Search

Corallocoris marksae (Woodward, 1958) (Figs. 5M–R, 7, 9) Omania marksae Woodward, 1958: 104. Holotype: ♂, Australia, Heron Island; QM. Kellen (1960):494, 495 (biology, distribution, habitat); Drake (1961): 302 (figures); Schuster (1965): 340 (distribution); Cobben (1968): 42, 43 (egg morphology), 43 (figure); Cobben (1970): 66, 68 (diagnosis, distribution, synonymy, morphology), 63, 67, 70, 72, 76, 79, 82 (figures); 69 (in key); 71, 73 (biology, habitat); Polhemus (1976): 244 (figures); Schuh et al. (1987): 260 (catalogue, distribution); Cobben (1987): 25 (distribution); Polhemus & Polhemus (2012): 333 (color figure, distribution). Omania samoensis Kellen, 1960: 495. Holotype: ♂, Samoa, Tutuila Island; USNM. (synonymized by Cobben 1970: 66). Schuster (1965): 340 (distribution). Material examined: AUSTRALIA: Queensland: Qld. Great Barrier Reef, Heron Island, 10-14. XI. 1957, leg. T.E. Woodward, paratypes 2 ♀♀ [lablled as Omania marksae]; Heron Island, 16-20.XII.1967, leg. S.R. Curtis, 4 exx; N. Qld. Great Barrier Reef, Lowe Isles, 19. VIII. 1954, leg. H. Marks, 1♂; “Australia Woodward” met melkz. beh. [= treated with lacic acid; no identification label]. NEW CALEDONIA: Îlot Croissant, dans empilement de madréporites mort [in a pile of dead Madrepora coral], 19.XI.1980, leg. L. Bigot, 1♂ 1♀, [îlot = a small island, which actually is a small coral reef]. SAMOA: Tutuila, 10.IV.1958, leg. W.R. Kellen, 3 exx.; “ Omania Samoa”, 1 ex. SINGAPORE: Pulau Blakang, Mati, 26.V.1969, leg. H.D. Murphy 34 exx.; Labrador on rock seashore, Feb.1961, leg. Murphy, 2 exx. [Labrador is a shorthand for Labrador Park, a nature reserve in SW Singapore]; (all in RMNH). Diagnosis: Body length of male (1.15–1.44), female (1.48–1.59). It can be distinguished from other species of Corallocoris by the following characteristics: the body is nearly black, the forewing has a distinct grey transverse fascia and grey apical margin (Fig. 7A–B, F). Habitat and biology. Corallocoris marksae was collected by searching volcanic rocks lying in the intertidal zone of a shallow, protected lagoon, which exposed 25 feet of beach at low tide and was covered by up to 2 feet of water at high tide (Kellen 1960). The motor behaviors of this species in the field were recorded by Kellen (1960) and he reared entire generations in the laboratory. Woodward (1958) recorded finding C. marksae mainly on the moist undersurface of the slabs of porous coral rock and on the sand and rock beneath when these rocks were exposed, together with Collembola, mites (Microtrombidium sp. and Eupodes sp.) and small beetles. Distribution: Australia (Great Barrier Reef); Malaysia (Malaya); New Caledonia (Îlot Croissant); Philippines (Badian, Luzon); Samoa (Tutuila Island); and Singapore (Sentosa Island) (Fig. 9A, Table 3)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on page 462, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Woodward, T. W. (1958) Studies on Queensland Hemiptera. Part III. A remarkable new intertidal saldid. University of Queensland Papers, 1, 101 - 110.","Kellen, W. R. (1960) A new species of Omania from Samoa, with notes on its biology (Heteroptera: Saldidae). Annals of the Entomological Society of America, 53, 494 - 499. https: // doi. org / 10.1093 / aesa / 53.4.494","Drake, C. J. (1961) Insects of Micronesia-Hemiptera: Saldidae. Bernice P. Bishop Museum, Honolulu, 7, 287 - 305.","Schuster, R. (1965) Faunistische Studien am Roten Meer (im Winter 1961 / 62). Teil 1: Litoralbewohnende Arthropoden terrestrischer Herkunft. Zoologische Jahrbuecher (Systematik), 92, 327 - 343.","Cobben, R. H. (1968) Evolutionary trends in Heteroptera. Part I. Eggs, architecture of the shell, gross embryology and eclosion. Center for Agricultural Publishing & Documentation, Wageningen, 475 pp.","Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Polhemus, J. T. (1976) Shore bugs (Hemiptera: Saldidae, etc.). In: Cheng, L. (Ed.), Marine insects. North-Holland Publishing Company, Amsterdam & Oxford & American Elsevier Publishing Company, New York, New York, pp. 225 - 262.","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Cobben, R. H. (1987) New African Leptopodomorpha (Heteroptera: Saldidae, Omaniidae, Leptopodidae), with an annotated checklist of Saldidae from Africa. II. New taxa of Saldidae (except the genus Saldula), Omaniidae, Leptopodidae, and a checklist of African shorebug. Revue de Zoologie Africaine, 101, 3 - 30.","Polhemus, J. T. & Polhemus, D. A. (2012) Guide to the aquatic Heteroptera of Singapore and Peninsular Malaysia. VIII. Leptopodomorpha, families Saldidae, Leptopodidae, and Omaniidae. The Raffles Bulletin of Zoology, 60, 329 - 341."]}

Omania coleoptrata Horváth, 1915 (Figs. 8A–B, 9A, Table 3) Omania coleoptrata Horváth, 1915: 598. Holotype: ♂, Oman, Muscat; HNHM. Poisson and Poisson (1943):123. (distribution, synonymy); Drake & Hoberlandt (1951): 5 (catalogue, distribution); Filippi (1957): 41 (distribution); Kellen (1960): 494 (distribution, habitat); Stichel (1960): 125 (catalogue, distribution); Schuster (1965): 125 (distribution); Cobben (1970): 63, 67, 70, 72, 76, 79, 85 (figures), 64 (diagnosis, distribution, morphology), 69 (in key), 71 (habitat); Cobben (1987): 25 (distribution); Schuh & Polhemus (1980): 6 (figures); Schuh et al. (1987): 261 (catalogue, distribution); Lindskog (1995): 141 (catalogue, distribution). Dollfusella minutissima China, 1938: 257. Holotype: ♂, Saudi Arabia, Sanafir Island; MNHN (synonymized by Poisson & Poisson 1943: 123). Material examined: EGYPT: 50 km N. of M. Alam, leg. Linnavuori, 1 ex.; Rotes Meer, Gubal strait, winter 1961/1962, 1 ♀; “ Omania coleoptrata ” 1 ex. [no further information]; (all in RMNH). Diagnosis: Body length of male (1.28–1.38), female (1.44–1.48). This taxon can be distinguished from the species of Corallocoris by the following characteristics: the body surface is uniformly black (Fig. 8A–B); a collumlike area of the head is demarcated from the vertex by a line that makes a sharp triangular incision anteromedially; the forewing lacks claval and R+M sutures (Fig. 8A–B); the grasping structure of the male is simple; the ponticulus basalis of the phallus has a dorsal bridge-like projection (see Cobben 1970). Habitat. Omania coleoptrata was found in rock crevices below the high tide level of the intertidal zone (China 1938), and in one instance was found living with a species of Ptiliidae (Coleoptera) and Halophiloscia sp. (Crustacea: Isopoda) in the same rock pieces (Schuster 1965). Distribution: Egypt (African Egypt, Sinai Peninsula); Oman (Muscat); Pakistan (the coast of Pakistan); and Saudi Arabia (Sanafir Island) (Fig. 9A, Table 3)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on pages 457-458, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Horvath, G. (1915) Novum genus peculliare Acanthiidarum. Annales Musei Nationalis Hungarici, 13, 598 - 599.","Poisson, R. A. & Poisson, A. (1943) Sur la synonymie de Dollfusella minutissima China 1938 et Omania coleoptrata Horvath 1915 (Hem. Acanthiidae). Bulletin de la Societe entomologique de France, 48, 123. [https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1943 _ num _ 48 _ 8 _ 15691] https: // doi. org / 10.3406 / bsef. 1943.15691","Drake, C. J. & Hoberlandt, L. (1951) Catalogue of genera and species of Saldidae (Hemiptera). Acta Entomologica Musei Nationalis Pragae, 26, 1 - 12. [1950]","Kellen, W. R. (1960) A new species of Omania from Samoa, with notes on its biology (Heteroptera: Saldidae). Annals of the Entomological Society of America, 53, 494 - 499. https: // doi. org / 10.1093 / aesa / 53.4.494","Stichel, W. (1960) Familia Saldidae, Leptopodidae. In Verzeichnis der Paliiarktischen Hemiptera-Heteroptera, III, 123 - 129.","Schuster, R. (1965) Faunistische Studien am Roten Meer (im Winter 1961 / 62). Teil 1: Litoralbewohnende Arthropoden terrestrischer Herkunft. Zoologische Jahrbuecher (Systematik), 92, 327 - 343.","Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Cobben, R. H. (1987) New African Leptopodomorpha (Heteroptera: Saldidae, Omaniidae, Leptopodidae), with an annotated checklist of Saldidae from Africa. II. New taxa of Saldidae (except the genus Saldula), Omaniidae, Leptopodidae, and a checklist of African shorebug. Revue de Zoologie Africaine, 101, 3 - 30.","Schuh, R. T. & Polhemus, J. T. (1980) Analysis of taxonomic congruence among morphological, ecological, and biogeographic data sets for the Leptopodomorpha (Hemiptera). Systematic Zoology, 29, 1 - 26. https: // doi. org / 10.2307 / 2412623","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Lindskog, P. (1995) Infraorder Leptopodomorpha. In: Auckema, B. & Rieger, C. (Eds.), Catalogue of Palearctic Heteroptera. Vol. 1. General introduction: Enicocephalomorpha & Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha. Netherlands Entomological Society, Amsterdam, pp. 115 - 141.","China, W. E. (1938) A new genus and species of Acanthiidae (Saldidae) from Sinai Peninsula. Memoires de l'Institut d'Egypte, 37, 255 - 257."]}

Corallocoris nauruensis (Herring & Chapman, 1967) (Figs. 8C–D, 9) Omania nauruensis Herring & Chapman 1967: 355. Holotype: ♂, Nauru Island; USNM. Cobben (1970): 68 (diagnosis), 69 (in key); Schuh (1987): 261 (catalogue, distribution); Cobben (1987): 25 (distribution); Schuh et al. (1987): 260 (catalogue, distribution); Schuh & Polhemus (1980): 5, 6, 7 (figures). Material examined: NAURU: Central Pacific, Nauru Island, 16.XI.1966, leg. H.C. Chapman, paratypes 2♂♂ 2♀♀ (RMNH). Diagnosis: Body length of male (1.21), female (1.30–1.40). C. nauruensis can be distinguished from other species of Corallocoris by the following characteristics: the pronotum and scutellum have a light pattern (Fig. 8A); the median fovea of the pronotal dome continues as a sulcus, which almost completely divides the dome into two lobes (Fig. 8A); a large basal and preapical blue-grey band is on the clavus, an irregular blue-grey patch on the base of corium extends narrowly down the lateral margin of the forewing, widening at the middle third and extending inward almost to the commissure, and a grey band is at the apical margin (Fig. 8A–B). Habitat. Corallocoris nauruensis was collected at low tide from pinnacles that were common in the reef, which completely surrounds Nauru Island (Herring & Chapman 1967). Distribution: Nauru Island (Fig. 9A, Table 3)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on page 462, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Herring, J. L. & Chapman, H. C. (1967) A new species of Omania from Micronesia (Hemiptera: Saldidae). Proceedings of the Entomological Society of Washington, 69, 353 - 359. [https: // www. biodiversitylibrary. org / partpdf / 56261]","Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Cobben, R. H. (1987) New African Leptopodomorpha (Heteroptera: Saldidae, Omaniidae, Leptopodidae), with an annotated checklist of Saldidae from Africa. II. New taxa of Saldidae (except the genus Saldula), Omaniidae, Leptopodidae, and a checklist of African shorebug. Revue de Zoologie Africaine, 101, 3 - 30.","Schuh, R. T. & Polhemus, J. T. (1980) Analysis of taxonomic congruence among morphological, ecological, and biogeographic data sets for the Leptopodomorpha (Hemiptera). Systematic Zoology, 29, 1 - 26. https: // doi. org / 10.2307 / 2412623"]}

Corallocoris xishaensis Luo, Wang & Chen, sp. nov. http://zoobank.org/ FFEBDBCB-050D-4742-85B3-2409D5F2E9BF (Figs. 1–5, 6, 9) Type material. Holotype (♂) CHINA: Hainan Province, Sansha Prefecture, Xisha, Lingyang Jiao Reef: 16°28'03"N, 111°35'58"E, 0 m.a.s.l., 2019-XI-12, leg. Jiu-Yang LUO, mounted on card (SYSBM). Paratypes: same data as holotype, mounted on cards (5♂♂, 3♀♀, SYSBM); same data as holotype, mounted on cards (1♂, 1♀, RMNH); CHINA: Hainan Province, Sansha Prefecture, Xisha, Lingyang Jiao Reef: 16°28'03"N, 111°35'58"E, 0 m.a.s.l., 2019-XI-12, leg. Yan-Hui WANG, mounted on card (4♂♂, 8♀♀, SYSBM); CHINA: Hainan Province, Sansha Prefecture, Xisha, Yongxing Dao: 2019-IV-18, leg. Qiang XIE, mounted on cards (1♂, SYSBM). Diagnosis: Corallocoris xishaensis sp. nov. is similar to C. marksae in size and habitus, but it can be distinguished from the latter species by following characteristics: (1) body greyish-black, without distinct grey fascia on the the middle of forewing (Fig. 1A–B, D–E), (body nearly black, with a transverse grey fascia near the middle of forewing in latter species [Fig. 7A–B, F]); (2) cephalic trichobothria pairs 1 and 2 subequal in length with pair 3 (Fig. 1B, E), (distinctly shorter than pairs 3 in latter species [Fig. 7B]); (3) the coxae are yellowish-brown, entire trochanters and femora, except for subapices, are whitish-yellow (Fig. 1B–C, E–F), (coxae, entire trochanters and femora, except for apices, are dark-brown in latter species [Fig. 7B–C]); (4) the proepisternum is whitish-yellow (Fig. 1B–C, E–F), (the proepisternum is nearly black in latter species [Fig. 7B–C]); (5) the pygophore is round in caudal view (Fig. 5D), (pygophore is high in caudal view in latter species [Fig. 5M]); (6) the paramere is curved near the middle (Fig. 5H–J), (the paramere is slightly curved near the middle in latter species [Fig. 5P]). Description: Size: Male, length 1.13–1.22 mm, greatest width 0.54–0.59 mm; female, length 1.24–1.38 mm, greatest width 0.63–0.70 mm. Color and body surface: Body and collum-like area of head greyish-black and dull (Fig. 1A–E); most of head and abdomen black; eyes red, ocelli dark red; elevated spots on head whitish-yellow; antennal segment I whitish-yellow, segment II slightly darker than segment I, and darkening from base to apex, segment III and IV blackish-brown, segment III slightly lighter than segment IV; labrum and labium light-brown to brown, apex of labial segment IV blackish-brown; pronotum, scutellum and upper surface of forewing with moderately dense, semierect brown setae (Fig. 1A, D); apical portion and hypocostal lamina of forewing yellowishbrown to dark-brown (Fig. 1B, E); proepisternum whitish-yellow; coxae yellowish-brown, entire trochanters and femora except for subapices whitish-yellow; tibiae and tarsi whitish-yellow, with blackish-brown distal end. Most of head with dense, relatively thin, silver setae (Figs. 3A–B, 4A); collar of head, surface of thorax and upper surface of forewing with dense, grey microtrichia (Fig. 4A–B), lower surface of forewing with dense, short microtrichia (Fig. 3F–H); antennae, labium, legs and abdomen with moderately dense, silver setae (Fig. 2B, E, H–J). Brachypterous male. Structures: Body oval. Body length 2.03–2.09× greatest width. Head. Wider than long, greatest width across eyes 2.63–2.85× median length, with three pairs of black cephalic trichobothria, two pairs on vertex behind ocelli, and one pairs arising from light, elevated spots before ocelli (Figs. 2B, E, 3B, 4A), cephalic trichobothria pairs 1 and 2 subequal in length with pair 3 (Fig. 1B, E); eyes large, width of eyes 1.14× vertex width of across ocelli, with three pairs of dark-brown trichobothria (Figs. 2B, E, 4A), trichobothria on eyes shorter and thinner than cephalic trichobothria; antennal length subequal to body width; segments I and III subequal in length; segment I thickest and segment IV longest (Fig. 2B, E); ratio of antennal segments I to IV = 1: 1.36: 1.10: 1.71. Apex of labium reaching to hind coxa (Fig. 1B–C, E–F); labial segment I widest, subequal to segment II in length, segment III longest, ratio of labial segments I to IV = 1.28: 1: 3.59: 2.69. Thorax. Pronotum short and trapezoid (Figs. 2B, 3A), median length about 0.5× humeral width; collar distinct, narrower than posterior lobe; anterior lobe elevated and deeply concave at fovea (Figs. 2B, 3A, D, 4A, C); with three pairs of black pronotal trichobothria (Figs. 2B, 3D); scutellum triangular, wider than long, ratio of length / width = 1: 1.44; lateral margin of scutellum with small concavity centrally (Figs. 2A, 3A, E); middle of scutellum base ridge-like, longitudinally convex (Fig. 2A); mesosternum with two small sub-transverse ridges (Fig. 4E–F), metasternum with high middle longitudinal ridge (Fig. 4E); forewing with claval and R+M sutures, bases of sutures with deep round pit, R+M suture ends at about two-thirds of forewing, claval suture ends at about four-sevenths of forewing (Fig. 2D); legs stout, coxae and femora thick, tibiae and tarsi slender; femora and tibiae with thick spines; tarsal formula 3-3-3 (Fig. 2H–J); forefemur and middle femur longer than tibia, forefemur about 1.14× length of foretibia, middle femur about 1.16× length of middle tibia; hind coxae with setose area (Figs. 2J, 4G–H); hind femur shorter than hind tibia, tibia about 1.18× length of femur. Pregenital abdomen. Mediotergites less sclerotized than sterna, with broad membranous area (Fig. 2G, K); mediotergites V–VII more sclerotized than I–IV; laterotergites less sclerotized than sterna, well-delimited with mediotergites and sterna (Fig. 2G, H); laterotergite III expanded, with strongly sclerotized margin (Fig. 2G); sterna strongly sclerotized; middle of basal margin of sternum I with cleft; segment VIII cylindrical shaped. External genitalia. Pygophore cylindrical (Fig. 5A–D), posterior area of venter expanded (Fig. 5B); with parandria protruding posterior margin of genital capsule opening (Fig. 5B); parameres club-shaped, with slender and distally curved bases (Fig. 5H–J); articulatory apparatus sclerotized (Fig. 5E–G); ligamentous processes with wide base (Fig. 5E); caudal side of phallotheca with swellings (Fig. 5E); more than half of endosoma with spicule (Fig. 5E–F). Brachypterous female. Structures: Similar to male, except for the following: Body length 1.94–2.00× greatest width. Head. Greatest width across eyes 2.63–2.90× median length; width of eyes 1.22× vertex width of across ocelli; ratio of antennal segments I to IV = 1: 1.42: 1.10: 1.82; ratio of labial segments I to IV = 1.06: 1: 2.91: 2.29. Thorax. Pronotum short and trapezoid, median length about 0.47× humeral width; forefemur about 1.16× length of foretibia. Pregenital abdomen. Laterotergite III normally developed. External genitalia. Gynatrium ellipsoidal; spermatheca oval; accessory canal more sclerotized (Fig. 5K–L). Measurements see Table 1. Etymology: The specific epithet refers to the type locality area, the Xisha Islands in the South China Sea; adjective. Biology: Specimens of Corallocoris xishaensis sp. nov. were collected in the intertidal zone. They spend periods submerged by high tides in rock crevices and then emerge at low tide, and were frequently found in crevices and holes of coral rocks (Fig. 6A–B). Distribution: This species is so far known only from Lingyang Jiao and Yongxing Dao, Xisha Islands in the South China Sea (Figs. 9A–B; Table 3)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on pages 464-465, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747

Corallocoris satoi (Miyamoto, 1963) (Fig. 9) Omania satoi Miyamoto, 1963: 39. Holotype: ♀, Japan, Takarajima; ELKU. Cobben (1970): 68 (diagnosis), 69 (in key); Schuh et al. (1987): 261 (catalogue, distribution); Cobben (1987): 25 (distribution); Lindskog (1995): 140 (catalogue, distribution); Hayashi et al. (2016): 376 (catalogue, distribution). Diagnosis: Body length of female (1.45). It can be distinguished from other species of Corallocoris by the following characteristics: cephalic trichobothria pair 2 is in front of pair 1; cephalic trichobothria pair 3 does not arise from light spots; the fourth antennal segment is twice as long as the third segment. Distribution: Japan (Ryukyu Islands) (Fig. 9A, Table 3)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on page 464, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Miyamoto, S. (1963) New halophilous saldids from the Tokara Islands. Sieboldia, 3, 39 - 49.","Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Cobben, R. H. (1987) New African Leptopodomorpha (Heteroptera: Saldidae, Omaniidae, Leptopodidae), with an annotated checklist of Saldidae from Africa. II. New taxa of Saldidae (except the genus Saldula), Omaniidae, Leptopodidae, and a checklist of African shorebug. Revue de Zoologie Africaine, 101, 3 - 30.","Lindskog, P. (1995) Infraorder Leptopodomorpha. In: Auckema, B. & Rieger, C. (Eds.), Catalogue of Palearctic Heteroptera. Vol. 1. General introduction: Enicocephalomorpha & Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha. Netherlands Entomological Society, Amsterdam, pp. 115 - 141.","Hayashi, M., Tomokuni, M., Yoshizawa, K. & Ishikawa, T. (Eds.) (2016) Catalogue of the Insects of Japan. Vol. 4. Paraneoptera. Touka-shobo, Fukuoka, 376 pp. [in Japanese]"]}

Genus Omania Horváth, 1915 Omania Horváth, 1915: 598. Type species: Omania coleoptrata Horváth, 1915 by monotypy. Drake & Hoberlandt (1951): 5 (catalogue, distribution); Kellen (1960): 494 (distribution); Stichel (1960): 125 (catalogue); Drake (1961): 302 (distribution, ecology, habitat); Herring & Chapman (1967): 354 (distribution, in key); Cobben (1970): 64, 66 (distribution, morphology), 69 (in key); Polhemus (1976): 239 (distribution, diagnosis); Schuh & Polhemus (1980): 19 (distribution, habitat, phylogenetic relationships); Schuh et al. (1987): 261 (catalogue, distribution); Lindskog (1995): 140 (catalogue, distribution). Dollfusella China 1938: 255. Type species: Dollfusella minutissima China 1938 by original designation; (synonymized by Poisson & Poisson 1943: 123 (synonymy)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on page 456, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Horvath, G. (1915) Novum genus peculliare Acanthiidarum. Annales Musei Nationalis Hungarici, 13, 598 - 599.","Drake, C. J. & Hoberlandt, L. (1951) Catalogue of genera and species of Saldidae (Hemiptera). Acta Entomologica Musei Nationalis Pragae, 26, 1 - 12. [1950]","Kellen, W. R. (1960) A new species of Omania from Samoa, with notes on its biology (Heteroptera: Saldidae). Annals of the Entomological Society of America, 53, 494 - 499. https: // doi. org / 10.1093 / aesa / 53.4.494","Stichel, W. (1960) Familia Saldidae, Leptopodidae. In Verzeichnis der Paliiarktischen Hemiptera-Heteroptera, III, 123 - 129.","Drake, C. J. (1961) Insects of Micronesia-Hemiptera: Saldidae. Bernice P. Bishop Museum, Honolulu, 7, 287 - 305.","Herring, J. L. & Chapman, H. C. (1967) A new species of Omania from Micronesia (Hemiptera: Saldidae). Proceedings of the Entomological Society of Washington, 69, 353 - 359. [https: // www. biodiversitylibrary. org / partpdf / 56261]","Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Polhemus, J. T. (1976) Shore bugs (Hemiptera: Saldidae, etc.). In: Cheng, L. (Ed.), Marine insects. North-Holland Publishing Company, Amsterdam & Oxford & American Elsevier Publishing Company, New York, New York, pp. 225 - 262.","Schuh, R. T. & Polhemus, J. T. (1980) Analysis of taxonomic congruence among morphological, ecological, and biogeographic data sets for the Leptopodomorpha (Hemiptera). Systematic Zoology, 29, 1 - 26. https: // doi. org / 10.2307 / 2412623","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Lindskog, P. (1995) Infraorder Leptopodomorpha. In: Auckema, B. & Rieger, C. (Eds.), Catalogue of Palearctic Heteroptera. Vol. 1. General introduction: Enicocephalomorpha & Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha. Netherlands Entomological Society, Amsterdam, pp. 115 - 141.","China, W. E. (1938) A new genus and species of Acanthiidae (Saldidae) from Sinai Peninsula. Memoires de l'Institut d'Egypte, 37, 255 - 257.","Poisson, R. A. & Poisson, A. (1943) Sur la synonymie de Dollfusella minutissima China 1938 et Omania coleoptrata Horvath 1915 (Hem. Acanthiidae). Bulletin de la Societe entomologique de France, 48, 123. [https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1943 _ num _ 48 _ 8 _ 15691] https: // doi. org / 10.3406 / bsef. 1943.15691"]}

Genus Corallocoris Cobben, 1970 Corallocoris Cobben, 1970: 62. Type species: Omania marksae Woodward, 1958 by original designation. Polhemus (1976): 240 (distribution, note); Schuh & Polhemus (1980): 19 (distribution, habitat, phylogenetic relationships); Schuh et al. (1987): 260 (catalogue, distribution); Lindskog (1995): 140 (catalogue, distribution)., Published as part of Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping & Xie, Qiang, 2022, First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha), pp. 449-467 in Zootaxa 5195 (5) on page 458, DOI: 10.11646/zootaxa.5195.5.4, http://zenodo.org/record/7223747, {"references":["Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90.","Woodward, T. W. (1958) Studies on Queensland Hemiptera. Part III. A remarkable new intertidal saldid. University of Queensland Papers, 1, 101 - 110.","Polhemus, J. T. (1976) Shore bugs (Hemiptera: Saldidae, etc.). In: Cheng, L. (Ed.), Marine insects. North-Holland Publishing Company, Amsterdam & Oxford & American Elsevier Publishing Company, New York, New York, pp. 225 - 262.","Schuh, R. T. & Polhemus, J. T. (1980) Analysis of taxonomic congruence among morphological, ecological, and biogeographic data sets for the Leptopodomorpha (Hemiptera). Systematic Zoology, 29, 1 - 26. https: // doi. org / 10.2307 / 2412623","Schuh, R. T., Galil, B. & Polhemus, J. T. (1987) Catalog and bibliography of Leptopodomorpha (Heteroptera). Bulletin of the American Museum of Natural History, 185, 243 - 406.","Lindskog, P. (1995) Infraorder Leptopodomorpha. In: Auckema, B. & Rieger, C. (Eds.), Catalogue of Palearctic Heteroptera. Vol. 1. General introduction: Enicocephalomorpha & Dipsocoromorpha, Nepomorpha, Gerromorpha & Leptopodomorpha. Netherlands Entomological Society, Amsterdam, pp. 115 - 141."]}

Kokeshia drepanoides sp. nov. urn:lsid:zoobank.org:act: B310837E-53EA-4E34-B9E8-0E16DE44D4CA Figs 9��� 10, 26E, 29A, 32 Diagnosis Kokeshia drepanoides sp. nov. can be recognized from all other species of Kokeshia by tergite VIII subdivided into two hemitergites, left hemitergite VIII with a flattened, curved, sickle-shaped process. (Figs 9D, E, G, 10 A���B). Etymology The species epithet is derived from the Greek adjective �� drepanoides ��, means ��sickle-shaped��, and refers to the curved, sickle-shaped process of tergite VIII. Material examined Holotype CHINA ��� ♂; Yunnan Province, Honghe, Jinping County, Adebo Town; 22��54���21��� N, 103��13���49��� E; ca 1480 m a.s.l.; 30 Jul. 2016; Jiu-Yang Luo leg.; NKUM. Paratypes CHINA ��� 7 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 5). Total body length 1.27 / 1.20���1.31; length of head 0.22/ 0.19���0.23, maximum width across eyes 0.36 / 0.36���0.37, interocular distance 0.25 / 0.24���0.25; length of antennal segment I 0.07 / 0.06���0.07, segment II 0.07 / 0.07, segment III 0.27 / 0.29���0.30, segment IV 0.33 / 0.32���0.33; length of labial segment I 0.07 / 0.07���0.08, segment II 0.05 / 0.04���0.05, segment III 0.03 / 0.04, segment IV 0.11 / 0.10���0.12; middle length of pronotum 0.34 / 0.35���0.37, length of collar 0.05 / 0.05���0.06, humeral width 0.60 / 0.61���0.63; length of forewing 1.16 / 1.07���1.15; length of fore femur 0.27 / 0.29���0.30, fore tibia 0.33 / 0.31���0.33, fore tarsus 0.12 / 0.12���0.13; length of middle femur 0.34 / 0.32���0.34, middle tibia 0.34 / 0.30���0.35, middle tarsus 0.11 / 0.11���0.12; length of hind femur 0.33 / 0.36���0.38, hind tibia 0.56 / 0.54���0.56, hind tarsus 0.17 / 0.17���0.18; greatest width of abdomen 0.55 / 0.52���0.55. MACROPTEROUS. Small sized (about 1.2���1.31 mm). COLORATION. Body light brown to brown, appendages yellowish brown (Fig. 9A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.67���0.69 �� length of maximum width across eyes. THORAX. Humeral width 1.69���1.77 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 26E). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, left hemitergite VIII with a flattened, curved, sickle-shaped process, and horizontally directed mesad; prominent setae near its inner margin (Figs 9D���E, G, 10 A���B). Right hemitergite VIII simple, posterolateral portion elongated subtrapezoidal, projecting posteriad, prominent setae near posterior margin (Figs 9E���F, 10C). GENITALIA. Pygophore asymmetrical, dorsal surface with rows of microtrichia on transverse projections, posterior margin with curved setae. Parameres strongly asymmetrical, left paramere (Figs 9E, 10D���E) with broadly rounded laminate extension directed dorsal near base, and with flattened and elongate distal projection, and curved inward from middle of it; right paramere (Figs 9E, 10F���G) with flattened, blunt and relatively slender distal projection. Apical portion of phallus (Figs 9D���E, 10E) tubular, short, forming half coil outside of pygophore, with triangular laminate at middle portion. Female Unknown. Distribution China: Yunnan (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 18-21, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Key to the species of Kokeshia Miyamoto, 1960 1. Tergite VIII not subdivided into two hemitergites, only a long,flattened, acuminate process on the right portion of tergite VIII, bent at about ⅓ from its apex (Fig. 28A) (China)..... K. acutiformis sp. nov. ��� Tergite VIII subdivided into left and right hemitergites................................................................... 2 2. Process of left hemitergite VIII elongated, needle-shaped, flagelliform or filiform......................... 3 ��� Process of left hemitergite VIII not needle-shaped, flagelliform or filiform.................................... 6 3. Process of left hemitergite VIII significantly long, almost reaching the width of tergite VII, the process with an appendage split into three to five branches (Fig. 17D; Ren & Zheng 1992: 191, figs 21���23) (China).................................................................................... K. hsiaoi Ren & Zheng, 1992 ��� Process of left hemitergite VIII relatively long, more than half the width of tergite VII, the process without any appendage..................................................................................................................... 4 4. Process of left hemitergite VIII filiform, rectangularly bent, with spiniform microtrichia on its apex (��tys 1985: 200, figs 37���38, 40) (Nepal)........................................................... K. similis ��tys, 1985 ��� Process of left hemitergite VIII needle-shaped or flagelliform, not filiform, without spiniform microtrichia on its apex.................................................................................................................... 5 5 Process of left hemitergite VIII slender, flagelliform, curved from approximal middle portion (R��dei 2008: 244, fig. 6; 248, fig. 14) (Thailand)....................................................... K. oroszi R��dei, 2008 _ Process of left hemitergite VIII slender, needle-shaped, curved from base (Fig. 29D) (China)................................................................................................................... K. pengae sp. nov. 6. Process of left hemitergite VIII with a broad brush-like structure formed by hair-like setae.......... 7 ��� Process of left hemitergite VIII without broad brush-like structures formed by hair-like setae...... 8 7. Brush-like structure on left hemitergite VIII formed by several narrow, hair-like setae; apical portion of phallus forming about one incomplete coil outside of pygophore (��tys 1985: 192, figs 15, 17; 194, figs 19���20; 196, fig. 26) (Nepal).................................................................... K. martensi ��tys, 1985 ��� Brush-like structure on left hemitergite VIII formed by less numerous and distinctly thicker setae; apical portion of phallus forming more than a half coil outside of pygophore (R��dei 2008: 244, fig. 5; 246, fig. 7) (India...................................................................................... K. stysi R��dei, 2008 8. Process of left hemitergite VIII triangular or finger-shaped............................................................. 9 ��� Process of left hemitergite VIII not triangular or finger-shaped..................................................... 14 9. Process of left hemitergite VIII triangular or subtriangular............................................................ 10 ��� Process of left hemitergite VIII finger-shaped................................................................................ 12 10. Process of left hemitergite VIII with a broad, thick near triangular process, with about 8 thick spiniform microtrichia at its right posterolateral margin (R��dei et al. 2012: 30, fig. 2; 31, fig. 4; 33, figs 8���10) (China)......................................................................... K. zhengi R��dei, Ren & Bu, 2012 ��� Process of left hemitergite VIII near triangular, not broad..............................................................11 11. Process of left hemitergite VIII elongated triangular, right margin with about nine, thick, nearly parallel spiniform microtrichia (Fig. 28C) (China)..................................................... K. bui sp. nov. ��� Process of left hemitergite VIII subtriangular, without thick spiniform microtrichia (Fig. 30B) (China)..................................................................................................................... K. renae sp. nov. 12. Process of left hemitergite VIII elongate, curved, finger-shaped, with one in apex, and two more thick spiniform microtrichia in ventral side of distal portion, right hemitergite VIII with lamellate process above tergite VII (Fig. 28B) (China)............................................................. K. baii sp. nov. ��� Process of left hemitergite VIII moderate long, finger-shaped, right hemitergite VIII without lamellate process above tergite VII................................................................................................. 13 13. Process of left hemitergite VIII curved, finger-shaped, with about six thick spiniform microtrichia at ventral side of apex; apical portion of phallus forming about two and a half coils outside of pygophore (Fig. 30C) (China)....................................................................... K. weirauchae sp. nov. ��� Process of left hemitergite VIII not curved, finger-shaped, with out thick spiniform setae;apical portion of phallus forming incomplete one coil outside of pygophore (Fig. 28D) (China).... K. caii sp. nov. 14. Process of left hemitergite VIII long, thick and horizontally directed, with numerous thick spiniform microtrichia at its ventral surface; apical portion of phallus forming about one coil outside of pygophore (Fig. 30D; R��dei et al. 2012: 30, fig. 1; 31, fig. 3; 33, figs 5���7) (China)............................................................................................... K. xiei R��dei, Ren & Bu, 2012 ��� Process of left hemitergite VIII not as above.................................................................................. 15 15. Process of left hemitergite VIII short and thick, hook-shaped, rectangularly bent; apical portion of phallus forming about one and a half coils outside of pygophore (Miyamoto 1960: 171, pl. 19h���i) (Japan)....................................................................................................... K. esakii Miyamoto, 1960 ��� Process of left hemitergite VIII not as above.................................................................................. 16 16. Process of left hemitergite VIII flattened, broad and curved, sickle-shaped; apical portion of phallus forming about a half coil outside of pygophore (Fig. 29A) (China)............ K. drepanoides sp. nov. ��� Process of left hemitergite VIII not as above.................................................................................. 17 17. Process of left hemitergite VIII flattened, slender and curved, sword-shaped; apical portion of phallus forming about one coil outside of pygophore (Fig. 30A) (China)........................... K. redeii sp. nov. ��� Process of left hemitergite VIII not as above.................................................................................. 18 18. Process of left hemitergite VIII nearly quadrangular, left portion with a raised projection, which bearing about seven thick basal-curved spiniform microtrichia, posterior margin of the process with about seven thick basal-curved spiniform microtrichia; apical portion of phallus forming more than one coil outside of pygophore (Fig. 29B) (China)...................................................... K. hilli sp. nov., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 45-48, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878, {"references":["Miyamoto S. 1960. A new genus of Schizopterinae from Japan (Heteroptera, Dipsocoridae). Sieboldia 2 (3): 163 - 170.","Redei D., Ren S. Z. & Bu W. J. 2012. Two new species of Kokeshia from China (Hemiptera: Heteroptera: Schizopteridae). Zootaxa 3497 (1): 29 - 36. https: // doi. org / 10.11646 / zootaxa. 3497.1.3","Ren S. Z. & Zheng L. Y. 1992. New species and new records of Dipsocoromorpha (Hemiptera: Heteroptera) from China. Entomotaxonomia 14: 187 - 196.","Stys P. 1985. Two new species of Kokeshia (Heteroptera, Schizopteridae) from Nepal and appraisal of alleged synapomorphies of Paraneoptera. Acta Entomologica Bohemoslovaca 82: 187 - 205.","Redei D. 2008. Two new species of Kokeshia from India and Thailand (Hemiptera: Heteroptera: Schizopteridae). Acta Entomologica Musei Nationalis Pragae 48 (2): 241 - 250. Available from https: // www. aemnp. eu / data / article- 1172 / 1153 - 48 _ 2 _ 241. pdf [accessed 17 Feb. 2022]."]}

Kokeshia bui sp. nov. urn:lsid:zoobank.org:act: 2FE947B4-B43C-4039-9372-2F0BC071C017 Figs 5���6, 26C, 28C, 32 Diagnosis Kokeshia bui sp. nov. can be recognized from other species of Kokeshia by male hemitergite VIII (Figs 5D���G, 6A���B) elongate, the left lateral margin slightly concave from about ⅓ of its apex, the posterior portion with an elongated triangular process, surpassing posterior margin of pygophore, right lateral margin of it with about nine subparallel, thick spiniform microtrichia. Etymology The species epithet is derived from and dedicated to Prof. Wenjun Bu (Institute of Entomology, Nankai University, China), recognizing his contributions to the study of taxonomy, phylogeny and biogeography of Heteroptera, and the taxonomy of Cecidomyiidae (Diptera). Material examined Holotype CHINA ��� ♂; Yunnan Province, Lincang City, Cangyuan County, Banhong Town; 23��13���53.8��� N, 99��2���42.94��� E; ca 920 m a.s.l.; 4 Aug. 2019; Qiang Xie and Yu Men leg.; light trap; SYSBM. Paratypes CHINA ��� 10 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 5). Total body length 1.20 / 1.17���1.24; length of head 0.20 / 0.19���0.21, maximum width across eyes 0.41 / 0.37���0.39, interocular distance 0.26 / 0.25���0.26; length of antennal segment I 0.07 / 0.07���0.08, segment II 0.07 / 0.07���0.08, segment III 0.27 / 0.24���0.29, segment IV 0.31 / 0.28���0.35; length of labial segment I 0.07 / 0.07, segment II 0.06 / 0.04���0.05, segment III 0.05 / 0.03���0.05, segment IV 0.11 / 0.10���0.11; middle length of pronotum 0.35 / 0.34���0.36, length of collar 0.06 / 0.06���0.07, humeral width 0.62 / 0.60���0.64; length of forewing 0.95 / 0.88���1.00; length of fore femur 0.28 / 0.27���0.30, fore tibia 0.32 / 0.31���0.32, fore tarsus 0.12 / 0.11���0.14; length of middle femur 0.33 / 0.29���0.34, middle tibia 0.31 / 0.31���0.34, middle tarsus 0.12 / 0.11���0.12; length of hind femur 0.37 / 0.36���0.38, hind tibia 0.51 / 0.47���0.55, hind tarsus 0.18 / 0.17���0.19; greatest width of abdomen 0.51 / 0.51���0.54. MACROPTEROUS. Small sized (about 1.17���1.24 mm). COLORATION. Body brown, appendages yellowish brown (Fig. 5A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.63���0.68 �� length of maximum width across eyes. THORAX. Humeral width 1.67���1.78 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An distinctly darker than wing surface, other veins darker than wing surface (Fig. 26C). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, left hemitergite VIII with elongated triangular process, right margin with about 9 subparallel, thick spiniform microtrichia (Figs 5E, G, 6 A���B). The basal portion of process bearing numerous small round projections. Short to moderate length microtrichia located at surface of process, ends of these microtrichia split into 2 to 5 branches (Fig. 5G). Right hemitergite VIII subtriangular, posterolateral portion projecting posteriad, with thin setae at its posterolateral margin. GENITALIA. Pygophore slightly asymmetrical, dorsal surface densely covered with groups of setae on small, tightly connected projections, posterior area with relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 5E, 6E���F) with broadly rounded laminate extension directed dorsal near the base, and with flattened and elongate distal projection slightly curved inward; right paramere (Figs 5E, 6G���H) with flattened and blunt distal projection. Apical portion of phallus (Figs 5D���E, 6D) tubular, relatively long, forming one complete coil outside of pygophore. Female Unknown. Distribution China: Yunnan (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 12-15, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Kokeshia weirauchae sp. nov. urn:lsid:zoobank.org:act: 884C3441-266D-4AB0-BA4C-2E1FE87BB781 Figs 21���22, 27C, 30C, 32 Diagnosis The male of K. weirauchae sp. nov. is similar to K. caii sp. nov., but can be recognized from the latter by tergite VIII subdivided into two hemitergites, left hemitergite VIII with a curved, finger-shaped process, with about six thick spiniform microtrichia at ventral side (Figs 21D���G, 22A, H). Etymology The species epithet is derived from and dedicated to Prof. Christiane Weirauch (Department of Entomology, University of California, Riverside, USA), recognizing her contributions to the study of taxonomy, morphology and phylogeny of Heteroptera and Dipsocoromorpha. Material examined Holotype CHINA ��� ♂; Yunnan Province, Honghe, Jinping County, Adebo Town; 23��18���44.65��� N, 99��4���39.48��� E; ca 1480 m a.s.l.; 30 Jul. 2016; Jiu-Yang Luo leg.; NKUM. Paratypes CHINA ��� 4 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 4). Total body length 1.23 / 1.22���1.30; length of head 0.23 / 0.20���0.22, maximum width across eyes 0.39 / 0.39���0.40, interocular distance 0.24 / 0.24���0.25; length of antennal segment I 0.07 / 0.07, segment II 0.08 / 0.07, segment III 0.25 / 0.27, segment IV 0.32 / 0.31���0.33; length of labial segment I 0.07 / 0.06���0.07, segment II 0.05 / 0.05, segment III 0.04 / 0.03���0.05, segment IV 0.10 / 0.09���0.11; middle length of pronotum 0.35 / 0.34���0.36, length of collar 0.06 / 0.05���0.06, humeral width 0.63 / 0.61���0.63; length of forewing 1.09 / 1.00���1.11; length of fore femur 0.29 / 0.29���0.30, fore tibia 0.31 / 0.30���0.32, fore tarsus 0.11 / 0.11���0.13; length of middle femur 0.33 / 0.31���0.34, middle tibia 0.33 / 0.30���0.33, middle tarsus 0.11 / 0.11���0.13; length of hind femur 0.35 / 0.34���0.37, hind tibia 0.52 / 0.50���0.54, hind tarsus 0.17 / 0.16���0.18; greatest width of abdomen 0.49 / 0.52���0.53. MACROPTEROUS. Small sized (about 1.22���1.30 mm). COLORATION. Body brown, appendages yellowish brown (Fig. 21A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.60���0.64 �� length of maximum width across eyes. THORAX. Humeral width 1.72���1.85 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 27C). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, left hemitergite VIII with curved, finger-shaped process, with about six thick spiniform microtrichia at its ventral side; besides, numerous semi-erect microtrichia at its surface, and ends of microtrichia unbranched or split into two to four branches. Basal portion of process with sparsely short microtrichia (Fig. 21D���E, G). Right hemitergite VIII with triangular posterolateral portion projecting posteriad (Figs 21F, 22G). GENITALIA. Pygophore simple, asymmetrical, dorsal surface with small, relative dense round projections and short setae, posterior area with relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 21E, 22C���D) with near round laminate extension directed dorsal near base, and with flattened and broad distal projection, and slightly curved inward; right paramere (Figs 21E, 22E���F) with flattened, blunt distal projection. Apical portion of phallus (Figs 21D���E, 22G) slender and tubular, forming more than two and a half coils outside of pygophore. Female Unknown. Distribution Yunnan Province, China (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 38-41, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Kokeshia pengae sp. nov. urn:lsid:zoobank.org:act: 85B2D5C7-6449-4606-BF7E-19DA4285CCBF Figs 15���16, 26H, 29D, 32 Diagnosis The male of K. pengae sp. nov. is similar to K. oroszi, but can be distinguished from the latter species by an obviously large body size 1.20���1.29 mm, whereas 1.04 in K. oroszi; process of left hemitergite VIII flagelliform, produced from left margin of left hemitergite VIII, whereas in K. oroszi, process of left hemitergite VIII flagelliform, produced from posterior margin of left hemitergite VIII; apical portion of phallus short, slightly curved, forming about a half coil outside of pygophore, whereas in K. oroszi, apical portion of phallus longer and curved, forming about one incomplete coil outside of pygophore. Etymology The species epithet is derived from and dedicated to Prof. Yanqiong Peng (Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, China), appreciating her kind help during our field works in Xishuangbanna and recognizing her contributions to the study of co-evolution of fig and fig wasps. Material examined Holotype CHINA ��� ♂; Yunnan Province, Xishuangbanna, Mengla County, Menglun Town, XTBG; 21��56���6.06��� N, 101��15���8.18��� E; ca 570 m a.s.l.; 19 Jul. 2019; Jiu-Yang Luo leg.; SYSBM. Paratypes CHINA ��� 3 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 3). Total body length 1.20 / 1.20���1.29; length of head 0.21 / 0.18���0.19, maximum width across eyes 0.35 / 0.34���0.35, interocular distance 0.25 / 0.22���0.25; length of antennal segment I 0.08 / 0.07, segment II 0.08 / 0.07���0.08, segment III 0.30 / 0.29, segment IV 0.30 / 0.31���0.35; length of labial segment I 0.06 / 0.07, segment II 0.04 / 0.05, segment III 0.04 / 0.04���0.05, segment IV 0.10 / 0.11; middle length of pronotum 0.37 / 0.32���0.33, length of collar 0.06 / 0.05���0.06, humeral width 0.60 / 0.56���0.59; length of forewing 1.05 / 1.04���1.10; length of fore femur 0.30 / 0.29���0.32, fore tibia 0.33 / 0.31���0.33, fore tarsus 0.11 / 0.12���0.13; length of middle femur 0.31 / 0.30���0.32, middle tibia 0.30 / 0.33���0.34, middle tarsus 0.13 / 0.11���0.12; length of hind femur 0.38 / 0.34���0.37, hind tibia 0.55 / 0.52���0.53, hind tarsus 0.18 / 0.17���0.18; greatest width of abdomen 0.51 / 0.50���0.55. MACROPTEROUS. Small sized (about 1.20���1.29 mm). COLORATION. Body light brown to brown, appendages yellowish brown (Fig. 15A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.63���0.74 �� length of maximum width across eyes. THORAX. Humeral width 1.62���1.84 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 26H). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, left hemitergite VIII with slender needle-shaped process, curved from base (Figs 15D��� E, G, 16A, C). Basal portion of process with sparsely microtrichia. Right hemitergite VIII with long, triangular posterolateral portion projecting posteriad, bearing microtrichia at apex (Figs 15E���F, 16B). GENITALIA. Pygophore simple, asymmetrical, dorsal surface with dense microtrichia, posterior area with relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 15E, 16D���E) with broadly rounded laminate extension directed dorsal near base, and with flattened and tapering distal projection, and curved inward; right paramere (Figs 15E, 16F���G) with flattened, round and relatively slender distal projection. Apical portion of phallus (Figs 15D���E, 16H) tubular, short, forming about �� coil outside of pygophore. Female Unknown. Distribution Yunnan Province, China (Fig. 20)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 28-31, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Kokeshia redeii sp. nov. urn:lsid:zoobank.org:act: B7AC7720-77E7-4DB3-9FD3-BB516BAEDE60 Figs 17���18, 27A, 30A, 32 Diagnosis Kokeshia redeii sp. nov. can be recognized from all other species of Kokeshia by tergite VIII subdivided into two hemitergites, left hemitergite VIII with a flattened, slender, curved, sword-shaped process, horizontally directed mesad (Figs 17D���E, G, 18A). Etymology The species epithet is derived and dedicated to Dr R��dei D��vid (NCHU) recognizing his contributions to the study of taxonomy and morphology of Dipsocoromorpha and Heteroptera. Material examined Holotype CHINA ��� ♂; Yunnan Province, Xishuangbanna, Mengla County, Menglun Town, Baka village; 21��57���58��� N, 101��12���33��� E; ca 580 m a.s.l.; 13 May. 2019; Qiang Xie leg.; SYSBM. Paratypes CHINA ��� 3 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratype, N = 3). Total body length 1.23 / 1.19���1.25; length of head 0.24 / 0.23���0.26, maximum width across eyes 0.32 / 0.31���0.32, interocular distance 0.25 / 0.24���0.25; length of antennal segment I 0.06 / 0.06, segment II 0.07 / 0.06���0.07, segment III unknown, segment IV unknown; length of labial segment I 0.08 / 0.06���0.08, segment II 0.05 / 0.05���0.06, segment III 0.05 / 0.04, segment IV 0.10 / 0.10; middle length of pronotum 0.31 / 0.31 / 0.32, length of collar 0.06 / 0.05���0.6, humeral width 0.57 / 0.56���0.58; length of forewing 1.10 / 1.06���1.08; length of fore femur 0.30 / 0.28���0.29, fore tibia 0.29 / 0.27���0.29, fore tarsus 0.12 / 0.11���0.12; length of middle femur 0.31 / 0.29���0.32, middle tibia 0.27 / 0.28���0.30, middle tarsus 0.12 / 0.11���0.12; length of hind femur 0.37 / 0.35���0.36, hind tibia 0.47 / 0.46���0.49, hind tarsus 0.18 / 0.17; greatest width of abdomen 0.47 / 0.46���0.52. MACROPTEROUS. Small sized (about 1.19���1.25 mm). COLORATION. Body yellowish brown to light brown, appendages yellowish brown (Fig. 17A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.75���0.78 �� length of maximum width across eyes. THORAX. Humeral width 1.75���1.84 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 27A). ABDOMEN. Tergite VII slightly asymmetrical, right portion wider than left portion, sternite VII nearly symmetrical. Tergite VIII subdivided into two hemitergite, left hemitergite with slender, flattened, sword-shaped process horizontally directed mesad, and without thick spiniform microtrichia on its surface (Figs 17D���E, G, 18A). Right hemitergite subtriangular (Fig. 18B). GENITALIA. Pygophore slightly asymmetrical, dorsal surface with rows of micro teeth, posterior area with about 12 relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 17E, 18D���E) with broadly rounded laminate extension directed dorsal near base, and with flattened and tapering distal projection, and nearly straight; right paramere (Figs 17E, 18G���H) with flattened, round distal projection. Apical portion of phallus (Figs 17D���E, 18F) tubular, short, forming one incomplete coil outside of pygophore. Female Unknown. Distribution China: Yunnan (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 32-35, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Kokeshia acutiformis sp. nov. urn:lsid:zoobank.org:act: 063E5A54-D193-4A58-BA44-A7AC26B1121E Figs 1���2, 26A, 28A, 32 Diagnosis Kokeshia acutiformis sp. nov. can be recognized from all other species of Kokeshia by male tergite VIII not subdivided into two hemitergites, only a long, flattened, acuminate process at right portion of tergite VIII, the process bent at about ⅓ from its apex (Figs 1D���G, 2A���B). Etymology The species epithet is derived from the Latin prefix ��acut -�� and suffix ��- formis ��, means ���acuminate shaped���, refers to the acuminate process of tergite VIII. Material examined Holotype CHINA ��� ♂; Yunnan Province, Lincang City, Cangyuan County, Banhong Town; 23��18���44.65��� N, 99��4���39.48��� E; ca 920 m a.s.l.; 6 Aug. 2019; Jiu-Yang Luo leg.; light trap; SYSBM. Paratypes CHINA ��� 2 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 2). Total body length 1.32 / 1.28���1.29; length of head 0.22 / 0.23���0.25, maximum width across eyes 0.41 / 0.40, interocular distance 0.24 / 0.22; length of antennal segment I 0.08 / 0.07���0.08, segment II 0.08 / 0.08, segment III 0.28 / 0.28, segment IV 0.30 / 0.30; length of labial segment I 0.07 / 0.07���0.08, segment II 0.05 / 0.05, segment III 0.04 / 0.04, segment IV 0.11 / 0.10; middle length of pronotum 0.35 / 0.36, length of collar 0.06 / 0.06���0.07, humeral width 0.59 / 0.60; length of forewing 1.07 / 0.95���1.08; length of fore femur 0.33 / 0.29���0.31, fore tibia 0.31 / 0.32, fore tarsus 0.12 / 0.11���0.12; length of middle femur 0.31 / 0.33, middle tibia 0.32 / 0.31, middle tarsus 0.12 / 0.12; length of hind femur 0.36 / 0.35���0.36, hind tibia 0.50 / 0.53���0.55, hind tarsus 0.17 / 0.17���0.18; greatest width of abdomen 0.53 / 0.53. MACROPTEROUS. Small sized (about 1.28���1.32 mm). COLORATION. Body yellowish brown to light brown, appendages yellowish brown (Fig. 1A���C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.55���0.59 �� length of maximum width across eyes. THORAX. Humeral width 1.66���1.68 �� as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 26A). ABDOMEN. Tergite VII slightly asymmetrical, right portion wider than left portion, sternite VII nearly symmetrical. Tergite VIII not subdivided into two hemitergites, only a long, flattened, acuminate process at right portion of tergite VIII, process bent at about ⅓ from its apex (Figs 1D���G, 2A���B). GENITALIA. Pygophore slightly asymmetrical, dorsum with rows of tooth like microtrichia, posterior area with about twelve relatively long setae (Fig. 1D���G). Parameres strongly asymmetrical, left paramere (Figs 1E, 2D���E) with broadly squarish laminate extension directed dorsal near base, and with flattened and elongate distal projection; right paramere (Figs 1E, 2F���G) with flattened and acute, near triangular distal projection, curved inward at middle portion. Apical portion of phallus (Figs 1D���E, 2C) tubular, short, forming one incomplete coil outside of pygophore. Female Unknown. Distribution China: Yunnan (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on pages 5-8, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878

Kokeshia caii sp. nov. urn:lsid:zoobank.org:act: F68DE9D8-F3D7-4583-9224-2807B7234F5C Figs 7–8, 26D, 28D, 32 Diagnosis The male of K. caii sp. nov. is similar to K. weirauchae sp. nov., but can be recognized from the latter by the left lateral margin of the hemitergite VIII strongly concave from about ⅓ of its apex, and the apical portion of hemitergite VIII with a finger-shaped process, which without any thick spiniform microtrichia, and with numerous microtrichia which the apex split into 2 to 5 branches (Figs 7D‒G, 8A‒B). Etymology The species epithet is derived from and dedicated to Prof. Wanzhi Cai (College of Plant Protection, China Agricultural University, China), recognizing his contributions to the study of taxonomy, morphology and phylogeny of Heteroptera. Material examined Holotype CHINA • ♂; Yunnan Province, Xishuangbanna, Menghai County, Gelanghe Town, Pazhen village; 21°51′10.37″ N, 100°35′26.67″ E; ca 1610 m a.s.l.; 27 Jul. 2019; Jiu-Yang Luo leg.; light trap; SYSBM. Paratypes CHINA • 5 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 5). Total body length 1.36 / 1.24–1.36; length of head 0.25 / 0.23–0.25, maximum width across eyes 0.39 / 0.37‒0.40, interocular distance 0.27 / 0.26‒0.28; length of antennal segment I 0.08 / 0.07–0.08, segment II 0.08 / 0.07‒0.08, segment III 0.30 / 0.28‒0.30, segment IV 0.34 / 0.31‒0.34; length of labial segment I 0.07 / 0.07‒0.08, segment II 0.06 / 0.05‒0.06, segment III 0.04 / 0.04‒0.05, segment IV 0.12 / 0.11‒0.12; middle length of pronotum 0.41 / 0.33‒0.40, length of collar 0.06 / 0.06, humeral width 0.70 / 0.60‒0.69; length of forewing 1.16 / 0.93–1.19; length of fore femur 0.30 / 0.31–0.33, fore tibia 0.35 / 0.33‒0.35, fore tarsus 0.13 / 0.13; length of middle femur 0.35 / 0.34‒0.38, middle tibia 0.36 / 0.33‒0.35, middle tarsus 0.12 / 0.12‒0.14; length of hind femur 0.37 / 0.38–0.41, hind tibia 0.58 / 0.55–0.58, hind tarsus 0.19 / 0.18–0.19; greatest width of abdomen 0.59 / 0.56‒0.58. MACROPTEROUS. Small sized (about 1.24‒1.36 mm). COLORATION. Body brown, appendages yellowish brown (Fig. 7A–C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.68–0.71 × length of maximum width across eyes. THORAX. Humeral width 1.69‒1.82 × as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An distinctly darker than wing surface, other veins darker than wing surface (Fig. 26D). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, left hemitergite VIII with finger-shaped process, and with numerous semi-erect, short to moderate long microtrichia on surface, and ends of these microtrichia split into 2 to 5 branches (similar to setae on process of hemitergite VIII of K. bui). Basal portion of process with scale-like projections, with 1 to 6 short microtrichia located on posterior margin of each one of them. Right hemitergite VIII simple, posterolateral portion round, projecting posteriad, outer margin with microtrichia (Figs 7E‒F, 8H). GENITALIA. Pygophore slightly asymmetrical, dorsal surface with groups of microtrichia on small, tightly connected projections, posterior area with relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 7E, 8C‒D) with broadly rounded laminate extension directed dorsal near base, and with flattened and elongate distal projection; right paramere (Figs 7E, 8F‒G) with flattened and blunt distal projection. Apical portion of phallus (Figs 7D‒E, 8E) tubular, forming nearly one complete coil outside of pygophore, apex thickened and forming a depression on one side. Female Unknown. Distribution China: Yunnan (Fig. 32).

Kokeshia zhengi R��dei, Ren & Bu, 2012 Figs 25, 31���32 Kokeshia zhengi R��dei, Ren & Bu, 2012: 32 (original description). Diagnosis Kokeshia zhengi can be recognized from all other species of Kokeshia by tergite VIII subdivided into two hemitergites, left hemitergite VIII with a broad, thick near triangular process, with about 8 thick spiniform microtrichia at its right posterolateral margin (Figs 25C, 31). Material examined Holotype CHINA ��� ♂; Hubei Province, Fang County, Tucheng Town; 16���23 Jul. 2011; Xi-Ping Zhang, Chuan- Ren Li, Jing Zhong and Chang-Liang Hou leg.; tobacco field; NKUM. Paratypes CHINA ��� 2 ♂♂; same collection data as for holotype; NKUM. Distribution Yunnan Province, China (Fig. 32)., Published as part of Luo, Jiu-Yang & Xie, Qiang, 2022, Taxonomic review of Kokeshia Miyamoto, 1960 from China, with description of ten new species (Hemiptera: Heteroptera: Schizopteridae), pp. 1-57 in European Journal of Taxonomy 802 on page 42, DOI: 10.5852/ejt.2022.802.1687, http://zenodo.org/record/6358878, {"references":["Redei D., Ren S. Z. & Bu W. J. 2012. Two new species of Kokeshia from China (Hemiptera: Heteroptera: Schizopteridae). Zootaxa 3497 (1): 29 - 36. https: // doi. org / 10.11646 / zootaxa. 3497.1.3"]}

Kokeshia hilli sp. nov. urn:lsid:zoobank.org:act: F4BD7B25-53B1-4726-9925-AE8E51FA4210 Figs 11–12, 26F, 29B, 32 Diagnosis Kokeshia hilli sp. nov. can be recognized from all other species of Kokeshia by tergite VIII subdivided into two hemitergites, left hemitergite VIII introduced into a flattened, nearly quadrangular process, left posterolateral margin of the process with a raised projection, which bearing about 7 thick, basal-curved spiniform microtrichia; right posterolateral margin of the process with about seven parallel, thick, basalcurved spiniform microtrichia. Right hemitergite VIII with a long, rodlike process projecting posteriad (Figs 11D–E, G, 12 A−B). Etymology The species epithet is derived from and dedicated to Mr Lionel Hill (Tasmanian Department of Primary Industry, Water and Environment, Australia), recognizing his contributions to the study of taxonomy of Dipsocoromorpha (mainly in the Australian region). Material examined Holotype CHINA • ♂; Yunnan Province, Honghe, Jinping County, Adebo Town; 22°54′21″ N, 103°13′49″ E; ca 1480 m a.s.l.; 31 Jul. 2016; Jiu-Yang Luo leg.; NKUM. Paratypes CHINA • 4 ♂♂; same collection data as for holotype; SYSBM. Description Male MEASUREMENTS (in mm) (male holotype / male paratypes, N = 4). Total body length 1.22 / 1.25–1.36; length of head 0.24/ 0.21–0.24, maximum width across eyes 0.37 / 0.38‒0.40, interocular distance 0.25 / 0.23‒0.24; length of antennal segment I 0.06 / 0.07–0.08, segment II 0.07 / 0.07‒0.08, segment III 0.30 / 0.30, segment IV 0.33 / 0.32; length of labial segment I 0.07 / 0.07‒0.08, segment II0.06 / 0.04‒0.05, segment III 0.05 / 0.03‒0.05, segment IV 0.11 / 0.09‒0.12; middle length of pronotum 0.35 / 0.33‒0.36, length of collar 0.06 / 0.06‒0.07, humeral width 0.62 / 0.62; length of forewing 1.07 / 1.08–1.14; length of fore femur 0.32 / 0.29–0.31, fore tibia 0.34 / 0.32‒0.34, fore tarsus 0.12 / 0.12‒0.13; length of middle femur 0.34 / 0.32‒0.35, middle tibia 0.34 / 0.34‒0.35, middle tarsus 0.12 / 0.11‒0.12; length of hind femur 0.36 / 0.35–0.41, hind tibia 0.56 / 0.52–0.57, hind tarsus 0.18 / 0.17–0.19; greatest width of abdomen 0.55 / 0.54‒0.56. MACROPTEROUS. Small sized (about 1.22‒1.36 mm). COLORATION. Body brown, appendages yellowish brown (Fig. 11A–C). Surface, vestiture and general structures as in generic description, except mentioned below. HEAD. Interocular distance 0.58–0.68 × length of maximum width across eyes. THORAX. Humeral width 1.72‒1.88 × as long as middle length of pronotum. Venation of forewing in accordance with general pattern of genus. C+Sc, R+M, R and basal portion of 1An darker than wing surface, other veins slightly darker than wing surface (Fig. 26F). ABDOMEN. Tergite VII and sternite VII slightly asymmetrical. Tergite VIII subdivided into two hemitergites, posterior margin of left hemitergite VIII (Figs 11D–E, G, 12 A−B) introduced into flattened, nearly quadrangular process, left posterolateral margin with raised projection, bearing about 7 thick, basal-curved spiniform microtrichia; right posterolateral margin of process with about 7 parallel, thick, basal-curved spiniform microtrichia. Basal portion of process with numerous small round pits and micro teeth. Right hemitergite VIII with long, stick shaped process projecting posteriad (Figs 11 E−F, 12E). GENITALIA. Pygophore slightly asymmetrical, dorsal surface with dense microtrichia, posterior area with relatively long setae. Parameres strongly asymmetrical, left paramere (Figs 11E, 12G‒H) with broadly blunt laminate extension directed dorsal near base, and with flattened and relatively broad distal projection, and slightly curved inward from middle of it; right paramere (Figs 11E, 12C‒D) with flattened, round and relatively slender distal projection. Apical portion of phallus (Figs 11D‒E, 12F) tubular, slender, forming less than one and a half coils outside of pygophore. Female Unknown. Distribution China: Yunnan (Fig. 32).

Kokeshia xiei Rédei, Ren & Bu, 2012 Figs 23–24, 27D, 30D, 32 Kokeshia xiei Rédei, Ren & Bu, 2012: 30 (original description). Diagnosis Kokeshia xiei can be recognized from all other species of Kokeshia by tergite VIII subdivided into two hemitergites, left hemitergite VIII with a long, thick and horizontally directed process, with numerous thick spiniform microtrichia at its ventral surface (Figs 23D‒E, G, 24A‒B). Material examined Holotype CHINA • ♂; Hainan Province, Lingshui County, Diaoluoshan; ca 300 m a.s.l.; 10 Aug. 2008; Qiang Xie leg.; at light; NKUM. Paratypes CHINA • 12 ♂♂; same collection data as for holotype; NKUM. Non-type specimens CHINA • 67 ♂♂; Hainan Province, Baisha County, Yinggeling National Nature Reserve, Nankai Protection Station; 19°4′45″ N, 109°24′40″ E; ca 260 m a.s.l.; 20 Jul. 2013; Yan-Hui Wang leg.; at light; NKUM. Distribution Yunnan Province, China (Fig. 32).