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Woody species encroachment into herbaceous and shrub-dominated vegetations is a concern in many rangeland ecosystems of the world. Arrival of woody species into affected rangelands leads to changes in the spatial structure of vegetation and alterations of biophysical processes. In the western USA, encroachment of pinyon (Pinus spp.) and juniper (Juniperus spp.) tree species into sagebrush steppes poses a threat to the proper ecohydrological functioning of these ecosystems. Prescribed fire has been proposed and used as one rangeland improvement practice to restore sagebrush steppe from pinyon-juniper encroachment. Short-term effects of burning on the ecohydrologic response of these systems have been well documented and often include a period of increased hydrologic and erosion vulnerability immediately after burning. Long-term ecohydrologic response of sagebrush steppe ecosystems to fire is poorly understood due to lack of cross-scale studies on treated sites. The aim of this study is to evaluate long-term vegetation, hydrologic, and erosion responses at two pinyon-juniper-encroached sagebrush sites 9 years after prescribed fire was applied as a restoration treatment. Thirty-six rainfall simulation experiments on 6 m × 2 m plots were conducted for 45 min under two conditions: a dry run (70 mm h−1; dry antecedent soils) and a wet run (111 mm h−1; wet antecedent soils). Runoff and erosion responses were compared between burned and unburned plots. Overall, increases in herbaceous cover in the shrub-interspace areas (intercanopy area between trees) at both sites 9 years post-burn resulted in runoff- and erosion-reduction benefits, especially under the wet runs. While the initially more degraded site characterized by 80% bare ground pre-burn, registered a higher overall increase (40% increase) in canopy cover, greater post-fire reductions in runoff and erosion were observed at the less degraded site (57% bare ground pre-burn). Runoff and erosion for the wet runs decreased respectively by 6.5-fold and 76-fold at the latter site on the burned plots relative to control plots, whereas these decreases were more muted at the more degraded site (2.5 and 3-fold respectively). Significant fragmentation of flow paths observed at the more-degraded site 9 years post-fire, suggests a decreased hydrologic connectivity as a mechanism of runoff and erosion reduction during post-fire recovery.

Encroachment of sagebrush (Artemisia spp.) shrublands by pinyon (Pinus spp.) and juniper (Juniperus spp.) conifers (woodland encroachment) induces a shift from biotic‐controlled resource retention to abiotic‐driven loss of soil resources. This shift is driven by a coarsening of the vegetation structure with increasing dominance of site resources by trees. Competition between the encroaching trees and understory vegetation for limited soil and water resources facilitates extensive bare intercanopy area between trees and concomitant increases in run‐off and erosion that, over time, propagate persistence of the shrubland‐to‐woodland conversion. We evaluated whether tree removal by burning can decrease late‐succession woodland ecohydrologic resilience by increasing vegetation and ground cover over a 9‐year period after fire and whether the soil erosion feedback on late‐succession woodlands is reversible by burning. To address these questions, we employed a suite of vegetation and soil measurements and rainfall simulation and concentrated overland flow experiments across multiple plot scales on unburned and burned areas at two sagebrush sites in the later stages of woodland succession. Prior to burning, tree cover was approximately 28% at the sites, and more than 70% of the area at the sites was intercanopy with depauperate understory vegetation and extensive bare ground (52–60% bare soil and rock). Burning initially increased bare ground across fine (

Land managers across the western United States are faced with selecting and applying tree-removal treatments on pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland-encroached sagebrush (Artemisia spp.) rangelands, but current understanding of long-term vegetation and hydrological responses of sagebrush sites to tree removal is inadequate for guiding management. This study applied a suite of vegetation and soil measures (0.5 − 990 m2), small-plot rainfall simulations (0.5 m2), and overland flow experiments (9 m2) to quantify the effects of mechanical tree removal (tree cutting and mastication) on vegetation, runoff, and erosion at two mid- to late-succession woodland-encroached sagebrush sites in the Great Basin, United States, 9 yr after treatment. Low amounts of hillslope-scale shrub (3 − 15%) and grass (7 − 12%) canopy cover and extensive intercanopy (area between tree canopies) bare ground (69 − 88% bare, 75% of area) in untreated areas at both sites facilitated high levels of runoff and sediment from high-intensity (102 mm • h− 1, 45 min) rainfall simulations in interspaces (~ 45 mm runoff, 59 − 381 g • m− 2 sediment) between trees and shrubs and from concentrated overland flow experiments (15, 30, and 45 L • min− 1, 8 min each) in the intercanopy (371 − 501 L runoff, 2 342 − 3 015 g sediment). Tree cutting increased hillslope-scale density of sagebrush by 5% and perennial grass cover by twofold at one site while tree cutting and mastication increased hillslope-scale sagebrush density by 36% and 16%, respectively, and perennial grass cover by threefold at a second more-degraded (initially more sparsely vegetated) site over nine growing seasons. Cover of cheatgrass (Bromus tectorum L.) was < 1% at the sites pretreatment and 1 − 7% 9 yr after treatment. Bare ground remained high across both sites 9 yr after tree removal and was reduced by treatments solely at the more degraded site. Increases in hillslope-scale vegetation following tree removal had limited impact on runoff and erosion for rainfall simulations and concentrated flow experiments at both sites due to persistent high bare ground. The one exception was reduced runoff and erosion within the cut treatments for intercanopy plots with cut-downed-trees. The cut-downed-trees provided ample litter cover and tree debris at the ground surface to reduce the amount and erosive energy of concentrated overland flow. Trends in hillslope-scale vegetation responses to tree removal in this study demonstrate the effectiveness of mechanical treatments to reestablish sagebrush steppe vegetation without increasing cheatgrass for mid- to late-succession woodland-encroached sites along the warm-dry to cool-moist soil temperature − moisture threshold in the Great Basin. Our results indicate improved hydrologic function through sagebrush steppe vegetation recruitment after mechanical tree removal on mid- to late-succession woodlands can require more than 9 yr. We anticipate intercanopy runoff and erosion rates will decrease over time at both sites as shrub and grass cover continue to increase, but follow-up tree removal will be needed to prevent pinyon and juniper recolonization. The low intercanopy runoff and erosion measured underneath isolated cut-downed-trees in this study clearly demonstrate that tree debris following mechanical treatments can effectively limit microsite-scale runoff and erosion over time where tree debris settles in good contact with the soil surface.

Pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland encroachment has imperiled a broad ecological domain of the sagebrush steppe (Artemisia spp.) ecosystem in the Great Basin Region, USA. As these conifers increase in dominance on sagebrush rangelands, understory vegetation declines and ecohydrologic function can shift from biotic (vegetation) controlled retention of soil resources to abiotic (runoff) driven loss of soil resources and long-term site degradation. Scientists, public land management agencies, and private land owners are challenged with selecting and predicting outcomes to treatment alternatives to improve ecological structure and function on these rangelands. This study is the first of a two-part study to evaluate effectiveness of prescribed fire to re-establish sagebrush steppe vegetation and improve ecohydrologic function on mid- to late-succession pinyon-and juniper-encroached sagebrush sites in the Great Basin. We used a suite of vegetation and soil measures, small-plot (0.5 m2) rainfall simulations, and overland flow experiments (9 m2) to quantify the effects of tree removal by prescribed fire on vegetation, soils, and rainsplash, sheetflow, and concentrated flow hydrologic and erosion processes at two woodlands 9-yr after burning. For untreated conditions, extensive bare interspace (87% bare ground) throughout the degraded intercanopy (69–88% bare ground) between trees at both sites promoted high runoff and sediment yield from combined rainsplash and sheetflow (~45 mm, 59–381 g m−2) and concentrated flow (371–501 L, 2343–3015 g) processes during high intensity rainfall simulation (102 mm h−1, 45 min) and overland flow experiments (15, 30, and 45 L min−1, 8 min each). Burning increased canopy cover of native perennial herbaceous vegetation by >5-fold, on average, across both sites over nine growing seasons. Burning reduced low pre-fire sagebrush canopy cover (30 yr. Enhanced herbaceous cover in interspaces post-fire reduced runoff and sediment yield from high intensity rainfall simulations by >2-fold at both sites. Fire-induced increases in herbaceous canopy cover (from 34% to 62%) and litter ground cover (from 15% to 36%) reduced total runoff (from 501 L to 180 L) and sediment yield (from 2343 g to 115 g) from concentrated flow experiments in the intercanopy at one site. Sparser herbaceous vegetation (49% cover) and litter cover (8%) in the intercanopy at the other, more degraded site post-fire resulted in no significant reduction of total runoff (371 L to 266 L) and sediment yield (3015 g to 1982 g) for concentrated flow experiments. Areas underneath unburned shrub and tree canopies were well covered by vegetation and ground cover and generated limited runoff and sediment. Fire impacts on vegetation, ground cover, and runoff and sediment delivery from tree and shrub plots were highly variable. Burning litter covered areas underneath trees reduced perennial herbaceous vegetation and increased invasibility to the fire-prone annual cheatgrass (Bromus tectorum L.). Cheatgrass cover increased from

Hydrologic response to rainfall on fragmented or burnt hillslopes is strongly influenced by the ensuing connectivity of runoff and erosion processes. Yet cross-scale process connectivity is seldom evaluated in field studies owing to scale limitations in experimental design. This study quantified surface susceptibility and hydrologic response across point to hillslope scales at two degraded unburnt and burnt woodland sites using rainfall simulation and hydrologic modelling. High runoff (31–47 mm) and erosion (154–1893 g m–2) measured at the patch scale (13 m2) were associated with accumulation of fine-scale (0.5-m2) splash-sheet runoff and sediment and concentrated flow formation through contiguous bare zones (64–85% bare ground). Burning increased the continuity of runoff and sediment availability and yield. Cumulative runoff was consistent across plot scales whereas erosion increased with increasing plot area due to enhanced sediment detachment and transport. Modelled hillslope-scale runoff and erosion reflected measured patch-scale trends and the connectivity of processes and sediment availability. The cross-scale experiments and model predictions indicate the magnitude of hillslope response is governed by rainfall input and connectivity of surface susceptibility, sediment availability, and runoff and erosion processes. The results demonstrate the importance in considering cross-scale structural and functional connectivity when forecasting hydrologic and erosion responses to disturbances.

Extensive woodland expansion in the Great Basin has generated concern regarding ecological impacts of tree encroachment on sagebrush rangelands and strategies for restoring sagebrush steppe. This study used rainfall (0.5 m2 and 13 m2 scales) and concentrated flow simulations and measures of vegetation, ground cover, and soils to investigate hydrologic and erosion impacts of western juniper (Juniperus occidentalis Hook.) encroachment into sagebrush steppe and to evaluate short-term effects of burning and tree cutting on runoff and erosion responses. The overall effects of tree encroachment were a reduction in understory vegetation and formation of highly erodible, bare intercanopy between trees. Runoff and erosion from high-intensity rainfall (102 mm · h‒1, 13 m2 plots) were generally low from unburned areas underneath tree canopies (13 mm and 48 g · m‒2) and were higher from the unburned intercanopy (43 mm and 272 g · m‒2). Intercanopy erosion increased linearly with runoff and exponentially where bare ground exceeded 60%. Erosion from simulated concentrated flow was 15- to 25-fold greater from the unburned intercanopy than unburned tree canopy areas. Severe burning amplified erosion from tree canopy plots by a factor of 20 but had a favorable effect on concentrated flow erosion from the intercanopy. Two years postfire, erosion remained 20-fold greater on burned than unburned tree plots, but concentrated flow erosion from the intercanopy (76% of study area) was reduced by herbaceous recruitment. The results indicate burning may amplify runoff and erosion immediately postfire. However, we infer burning that sustains residual understory cover and stimulates vegetation productivity may provide long-term reduction of soil loss relative to woodland persistence. Simply placing cut-downed trees into the unburned intercanopy had minimal immediate impact on infiltration and soil loss. Results suggest cut-tree treatments should focus on establishing tree debris contact with the soil surface if treatments are expected to reduce short-term soil loss during the postcut understory recruitment period.