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Many studies recommend no-till (NT) to increase soil organic carbon (SOC) in the topsoil (input) from crop residues were further analyzed. The systems were either non-fertilized (N0) or fertilized with mineral N (N1) according to a demand of 180 and 380 kg N ha−1 year−1 in the silage maize and grassland systems, respectively. For the 7-year period, the renovated grassland using NT ensured maintenance of the initial SOC in the topsoil, while a conversion toward arable cropping resulted in SOC losses, regardless of the tillage method. The use of NT during conversion significantly reduced these losses from 2.5 Mg ha−1 year−1 to 1.8 Mg ha−1 year−1, for a 28% reduction compared to CT. In the subsoil (30–90 cm), SOC remained stable and was not affected by the cropping systems nor by the tillage method. Reduced annual Cinput was found as the main factor affecting SOC losses after grassland removal, regardless of the tillage method. Our findings highlight the potential of NT to mitigate annual SOC losses after grassland conversion if annual Cinput remains high.

Integrating leys, cover crops, and animal manures constitute promising avenues to reach annual soil organic carbon changes (ΔSOC) >0.4% in forage and grain-based crop rotations, rates required to offset the increasing C emissions from fossil fuels (“4 per mille” initiative). How these practices and rotations perform in reaching this aim was object of analysis in this paper. Five cropping systems (CS), including three three-year forage and grain-based crop rotations containing annual grass-clover leys (FR and MR) or cover crops (GR), and two contrasting controls (continuous silage maize (CM), and permanent grassland (PG)) were compared for their impact on SOC stocks over eight years (2010–2018). The CS were unfertilized (N0) or fertilized using cattle slurry (N1) at a rate of 240 kg N ha−1 yr−1 applied in the non-leguminous crops. The ΔSOC of the top 30 cm soil layer and the annual carbon inputs (Cin) from slurry applications and plant residues were estimated, their relationship established, and the slurry-induced C retention coefficient was determined. The FR and MR SOC stocks remained stable at N1, while the GR and CM SOC decreased over time by tendency even at N1. Only the PG reached ΔSOC >0.4%. Differences in ΔSOC between CS and N rates were highly associated with the system-specific increase in belowground Cin, induced by slurry applications. Slurry-induced C retention coefficients differed strongly between CS: CM (3%) followed by GR (12%), and by FR and MR (20–15%), and lastly by PG (24%). Promoting belowground carbon inputs was identified as an efficient way to reach significant increases in ΔSOC. We conclude that a ley in only one out of three years is not sufficient to significantly increase SOC stocks in arable crop rotations of the study region.