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The Sulphur Mountain Formation in Alberta, Canada, comprises shales, siltstones, and limestones laid down on the continental shelf and shoreline along the western margin of the North American Craton during the Early and Middle Triassic. At least 13 taxa of fossil marine fishes have been found at a number of Triassic localities along the Front Ranges and foothills in Alberta. Common taxa (Boreosomus, Birgeria, Bobasatrania, Australosomus, Saurichthys, parasemionotids, and coelacanths) found at localities in Alberta are similar to those found at the classic Lower and Middle Triassic localities near Wapiti Lake, British Columbia, and in East Greenland, Spitzbergen, Madagascar, and China. The occurrence of Lower Triassic fishes in western Alberta shows that this fauna was widespread, occurring over at least 1000 km of shoreline, and that ecological and environmental factors were similar along the western edge of the Panthalassic Ocean during the Early Triassic. [ABSTRACT FROM AUTHOR]

A new species of stem actinopterygian, Helmolepis cyphognatus sp. nov., is reported from the Lower Triassic Sulphur Mountain Formation of western Canada (probably Smithian). This taxon differs from the only other known Early Triassic platysiagid, H. gracilis from the Lower Triassic Wordie Creek Formation of East Greenland (Griesbachian), in counts of branchiostegal rays, shape of the maxilla, shape (and possibly counts) of extrascapulars, and the size ratio of major opercular bones. In spite of their overall unfavorable preservation, the numerous available specimens amend our knowledge of the little known genus Helmolepis considerably: it has become evident that the morphology of Helmolepis cyphognatus sp. nov. comes closest to Platysiagum minus (Middle Triassic Besano Formation of central Europe). This study suggests placement of the two genera in the family Platysiagidae. Investigation of this new species also shows certain features of the cheek and the caudal fin are more primitive than previously believed, whereas the snout region is probably derived but of yet uncertain affinities in Helmolepis. [ABSTRACT FROM AUTHOR]

Myledaphus bipartitus is a Late Cretaceous (75 Ma) freshwater guitarfish, commonly represented by teeth and vertebral centra. The species was first described in 1876, but until now, very little has been known about its ecology and life history. We analyzed the age and growth using 117 fossilized centra (74 hemisectioned) from the Oldman and Dinosaur Park formations of southern Alberta. The resulting estimated maximum age was 16 years. Radial distance (RD) at the estimated asymptotic growth was calculated, and average RD at birth and 3 years were compared with two extant marine species of Rhinobatos ( R. productus and R. rhinobatos). Birth ring size ranged from 0.9 to 1.8 mm RD. At age three, centrum RD was between 2.8 and 4.6 mm RD for M. bipartitus and significantly less than R. productus and R. rhinobatos (5.0-6.6 mm RD). Myledaphus approached 'asymptotic' growth at 7.8 mm RD in the growth equation. The growth trajectory of Myledaphus had a similar slope to the Rhinobatos species, but with significantly smaller centrum RD. Rhinobatos rhinobatos also outlived Myledaphus by 8 years. [ABSTRACT FROM AUTHOR]

Horseshoeichthys armaserratus, gen. et sp. nov., (Clupeomorpha: Ellimmichthyiformes: Sorbinichthyidae) is described from the Horseshoe Canyon Formation (Maastrichtian), Albertosaurus bonebed locality, Alberta, Canada. Horseshoeichthys armaserratus is classified as an ellimmichthyiform based on the following characters: the presence of a sixth infraorbital with a sensory canal that leads to the fifth infraorbital, absence of a supraorbital bone, subrectangular predorsal scutes, parietals in contact with each other at the midline, and two supramaxillae. The specimen is classified in the Sorbinichthyidae Family as it has abdominal ribs articulating in pits on the centra, posterior spines on predorsal scutes, and absence of a median spine on predorsal scutes. A new genus and species is proposed based on the presence of (anteriorly) Y-shaped mesethmoid, supraorbital, subrectangular predorsal scutes with coarse, rounded serrae on the posterior margin and a large anterior projection, scales with serrae on the circuli, and two postcleithra. This specimen represents the first freshwater ellimmichthyiform from the Upper Cretaceous of North America and the highest paleolatitude (59°N) occurrence known for the family. Furthermore, the dentary and centra have distinctive morphologies that are matched by specimens in microvertebrate localities from three underlying formations, including the Milk River Formation (Santonian), which indicates at least a 14 million year history for this lineage. Horseshoeichthys armaserratus, gen. et sp. nov., (Clupeomorpha: Ellimmichthyiformes: Sorbinichthyidae), de la localité du gisement d'ossements d'Albertosaurus de la Formation de Horseshoe Canyon (Maastrichtien), en Alberta, au Canada, est décrit. Horseshoeichthys armaserratus est affecté aux ellimmichthyiformes à la lumière des caractères suivants : la présence d'un sixième sous-orbitaire présentant un canal sensoriel qui mène au cinquième sous-orbitaire, l'absence d'un os sus-orbitaire, des plaques osseuses pré-dorsales sub-rectangulaires, des pariétaux en contact les uns avec les autres au niveau de la ligne médiane et deux sus-maxillaires. Le spécimen est affecté aux sorbinichthyidés puisqu'il présente des côtes abdominales s'articulant dans des fosses sur le centrum et des épines postérieures sur les plaques osseuses pré-dorsales, et que ces dernières n'ont pas d'épine médiane. Un nouveau genre et une nouvelle espèce sont proposés à la lumière de la présence d'un mésethmoïde (antérieurement) en forme de Y, d'un sus-orbitaire, de plaques osseuses pré-dorsales sub-rectangulaires présentant des stries arrondies et épaisses sur leur bord libre postérieur et une grande projection antérieure, d'écailles avec des stries sur le circulus et de deux postcleithra. Ce spécimen représente le premier ellimmichthyiforme d'eau douce du Crétacé supérieur de l'Amérique du Nord et l'exemple connu de cette famille provenant de la plus haute paléolatitude (59oN). En outre, le dentaire et le centrum présentent des morphologies particulières semblables à celles de spécimens trouvés dans des localités de microvertébrés de trois formations sous-jacentes, dont la Formation de Milk River (Santonien), ce qui indique une histoire d'au moins 14 millions d'années pour cette lignée. [ABSTRACT FROM AUTHOR]