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The rotation forestry system provides high biomass production, but could also have a negative impact on species sensitive to disturbance. Continuous cover forestry (CCF) could contribute to solving these conflicting goals, but its feasibility in nutrient limited boreal forests is yet unresolved. In a unique experiment, we simultaneously assessed the short-term effect of single-tree selection on both biomass production and biodiversity (vascular plants, bryophytes, wood-inhabiting fungi), and tested fertilization as a way to mediate growth-biodiversity trade-offs. We found that unharvested stands and stands subjected to single-tree selection had a similar species assemblage of vascular plants, bryophytes, and wood-inhabiting fungi. Fertilization increased growth by 37% and induced shifts in two understory species (favoring the grass Avenella flexuosa and disfavoring the bryophyte Hylocomium splendens). We conclude that single-tree selection may become a useful tool to enhance biodiversity in managed forests. [ABSTRACT FROM AUTHOR]

Conservation of biodiversity requires in-depth knowledge of trait-environment interactions to understand the influence the environment has on species assemblages. Saproxylic beetles exhibit a wide range of traits and functions in the forest ecosystems. Understanding their responses to surrounding environment thus improves our capacity to identify habitats that should be restored or protected. We investigated potential interactions between ecological traits in saproxylic beetles (feeding guilds and habitat preferences) and environmental variables (deadwood, type and age of surrounding forest). We sampled beetles from 78 plots containing newly created high stumps of Scots pine and Silver birch in boreal forest landscapes in Sweden for three consecutive years. Using a model based approach, our aim was to explore potential interactions between ecological traits and the surrounding environment at close and distant scale (20 m and 500 m radius). We found that broadleaf-preferring beetle species are positively associated with the local broadleaf-originated deadwood and broadleaf-rich forests in the surrounding landscapes. Conifer-preferring species are positively associated with the local amount of coniferous deadwood and young and old forests in the surrounding landscape. Fungivorous and predatory beetles are positively associated with old forests in the surrounding landscapes. Our results indicate that both local amounts of deadwood and types of forests in the landscape are important in shaping saproxylic beetle communities. We particularly highlight the need to increase deadwood amounts of various qualities in the landscape, exempt older forests from production and to increase broadleaf-rich habitats in order to meet different beetle species' habitat requirements. Trait responses among saproxylic beetles provide insights into the significance of broadleaf forest and dead wood as essential attributes in boreal forest restoration, which helps conservation planning and management in forest landscapes. • Trait – environment interactions of saproxylic beetles are investigated. • Broadleaf-preferring beetles thrive with local broadleaved deadwood. • Broadleaf-rich forests contribute to the abundance of broadleaf-preferring beetles. • Fungivores and predators positively interact with old forests in the landscape. • Deadwood increase and a 'broadleafication' of boreal forests need to be implemented. [ABSTRACT FROM AUTHOR]

• Continuous Cover Forestry is highly debated. • We reviewed long-term yield and biodiversity between two silviculture systems. • Comparative studies between silviculture systems are inconclusive. • Species sensitive to clearcutting may benefit from Continuous Cover Forestry. • More long-term experiments are needed, especially on conservation species. There is an increasing interest in implementing Continuous Cover Forestry (CCF) as a tool to mitigate negative effects of the traditional rotation forestry system on biodiversity. However, the effects of CCF on forest growth and yield and on biodiversity is still poorly known. In this qualitative review, we compare biodiversity and long-term yield between the selection system, which is a type of CCF practiced in full-storied forests, and the traditional rotation forestry system. We specifically focus on forests dominated by Picea abies , which is a tree species of high economic relevance. Our literature search resulted in 17 publications on stand growth and yield and 21 publications on biodiversity. A majority of simulation studies found a higher long-term yield in the rotation forestry system, but it is challenging to conclude which system is the most productive. The magnitude of the difference in yield between systems, and how it varies across different environmental conditions, remains to be determined. For biodiversity, comparisons of species assemblage and individual species were only made to certain phases of the rotation cycle (recent clearcuts and middle-aged stands). Nevertheless, two aspects can be highlighted: i) the species assemblage in clearcuts differ substantially from stands managed with the selection system. Some of these effects may however be short lasting as examplified by studies on beetle assamblages showing that middle-aged rotation forestry stands become more similar to stands managed with the selection system, ii) the selection system maintains a similar species assemblage as the uncut control during the first years after cutting. In conclusion, management with the selection system may come with a loss in long-term stand yield, but much of the species assemblage is maintained after logging. We recommend future studies to specifically focus on long-term effects on biodiversity – in particular on species of conservation concern. There is also a need to establish a long-term research infrastructure to further develop the field. [ABSTRACT FROM AUTHOR]

Management of forest for wood production has altered ecosystem structures and processes and led to habitat loss and species extinctions, worldwide. Deadwood is a key resource supporting forest biodiversity, and commonly declines following forest management. However, different forest management methods affect dead wood differently. For example, uneven-aged silviculture maintains an age-stratified forest with ongoing dead wood production, while even-aged silviculture breaks forest continuity, leading to long periods without large trees. We asked how deadwood-dependent beetles respond to different silvicultural practices and if their responses depend on deadwood volume, and beetles preference for decay stages of deadwood. We compared beetle assemblages in five boreal forest types with different management strategies: clearcutting and thinning (both representing even-aged silviculture), selective felling (representing uneven-aged silviculture), reference and old growth forest (both uneven-aged controls without a recent history [~50 years] of management, but the latter with high conservation values). We collected beetles using window traps and by sieving the bark from experimental logs (bolts). Beetle assemblages on clear-cuts differed from all other stand types, regardless of trapping method or decay stage preference. Thinning differed from reference stands, indicating incomplete recovery after clear-cutting, while selective felling differed only from clear-cuts. In contrast to our predictions, early and late successional species responded similarly to different silvicultural practices. However, there were indications of marginal assemblage differences both between thinned stands and selective felling and between thinned and old growth stands (p = 0.10). The stand volume of early decay stage wood influenced assemblage composition of early, but not late successional species. Uneven-aged silviculture maintained species assemblages similar to those of the reference and old growth stands and might therefore be a better management option when considering biodiversity conservation. [ABSTRACT FROM AUTHOR]

Uneven-aged silviculture may better maintain species assemblages associated with old-growth forests than clear felling in part due to habitat heterogeneity created by maintaining standing retention strips adjacent to harvest trails. Retention strips and harvest trails created at the time of tree removal will likely have different microclimate and may harbor different assemblages. In some cases, the resultant stand heterogeneity associated with uneven-aged silviculture may be similar to natural small-scale disturbances. For beetles, increased light and temperature as well as potential access to young vegetation and deadwood substrates present in harvset trails may harbor beetle assemblages similar to those found in natural gaps. We sampled saproxylic beetles using flight intercept traps placed in harvest corridors and retention strips in 9 replicated uneven-aged spruce stands in central Sweden. We compared abundance, species richness and composition between harvest corridors and retention strips using generalized linear models, rarefaction, permutational multivariate analysis of variance and indicator species analysis. Canopy openness doubled, mean temperature and variability in daily temperature increased and humidity decreased on harvest trails. Beetle richness and abundance were greater in harvests trails than in retention strips and the beetle species composition differed significantly between habitats. Twenty-five species were associated with harvest trails, including three old-growth specialists such as Agathidium discoideum (Erichson), currently red-listed. We observed only one species, Xylechinus pilosus (Ratzeburg) that strongly favored retention strips. Harvest trails foster both open habitat species and old-growth species while retention strips harbored forest interior specialists. The combination of closed canopy, stratified forest in the retention strips and gap-like conditions on the harvest trails thus increases overall species richness and maintains more diverse assemblages at the stand level than would otherwise be seen in less heterogeneous stand types. This suggests that uneven-aged silviculture may provide added conservation benefits for both open habitat and old-growth specialists than silvicultural approaches that reduce stand heterogeneity. [ABSTRACT FROM AUTHOR]

Large scale use of even-aged silviculture (clear-cutting) commencing in the mid-20th century has had negative impacts on forest biodiversity. As a consequence, uneven-aged silviculture is currently being considered to help meet the ecological and social criteria required for sustainable forest management. Uneven-aged silviculture (e.g. selective felling) involves selective removal of some older trees in a stand which may to some extent mimics natural small scale stand dynamics and thus potentially benefit species associated with old forests. Here we test whether selective felling benefits beetle biodiversity by producing beetle assemblages that better resemble those of old growth stands than those found in uncut production stands. We conducted a field study in northern Sweden, comparing beetles assemblages collected with window traps in three spruce dominated stand types: (1) Stands recently (on average 7 years prior to the study) subjected to selective felling ( Selective felling ), (2) mature uneven-aged stands without recent history of management, resembling selective felling stands prior to management ( Uncut ), and (3) old-growth stands with high conservation values ( Old growth ). As predicted, we found that assemblage composition was similar in selective felling and old growth stands, and that assemblages of cambivores and obligate saproxylics (marginally significant) differed between these two stand types and uncut stands. The differences were largely explained by a higher abundance of saproxylic species presumably associated with old growth conditions and large volumes of deadwood. Thus, although overall assemblage composition did not differ between stand types, part of the beetle community seemingly benefited from selective felling. We therefore recommend that selective felling is considered as an alternative to clear-felling to maintain biodiversity values. [ABSTRACT FROM AUTHOR]

Decline in biodiversity have increased the interest in alternative forest management approaches. Uneven-aged silviculture has been proposed as a mean to maintain continuity of forest canopy cover, mimic small-scale disturbances and provide a stratified forest structure similar to that of old-growth forests and therefore better maintain species associated with unmanaged forest. We used a large-scale chronosequence study spanning 50 years to study beetle diversity in uneven-aged silviculture compared with both short-term impacts and the longer-term legacy of even-aged silviculture. We compared: (1) even-aged recently clear-felled stand, (2) even-aged recently thinned stand, (3) uneven-aged stands subjected to selective felling with (4) uneven-aged reference stands to evaluate whether abundance, species richness and composition of beetles (Coleoptera) were affected differently by even-aged than by uneven-aged management. We collected 15,147 beetles from 461 species using flight interception traps in 30 stands. Beetle composition was maintained in uneven-aged managed stands; composition did not differ from unmanaged reference stands, the exception being cambium consumers. Both even-aged silviculture treatments (clear-felling and thinning) had different beetle composition compared to the reference stands, indicating that assemblages had yet to recover even 50 years into the rotation. However, beetle composition did not differ between uneven-aged managed and thinned stands. The result supports our prediction that uneven-aged silviculture better maintains beetles assemblages associated with semi-natural mature forest than even-aged silviculture. The greater temporal continuity in selectively felled stands could benefit species dependent of mature or old growth forest since some of the needed habitat qualities are continuously available. Uneven-aged silviculture could therefore serve as an important tool for landscape planning to benefit biodiversity and thus help fulfil environmental commitments. However, uneven-aged silviculture may still alter the forest and should therefore be viewed as an alternative to even-aged silviculture, rather than to set-asides. [ABSTRACT FROM AUTHOR]

Management of boreal forests for timber production has caused changes in forest structures and disturbance regimes, which have influenced a wide range of organisms. The aim of this study was to evaluate how composition of epigaeic (ground-living) beetle assemblages is influenced by stand age and management history in a heavily managed boreal forest landscape. We compared the epigaeic assemblages among stands of three ages: (1) young (8–25 years) and (2) middle-aged (40–58 years) stands regrown after clear-cutting, and mature stands (80–130 years) that had been selectively cut historically but never clear felled. We sampled epigaeic beetles in each of 42 stands, using 10 pitfall traps during seven summer weeks. More than 9000 specimens were collected and identified. The assemblages in young stands differed from those in middle-aged and mature stands, both for the Staphylinidae (rove beetles) and all beetle families combined. Carabidae (ground beetles) composition differed between young and middle aged stands only, and assemblages of Curculionidae (weevils, bark beetles and allies) differed between young and mature stands only. Assemblages of Leiodidae (round fungus beetles) had similar composition in all three stand types. Considering all families, young stands generally harbored fewer species and lower abundances compared with middle aged and mature stands. However, the Leiodidae had similar species richness in all three stand types. The lack of differences in assemblage composition, species richness and abundance between middle aged and mature stands suggests that epigaeic beetle assemblages recolonize following clear-felling. However, our collections included large numbers of unique and usually rare species in mature stands, indicating that old forest is important for the conservation of epigaeic beetles. Furthermore, the lower abundance of these beetles in young stands indicates that an increasing proportion of young stands on managed landscapes will reduce the overall abundances of epigaeic beetle species, with potentially negative impacts on recolonization. [ABSTRACT FROM AUTHOR]

Intensive management of boreal forests has replaced natural disturbances, leading to losses of residual deadwood and saproxylic biodiversity. Declining population sizes have been well documented for insect groups such as bark and wood-boring beetles, whilst other groups such as flat bugs (Heteroptera: Aradidae) are less well-studied and little is known about their response to forest restoration. We conducted two restoration experiments addressing this lack of knowledge. We predicted the following responses to restoration: (1) artificial gap-creation and prescribed burning will attract more flat bugs than control stands, (2) increasing pre-burning fuel loads will have a positive influence on flat bug response to fire restoration. To test these predictions we conducted two field experiments; in Sweden we compared prescribed burning of standing forest with artificial gap-creation and in Finland we compared differing levels of fuel-load prior to burning. We found partial support for our first prediction, total numbers of flat bugs collected was higher in burned forest stands compared to controls in both countries. By contrast, gap-cutting only resulted in a marginal increase in the numbers and species of flat bugs collected. We found no support for our second hypothesis. The red-listed Aradus angularis and A. laeviusculus were primarily found in burned stands. Aradus betulae, A. betulinus and A. lugubris were more frequent in burned stands than gap-cuttings and untreated controls. Our study therefore suggests that ecological restoration, especially burning, is an efficient way of attracting flat bugs to suitable habitat, we therefore recommend ecological restoration to be incorporated in boreal forest management. [ABSTRACT FROM AUTHOR]

The increasing demand for biofuels from logging residues require serious attention on the importance of dead wood substrates on clear-cuts for the many forestry-intolerant saproxylic (wood-inhabiting) species. In particular, the emerging harvest of low stumps motivates further study of these substrates. On ten clear-cuts we compared the species richness, abundance and species composition of saproxylic beetles hatching from four to nine year old low stumps, high stumps and logs of Norway spruce. By using emergence traps we collected a total of 2,670 saproxylic beetles among 195 species during the summers of 2006, 2007 and 2009. We found that the species assemblages differed significantly between high stumps and logs all three years. The species assemblages of low stumps, on the other hand, were intermediate to those found in logs and high stumps. There were also significant difference in species richness between the three examined years, and we found significant effect of substrate type on richness of predators and fungivores. As shown in previous studies of low stumps on clear-cuts they can sustain large numbers of different saproxylic beetles, including red-listed species. Our study does, in addition to this fact, highlight a possible problem in creating just one type of substrate as a tool for conservation in forestry. Species assemblages in high stumps did not differ significantly from those found in low stumps. Instead logs, which constitute a scarcer substrate type on clear-cuts, provided habitat for a more distinct assemblage of saproxylic species than high stumps. It can therefore be questioned whether high stumps are an optimal tool for nature conservation in clear-cutting forestry. Our results also indicate that low stumps constitute an equally important substrate as high stumps and logs, and we therefore suggest that stump harvesting is done after carefully evaluating measures to provide habitat for saproxylic organisms. [ABSTRACT FROM AUTHOR]

Highlights: [•] We review environmental considerations in forestry with Sweden as a case study. [•] Environmental considerations in forestry have support in the scientific literature. [•] Retention levels in Sweden are low compared with ecological thresholds. [•] There is a need to better adjust the levels above thresholds from empirical studies. [•] The allocation of consideration in and among landscapes needs to be improved. [ABSTRACT FROM AUTHOR]

Abstract: Intensive forest management has dramatically reduced the area of old-growth forest in Fennoscandia. We examined if the proportion of old forest in a landscape affects species composition, richness and abundance of saproxylic beetles. We used tube-shaped window traps in five pairs of sites, selected so that the sites within each pair differed with respect to the proportion of old forest (>125years) in the surrounding landscape. A landscape level inventory of the wood fungi Fomitopsis rosea, as a proxy for forest with high conservation values, was used to complement the data on old forests. In addition, to testing whether mycelia-colonised wood may attract saproxylic beetles, the tube-shaped window traps were baited with wood colonised by Fomitopsis pinicola or F. rosea. Old-forest-rich landscapes supported significantly more species and a higher abundance of saproxylic beetles than old-forest-poor landscapes. The analysis revealed a clear connection between the community composition of saproxylic beetles and the proportion of old forest and number of F. rosea fruiting bodies in the surrounding landscape (radius 3km). The local landscape species pool thus appears to be important for the beetle species that are trapped since the composition of saproxylic beetles differed between the two landscape types. The effects of the different baits were less pronounced than the effect of landscape type, although species-specific responses to the two mycelia baits were observed. This indicates that volatiles from mycelia of wood-decaying fungi and the mycelial community may affect colonisation patterns of saproxylic beetles. Our results suggest that forest fragmentation and habitat loss have resulted in depauperate beetle faunas in old-forest-poor landscapes. Our results highlight the need to invoke a landscape scale approach for preserving biodiversity, in this case the need to maintain a sufficient proportion of forest with old growth characteristics in the managed landscape. [Copyright &y& Elsevier]

1. Suppression of wildfires in boreal landscapes has become widespread and has seriously affected many fire favoured species. However, little is known about the response of organism assemblages to large wildfires in regions with a long history of effective fire suppression, such as Scandinavia. 2. We studied the short-term effects of a >1600 ha wildfire on beetle assemblages in northern Sweden. The first summer after fire, beetles were sampled in 12 sites using 36 large window traps, half in old pine forest stands in the burned area and half in similar, but unburned control stands. The entire beetle assemblage and eight subgroups were analysed: saproxylics, non-saproxylics, moderately fire favoured, strongly fire favoured, fungivores, predators, cambium consumers and red-listed species. 3. Species composition differed markedly between burned and unburned forests in all nine groups. Furthermore, beetle abundance was higher in the burned area for the entire assemblage and for saproxylics, both groups of fire favoured species, predators and cambium consumers. Species number was higher only for non-saproxylics, strongly fire favoured species and cambium consumers. 4. Our results show that wildfire has rapid and strong effects on a wide range of beetles. However, we only trapped two individuals of fire-dependent beetles, which may suggest a lack of such species in the region, possibly due to >100 years of fire suppression. At the regional scale, the studied wildfire may potentially increase the abundance of these beetles after a longer period of reproduction in the burned area. [ABSTRACT FROM AUTHOR]

Abstract: Predatory beetles are important natural enemies of early successional cambium feeding beetles and knowledge about their ecology is of paramount importance for biological control and sustainable forest management. We used a large-scale experiment including 3 stand types (clear-cut, mature managed and old-growth stand) and 6 dead wood substrates (burned, shaded, white rot inoculated, brown rot inoculated, untreated control logs and created snags) to test substrate preferences of 10 common predatory beetle species. We collected insects in eclector traps and measured dead wood availability within 100m from the experimental logs at each site. We also tested the relationships between predators and prey at the trap level. Stand type was important for all predators; Nudobius lentus and Ampedus tristis were more abundant on clear-cuts while Phloeonomus sjoebergi, Quedius plagiatus, Epuraea boreella, Epuraea pygmaea, Epuraea angustula, Rhizophagus dispar and Epuraea laeviscula were more abundant in mature managed and old-growth stands. Snags differed most from the control logs supporting significantly more E. boreella and E. angustula and significantly less P. sjoebergi, E. pygmaea, A. tristis and R. dispar. Prey abundance proved to be more important than dead wood availability at the stand level for most species, although the response varied among species. At the trap level, the abundances of P. sjoebergi, E. laeviscula and Q. plagiatus were correlated with the abundances of their known prey species. P. sjoebergi, E. boreella, E. pygmaea, R. dispar and N. lentus were correlated with total bark beetle abundance suggesting that many predators might be less specific in their choice of prey than previously thought. The variable response of the 10 examined predatory beetles at different levels of scale suggests complex relationships with their prey and environment. The conservation of this group of beetles is important both to achieve environmental goals but also for biological control of potential pest species. Thus, conservation oriented measures needs to be performed aiming at improving both substrate and stand qualities. [Copyright &y& Elsevier]

Abstract: Different functional groups of beetles are likely to exhibit different responses to forest management and it is essential that we gain knowledge on these responses to understand and minimize the negative impacts of forestry in boreal ecosystems. In a large-scale manipulative experiment in nine sites (including 27 stands) in northern Sweden, we tested how saproxylic beetles, fire-favoured beetles and functional groups of beetles were affected by log treatment (shaded, burnt and fungi inoculated logs) and stand type (clear-cuts, mature managed and old-growth stands). All sites included one clear-cut, one mature managed stand and one old-growth stand, which give in total 27 stands. Fresh logs were placed out in the stands and exposed to different treatments. Insects were caught in eclector traps attached to the experimental logs. Log treatment, stand type and their interactions affected abundance, species richness and assemblages of saproxylic beetles, fire-favoured beetles and functional groups of beetles. Cambium consumers were less abundant on burnt logs compared with control logs. Furthermore, the assemblage composition differed between burnt logs and control logs for fire-favoured species and cambium consumers. On the clear-cuts burnt logs hosted lower abundance of all saproxylic beetles and fire-favoured species. For predators, the assemblage composition differed between burnt logs and control logs on clear-cuts. Shaded logs hosted lower abundance of wood borers compared to control logs. Shaded logs also supported different assemblages of saproxylic beetles and predators compared with the control logs on clear cuts. The effect of stand type was stronger than the effect of log treatment and for most of the examined groups; the clear-cuts differed from the mature managed and old-growth stands, while few differences was detected between mature managed and old-growth stands. The results indicate that in an area with intense forestry, both old-growth forest reserves and the creation of suitable substrates in managed forests are needed to conserve functionally intact beetle assemblages. [Copyright &y& Elsevier]

Abstract: Many saproxylic beetles do not feed on wood directly but on fungi colonizing the wood. The volume of decaying wood has decreased drastically in Scandinavian managed forest landscapes in recent years, so improved knowledge on the interactions between beetles and wood-decaying fungi is important for the long-term persistence of these trophic partners. Sporocarps of polypores are known to emit volatiles attracting both fungivorous and predatory beetles, but it is unknown whether some beetles are also attracted to odours from the mycelia. The aim of this experiment was to test the attraction of beetles to volatiles from the sporocarps and mycelia of wood-decaying fungi. In a randomized block design, six substrate types: Fomitopsis pinicola sporocarp, F. pinicola mycelium-infected wood, Fomitopsis rosea sporocarp, F. rosea mycelium-infected wood, Phellinus chrysoloma sporocarp and Phlebia centrifuga mycelium-infected wood were attached separately to specially designed window traps in four old-growth spruce forests in northern Sweden. Empty traps and traps with sterilised wood were used as controls. We found no significant differences in the species richness or abundance of saproxylic beetles between the control and sterilised wood and the fungal substrates. However, two abundant species showed significant preferences for one substrate type. The bark beetle Dryocoetes autographus preferred F. rosea mycelium-infected wood and the rove beetle Lordithon lunulatus preferred fruiting bodies of F. pinicola. The results indicate that some species do discriminate between volatiles emitted by different polypore species and also between volatiles emitted by the sporocarps and mycelia from the same species. Our data indicate a hitherto unknown interdependence between D. autographus and F. rosea. We conclude that present knowledge on interactions between beetles and wood-decaying fungi is limited and further studies are needed to enhance our ability to design appropriate conservation strategies in the forest landscape. [Copyright &y& Elsevier]

Forest management practices have led to a reduction in the volume and a change in the composition of coarse woody debris (CWD) in many forest types. This study compared CWD volume and composition in reserves and two types of managed forest in the central boreal zone of Sweden. Ten areas were surveyed, each containing clear-cut, mature managed and old-growth stands, to determine the volume of standing and lying CWD in terms of species composition, decay class and size class. Volumes of CWD on clear-cuts and in mature managed forests were high compared with previous studies. Old-growth forests (72.6 m 3 ha -1 ) contained a greater volume of CWD than mature managed forests (23.3 m 3 ha -1 ) and clear-cuts (13.6 m 3 ha -1 ). Differences were greatest for the larger size classes and intermediate decay stages. Despite stand ages being up to 144 years, CWD volume and composition in managed forests was more similar to clear-cuts than to old-growth forests. [ABSTRACT FROM AUTHOR]

• Forest management has promoted conifers and altered stream and riparian function. • We suggest delineating and managing riparian buffers throughout the rotation cycle. • Make multi-layered, mixed-species forests with more deciduous species the goal. • Riparian zones will provide functions throughout the rotation, not just at final-felling. Riparian buffers are the primary tool in forest management for protecting the habitat structure and function of streams. They help protect against biogeochemical perturbation, filter sediments and nutrients, prevent erosion, contribute food to aquatic organisms, regulate light and hence water temperature, contribute deadwood, and preserve biodiversity. However, in production forests of Sweden and Finland, many headwater streams have been straightened, ditched, and/or channelized, resulting in altered hydrology and reduced natural disturbance by floods, which in turn affects important riparian functions. Furthermore, in even-aged management systems as practiced in much of Fennoscandia, understory trees have usually been cleared right up to the stream's edge during thinning operations, especially around small, headwater streams. Fire suppression has further favored succession towards shade tolerant species. In the regions within Fennoscandia that have experienced this combination of intensive management and lack of natural disturbance, riparian zones are now dominated by single-storied, native Norway spruce. When the adjacent forest is cut, thin (5 - 15m) conifer-dominated riparian buffers are typically left. These buffers do not provide the protection and subsidies, in terms of leaf litter quality, needed to maintain water quality or support riparian or aquatic biodiversity. Based on a literature review, we found compelling evidence that the ecological benefits of multi-layered, mixed-species riparian forest with a large component of broadleaved species are higher than what is now commonly found in the managed stands of Fennoscandia. To improve the functionality of riparian zones, and hence the protection of streams in managed forest landscapes, we present some basic principles that could be used to enhance the ecological function of these interfaces. These management actions should be prioritized on streams and streamside stands that have been affected by simplification either through forest management or hydrological modification. Key to these principles is the planning and managing of buffer zones as early as possible in the rotation to ensure improved function throughout the rotation cycle and not only at final felling. This is well in line with EU and national legislation which can be interpreted as requiring landscape planning at all forest ages to meet biodiversity and other environmental goals. However, it is still rare that planning for conservation is done other than at the final felling stage. Implementing this new strategy is likely to have long-term positive effects and improve the protection of surface waters from negative forestry effects and a history of fire suppression. By following these suggested management principles, there will be a longer time period with high function and greater future management flexibility in addition to the benefits provided by leaving riparian buffers at the final felling stage. [ABSTRACT FROM AUTHOR]

• Even-aged silviculture (thinning) vs uneven-aged silviculture (selective felling) • Species richness did not differ between management type. • Thinned stands supported different bird assemblages than selectively felled stands. • Independent of management type, time since treatment effect assemblage structures. Biodiversity is in decline and therefore alternative forest management approaches have gained interest. One of such approaches is uneven-aged silviculture, which has been suggested to better maintain mature or late-successional forest characteristics and species assemblages than even-aged silviculture. Therefore, it is assumed that uneven-aged silviculture can be a useful tool for landscape planning to benefit biodiversity. Nevertheless, there is a lack of empirical studies regarding bird responses to uneven-aged silviculture in north European boreal ecosystems. Here we test the similarity of bird assemblage structure between mature forests within even-aged silviculture ('thinning') and uneven-aged silviculture ('selective felling'). In spring 2018 we censused breeding birds using territory mapping in 14 thinned stands and 14 selectively felled stands. We found higher abundance and different bird assemblages in thinned stands compared to selectively felled stands. The pied flycatcher, tree pipit and great tit contributed most to the variation of bird assemblages between the two management types. None of the species were more abundant in selectively felled than in thinned stands. According to functional guilds, the abundance of ground breeders, ground feeders and generalists was higher in thinned stands than in selectively felled stands, similar results were found in the species richness of long-distance migrants, ground nesters, secondary cavity nesters and generalists. Independent of management type, time since treatment had an overall effect on assemblage structures, the mistle thrush and wren were negatively correlated with time since treatment, while the chiffchaff showed the opposite trend. Our results suggest that at these locations and given this particular type of uneven-age management, selective felling is less suitable for some abundant generalists than even-aged forest stands reaching the thinning age. However, the results from this study does not provide clear management recommendations aiming to maintain biodiversity, as management guidelines should be based on red-listed species and not on common generalists. Nevertheless, our results stresses the urgent need for more long-term studies comparing the effect of these different silvicultural strategies on bird assemblages. [ABSTRACT FROM AUTHOR]

Abstract: The habitat requirements and effects of forest management on insects belonging to higher trophic levels are relatively unknown in forest ecosystems. We tested the effect of forest successional stage and dead wood characteristics on the saproxylic parasitoid (Hymenoptera, Ichneumonoidea) assemblage in boreal spruce-dominated forests in northern Sweden. Within each of nine areas, we selected three sites with different management histories: (1) a clear-cut (2) a mature managed forest and (3) an old-growth forest. Parasitoids were collected in 2003 using eclector traps mounted on fresh logs, which were either untreated (control), burned, inoculated with fungi, or naturally shaded, and on artificially-created snags. Both forest type and dead wood characteristics had a significant effect on parasitoid assemblages. Grouped idiobionts and some species, such as Bracon obscurator and Ontsira antica, preferred clear-cuts, while others, such as Cosmophorus regius (Hym., Braconidae) and other koinobionts, were associated with older successional forest stages. No single dead wood substrate was sufficient to support the entire community of parasitoids in any forest type, even when the regular host was present. In particular, snags hosted a different assemblage of species from other types of dead wood, with parasitoids of Tetropium spp. such as Rhimphoctona spp. (Hym., Ichneumonidae) and Helconidea dentator (Hym., Braconidae) being abundant. These results indicate that a diversity of dead wood habitats is necessary to support complete assemblages of beetle-associated parasitoids from early successional stages of dead wood and that parasitoids may be more sensitive to habitat change than their hosts. [Copyright &y& Elsevier]