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Factors such as soil type and precipitation vary across rangeland landscapes, and these factors affect restoration outcomes and ultimately mean that “one size fits all” management strategies are not effective across large, complex landscapes. Big sagebrush (Artemisia tridentata) is a foundational rangeland species that is important to wildlife habitat across the western U.S. On the Colorado Plateau, sagebrush is important browse for ungulates, such as mule deer and pronghorn, which motivates a great deal of restoration effort. However, most scientific knowledge of big sagebrush comes from the Great Basin, and we know much less about how to restore sagebrush on the Colorado Plateau, where soils and precipitation patterns are different and conditions are warmer and drier. This fact sheet describes research about establishing and restoring sagebrush seedlings on the Colorado Plateau.

This handbook is a guide to developing realistic project plans and implementing appropriate management strategies by enhancing your understanding of basic shrub biology, ecological concepts,and management principles.

In the western United States, fire has become a significant concern in the management of big sagebrush (Artemisia tridentata Nutt.) ecosystems. This is due to large‐scale increases in cover of the fire‐prone invasive annual cheatgrass (Bromus tectorum L.) and, concurrently, concerns about declining quantity and quality of habitat for Greater Sage‐grouse (Centrocercus urophasianus). The prevailing paradigm is that fire results in a loss of sage‐grouse habitat on timescales relevant to conservation planning (i.e., 1–20 yr), since sagebrush cover can take many more years to recover post‐fire. However, fire can have effects that improve sage‐grouse habitat, including stimulating perennial grass and forb production. The conditions under which fire results in the permanent loss or enhancement of sage‐grouse habitat are not well understood. We used long‐term data from the Utah Division of Wildlife Resources Range Trend Project to assess short‐term (1–4 yr post‐treatment) and long‐term (6–10 yr post‐treatment) effects of fire on vegetation cover at 16 sites relative to sage‐grouse habitat vegetation guidelines. Sagebrush cover remained low post‐fire at sites considered historically unsuitable for sage‐grouse (10%) pre‐fire sagebrush cover, sagebrush cover decreased to10% cover. Post‐fire sagebrush cover was positively related to elevation. Across all sites, perennial grasses and forbs increased in cover to approximately meet the habitat vegetation guidelines for sage‐grouse. Cheatgrass cover did not change in response to fire, and increased perennial grass cover appears to have played an important role in suppressing cheatgrass. Our results indicate that, while fire poses a potential risk for sage‐grouse habitat loss and degradation, burned sites do not necessarily need to be considered permanently altered, especially if they are located at higher elevation, have high sagebrush cover pre‐fire, and are reseeded with perennial grasses and forbs post‐fire. However, our results confirm that fire at more degraded sites, for example, those with

This fact sheet provides suggestions for how cattle grazing can be integrated into Phragmites management programs in Northern Utah.

Rainfall and herbivory are fundamental drivers of grassland plant dynamics, yet few studies have examined long‐term interactions between these factors in an experimental setting. Understanding such interactions is important, as rainfall is becoming increasingly erratic and native wild herbivores are being replaced by livestock. Livestock grazing and episodic low rainfall are thought to interact, leading to greater community change than either factor alone. We examined patterns of change and stability in herbaceous community composition through four dry periods, or droughts, over 15 years of the Kenya Long‐term Exclosure Experiment (KLEE), which consists of six different combinations of cattle, native wild herbivores (e.g., zebras, gazelles), and mega‐herbivores (giraffes, elephants). We used principal response curves to analyze the trajectory of change in each herbivore treatment relative to a common initial community and asked how droughts contributed to community change in these treatments. We examined three measures of stability (resistance, variability, and turnover) that correspond to different temporal scales and found that each had a different response to grazing. Treatments that included both cattle and wild herbivores had higher resistance (less net change over 15 years) but were more variable on shorter time scales; in contrast, the more lightly grazed treatments (no herbivores or wild herbivores only) showed lower resistance due to the accumulation of consistent, linear, short‐term change. Community change was greatest during and immediately after droughts in all herbivore treatments. But, while drought contributed to directional change in the less grazed treatments, it contributed to both higher variability and resistance in the more heavily grazed treatments. Much of the community change in lightly grazed treatments (especially after droughts) was due to substantial increases in cover of the palatable grass Brachiaria lachnantha. These results illustrate how herbivory and drought can act together to cause change in grassland communities at the moderate to low end of a grazing intensity continuum. Livestock grazing at a moderate intensity in a system with a long evolutionary history of grazing contributed to long‐term stability. This runs counter to often‐held assumptions that livestock grazing leads to directional, destabilizing shifts in grassland systems.

Recent elevated temperatures and prolonged droughts in many already water-limited regions throughout the world, including the southwestern United States, are likely to intensify according to future climate-model projections. This warming and drying can negatively affect perennial vegetation and lead to the degradation of ecosystem properties. To better understand these detrimental effects, we formulate a conceptual model of dryland ecosystem vulnerability to climate change that integrates hypotheses on how plant species will respond to increases in temperature and drought, including how plant responses to climate are modified by landscape, soil and plant attributes that are integral to water availability and use. We test the model through a synthesis of fifty years of repeat measurements of perennial plant species cover in large permanent plots across the Mojave Desert, one of the most water-limited ecosystems in North America. Plant species ranged in their sensitivity to precipitation in different seasons, capacity to increase in cover with high precipitation and resistance to decrease in cover with low precipitation. Our model successfully explains how plant responses to climate are modified by biophysical attributes in the Mojave Desert. For example, deep-rooted plants were not as vulnerable to drought on soils that allowed for deep-water percolation, whereas shallow-rooted plants were better buffered from drought on soils that promoted water retention near the surface. Synthesis. Our results emphasize the importance of understanding climate–vegetation relationships in the context of biophysical attributes that influence water availability and provide an important forecast of climate-change effects, including plant mortality and land degradation in dryland regions throughout the world.

Public land management agencies, such as the Bureau of Land Management (BLM), are charged with managing rangelands throughout the Western United States for multiple uses such as livestock grazing and conservation of sensitive species and their habitats. Monitoring of condition and trends of these rangelands, particularly with respect to effects of livestock grazing, provide critical information for effective management of these multi-use landscapes. We therefore investigated the availability of livestock grazing-related quantitative monitoring data and qualitative region-specific Land Health Standards (LHS) data across BLM grazing allotments in the Western United States. We then queried university and federal rangeland science experts about how best to prioritize rangeland monitoring activities. We found that the most commonly available monitoring data were permittee-reported livestock numbers and season-of-use data (71% of allotments) followed by repeat photo points (58%), estimates of forage utilization (52%), and finally, quantitative vegetation measurements (37%). Of the 57% of allotments in which LHS had been evaluated as of 2007, BLM indicated 15% had failed to meet LHS due to livestock grazing. A full complement of all types of monitoring data, however, existed for only 27% of those 15%. Our data inspections, as well as conversations with rangeland experts, indicated a need for greater emphasis on collection of grazing-related monitoring data, particularly ground cover. Prioritization of where monitoring activities should be focused, along with creation of regional monitoring teams may help improve monitoring. Overall, increased emphasis on monitoring of BLM rangelands will require commitment at multiple institutional levels.

Empirical and theoretical evidence suggests that facilitation between plants, when it occurs, is more likely during periods of abiotic stress, while competition predominates under more moderate conditions. Therefore, one might expect the relative importance of competition vs. facilitation to vary seasonally in ecosystems characterized by pronounced dry (abiotically stressful) and wet (benign) seasons. Herbivory also varies seasonally and can affect the net outcome of plant-plant interactions, but the interactive effects of seasonality and herbivory on the competition-facilitation balance are not known. I experimentally manipulated neighboring plants and herbivory during wet and dry periods for two species of grass: Cynodon plectostachyus and Pennisetum stramineum, in the semiarid Laikipia District of Kenya. These experiments indicate that Pennisetum was competitively dominant during the wet season and that it responded negatively to grazing, especially during the dry season. Cynodon showed more complex season- and herbivore-dependent responses. Cynodon experienced facilitation that was simultaneously dependent on presence of herbivores and on dry season. During the wet season Cynodon experienced net competition. These results illustrate how herbivory and seasonality can interact in complex ways to shift species-species competition-facilitation balance. Additionally, because Cynodon and Pennisetum are key players in a local successional process, these results indicate that herbivory can affect the direction and pace of succession.