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Over the past decade, a growing interest has developed on the archaeology, palaeontology, and palaeoenvironments of the Arabian Peninsula. It is now clear that hominins repeatedly dispersed into Arabia, notably during pluvial interglacial periods when much of the peninsula was characterised by a semiarid grassland environment. During the intervening glacial phases, however, grasslands were replaced with arid and hyperarid deserts. These millennial-scale climatic fluctuations have subjected bones and fossils to a dramatic suite of environmental conditions, affecting their fossilisation and preservation. Yet, as relatively few palaeontological assemblages have been reported from the Pleistocene of Arabia, our understanding of the preservational pathways that skeletal elements can take in these types of environments is lacking. Here, we report the first widespread taxonomic and taphonomic assessment of Arabian fossil deposits. Novel fossil fauna are described and overall the fauna are consistent with a well-watered semiarid grassland environment. Likewise, the taphonomic results suggest that bones were deposited under more humid conditions than present in the region today. However, fossils often exhibit significant attrition, obscuring and fragmenting most finds. These are likely tied to wind abrasion, insolation, and salt weathering following fossilisation and exhumation, processes particularly prevalent in desert environments. Introduction Previous investigations Methods - Surveys - Systematic palaeontology - Taphonomic analysis Results - Systematic Palaeontology - Taphonomic analysis Discussion - Species representation and palaeoenvironmental implications - Taphonomic analysis Conclusions

The Middle Palaeolithic record of the Arabian Peninsula can provide crucial evidence for understanding human dispersal. The authors summarise the archaeological evidence and suggest some of the routes taken by the earliest humans coming out of Africa, including one implying the use of boats. Early populations adapted to a hospitable environment, but had later to adapt to the advance of the desert.