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Unsettled, dubious Oecetis tripunctata taxa are studied applying the principles and practices of fine phenomics relying on the ventral profile of the gonopods and the lateral profile of the phallic organ. The distribution of Oecetis tripunctata (Fabricius, 1793) is restricted to the Palaearctic Faunal Region. 14 new species were described from the Oriental Faunal Region: Oecetis anhua s. nov., O. baneswara sp. nov., O. baoloca sp. nov., O. bhubaneswa sp. nov., O. dagna sp. nov., O. dhaulia sp. nov., O. gujara sp. nov., O. halonga sp. nov., O. hoabinha sp. nov., O. kamba sp. nov., O. orissa sp. nov., O. peraka sp. nov., O. prenna sp. nov., O. sabarma sp. nov. and one species Oecetis lata Johanson, Pham, Malm & Sjöberg, 2020 was recorded from Vietnam. 2 new species were described from Australasian Faunal Region: O. kokopa sp. nov., O. parom sp. nov. as new records of the Oecetis tripunctata species group. 10 new species were described from the Afrotropical Faunal Region, including mailand Africa and Madagascar: O. anka sp. nov., O. banda sp. nov., O. bua sp. nov., O. conga sp. nov., O. congana sp. nov., O. ghana sp. nov., O. manta sp. nov., O. maroa sp. nov., O. volta sp. nov., O. zoa sp. nov., and one species Oecetis kagerana Kimmins, 1956 was recorded from Ghana, as well as O. maculipennis Ulmer, 1922 was placed to the Oecetis tripunctata species group. This replacement has necessitated revising and renaming the Oecetis maculipennis species group describing 2 new species: O. kimminsiana sp. nov. and O. nkwanta sp. nov. Oecetis maculipennis species group was renamed as Oecetis kimminsiana species group.

We have recognised significant incongruences among the most commonly used taxonomic characters in the European species of Wormaldia genus of the Philopotamidae caddisfly family. During taxonomical analysis and ranking procedures we have recorded incongruent, discorcordant characters also in the taxa in Rhyacophilidae, Hydropsychidae and Limnephilidae caddisfly families. Based on theoretical background we concluded that taxa of examined caddisflies and probably all living creatures are chimeric entities composed of components of different origin. Genomes and phenomes are tree-like on the surface but more reticulated in the deep. We understand chimerism with universal consequences, expanding well beyond the evolutionary tree-thinking of reductionism and determinism. Taxa are chimeric or at least chimerical in a stochastic universe under the permanent fluxes of the external and internal impacts created by intercourses between entropy and energy gradients. We have surveyed how to create and correct synonymies in the splitter/lumper perspectives along the principles of compositional and specification hierarchies understood as quantitative variability of non-adaptive neutral and qualitative stability of adaptive, non-neutral traits. We outlined how the apophantic (declaratory) hybris creates synonymies and underestimates biodiversity. After redrawing the diverging genitalic structures, particularly the speciation traits we have reinstated species status of eight taxa: W. trifida Andersen, 1983 stat.restit, stat. nov., W. albanica Oláh, 2010 stat. restit., W. bulgarica Novak, 1971 stat. nov., W. daga Oláh, 2014 stat. restit., W. graeca Oláh, 2014 stat. restit., W. busa Oláh, 2014 stat. restit., W. homora Oláh, 2014 stat. restit. W. nielseni Moretti, 1981 stat. nov. Character selection and lineage sorting procedures established the following species groups, species complexes and species clades in the European species of Wormaldia: W. occipitalis species group: W. occipitalis species complex; W. charalambi species group; W. copiosa species group; W. triangulifera species group: W. bulgarica species complex, W. khourmai species complex, W. subnigra species complex: W. asterusia species clade, W. subnigra species clade, W. vercorsica species clade; W. triangulifera species complex, W. variegate species complex. Unplaced species: W. ambigua, W. algirica, W. sarda. In this revision we have described fourteen new species: W. longiseta, W. carpathica, W. kurta, W. parba, W. foslana, W. kumanskii, W. libohova, W. silva, W. gorba, W. kera, W. rona, W. sima, W. granada, W. telva

The taxonomic potential of speciation traits, the adaptive structural components of the phallic head that build the early stages of reproductive barriers was applied in our first revision of the Aethaloptera genus with unsettled obscured taxonomy. The present paper complements the first revision and is based on the examination of the historical Aethaloptera collection of the Natural History Museum, London. The difficulty of presenting the tiny, subtle, but stable divergences in the speciation traits is discussed. The presented graphical copy of phenomic reality delineates species reliably if the matrices of the drawings from various populations, clearly visualizes empirically, without geometric morphometrics, that the detected shape divergences are larger than the combined variation ranges of the artefacts and of the infra-populations. Based upon the lateral profiles of the phallic head two new species complexes: A. dispar and A. maxima and four new species have been described: A. nulta sp. nov., A. pipa sp. nov., A. feltora sp. nov. and A. pricei sp. nov.

Taxonomy of the “widely distributed and highly varying species” in the small Aethaloptera genus was questioned long ago. Relying on the macro morphology, detectable by the routine resolution of the stereomicroscope, these species look very similar. However, applying the higher resolution of compound microscope with focusing on profile stability, significant shape divergences have been recognised in the fine structure of the speciation trait that is in the much specialised head structure of the phallic organ. The ancestral prototype of caddisfly genitalia is highly modified in the Hydropsychidae family. The hydropsychid prototype is further organised in the macronematine Aethaloptera genus: the abbreviated primordial genital structures of Hydropsychidae family are retracted into the invaginated tip of the phallotheca with apomorphic organisation of complexity by reduction or simplification, a complexity integrated by incremental subtraction. Examining the fine phenomics of the phallic head we have tripled the species number in this small genus and delineated altogether 12 species. We expect many more species to be collected and distinguished in this neglected genus by the fine structure of this speciation trait. Here we established two new species groups: Aethaloptera dispar new species group and A. meyi new species group and described five new species: Aethaloptera felalla, A. karima, A. malickyi, A. meyi and A. wellsi spp. nov.

The subspecies of the biological species concept with incomplete reproductive isolation versus the incipient sibling species of the phylogenetic species concept with permeable reproductive barrier are still applied side by side in the everyday practice of taxonomy. Both terms refer to the same organisms diverged mostly in allopatry with various stages of repro-ductive isolation. Question remained: how human ranks these entities organised by nature? The reliable ranking of living hierarchies is retarded and even obscured by the suppressed state of taxonomy. Disappointing scenario: the science of biodiversity is stuck in century old macromorphologies without innovation of fine phenomics and without exploring its high-tech and high-throughput potential. The empirical science of taxonomy is “modernised” by the neutral DNA marker industry diverting the epistemological focus from empirical to virtual. Virtuality of noumenon is used to camouflage the phenomenon of the adverse environmental processes, the wasteful byproducts of the profit oriented liberalized economy. The sensual reality of species and the accelerated species extinction is effectively masked by the virtual sciences of the abstract: numbers, data, statistics, algorithms, equations, models and ideas. To understand the birth of a young incipient species we have briefly reviewed the postmodern development of the unified phylogenetic species concept. (1) The reality of species and higher phylogenetic taxa. (2) The biological and phylogenetic species. (3) How to delineate phylogenetic species? (4) The infinite versus finite division of phylogenetic species. (5) The construct of the unified species concept. (6) Taking subspecies and race out of science. Without recognition of incipient siblings of the phylogenetic species the biodiversity remains under-estimated and the pharisaic anti-science ranking of humans remains with us. The discovery of speciation trait that is the sexual adaptive structures in reproductive barrier building, which are detectable by fine phenomics, gives perspective to find the finite division, the dynamic initial split in the continuous process of diversification. The speciation traits produced by integrative organisation, as opposed to competitive selection, help to unify the operational criteria of the biological species concept that is the speciation by reproductive isolation with the general concept of phylogenetic species that is the causal process of the separately evolving metapopulation lineages. The subspecies and racial ranking is untenable anymore, we suggest taking subspecies and race out of science: the finite division of the initial split detected by speciation traits is the birth of the phylogenetic incipient sibling species. There is no “subspecies”and “races”, as there is no “subindividual” in the biological organisation. In the present caddisfly taxonomy the subspecies remained as a valid status in the Potamophylax cingulatus caddisfly species group. With a clear distinction between the neutral and adaptive traits in the P. cingulatus species group and applying the subtle and stable shape divergences in phallic fine structures we have proposed to change the taxonomic status of subspecies to incipient sibling phylogenetic species rank: Potamophylax alpinus stat. nov., P. depilis stat nov., P. ibericus stat. nov., P. inermis stat. nov., reinstated the species status of P. cingulatus stat. restit. and we have described three new species: P. fesus Oláh, P. portugalicus Oláh et Szczesny, and P. transalpinus Oláh & Coppa, spp. nov.

In the last few years we have described over 70 new incipient sibling limnephild species applying the discovered Trichoptera speciation traits of the paraproct and paramere for species recognition and delimitation. In this revision on Drusinae subfamily, comprising 177 species, we have applied these subtle, but rapid and stable speciation traits and described 49 new sibling species from the “well studied” European mountain ranges. Discussing the theoretical background we have elaborated and adapted a new character state ranking system of phenomics to revise the long-neglected taxonomy of the Drusinae subfamily and synonymised the Cryptothrix, Monocentra, Metanoea, Leptodrusus, Anomalopterygella, Hadimina genera with the Drusus genus. These old genera of artificial constructs were established exclusively by divergences of secondary sexual traits known already to have only species level ranking value. According to our new character ranking system in the Drusinae subfamily, beside the Drusus genus, only the Ecclisopteryx genus has been retained having robust generic level divegences of paraproct loss and ancestral duplication of spine organising centre on the paramere pattern. Speciation trait function of the peg-packed surface on the paraproct head in Drusus genus moved to the gonopod apices and integrated into variously shaped stimulatory organ in the Ecclisopteryx genus. In the Drusus genus the ancestral divergence of the single spine organising centre has integrated 11 species groups with remarkably stable paramere spine pattern. Based upon ancestral divergences in the paraproct architecture we have differenciated 28 species complexes inside the 11 species groups. The delineation of the 163 mostly incipient siblings species, inside the 28 species complexes with 44 new Drusus species, was based primarily on the divergences of speciation trait, that is in the stimulatory head shape of the apical arms on the dorsal branches of the paraproct. In the Ecclisopteryx genus with 14 species we have established two independent lineages both with a single species, as well as two species complexes with five new species applying the speciation trait of the genus, that is the shape divergence of the stimulatory organ on the dorsoapical surface of the gonopods. Based on the Darwinian natural selection, we do not understand how the discovered 70+49 new European incipient phylogenetic species of limnephilid caddisflies have been evolved in the isolated sky island habitats of high mountain ranges. This isolation induced speciation represents a challenge to the mechanistic reductionist concept of the natural selection. Our first trial to extract information from various disciplines to answer this question is presented in a brief theoretical discourse: (1) rethinking the status of natural selection towards postdarwinism; (2) teleology or teleonomy; (3) limits and potentials in understanding reality; (4) organisation of universe by integration; (5) what are and how the organising forces are powered to work in the emerging energy mechanisms; (6) divergence by integration; (7) divergence in isolation; (8) reproductive isolation by sexual selection; (9) shape divergence; (10) speciation traits; (11) generic ranking characters. Assessing the limits and potentials, the humility and hybris attitudes towards understanding reality, we hypothesise an integrative power of organisation, instead of simplistic natural selection, that works both in isolated and sympatric populations to maintain the integer state of the integrated autonomic entities against mutations, permutations, transmutations and perturbations. The permanent flux of these adverse, neutral or advantegous external and internal effects generates and gives rise to a responsive and balancing integrative mechanisms with comparative magnitude and multitude. This permanently balancing organisation process develops and drives a large number of interactions along various patterns of supervenient emergent mechanisms in order to integrate them into an oscillating autonomy. In isolation of the sky-islands on high altitudes, the divergences of new taxa are the adaptive autonomic byproducts of this integrative organisation. Integration and not the selection is the real force mechanism that harmonises the concerted flux of stochastic processes into reproductive isolation by sexual selection. The primacy of integration is clearly demonstrated by empirical evidences in the causes and consequences of the integrated speciation traits. Any kind of selection, artificial, natural, social or sexual, is only an emergent perturbating mechanism forced to integrate into the autonomy of an entity. This is how the entanglement of the quantum world and any supervenient integer work together through energeticism of the interactive realism. This is a must in the ontic, epistemic and semantic structural realism.