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High-resolution pollen and dinoflagellate cyst records from sediment core M72/5-25-GC1 were used to reconstruct vegetation dynamics in northern Anatolia and surface conditions of the Black Sea between 64 and 20 ka BP. During this period, the dominance of Artemisia in the pollen record indicates a steppe landscape and arid climate conditions. However, the concomitant presence of temperate arboreal pollen suggests the existence of glacial refugia in northern Anatolia. Long-term glacial vegetation dynamics reveal two major arid phases ~64-55 and 40-32 ka BP, and two major humid phases ~54-45 and 28-20 ka BP, correlating with higher and lower summer insolation, respectively. Dansgaard-Oeschger (D-O) cycles are clearly indicated by the 25-GC1 pollen record. Greenland interstadials are characterized by a marked increase in temperate tree pollen, indicating a spread of forests due to warm/wet conditions in northern Anatolia, whereas Greenland stadials reveal cold and arid conditions as indicated by spread of xerophytic biomes. There is evidence for a phase lag of ~500 to 1500 yr between initial warming and forest expansion, possibly due to successive changes in atmospheric circulation in the North Atlantic sector. The dominance of Pyxidinopsis psilata and Spiniferites cruciformis in the dinocyst record indicates brackish Black Sea conditions during the entire glacial period. The decrease of marine indicators (marine dinocysts, acritarchs) at ~54 ka BP and increase of freshwater algae (Pediastrum, Botryococcus) from 32 to 25 ka BP reveals freshening of the Black Sea surface water. This freshening is possibly related to humid phases in the region, to connection between Caspian Sea and Black Sea, to seasonal freshening by floating ice, and/or to closer position of river mouths due to low sea level. In the southern Black Sea, Greenland interstadials are clearly indicated by high dinocyst concentrations and calcium carbonate content, as a result of an increase in primary productivity. Heinrich events show a similar impact on the environment in the northern Anatolia/Black Sea region as Greenland stadials. [ABSTRACT FROM AUTHOR]

High-resolution pollen and dinoflagellate cyst records from marine sediment core 25-GC1 were used to reconstruct vegetation dynamics in Northern Anatolia and surface conditions of the Black Sea between 64 and 25 ka BP. During this period, the dominance of Artemisia in the pollen record indicates a steppe landscape and arid climate conditions. However, the presence of temperate and warm-temperate arboreal pollen suggests the existence of glacial refugia in Northern Anatolia. A general cooling trend towards 25 ka BP is evidenced by the decrease of Quercus and increase of Pinus. There is evidence of orbital-driven vegetation dynamics in Northern Anatolia during 64-25 ka BP with spread of steppe during precession minima (insolation maxima) and development of forests during precession maxima (insolation minima). Dansgaard-Oeschger (D-O) events are characterized by a marked increase in temperate tree pollen, indicating a spread of forests due to warm and wet conditions in Northern Anatolia. The dominance of Pyxidinopsis psilata and Spiniferites cruciformis in the dinocyst record indicates a rather brackish Black Sea during the last glacial period. The decrease of marine indicators (marine dinocysts, acritachs) at ~ 54 ka BP and increase of freshwater algae (Pediastrum, Botryococcus) from 32 to 25 ka BP reveals freshening of the Black Sea surface water, related to orbital-driven arid/humid phases in the region, influencing hydrology and level changes of the Black Sea. D-O interstadials are characterized by high dinocyst concentrations and calcium carbonate content, as a result of an increase in primary productivity in the Black Sea. Heinrich events show a similar impact on the environment in Northern Anatolia/Black Sea region as D-O stadials. [ABSTRACT FROM AUTHOR]

This paper presents two dinoflagellate cyst records from the south-western shelf of the Black Sea. A new site, MAR05-13, from the Sakarya shelf is described and placed into context with site MAR02-45, ∼250 km distant on the Thracian shelf. The records provide a centennial resolution of surface water conditions in the Holocene. Analysis of the data suggests that the surface salinity of the south-western shelf increased in a gradual and progressive manner. In the period ∼11 000-9000 cal a BP the assemblages suggest surface-water salinities between 7-13 psu. The initial arrival of euryhaline species, ∼8100 cal a BP, is linked to the reconnection of the Black Sea and Marmara Sea. The suggested surface water changes related to the reconnection took approximately 1000 years. Following this initial change in assemblages, a further increase in the number of euryhaline species is noted between 5000 and 4000 cal a BP. This is linked to the establishment of more saline surface-water conditions, close to present-day values. The record for MAR05-13 highlights the complexity of the changes in cyst assemblages during the mid-Holocene. Copyright © 2012 John Wiley & Sons, Ltd. [ABSTRACT FROM AUTHOR]

The rapid climatic variability characterising the Marine Isotopic Stage (MIS) 3 (~60-30 cal ka BP) provides key issues to understand the atmosphere-ocean-cryosphere dynamics. Here we investigate the response of sea-surface paleoenvironments to the MIS3 climatic variability through the study of a high resolution oceanic sedimentological archive (core MD99-2281, 60°21' N; 09°27'W; 1197m water depth), retrieved during the MD114-IMAGES (International Marine Global Change Study) cruise from the southern part of the Faeroe Bank. This sector was under the proximal influence of European ice sheets (Fennoscandian Ice Sheet to the East, British Irish Ice Sheet to the South) during the last glacial and thus probably responded to the MIS3 pulsed climatic changes. We conducted a multi-proxy analysis of core MD99- 2281, including magnetic properties, x-ray fluorescence measurements, characterisation of the coarse (>150 µm) lithic fraction (grain concentration) and the analysis of selected biogenic proxies (assemblages and stable isotope ratio of calcareous planktonic foraminifera, dinoflagellate cyst - e.g. dinocyst - assemblages). Results presented here are focussed on the dinocyst response, this proxy providing the reconstruction of past sea-surface hydrological conditions, qualitatively as well as quantitatively (e.g. transfer function sensu lato). Our study documents a very coherent and sensitive oceanic response to the MIS3 rapid climatic variability: strong fluctuations, matching those of stadial/interstadial climatic oscillations as depicted by Greenland ice cores, are recorded in the MD99-2281 archive. Proxies of terrigeneous and detritical material suggest increases in continental advection during Greenland Stadials (including Heinrich events), the latter corresponding also to southward migrations of polar waters. At the opposite, milder sea-surface conditions seem to develop during Greenland Interstadials. After 30 ka, reconstructed paleohydrological conditions evidence strong shifts in SST: this increasing variability seems consistent with the hypothesised coalescence of the British and Fennoscandian ice sheets at that time, which could have directly influenced sea-surface environments in the vicinity of core MD99-2281. [ABSTRACT FROM AUTHOR]

The rapid climatic variability characterising the Marine Isotopic Stage (MIS) 3 (~ 60-30 CAL-ka BP) provides key issues to understand the atmosphere-ocean-cryosphere dynamics. Here we investigate the response of sea-surface paleoenvironments to the MIS3 climatic variability through the study of a high resolution oceanic sedimentological archive (core MD99-2281, 60°21' N; 09°27' W; 1197 m water depth), retrieved during the MD114-IMAGES (International Marine Global Change Study) cruise from the Southern part of the Faeroe Bank. This sector was under the proximal influence of European Ice Sheets (Fennoscandian Ice Sheet to the East, British Irish Ice Sheet to the South) and thus probably recorded their response to the MIS3 pulsed climatic changes. We conducted a multi-proxy analysis on core MD99-2281, including magnetic properties, X-Ray Fluorescence measurements, characterisation of the coarse (> 150 µm) lithic fraction (grain concentration) and the analysis of selected biogenic proxies (assemblages and stable isotope ratio of calcareous planktonic foraminifera, dinoflagellate cyst - e.g. dinocyst -- assemblages). Results presented here are focussed on the dinocyst response, this proxy providing the reconstruction of past sea-surface hydrological conditions, qualitatively as well as quantitatively (e.g. transfer function sensu lato). Our study documents a very coherent and sensitive oceanic response to the MIS3 rapid climatic variability: strong fluctuations, matching those of stadial/interstadial climatic oscillations as depicted by Greenland Ice Cores, are recorded in the MD99-2281 archive. Proxies of terrigeneous and detritical material typify increases in continental advection during Greenland Stadials (including Heinrich events), the latter corresponding also to southward migrations of polar waters. At the opposite, milder sea-surface conditions seem to develop during Greenland Interstadials. After 30 ka, reconstructed paleohydrological conditions evidence strong shifts in SST: this increasing variability seems consistent with the hypothesised coalescence of the British and Fennoscandian ice sheets at that time, which could have directly influenced sea-surface environments in the vicinity of core MD99-2281. [ABSTRACT FROM AUTHOR]

New dinocyst analyses were conducted on core MD99-2339 retrieved from the central Gulf of Cadiz. Dinocyst and foraminiferal assemblages from this core are combined with existing data off SW Portugal and NW Morocco to investigate past hydrological and primary productivity regimes in the subtropical NE Atlantic Ocean over the last 30 ka. Our results have revealed highest upwelling intensity during Heinrich Stadial 1 (HS 1) and the Younger Dryas and weaker upwelling cells during the Last Glacial Maximum (LGM) and HS 2, off the SW Iberian and NW Moroccan margins. Similar assemblages between the Gulf of Cadiz and the NW Moroccan margin, and distinct species off Portugal, were observed during the cold climatic extremes that punctuated the last 30 ka. This pattern has been linked to the occurrence of a hydrological structure between SW Iberia and Cadiz during the last glacial period, perhaps similar to the modern Azores Front. This front was probably responsible locally for heterotrophic dinocysts found in the Gulf of Cadiz during the last glacial period, even if this sector is not conductive to upwelling phenomena by Ekman transport. Regional reconstructions of paleo-sea-surface temperatures (SSTs) using dinocyst and foraminiferal transfer functions, as well as alkenones, are also discussed and depict coherent scenarios over the last 30 ka. Seasonal reconstructions of LGM SSTs obtained with this multi-proxy panel are discussed jointly with model outputs in order to contribute to ongoing efforts in model-data comparison. [ABSTRACT FROM AUTHOR]

New dinocyst analyses were conducted on core MD99-2339 retrieved from the central Gulf of Cadiz. Dinocyst and foraminiferal assemblages from this core are combined with existing data off SW Portugal and NW Morocco to investigate past hydrological and primary productivity regimes in the subtropical NE Atlantic Ocean over the last 30 ka. Our results have revealed highest upwelling intensity during Heinrich Stadial 1 (HS 1) and the Younger Dryas and weaker upwelling cells during the Last Glacial Maximum and HS 2, off the SW Iberian and NW Moroccan margins. Similar assemblages between Cadiz and Morocco and distinct species off Portugal were observed during the cold climatic extremes that punctuated the last 30 ka. This pattern has been linked to the occurrence of a hydrological structure between SW Iberia and Cadiz during the last glacial period, probably similar to the modern Azores Front. This front was probably responsible locally for heterotrophic dinocysts found in the Gulf of Cadiz during the last glacial period, even if this sector is not conductive to upwelling phenomena by Ekman transport. Regional reconstructions of paleo-sea-surface temperatures using dinocyst and foraminiferal transfer functions, as well as alkenones, are also discussed and depict coherent scenarios over the last 30 ka. However, some mismatches are observed between the different quantitative reconstructions such as during HS 1 in the Gulf of Cadiz and during the LGM at the three core locations. [ABSTRACT FROM AUTHOR]

High-temporal resolution pollen record from the Alboran Sea ODP Site 976, pollen-based quantitative climate reconstruction and biomisation show that changes of Mediterranean vegetation have been clearly modulated by short and long term variability during the last 25 000 years. The reliability of the quantitative climate reconstruction from marine pollen spectra has been tested using 22 marine core-top samples from the Mediterranean. The ODP Site 976 pollen record and climatic reconstruction confirm that Mediterranean environments have a rapid response to the climatic fluctuations during the last Termination. The western Mediterranean vegetation response appears nearly synchronous with North Atlantic variability during the last deglaciation as well as during the Holocene. High-resolution analyses of the ODP Site 976 pollen record show a cooling trend during the Bölling/Allerød period. In addition, this period is marked by two warm episodes bracketing a cooling event that represent the Bölling-Older Dryas-Allerød succession. During the Holocene, recurrent declines of the forest cover over the Alboran Sea borderlands indicate climate events that correlate well with several events of increased Mediterranean dryness observed on the continent and with Mediterranean Sea cooling episodes detected by alkenonebased sea surface temperature reconstructions. These events clearly reflect the response of the Mediterranean vegetation to the North Atlantic Holocene cold events. [ABSTRACT FROM AUTHOR]

The Sea of Marmara is the connection between the vast Black Sea-Caspian Sea basin (Pontocaspian) and the Global Ocean via the Mediterranean Sea. Its water levels and water conditions has widely varied over times. Combining two cores in the Sea of Marmara (Turkey) and using organic-walled dinoflagellate cyst assemblages as the main proxy (combined with alkenones, diatoms and benthic foraminifera), allow qualitatively reconstructing water conditions during Marine Isotopic Stage (MIS) 6 and 5, such as salinity and oxygen level. A clear main marine phase is illustrated in MIS 5e. A minor marine incursion occurred during MIS 5c, mostly supported by alkenone data. The rest of the record indicates brackish Pontocaspian conditions, with more Spiniferites inaequalis in MIS 6 and more S. cruciformis in the non-marine parts of MIS 5. At the MIS 6/MIS 5 transition, an earlier initial marine flooding in the Sea of Marmara (dinocyst assemblages) in comparison to the Black Sea was highlighted. The marine reconnection occurred at different moments as seen in the terrestrial vegetation reconstructed from pollen analysis linking the two seas. The sapropels of the Sea of Marmara form when marine water penetrates at depth from the Aegean Sea beneath a layer of lower salinity water. Variations of the residence time of the marine deep water in a stratified Sea of Marmara are potentially an important factor underlying hypoxia/anoxia and sapropel deposition. When combining surface water proxies with benthic foraminifera (test assemblages and presence of inner organic lining), it appears that the successive MIS 5 sapropels formed under decreasing salinity and oxygen availability conditions as the marine inflow was increasingly restricted. Understanding the hydrologic evolution of the Sea of Marmara during MIS 5 implies taking into account that the Bosphorus and Dardanelle straits are hydrodynamically coupled and may not be simplified as on/off switches based on present day sill depths. • Dinocyst analyses of last interglacial and previous glacial in the Sea of Marmara on two long cores. • Marine phase in MIS5e (dinocysts, benthic foraminifera, alkenones) minor in MIS5c • BrackishPontocaspian conditions for rest of sequence although different in MIS6 & 5. • Earlier marine flooding at beginning of MIS5e in Sea of Marmara than in Black Sea. • Three sapropels highlight decreasing anoxic conditions over time. [ABSTRACT FROM AUTHOR]

Abstract: The interglacial periods of the late Quaternary are frequently investigated as they constitute potential analogues for our modern climate and may shed light on the key questions of natural climate variability and future developments. The aim of this work is to reconstruct the paleoenvironmental history of Marine Isotope Stages (MIS) 5 and 7 in deep-sea sediments of the northern Bay of Biscay (North East Atlantic Ocean). On the basis of a multiproxy compilation, including analysis of dinoflagellate cyst assemblages, we discuss the nature of the paleoenvironmental and paleoclimate changes that took place in this region. Recurrent successions of species marking the beginning and the Termination of the Interglacial Complexes (MIS 5 and MIS 7) revealed a coherent scheme of water mass migration during these key transitional periods. Moreover, our data extend knowledge about the ecology of several dinoflagellate taxa. In particular, we discuss the ecology of Spiniferites septentrionalis and note that this species can be used as a biostratigraphical tracer in North Atlantic Quaternary sediments, until a major event of iceberg calving occurred during MIS 6, dated to around 150 ka BP. The dinocyst Spiniferites mirabilis appears to be an important proxy for recognizing warm intervals within Interglacial Complexes. During MIS 5e, the Last Interglacial, this species is represented by the highest percentages ever recorded in sediments from the North Atlantic region. Peak occurrence of this species during MIS 7 indicates that substage MIS 7c, the second warm interval of the Penultimate Interglacial, represents the climatic optimum during the MIS 7 Interglacial Complex. [Copyright &y& Elsevier]

In the current circumscription, the Thoracosphaeraceae comprise all dinophytes exhibiting calcified coccoid cells produced during their life-history. Species hitherto assigned to Ensiculifera and Pentapharsodinium are mostly based on the monadoid stage of life-history, while the link to the coccoid stage (occasionally treated taxonomically distinct) is not always resolved. We investigated the different life-history stages and DNA sequence data of Ensiculifera mexicana and other species occurring in samples collected from all over the world. Based on concatenated ribosomal RNA gene sequences Ensiculiferaceae represented a distinct peridinalean branch, which showed a distant relationship to other calcareous dinophytes. Both molecular and morphological data (particularly of the coccoid stage) revealed the presence of three distinct clades within Ensiculiferaceae, which may include other dinophytes exhibiting a parasitic life-history stage. At a higher taxonomic level, Ensiculiferaceae showed relationships to parasites and endosymbionts (i.e., Blastodinium and Zooxanthella) as well as to dinophytes harbouring diatoms instead of chloroplasts. These unexpected phylogenetic relationships are corroborated by the presence of five cingular plates in all such taxa, which differs from the six cingular plates of most other Thoracosphaeraceae. We herein describe Ensiculiferaceae, emend the descriptions of Ensiculifera and Pentapharsodinium, erect Matsuokaea and provide several new combinations at the species level., (Copyright © 2020 Elsevier GmbH. All rights reserved.)