easily discerned from landform type or position alone, but rather from filtering that operates and interacts with biota over multiple spatial scales. Our results imply that it is important to characterize flood dynamics in ways that can be clearly linked to ecological processes and that treatment of floodplain landforms as transferable units across river‐valley segments is problematic, even within a single basin. [ABSTRACT FROM AUTHOR]

Mounting evidence suggests that climate change will cause shifts of tree species range and abundance (biomass). Abundance changes under climate change are likely to occur prior to a detectable range shift. Disturbances are expected to directly affect tree species abundance and composition, and could profoundly influence tree species spatial distribution within a geographical region. However, how multiple disturbance regimes will interact with changing climate to alter the spatial distribution of species abundance remains unclear. We simulated such forest demographic processes using a forest landscape succession and disturbance model (LANDIS-II) parameterized with forest inventory data in the northeastern United States. Our study incorporated climate change under a high-emission future and disturbance regimes varying with gradients of intensities and spatial extents. The results suggest that disturbances catalyze changes in tree species abundance and composition under a changing climate, but the effects of disturbances differ by intensity and extent. Moderate disturbances and large extent disturbances have limited effects, while high-intensity disturbances accelerate changes by removing cohorts of mid-and late-successional species, creating opportunities for early-successional species. High-intensity disturbances result in the northern movement of early-successional species and the southern movement of late-successional species abundances. Our study is among the first to systematically investigate how disturbance extent and intensity interact to determine the spatial distribution of changes in species abundance and forest composition. [ABSTRACT FROM AUTHOR]

Forests are currently a substantial carbon sink globally. Many climate change mitigation strategies leverage forest preservation and expansion, but rely on forests storing carbon for decades to centuries. Yet climate‐driven disturbances pose critical risks to the long‐term stability of forest carbon. We quantify the climate drivers that influence wildfire and climate stress‐driven tree mortality, including a separate insect‐driven tree mortality, for the contiguous United States for current (1984–2018) and project these future disturbance risks over the 21st century. We find that current risks are widespread and projected to increase across different emissions scenarios by a factor of >4 for fire and >1.3 for climate‐stress mortality. These forest disturbance risks highlight pervasive climate‐sensitive disturbance impacts on US forests and raise questions about the risk management approach taken by forest carbon offset policies. Our results provide US‐wide risk maps of key climate‐sensitive disturbances for improving carbon cycle modeling, conservation and climate policy. [ABSTRACT FROM AUTHOR]

On a warming planet, a key challenge natural resource managers face is protecting wildlife while mitigating climate change—as through forest carbon storage—to the greatest extent possible. But in some ecosystems, habitat restoration for imperiled species may be incompatible with maximizing carbon storage. For example, promoting early successional forest conditions does not maximize stand‐level carbon storage, whereas uniformly promoting high stocking or mature forest conditions in the name of carbon storage excludes species that require open or young stands. Here, we briefly review the literature regarding carbon and wildlife trade‐offs and then explore four case studies from the Northern Forest region of the United States. In each case, human activities have largely dampened the influence of natural disturbances; restoring or emulating these disturbances is typically required for habitat restoration even when doing so equates to less carbon storage at the stand level. We propose that applying a climate adaptation lens can help managers and planners navigate these trade‐offs and steer away from maladaptive practices that may ultimately reduce adaptive capacity. Instead, critically evaluating the consequences of stand‐level management actions on both carbon and wildlife can then facilitate landscape‐scale climate adaptation planning that supports a diversity of habitats alongside opportunities to invest in maximizing forest carbon. [ABSTRACT FROM AUTHOR]

In the context of widespread ecological changes, land managers and policymakers confront the need to prioritize ecosystem restoration and fuel management activities across large areas to sustain ecosystem services. Reference conditions inform prioritization efforts by providing a baseline from which to measure where and how vegetation and fuels have changed, but until recently the USA lacked a complete set of reference conditions. We describe the ongoing development of a comprehensive set of vegetation reference conditions based on over 900 quantitative vegetation dynamic models and accompanying description documents for terrestrial ecosystems in the USA. These models and description documents, collaboratively developed by more than 800 experts around the country through the interagency LANDFIRE Program, synthesize fundamental ecological information about ecosystem dynamics, structure, composition, and disturbance regimes before European‐American settlement. These products establish the first comprehensive national baseline for measuring vegetation change in the USA, providing land managers and policymakers with a tool to support vegetation restoration and fuel management activities at regional to national scales. Users have applied these products to support a variety of land management needs including exploring ecosystem dynamics, assessing current and desired conditions, and simulating the effects of management actions. In an era of rapid ecological change, these products provide land managers with an adaptable tool for understanding ecosystems and predicting possible future conditions. [ABSTRACT FROM AUTHOR]

Many forests in dry mountain regions are characterized by a lower elevational treeline. Understanding the controls on the position of lower treeline is important for predicting future forest distributional shifts in response to global environmental change. Lower treelines currently at their climate limit are expected to be more sensitive to changing climate, whereas lower treelines constrained by non‐climatic factors are less likely to respond directly to climate change but may be sensitive to other global change agents. In this study, we used existing vegetation classifications to map lower treelines for our 1.7 million km2 study region in the U. S. Intermountain West. We modeled topoclimatic drivers of lower treeline position for each of three dominant forest types to identify topoclimatically limited treelines. We then used spatial data of edaphic properties, recent fire, and land use to identify lower treelines potentially constrained above their ecophysiological limits by non‐climatic processes. We found that the lower treeline ecotone of pinyon–juniper woodlands is largely limited by topoclimate and is likely to be sensitive to increasing temperatures and associated droughts, though these effects may be heterogeneously distributed across the landscape. In contrast, dry mixed‐conifer lower treelines in the northern portion of the study area rarely reached their modeled topoclimatic limit, suggesting that non‐climatic processes, including fire and land use, constrain the lower treeline above its ecophysiological limits in this forest type. Our results suggest that much of the lower treeline in the Intermountain West is currently climate limited and will thus be sensitive to ongoing climate changes. Lower treelines in other arid or semi‐arid mountainous regions around the globe may also be strongly sensitive to climate, though treeline response to climate change will be mediated at the local scale by soil properties, biotic interactions, and natural or anthropogenic disturbances. Our regional study of lower treeline provides a framework for identifying the drivers of lower treeline formation and allows for more robust projections of future treeline dynamics, which are needed to anticipate shifting global distributions of the forest biome. [ABSTRACT FROM AUTHOR]

Despite widespread interest in drought legacies—multiyear impacts of drought on tree growth—the key implication of reported drought legacies remains unaddressed: as impaired growth and slow recovery associated with drought legacies are pervasive across forest ecosystems, what is the impact of more frequent drought conditions? We investigated the assumption that either multiple drought years occurring during a short period (multiyear droughts), or droughts occurring during the recovery period from previous drought (compounded droughts), are detrimental to subsequent growth. There is evidence that drought responses may vary among populations of widespread species, leading us to examine regional differences in responses of the conifer Pinus ponderosa to historic drought frequency in the western United States. More frequent drought conditions incurred additional growth declines and shifts in growth–climate sensitivities in the years following drought relative to single‐drought events, with 'triple‐droughts' being worse than 'double‐droughts'. Notably, prediction skill was not strongly reduced when ignoring compounded droughts, a consequence of the temporally comprehensive formulation of our stochastic antecedent model that accounts for the climatic memory of tree growth. We argue that incorporating drought‐induced temporal variability in tree growth sensitivities can aid inference gained from statistical models, where more simplistic models could overestimate the severity of drought legacies. We also found regional differences in response to repeated drought, and suggest plastic post‐drought sensitivities and climatic memory may represent beneficial physiological adjustments in interior regions. Within‐species variability may thus mediate forest responses to increasing drought frequency under future climate change, but experimental approaches using more species are necessary to improve our understanding of the mechanisms that underlie drought legacy effects on tree growth. [ABSTRACT FROM AUTHOR]

Tree harvest and climate change can interact to have synergistic effects on tree species distribution changes. However, few studies have investigated the interactive effects of tree harvest and climate change on tree species distributions.We assessed the interactive effects of tree harvest and climate change on the distribution of 29 dominant tree species at 270 m resolution in the southern United States, while accounting for species demography, competition, urban growth and natural fire. We simulated tree species distribution changes to year 2100 using a coupled forest dynamic model (LANDIS PRO), ecosystem process model (LINKAGES) and urban growth model (SLEUTH).The distributions of 20 tree species contracted and nine species expanded within the region under climate change by end of 21st century. Distribution changes for all tree species were very slow and lagged behind the changes in potential distributions that were in equilibrium with new climatic conditions.Tree harvest and climate change interacted to affect species occurrences and colonization but not extinction. Occurrence and colonization were mainly affected by tree harvest and its interaction with climate change while extinctions were mainly affected by tree harvest and climate change.Synthesis and applications. Interactive effects of climate and tree harvest acted in the same direction as climate change effects on species occurrences, thereby accelerating climate change induced contraction or expansion of distributions. The overall interactive effects on species colonization were negative, specifically with positive interactive effects at leading edges of species ranges and negative interactive effects at trailing edges. Tree harvest generally did not interact with climate change to greatly facilitate or ameliorate species extinction. Our modelling results highlight the importance of considering disturbances and species demography (e.g. post‐harvest regeneration dynamics) when predicting changes in tree distributions. [ABSTRACT FROM AUTHOR]

Aim: Climate and disturbance alter forest dynamics, from individual trees to biomes and from years to millennia, leaving legacies that vary with local, meso‐ and macroscales. Motivated by recent insights in temperate forests, we argue that temporal and spatial extents equivalent to that of the underlying drivers are necessary to characterize forest dynamics across scales. We focus specifically on characterizing mesoscale forest dynamics because they bridge fine‐scale (local) processes and the continental scale (macrosystems) in ways that are highly relevant for climate change science and ecosystem management. We revisit ecological concepts related to spatial and temporal scales and discuss approaches to gain a better understanding of climate–forest dynamics across scales. Location: Eastern USA. Time period: Last century to present. Major taxa studied: Temperate broadleaf forests. Methods: We review regional literature of past tree mortality studies associated with climate to identify mesoscale climate‐driven disturbance events. Using a dynamic vegetation model, we then simulate how these forests respond to a typical climate‐driven disturbance. Results: By identifying compound disturbance events from both a literature review and simulation modelling, we find that synchronous patterns of drought‐driven mortality at mesoscales have been overlooked within these forests. Main conclusions: As ecologists, land managers and policy‐makers consider the intertwined drivers of climate and disturbance, a focus on spatio‐temporal scales equivalent to those of the drivers will provide insight into long‐term forest change, such as drought impacts. Spatially extensive studies should also have a long temporal scale to provide insight into pathways for forest change, evaluate predictions from dynamic forest models and inform development of global vegetation models. We recommend integrating data collected from spatially well‐replicated networks (e.g., archaeological, historical or palaeoecological data), consisting of centuries‐long, high‐resolution records, with models to characterize better the mesoscale response of forests to climate change in the past and in the future. [ABSTRACT FROM AUTHOR]

Biotic disturbance agents such as insects can be highly responsive to climatic change and have widespread ecological and economic impacts on forests. Quantifying the responses of introduced and native insects to climate, including how dynamics of one agent may mediate those of another, is important for forecasting disturbance and associated impacts on forest structure and function. We investigated drivers of outbreaks by larch casebearer Coleophora laricella, an invasive defoliator, and eastern larch beetle Dendroctonus simplex, a native, tree‐killing bark beetle, on tamarack Larix laricina from 2000 to in Minnesota, USA. We evaluated the utility of temporal, spatial and climatic variables in predicting the presence/absence of outbreaks of each insect in cells of rasterized aerial survey data. The role of defoliation by larch casebearer in outbreaks of eastern larch beetle was also investigated. For both species, the most important predictors of outbreak occurrence were proximity of conspecific outbreaks in space and time. For larch casebearer, outbreak occurrence was positively associated with spring precipitation and warmer growing seasons. Outbreak occurrence of eastern larch beetle was positively associated with warmer and dryer years and was more likely in cells with prior defoliation by larch casebearer. Our results demonstrate that climate can drive large scale outbreaks of introduced and non‐native disturbance agents on a single host species, and that interactions at the tree level between such agents may scale up to manifest across large temporal and spatial scales. [ABSTRACT FROM AUTHOR]

Abstract: The Alaskan Matanuska‐Susitna Basin (MSB) provides habitat for all five Pacific salmon species, and their large seasonal spawning runs are important both ecologically and economically. However, the encroachment of human development through urbanization and extractive industries poses a serious risk to salmon habitat in the MSB. Using systematic conservation planning techniques, different methods of incorporating anthropogenic risks were assessed to determine how to conserve salmon habitat in the area cost‐effectively. The consequences of four distinct conservation scenarios were quantified: no consideration either of urbanization or extractive industries (‘Risk ignored’ scenario); accounting for the risk of urbanization, and avoiding conservation in all areas rich in fossil fuels (‘Urbanization accounted, all extraction avoided’ scenario); accounting for urbanization and oil and gas development, but avoiding conservation in coal‐rich areas (‘Urbanization accounted, coal areas avoided’ scenario); and accounting for all anthropogenic risks to habitat, and allowing conservation in oil, gas, or coal‐rich areas (‘All risks accounted’ scenario). To compare conservation success and resilience, the impacts of these risks were estimated using Monte Carlo simulations. The final cost of each solution was then divided by the number of conservation targets met to determine a return on investment. Results from scenarios that avoided all extractive activities, or only coal, suggest that conservation targets cannot be met simply by avoiding areas rich in fossil fuels, and these scenarios resulted in lower returns on investment than those in which risks from extraction were incorporated into the solution. By developing a method for setting priorities that are economically based, this study provides a method for local managers and conservation groups to identify conservation opportunities in MSB river basins. [ABSTRACT FROM AUTHOR]

Questions Does the increase in disturbance associated with removing harvest residues negatively impact biodiversity and resilience in aspen-dominated forest ecosystems? How do responses of functional diversity measures relate to community recovery and standing biomass? Location Aspen ( Populus tremuloides, Michx.) mixedwood forests in Minnesota and Michigan, USA. Methods Three levels for two factors, organic matter removal and compaction, were fully crossed, resulting in nine experimental treatments that spanned a range of disturbance severity. Each treatment was replicated three times at each of three sites dominated by the same tree species but having different soil textures (clay, silty loam, sandy). Community composition and taxonomic diversity (species richness, species evenness, Shannon diversity index) were quantified based on woody species abundance sampled 5, 10 and 15 yr after disturbance. Community composition response was assessed using non-metric multidimensional scaling. Functional diversity (functional richness, evenness, dispersion and divergence) was also estimated using eight species effect and response traits. Finally, we examined community recovery as well as responses of species and functional diversity to disturbance severity over time using repeated measures ANOVA. Results Two responses indicated a potentially negative impact of whole-tree harvest relative to conventional, stem-only harvest: functional richness on silty loam soils and species evenness on clayey soils. Otherwise, negative impacts were restricted to forest floor removal or increased compaction. Recovery in community composition was reduced by the most severe treatments, particularly forest floor removal, across the study, but the responses of functional and taxonomic diversity varied among sites, with some measures increasing as a result of severe disturbance. Conclusions Maximization of standing biomass may mean a short-term sacrifice in species and functional diversity. Also, examinations of forest management impacts on species and functional diversity and composition should apply multiple metrics and indices to ensure potential impacts are not obscured by the reliance on a single approach. [ABSTRACT FROM AUTHOR]

The hydrologic regime is regarded as the primary driver of freshwater ecosystems, structuring the physical habitat template, providing connectivity, framing biotic interactions, and ultimately selecting for specific life histories of aquatic organisms. In the present study, we tested ecological theory predicting directional relationships between major dimensions of the flow regime and life history composition of fish assemblages in perennial free-flowing rivers throughout the continental United States. Using long-term discharge records and fish trait and survey data for 109 stream locations, we found that 11 out of 18 relationships (61%) tested between the three life history strategies (opportunistic, periodic, and equilibrium) and six hydrologic metrics (two each describing flow variability, predictability, and seasonality) were statistically significant (P ⩽ 0.05) according to quantile regression. Our results largely support a priori hypotheses of relationships between specific flow indices and relative prevalence of fish life history strategies, with 82% of all significant relationships observed supporting predictions from life history theory. Specifically, we found that (I) opportunistic strategists were positively related to measures of flow variability and negatively related to predictability and seasonality, (2) periodic strategists were positively related to high flow seasonality and negatively related to variability, and (3) the equilibrium strategists were negatively related to flow variability and positively related to predictability. Our study provides important empirical evidence illustrating the, value of using life histor.y theory to understand both the patterns and processes by which fish assemblage structure is shaped by adaptation to natural regimes of variability, predictability, and seasonality of critical flow events over broad biogeographic scales. [ABSTRACT FROM AUTHOR]

Current global models predict a hotter and drier climate in the southwestern United States with anticipated increases in drought frequency and severity coupled with changes in flash flood regimes. Such changes would likely have important ecological consequences, particularly for stream and riparian ecosystems already subject to frequent hydrologic disturbance. This study assessed the potential response of aquatic macroinvertebrates to interannual variation in hydrology in a spatially intermittent desert stream (Sycamore Creek, AZ). We compiled data on the recovery of macroinvertebrate communities following spring floods, with successional sequences captured 11 times over a 16-year period (1983–1999). This period encompassed a transition from perennial to intermittent flow in this system, and included a record drought in 1989–1990. Results show that while the size of floods initiating sequences had little explanatory power, changes in macroinvertebrate community structure during postflood succession were closely associated with antecedent flooding and drought. Year-to-year differences in benthic communities integrated taxon-specific responses to antecedent disturbance, including differential resistance to channel drying, use of hyporheic refugia, and variable rates of recovery once stream flow resumed. The long-term consequences of drying on community structure were only evident during later stages of postflood succession, illustrating an interaction between flood and drought recovery processes in this system. Our observations highlight the potential for predicted climate changes in this region to have marked and long-lasting consequences for benthic communities in desert streams. [ABSTRACT FROM AUTHOR]

1. Understanding species-specific habitat requirements is important in insect ecology. Co-occurrence of closely related species can provide insight into microhabitat segregation. Tiger beetles are well suited for such studies, as they occur in multi-species assemblages, exhibit species-specific habitat requirements, and are sensitive to anthropogenic modifications of the environment. 2. This study investigated ovipositional microhabitat choice of two sympatric tiger beetles at Point Reyes National Seashore, California, U.S.A., and evaluates how human activity affects that choice. Cicindela oregona is widespread and common whereas C. hirticollis, an undescribed subspecies, is locally threatened. Laboratory experiments tested oviposition-site selection of the two species in response to variations in moisture, pH, salinity, and grain size. Field experiments tested larval burrow location in response to experimental shading and experimental human disturbance. 3. Laboratory results indicated that while C. hirticollis females preferred a narrow range of moisture levels and fine sands, C. oregona females had more specific pH and salinity requirements and preferred coarser sands. They also differed in timing of activity. Preferences are probably related to prevention of fungal or bacterial infection and avoidance of desiccation and immersion. 4. In the field, significantly more larval burrows were found under experimental shading for both species. In addition, repeated human compaction of the sand significantly decreased the number of larval burrows. 5. Results suggest that protection of microhabitat, restoration of native plants, and limitation of human traffic are vital in the conservation management of tiger beetles. [ABSTRACT FROM AUTHOR]

To evaluate anthropogenic sedimentation in United States (U.S.) Pacific Northwest coastal streams, we applied an index of relative bed stability ( LRBS*) to summer low flow survey data collected using the U.S. Environmental Protection Agency’s Environmental Monitoring and Assessment Program field methods in a probability sample of 101 wadeable stream reaches. LRBS* is the log of the ratio of bed surface geometric mean particle diameter ( Dgm) to critical diameter ( D*cbf) at bankfull flow, based on a modified Shield’s criterion for incipient motion. We used a formulation of LRBS* that explicitly accounts for reductions in bed shear stress that result from channel form roughness due to pools and wood. LRBS* ranged from −1.9 to +0.5 in streams within the lower quartile of human riparian and basin disturbance, and was substantially lower (−4.2 to −1.1) in streams within the upper quartile of human disturbance. Modeling results suggest that the expected range of LRBS* in streams without human disturbances in this region might be generally between −0.7 and +0.5 in either sedimentary or volcanic lithology. However, streams draining relatively soft, erodible sedimentary lithology showed greater reductions in LRBS* associated with disturbance than did those having harder, more resistant volcanic (basalt) lithology with similar levels of basin and riparian disturbance. At any given level of disturbance, smaller streams had lower LRBS* than those with larger drainages. In sedimentary lithology (sandstone and siltstone), high-gradient streams had higher LRBS* than did low-gradient streams of the same size and level of human disturbance. High gradient streams in volcanic lithology, in contrast, had lower LRBS* than low-gradient streams of similar size and disturbance. Correlations between Dgm and land disturbance were stronger than those observed between D*cbf and land disturbance. This pattern suggests that land use has augmented sediment supplies and increased streambed fine sediments in the most disturbed streams. However, we also show evidence that some of the apparent reductions in LRBS*, particularly in steep streams draining small volcanic drainages, may have resulted in part from anthropogenic increases in bed shear stress. The synoptic survey methods and designs we use appear adequate to evaluate regional patterns in bed stability and sedimentation and their general relationship to human disturbances. More precise field measurements of channel slope, cross-section geometry, and bed surface particle size would be required to use LRBS* in applications requiring a higher degree of accuracy and precision, such as site-specific assessments at individual streams. [ABSTRACT FROM AUTHOR]

The article presents a study on the examination of effects of catastrophic flooding on stream biogeochemistry changes in Staunton River in Shenandoah National Park in Virginia. It relates that the flood caused large-scale debris flows and complete scouring of riparian soils down to bedrock in the lower 2 kilometer of the stream. It also discusses that the response was elevations in the stream concentration of Calcium, Magnesium, and Potassium by 59 per cent, 87 per cent, and 49 per cent, respectively, for 6 months immediately following the flood.

Aim The local distribution of woody vegetation affects most functional aspects of semiarid landscapes, from soil erosion to nutrient cycling. With growing concern about anthropogenic climate change, it has become critical to understand the ecological determinants of woody plant distribution in semiarid landscapes. However, relatively little work has examined the determinants of local variation in woody cover. Here we examine water balance controls associated with patterns of tree cover density in a topographically complex semiarid woodland. Location Los Pinos Mountains, Sevilleta National Wildlife Refuge LTER, New Mexico, USA. Methods To explore the relationship between local water balance variation and tree cover density, we used a combination of high-resolution (1 × 1 m), remotely sensed imagery and quantitative estimates of water balance variation in space and time. Regression tree analysis (RTA) was used to identify the environmental parameters that best predict variation in tree cover density. Results Using six predictor variables, the RTA explains 39% of the deviance in tree cover density over the landscape. The relationship between water balance conditions and tree cover density is highly contingent; that is, similar tree cover densities occur under very different combinations of water balance parameters. Thus, the effect of one environmental parameter on tree cover density depends on the values of other parameters. After tree cover density is adjusted for water balance conditions, residual variation is related to tree cover density in the neighbourhood of a particular location. Conclusion In semiarid landscapes, vegetation structure is largely controlled by water supply and demand. Results presented here indicate that localized feedbacks and site-specific historical processes are critical for understanding the responses of semiarid vegetation to climate change. [ABSTRACT FROM AUTHOR]

Abstract Aim There is increasing research attention being given to the role of interactions among natural disturbances in ecosystem processes. We studied the interactions between fire and spruce beetle (Dendroctonus rufipennis Kirkby) disturbances in a Colorado subalpine forest. The central questions of this research were: (1) How does fire history influence stand susceptibility to beetle outbreak? And conversely, (2) How does prior occurrence of a beetle outbreak influence stand susceptibility to subsequent fire? Methods We reconstructed the spatial disturbance history in a c. 4600 ha area by first identifying distinct patches in the landscape on aerial photographs. Then, in the field we determined the disturbance history of each patch by dating stand origin, fire scars, dates of mortality of dead trees, and releases on remnant trees. A geographical information system (GIS) was used to overlay disturbance by fire and spruce beetle. Results and main conclusions The majority of stands in the study area arose following large, infrequent, severe fires occurring in c . 1700, 1796 and 1880. The study area was also affected by a severe spruce beetle outbreak in the 1940s and a subsequent low-severity fire. Stands that originated following stand-replacing fire in the late nineteenth century were less affected by the beetle outbreak than older stands. Following the beetle outbreak, stands less affected by the outbreak were more affected by low-severity fire than stands more severely affected by the outbreak. The reduced susceptibility to low-severity fire possibly resulted from increased moisture on the forest floor following beetle outbreak. The landscape mosaic of this subalpine forest was strongly influenced by the interactions between fire and insect disturbances. [ABSTRACT FROM AUTHOR]

Summary 1 Using 18 years of permanent-plot data from a 4-ha stand, we investigated whether stand dynamics is predictable, i.e. driven by competition (inferred from species shade tolerances), or unpredictable (driven by frequent exogenous disturbance or other factors). We also considered whether small disturbance might accelerate or retard succession. The study involved dynamics of stems ≥ 4.5 cm diameter at breast height (d.b.h.) for 10 important species in a southern mixed hardwood forest 80 years after selective removal of Pinus taeda. 2 Shade-tolerant species (Ilex opaca , Fagus grandifolia , Magnolia grandiflora , Acer rubrum , Nyssa sylvatica ) increased in small d.b.h. classes, whereas intermediate (Quercus alba ) and shade-intolerant species (Q. nigra , Q. hemisphaerica , Liquidambar styraciflua , Pinus taeda ) did not. 3 Per-capita ingrowth was higher for shade-tolerant species (0.016–0.024) than for intermediate and shade-intolerant species (0–0.006), whereas mortality was lower ((0.006–0.015 vs. 0.019–0.035). 4 Over 18 years, ingrowth exceeded mortality loss slightly, resulting in an increase in number of stems of about 1%, from 995 to 1029 stems ha-1 ; average live basal area (BA; 35.9 m2 ha-1 ) varied by less than 2% over eight surveys spanning 18 years. Basal area increased substantially for three species (Ilex opaca , Pinus taeda , Acer rubrum ), and declined strongly for three others (Quercus nigra , Q. hemisphaerica and Fagus grandifolia ). 5 The stand d.b.h. distribution had a rotated sigmoid form which became flatter over time. Species d.b.h. distributions were mostly bell-shaped for intolerant species, and irregular-to-monotonic declining for shade-tolerant species. Over time, bell-shaped distributions moved to the right and irregular distributions remained irregular. 6 A hurricane in 1986 caused loss of 4.5% of standing BA. Compositional change was most rapid during the... [ABSTRACT FROM AUTHOR]

Abstract Aim The aim of this study was to document changes in forest composition, structure and distribution across Massachusetts, USA, from the time of European settlement (seventeenth century) to the present, and to investigate environmental and historical influences on regional patterns of variation. Location The study area encompasses the State of Massachusetts (69.9–73.5°E, 41.3–42.9°N), a 21,000-km2 area in the north-eastern United States. Methods A wide range of historical sources was used to document changes in land use and land cover for the historical period. Witness trees from early land surveys enabled us to evaluate vegetation patterns prior to widespread European settlement, and to compare historical and modern species composition. Nineteenth century maps of forest cover and contemporary agricultural censuses documented forest patterns during the peak agricultural period. Geographic Information System analyses were used to relate variation in climate, geology and land-use history to historical and modern forest composition. Results Massachusetts has a complex east-to-west environmental gradient involving changes in physiography, climate, geology and natural disturbance. Until the middle of the twentieth century, agriculture was the most important land-use across the region; although the percentage of land in agriculture and the timing of major land-use changes were remarkably consistent across the state, historical forest patch sizes varied locally and regionally in relation to physiography. Forest composition of both early historical and modern forests is most strongly related to environmental conditions, especially variation in climate. Historical land-use resulted in a state-wide increase in early successional tree species and a dramatic, although recovering, change in forest structure. Main conclusions At a regional scale, environmental conditions apparently control broad patterns of variation in vegetation... [ABSTRACT FROM AUTHOR]

Abstract Aim This study documents the vegetation history and age-structure of a rare, ridgetop dwarf pine–oak community and compares the dynamics of this unusual vegetation with similar dwarf pine communities found elsewhere in the north-eastern United States (US). Location The study area is located on the summit of Mt Everett in the Taconic Mountains of south-western Berkshire County, Massachusetts, USA (42°06′N 73°26′W). Methods Vegetation composition, tree age-structure, physical site characteristics, and evidence of fire and other disturbances were determined for twelve 15 × 15 m plots in dwarf pine–oak vegetation and two plots in oak forests on the summit. Age-structure analyses, tree-ring patterns, and historical records of human and natural disturbance were used to investigate the long-term history and dynamics of the summit vegetation. Results The summit of Mt Everett has been dominated by dwarf pines (1–3 m tall) and ericaceous shrubs similar to the modern vegetation throughout the historical period; there is no evidence that tall-stature forests occurred on the site at any point in the past few centuries. The summit supports uneven-aged stands; pitch pine (Pinus rigida ) recruitment began in the 1830s and occurred in every decade since the 1860s. Average pitch pine age is seventy-eight with a range of 12–170 years. Red oak (Quercus rubra ) and red maple (Acer rubrum ) increased in importance in the twentieth century, with most stems establishing from 1940 to 1980. Pitch pine radial growth rates averaged <0.5 mm year-1 while red oak and red maple averaged 1.0 and 0.8 mm year-1 , respectively. In some areas, hardwoods have overtopped pitch pines, apparently resulting in pitch pine mortality. Whereas most dwarf pitch pine communities occur on sites that burn frequently and have a high degree of cone serotiny, we found no evidence of recent fires or cone serotiny. Small... [ABSTRACT FROM AUTHOR]

Summary 1 In 1997, a major windstorm blew down over 10 000 hectares of subalpine forest in Routt National Forest in north-western Colorado. We tested whether fire history and topographic variation, across the landscape, determined subsequent susceptibility to damage from this windstorm. 2 We combined dendrochronological techniques with a geographical information system (GIS) to examine the relationship between the effects of the blowdown and the spatial heterogeneity of the vegetation. We reconstructed the spatial fire history in a c. 4400 ha area by first identifying distinct patches in the landscape on aerial photographs, and then in the field by determining the disturbance history of each patch by dating fire scars, ages of post-fire cohorts and releases of remnant trees. 3 Both topographic position and fire history contribute to susceptibility to wind damage. Stands at higher elevations, on easterly slopes and closer to ridges were most affected. Younger stands (i.e. more recently affected by stand-initiating fires) were less affected particularly in areas affected by low–moderate severity blowdown (< 80% damage). The ecological effects of the very severe 1997 windstorm were mediated by the legacies of fires which occurred several centuries ago. Interactions between natural disturbances, such as fire and blowdown, lead to synergistic effects on forest dynamics, even when disturbance is infrequent. [ABSTRACT FROM AUTHOR]

AimIn this study, we evaluated the fire-forest mosaic of a mixed conifer forest landscape by testing the hypothesis that pre-fire suppression fire regime parameters vary with species composition (tree species), and environment (i.e. slope aspect, slope position, elevation). LocationOur study was conducted in the 1587 ha Cub Creek Research Natural Area (CCRNA), Lassen National Forest, CA, USA. MethodsWe quantified the return interval, seasonal occurrence, size, rotation period, and severity of fires using dendroecology. ResultsSlope aspect, potential soil moisture, forest composition, and fire regime parameters in our study area co-vary. Median composite and point fire return intervals (FRI) were longest on higher, cooler, more mesic, north-facing (NF) slopes covered with white fir (Abies concolor), Douglas fir (Pseudotsuga menziesii)–white fir, and red fir (A. magnifica)–white fir forests, shortest on the dry, south-facing (SF) slopes covered with ponderosa pine (Pinus ponderosa)–white fir forests and intermediate on west-facing slopes dominated by white fir–sugar pine (P. lambertiana)–incense cedar (Libocedrus decurrens) forests. The spatial pattern for length of fire rotation (FR) was the same as that for FRI. Fires in CCRNA mixed conifer forests occurred mainly (90%) in the dormant season. Size of burns in CCRNA mixed conifer forests were generally small (mean=106 ha), however, during certain drought years widespread fires burned across fuel breaks and spread throughout the watershed. Fire severity was mainly high on upper slopes, low on lower slopes and moderate and low severity on middle slopes. Patterns of fire severity also varied with slope aspect. Fire frequency decreased dramatically in CCRNA after 1905. ConclusionsIn CCRNA, fire regime parameters [e.g. FRI, fire extent, FR, fire severity] varied widely with species composition, slope aspect and slope position. There was also temporal variation in fire extent with the most widespread fires occurring during drought years. The important contributions of topography and climate to variation in the fire regime indicates that exogenous factors play a key role in shaping the fire-forest structure mosaic and that the fire-forest structure mosaic is more variable, less predictable and less stable than previously thought. Finally, some characteristics of the fire regime (i.e. fire severity, season of burn) in CCRNA are different than those described for other mixed conifer forests and this suggests that there are geographical differences in mixed conifer fire regimes along the Pacific slope. [ABSTRACT FROM AUTHOR]

Abstract Aim Spatial and temporal variation in fire regime parameters and forest structure were assessed. Location A 2630-ha area of mid- and upper montane forest in Lassen Volcanic National Park (LVNP). Methods Two hypotheses were tested concerned with fire-vegetation relationships in southern Cascades forests: (1) fire regime parameters (return interval, season of burn, fire size, rotation period) vary by forest dominant, elevation and slope aspect; and (2) fire exclusion since 1905 has caused forest structural and compositional changes in both mid- and upper montane forests. The implications of the study for national park management are also discussed. Results Fire regime parameters varied by forest compositional group and elevation in LVNP. Median composite and point fire return intervals were shorter in low elevation Jeffrey pine (Pinus jeffreyi) (JP) (4–6 years, 16 years) and Jeffrey pine–white fir (Abies concolor) (JP-WF) (5–10 years, 22 years) and longer in high elevation red fir (Abies magnifica)— western white pine (Pinus monticola) (RF-WWP) forests (9–27 years, 70 years). Median fire return intervals were also shorter on east-facing (6–9 years, 16.3 years) and longer on south- (11 years, 32.5 years) and west-facing slopes (22–28 years, 54-years) in all forests and in each forest composition group. Spatial patterns in fire rotation length were the same as those for fire return intervals. More growing season fires also occurred in JP (33.1%) and JP-WF (17.5%) than in RF-WWP (1.1%) forests. A dramatic decline in fire frequency occurred in all forests after 1905. Conclusions Changes in forest structure and composition occurred in both mid- and upper montane forests due to twentieth-century fire exclusion. Forest density increased in JP and JP-WF forests and white fir increased in JP-WF forests and is now replacing Jeffrey pine. Forest density only increased in some RF-WWP stands, but not others. Resource managers restoring fire to these now denser forests need to burn larger areas if fire is going to play its pre-settlement role in montane forest dynamics. [ABSTRACT FROM AUTHOR]

Fire is a critical force in structuring ecosystems, but it also removes substantial amounts of nitrogen (N), which can limit plant growth. Biological N fixation (BNF) may alleviate fire‐induced N deficiencies that inhibit ecosystem recovery, yet if and how BNF achieves this under frequent fire is unclear. This problem is further complicated in the context of modern human influences (such as land‐use history and atmospheric N deposition), which may confound the relationship between fire and fixation. Here, we investigate whether BNF supplies the N necessary to replace fire‐induced N losses in restored longleaf pine savannas, and, if so, what factors control fixation. We established 54 1‐ha plots of longleaf pine capturing 227 yr of forest recovery and a broad gradient of fire return interval (1.5–20 yr) at two sites in the southeastern United States. We quantified N fixation from three functional groups (herbaceous legumes, soil crusts, and asymbiotic N fixing bacteria), N losses from individual fire events and ecosystem dynamics of N supply and demand. We found that BNF rates were low but sustained over stand age but were substantially below estimated rates of atmospheric N deposition. While fire temporarily stimulated BNF from herbaceous legumes, neither BNF nor atmospheric N deposition were sufficient to balance N losses from fire and soil N stocks declined over stand age. However, rates of N mineralization were surprisingly high and tree productivity was unrelated to N availability, questioning the importance of N limitation in these temperate savannas. While it is possible that progressive N losses signal a decline in ecosystem resiliency, N enrichment from multiple land‐use transitions and anthropogenic N deposition may suppress rates of BNF or diminish its importance as a long‐term N balancing source in these pyrogenic ecosystems. In this case, fire may be acting as relief mechanism, critical for returning the modern longleaf pine landscape to its historical oligotrophic condition. [ABSTRACT FROM AUTHOR]

To better understand forest carbon (C) storage patterns, C budgets were constructed and contrasted among four stands of different vegetation, elevation, and disturbance histories from the northern Sierra Nevada of California, USA. The True‐fir stand, considered to be undisturbed and at a higher elevation (1820 m), stored substantially more C than the three stands at lower elevation (1360–1430 m) and that had sustained various levels of disturbance. The Mixed Conifer stand was minimally disturbed by thinning 27 yr ago. The Oak stand was moderately disturbed by two fires in the past 17 yr. The Pine Plantation stand was considered the most disturbed having been cleared and subjected to site preparation and windrowing 50 yr ago and thinned again 17 yr ago. Total C mass to a 30 cm soil depth (Mg C/ha) was True‐fir 531, Mixed Conifer 317, Oak 310, and Pine Plantation 253. Adding in soil C to 1 m depth increased total C mass to 595, 349, 404, and 345, respectively. True‐fir had significantly larger C mass than the other three stands in nearly all pools measured. The three low‐elevation stands, while similar to each other in total C, stored C differently in patterns that reflected their disturbance histories. Oak and Pine Plantation were very similar in detrital C stores, whereas Mixed Conifer had at least twice the C mass in both the forest floor and woody debris pools. Mixed Conifer had higher concentrations of soil C but had very low soil C mass due to considerably lower fine‐soil bulk densities and higher rock contents. C pools in the forest floor and soil showed observable patterns over distances of 10–100 m that paralleled the patterns of C in the live vegetation. Total C in True‐fir was equal to the large masses of old‐growth conifer forests of the U.S. Pacific Northwest and likely represents a high‐equilibrium level of C storage for forests of this region. The three low‐elevation stands represent typical forests in various states of recovery from disturbance that are in C accumulating stages. [ABSTRACT FROM AUTHOR]

We studied the effects of windthrow on the understory plant species composition of a pine forest (dominated by Pinus strobus) and an oak forest (dominated by Quercus ellipsoidalis). We recorded the presenceof vascular plant species in randomly located quadrats in the two forests, and in three microsite types associated with tipup mounds (pit, old soil and new soil) in the pine forest at irregular intervals over the course of 14 years. The understories of the two forests remained distinct throughout the study. The frequency of occurrence of a number of forest floor species considerably increased; few species decreased. The disturbance specialists Rubus idaeus and Polygonum cilinode increased in frequency throughout the study in the pine forest, butare beginning to decline in the less disturbed oak forest. Annuals and biennials preferentially colonized the disturbed soil of microsites on tipups, and declined in frequency after about 7 yr. Both forestshave increased in understory species richness, but have not changed substantially in the distribution of growth forms. Despite early differences in species composition, microsite types associated with tipupmounds became more similar through time. Although small in magnitude, there was a directional change in understory composition at both forests, with no apparent sign of a return to pre-disturbance conditions. [ABSTRACT FROM AUTHOR]