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Species vary in their responses to urban habitat; most species avoid these environments, whereas others tolerate or even thrive in them. To better characterize the extent to which species vary in their responses to urban habitat (from this point forwards "urban tolerance"), we used several methods to quantify these responses at a continental scale across all birds. Using open access community science‐derived data from the eBird Status and Trends Products and two different types of high‐resolution geospatial data that quantify urbanization of landscapes, we calculated urban tolerance for 432 species with breeding ranges that overlap large cities in Canada or the USA. We developed six different calculations to characterize species‐level urban tolerance, allowing us to assess how each species' relative abundance across their breeding range varied with estimates of urban habitat use and intensity. We assessed correlations among these six indices, then compressed the two best‐performing indices into a single principal component (multivariate urban tolerance index) that captured variation in urban tolerance among species. We assessed the accuracy of our single and multivariate urban tolerance indices using 24 test species that have been well characterized for their tolerance or avoidance of the urban habitat, as well as with previously published, independent urban tolerance estimates. Here, we provide this new dataset of species‐level urban tolerance estimates that improves upon previous metrics by incorporating continental‐scale, continuous estimates that better differentiate species' tolerance of urban habitat compared with existing, categorical methods. These refined metrics can be used to test hypotheses that link ecological, life history, and behavioral traits to avian urban tolerance. The dataset is licensed as CC‐By Attribution 4.0 International. Users must appropriately cite the data paper and dataset if used in publications and scientific presentations. [ABSTRACT FROM AUTHOR]

Farmland comprises an important opportunity for biodiversity conservation worldwide. The likelihood of a farmer fostering habitat for biodiversity has been studied from the perspective of personal or economic drivers. Beyond farm area, the geographic characteristics of the farmland itself—such as parcel sizes, numbers, and distribution—are rarely considered. This paper uses a landholder survey (n = 350, 37% response rate) to explore the variety of farmland management fragmentation and its implications for habitat on farms, specifically ponds, wetlands, and woodlands. This exploratory research was implemented in Nova Scotia, a relatively long‐settled province of Canada, which is subject to inheritance, farmland abandonment, and farm expansion. An index and a typology of management fragmentation were developed, based on geographical impacts (increased distance to travel and edge per unit area), drawing on hypotheses about how geography may affect farmer decision making about habitat. Results suggest that farm size matters more than fragmentation, but also that perceptions of ecosystem services from each ecosystem may have an impact. Suggestions for further research are provided, including alternative methods that could be used and testing these insights in more homogenous agricultural landscapes. [ABSTRACT FROM AUTHOR]

Grizzly bears inhabit wilderness areas in the northwestern region of the lower forty-eight states, western Canada, and areas of Alaska. Because of the settlement of the west and loss of prime habitat, populations declined rapidly in the nineteenth century, and in 1975 federal action was taken to protect grizzlies under the Endangered Species Act. Since 1950 about 722 technical papers have been written on the grizzly bear. Major research has focused on ecology, conservation, reproductive biology, behavior, dietetics, anatomy, and physiology, among other topics. Due to geographic distribution of the species, much of the research has been carried out by authors and organizations in western regions of the United States and Canada where major grizzly populations exist. A significant number of technical papers appear in three key journals: Ursus, the Journal of Wildlife Management, and the Canadian Journal of Zoology. According to data in WorldCat, about 1,167 records, covering monographs and technical reports, contain information on grizzlies and present research findings. The bulk of monographs appeal mainly to a general audience. However, citation analysis reveals a core of highly cited technical papers, many written with an emphasis on special themes or topics, whereas others focus on the grizzly itself, all together advancing the science on this species. [ABSTRACT FROM PUBLISHER]

Despite its role as a keystone species, few studies have investigated the habitat requirements of the Yellow-bellied Sapsucker (Sphyrapicus varius (L., 1766)) in the northwestern part of its range, where the current forest harvesting rotation schedule and targets for reducing the prevalence of old and mixedwood stands may reduce the optimal habitat for this species. We studied nesting and foraging habitat requirements of nesting sapsuckers in a boreal mixedwood forest of northwestern Canada by collecting data on nesting sites and foraging substrates on twenty-four 16- to 56-ha plots distributed among four habitat types in 1998 and 2002. Nests (n = 68) were generally found in large (≥22 cm diameter at breast height) trembling aspen (Populus tremuloides Michx) that were alive but declining and that showed high incidence (81.1%) of heartwood rot infection (Phellinus tremulae (Bondarzev) Bondarzev & Borisov in Bondarzev). Nest-site use by sapsuckers was predicted mainly by the presence of external fungal conks and tree diameter. Among tree species used as foraging substrates, paper birch (Betula papyrifera Marsh.) (65.6%) and green alder (Alnus viridis (Vill.) Lam. & DC.) (21.3%) were used most frequently. The use of birch was strongly correlated with its availability. Mature forests had higher densities of nesting sites and foraging substrates than immature aspen stands. In addition to reconfirming the importance of mature mixedwood forests for Yellow-bellied Sapsuckers, our study also indicates that forest harvest rotations in northwestern Canada should exceed 90 years to promote the keystone role of this species. [ABSTRACT FROM AUTHOR]

Much of Canada's terrestrial biodiversity is supported by boreal forests. Natural resource development in boreal forests poses risks to this biodiversity. This paper reviews the scientific literature to assess the effects of natural resource development on terrestrial biodiversity in Canadian boreal forests. We address four questions: (1) To what extent have Canadian boreal forests changed due to natural resource development? (2) How has biodiversity responded to these changes? (3) Will the biodiversity of second-growth forests converge with that of primary boreal forests? (4) Are we losing species from boreal forests? We focus on trees, understory plants, insects, fungi, selected mammals, and songbirds because these groups have been most studied. We review more than 600 studies and found that changes in community composition are prevalent in response to large-scale conversion of forest types, changes in stand structures and age distributions, and altered landscape structure resulting from forest management and habitat loss associated with other developments such as oil and gas, hydroelectric, and mining. The southern boreal forest has been more highly impacted than the north due to more extensive forest management and the cumulative effects of multiple forms of development. There is abundant evidence that most species are not in danger of being extirpated from the boreal forest due to these anthropogenic changes. A few species, including woodland caribou ( Rangifer tarandus) and grizzly bear ( Ursus arctos), have, however, undergone long-term range contractions. Significant gaps in our ability to assess the effects of natural resource development on biodiversity in the boreal zone are the lack of long-term spatial and population data to monitor the impact of forest changes on ecosystems and species. [ABSTRACT FROM AUTHOR]

Background: Feral populations of cultivated crops can pose challenges to novel trait confinement within agricultural landscapes. Simulation models can be helpful in investigating the underlying dynamics of feral populations and determining suitable management options. Methodology/Principal Findings: We developed a stage-structured matrix population model for roadside feral alfalfa populations occurring in southern Manitoba, Canada. The model accounted for the existence of density-dependence and recruitment subsidy in feral populations. We used the model to investigate the long-term dynamics of feral alfalfa populations, and to evaluate the effectiveness of simulated management strategies such as herbicide application and mowing in controlling feral alfalfa. Results suggest that alfalfa populations occurring in roadside habitats can be persistent and less likely to go extinct under current roadverge management scenarios. Management attempts focused on controlling adult plants alone can be counterproductive due to the presence of density-dependent effects. Targeted herbicide application, which can achieve complete control of seedlings, rosettes and established plants, will be an effective strategy, but the seedbank population may contribute to new recruits. In regions where roadside mowing is regularly practiced, devising a timely mowing strategy (early- to mid-August for southern Manitoba), one that can totally prevent seed production, will be a feasible option for managing feral alfalfa populations. Conclusions/Significance: Feral alfalfa populations can be persistent in roadside habitats. Timely mowing or regular targeted herbicide application will be effective in managing feral alfalfa populations and limit feral-population-mediated gene flow in alfalfa. However, in the context of novel trait confinement, the extent to which feral alfalfa populations need to be managed will be dictated by the tolerance levels established by specific production systems for specific traits. The modelling framework outlined in this paper could be applied to other perennial herbaceous plants with similar life-history characteristics. [ABSTRACT FROM AUTHOR]

No net loss of productive capacity (PC) of fish habitat has been the central concept guiding Canadian fish habitat management policy since 1986. The purpose of this paper is to describe the concept of PC, to review the history and application of the fish habitat management policy in Canada, and to provide a critical review of the range of potential approaches to estimating PC. The approaches were grouped by their central focus: habitat, individual, population, and community-ecosystem. A set of case studies is used to illustrate the use of some approaches drawn from freshwater and marine contexts. Ten components to assessing no net loss of PC were developed and used in the review of approaches for evaluating potential limitations. The review also highlighted the likely future direction of method development, with increasing emphasis on dynamic models integrating population responses to habitat supply characteristics. More work needs to be done to turn research-based metrics of PC into practical operational management assessment tools and to better quantify the link between habitat structure and function and fisheries productivity. The evolving approaches to measure PC reinforce the ties that fish habitat management has to the emerging practices in ecosystem-based management. [ABSTRACT FROM AUTHOR]

Abstract: Birds can be used as indicators to monitor success of programs encouraging prairie landowners to increase biodiversity. Using a case study from Alberta, Canada, this paper compares bird diversity measures at the farm scale and examines their consistency across different habitat types to test for design, output, and end use validation. Based on 2005 point count data (two types) from 178 sites at 22 farms, we calculated bird species richness, abundance, Shannon index, and Inverse Simpson index. The 50m radius data produced species richness and abundance measures about half the size as those produced by the unlimited radius data. The bird diversity measures were consistent across habitat types. The 50m radius data showed differences among 3–5 habitat types, whereas the unlimited radius data showed differences between only two habitat types. Using any bird diversity measure, the wetland/riparian habitat scored highest, followed by homestead, upland forest, native prairie, tame pasture, and cultivated land habitats. Bird monitoring methods should favor fixed over unlimited radius point counts, because of the former''s greater discriminating ability. Given that diversity measures are consistent across habitats and are highly correlated, the species richness measure, which is simple and easy to understand, can be used in conversations with landowners and policy-makers. [Copyright &y& Elsevier]

Background: One of the greatest obstacles to moving ecosystem-based management (EBM) from concept to practice is the lack of a systematic approach to defining ecosystem-level decision criteria, or reference points that trigger management action. Methodology/Principal Findings: To assist resource managers and policymakers in developing EBM decision criteria, we introduce a quantitative, transferable method for identifying utility thresholds. A utility threshold is the level of human-induced pressure (e.g., pollution) at which small changes produce substantial improvements toward the EBM goal of protecting an ecosystem's structural (e.g., diversity) and functional (e.g., resilience) attributes. The analytical approach is based on the detection of nonlinearities in relationships between ecosystem attributes and pressures. We illustrate the method with a hypothetical case study of (1) fishing and (2) nearshore habitat pressure using an empirically-validated marine ecosystem model for British Columbia, Canada, and derive numerical threshold values in terms of the density of two empirically-tractable indicator groups, sablefish and jellyfish. We also describe how to incorporate uncertainty into the estimation of utility thresholds and highlight their value in the context of understanding EBM trade-offs. Conclusions/Significance: For any policy scenario, an understanding of utility thresholds provides insight into the amount and type of management intervention required to make significant progress toward improved ecosystem structure and function. The approach outlined in this paper can be applied in the context of single or multiple human-induced pressures, to any marine, freshwater, or terrestrial ecosystem, and should facilitate more effective management. [ABSTRACT FROM AUTHOR]

1. Elevational gradients create environmental variation that is hypothesized to promote variation in life-history strategies. We tested whether differences in life-history strategies were associated with elevation in a songbird, the dark-eyed junco ( Junco hyemalis; Aves; A.O.U. 1998 ). 2. We monitored birds in four replicated sites per elevation, at 2000 m a.s.l. (high elevation) and 1000 m a.s.l. (low elevation), in the Rocky Mountains of Canada. 3. Over 5 years, we measured the following traits and vital rates: egg-laying schedules, morphological indicators of reproductive stage, seasonal reproductive success, indicators of competitive class (age, size, arrival time), and survival rates. 4. We found two main patterns: with an increase in breeding elevation, dark-eyed juncos delayed the development of structures necessary for reproduction (e.g. cloacal protuberance in males) and reduced the duration of their reproductive period to less than half of the time used by low-elevation birds; and 5. Juncos at high-elevation sites had 55–61% lower annual reproductive success and 15 to 20% higher survival rates. While adult juncos at high elevations produced fewer offspring, those offspring were in better condition. Proportions of age and size classes in high- compared to low-elevation populations were similar, suggesting that a life-history trade-off is present, rather than competition forcing inferior competitors to breed in a peripheral habitat. The apparent trade-off between reproduction and survival corresponded to a shorter period of favourable weather and available food in high- compared to low-elevation habitats. 6. Thus, elevation had a strong influence on life-history characteristics of a single species over a short spatial distance, suggesting a shift in life history from a high reproductive strategy at lower elevations to a high survivor strategy at high elevations. 7. This is the first paper to show a shift in avian life-history strategies along an elevational gradient (in both genders, of multiple age classes) when region (latitude) and phylogenetic histories are controlled for. [ABSTRACT FROM AUTHOR]

This paper discusses local understanding of plants and habitats, based on the linguistic evidence [terms for plants and (or) habitats] gathered from ethnobotanical and ethnoecological field work conducted with several Dene Nations of the Canadian northwestern boreal forest and adjacent regions. Nations involved in the study include Gwich’in (Mackenzie Delta Region), Sahtú’otine’ (Great Bear Lake), Kaska Dena (southern Yukon), and Witsuwit’en (northwest British Columbia). Key plant-related habitats include meadow, “swamp”, forest, “willows”, and “brush”. The ethnobotanical classification of willows is explored in conjunction with the explanation of the Dene habitat concept. In local classifications, ‘willow’ is not co-extensive with the genus Salix, but includes a variety of medium to tall woody shrubs that lack either conspicuous flowers, ‘berries’, or thorns; these may include shrubby species of Salix, Alnus, Cornus, and Betula. Shoreline and alpine environments are also discussed as plant habitats. Dene use of alpine environments and resources is ancient, according to the results of recent alpine ice patch research in the Yukon region. The Human dimensions of habitat knowledge are presented. Indigenous concepts of plant taxa and of landscape associations or habitats may differ substantially from those of scientific botany and ecology, and are based in a holistic and interactive ethnoecology. [ABSTRACT FROM AUTHOR]

This paper presents a sampling design for monitoring spatial and temporal changes in forest health in the Upper Elbow River Basin, in Alberta, Canada. The procedure involved a combination of cumulative effects assessment and remote sensing techniques for selecting sampling sites based on physical and anthropogenic features. Normalized difference vegetation index (NDVI) was the indicator of forest health. Unique combinations of slope, aspect, soil moisture, NDVI, vegetation type, and the zone of influence of human activities were used to select pairs of sampling sites. Each pair consisted of a site within the zone of influence and one outside the zone. Spatial discrimination analysis was the method used for reclassification. The analysis suggested that 58 pairs would be appropriate for monitoring. NVDI was negatively correlated with dry soils and increased with the slope. To various extents, most of the species displayed NDVI values between 0.20 and 0.59. The density of linear disturbances (km/km2) was estimated and it showed that one of the three sub-catchments, the Bragg Creek, has levels of human disturbance above the value considered optimal for wildlife habitat suitability. [ABSTRACT FROM AUTHOR]

Moose (AIces alces)-vehicle collisions (MVC) can be costly ecologically by affecting population numbers, economically by vehicle damage, and socially through human injury or mortality. The purpose of this paper is to identify factors related to moose ecology, driver behaviour, and road design that are useful for predicting the spatial location of MVC on the Trans Canada Highway dissecting Mount Revelstoke and Glacier National Parks. We identified 6 subsets of logistic regression models and used Akaike's information Criteria (AIC) to determine the most parsimonious model within each subset. In addition to this study being the first to examine collisions within these 2 parks, each of these 6 modelling procedures is unique in predicting MVC. Five of the six subsets modelled local-scale/field-based hypotheses of driver visibility, moose evidence, highway design, roadside vegetation, and moose habitat, while the sixth subset examined landscape-scale hypotheses through the use of a Geographic Information System (GIS). The Receiver Operator Characteristic (ROC) discriminated between MVC and random reference sites in order to validate the best fit model from each of the 6 subsets. A MVC probability map along the highway was created using the GIS model, providing a powerful and relatively efficient and inexpensive planning tool. The moose evidence model correctly classified the most MVC among the local-scale models. In relation to the spatial analysis, highway planning to reduce MVC risk within the parks should begin by assessing landscape-scale variables with emphasis on distance to wetland and landscape slope. This landscape-scale analysis should be followed by field-based modelling using moose evidence and habitat-related modelling with important predictors of moose tracks, game trails, and coniferous forest habitat. If highway planning cannot be effective in decreasing MVC, mitigation measures should include a public awareness program, speed reduction, and consideration of an alternative intercept foraging plan. [ABSTRACT FROM AUTHOR]

Hydro-Que´bec wants to divert the Kipawa River in northwest Que´bec from its natural streambed. While the first time visitor is likely to emphatically proclaim the Kipawa River as the most beautiful, most serene place they have ever encountered, hydro consultants and engineers, disconnected from the attractiveness of that place, are making cost/benefit recommendations that marginalize the inherent value of a free-flowing Kipawa. This paper will discuss the following points: (1) The Kipawa River has its own inherent value, which is related to the cost of simulating threatened white-water habitats in general. (2) The costs of recreating white-water habitats are more understandable through the study of man-made white-water venues. (3) The cost to recreate or simulate a threatened white-water habitat should be factored into the cost of the hydro-project feasibility. The Kipawa River's own inherent value should be factored into the cost of the Tabaret Diversion Project. (4) Methods of gaining community acceptance should be public and open: independent third-party arbitartion is recommended. Use of monetary incentives to encourage public acceptance is unethical, immoral and unjustly biased against the survival of white-water habitats. (5) Recreational use of white-water habitats, like the Kipawa River are increasingly important engines of economic growth in Canada and around the world. [Copyright &y& Elsevier]

Most habitat maps are presented as if they were a certain fact, with no indication of uncertainties. In many cases, researchers faced with the task of constructing such maps are aware of problems with the modelling data and of decisions that they make within the modelling process that are likely to affect the output, but they find it difficult to quantify this information. In some cases they attempt to evaluate the modelled predictions against independent data, but the summary statistics have no spatial component and do not address errors in the predictions. It is proposed that maps of uncertainty would help in the interpretation of these summaries, and to emphasize patterns in uncertainty such as spatial clustering or links with particular covariates. This paper reviews the aspects of uncertainty that are relevant to habitat maps developed with logistic regression, and suggests methods for investigating and communicating these uncertainties. It addresses the problems of subjective judgement, model uncertainty and vague concepts along with the more commonly considered uncertainties of random and systematic error. Methods for developing realistic confidence intervals are presented along with suggestions on how to visualize the information for use by decision-makers. [Copyright &y& Elsevier]

Linear developments such as roads, seismic lines, and pipeline rights-of-way are common anthropogenic features in the boreal forest of Alberta. These features may act as barriers to the movement of threatened woodland caribou, Rangifer tarandus caribou. Seismic lines were not barriers to caribou movements, whereas roads with moderate vehicle traffic acted as semipermeable barriers to caribou movements. Twenty-two adult female and 1 adult male woodland caribou were captured and collared on February 17–19, 1998. Each caribou was fitted with a global positioning system collar. The greatest barrier effects were evident during late winter, when caribou crossed actual roads 6 times less frequently than simulated road networks. Semipermeable barrier effects may exacerbate functional habitat loss demonstrated through avoidance behavior. Caribou may cross roads less frequently than expected for a number of reasons, including an aversion to the physical barrier presented by a road and associated forest opening, vehicle traffic, or predation by humans. The effects of habitat fragmentation through habitat loss, avoidance, and the semipermeable barrier effects of roads should be considered in developing strategies to maintain woodland caribou populations in Alberta. Roads that act as semipermeable barriers to caribou movements may make the presence of caribou more predictable in space and time, and hence compromise the "spacing out" strategy that caribou adopt to reduce predation. The approach we have outlined in this paper is a useful method to quantify the effects of human developments acting as semipermeable barriers to animal movements and should be considered in the development of animal-movement models.

Pennatulacean octocorals (sea pens), one of the primary conservation targets of the Laurentian Channel Marine Protected Area (MPA) in eastern Canada, occur at high densities in some deep-sea sedimentary habitats. Considered important habitat-forming organisms for many megafaunal organisms, the effect of sea pens on nearby sedimentary macrofaunal communities remains unexplored. During two cruises in September 2017 and July 2018, we sampled 9 stations within the MPA (336-445 m depth), including sea pen fields and bare sedimentary habitats, targeting individual sea pens in a subset of the cores to assess small-scale effects. We evaluated macrofaunal density, taxonomic diversity, vertical distribution, community composition, and biological trait expression, and investigated variation between sea pen fields and other (bare) sedimentary habitats, as well as between cores with and without sea pen specimens. Using multivariate analyses, we identified abiotic and biotic drivers of macrofaunal community composition and biological trait expression. Enhancement of macrofaunal density and taxonomic diversity and higher percentages of organisms in the upper sediment layers characterized sea pen fields in autumn, with more variable results in summer. Community composition and biological trait expression consistently differed in sea pen fields compared to bare sedimentary habitats, with Pennatula sea pen density as one of the primary drivers of variation especially for community composition, along with other environmental drivers (depth, grain size, and organic matter quantity and quality). We also detected small-scale enhancement of macrofaunal diversity in cores containing sea pens at stations characterized by predominantly bare sediments. Our results indicate that macrofaunal communities within sea pen fields differ from those in bare sediments and we propose sea pens play a role in influencing those patterns by increasing food availability, stability, and small-scale heterogeneity in sedimentary habitats, acknowledging that other environmental factors might also play a role in determining the observed patterns. Characteristics of macrofaunal communities within sea pen fields also suggested potentially higher sensitivity to disturbance, which amplifies the need for protection of sea pen fields in deep-sea sedimentary environments. [ABSTRACT FROM AUTHOR]

In order to characterize spatial patterns and environmental and biological drivers of organic matter remineralization and nutrient regeneration in deep-sea sedimentary habitats, we measured fluxes of nitrate, nitrite, ammonium, phosphate, and silicate at the sediment-water interface during 48-h ex situ incubation of sediment cores. We sampled a total of 6 stations (351–445 m depth) inside the Laurentian Channel Marine Protected Area (MPA), on the outer continental shelf of Newfoundland (Canada). We assessed the potential effect of octocoral sea pens on uni- and multivariate benthic nutrient fluxes at large- and small-scale by comparing sea pen fields and other sedimentary habitats, and cores with and without sea pens. For each station, we evaluated a wide range of environmental variables, including physico-chemical and sedimentary factors, and we identified macrofaunal organisms inhabiting the cores, assessed their taxonomic diversity, community composition, and biological trait expression. Our analysis identified macrofaunal species richness and the density of a few key taxa, as well as environmental factors such as the quantity of sedimentary organic matter as the primary drivers of variation of multivariate benthic fluxes. Macrofauna explained up to 41% of the variation in benthic fluxes, whereas environmental variables only explained up to 19%, highlighting the importance of biodiversity for ecosystem functioning. Uni- and multivariate analysis of fluxes did not reveal clear spatial patterns within the MPA habitats and stations, and fluxes showed high small-scale variability. We found enhanced ammonium efflux rates associated with the presence of sea pens at both small- and large-scale, likely reflecting both direct and indirect effects of these soft corals on organic matter deposition and sedimentary biogeochemical processes. Sea pens, however, did not appear to influence other fluxes, leaving their role for organic matter remineralization unclear. The extreme complexity and small-scale heterogeneity of benthic processes, particularly within what appears to be a relatively homogeneous environment, underscores the need for further studies to facilitate generalizations of patterns and drivers of benthic nutrient fluxes at larger scales, which will ultimately enable the effective integration of ecosystem functioning into conservation strategies. [ABSTRACT FROM AUTHOR]

The southern extent of the range of Canada lynx (Lynx canadensis) extends into the contiguous United States in locations from Washington State in the West to northern Maine in the East. Lynx persist in various habitats across this range from high-elevation wilderness to intensively managed industrial forests. Lynx habitat use at the species' southern range boundary was poorly understood before the species was listed as Threatened under the Endangered Species Act in 2000, because most research had been conducted in boreal forest. The papers in this special section outline a variety of questions regarding lynx populations at the southern extent of their range, address topics on lynx space use and denning habitat, and provide generally consistent results in terms of features important to lynx habitat use despite the wide variation in specific habitats among the study areas. [ABSTRACT FROM AUTHOR]