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Competing Interests: We declare no competing interests. FYL, OCC, and MZT conceptualised the study, analysed data, and wrote the manuscript. ZG, GX, CWK, and HL performed computational simulation experiments, analysed data, and reviewed the manuscript. LHL, S-WF, YSG, and JCCW conceptualised the study, analysed data, and reviewed the manuscript. All authors approved the final version of the manuscript. All authors had full access to all the data in the study and had final responsibility for the decision to submit for publication.