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A key to extinct Eocene species of Atomaria 1. Body size smaller than 1.5 mm.......................................................................... 2 - Body size larger than 1.5 mm........................................................................... 4 2. Body size 1.0– 1.2 mm. Antennomere 3 short, subquadrate, as long as wide, much shorter than antennomere 2. Pronotal disc convex.............................................................................................. 3 - Body size 1.35 mm. Antennomere 3 elongated, 1.5 times as long as wide, slightly shorter than antennomere 2. Antennomere 5 conical, elongated, 1.5× as long as wide. Pronotal disc flat. Lateral side borders of pronotum visible from above only in basal half of its length. Bitterfeld amber......................................... A. saxonica Lyubarsky et Perkovsky 3. Lateral side borders of pronotum visible from above along entire its length. Pronotum not narrowed toward base. Pronotum maximum width before the middle of its length. Antennomere 5 conical, elongated, 1.5× as long as wide. Body size 1.0 mm. Baltic amber............................................................ A. groehni Perkovsky et Lyubarsky - Lateral side borders of pronotum visible from above only in basal half of its length. Pronotum narrowed toward base. Pronotum maximum width in the middle of its length. Antennomere 5 conical, nearly as long as wide, 1.1× as long as wide. Body size 1.19 mm. Rovno amber............................................................... A. perkovskyi sp. nov. 4. Pronotal disc convex.Antennomere 3 about 2× as long as wide.Antennomeres 9 and 10 strongly transverse. Pronotum maximum width nearly in the middle of its length. Base of pronotum with shallow depression. Body size 1.54 mm. Baltic amber......................................................................................... A. alekseevi sp. nov - Pronotal disc flat. Antennomere 3 about 1.5× as long as wide. Antennomere 10 subquadrate, nearly as long as wide. Pronotum maximum width much behind the middle of its length. Base of pronotum without depression. Body size 1.65 mm. Baltic amber.............................................................. A. gedanicola Lyubarsky et Perkovsky, Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on page 288, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693

Atomaria (Anchicera) perkovskyi sp. nov. (Figs 3–4) Type material. Holotype: collection number No 6832 [MAIG] (ex coll. Jonas Damzen JDC-10149 R); adult, sex unkown. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 38× 52 mm and a maximum thickness of 15 mm; preserved without supplementary fixation. Ventral side of specimen completely obscured by milky opacity. Syninclusions: four small Nematocera (Diptera) specimens. Type stratum. Rovno amber, Late Eocene (Perkovsky et al. 2007). Type locality. Rivne Oblast (region), Ukraine. Description. Measurements: body length (from anterior margin of head to elytral apex along midline) about 1.19 mm, body maximum width across both elytron 0.50 mm; head length 0.16 mm, head maximum width across eyes 0.27 mm; pronotum length 0.30 mm, pronotum maximum width 0.38 mm; elytra length 0.73 mm, elytra maximum width 0.50 mm. Body broadly elongate, moderately arched, slightly convex; integument uniformly darkbrown (as preserved); sparsely covered with semierect pubescence. Head transverse, about 1.7× as wide as long; coarsely punctate, distance between punctures equal to 0.5-1.0× diameter of one puncture, punctation denser laterally. Compound eyes hemispherical, convex, rather large, with large facets. Antennae inserted close to eyes, under lateral margin of forehead; antennal insertions widely separated; distance between antennal insertions larger than distance between antennae and аnterior margin of eye. Antennae 11-segmented with 3-segmented loose club, slender, short, reaching about to posterior one-third of pronotal length; antennomere 1 subcylindrical, elongate, 1.6× as long as wide; antennomere 2 conical, elongate, 1.5× as long as wide, slightly shorter than antennomere 1; antennomere 3 conical, short, nearly as long as wide, narrower and distinctly shorter than antennomere 2; antennomere 4 nearly as long as wide; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; antennomeres 6–8 trapezoidal, as long as wide; antennomeres 9–10 trapezoidal, strongly transverse, 1.5× as wide as long; antennomere 11 widely oval, with rounded apex, nearly as long as wide; relative length ratios of antennomeres 1–11 equal to 16:12:7:7:8:5:5:6:8:10:16. Pronotum transverse, 1.2× as wide as long, slightly narrowed posteriad and anteriad, with maximum width in middle of its length; with slightly convex disc; with finely margined posterior and lateral sides; lateral side borders visible from above only in basal half. Pronotal punctation moderately coarse and dense, distance between punctures equal to 1.5-2.0× diameter of one puncture; puncture diameter slightly smaller than eye facet diameter. Posterior and anterior angles obtuse. Anterior edge weakly sinuate, without excision; posterior edge strongly bisinuate, lobed, with shallow depression and basal pits; lateral edges widely rounded, without callosity and teeth. Scutellar shield small, oval, strongly transverse, 2.0× as wide as long, covered with fine punctation. Elytra short-oval, moderately convex, with maximum width about in anterior one-third of its length, 1.5× as long as wide and 2.4 times as long as pronotum length; completely covering the abdomen; lateral sides widely rounded; humeri rounded, slightly prominent. Elytral punctation irregular, moderately coarse and dense; (nearly as pronotal punctation), distance between punctures equal to 1.5-2.0× diameter of one puncture. Epipleura incomplete. Macropterous, metathoracic wings fully developed. Legs slender, long, relatively similar in shape, finely and densely punctate. Femora elongate-oval, flattened. Pretarsal claws simple, thin. Etymology. The species is named in honor of our dear colleague Dr. Evgeny E. Perkovsky (Kiev, Ukraine), leading researcher of inclusions in Rovno amber. Differential diagnosis. Atomaria perkovskyi sp. nov. can be distinguished from related species by a combination of characters: antennomere 3 short, nearly as long as wide, distinctly shorter than antennomere 2; antennomeres 9 and 10 strongly transverse; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; pronotum narrowed toward base; lateral side borders of pronotum visible from above only in basal half of its length; and pronotum maximum width in middle of its length. A new species can distinguished from all other extinct Eocene Atomaria representatives using the identification key provided below., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on pages 286-288, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693, {"references":["Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Taraschuk, M. V. (2007) A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples. African Invertebrates, 48, 229 - 245. [https: // hdl. handle. net / 10520 / EJC 84578]"]}

Atomaria (Anchicera) alekseevi sp. nov. (Figs 1–2) Type material. Holotype: collection number No 6831 [MAIG] (ex coll. Jonas Damzen JDC-10145); adult, sex unkown. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 17× 25 mm and a maximum thickness of 5 mm; preserved without supplementary fixation. Syninclusions: one Carabidae (Coleoptera) specimen, one small Nematocera (Diptera) specimen, and numerous stellate trichomes of Fagaceae. Type stratum. Baltic amber; Middle–Late Eocene (Sadowski et al. 2017; Seyfullah et al. 2018; Bukejs et al. 2019; Kasiński et al. 2020; Iakovleva et al. 2021). Type locality. Yantarny village (formerly Palmnicken), the Kaliningrad Region, Russia. Description. Measurements: body length (from anterior margin of head to elytral apex along midline) about 1.54 mm, body maximum width across both elytron 0.69 mm; head length 0.23 mm, head maximum width across eyes 0.38 mm; pronotum length 0.35 mm, pronotum maximum width 0.54 mm; elytra length 0.96 mm, elytra maximum width 0.69 mm. Body broadly elongate, moderately arched, slightly convex; integument uniformly lightbrown (as preserved); sparsely covered with semierect pubescence. Head transverse, about 1.7× as wide as long; coarsely punctate, distance between punctures nearly equal to diameter of one puncture. Compound eyes hemispherical, convex, rather large, with large facets. Antennae inserted close to eyes, under lateral margin of forehead; antennal insertions widely separated; distance between antennal insertions larger than distance between antennae and аnterior margin of eye. Antennae 11-segmented with 3- segmented loosed club, slender, rather long, reaching slightly beyond pronotal base; antennomere 1 subcylindrical, slightly elongate, 1.3× as long as wide; antennomere 2 conical, elongate, 1.6× as long as wide; antennomere 3 conical, elongate, about 2× as long as wide, distinctly narrower and slightly shorter than antennomere 2; antennomere 4 slightly elongate, 1.2× as long as wide; antennomere 5 conical, elongate, 1.5× as long as wide; antennomeres 6–8 trapezoidal, nearly as long as wide; antennomeres 9–10 trapezoidal, strongly transverse, 1.3–1.4× as wide as long; antennomere 11 widely oval, with rounded apex, 1.2× as long as wide; relative length ratios of antennomeres 1–11 equal to 10:10:7:5:6:5:5:5:6:7:13. Pronotum transverse, 1.5× as wide as long, narrowed posteriad and anteriad, with maximum width slightly beyond middle; with convex disc; with finely margined posterior and lateral sides; lateral side borders visible from above only in basal half. Pronotal punctation coarse and dense, distance between punctures equal to 0.5-1.5× diameter of one puncture, punctation distinctly sparser in posterior half; puncture diameter subequal to eye facet diameter; pronotal punctation larger than punctation of head. Posterior angles obtuse; anterior angles obtuse, slightly projecting anteriad. Anterior edge weakly sinuate, without excision; posterior edge strongly bisinuate, lobed, with shallow depression and basal pits; lateral edges widely rounded, without callosity and teeth, with distinct crenulation. Prohypomera slightly impressed; with sparse, fine punctation. Prosternum with disc convex; sparsely covered with coarse punctation, distance between punctures distinctly larger than diameter of one puncture. Prostenal process convex, rather narrow, with straight posterior margin, slightly dilated apically. Procoxal cavities apparently closed externally. Scutellar shield small, suboval, strongly transverse, 2.2× as wide as long, covered with fine punctation. Elytra short-oval, moderately convex, with maximum width in anterior one-third of its length, 1.4× as long as wide and 2.7 times as long as pronotum length; completely covering abdomen; lateral sides widely rounded; humeri rounded, slightly prominent. Elytral punctation irregular, coarse and dense; punctures in basal part slightly larger than punctures on pronotal disc, and distance between punctures equal to 1.5-2.0× diameter of one puncture. Epipleura narrowed posteriorly, incomplete, extending about to abdominal ventrite 2. Metaventrite with sparse, coarse punctation; disc convex. Macropterous, metathoracic wings fully developed. Legs slender, long, relatively similar in shape, finely and densely punctate. Procoxae large, widely suboval, transverse; mesocoxae round; metacoxae suboval, strongly transverse. Femora elongate-oval, flattened. Tibiae thin, almost straight, nearly as long as femora. Tarsi long and thin; tarsomeres simple, not lobed; tarsal formula 5-5-5. Abdomen with five visible, freely articulated ventrites; ventrite 1 sparsely covered with rather coarse punctation, ventrites 2–5 with small and denser punctures. Relative length ratios of ventrites 1–5 equal to 31:10:10:9:11 (measured medially). Etymology. The species is named in honor of our dear colleague Dr. Vitalii I. Alekseev (Kaliningrad, Russia), leading researcher of Coleoptera inclusions in Baltic amber. Differential diagnosis. Atomaria alekseevi sp. nov. can be distinguished from related species by a combination of characters: antennomere 3 elongated, about 2× as long as wide, slightly shorter than antennomere 2; antennomeres 9 and 10 strongly transverse; pronotal disc convex; pronotum narrowed toward base; lateral side borders of pronotum visible from above only in basal half of its length; pronotum maximum width nearly in middle of its length. A new species can distinguished from all other extinct Eocene Atomaria representatives using the identification key provided below., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on pages 284-286, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693, {"references":["Sadowski, E. - M., Seyfullah, L. J., Schmidt, A. R. & Kunzmann, L. (2017) Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia, 106, 1 - 73.","Seyfullah, L. J., Beimforde, C., Dal Corso, J., Perrichot, V., Rikkinen, J. & Schmidt, A. R. (2018) Production and preservation of resins - past and present. Biological Reviews, 93, 1684 - 1714. https: // doi. org / 10.1111 / brv. 12414","Bukejs, A., Alekseev, V. I. & Pollock, D. A. (2019) Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa, 4664 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4664.2.8","Kasinski, J. R., Kramarska, R., Slodkowska, B., Sivkov, V. & Piwocki, M. (2020) Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 bore hole, Kaliningrad region, Russia). Geological Quarterly, 64, 29 - 53. https: // doi. org / 10.7306 / gq. 1513","Iakovleva, A. I., Mychko, E. V. & Aleksandrova, G. N. (2021) New dinocyst data from amber quarry \" Primorsky \" (Kaliningrad region): clarification of the deposits age as base for the revision of paleogeographic reconstructions of Danish-Polish strait in late Paleogene. In: Golubev, V. K. & Nazarov, V. N. (Eds), Paleostrat- 2021. Annual meeting (scientific conference) of the Paleontological group of the Moscow Society of Naturalists and the Moscow branch of the Russian Paleontological Society, Russian Academy of Sciences (Moscow, 25 - 26 January 2021). Abstracts. Borissiak Paleontological Institute Press, Moscow, pp. 77 - 78. [in Russian]"]}

Subgenus Anchicera Thomson, 1863 Note. The fossil specimens under consideration are placed in Cryptophagidae based on antennae 11-segmented with 3-segmented loose club; tarsal formula 5-5-5; incomplete epipleura; elytra completely covering abdomen, with irregular punctation; procoxal cavities closed externally; tarsomeres simple, not lobed; and abdominal ventrite 1 longest. A new extinct species are assigned to the subgenus Anchicera within the genus Atomaria based on the combination of the following characters: distance between antennal insertions larger than distance between antennae and аnterior margin of eye; and lateral sides of elytra widely rounded, not parallel., Published as part of Lyubarsky, Georgy Yu. & Bukejs, Andris, 2022, TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins, pp. 283-289 in Zootaxa 5188 (3) on page 284, DOI: 10.11646/zootaxa.5188.3.5, http://zenodo.org/record/7091693

Two new species of Atomaria Stephens, 1829 from China and Taiwan are described and illustrated: Atomaria (Atomaria) cooterisp. nov. and Atomaria (Atomaria) johnsoni sp. nov. New records for China and Taiwan are added: Atomaria (Atomaria) klapperichi Johnson, 1971 and Atomaria (Atomaria) plecta Lyubarsky, 1995. New subgeneric combinations are proposed for Atomaria (Atomaria) klapperichi, Atomaria (Atomaria) plecta and Atomaria (Atomaria) torrida Johnson, 1971 SI