Search

Blacus (Contochorus) capillosus sp. nov. (Figs 1–2) Material examined. Holotype: ♀, China, Shaanxi, Qinling, Tiantai Mtn, 4.IX.1999, Chen Xuexin, No. 991398 (ZJUH). Paratype: 1♀, China, Shaanxi, Qinling, Tiantai Mtn, 3.IX.1999, He Junhua, No. 990176 (ZJUH). Description. Female. Length of body 2.4 mm, length of fore wing 2.35 mm. Body setosity. Body with dense setae, although sparser on mesopleuron. Head. Width of head in dorsal view 1.4 × its length (Fig. 2A); antennomeres 16 (Fig. 2C); length of first flagellomere 1.3 × length of second flagellomere; length of first, second and penultimate flagellomeres 4.6, 4.2 and 1.6 × their width, respectively; length of maxillary palp 1.0 × height of head; OOL: diameter of posterior ocellus: POL = 22:9:11; eyes with short dense setae (Fig. 2A); length of eye in dorsal view 0.85 × length of temple; occiput almost level (Fig. 2A); face with dense setae; temple with dense setae; malar suture present, length of malar space 1.75 × basal width of mandible. Mesosoma. Length of mesosoma 1.45 × its height; pronotal sides coarsely reticulate ventrally, dorsally smooth; area behind prepectal carina less conspicuously crenulate (Fig. 2E); precoxal sulcus narrow, with a few very short crenulae (Fig. 2E); notauli absent (Fig. 2D), without transverse carinae anteriorly; scutellar sulcus with medio-longitudinal carina (Fig. 2D); scutellum smooth, its lateral carina present, not protruding apically; surface of propodeum smooth between carinae, its medial area rectangular (Fig. 2F). Wings. Fore wing: first discal cell widely truncate anteriorly (Fig. 2H); vein r 1.2 × longer than maximum width of pterostigma; 1-CU1:2-CU1 = 13:21 (Fig. 2H). Legs. Hind coxa smooth, but with a distinct carina dorsally (Fig. 2F); length of femur, tibia and basitarsus of hind leg 6.2, 9.9 and 11.0 × their width, respectively; fore and mid claws with bristles (Figs 2I, 2J). Metasoma. Length of first tergite 2.6 × its apical width, almost parallel-sided, its surface rugulose, dorsal carinae complete (Fig. 2F); second tergite rugose; length of ovipositor sheath 0.10 × as long as fore wing. Colour. Body reddish-brown; palpi, mandible, tegulae, fore and middle legs, hind coxa and trochanter yellow; pterostigma, parastigma, veins, ovipositor, ovipositor sheath and all basitarsus brown; bristles of fore and mid claws blackish; wing membrane slightly infuscate. Variation. Length of first tergite 2.4–2.6 × its apical width. Male. Unknown. Biology. Unknown. Distribution. China (Shaanxi). Etymology. The specific name capillosus refers to the setose body and is derived from “ capillosus ” (Latin for hairy). Notes. The new species is similar to B. (C). turbidus Papp, 1985. Both species have medial area of propodeum rectangular, first discal cell of fore wing widely truncate anteriorly, and area behind prepectal carina less conspicuously crenulate than B. (C). mellitarsis and B. (C). epomidus but B. (C). capillosus differ by having 16 antennomeres of female (21–23 in B. (C). turbidus), lateral carina of scutellum not protruding apically (strongly protruding apically in B. (C). turbidus), mid claws of female with blackish bristles (only setose in B. (C). turbidus), length of eye of ♀ in dorsal view 0.85 × temple (1.3–1.6 × in B. (C). turbidus) and notauli absent (complete in B. (C). turbidus)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on pages 278-279, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232."]}

Blacus (Contochorus) planus sp. nov. (Figs 5–6) Material examined. Holotype: ♀, China, Hunan, Shimen, Huping Mtn, Xiangbizigou, 8.VII.2009, Ma Li, No. 200903023 (ZJUH). Paratype: 1♀, China, Zhejiang, Anji, Longwang Mtn, 21.VII.1995, Wu Hong, No. 971319 (ZJUH). Description. Female. Length of body 2.1 mm, length of fore wing 1.8 mm. Body setosity. Body with long pubescence, pubescence on head, side of pronotum and mesopleuron sparser than on remainder of body. Head. Head longitudinal and flat in dorsal view, width of head in dorsal view 1.1 × its length (Fig. 6A); antennomeres 16 (Fig. 6C); length of first flagellomere 1.2 × length of second flagellomere; length of first, second and penultimate flagellomeres 4.0, 3.3 and 1.4 × their width, respectively; length of maxillary palp 0.7 × height of head; OOL: diameter of posterior ocellus: POL = 16:6:7; eyes with short sparse setae (Fig. 6A); length of eye in dorsal view 0.8 × length of temple; occiput concave (Fig. 6A); face with dense long pubescence; temple smooth; malar suture weakly developed, length of malar space 1.5 × basal width of mandible. Mesosoma. Length of mesosoma 1.5 × its height; pronotal sides coarsely reticulate ventrally, dorsally smooth; area behind prepectal carina less conspicuously crenulate (Fig. 6E); precoxal sulcus narrow, with short crenulae (Fig. 6E); notauli absent posteriorly (Fig. 6D), impressed and crenulate anteriorly (Fig. 6D), without transverse carinae anteriorly; scutellar sulcus with medio-longitudinal carina (Fig. 6D); scutellum smooth, its lateral carina lamelliform, not protruding apically; surface of propodeum smooth between carinae, its medial area almost rectangular (Fig. 6F). Wings. Fore wing: first discal cell widely truncate anteriorly (Fig. 6I); vein r 0.6 × as long as maximum width of pterostigma; 1-CU1:2-CU1 = 1:3. Legs. Hind coxa smooth, but with a distinct carina dorsally (Figs 6G, 6H); length of femur, tibia and basitarsus of hind leg 5.0, 9.1 and 8.4 × their width, respectively; fore and mid claws with bristles (Figs 6J, 6L). Metasoma. Length of first tergite 2.9 × its apical width, parallel-sided, its surface rugulose, dorsal carinae complete (Fig. 6G); second tergite smooth; length of ovipositor sheath 0.13 × fore wing. Colour. Reddish-brown; palpi, mandible, tegulae, three basal segments of antenna, fore and middle legs, hind coxa and trochanter yellow; remainder of antenna, pterostigma, parastigma, veins, ovipositor, ovipositor sheath and all basitarsi brown; bristles of fore and mid claws blackish; wing membrane slightly infuscate. Variation. Precoxal sulcus with short to medium-sized striae. Male. Unknown. Biology. Unknown. Distribution. China (Hunan, Zhejiang). Etymology. The specific name planus refers to the rather flattened head and is derived from “ planus ” (Latin for flat). Notes. The new species is similar to B. (C). glaber van Achterberg, 1976.In both species the middle lobe of the mesoscutum lacks the strong carinae anteriorly, lateral carina of scutellum not strongly protruding apically, and middle claws with blackish bristles, but the two species differ by having 16 antennomeres of ♀ (17 in B. (C). glaber), notauli incomplete (complete in B. (C). glaber), medial area of propodeum rectangular and somewhat longer than wide (quadrangular, its height subequal to its width in B. (C). glaber), width of head in dorsal view 1.1 × its length (1.4 × in B. (C). glaber), length of eye in dorsal view 0.8 × length of temple (1.1 × in B. (C). glaber) and length of hind femur 5.0 × its width (7.0 × in B. (C). glaber)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on pages 281-283, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324.","van Achterberg, C. (1976) A revision of the tribus Blacini (Hym., Braconidae, Helconinae). Tijdschrift voor Entomologie, 118, 159 - 322."]}

Blacus (Contochorus) turbidus Papp, 1985 (Figs 7–8) Blacus (Ganychorus) turbidus Papp, 1985: 229. Blacus (Contochorus) turbidus: van Achterberg, 1988: 78; Papp, 1993: 159; Ahmad & Ahmed, 2008: 84. Material examined. 3♀♀, China, Hainan, Jianfengling, Tianchi, 25. XI.2008, Wang Manman, No. 200903082; Tan Jiangli, No. 200903085; Bawangling, 26.XI.2008, Wang Manman, No. 200805647; 1♀, China, Yunnan, Xishuangbanna, 28.XI.1993, Fan Xiang, No. 940417 (ZJUH). Distribution. China (Hainan, Yunnan); India; Nepal; Thailand. New record for China., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on pages 285-287, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232.","van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324.","Papp, J. (1993) New braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 4. Annales historico-naturales Musei nationalis hungarici, 84, 155 - 180.","Ahmad, Z. & Ahmed, Z. (2008) Description of a new species of the genus Blacus Nees (Hymenoptera: Braconidae), along with a key to Indian species. Journal of the Bombay Natural History Society, 105 (1), 84 - 85."]}

Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu, Chen, Xue-Xin (2023): The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China. Zootaxa 5306 (2): 277-287, DOI: 10.11646/zootaxa.5306.2.7, URL: http://dx.doi.org/10.11646/zootaxa.5306.2.7

Key to world species of subgenus Contochorus van Achterberg 1 Posterior half of notauli absent (Figs 2D, 6D); antennomeres of ♀ 16; length of eye of ♀ in dorsal view 0.8 × length of temple.............................................................................................. 2 - Posterior half of notauli complete (Figs 4D, 8D); antennomeres of ♀ 17–23; length of eye of ♀ in dorsal view 1.1–2.1 × length of temple............................................................................................ 3 2 Vein r of fore wing much shorter than maximum width of pterostigma (Fig. 6I); third antennomere yellow; head longitudinal and flat in dorsal view, width of head in dorsal view 1.1 × its length; notauli present anteriorly; vein 2-CU1 of fore wing 3 × longer than vein 1-CU1 (Fig. 6I); second tergite smooth...................................... B. ( C.) planus sp. nov. - Vein r of fore wing about as long as maximum width of pterostigma (Fig. 2H); third antennomere dark brown; width of head in dorsal view 1.4 × its length; notauli absent anteriorly; vein 2-CU1 of fore wing 1.6 × longer than vein 1-CU1 (Fig. 2H); second tergite rugose.................................................................... B. ( C.) capillosus sp. nov. 3 Antennomeres of ♀ 17; notauli anteriorly smooth, without transverse carinae; middle claw of ♀ with blackish bristles; medial area of propodeum quadrangular, its height subequal to its width; width of head in dorsal view 1.4 × its length; lateral carina of scutellum slightly protruding apically........................................ B. ( C.) glaber van Achterberg, 1976 - Antennomeres of ♀ 21–23; notauli with two transverse carinae anteriorly; middle claw of ♀ without blackish bristles; medial area of propodeum rectangular somewhat longer than wide; head transverse and flat in dorsal view, width of head in dorsal view 1.6–1.8 × its length; lateral carina of scutellum strongly protruding apically....................................... 4 4 First discal cell of fore wing widely truncate anteriorly; length of eye of ♀ in dorsal view 1.3–1.6 × length of temple; notauli comparatively narrow (Fig. 8D); area behind prepectal carina less conspicuously crenulate..... B. ( C.) turbidus Papp, 1985 - First discal cell of fore wing narrowly truncate anteriorly; length of eye of ♀ in dorsal view 1.7–2.1 × length of temple; notauli comparatively wide (Fig. 4D); conspicuously crenulate behind prepectal carina.................................... 5 5 Vein M+CU of hind wing shorter than vein 1-M; notauli widened anteriorly; eye of ♀ in dorsal view 2.0–2.1 × length of temple; hind tarsus of ♀ somewhat shorter than hind tibia; tarsi (pale) brownish-yellow....................................................................................................... B. ( C.) mellitarsis van Achterberg, 1988 - Vein M+CU of hind wing subequal to vein 1-M; notauli evenly wide anteriorly and posteriorly; eye of ♀ in dorsal view approx. 1.7 × length of temple; hind tarsus of ♀ as long as hind tibia; tarsi whitish-yellow...................................................................................................... B. ( C.) epomidus van Achterberg, 1988, Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on page 278, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["van Achterberg, C. (1976) A revision of the tribus Blacini (Hym., Braconidae, Helconinae). Tijdschrift voor Entomologie, 118, 159 - 322.","Papp, J. (1985) Three new Blacus Nees, 1818 species from India (Hym., Braconidae: Blacinae). Annales historico-naturales Musei nationalis hungarici, Zoology, 77, 227 - 232.","van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324."]}

Blacus (Contochorus) mellitarsis van Achterberg, 1988 (Figs 3–4) Blacus (Contochorus) mellitarsis van Achterberg, 1988: 77. Material examined. 1 ♀, China, Hainan, Diaoluo Mtn., 16–17. XII.2006, Weng Liqiong, No. 200802539 (ZJUH). Distribution. China (Hainan); Indonesia. New record for China., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Contochorus van Achterberg, 1976 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, pp. 277-287 in Zootaxa 5306 (2) on page 281, DOI: 10.11646/zootaxa.5306.2.7, http://zenodo.org/record/8058845, {"references":["van Achterberg, C. (1988) Revision of the subfamily Blacinae Foerster (Hymenoptera: Braconidae). Zoologische Verhandelingen Leiden, 249, 1 - 324."]}

Blacus (Hysterobolus) xui sp. nov. (Figs. 15–16) Material examined. Holotype: ♀, Guangdong, Fuyuan, Nanling, 4.VIII.2004, Xu Zaifu, No. 20049864 (ZJUH). Paratypes: 3 ♀♀, Guangdong, Fuyuan, Nanling, 4.VIII.2004, Xu Zaifu, No. 20049813, 20049819, 20049936 (ZJUH). Description. Female. Length of body 2.9 mm, length of fore wing 2.4 mm. Body setosity. Body with long pubescence, pubescence on propodeum denser than the remainder. Head. Antennomeres 19; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 2.5, 1.5 and 2.7 × their width, respectively; length of maxillary palp 1.0 × height of head; OOL: diameter of posterior ocellus: POL = 11:4:5; frons smooth, with a longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.0 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.6 × its medial height; tentorial pits large, distance between pits 2.1 × distance from pit to eye; malar suture absent, length of malar space 2.4 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with distinct median carina and four carinae on both sides; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.1 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 4:5. Hind wing: 1r-m:1-M = 19:25; M+CU subequal to1-M; 2-1A present and short, slightly branched down. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 4.8, 8.9 and 8.0 × their width, respectively; fore tarsal claws with brownish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.6; remainder of tergites smooth; length of ovipositor sheath 0.21 × length of fore wing. Colour. Brown; dorsal head (stemmaticum darker), scapus and dark brown; palpi, labrum, tegulae, fore and middle legs yellow; several basal flagellomeres brown, gradually paler towards apex, but several apical flagellomeres dark brown; parastigma and pterostigma basally white; scutellum reddish-brown; most mesoscutum black, partly yellowish-red; mesopleuron, propodeum and metasomal first tergite black; remainder of metasomal, ovipositor and sheath, veins and pterostigma brown; hind leg blackish-brown; all basitarsi apically brown; wing membrane slightly infuscate. Variation. Length of first tergite 1.7–1.8 × its apical width; length of apical flagellomeres 2.7–2.9 × their width, dorsal head dark brown to blackish; most mesoscutum dark brown to black, partly yellowish-red to reddish-brown; hind leg dark brown to blackish-brown. Male. Unknown. Biology. Unknown. Distribution. China (Guangdong). Etymology. The specific name refers to the collector of the species, the late Prof. Zai-fu Xu, who was a wellknown specialist of Chrysidoidea. Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg,1988 because the scutellum is coarsely rugose, the hind tibia infuscated medially, the first metasomal tergite widened apically, the face transversely rugose, length of eye in dorsal view approx. 1.1 times temple, and fore claw with brownish bristles, but differs by having the precoxal sulcus with long striae (with a few medium-sized striae in B. fuscitibialis), dorsal carinae distinct in basal 0.6 of first tergite (basal 0.7), length of ovipositor sheath 0.21 × length of fore wing (0.18 ×), vein 2-SR of the fore wing 1.9 × longer than vein r (1.6 x), vein 1-CU1 oblique and nearly as long as vein 2-CU1 (horizontal and shorter than vein 2-CU1), and most of mesoscutum dark brown to black (mesoscutum reddish-brown)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 491-495, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) niger sp. nov. (Figs. 11–12) Material examined. Holotype: ♀, China, Yunnan, Baoshan, Gaoligongshan Nature park (24º50′N, 98º46′E), 11.V.2009, Wang Manman, No. 200904572 (ZJUH). Description. Female. Length of body 2.35 mm, length of fore wing 2.4 mm. Body setosity. Body with long pubescence, pubescence on mesopleuron sparse, propodeum and side of pronotum ventrally densely setose. Head. Antennomeres 19; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 3.0, 1.5 and 3.0 × their width, respectively; length of maxillary palp 1.1 × height of head; OOL: diameter of posterior ocellus: POL = 20:7:9; frons smooth, with a shallow longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.0 × temple; occiput slightly concave; face with shallow transversely rugae; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.7 × its medial height; tentorial pits large, distance between pits 1.7 × distance from pit to eye; malar suture absent, length of malar space 1.8 ×basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.35 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some medium-sized striae, only one stria reaching the anterior edge of mesopleuron; notauli complete and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with distinct median carina and two weakly developed carinae (Fig. 12D); scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, slightly acute apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.1 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 2:3. Hind wing: 1r-m:1-M = 4:7; M+CU slightly less than 1-M; 2-1A absent. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.55, 9.6 and 7.1 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; remainder of tergites smooth; length of ovipositor sheath 0.23 × length of fore wing. Colour. Dark reddish-brown; dorsal head black; palpi, tegulae, ovipositor, fore and middle legs yellow; scape, clypeus reddish-brown; several basal flagellomeres yellowish-brown, gradually paler towards apex, but several subapical flagellomeres brown; mandible yellow; parastigma and base of pterostigma white; most of mesonotum black; metasomal tergite (except first) brown; ovipositor sheath, veins and pterostigma reddish-brown; hind legs coxa brown, femur and tibia blackish-brown; all apical basitarsus (basal yellow) and claws brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan) Etymology. The specific name refers to the black mesonotum and is derived from “ niger ” (Latin for black). Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg,1988 because the scutellum is coarsely rugose, the malar suture absent, the hind tibia infuscated medially, the first metasomal tergite widened apically, the face transversely rugose, and the precoxal sulcus with medium-sized striae, but differs by having the fore claw with blackish bristles (brownish), dorsal carinae distinct in basal 0.5 (0.7), length of ovipositor sheath 0.23 × length of fore wing (0.18 ×), and the mesonotum black (mesoscutum reddish-brown)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 486-488, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) variegatus sp. nov. (Figs. 13-14) Material examined. Holotype: ♀, China, Chongqing, Jinfo Mtn, (29º3′N, 107º7′E), 18.VIII.2012, Huang Xinlei, No. 201207984 (ZJUH). Paratypes: 5 ♀ ♀, China, Zhejiang, Tianmu Mtn, 3-5.X.1983, Ma Yun, No. 834329; 25-29.VII.2011, yellow pan trap, No. 201501198; Chanyuan Temple, 9. V.1998, Zhao Mingshui, No. 20003401; Zhejiang, Fengyang Mtn, 1. VI.2005, Liu Jingxian, No. 200809039; Zhejiang, Longtang Mtn, 29. V.2012, Tang Pu, No. 201204960; 1 ♀, China, Hunan, Shimen, Huping Mtn, 12.VII.2009, Zeng Jie, No. 200903015; 1 ♀, China, Guizhou, Leigong Mtn, Tree Farm, 12.VII.2009, Zhang Hongying, No. 200606873 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.65 mm. Body setosity. Body with dense pubescence, pubescence on head and mesopleuron sparser, propodeum denser. Head. Antennomeres 18; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.0, 2.4, 1.2 and 2.8 × their width, respectively; length of maxillary palp 1.1 × height of head; OOL: diameter of posterior ocellus: POL = 24:8:9; frons smooth, with a shallow longitudinal groove in upper second and five groove between antennal sockets; eyes with short setose, sparse; length of eye in dorsal view 0.9 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, smooth; width of clypeus 2.4 × its medial height; tentorial pits large, distance between pits 1.55 × distance from pit to eye; malar suture absent, length of malar space 2.0 × basal width of mandible, mandible with long setae. Mesosoma. Pubescence on mesonotum sparser than remainder. Length of mesosoma 1.25 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate, with short transversely carina in anterior third; mesoscutal lobes rather convex, densely pubescent; scutellar sulcus with distinct median carina, and distinctly crenulate laterally; scutellum sculpture reduced and smooth medially, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, obtuse apically, lateral apical angle almost 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 5.0 × its maximum width; length of vein r 1.4 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 13:15. Hind wing: 1r-m:1-M = 9:17; M+CU about subequal to 1-M; 2-1A absent. Legs. Fore leg tibial spur large and long; hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 4.8, 9.3 and 8.0 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 2.0 × its apical width, widened apically, medio-dorsally rugose, both sides with longitudinally striae; dorsal carinae distinct in basal 0.4; remainder of tergites smooth; length of ovipositor sheath 0.16 × length of fore wing. Colour. Dark reddish-brown; dorsal head, propodeum and metasomal first tergite black; palpi, tegulae, fore and middle legs yellow; several basal flagellomeres reddish-brown, gradually paler towards apex, but two apical flagellomeres dark brown; clypeus and mandible yellowish-red; parastigma and bottom margin of pterostigma white; side of pronotum yellowish-red, remainder reddish-brown; most mesonotum and scutellum yellowish-red, mesoscutum partly brown; metasomal second tergite yellowish-red, remainder of metasomal tergites and sternum, ovipositor and sheath, veins and pterostigma brown; hind leg (except second, third and fourth tarsus) reddish-brown; all basitarsi apically brown; wing membrane slightly infuscate. Variation. Length of first tergite 1.8–2.1 × its apical width; penultimate flagellomere 1.25–1.3 (occasionally 1.4) × its width; length of fore wing 2.4–2.65 mm; length of ovipositor sheath 0.16–0.18 × length of fore wing; propodeal tubercles medium size to large; scutellum sculpture reduced, at most slightly rugose; head and propodeum dark reddish-brown; two (or at most three) apical flagellomeres dark brown. Male. Unknown. Biology. Unknown. Distribution. China (Chongqing, Hunan, Zhejiang, Guizhou) Etymology. The specific name refers to the particular different difference in colour of head and mesonotum and is derived from “ varius” (Latin for different). Notes. The new species is similar to B. (H.) fuscitibialis van Achterberg, 1988 because the malar suture is absent, the face transversely rugose, and the first metasomal tergite widened apically, but differs by having the precoxal sulcus with long striae (with a few medium-sized striae in B. fuscitibialis), hind leg (except second, third and fourth tarsal flagellomeres) reddish-brown (only hind tibia infuscated medially), fore claw with blackish bristles (brownish bristles), length of penultimate flagellomere 1.2 × its width (1.5 ×), dorsal carinae distinct in basal 0.4 of tergite (basal 0.7), and length of ovipositor sheath 0.16 × length of fore wing (0.18 ×). This new species is also similar to B. (H.) melanoapicalis sp. nov., but differs by having 18 antennomeres (19 antennomeres in B. melanoapicalis), occiput comparatively slightly concave (distinctly concave), and the colour of head and mesonotum particularly different (similar)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 488-491, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) chabarovi Belokobylskij, 1995 (Figs. 19–20) Hysterobolus chabarovi Belokobylskij, 1995: 458; Lee et al., 2020: 32. Material examined. 1 ♀, China, Chongqing, Jinfo Mt. (29º3′N, 107º7′E), 19.VIII.2012, Huang Xinlei, No. 201209624; 2 ♀♀, China, Ningxia, Longde, Liupanshan Nature Park, Sutai Tree Farm, 24. VI.2008, No. 200903090, 200903097; 1 ♀, Ningxia, Jingyuan, Liupanshan Nature Reserve, Hongxia Tree Farm, 1.VII.2008, Liu Jingxian, No. 200902112 (ZJUH). Distribution. China (Chongqing, Ningxia); Korea; Russia. New record for China. Remarks. The characters are largely consistent as described by Belokobylskij (1995), but the fore tarsal claws have weakly developed bristles., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on page 476, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467.","Lee, H. - R., Belokobylskij, S. A., Ku, D. - S. & Byun, B. - K. (2020) Eight newly recorded species of the genus Blacus (Hymenoptera: Braconidae: Blacinae) in Korea. Animal Systematics Evolution and Diversity, 36 (1), 31 - 34."]}

Blacus (Hysterobolus) flaviceps sp. nov. (Figs. 3-4) Material examined. Holotype: ♀, China, Yunnan, Baoshan, Gaoligongshan Nature Park (24º50′N, 98º46′E), 10.V.2009, Wang Manman, No. 200904546 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.6 mm. Body setosity. Body with long pubescence, entire head with long setae, pubescence on mesonotum sparser than on remainder of mesosoma. Head. Antennomeres 20; length of first flagellomere 1.7 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.4, 2.9, 1.75 and 3.3 × their width, respectively; length of maxillary palp subequal to height of head; OOL: diameter of posterior ocellus: POL = 11:4:3; frons smooth, with a shallow longitudinal groove in upper half; stemmaticum smooth; eyes with short sparse setae; length of eye in dorsal view 0.8 × temple; occiput concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate; width of clypeus 2.5 × its medial height; tentorial pits large, distance between pits twice distance from pit to eye; malar suture absent, length of malar space 2.8 × basal width of mandible. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex, densely pubescence; scutellar sulcus with median carina; scutellum reticulate, its lateral carina lamelliform; propodeal tubercles small, obtuse apically, lateral apical angle about 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.75 × its maximum width; length of vein r 1.0 × width of pterostigma; vein r arising almost from middle of pterostigma; 1-CU1:2-CU1 = 6:7. Hind wing: 1r-m:1-M = 1:2; M+CU slightly short than 1-M; 2-1A present and short, branched down. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.8, 11.4 and 10.0 × their width, respectively; fore tarsal claws missing. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; second tergite smooth; length of ovipositor sheath 0.17 × length of fore wing. Colour. Dark reddish-brown; head (dorsal and anterior), scape, palpi, tegulae, hind leg (except tibia) and ovipositor yellow; antenna yellow, gradually darkened towards apex; stemmaticum, pterostigma (basally paler), parastigma, veins, ovipositor sheath brown; edges of mesoscutum yellow, remainder of mesoscutum and scutellum reddish-brown, reddish-brown area on middle lobes remain separated from each other; dorsal side of pronotum yellow, remainder of pronotum, mesopleuron, propodeum and metasomal first tergite reddish-brown; second metasomal tergite and sternum yellow, remainder of metasomal tergites blackish; fore and middle legs light yellow; all basitarsi dark brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan). Etymology. The specific name refers to the yellow head and is derived from “ flavus ” (Latin for yellow) and “ caput ” (Latin for head). Notes. The new species differs from other species by having a yellow head, 20 antennomeres, length of basitarsus of hind leg 10.0 × its width, and length of vein r of fore wing almost equal to width of pterostigma., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 476-477, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) yunnanensis sp. nov. (Figs. 11–12) Material examined. Holotype: ♀, China, Yunnan, Ruili (24º1′N, 97º51′E), 29. IV.1981, He Junhua, NO. 812372 (ZJUH). Description. Female. Length of body 2.6 mm, length of fore wing 2.5 mm. Body setosity. Body with long setae, mesopleuron and side of pronotum glabrous, propodeum denser setose than head and mesonotum. Head. Antenna with at least 18 antennomeres (apex missing); length of first flagellomere 1.8 × second flagellomere; length of first and second flagellomere 4.9 and 2.7 × their width, respectively; length of maxillary palp 1.1 ×height of head; OOL:diameter of posterior ocellus:POL=20:8:11;frons with sparse setae and a deep longitudinal groove in upper second; eyes with short setose, sparse; length of eye in dorsal view 1.2 × temple; occiput slightly concave; face almost entirely smooth; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 2.5 × its medial height; tentorial pits large, distance between pits 2.7 × distance from pit to eye; malar suture present, length of malar space 2.0 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.35 × its height; side of pronotum largely smooth dorsally; precoxal sulcus narrow, with a few short striae, far from the anterior edge of mesopleuron (Fig. 18G); notauli complete, wide and crenulate; mesoscutal lobes rather convex, with long and soft pubescence; scutellar sulcus with median carina; scutellum (superficially) rugulose, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, acute apically, lateral apical angle slightly more than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.75 × its maximum width; length of vein r 1.2 × width of pterostigma; vein r arising from apical of pterostigma; 1-CU1:2-CU1 = 12:13. Hind wing: 1r-m:1-M = 20:27; M+CU almost subequal to 1-M; 2-1A absent. Legs. Hind coxa slightly rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.9, 11.5 and 7.1 × their width, respectively, fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 1.6 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.7; remainder of tergites smooth; length of ovipositor sheath 0.14 × length of fore wing. Colour. Reddish-brown; dorsal head black; palpi, tegulae, parastigma, fore and middle legs yellow; medial flagellomeres lighter in colour; mandible yellow, apex red; veins, pterostigma, ovipositor sheath, and metasomal tergite (except first) brown; ovipositor, hind legs coxa, femur and telotarsus yellowish-brown, tibia brown; all apical basitarsus (basal yellow) and claws brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Yunnan) Etymology. The specific name refers to the locality of the holotype. Notes. The new species is similar to B. (H.) nixoni Haeselbarth, 1973 because the malar suture is present, the scutellum (superficially) rugulose, the propodeal tubercle comparatively small, length of first tergite 1.6 × its apical width, flagellomeres long setose, and face almost entirely smooth, but differs by having the eighth flagellomere 2.2 × longer than wide (about 1.7 × in B. nixoni), length of malar space of 2.0 × basal width of mandible (1.5 ×), dorsal carinae distinct in basal 0.7 (0.9 ×), precoxal sulcus with less striae (with more striae), and body reddish-brown (black)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 495-497, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467."]}

Key to Chinese species of subgenus Hysterobolus Viereck 1 Vein M+CU of hind wing distinctly shorter than vein 1-M; penultimate flagellomere quadrate, about as long as wide (Fig. 6C); length of malar space 3.0 × to basal width of mandible; first metasomal tergite about 2.1 × longer than its apical width (Fig. 6F)......................................................................... B. ( H.) laculaticornis sp. nov. - Vein M+CU of hind wing about subequal to vein 1-M; penultimate flagellomere distinctly longer than wide (Fig. 8D); length of malar space at most 2.8 × basal width of mandible; first tergite usually 1.5–1.8 × longer than wide apically (Figs 2H, 4H, 8E, 10J, 12F, 16F, 18E), except in B. variegatus (Fig.14E)........................................................ 2 2 Length of vein r of fore wing almost equal to width of pterostigma; antennomeres 20; length of hind basitarsus 10.0 × its width; head yellow....................................................................... B. ( H.) flaviceps sp. nov. - Length of vein r of fore wing 1.1–1.5 × width of pterostigma; antennomeres less than 20; length of hind basitarsus 7.1–8.8 × its width; colour of head variable........................................................................... 3 3 Malar suture present................................................................................... 4 - Malar suture absent.................................................................................... 5 4 Vein 2-CU1 of fore wing 1.2 × longer than vein 1-CU1 (Fig. 18F); face largely smooth; precoxal sulcus narrow, striae very short (Fig. 18G)..................................................................... B. ( H.) yunnanensis sp. nov. - vein 2-CU1 of fore wing 2.2 × longer than vein 1-CU1(Fig. 20I); face transversely rugose; precoxal sulcus wide, striae long (Fig. 20F)............................................................ B. ( H.) chabarovi Belokobylskij, 1995 5 Hind tibia yellow or reddish-brown....................................................................... 6 - Hind tibia dark brown or blackish-brown.................................................................. 8 6 First metasomal tergite 1.5 × longer than its apical width (Fig. 8E); length of pterostigma 3.5 × its maximum width; length of terminal flagellomere 3.0 × its width; body generally black............................. B. ( H.) longiapicalis sp. nov. - First tergite 1.7–2.1 × longer than its apical width; length of pterostigma 5.0 × its maximum width; length of terminal flagellomere 2.8 × its width; body generally reddish-brown............................................................... 7 7 Length of first flagellomere 1.7 × second flagellomere; scutellum smooth medially; antennomeres 18; occiput slightly concave (Fig. 14A); hind tibia reddish-brown; 2–3 apical flagellomeres dark brown; first metasomal tergite about 2.1 × longer than its apical width (Fig. 14E)............................................................ B. ( H.) variegatus sp. nov. - Length of first flagellomere 1.4 × second flagellomere; scutellum rugose medially; antennomeres 19; occiput distinctly concave (Fig. 10A); hind tibia yellow; 5–6 apical flagellomeres dark brown; first tergite 1.7–1.8 × longer than its apical width (Fig. 10J)........................................................................ B. (H.) melanoapicalis sp. nov. 8 Fore claw of ♀ with blackish bristles (Fig. 12I); face weakly rugose (Fig. 12B); striae of precoxal sulcus medium-sized; vein 2-1A of hind wing absent (Fig. 12H)...................................................... B. ( H.) niger sp. nov. - Fore claw of ♀ with brownish bristles (Figs 2J, 16I); face coarsely rugose (Figs 2B, 16B); striae of precoxal sulcus long; vein 2-1A of hind wing present (Figs 2M, 18F).................................................................. 9 9 Terminal flagellomere 2.7–2.9 × longer than wide; first discal cell of fore wing acute (Fig. 16H); setose part of ovipositor sheath slightly longer, 0.21 × length of fore wing; mesoscutum largely dark brown to black; hind femur dark brown to blackish-brown............................................................................... B. ( H.) xui sp. nov. - Terminal flagellomere about 2.2 × longer than wide; first discal cell of fore wing truncate (Fig. 2L); setose part of ovipositor sheath slightly shorter, 0.18 × length of fore wing; mesoscutum yellowish-brown; hind femur yellow............................................................................................. B. ( H.) baishanzuensis sp. nov., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on page 473, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330, {"references":["Belokobylskij, S. A. (1995) New and rare species of the genus Blacus (Hymenoptera: Braconidae) from the Russian Far East. European Journal of Entomology, 92 (2), 449 - 467."]}

Blacus (Hysterobolus) laculaticornis sp. nov. (Figs. 5-6) Material examined. Holotype: ♀, China, Ningxia, Longde, Liupanshan Nature Park, Sutai Tree Farm, 22.VI.2008, Liu Jingxian, No. 200902135(ZJUH). Description. Female. Length of body 2.5 mm, length of fore wing 2.25 mm. Body setosity. Body with dense setae, setae on pronotum and propodeum denser than on mesonotum, mesopleuron and metasoma. Head. Antennomeres 18; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 3.6, 2.2, 1.0 and 2.2 × their width, respectively; length of maxillary palp 0.95 × height of head; OOL: diameter of posterior ocellus: POL = 4:1:2; frons smooth, with a deep longitudinal groove in upper half; stemmaticum with a distinct furrow from triangle centre to occipital carina; eyes with short sparse setae; length of eye in dorsal view 0.9 × temple; occiput slightly concave; face transversely rugose; temple smooth; clypeus convex, smooth; width of clypeus 2.3 × its medial height; tentorial pits large, distance between pits 1.8 × distance from pit to eye; malar suture absent, length of malar space 3.0 × basal width of mandible, mandible with long setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching anterior edge of mesopleuron; notauli complete, wide and simple; mesoscutal lobes convex; scutellar sulcus with distinct median carina; scutellum sculpture reduced, its lateral carina lamelliform; propodeal tubercles large (Figs 6F and 6H), obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 3.5 × its maximum width; length of vein r 1.3 × width of pterostigma; vein r arising slightly from apical of pterostigma; 1-CU1:2-CU1 = 2:3. Hind wing: 1r-m:1-M =1:2; M+CU distinctly less than 1-M; 2-1A present and short, branched posteriorly. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.0, 8.75 and 8.3 × their width, respectively; fore tarsal claws with blackish bristles. Metasoma. Length of first tergite 2.1 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.9; remainder of tergites smooth; length of ovipositor sheath 0.16 × length of fore wing. Colour. Reddish-brown; stemmaticum dark reddish-brown; palpi, tegulae, fore and middle legs yellow; antenna yellow (scape and apical flagellomeres reddish-brown); parastigma and base of pterostigma white; mandible, clypeus, side of pronotum dorsally and mesonotum yellowish-brown; side of pronotum ventrally, mesopleuron, propodeum and first metasomal tergite black; second and third tergite light yellow, remainder of metasomal tergites, ovipositor and sheath, veins and pterostigma brown; hind leg reddish-brown; all basitarsi and claws brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Ningxia). Etymology. The specific name laculaticornis refers to the square penultimate flagellomere and is derived from “ laculatus ” (Latin for four-cornered) and “ cornu ” (Latin for horn). Notes. The new species is similar to B. (H). mamillanus Ruthe, 1861 because the malar suture is absent, the propodeal tubercle rather large, the fore claw has blackish bristles, and length of the penultimate flagellomere1.3 times its width or less, but differs by having the malar space 3.0 × longer than basal width of mandible (2.2 × in B. mamillanus), second flagellomere 3.6 longer than wide (3.0 ×), length of first tergite 2.1 × its apical width (1.4 ×), and length of setose part of ovipositor sheath 0.16 × length of fore wing (0.18 ×)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 477-481, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Blacus (Hysterobolus) baishanzuensis sp. nov. (Figs. 1–2) Material examined. Holotype: ♀, China, Zhejiang, Qingyuan, Baishanzu, 18.VII.1994, Wu Hong, No. 946835 (ZJUH). Description. Female. Length of body 2.3 mm, length of fore wing 2.35 mm. Body setosity. Body with dense pubescence, pubescence on propodeum denser than on remainder. Head. Antennomeres 19; length of first flagellomere 1.6 × second flagellomere; length of first, second, penultimate and terminal flagellomere 4.75, 3.0, 1.3 and 2.2 × their width, respectively; length of maxillary palp 0.9 × height of head; OOL: diameter of posterior ocellus: POL = 11:3:5; frons smooth, with a very shallow longitudinal groove in upper half; eyes with short sparse setae; length of eye in dorsal view 1.0 × temple; occiput slightly concave; face transversely rugose; temple smooth; clypeus convex, slightly punctate, smooth; width of clypeus 3.0 × its medial height; tentorial pits large, distance between pits 2.3 × distance from pit to eye; malar suture absent, length of malar space 2.0 × basal width of mandible, mandible with setae. Mesosoma. Length of mesosoma 1.4 × its height; side of pronotum smooth dorsally, remainder coarsely reticulate; precoxal sulcus wide, with some long striae, reaching the anterior edge of mesopleuron; notauli complete, wide and crenulate; mesoscutal lobes rather convex; scutellar sulcus with distinct median carina and four weakly developed carinae; scutellum convex, reticulate, its lateral carina lamelliform, distinctly protruding apically; propodeal tubercles small, the left acute apically, the right obtuse apically, lateral apical angle slightly less than 90°; surface of propodeum rugulose. Wings. Fore wing: parastigma small; length of pterostigma 4.7 × its maximum width; length of vein r 1.3 × width of pterostigma; vein r arising slightly from apical of pterostigma; 1-CU1:2-CU1 = 11:15. Hind wing: 1r-m:1- M = 17:23; M+CU about subequal to 1-M; 2-1A present and short, branched posteriorly. Legs. Hind coxa coarsely rugose dorsally, with distinct dorsal carina; length of femur, tibia and basitarsus of hind leg 5.0, 9.2 and 7.7 × their width, respectively; fore tarsal claws with brownish bristles. Metasoma. Length of first tergite 1.8 × its apical width, widened apically, longitudinally rugose, dorsal carinae distinct in basal 0.5; remainder of tergites smooth; length of ovipositor sheath 0.18 × length of fore wing. Colour. Brown; scape, head (dorsal and frontal) dark brown, palpi, labrum, mandible, tegulae, all legs (except hind coxa and tibia, all basitarsi) yellow; flagellomeres dark brown, but medial flagellomeres yellow; parastigma and base of pterostigma white; side of pronotum and mesopleuron reddish-brown; mesoscutum yellowish-brown; propodeum and first metasomal tergite black; remainder of metasoma, ovipositor and sheath, veins and pterostigma brown; hind tibia apically dark brown; all basitarsi dark brown; wing membrane slightly infuscate. Male. Unknown. Biology. Unknown. Distribution. China (Zhejiang). Etymology. The specific name refers to the name of the collection site. Notes. The new species is similar to B. (H.) robustus Haeselbarth, 1973 because the scutellum is coarsely rugose, the malar suture absent, length of eye in dorsal view as long as temple, length of the penultimate flagellomere 1.3 × its width, and the hind tibia darkened apically, but differs by having a more setose body (sparser setose in B. robustus), the temples more narrowed behind eyes in dorsal view and also in anterior view, the notauli completely crenulate (at least partly simple), the fore claw with brownish bristles (blackish), length of the malar space 2.0 × basal width of mandible (1.7 ×), length of the first tergite 1.8 × its apical width (1.4–1.5 ×), and dorsal carinae of first tergite distinct in basal 0.5 (in basal 0.3)., Published as part of Lu, Qian-Yu, Achterberg, Cornelis Van, Tang, Pu & Chen, Xue-Xin, 2023, The discovery of the subgenus Hysterobolus Viereck, 1913 (Hymenoptera, Braconidae, Brachistinae, Blacini, Blacus) in China, with description of nine new species, pp. 472-498 in Zootaxa 5293 (3) on pages 473-476, DOI: 10.11646/zootaxa.5293.3.3, http://zenodo.org/record/7961330

Chelonus (Megachelonus) macrodontus sp. nov. Figs 9–10 Material examined. Holotype: female, China, Yunnan Prov., Tongbiguan, 23.VI.2005, Jinxian Liu, No. 200609605(ZJUH). Diagnosis. This species differs from the other species in the subgenus Megachlonus by having a pair of long and large teeth at apex of metasomal carapace (Figs 10G, 10H). Description. Holotype, female, length of body 5.3 mm, of fore wing 3.9 mm. Head. Antennomeres 28 (remaining, broken in holotype), basal half of antenna distinctly broadened, then rapidly tapering to apex; third antennomere 1.3 × longer than fourth; third, fourth and terminal antennomeres 4.7 ×, 3.7 × and 3.2 × longer than wide, respectively; 9-20 th antennomeres flattened, distinctly shorter than wide; head transverse, in dorsal view roundly contracted behind eyes; OOL: OD: POL= 15: 6: 10; longitudinal diameter of eye 2.0 × height of temples; vertex coarsely striate-rugose; frons distinctly concave, reticulate-rugose but smooth near antennal sockets; temple irregularly reticulate-rugose (Fig. 10E); face coarsely reticulate-rugose with longitudinal elements, clypeus flat and densely punctate, smooth and shining, length of malar space 1.6 × basal width of mandible (Fig. 10D); face densely setose in lateral view (Fig. 10F). Mesosoma. Length of mesosoma 1.4 × its height; mesoscutum areolate-rugose, but smooth and shiny medially; scutellum punctate medially, rugose laterally; scutellar suture comparatively deep; mesopleuron evenly convex, coarsely reticulate-rugose (Fig. 10B); propodeum coarsely reticulaterugose, with a distinct transversal carina connected with 4 blunt tubercles, lateral tubercles larger than medial ones (Fig. 10C). Wings. Length of fore wing 2.7 × its width; length of marginal cell 0.8 × pterostigma; vein 2-Rl just present, vein r shorter than 3-SR; r: 3-SR: SRl = 1: 1: 5; SRl more or less curved; parastigma distinctly swollen; 1-CU1: 2- CU1= 1: 2; vein CUlb present (Fig. 10A). Legs. Hind coxa densely short setose and striate-rugose; hind femur robust; length of hind femur, tibia and basitarsus 3.2 ×, 4.5 × and 6.4 × their maximum width, respectively. Metasoma. Carapace elongate-oval in dorsal view, coarsely reticulaterugose basally, longitudinal striate-rugose medially, densely reticulaterugose apically; length of carapace 2.4 × its maximum width in dorsal view (Fig. 10H); ventral cavity (as a narrow channel) open to apex, with a pair of long and large teeth; ovipositor long, slender and the ovipositor sheath 0.16 × as long as fore wing, 0.44 × as long as hind tibia (Figs 10G, 10H). Colour. Black; antennomeres blackish-brown; basal half of clypeus reddish-brown; pterostigma and veins blackish-brown; fore wing more or less infuscate; basal third of carapace yellowish; legs almost entirely yellowishbrown, hind tibia light yellow basally and black apically; ovipositor sheath brown. Male. Unknown. Biology. Unknown. Etymology. From macro (Greek for long) and odontos (Greek for teeth), because of the apex of the metasomal carapace bears a pair of long teeth., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on pages 538-540, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024

Chelonus (Arichelonus) kenosetrus sp. nov. Figs 3–4 Material examined. Holotype: female, China, Zhejiang Prov., Deqing, 27.V.1995, Junhua He, No. 965092 (ZJUH); Paratypes, 2 females, Deqing, Zhejiang province, 27.V.1995, Junhua He, Nos. 954349, 954350. Diagnosis. This new species is similar to C. (A.) olgae Kokujev, 1895, but can be separated from the latter by the following characters: ovipositor obviously extended beyond apex of metasomal carapace and covered with a semi-round and semi-hyaline lamella (the ovipositor not or weakly projecting beyond apex of metasoma in C. (A.) olgae Kokujev, 1895); metasoma entirely brown (metasoma with pair of yellow basal spots); about 22 antennomeres of female, distinctly shorter than body length (antennomeres of female about 45, equal to body length). Description. Holotype, female, length of body 4.6 mm, of fore wing 3.3 mm. Head. Antennomeres 22 (Fig. 4I); third antennomere 1.1 × longer than fourth; third, fourth and terminal antennomeres 4.0 ×, 3.5 × and 1.75 × longer than wide; head transverse, in dorsal view roundly contracted behind eyes; OOL: OD: POL=19: 15: 9; longitudinal diameter of eye 2.1 × height of temples; vertex behind ocelli and temple transversely striate; frons distinctly concave (Fig. 4G); face evenly convex, coarsely transversely striate; clypeus hardly convex and densely punctate, slightly shiny; length of malar space 1.5 × basal width of mandible (Fig. 4F); face and eyes densely setose in lateral view (Fig. 4H). Mesosoma. Length of mesosoma 1.3 × its height; mesonotum evenly convex, irregular areolate-rugose and more strongly rugose postero-medially; scutellum hardly convex, punctate medially, rugose laterally; scutellar suture comparatively deep; propodeum coarsely reticulaterugose (Fig. 4C); mesopleuron evenly convex, coarsely reticulate-rugose (Fig. 4B). Wings. Length of fore wing 2.9 × its width; length of marginal cell 1.3 × pterostigma; r: 3-SR: SRl= 9: 9: 43; SRl straight; parastigma distinctly swollen; 1-CU1: 2-CU1= 10: 27 (Fig. 4A). Legs. Hind coxa completely smooth and punctate; hind femur pretty robust; length of hind femur, tibia and basitarsus 3.1 ×, 5.2 × and 4.75 × their maximum width, respectively. Metasoma. Carapace elongate-oval, a narrow lamella dorsally of apical carapace in dorsal view, posterior part of carapace with transversely curved deepened furrow; length of carapace 2.6 × its maximum width in dorsal view (Fig. 4D); ventral cavity completely open to apex; ovipositor extended beyond apex of metasomal carapace and covered with a semi-round and semi-hyaline short lamella, ovipositor slender and the ovipositor sheath 0.11 × as long as fore wing, 0.32 × as long as hind tibia (Fig. 4E). Colour. Black; antennomeres blackish-brown; pterostigma and veins blackish-brown; fore wing more or less darkened; metasoma entirely brown; legs almost entirely yellowish-brown; ovipositor sheath brown. Variation. Body length 3.7–4.6 mm, and length of ovipositor 0.2–0.4 × of carapace length. Male. Unknown. Biology. Unknown. Etymology. From kenos (Greek for empty) and etron (Greek for belly) because its ventral cavity completely open to the apex., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on pages 531-533, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024

Chelonus (Megachelonus) irilaminatus sp. nov. Figs 5–6 Material examined. Holotype: female, China, Yunnan Prov., Mt. Gaoligong, 16-17.VII.2006, Jie Zeng, No. 200611349 (ZJUH). Paratypes: 1 female, China, Guangdong Prov., Mt. Guanyin, 15.IX.2007, Jie Zeng, No. 200807931. Diagnosis. This new species is similar to C. (M.) macros Zhang, Chen et He, but can be separated from the latter by the following characters: carapace without lamella (carapace with long and wide lamella in C. macros); apex of carapace narrowed in dorsal view(apex of carapace rounded); apex of propodeum strongly concaved with two pairs of blunt tubercles, lateral tubercles distinctly bigger and longer than medial tubercles (apex of propodeum weakly concaved with four comparably average size blunt tubercles). Description. Holotype, female, length of body 4.4 mm, of fore wing 3.0 mm. Head. Antennomeres 16 in female, average length of 3-7 th antennomeres 3.3 × their width, 8-11 th antennomeres 1.5 × longer than wide; head transverse, width of head 1.9 × its dorsal length, roundly contracted behind eyes; temple rounded posteriorly; OOL: OD: POL= 20: 6: 19, vertex evenly convex and with transverse rugae behind ocelli, temple finely and longitudinally striate- rugose, frons concave, striate-rugose around antennal sockets; longitudinal diameter of eye 2.1× height of temples (Fig. 6G); face striate-rugose, clypeus densely setose, shiny (Fig. 6F). Mesosoma. Length of mesosoma 1.4 × its height; pronotum finely reticulate-rugose; mesoscutum evenly convex, reticulate-rugose, more strongly and coarsely rugose postero-medially than anteriorly; scutellum punctate medially, rugose laterally (Fig. 6B); scutellar suture comparatively deep with 4 carinae (Fig. 6C); propodeum coarsely reticulate-rugose, strongly concave with two pairs of blunt tubercles apically, lateral tubercles distinctly bigger and longer than medial ones (Fig. 6J). Wings. Length of fore wing 3.9 × its width; length of marginal cell 2.9 × pterostigma; vein 2-Rl just present; r: 3-SR: SRl= 1: 1: 7; SRl complete and straight; parastigma not swollen; 1-CUl: 2-CUl= 1: 4; vein CUlb present (Fig.6A). Legs. Hind соха completely smooth and setose, hind femur robust, length of hind femur, tibia and basitarsus 3.7 ×, 4.3 × and 6.2 × their maximum width, respectively; length of hind tibial spurs 0.5 × hind basitarsus. Metasoma. Carapace elongate-oviform, length of metasoma 2.2 × its maximum width in dorsal view; metasomal carapace much more strongly narrowed towards apex than towards base, with two ventro-apical tooth-like protuberances, coarsely longitudinally rugulose, connected by transverse elements (Fig. 6E), carapace without lamella around ventral opening (Fig. 6D); ovipositor long, slender and curved, the ovipositor sheath 0.19 × as long as fore wing, 0.59 × as long as hind tibia (Fig. 6H). Colour. Black; 3 basal antennomeres yellowish-brown, remainder of antennomeres dark brown; palp light yellow; mandible brown but basally blaсk, pterostigma and most of veins blackish brown, veins of basal half of wing light yellow; parastigma light yellow; margination of metasomal carapace yellowish; legs yellowish-brown but hind tibia apically and base of hind coxa dark brown. Male. Unknown. Biology. Unknown. Etymology. From ir (Latin for without) and lamina (Latin for thin plate), because metasoma lacks the lateral lamella., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on pages 533-536, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024

Subgenus Megachelonus Baker, 1926 Chelonus (Megachelonus) Baker, 1926: 451. Type-species: Megachelonus bidentatus Baker, 1926. Megachelonus: De Saeger, 1948: 88 (as syn of Chelonus Panzer). Diagnosis. Head transverse (Figs 6G, 8B, 10B, 12E); antennomeres of apical half of antenna distinctly flattened and broadened; face striate to reticulate rugose, clypeus semi-elliptical (Figs 6F, 8A, 10A, 12D); eyes long elliptical or ovoid in lateral view (Figs 6I, 8G, 10G, 12F); mesosoma coarsely reticulate-rugose, transverse suture distinct; scutellum distinctly protruding above level of mesoscutum, with same sculpture as mesosoma; propodeum with a transverse carina and distinct tooth-like tubercles(Figs 6C, 6J, 8F, 10F, 12C); metasomal carapace with two ventroapical tooth-like protuberances (Figs 6E, 8C, 10C, 12H). Biology. Unknown. Distribution. Oriental. Key to species of subgenus Megachelonus 1 Carapace broadly yellowish basally....................................................................... 2 - Carapace completely black.............................................................................. 6 2 Antennomeres 28 in female; apex of metasomal carapace with two large teeth (Fig. 10H)..... C. (M.) macrodontus sp. nov - Antennomeres 16 in female; metasomal carapace with two comparatively small ventro-apical tooth-like protuberances (Figs 6E, 8C, 10C)......................................................................................... 3 3 Carapace without lamella (Fig. 6D)................................................ C. (M.) irilaminatus sp. nov. - Carapace with lamella (Figs 8D, 12D)..................................................................... 4 4 Lamella of carapace ventrally long and wide (Figs 11, 12D)........................ C. (M.) macros Zhang, Chen & He - Lamella of carapace ventrally short and narrow (Figs 7, 8D).................... C. (M.) longqiensis Zhang, Chen & He 5 Ventral opening of metasomal carapace reaching apex of carapace.............................................. 6 - Ventral opening remains far away from apex of metasomal carapace; apex of metasomal carapace distinctly swollen in lateral view............................................................................................... 8 6 Base of metasomal carapace entirely yellowish................................... C. (M.) bicoloricomis Baker, 1926 - Base of metasomal carapace black, without spots............................................................ 7 7 Metasomal carapace with a strongly transverse depression dorsally; body length 5.5 mm.. C. (M.) rectangularis Baker, 1926 - Metasomal carapace without dorsal depression body; length 6.0 mm................. C. (M.) megacephalus Baker, 1926 8 Antennomeres dark brown; hind femur entirely dark brown or blackish; body length 3.5 mm..................................................................................................... C. (M.) uniformis Baker, 1926 - Basal half of antennomeres pale brown and apical half dark brown; hind femur only apically blackish; body length 4 mm........................................................................... C. (M.) bicoloricomis Baker, 1926, Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on page 533, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024, {"references":["Baker, C. F. (1926) Braconidae-Cheloninae of the Philippines, Malaya, and Australia. I. Chelonini (except Chelonus). The Philippine Journal of Science, 31 (4), 451 - 499."]}

Chelonus (Scabrichelonus) nigrifemoratus sp. nov. Figs 13–16 Material examined. Holotype:female, China, Hainan Prov., Mt.Wuzhi, 16–20.V.2007, Liqiong Wen,No. 200705133 (ZJUH). Paratype, 1 male, China, Hainan Prov., Mt. Wuzhi, 17. V.2007, Bin Xiao, No. 200705132 (ZJUH). Diagnosis. This new species is similar to C. (S.) sinensis He, Chen & van Achterberg, but can be separated from the latter by the following characters: hind coxa and hind femur black in both sexes (hind coxa apically and hind femur basally more or less yellowish-brown in C. (S.) sinensis); apical opening of male carapace is relatively large and close to lateral margins of carapace (apical opening of male carapace relatively small and remaining removed from lateral margins of carapace). Description. Holotype, female, length of body 4.5 mm, of fore wing 3.4 mm. Head. Antennomeres more than 8 (broken in holotype), scapus robust, length of third antennomere 1.2 × fourth, length of third and fourth antennomeres 3.8 × and 3.3 × their width, respectively; in dorsal view its maximum width and width between eyes 1.2 × and 1.8 × its median length, respectively; temple strongly swollen posteriorly; frons slightly concave; temple and vertex distinctly transversely rugose; OOL: OD: POL= 21: 9: 11; longitudinal diameter of eye 0.8 × height of temples (Fig. 14G); face as long as its width, evenly convex; clypeus flat, sparsely and finely punctate; length of malar space 0.5 × basal width of mandible (Fig. 14F); face and eyes with densely setose in lateral view (Fig. 14I). Mesosoma. Length of mesosoma 2.2 × its height; pronotum densely irregularly reticulate; mesoscutum evenly convex, irregularly finely reticulate-rugose, more strongly and coarsely rugose postero-medially; scutellum finely punctate-rugose; scutellar sulcus narrow, with 7 carinae; propodeum densely reticulate-rugose (Fig. 14C); mesopleuron evenly convex, densely finely reticulate-rugose, more strongly and coarsely so antero-dorsally; prepectal carina just extending to level of postero-ventral margin of pronotum; metapleuron coarsely rugose (Fig. 14B). Wings. Fore wing (Fig. 14A): length of fore wing 3.0 × its width; length of vein 1-R1 0.9 × length of pterostigma; vein 2-R1 just present; r: 3-SR: SRl =11: 19: 70; parastigma slightly swollen; 1-CU1: 2-CU1 = 13:46; vein CUlb present. Legs. Hind coxa shallowly punctate-rugose; hind femur robust, its length 3.1 × its width; hind tibia rather thick apically, its length 1.2 × length of tarsus; hind tarsus narrowed towards apex; length of basitarsus 0.4 × length of tarsus, 2.0 × length of inner tibial spur (Fig. 14E). Metasoma. Length of metasoma 2.8 × its maximum width in dorsal view; metasomal carapace longitudinally densely reticulate-rugose (Fig. 14H); ovipositor is hidden under the ventral opening of metasomal carapace, the ovipositor sheath 0.12 × as long as fore wing, 0.31 × as long as hind tibia (Fig. 14D). Colour. Black; antennomeres blackish-brown; palp yellowish brown; scapus (except for base) and mandibles basally reddish-brown; pterostigma and most of veins blackish-brown; basal third of carapace yellowish; legs almost yellowish-brown totally; hind соха blасk but apically yellowish-brown; hind femur blackish-brown; basal hind tibia light yellow and apically brown. male. Antennomeres 30; similar to female, but differs in entirely black carapace (Figs 16D, 16E); carapace with a small apical opening which transverse elongate-oval in posterior view, 2.3 × wider than high, and height of opening 0.5 × ventral height of carapace (Fig. 16K). Biology. Unknown Etymology. Named after its blackish-brown hind femur: niger is Latin for black, femora is Latin for femur., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on pages 544-546, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024

Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu, Chen, Xue-Xin (2022): A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China. Zootaxa 5165 (4): 527-547, DOI: https://doi.org/10.11646/zootaxa.5165.4.5

Subgenus Scabrichelonus He, Chen & van Achterberg, 1997 Chelonus ( Scabrichelonus) He, Chen & van Achterberg, 1997: 53. Type-species: Chelonus (Scabrichelonus) sinensis He, Chen & van Achterberg, 1997 (Monotype). Microchelonus (Scabrichelonus): Tobias, 2000: 481. Scabrichelonus: Chen & Ji, 2003: 111 (as syn. of Microchelonus Szépligeti). Diagnosis. Head triangular in anterior view (Figs 14F, 16G), trapezoid in dorsal view (Figs 14G, 16H); temple distinctly swollen, with 3 strong ridge-like transverse carinae; face evenly convex, distinctly transversely rugose, with a longitudinal carina dorso-medially, mandibles robust, not twisted apically, upper tooth longer than lower tooth; (Figs 14F, 16G); occipital carina thin, arched dorso-medially, ventrally connected to hypostomal carina above base of mandible (Figs 14G, 16H); mesoscutum evenly convex, transverse suture distinct; notauli superficially impressed; propodeum short, near posterior margin of propodeum (Figs 14C, 16C); metapleuron coarsely rugose, without metapleural flange; apical transverse carina of (Figs 14B, 16B) length of metasoma equal to combined length of head and mesosoma; metasomal carapace elongate, without sutures, only slightly widened apically; ventral metasomal opening close to posterior margin of metasomal carapace; metasoma of male with an apical opening. Biology. Unknown. Distribution. Oriental (China)., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China, pp. 527-547 in Zootaxa 5165 (4) on pages 540-542, DOI: 10.11646/zootaxa.5165.4.5, http://zenodo.org/record/6854024, {"references":["He, J. H., Chen, X. X. & van Achterberg, C. (1997). Scabrichelonus, a new subgenus of Chelonus Panzer (Hymenoptera: Braconidae: Cheloninae) from China. Zoologische Mededelingen, Leiden, 71, 1 - 18."]}

Chelonus (Mirachelonus) miraventris sp. nov. Material examined. Holotype: ♀, China, Yunnan Prov., Yingjiang, 1.X.2004, Jinxian Liu, No. 20059097 (ZJUH). Paratypes: (ZJUH): 1♀, China, Yunnan Prov., Yingjiang, 23. VI.2005, Jinxian Liu, No. 200609601; 2♀♀, China, Hainan, Mt. Wuzhi, 16-18. V.2007, Jie Zeng, Nos. 200807590, 200705120; 1♂, China, Hainan, Mt. Wuzhi, 6- 18. V.2007, Jie Zeng, No. 200807647; 1♀, China, Hainan, Mt. Wuzhi, 16-20. V.2007, Liqiong Wen, No. 200705135; 1♀, China, Hainan, Yinggeling, 25. V.2007, Jinxian Liu, No. 200702332; 2♂♂, China, Taiwan, Kaohsiung, Mt. Wugong, 31. V.2011, Pu Tang, Nos. 201104855, 201104842; 15♂♂, China, Taiwan, Kenting National Park, 31. V.2011, Pu Tang, Nos. 201104822, 201104823, 201104830, 201104833, 201104906, 201104908, 201104910, 201104913, 201104916, 201104918, 201104919, 201104922, 201104924, 201104928, 201104935. Description. Holotype, female, length of body 3.83 mm, of fore wing 3 mm. Head. Antennomeres 16, distinctly shorter than body, average length of 3 rd ���7 th segments 3.2 �� their width, 8 th ���11 th segments 1.6 �� longer than wide; head transverse, width of head 2.2 �� its dorsal length, roundly contracted behind eyes, length of eyes 15.2 �� temple in dorsal view, OOL: OD: POL= 16: 5: 14, longitudinal diameter of eye 2.3 �� height of gena; vertex evenly convex and with transverse rugae behind ocelli, temple finely and longitudinally striate-rugose, frons concave, striate-rugose around antennal sockets (Fig. 2G); length maxillary palp 0.5 �� height of head in anterior view, face striate-rugose, clypeus shiny, finely punctate, length of malar space 2.2 �� basal width of mandible (Fig. 2F). Mesosoma. Length of mesosoma 1.3 �� its height; pronotum finely reticulate-rugose; mesoscutum evenly convex, reticulate-rugose, more strongly and coarsely rugose postero-medially; scutellum punctate medially, rugose laterally, more or less protruding above level of mesoscutum; scutellar suture comparatively deep and with 4 carinae; mesopleuron coarsely reticulate-rugose; propodeum coarsely reticulate-rugose with transversal carina and four blunt tubercles (Fig. 2B). Wings. Length of fore wing 2.5 �� its width; length of marginal cell 0.9 �� pterostigma; vein 2-Rl short (Fig. 1); r: 3-SR: SRl = 1: 1: 6; SRl complete and straight; parastigma not or weakly swollen; 1-CUl: 2-CUl = 3: 7; vein CUlb short (Fig. 2A). Legs. Hind соха completely smooth and setose; hind femur robust, length of hind femur, tibia and basitarsus 3.2, 4.5 and 6.4 �� their maximum width, respectively; length of hind tibial spurs 0.5 �� hind basitarsus (Fig.2I). Metasoma. Length of carapace 2.5 �� its maximum width in dorsal view (Fig. 2E); third tergite of carapace almost straightly narrowed, longitudinally densely reticulate-rugose; apex of carapace truncate and narrow in dorsal view, with a wide ventral lamella in lateral view (Fig. 2D); apex of carapace deeply incurved, about one fourth of total length of carapace. Colour. Black; six basal antennomeres yellowish-brown, remainder of antennomeres dark brown; palpi light yellow; mandible brown except black basally; pterostigma and most of veins blackish brown, veins of basal half of wing light yellow; parastigma light yellow; carapace yellowish in basal 1/3; legs mainly yellowish-brown except dark brown apex of hind tibia and all telotarsi black. Male. Length of body 3.0 mm, similar to female, but differs in having 23���26 antennomeres; length of metasoma 2.7 �� its maximum width in dorsal view (Fig. 3E); carapace with a small apical opening filled by white setae (Fig. 3J). Biology. Unknown. Distribution. Oriental China (Yunnan, Taiwan, Hainan). Etymology. Named after its special metasoma: mirabilis is Latin for wonderful and strange, venter is Latin for belly., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A new subgenus of Chelonus Panzer, 1806 (Braconidae: Cheloninae) from China, pp. 288-294 in Zootaxa 5115 (2) on pages 292-294, DOI: 10.11646/zootaxa.5115.2.8, http://zenodo.org/record/6352481

Key to subgenera of Chelonus Panzer 1 Temple strongly swollen and with three distinct ridge-like rugae........ Scabrichelonus He, Chen & van Acheterberg, 1997 - Temple hardly or not swollen and without distinct ridge-like rugae.............................................. 2 2 Metasomal carapace open ventro-apically (thus ventral opening reaching apex of carapace) and protruding apically, at least as long as combined length of head and metasoma........................................ Arichelonus Viereck, 1913 - Metasomal carapace more or less closed apically............................................................ 3 3 Metasomal carapace with one or two apical teeth............................................................ 4 - Metasomal carapace rounded dorso-apically (but minute apical tooth often present in Stylochelonus)................... 5 4 Metasomal carapace with a dorso-apical spine-like protuberance; vein SR1 of fore wing sometimes incomplete; carapace with narrow straight lamella ventrally; [apical half of antenna widened]............................ Areselonus Braet, 1999 - Metasomal carapace with two ventro-apical tooth-like protuberances; vein SR1 of fore wing complete; carapace with undulating lamella ventrally; [apical half of antenna distinctly widened and antennomeres longer than wide; antenna of female with 16 antennomeres]................................................................... Megachelonus Baker, 1926 5 Head cubical; in dorsal view interocular width slightly more than width of eye and distinctly convexly extended beyond eyes; posterior face of propodeum not separated from anterior face by a transverse rim.............. Cubochelonus Baker, 1926 - Head distinctly transverse; in dorsal view interocular width always distinctly greater than width of eye and at most slightly extended beyond eyes; posterior face of propodeum always separated from anterior face by a transverse rim............. 6 6 Third���fifth antennomeres of female 6.0���7.0 �� longer than wide; frons without distinct curved lateral carina; scutellum more or less protuberant above level of mesoscutum; vein 1-M usually comparatively close to vein C+SC+R); [female with 16 antennomeres]............................................................ Baculonus Braet & van Achterberg, 2001 - Third���fifth antennomeres of female distinctly less than 5.0 �� longer than wide, if about 5.0 �� then frons with distinct curved lateral carina; scutellum not protruding above level of mesoscutum; vein 1-M usually more removed from vein C+SC+R....................................................................................................... 7 7 Frons deeply excavated posteriorly and smooth; carapace very broadly truncate apically and ventral opening slightly longer than half length of carapace, opening surrounded by a lamelliform carina................... Carinichelonus Tobias, 2000 - Frons shallowly excavated posteriorly, and more or less sculptured; carapace more or less narrowed apically and ventral opening usually distinctly longer than half length of carapace, and opening usually without a lamelliform carina.............. 8 8 Third tergite of metasomal carapace almost straightly narrowed posteriorly in dorsal view (Fig. 2E); apex of metasomal carapace of both sexes truncate and narrow in dorsal view (Figs 2E, 4E) and oblique in lateral view (Figs 2D, 4D); carapace of male deeply incurved (about one fourth of total length of carapace) in lateral view (Fig. 4D)......... Mirachelonus subgen. nov. - Third tergite of metasomal carapace hardly or not narrowed posteriorly in dorsal view; apex of metasomal carapace round posteriorly or nearly so in dorsal and lateral view; carapace of male gradually rounded in lateral view.................. 9 9 Antennomeres of female 16; carapace of male with apical opening.................................................................................................................... Microchelonus Sz��pligeti, 1908 s. str. Notes. Tobias (2011) introduced a new monotypic subgenus Rasnichelonus for Microchelonus elongatus Papp, 1971, because of its elongate body, head expanding behind eyes and shallow apical aperture of male metasoma. In our opinion the differences with other species are too variable to deserve a subgeneric status. - Antennomeres of female more than 16; carapace of male variable, usually without distinct opening................... 10 10 Antennomeres of female 30���32; carapace of male with deep apical opening; body at least 5.0 mm long................................................................................................ Parachelonus Tobias, 1995 - Antennomeres of both sexes 17���26; carapace of male variable, often closed; length of body variable.................. 11 11 Metasomal carapace of male with small apical opening, which may be filled by point-like sclerite; carapace of female usually with an apical tooth; [mesosoma at least 1.5 �� longer than high]........................... Stylochelonus Hell��n, 1958 - Metasomal carapace of male closed apically and without opening or point-like sclerite posteriorly; carapace of female without apical tooth.................................................................... Chelonus Panzer, 1806 s. str., Published as part of Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu & Chen, Xue-Xin, 2022, A new subgenus of Chelonus Panzer, 1806 (Braconidae: Cheloninae) from China, pp. 288-294 in Zootaxa 5115 (2) on page 289, DOI: 10.11646/zootaxa.5115.2.8, http://zenodo.org/record/6352481, {"references":["Tobias, V. I. (2000) Cheloninae. In: Ler, P. A. (Ed.), Key to the insects of Russian Far East. Vol. IV. Neuropteroidea, Mecoptera, Hymenoptera. Pt 4. Opredelitel nasekomykh Dalnego Vostoka Rossii. T. IV. Setchatokryloobraznye, skorpionnitsy, pereponchatokrylye. Ch. 4. Dalnauka, Vladivostok, pp. 426 - 571."]}