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111. Isohypsibius sattleri (Richters, 1902) sensu lato [T] Hypsibius sattleRi (Richters) 1902 (da Cunha & do Nascimento 1964) Isohypsibius bakonyiensis (Iharos, 1964) (Binda 1971, 1984) Isohypsibius bakonyiensis Iharos, 1964 (Pilato & Pennisi 1976) Hypsibius bakonyiensis Iharos (Iharos 1978) Isohypsibius bakonyiensis, I. sattleRi (Ramløv & Kristensen 1985) Isohypsibius sattleRi (Richters, 1902) (Pilato et al. 2003) Terra typica: Germany (Europe) Angola: • 07°47′S, 15°01′E; 650 m asl: Uíge Province, near Carmona [Uíge], Estate at Pumba Loge, mosses. da Cunha & do Nascimento (1964) Democratic Republic of Congo: • 00°04′S, 29°18′E; 1,950 m asl: North Kivu Province, outskirts of Lukanga, moss (BRachythecium velutinum). Pilato et al. (2003) Libya: • 32°29′N, 20°50′E; 350 m asl: Marj District, Barce [Marj], moss. Pilato & Pennisi (1976) Morocco: • 32°56′N, 05°40′W; 1,700 m asl [850 m asl]: Meknès-Tafilalet Region, Khénifra, moss. Binda (1971) • 32°25′N, 05°13′W; 1,750 m asl: Meknès-Tafilalet Region, S of Azrou, cedar forest, moss on basalt and limestone boulders in the forest glade and moss on soil and fallen branches and stems (3 samples). Ramløv & Kristensen (1985) • Republic of South Africa: • 34°01′S, 23°55′E; 200 m asl: Eastern Cape Province, Tzitzikama [Tsitsikamma National Park], moss. Binda (1984) Tunisia: • 36°14′N, 08°46′E; 1,100 m asl: Kef Governorate, Kef Governorate, 6 km NE of El Kef, Djebel Eddir Mts., surroundings of Ferme Shitta, moss and lichen on rock. Iharos (1978) • 36°46'36''N, 08°42'45''E; 784 m asl: Jendouba Governorate, Beni M'tir, urban, moss on soil. Gąsiorek et al. (2017) Record numbers. Angola: 1, Democratic Republic of Congo: 1, Libya: 1, Morocco: 2, Republic of South Africa: 1, Tunisia: 2; total: 8. Remarks. This member of the tuberculatus -group has a history of major taxonomic confusion with I. bakonyiensis (now the junior synonym), but was re-described from a Richters’ 1905 slide in the Senckenberg Museum (Germany) by Dastych (1990). Nevertheless, the literature-based, supposed, cosmopolitan distribution (McInnes 1994), requires major taxonomic re-evaluation of all specimens attributed to I. sattleri., Published as part of Kaczmarek, Łukasz, 2017, Annotated zoogeography of non-marine Tardigrada. Part IV: Africa, pp. 1-74 in Zootaxa 4284 (1) on pages 39-40, DOI: 10.11646/zootaxa.4284.1.1, http://zenodo.org/record/1010400, {"references":["Richters, F. (1902) Neue Moosbewohner. BeRicht deR SenckenbeRgischen NatuRfoRschenden gesellschaft in FRankfoRt am Main, 1902, 23 - 26.","Da Cunha, A. X. da & Nascimento Ribeiro, F. (1964) Tardigrados de Angola. GaRcia de ORta, Lisboa, 12, 397 - 406.","Pilato, G. & Pennisi, G. (1976) Prime notizie sui Tardigradi della Cirenaica. Animalia, 3, 243 - 258.","Iharos, G. (1978) Data to the knowledge of the Tardigrada fauna of Tunisia. Folia Entomologica HungaRica, 21, 175 - 177.","Ramlov, H. & Kristensen, R. M. (1985) Bjornedyr, Tardigrada. Marokko - 1985. Dansk NatuRhistoRisk FoRenings Rejse til MaRokko, 8 - 22 ApRil 1985, pp. 27 - 32.","Gasiorek, P., Morek, W., Stec, D., Marnissi, J. & Michalczyk, L. (2017) The tardigrade fauna of Tunisia, with an integrative description of BRyodelphax maculatus sp. nov. (Tardigrada: Heterotardigrada: Echiniscoidea). African Zoology. http: // dx. doi. org / 10.1080 / 15627020.2017.1297688","McInnes, S. J. (1994) Zoogeographic distribution of terrestrial / freshwater tardigrades from current literature. JouRnal of NatuRal HistoRy, 28, 257 - 352. https: // doi. org / 10.1080 / 00222939400770131"]}

146. Isohypsibius sattleri (Richters, 1902) sensu lato [T] Hypsibius sattleri Richt. (Iharos 1963) H. bakonyiensis Iharos (Iharos 1969) H. (I.) sattleri (Mihelčič 1972) Hypsibius bakonyiensis (Iharos, 1964) (Hypsibius (I.) sattleri Auct., partim) (Maucci 1988) I. sattleri (Garitano-Zavala 1996) Terra typica: Germany (Europe) Argentina: ��� 41 �� 58 ���S, 71 �� 28 ���W; 1,460 m asl: Rio Negro Province, El Bols��n, Piltriquitron Mt., mosses on rocks and barks of trees near edge within Nothofagus pumilio forest. Iharos (1963) ��� Undefined localities cited according Mihelčič (1967), cypress grove, dry mosses and lichens on rocks in full sun and dry and wet tufts and rosette plants on rocks (3 samples). Mihelčič (1972) Bolivia: ��� 16 �� 13 ���S, 68 �� 13 ���W; 4,550���4,700 m asl: La Paz Department, Cordillera de la Real de los Andes, Tuni, subnival floor, cushion moss. Garitano-Zavala (1995, 1996) ��� 16 �� 18 ���S, 68 �� 17 ���W; 4,300���4,400 m asl: La Paz Department, Cordillera de la Real de los Andes, Wila Kunka, high Andean floor, cushion moss. Garitano-Zavala (1995, 1996) ��� 16 �� 20 ���S, 68 �� 23 ���W; 4,650���4,700m asl: La Paz Department, Cordillera de la Real de los Andes, La Cumbre, subnival floor, cushion moss. Garitano-Zavala (1995, 1996) Chile: ��� 33 ��04���S, 71 ��00���W; 1,000 m asl: Region RM Metropolitana (Regi��n Metropolitana de Santiago), Tiltil, Cuesta La Dormida, soil on dry forest. Iharos (1969) ��� 51 �� 34 ���S, 72 �� 36 ���W; 200 m asl: Region XII Magallanes (Regi��n de Magallanes y de la Ant��rtica Chilena), Ultima Esperanza, near Cueva del Milod��n, mosses on rocks, in sunlight. Maucci (1988) Colombia: ��� 11 ��06���20.0������N, 74 ��03���50.0������W; 2,200 m asl: Magdalena Department, Sierra Nevada de Santa Marta, between La Tagua and Cuchilla de San Lorenzo, sub-Andean forest, lichen on rock. Lisi et al. (2014) Ecuador: ��� 00�� 22 ���S, 78 �� 55 ���W, 1,100 m asl: Cotopaxi Province, Otonga (San Pedro de las Pampas) [Otonga Nature Reserve], liverwort (Plagiochila punctata). Pilato et al. (2001) Record numbers: Argentina: 2, Bolivia: 3, Chile: 2, Colombia: 1, Ecuador: 1; total: 9. Remarks: This member of the tuberculatus- group has a history of major taxonomic confusion, but been redescribed from contemporary material (Dastych 1990). Nevertheless, its cosmopolitan distribution remains questionable (McInnes, 1994 a), requiring major taxonomic revision., Published as part of Kaczmarek, ��ukasz, Michalczyk, ��ukasz & Mcinnes, Sandra J., 2015, Annotated zoogeography of non-marine Tardigrada. Part II: South America, pp. 1-107 in Zootaxa 3923 (1) on pages 58-59, DOI: 10.11646/zootaxa.3923.1.1, http://zenodo.org/record/241936, {"references":["Richters, F. (1902) Neue Moosbewohner. Bericht der Senckenbergischen Naturforschenden gesellschaft in Frankfort am Main, 23 - 26.","Iharos, G. (1963) The zoological results of Gy. Topal's collections in South Argentina, 3. Tardigrada. Annales Historico Naturales Musei Nationalis Hungarici, 55, 293 - 299.","Iharos, G. (1969) The scientific results of the Hungarian soil zoological expeditions to South America: 15. Tardigraden aus den Sammlungen der ersten und zweiten Expedition. Opuscula Zoologica, Budapest, 9, 279 - 289.","Mihelcic, F. (1972) Ein weiterer Beitrag zur Kenntnis der Tardigraden Argentiniens. Verhandlungen der Zoologisch- Botanischen Gesellschaft in Wien, 110 / 111, 47 - 52.","Iharos, G. (1964) Neuere Beitrage zur Kenntnis der Tardigraden- Fauna Ungarns. V. Opuscula Zoologica, Budapest 5, 57 - 67.","Maucci, W. (1988) Tardigrada from Patagonia (Southern South America) with description of three new species. Revista Chilena de Entomologia, 16, 5 - 13.","Garitano-Zavala, B. A. (1996) Efecto de la humedad microambiental y la altura, sobre la distribucion de la Tardigradifauna Muscicola (Phylum Tardigrada) en la vertiente occidental de la Cordillera Real de los Andes (La Paz, Bolivia). Ecologia en Bolivia, 26, 1 - 48.","Mihelcic, F. (1967) Ein Beitrag zur Kenntnis der Tardigraden Argentiniens. Verhandlungen der Zoologisch - Botanischen Gesellschaft in Wien, 107, 43 - 56.","Garitano-Zavala, A. B. (1995) Primer reporte de las especies de tardigrados muscicolas (Phylum Tardigrada) presentes en la vertiente occidental de Cordillera Real de los Andes (La Paz - Bolivia), Ecologia en Bolivia, 24, 1 - 39.","Lisi, O., Londono, R. & Quiroga, S. (2014) Tardigrada from a sub-Andean forest in the Sierra Nevada de Santa Marta (Colombia) with the description of Itaquascon pilatoi sp. nov. Zootaxa, 3841 (4), 551 - 562. http: // dx. doi. org / 10.11646 / zootaxa. 3841.4.5","Dastych, H. (1990) Isohypsibius sattleri (Richters 1902), a valid species (Tardigrada). Senckenbergiana Biologica, 71, 181 - 189.","McInnes, S. J. (1994 a) Zoogeographic distribution of terrestrial / freshwater tardigrades from current literature. Journal of Natural History, 28, 257 - 352. http: // dx. doi. org / 10.1080 / 00222939400770131"]}

146. Isohypsibius sattleri (Richters, 1902) sensu lato [T] Hypsibius sattleri Richt. (Iharos 1963) H. bakonyiensis Iharos (Iharos 1969) H. (I.) sattleri (Mihelčič 1972) Hypsibius bakonyiensis (Iharos, 1964) (Hypsibius (I.) sattleri Auct., partim) (Maucci 1988) I. sattleri (Garitano-Zavala 1996) Terra typica: Germany (Europe) Argentina: • 41 ° 58 ′S, 71 ° 28 ′W; 1,460 m asl: Rio Negro Province, El Bolsón, Piltriquitron Mt., mosses on rocks and barks of trees near edge within Nothofagus pumilio forest. Iharos (1963) • Undefined localities cited according Mihelčič (1967), cypress grove, dry mosses and lichens on rocks in full sun and dry and wet tufts and rosette plants on rocks (3 samples). Mihelčič (1972) Bolivia: • 16 ° 13 ′S, 68 ° 13 ′W; 4,550–4,700 m asl: La Paz Department, Cordillera de la Real de los Andes, Tuni, subnival floor, cushion moss. Garitano-Zavala (1995, 1996) • 16 ° 18 ′S, 68 ° 17 ′W; 4,300–4,400 m asl: La Paz Department, Cordillera de la Real de los Andes, Wila Kunka, high Andean floor, cushion moss. Garitano-Zavala (1995, 1996) • 16 ° 20 ′S, 68 ° 23 ′W; 4,650–4,700m asl: La Paz Department, Cordillera de la Real de los Andes, La Cumbre, subnival floor, cushion moss. Garitano-Zavala (1995, 1996) Chile: • 33 °04′S, 71 °00′W; 1,000 m asl: Region RM Metropolitana (Región Metropolitana de Santiago), Tiltil, Cuesta La Dormida, soil on dry forest. Iharos (1969) • 51 ° 34 ′S, 72 ° 36 ′W; 200 m asl: Region XII Magallanes (Región de Magallanes y de la Antártica Chilena), Ultima Esperanza, near Cueva del Milodón, mosses on rocks, in sunlight. Maucci (1988) Colombia: • 11 °06′20.0′′N, 74 °03′50.0′′W; 2,200 m asl: Magdalena Department, Sierra Nevada de Santa Marta, between La Tagua and Cuchilla de San Lorenzo, sub-Andean forest, lichen on rock. Lisi et al. (2014) Ecuador: • 00° 22 ′S, 78 ° 55 ′W, 1,100 m asl: Cotopaxi Province, Otonga (San Pedro de las Pampas) [Otonga Nature Reserve], liverwort (Plagiochila punctata). Pilato et al. (2001) Record numbers: Argentina: 2, Bolivia: 3, Chile: 2, Colombia: 1, Ecuador: 1; total: 9. Remarks: This member of the tuberculatus- group has a history of major taxonomic confusion, but been redescribed from contemporary material (Dastych 1990). Nevertheless, its cosmopolitan distribution remains questionable (McInnes, 1994 a), requiring major taxonomic revision.

10. Isohypsibius sattleri (Richters, 1902) Margoitsy (1 moss sample: 2 specimens). The species, already reported from Belarus, is considered cosmopolitan., Published as part of Pilato, Giovanni, Kiosya, Yevgen, Lisi, Oscar & Sabella, Giorgio, 2012, New records of Eutardigrada from Belarus with the description of three new species, pp. 39-60 in Zootaxa 3179 on page 42, DOI: 10.5281/zenodo.279945, {"references":["Richters, F. (1902) Beitrage zur Kenntnis der Fauna der Umgebung von Frankfurt a. M. 1 Forts. Bericht der Senckenberzischen Natuforschenden Gesellschaft in Frankfurt am Main, 3 - 21."]}

Isohypsibius sattleri (Richters, 1902) Material examined. Three specimens were collected from the Huangguoshu Fall. Remarks. This species was reported as a new record for China from Yunnan Province (Beasley et al., 2006). It is the first report of this species from Guizhou Province., Published as part of Wang, Hanping, Wang, Lizhi & Li, Xiaochen, 2007, Tardigrades from Guizhou Province (China) with description of a new species of genus Doryphoribius (Tardigrada: Hypsibiidae), pp. 59-67 in Zootaxa 1626 on page 66, DOI: 10.5281/zenodo.179303, {"references":["Beasley, C. W., Kaczmarek, L. & Michalczyk, L. (2006) New records of tardigrades from China, with zoogeographical remarks. Biological Letters, 43 (1), 3 - 10."]}

Isohypsibius sattleri (Richters, 1902) Material examined Two specimens from an undetermined moss sample developing on rock. Remarks The species is cosmopolitan., Published as part of Pilato, Giovanni & Lisi, Oscar, 2006, Notes on some tardigrades from southern Mexico with description of three new species, pp. 53-68 in Zootaxa 1236 on page 66, DOI: 10.5281/zenodo.273448, {"references":["Richters, F. (1902) Beitrage zur Kenntnis der Fauna der Umgebung von Frankfurt a. M. Berichte Senckenbergische Naturforschende Gesellschaft, 8 - 13."]}