Tethystrombus coronatus (Defrance, 1827) Plate 4 A * Strombus coronatus Defrance 1827: 124. Strombus italicus Duclos — Mayer 1864: 73, pl. 7, fig. 54. Strombus coronatus Defr. —Pereira da Costa 1867: 137, pl. 17, fig. 3. Strombus coronatus Defrance — Fontannes 1879: 151, pl. 9, fig. 1. Strombus coronatus (Defr.) — Sacco 1893: 7, pl. 1, figs. 19-27. Strombus coronatus Defrance, 1827 — Palla 1967: 959, pl. 72, fig. 2. Strombus (Strombus) coronatus (Defrance, 1827) — Malatesta 1974: 219, pl. 17, figs. 1-7. Strombus coronatus Defrance — Fekih 1975: 111, pl. 33, fig. 1. Strombus (S.) coronatus Defrance — Pavia 1976a, pl. 5, figs. 1-4. Strombus coronatus Defrance, 1827 — Meco 1977: 56, pl. 14, fig. 2, pl. 15, fig. 2, pl. 16, figs. 1-2, pl. 17, figs. 1-2, pl. 18, figs. 1-2, pl. 19, fig. 1, pl. 20, fig. 1, pl. 21, fig. 1, pl. 22, fig. 1, pl. 23, figs. 1-3, pl. 24, fig. 1, pl. 25, figs. 1-2, pl. 26, fig. 1, pl. 27, figs. 1-2. Strombus coronatus Defrance, 1827 — Martinell 1979: 123, pl. 3, figs. 5-6. Strombus coronatus (Defrance, 1827) — Ferrero Mortara et al. 1984: 138, pl. 21, figs. 6-7. Strombus coronatus Defrance, 1827 — Cavallo & Repetto 1992: 58, fig. 101. Strombus coronatus Defrance, 1827 — Solsona 1998: 169, pl. 8, figs. 1-6. Strombus (Strombus) coronatus Defrance, 1827 — Landau et al. 2004a: 63, pl. 14, fig. 6. Strombus coronatus (Defrance, 1827) [sic]—Chirli 2008: 6, pl. 2, figs. 1-6. Strombus coronatus Defrance, 1827 — Chirli & Richard 2008: 24, pl. 3, fig. 7. Strombus coronatus Defrance, 1827 — Sosso & Dell’Angelo 2010: 23, 33, unumbered figure second row left. Strombus coronatus (Defrance, 1827) [sic]—Brunetti 2014: 30, unumbered fig. Persististrombus coronatus (Defrance, 1827) — Ávila et al. 2016b: 915, fig. 2A-H. Persististrombus coronatus (Defrance, 1827) —Brunetti & Vecchi 2016: 62, pl. 1, fig. A. Persististrombus coronatus (Defrance, 1827) — Brunetti & Cresti 2018: 50, fig. 130. non Strombus coronatus Defr. —Ĥrnes 1853: 187, pl. 17, fig. 1 [Persististrombus inflexus (Eichwald, 1830)]. non Strombus coronatus Defr. —Hoernes & Auinger (partim) 1884: 163, pl. 18, figs. 4-5, pl. 19. fig. 1 [Persististrombus inflexus (Eichwald, 1830)]. non Strombus coronatus Defr. —Hoernes & Auinger (partim) 1884: 163, pl. 18, figs. 1-3 [Persististrombus lapugyensis (Sacco, 1893)]. non Strombus coronatus Defrance — Strausz 1966: 222, figs. 102-103 [Persististrombus inflexus (Eichwald, 1830)]. non Strombus coronatus Defrance — Schultz 1998: 60, pl. 23, fig. 6 [Persististrombus inflexus (Eichwald, 1830)]. Santa Maria material examined. Maximum height 119.1 mm, width 107.2 mm. Figueiral: CIADP-F 32 (1) Casa dos Fósseis Museum, Santa Maria (collection Dalberto Pombo); DBUA-F 1011 (2), DBUA-F 1236 (1), DBUA-F 533 (1), Malbusca; DBUA-F 928- A (1), Malbusca, Santa Maria Island, Azores, Touril Complex, Lower Pliocene. Discussion. Dekkers (2008) erected the genus Tethystrombus with the West African Strombus latus Gmelin, 1791 as type species. He included in his new genus the European fossil Strombus bubonius Lamarck, 1822 and S. coronatus Defrance, 1827. The ongoing heated discussion as to the phylogeny of this group (i.e., Dekkers, 2008, vs. Kronenberg & Vermeij 2002) will not be entered into here. The presence of Tethystrombus coronatus (Defrance, 1827) in the Santa Maria assemblages and its biogeographic, palaeoecological and palaeoclimatological implications were fully discussed in Ávila et al. (2016b). Distribution. Middle Miocene: Atlantic, Aquitaine Basin (Cossmann & Peyrot 1924), Loire Basin (Glibert 1949). Upper Miocene: Atlantic, Portugal (Pereira da Costa 1867); Proto-Mediterranean, Italy (Sacco 1893). Lower Pliocene: Atlantic, Santa Maria Island, Azores (Ávila et al. 2015b, 2016b, 2018a, 2020a, 2022; Rebelo et al. 2016; Uchman et al. 2017, 2020; Dávid et al. 2021), Madeira and Porto Santo (Mayer 1864), Canary Islands (Meco 1977), Morocco (Lecointre 1952); western Mediterranean, NE Spain, (Martinell 1979; Martinell & Marquina 1981), France (Fontannes 1879; Chirli & Richard 2008); central Mediterranean, Italy (Sacco 1893; Pavia 1976a; Cavallo & Repetto 1992; Chirli 2008; Sosso & Dell’Angelo 2010; Brunetti & Cresti 2018). Upper Pliocene: central Mediterranean, Italy (Brunetti & Vecchi 2016), western Mediterranean, Estepona Basin (Landau et al. 2004a)., Published as part of Sacchetti, Claudia, Landau, Bernard & Ávila, Sérgio P., 2023, The Lower Pliocene marine gastropods of Santa Maria Island, Azores: Taxonomy and palaeobiogeographic implications, pp. 1-150 in Zootaxa 5295 (1) on pages 56-57, DOI: 10.11646/zootaxa.5295.1.1, http://zenodo.org/record/7965273, {"references":["Mayer, K. (1864). Die Tertiar-fauna der Azoren und Madeiren. Zurich, published by the author. Z ¸ rich: vi + 107 pp. + 7 pls.","Fontannes, F. (1879 - 1880) Les inverte´bre´s du bassin tertiaire du Sud-Est de la France. Les mollusques plioce`nes de la Valle´edu Rho ˆ ne et du Roussillon. 1. Gaste´ropodes des formations marines et saumatres. Georg, Lyon & F. Savy, Paris, VIII + 276 pp., 12 pls. [pp. 1 - 76 published in 1879, remainder in 1880] https: // doi. org / 10.5962 / bhl. title. 13209","Sacco, F. (1893). I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte XIV. (Strombidae, Terebellidae, Chenopidae ed Haliidae). Carlo Clausen, Torino, 40 pp., 2 pls.","Palla, P. (1967) Gasteropodi pliocenici della Bassa Val d'Elsa (Toscana Occidentale). Rivista Italiana di Paleontologia e Stratigrafia, 73, 931 - 1020.","Malatesta, A. (1974) Malacofauna pliocenica Umbra. Memorie per Servire alla Carta Geologica d'Italia, 13, 1 - 498.","Fekih, M. (1975) Paleoecologie du Plioce`ne marin au nord de la Tunisie. Annales des Mines et de la Ge´ologie, 27, 1 - 195.","Pavia, G. (1976 a [1975]) I molluschi del Pliocene inferiore di Monteu Roero (Alba, Italia NW). Bollettino della Societa Paleontologica Italiana, 14, 99 - 175.","Meco, J. (1977) Los Strombus neogenos y cuaternarios del Atlantico euroafricano (taxonomia, biostratigrafia y paleoecologia). Paleontologia de Canarias vol. 1. Excmo. Cabildo Insular de Gran Canaria, Gran Canaria, 142 pp.","Martinell, J. (1979) Mesogastropoda del Plioceno del Emporda (Girona). 1. Descriptiva y sistematica. Studia Geologica Salmanticensia, 15, 85 - 165.","Ferrero Mortara, E. L., Montefameglio, L., Novelli, M., Opesso, G., Pavia, G. & Tampieri, R. (1984) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. 2. Museo Regionale di Scienze Naturali, Cataloghi 7, 1 - 484.","Cavallo, O. & Repetto, G. (1992) Conchiglie fossili del Roero. Atlante iconografico. Associazione Naturalistica Piemontese Memorie (Associazione Amici del Museo ' Federico Eusebio'), 2, 1 - 251.","Solsona, M. (1998) Paleobiologia dels mesogastero`podes del Plioce`del Mediterrani nord-occidental. Unpublished Tese de Doutoramento, Departamento d'Estratigrafia i Paleontologia, Universitat de Barcelona, Barcelona, 540 pp.","Landau, B. M, Marquet, R. & Grigis, M. (2004 a) The early Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. 2. Orthogastropoda, Neotaenioglossa. Palaeontos, 4, 1 - 108.","Chirli, C. & Richard, C. (2008) Les mollusques plaisanciens de la Co ˆ te d'Azur. Chirli, C., Tavarnelle, 128 pp., 25 pls.","Sosso, M. & Dell'Angelo, B. (2010) I fossili del Rio Torsero. Editing Marginalia, Cartotectonica Beusi srl, Prato, 95 pp.","Avila, S. P., Melo, C., Berning, B., Cordeiro, R., Landau, B. & Silva, C. M. da (2016 b) Persististrombus coronatus (Mollusca: Strombidae) in the lower Pliocene of Santa Maria Island (Azores, NE Atlantic): paleoecology, paleoclimatology and paleobiogeographic implications. Palaeogeography, Palaeoclimatology, Palaeoecology, 441, 912 - 923. https: // doi. org / 10.1016 / j. palaeo. 2015.10.043","Brunetti, M. M. & Cresti, M. (2018) I fossili di Orciano Pisano. Atlante iconografico. Edizioni Danaus, Palermo, 232 pp.","Eichwald, E. (1830) Naturhistorische Skizze von Lithauen, Volhynien und Podolien in geognostisch-mineralogischer, botanischer und zoologischer Hinsicht. Eichwald, Wilna, 256 pp.","Strausz, L. (1966) Die Mioza ¨ n-Mediterranen Gastropoden ungarns. Akade´miai Kiado´, Budapest, 692 pp.","Schultz, O. (1998) Tertia ¨ rfossilien o ¨ sterreichs, Wirbellose, niedere Wirbeltiere und marine Sa ¨ ugetiere; scho ¨ ne, interessante, ha ¨ ufige und wichtige Makrofossilien aus den Besta ¨ nden des Naturhistorischen Museums Wien und Privatsammlungen; eine Bilddokumentation. Golschneck-Verlag, Wien, 159 pp.","Dekkers, A. M. (2008) Revision of the family Strombidae (Gastropoda) on the supraspecific level. Part One. De Kreukel, 44 (3), 35 - 64.","Gmelin, J. F. (1791) Caroli a Linnei systema natura per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, disserentis, synonymis, locis etc. Editio decima tertia, aucta, reformata, cura J. F. Gmelin. 1 (6). Vermes testacea. G. E. Beer, Lipsiae, pp. 3021 - 4120.","Lamarck, J. B. P. A. de M. (1822) Histoire naturelle des animaux sans verte`bres, pre´sentant des characte`res ge´ne´raux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales espe`ces qui s'y rapportent, pre´ce´de´e d'une introduction offrant la de´termination des caracte`res essentiels de l'animal, sa distinction du ve´ge´tal et des autres corps naturels; enfin, l'exposition des principes fondamentaux de la zoologie, 7. Lamarck, Paris, 711 pp.","Kronenberg, G. C. & Vermeij, G. J. (2002) Terestrombus and Tridentarius, new genera of Indo-Pacific Strombidae (Gastropoda), with comments on included taxa and on shell characters in Strombidae. Vita Malacologica, 1, 49 - 54.","Glibert, M. (1949) Gastropodes du Miocene moyen du Bassin de la Loire. 1. Memoires de l'Institut Royal des Sciences Naturelles de Belgique, 2 (30), 1 - 240.","Avila, S. P., Ramalho, R., Habermann, J., Quartau, R., Kroh, A., Berning, B., Johnson, M., Kirby, M., Zanon, V., Titschack, J., Goss, A., Rebelo, A. C., Melo, C., Madeira, P., Cordeiro, R., Meireles, R., Bagaco, L., Hipolito, A., Uchman, A., Da Silva, C. M., Cach \" o, M. & Madeira, J. (2015 b) Palaeoecology, taphonomy, and preservation of a lower Pliocene shell bed (coquina) from a volcanic oceanic island (Santa Maria Island, Azores, NE Atlantic Ocean). Palaeogeography, Palaeoclimatology, Palaeoecology, 430, 57 - 73. https: // doi. org / 10.1016 / j. palaeo. 2015.04.015","Uchman, A., Quintino, V., Rodrigues, A. M., Johnson, M. E., Melo, C., Cordeiro, R., Ramalho, R. & Avila, S. P. (2017) The trace fossil Diopatrichnus santamariensis nov. sp. - a shell armored tube from Pliocene sediments of Santa Maria Island, Azores (NE Atlantic Ocean). Geobios, 50 (5 - 6), 459 - 469. https: // doi. org / 10.1016 / j. geobios. 2017.09.002","Uchman, A., Johnson, M. E., Ramalho, R., Quartau, R., Berning, B., Hipolito, A., Melo, C., Rebelo, A. C., Cordeiro, R. & Avila, S. P. (2020) Neogene marine sediments and biota encapsulated between lava flows on Santa Maria Island (Azores, northeast Atlantic): An interplay between sedimentary, erosional, and volcanic processes. Sedimentology, 67, 3595 - 3618. https: // doi. org / 10.1111 / sed. 12763","David, A., Uchman, A., Ramalho, R. S., Madeira, J., Melo, C., Madeira, P., Rebelo, A. C., Berning, B., Johnson, M. E. & Avila, S. P. (2021) Diverse bioerosion structures in lower Pliocene deposits from a volcanic oceanic island: Baia de Nossa Senhora section, Santa Maria Island, Azores (Central Atlantic Ocean). Palaeogeography, Palaeoclimatology, Palaeoecology, 569, 110 - 284. https: // doi. org / 10.1016 / j. palaeo. 2021.110284","Lecointre, G. (1952) Recherches sur le Ne´oge`ne et le Quaternaire marins de la co ˆ te atlantique du Maroc, 2, Pale´ontologie. Notes et Me´moires du Service Ge´ologique du Maroc, 99, 5 - 170.","Martinell, J. & Marquina, J. M. (1981) Malacofauna Pliocenica de St. Vicenc ¸ dels Horts (Baix Llobregat, Barcelona). Iberus, 1, 9 - 22."]}