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The Triassic Period (~252–201.5 Ma) saw a transformative radiation and reorganization of continental tetrapod diversity following the end-Permian Extinction, including an assemblage of diverse forms that do not survive the end-Triassic (herein termed the 'endemic Triassic fauna', =ETF), as well as the earliest fossil representatives of all major modern tetrapod groups (herein termed the 'Living [Triassic to Recent] Fauna', =LTF; i.e. Salientia, Caudata, Gymnophiona, Mammaliaformes, Squamata, Rhynchocephalia, Testudinata, Crocodylomorpha, and Dinosauria). With few exceptions, only the LTF assemblage survives the end-Triassic Extinction (~201.5 Ma), highlighting the Late Triassic (~227–201.5 Ma) record as essential for understanding this pivotal transition and the evolutionary and ecological origins of post-Triassic non-marine tetrapod faunas, including those of present day. Micro-microvertebrate bonebeds are arguably the best proxy for tracking continental vertebrate biodiversity, however gaps in their Late Triassic record obscure patterns and drivers of evolutionary, ecological, and environmental change during the rise of LTF communities. In my dissertation, I use new data collected from Upper Triassic microvertebrate bonebeds from North America, and particularly the Thunderstorm Ridge site (PFV 456) in Petrified Forest National Park, Arizona, U.S.A, to fill gaps in the evolutionary record of specific groups (e.g., lissamphibians and lepidosaurs), as well as the vertebrate paleocommunity record of Triassic equatorial Pangaea. My first chapter describes and analyzes an assemblage of gymnophionomorph (stem caecilian) bones from PFV 456 which represent the oldest-known caecilian fossils globally. As the oldest caecilian fossils, they provide new support for the dissorophoid temnospondyl affinities of caecilians and other living amphibians, evidence of a step-wise acquisition of caecilian anatomies associated with fossoriality, and evidence of an ancient pattern of equa

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Cratons are conventionally assumed to be areas of long-term stability. However, whereas Precambrian basement crops out across most of the Baltic Shield, Palaeozoic and Mesozoic sediments rest on basement in southern Sweden, and thus testify to a complex history of exhumation and burial. Our synthesis of published stratigraphic landscape analysis and new apatite fission-track analysis data reveals a history involving five steps after formation of the extremely flat, Sub-Cambrian Peneplain. (1) Cambrian to Lower Triassic rocks accumulated on the peneplain, interrupted by late Carboniferous uplift and exhumation. (2) Middle Triassic uplift removed the Palaeozoic cover along the south-western margin of the shield, leading to formation of a Triassic peneplain with a predominantly flat relief followed by deposition of Upper Triassic to Lower Jurassic rocks. (3) Uplift that began during the Middle Jurassic to earliest Cretaceous caused denudation leading to deep weathering that shaped an undulating, hilly relief that was buried below Upper Cretaceous to Oligocene sediments. (4) Early Miocene uplift and erosion produced the South Småland Peneplain with scattered hills. (5) Early Pliocene uplift raised the Miocene peneplain to its present elevation leading to reexposure of the sub-Cretaceous hilly relief near the coast. Our results thus provide constraints on the magnitude and timing of episodes of deposition and removal of significant volumes of Phanerozoic rocks across the southern portion of the Baltic Shield. Late Carboniferous, Middle Triassic and mid-Jurassic events of uplift and exhumation affected wide areas beyond the Baltic Shield, and we interpret them as epeirogenic uplifts accompanying fragmentation of Pangaea, caused by accumulation of mantle heat beneath the supercontinent. Early Miocene uplift affected north-west Europe but not East Greenland, and thus likely resulted from compressive stresses from an orogeny on the Eurasian plate. Early Pliocene uplift relat

Cratons are conventionally assumed to be areas of long-term stability. However, whereas Precambrian basement crops out across most of the Baltic Shield, Palaeozoic and Mesozoic sediments rest on basement in southern Sweden, and thus testify to a complex history of exhumation and burial. Our synthesis of published stratigraphic landscape analysis and new apatite fission-track analysis data reveals a history involving five steps after formation of the extremely flat, Sub-Cambrian Peneplain. (1) Cambrian to Lower Triassic rocks accumulated on the peneplain, interrupted by late Carboniferous uplift and exhumation. (2) Middle Triassic uplift removed the Palaeozoic cover along the south-western margin of the shield, leading to formation of a Triassic peneplain with a predominantly flat relief followed by deposition of Upper Triassic to Lower Jurassic rocks. (3) Uplift that began during the Middle Jurassic to earliest Cretaceous caused denudation leading to deep weathering that shaped an undulating, hilly relief that was buried below Upper Cretaceous to Oligocene sediments. (4) Early Miocene uplift and erosion produced the South Småland Peneplain with scattered hills. (5) Early Pliocene uplift raised the Miocene peneplain to its present elevation leading to reexposure of the sub-Cretaceous hilly relief near the coast. Our results thus provide constraints on the magnitude and timing of episodes of deposition and removal of significant volumes of Phanerozoic rocks across the southern portion of the Baltic Shield. Late Carboniferous, Middle Triassic and mid-Jurassic events of uplift and exhumation affected wide areas beyond the Baltic Shield, and we interpret them as epeirogenic uplifts accompanying fragmentation of Pangaea, caused by accumulation of mantle heat beneath the supercontinent. Early Miocene uplift affected north-west Europe but not East Greenland, and thus likely resulted from compressive stresses from an orogeny on the Eurasian plate. Early Pliocene upl, Publ online July 2015

Until recently the climate of the Early–Middle Triassic at low latitudes was broadly considered as generally temperate-warm with no major climate oscillations. This work examines the climate of this period through a detailed study of the sedimentary, plant, soil and mineral records of continental rocks (Buntsandstein facies) in eastern Iberian basins. Our findings indicate temporal climate variations for these near equator (10°–14°N) regions and unveil the significance of such variations in the southern Laurasian domain. The climate of Iberia's Early Triassic was mainly dominated by alternating brief (< 0.4 ma) arid and semi-arid climate periods, with two main arid periods documented at the end of the Smithian and middle Spathian. However, an initial short subhumid to semi-arid period was also observed in the late Spathian. Remarkably, this latter period appears just after an unconformity related to the tectonically induced Hardegsen Event in western Europe. It is also of interest that this short subhumid climate period is concurrent with the beginning of faunal and floral recovery in the basins examined. The Early Triassic ended again with a short very arid period. Although the beginning of the Anisian (Aegean) was represented by alternating arid and semi-arid to subhumid intervals, during the Bithynian and Pelsonian clearly wetter climates are recorded by the succession consisting of alternating semi-arid to semi-humid intervals. This general tendency was interrupted by three short but marked intervals, two humid intervals in the late Bithynian, and one arid period near the Bithynian/Pelsonian boundary., Ministerio de Economía y Competitividad (MINECO), Universidad Complutense de Madrid, Consejo Superior de Investigaciones Científicas (CSIC), Depto. de Mineralogía y Petrología, Fac. de Ciencias Geológicas, TRUE, pub

Until recently the climate of the Early–Middle Triassic at low latitudes was broadly considered as generally temperate-warm with no major climate oscillations. This work examines the climate of this period through a detailed study of the sedimentary, plant, soil and mineral records of continental rocks (Buntsandstein facies) in eastern Iberian basins. Our findings indicate temporal climate variations for these near equator (10°–14°N) regions and unveil the significance of such variations in the southern Laurasian domain. The climate of Iberia's Early Triassic was mainly dominated by alternating brief (< 0.4 ma) arid and semi-arid climate periods, with two main arid periods documented at the end of the Smithian and middle Spathian. However, an initial short subhumid to semi-arid period was also observed in the late Spathian. Remarkably, this latter period appears just after an unconformity related to the tectonically induced Hardegsen Event in western Europe. It is also of interest that this short subhumid climate period is concurrent with the beginning of faunal and floral recovery in the basins examined. The Early Triassic ended again with a short very arid period. Although the beginning of the Anisian (Aegean) was represented by alternating arid and semi-arid to subhumid intervals, during the Bithynian and Pelsonian clearly wetter climates are recorded by the succession consisting of alternating semi-arid to semi-humid intervals. This general tendency was interrupted by three short but marked intervals, two humid intervals in the late Bithynian, and one arid period near the Bithynian/Pelsonian boundary. Iberia was crossed by prominent irregular highs separating marked corridors or isolated areas. This palaeogeography, prevailing since Variscan tectonics, clearly conditioned dominant climates and their geographical distribution. No clear climate belts developed in these conditions. However, isolated internal climate zones separated by elevated areas are identified.

Incorporation of fossils into biogeographic studies can have a profound effect on the conclusions that result, particularly when fossil ranges are nonoverlapping with extant ranges. This is the case in archaeid spiders, where there are known fossils from the Northern Hemisphere, yet all living members are restricted to the Southern Hemisphere. To better understand the biogeographic patterns of archaeid spiders and their palpimanoid relatives, we estimate a dated phylogeny using a relaxed clock on a combined molecular and morphological data set. Dating information is compared with treating the archaeid fossil taxa as both node calibrations and as noncontemporaneous terminal tips, both with and without additional calibration points. Estimation of ancestral biogeographic ranges is then performed, using likelihood and Bayesian methods to take into account uncertainty in phylogeny and in dating. We find that treating the fossils as terminal tips within a Bayesian framework, as opposed to dating the phylogeny based only on molecular data with the dates coming from node calibrations, removes the subjectivity involved in assigning priors, which has not been possible with previous methods. Our analyses suggest that the diversification of the northern and southern archaeid lineages was congruent with the breakup of Pangaea into Laurasia and Gondwanaland. This analysis provides a rare example, and perhaps the most strongly supported, where a dated phylogeny confirms a biogeographical hypothesis based on vicariance due to the breakup of the ancient continental plates.

Incorporation of fossils into biogeographic studies can have a profound effect on the conclusions that result, particularly when fossil ranges are nonoverlapping with extant ranges. This is the case in archaeid spiders, where there are known fossils from the Northern Hemisphere, yet all living members are restricted to the Southern Hemisphere. To better understand the biogeographic patterns of archaeid spiders and their palpimanoid relatives, we estimate a dated phylogeny using a relaxed clock on a combined molecular and morphological data set. Dating information is compared with treating the archaeid fossil taxa as both node calibrations and as noncontemporaneous terminal tips, both with and without additional calibration points. Estimation of ancestral biogeographic ranges is then performed, using likelihood and Bayesian methods to take into account uncertainty in phylogeny and in dating. We find that treating the fossils as terminal tips within a Bayesian framework, as opposed to dating the phylogeny based only on molecular data with the dates coming from node calibrations, removes the subjectivity involved in assigning priors, which has not been possible with previous methods. Our analyses suggest that the diversification of the northern and southern archaeid lineages was congruent with the breakup of Pangaea into Laurasia and Gondwanaland. This analysis provides a rare example, and perhaps the most strongly supported, where a dated phylogeny confirms a biogeographical hypothesis based on vicariance due to the breakup of the ancient continental plates.

New tetrapod footprints belonging to the ichnogenus Hyloidichnus have been discovered in Turkey for the first time, in the lower part of the Cakraz Formation (Northwestern Anatolia) and together with macrofloral imprints of Annularia and Stigmaria. These discoveries confirm the Permian age of the fossiliferous red beds in which the coniferophyte Walchia was previously recorded. Based on the stratigraphic range of Annularia,Stigmaria and Hyloidichnus known elsewhere, a Cisuralian age is proposed for these beds. These new ichno- and macrofloral remains, together with the sedimentological data (mudcracks, rain drops) suggest the presence of captorhinid reptiles living in a palustrine floodplain environment, and under a warm, seasonal climate alternating between humid and relatively long dry seasons. These climatic conditions may have permitted the migration of these captorhinids through Laurasia during the Permian.