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The transition of sagebrush‐dominated (Artemisia spp.) shrublands to pinyon (Pinus spp.) and juniper (Juniperus spp.) woodlands markedly alters resource‐conserving vegetation structure typical of these landscapes. Land managers and scientists in the western United States need knowledge and predictive tools for assessment and effective targeting of tree‐removal treatments to conserve or restore sagebrush vegetation and associated hydrologic function. This study developed modeling approaches to quantify the hydrologic vulnerability and erosion potential of sagebrush rangelands in the later stages of woodland encroachment and in response to commonly applied tree‐removal treatments. Using experimental data from multiple sites in the Great Basin Region, USA, and process‐based knowledge from decade‐long vegetation and rainfall simulation studies at those sites, we (1) assessed the capability of the Rangeland Hydrology and Erosion Model (RHEM) to accurately predict patch‐scale (12 m2) measured runoff and erosion from tree canopy and intercanopy hydrologic functional units in untreated and burned woodlands 9 years postfire, and (2) developed and evaluated multiple RHEM approaches/frameworks to model aggregated effects of tree canopy and intercanopy areas on patch‐ and hillslope‐scale (50 m length) runoff and erosion processes in untreated and treated (burned, cut, and masticated) woodlands. The RHEM accurately predicted measured runoff and sediment yield from patch‐scale rainfall simulations as partitioned on untreated and treated tree canopy and intercanopy areas and effectively parameterized the dominant controls on runoff and erosion process in woodlands. With few exceptions, evaluated hillslope‐scale RHEM frameworks similarly predicted reduced hydrologic vulnerability and erosion potential for conditions 9 years following tree removal by burning, cutting, and mastication treatments. Regressions of RHEM‐predicted hillslope runoff, sediment, and hydraulic/erosion parameters with bare ground and ground cover attributes indicate all RHEM frameworks effectively represented the dominant controls on hydrologic and erosion processes for rangelands and woodlands. The results provide RHEM frameworks and recommendations for assessing hydrologic vulnerability and erosion potential on woodland‐encroached sites and predicting the effectiveness of tree removal to reestablish a water and soil resource‐conserving vegetation structure on sagebrush rangelands. We anticipate our RHEM or similar modeling approaches may be applicable to analogous water‐limited landscapes elsewhere subject to woody plant encroachment. [ABSTRACT FROM AUTHOR]

Wildfire is a natural component of sagebrush (Artemisia spp.) steppe rangelands that induces temporal shifts in plant community physiognomy, ground surface conditions, and erosion rates. Fire alteration of the vegetation structure and ground cover in these ecosystems commonly amplifies soil losses by wind‐ and water‐driven erosion. Much of the fire‐related erosion research for sagebrush steppe has focused on either erosion by wind over gentle terrain or water‐driven erosion under high‐intensity rainfall on complex topography. However, many sagebrush rangelands are geographically positioned in snow‐dominated uplands with complex terrain in which runoff and sediment delivery occur primarily in winter months associated with cold‐season hydrology. Current understanding is limited regarding fire effects on the interaction of wind‐ and cold‐season hydrologic‐driven erosion processes for these ecosystems. In this study, we evaluated fire impacts on vegetation, ground cover, soils, and erosion across spatial scales at a snow‐dominated mountainous sagebrush site over a 2‐year period post‐fire. Vegetation, ground cover, and soil conditions were assessed at various plot scales (8 m2 to 3.42 ha) through standard field measures. Erosion was quantified through a network of silt fences (n = 24) spanning hillslope and side channel or swale areas, ranging from 0.003 to 3.42 ha in size. Sediment delivery at the watershed scale (129 ha) was assessed by suspended sediment samples of streamflow through a drop‐box v‐notch weir. Wildfire consumed nearly all above‐ground live vegetation at the site and resulted in more than 60% bare ground (bare soil, ash, and rock) in the immediate post‐fire period. Widespread wind‐driven sediment loading of swales was observed over the first month post‐fire and extensive snow drifts were formed in these swales each winter season during the study. In the first year, sediment yields from north‐ and south‐facing aspects averaged 0.99–8.62 t ha−1 at the short‐hillslope scale (~0.004 ha), 0.02–1.65 t ha−1 at the long‐hillslope scale (0.02–0.46 ha), and 0.24–0.71 t ha−1 at the swale scale (0.65–3.42 ha), and watershed scale sediment yield was 2.47 t ha−1. By the second year post fire, foliar cover exceeded 120% across the site, but bare ground remained more than 60%. Sediment yield in the second year was greatly reduced across short‐ to long‐hillslope scales (0.02–0.04 t ha−1), but was similar to first‐year measures for swale plots (0.24–0.61 t ha−1) and at the watershed scale (3.05 t ha−1). Nearly all the sediment collected across all spatial scales was delivered during runoff events associated with cold‐season hydrologic processes, including rain‐on‐snow, rain‐on‐frozen soils, and snowmelt runoff. Approximately 85–99% of annual sediment collected across all silt fence plots each year was from swales. The high levels of sediment delivered across hillslope to watershed scales in this study are attributed to observed preferential loading of fine sediments into swale channels by aeolian processes in the immediate post‐fire period and subsequent flushing of these sediments by runoff from cold‐season hydrologic processes. Our results suggest that the interaction of aeolian and cold‐season hydrologic‐driven erosion processes is an important component for consideration in post‐fire erosion assessment and prediction and can have profound implications for soil loss from these ecosystems. © 2019 John Wiley & Sons, Ltd. [ABSTRACT FROM AUTHOR]

Abstract Land managers across the western United States are faced with selecting and applying tree-removal treatments on pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland-encroached sagebrush (Artemisia spp.) rangelands, but current understanding of long-term vegetation and hydrological responses of sagebrush sites to tree removal is inadequate for guiding management. This study applied a suite of vegetation and soil measures (0.5 − 990 m2), small-plot rainfall simulations (0.5 m2), and overland flow experiments (9 m2) to quantify the effects of mechanical tree removal (tree cutting and mastication) on vegetation, runoff, and erosion at two mid- to late-succession woodland-encroached sagebrush sites in the Great Basin, United States, 9 yr after treatment. Low amounts of hillslope-scale shrub (3 − 15%) and grass (7 − 12%) canopy cover and extensive intercanopy (area between tree canopies) bare ground (69 − 88% bare, 75% of area) in untreated areas at both sites facilitated high levels of runoff and sediment from high-intensity (102 mm • h− 1, 45 min) rainfall simulations in interspaces (~ 45 mm runoff, 59 − 381 g • m− 2 sediment) between trees and shrubs and from concentrated overland flow experiments (15, 30, and 45 L • min− 1, 8 min each) in the intercanopy (371 − 501 L runoff, 2 342 − 3 015 g sediment). Tree cutting increased hillslope-scale density of sagebrush by 5% and perennial grass cover by twofold at one site while tree cutting and mastication increased hillslope-scale sagebrush density by 36% and 16%, respectively, and perennial grass cover by threefold at a second more-degraded (initially more sparsely vegetated) site over nine growing seasons. Cover of cheatgrass (Bromus tectorum L.) was < 1% at the sites pretreatment and 1 − 7% 9 yr after treatment. Bare ground remained high across both sites 9 yr after tree removal and was reduced by treatments solely at the more degraded site. Increases in hillslope-scale vegetation following tree removal had limited impact on runoff and erosion for rainfall simulations and concentrated flow experiments at both sites due to persistent high bare ground. The one exception was reduced runoff and erosion within the cut treatments for intercanopy plots with cut-downed-trees. The cut-downed-trees provided ample litter cover and tree debris at the ground surface to reduce the amount and erosive energy of concentrated overland flow. Trends in hillslope-scale vegetation responses to tree removal in this study demonstrate the effectiveness of mechanical treatments to reestablish sagebrush steppe vegetation without increasing cheatgrass for mid- to late-succession woodland-encroached sites along the warm-dry to cool-moist soil temperature − moisture threshold in the Great Basin. Our results indicate improved hydrologic function through sagebrush steppe vegetation recruitment after mechanical tree removal on mid- to late-succession woodlands can require more than 9 yr. We anticipate intercanopy runoff and erosion rates will decrease over time at both sites as shrub and grass cover continue to increase, but follow-up tree removal will be needed to prevent pinyon and juniper recolonization. The low intercanopy runoff and erosion measured underneath isolated cut-downed-trees in this study clearly demonstrate that tree debris following mechanical treatments can effectively limit microsite-scale runoff and erosion over time where tree debris settles in good contact with the soil surface. [ABSTRACT FROM AUTHOR]

Increasing trends in wildfire activity on semi-arid rangelands necessitate advancement in understanding of fire impacts on vegetation, soils, and runoff and erosion processes. This study used artificially applied rainfall and concentrated overland flow experiments to evaluate the ecohydrologic response and recovery of a semi-arid shrubland in the Great Basin Region, USA, following fire. Rainfall experiments were conducted at the 0.5 m 2 plot scale to assess fire impacts on rainsplash and sheetflow processes. Concentrated flow experiments were applied on 9 m 2 plots to evaluate fire impacts on concentrated overland flow processes. Vegetation, soil, hydrologic, and erosion variables were assessed at each scale pre-fire and 1, 2, and 5 yr post-fire. Infiltration and runoff on rainfall simulation plots were affected more by measured background soil water repellency than fire effects on vegetation and soils. Runoff from rainfall on shrub-dominated plots was unchanged 1 yr post-fire, but runoff from interspace plots between shrubs declined 1 yr post-fire. Runoff increased on shrub and interspace rainfall plots 2 yr post-fire and then declined in the 5 yr post-fire. Bare ground generally declined across study years, implicating the temporal variability in soil water repellency as the causal factor for infiltration and runoff trends. Erosion on rainfall plots increased by factors of 8 to more than 10 following fire removal of vegetation and ground cover and declined with vegetation recovery through five growing seasons. Concentrated overland flow plots generated slightly more total runoff and 26-fold more total sediment 1 yr following burning relative to pre-fire measures. Erosion from concentrated overland flow remained greater on burned than unburned plots after five growing seasons even though ground cover returned to approximately 85%. The relative recovery of vegetation and total ground cover were typical for the shrubland community assessed, but elevated erosion with 85% ground cover 5 yr post-fire was unexpected. The persistent high sediment delivery from concentrated plots is attributed to the fine textured soils and thin litter accumulation. The importance of considering erodibility in context with sediment supply and vegetative recovery is discussed. The results demonstrate the complexity of post-fire ecohydrologic interactions, advance process understanding of post-fire ecohydrologic responses for semi-arid rangelands, and underscore the need for additional studies on post-fire recovery over time. [ABSTRACT FROM AUTHOR]

Hydrologic response to rainfall on fragmented or burnt hillslopes is strongly influenced by the ensuing connectivity of runoff and erosion processes. Yet cross-scale process connectivity is seldom evaluated in field studies owing to scale limitations in experimental design. This study quantified surface susceptibility and hydrologic response across point to hillslope scales at two degraded unburnt and burnt woodland sites using rainfall simulation and hydrologic modelling. High runoff (31-47 mm) and erosion (154-1893 g m-2) measured at the patch scale (13 m²) were associated with accumulation of fine-scale (0.5-m²) splash-sheet runoff and sediment and concentrated flow formation through contiguous bare zones (64-85% bare ground). Burning increased the continuity of runoff and sediment availability and yield. Cumulative runoff was consistent across plot scales whereas erosion increased with increasing plot area due to enhanced sediment detachment and transport. Modelled hillslope-scale runoff and erosion reflected measured patch-scale trends and the connectivity of processes and sediment availability. The cross-scale experiments and model predictions indicate the magnitude of hillslope response is governed by rainfall input and connectivity of surface susceptibility, sediment availability, and runoff and erosion processes. The results demonstrate the importance in considering cross-scale structural and functional connectivity when forecasting hydrologic and erosion responses to disturbances. [ABSTRACT FROM AUTHOR]

Pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland encroachment into sagebrush (Artemisia spp.) steppe communities throughout western North America has substantially altered the vegetation structure and hydrologic function of one of the most ecologically important rangeland ecosystems in the world. Various pinyon and juniper tree removal practices are employed to re-establish sagebrush steppe vegetation and an associated resource-conserving ecohydrologic function. The effectiveness of these practices is highly variable owing to the vast domain in which woodland encroachment occurs, climate fluctuations, differences in treatment applications, and myriads of pre-treatment conditions and post-treatment land uses. This study evaluated the long-term (13 years post-treatment) effectiveness of prescribed fire and mechanical tree removal to re-establish sagebrush steppe vegetation and associated spatial patterns in ground surface conditions and soil hydrologic properties of two woodland-encroached sites. Specifically, we assessed the effects of tree removal on: (1) vegetation and ground cover at the hillslope scale (990 m2 plots) and (2) associated spatial patterns in point-scale ground surface conditions and soil hydrologic properties along transects extending from tree bases and into the intercanopy areas between trees. Both sites were in mid to late stages of woodland encroachment with extensive bare conditions (~60–80% bare ground) throughout a degraded intercanopy area (~75% of the domain) surrounding tree islands (~25% of domain, subcanopy areas). All treatments effectively removed mature tree cover and increased hillslope vegetation. Enhanced herbaceous cover (4–15-fold increases) in burned areas reduced bare interspace (bare area between plants) by at least 4-fold and improved intercanopy hydraulic conductivity (> than 2-fold) and overall ecohydrologic function. Mechanical treatments retained or increased sagebrush and generally increased the intercanopy herbaceous vegetation. Intercanopy ground surface conditions and soil hydrologic properties in mechanical treatments were generally similar to those in burned areas but were also statistically similar to the same measures in untreated areas in most cases. This suggests that vegetation and ground surface conditions in mechanical treatments are trending toward a significantly improved hydrologic function over time. Treatments had limited impact on soil hydrologic properties within subcanopy areas; however, burning did reduce the soil water repellency strength and the occurrence of strong soil water repellency underneath trees by three- to four-fold. Overall, the treatments over a 13-year period enhanced the vegetation, ground surface conditions, and soil hydrologic properties that promote infiltration and limit runoff generation for intercanopy areas representing ~75% of the area at the sites. However, ecological tradeoffs in treatment alternatives were evident. The variations in woodland responses across sites, treatments, and measurement scales in this long-term study illustrate the complexity in predicting vegetation and hydrologic responses to tree removal on woodland-encroached sagebrush sites and underpin the need and value of multi-scale long-term studies. [ABSTRACT FROM AUTHOR]

Pinyon (Pinus spp.) and juniper (Juniperus spp.) woodland encroachment has imperiled a broad ecological domain of the sagebrush steppe (Artemisia spp.) ecosystem in the Great Basin Region, USA. As these conifers increase in dominance on sagebrush rangelands, understory vegetation declines and ecohydrologic function can shift from biotic (vegetation) controlled retention of soil resources to abiotic (runoff) driven loss of soil resources and long-term site degradation. Scientists, public land management agencies, and private land owners are challenged with selecting and predicting outcomes to treatment alternatives to improve ecological structure and function on these rangelands. This study is the first of a two-part study to evaluate effectiveness of prescribed fire to re-establish sagebrush steppe vegetation and improve ecohydrologic function on mid- to late-succession pinyon-and juniper-encroached sagebrush sites in the Great Basin. We used a suite of vegetation and soil measures, small-plot (0.5 m2) rainfall simulations, and overland flow experiments (9 m2) to quantify the effects of tree removal by prescribed fire on vegetation, soils, and rainsplash, sheetflow, and concentrated flow hydrologic and erosion processes at two woodlands 9-yr after burning. For untreated conditions, extensive bare interspace (87% bare ground) throughout the degraded intercanopy (69–88% bare ground) between trees at both sites promoted high runoff and sediment yield from combined rainsplash and sheetflow (~45 mm, 59–381 g m−2) and concentrated flow (371–501 L, 2343–3015 g) processes during high intensity rainfall simulation (102 mm h−1, 45 min) and overland flow experiments (15, 30, and 45 L min−1, 8 min each). Burning increased canopy cover of native perennial herbaceous vegetation by >5-fold, on average, across both sites over nine growing seasons. Burning reduced low pre-fire sagebrush canopy cover (<1% to 14% average) at both sites and sagebrush recovery is expected to take >30 yr. Enhanced herbaceous cover in interspaces post-fire reduced runoff and sediment yield from high intensity rainfall simulations by >2-fold at both sites. Fire-induced increases in herbaceous canopy cover (from 34% to 62%) and litter ground cover (from 15% to 36%) reduced total runoff (from 501 L to 180 L) and sediment yield (from 2343 g to 115 g) from concentrated flow experiments in the intercanopy at one site. Sparser herbaceous vegetation (49% cover) and litter cover (8%) in the intercanopy at the other, more degraded site post-fire resulted in no significant reduction of total runoff (371 L to 266 L) and sediment yield (3015 g to 1982 g) for concentrated flow experiments. Areas underneath unburned shrub and tree canopies were well covered by vegetation and ground cover and generated limited runoff and sediment. Fire impacts on vegetation, ground cover, and runoff and sediment delivery from tree and shrub plots were highly variable. Burning litter covered areas underneath trees reduced perennial herbaceous vegetation and increased invasibility to the fire-prone annual cheatgrass (Bromus tectorum L.). Cheatgrass cover increased from <1% pre-fire to 16–30%, on average, post-fire across the sites and was primarily restricted to areas around burned trees. High herbaceous cover (73%) under burned trees at the less degraded site resulted in similar low total runoff and sediment from concentrated flow experiments as pre-fire (136–228 L, 204–423 g). In contrast, fire-reduction of litter (from 79% to 49%) resulted in increased total runoff (from 103 L to 333 L) and sediment yield (from 619 g to 2170 g) from concentrated flow experiments in burned tree areas at the more degraded site. The experimental results demonstrate pinyon and juniper removal by prescribed fire can effectively re-establish a successional trajectory towards sagebrush steppe vegetation structure and thereby improve ecohydrologic function. Responses to burning at the more degraded site suggest results should be interpreted with caution however. Although burning substantially increased perennial grass cover and reduced fine-scale runoff and erosion at the more degraded site, poor sagebrush recovery, delayed litter recruitment, and persistent high concentrated flow erosion at that site suggest not all sites are good candidates for prescribed fire treatments. Furthermore, high levels of cheatgrass in burned tree areas (~30% of area) at both sites increases wildfire risk, but cheatgrass is expected to decline over time in absence of fire. Our results in context with the literature suggest fire-surrogate tree-removal treatments (e.g., tree cutting or shredding) may be more appropriate on degraded sites with limited pre-treatment sagebrush and perennial herbaceous vegetation and that seeding may be necessary to improve post-fire establishment of sagebrush steppe vegetation structure and associated ecohydrologic function under these conditions. Lastly, vegetation, runoff, and erosion responses in this study are not directly applicable outside of the Great Basin, but similar responses in woodland studies from the southwestern US suggest potential application of results to woodlands in that region. The concept of re-establishing vegetation structure to improve ecohydrologic function is broadly applicable to sparsely vegetated lands around the World. • Burning increased grass in the bare intercanopy at two woodlands 9 yr post-fire. • Grass increases improved infiltration and erosion in degraded interspaces (0.5 m2). • Litter regulated overland flow whereas runoff and flow velocity controlled erosion. • Increased grass and litter reduced intercanopy (9 m2) runoff and erosion at 1 site. • More degraded initial conditions limited ecological improvements at a 2nd site. [ABSTRACT FROM AUTHOR]

Encroachment of sagebrush (Artemisia spp.) shrublands by pinyon (Pinus spp.) and juniper (Juniperus spp.) conifers (woodland encroachment) induces a shift from biotic‐controlled resource retention to abiotic‐driven loss of soil resources. This shift is driven by a coarsening of the vegetation structure with increasing dominance of site resources by trees. Competition between the encroaching trees and understory vegetation for limited soil and water resources facilitates extensive bare intercanopy area between trees and concomitant increases in run‐off and erosion that, over time, propagate persistence of the shrubland‐to‐woodland conversion. We evaluated whether tree removal by burning can decrease late‐succession woodland ecohydrologic resilience by increasing vegetation and ground cover over a 9‐year period after fire and whether the soil erosion feedback on late‐succession woodlands is reversible by burning. To address these questions, we employed a suite of vegetation and soil measurements and rainfall simulation and concentrated overland flow experiments across multiple plot scales on unburned and burned areas at two sagebrush sites in the later stages of woodland succession. Prior to burning, tree cover was approximately 28% at the sites, and more than 70% of the area at the sites was intercanopy with depauperate understory vegetation and extensive bare ground (52–60% bare soil and rock). Burning initially increased bare ground across fine (<1 m2) to patch (tens of metres) scales, resulting in enhanced sediment availability at the fine scale, sustained high run‐off and erosion within degraded intercanopies, amplified run‐off and erosion from tree canopy areas, and amplified sediment delivery across fine to patch scales. However, fire‐induced increases in grass cover over nine growing seasons improved infiltration, limited run‐off and sediment delivery from the fine scale, and reduced intercanopy run‐off and erosion at the patch scale. These changes reflect a switch in vegetation structure, triggered by burning and subsequent vegetation re‐establishment, and a shift to biotic control on run‐off and erosion across spatial scales. The responses and persistence over the 9‐year period postfire at the two sites demonstrate that fire can decrease woodland ecohydrologic resilience by altering plant community physiognomy and thereby can reverse the soil erosion feedback on sagebrush shrublands in the later stages of woodland encroachment. [ABSTRACT FROM AUTHOR]

Pinyon (Pinus spp.) and juniper (Juniperus spp.) woodlands are an important vegetation type in the Great Basin, Colorado Plateau, and southwestern desert regions of the western US that is undergoing substantial changes associated with land management, altered disturbance regimes, and climate change. We synthesized literature on the ecohydrologic impacts of pinyon and juniper tree reductions across plot to watershed scales, short- and long-term periods, and regional climatic gradients. We found that the initial plot- to hillslope-scale ecohydrologic and erosion impacts of tree reduction on pinyon and juniper woodlands by fire, mechanical tree removal, or drought depend largely on: (1) the degree to which these perturbations alter vegetation and ground cover structure, (2) initial conditions, and (3) inherent site attributes. Fire commonly imparts an initial increased risk for hillslope runoff and erosion that degrades over time with vegetation and ground cover recovery whereas tree reductions by mechanical means pose fewer initial negative ecohydrologic impacts. Tree reduction by either approach can enhance understory vegetation and improve site-level ecohydrologic function over time, particularly on sites with an initially favorable cover of native herbaceous vegetation and a cool-season precipitation regime. Understory vegetation and ground cover enhancements appear to increase ecohydrologic resilience of some woodland communities to disturbances such as drought, fire, and insect infestations. In contrast, intensive land use, prolonged drought or repeated burning associated with invasions of fire-prone grasses can propagate long-term site degradation through persistent elevated runoff and erosion rates. Our synthesis suggests the annual precipitation requirement for increases in plot- to hillslope-scale soil water availability for herbaceous enhancement through tree removal likely ranges from 200–400 mm for sites in the Great Basin and northern Colorado Plateau (cool-season precipitation regimes), and, although suggested with great uncertainty, likely exceeds 400 mm for woodlands with rain-dominated precipitation regimes in the southwestern US. Overall, literature is inconclusive regarding tree reduction impacts on watershed-scale changes in groundwater and streamflow. To date, there is little evidence that drought-related changes to vegetation in pinyon and juniper woodlands substantially affect watershed-scale water availability and streamflow at the annual time scale. Our synthesis identifies key knowledge gaps to overcome in improving understanding of the ecohydrologic and erosion impacts of broadly occurring pinyon and juniper tree reductions in the western US. [ABSTRACT FROM AUTHOR]

Runoff and erosion estimates are needed for rangeland management decisions and evaluation of ecosystem services derived from rangeland conservation practices. The information on the effect of scale on runoff and erosion and on the choice of runoff and erosion predictors remains scarce. The objective of this work was to evaluate the effect of scale on the selection of runoff and erosion predictors with the data from rich National Range Study database containing data from 444 coupled large (3.05 m by 9.1 m) and small (0.61 m by 1.22 m) plot field experiments. With data from both plot sizes, we assessed the usefulness of adding site-specific soil surface information to basic soil and rainfall data in order to estimate runoff and erosion in rangelands. We observed the scale-dependence of the runoff coefficient and the sediment yield. Smaller values of both variables were found at large plots as compared to small plots. Regression trees were used to build predictive relationships and evaluate the relative importance of predictors. Rainfall and basic soil properties were identified as the major predictors of runoff coefficients and sediment yields at both scales. Differences in the importance of predictors were observed between the two plot sizes and between predictions of runoff and sediment yield at the same plot sizes. The antecedent soil water content was not as important as rainfall parameters. Overall, including site-specific soil surface properties did not improve the predictability of the runoff coefficient and the sediment yield. The difference in runoff and sediment yield between small and large plots was found most likely because the small plots only contained a single soil/vegetation expression, whereas there was a matrix of vegetation clumps and bare interspaces arranged in a non-uniform pattern at the large plots. The variability of runoff and sediment yield may depend on how the latter pattern expresses itself in each of the large plots. More research or a different approach is required to account for vegetation-driven spatial hydrologic processes and their influence on rangeland runoff and soil erosion processes. [ABSTRACT FROM AUTHOR]

• Prescribed fire promoted herbaceous growth in bare interspaces. • Litter enhanced infiltration and provided protection against erosion. • Increased vegetation cover altered flow paths and reduced runoff and soil loss. • Reduced soil surface connectivity post-fire improved hydrologic function. • Hydrologic function continues to improve 9 yr after fire. Woody species encroachment into herbaceous and shrub-dominated vegetations is a concern in many rangeland ecosystems of the world. Arrival of woody species into affected rangelands leads to changes in the spatial structure of vegetation and alterations of biophysical processes. In the western USA, encroachment of pinyon (Pinus spp.) and juniper (Juniperus spp.) tree species into sagebrush steppes poses a threat to the proper ecohydrological functioning of these ecosystems. Prescribed fire has been proposed and used as one rangeland improvement practice to restore sagebrush steppe from pinyon-juniper encroachment. Short-term effects of burning on the ecohydrologic response of these systems have been well documented and often include a period of increased hydrologic and erosion vulnerability immediately after burning. Long-term ecohydrologic response of sagebrush steppe ecosystems to fire is poorly understood due to lack of cross-scale studies on treated sites. The aim of this study is to evaluate long-term vegetation, hydrologic, and erosion responses at two pinyon-juniper-encroached sagebrush sites 9 years after prescribed fire was applied as a restoration treatment. Thirty-six rainfall simulation experiments on 6 m × 2 m plots were conducted for 45 min under two conditions: a dry run (70 mm h−1; dry antecedent soils) and a wet run (111 mm h−1; wet antecedent soils). Runoff and erosion responses were compared between burned and unburned plots. Overall, increases in herbaceous cover in the shrub-interspace areas (intercanopy area between trees) at both sites 9 years post-burn resulted in runoff- and erosion-reduction benefits, especially under the wet runs. While the initially more degraded site characterized by 80% bare ground pre-burn, registered a higher overall increase (40% increase) in canopy cover, greater post-fire reductions in runoff and erosion were observed at the less degraded site (57% bare ground pre-burn). Runoff and erosion for the wet runs decreased respectively by 6.5-fold and 76-fold at the latter site on the burned plots relative to control plots, whereas these decreases were more muted at the more degraded site (2.5 and 3-fold respectively). Significant fragmentation of flow paths observed at the more-degraded site 9 years post-fire, suggests a decreased hydrologic connectivity as a mechanism of runoff and erosion reduction during post-fire recovery. [ABSTRACT FROM AUTHOR]