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Putaoa titanoverpa new species (ĦṜƥgḓ) Figures 3, 4, 5D–F, 6C, D Holotype ♁ (HNU284): CHINA, Hunan Province, Liuyang City, Daweishan Mountain, 28°26.09'N, 114°10.09'E, 530m, 17 I 2018, Keke Liu, Luyu Wang, Guchun Zhou. Paratypes: 1♁ (HNU285) 4♀ (HNU286–289), same data as the holotype. Additional specimens examined: 8♀ (HNU290–297), same data as the types. Etymology. The species epithet refers to the robust and strongly sclerotized embolus and is derived from the Greek word titan (in the sense of large size) and the Latin word verpa (penis). Diagnosis. Males of this new species are similar to those of Putaoa huaping Hormiga and Tu, 2008 in having “U” shaped and sclerotized embolus (compare Figs 4A, 6D with fig. 3C in Hormiga and Tu, 2008), but can be distinguished by having the palpal tibia with one distal macroseta (cuspule-like), and the suprategulum with a lamelliform distal suprategular apophysis, while in P. huaping the palpal tibia has several distal macrosetae and the suprategulum has a stronger distal suprategular apophysis (compare Figs 4A–D, 6C, D with fig. 3 in Hormiga and Tu 2008). The diagnosis of the females of this new species is given under that of Putaoa annulata n. sp. Description. Male (holotype; HNU284, Figs 3A, 4). Total length 9.68. Cephalothorax 4.33 long, 3.56 wide. Sternum 2.22 long, 1.99 wide. Abdomen 5.07 long, 3.69 wide. Anterior median eye diameter 0.17. Clypeus height 1.5 times one anterior median eye diameter. Carapace with shallow and longitudinal fovea (Fig. 3A). Chelicerae with six prolateral and six retrolateral teeth (Fig. 5D); stridulatory striae absent. Legs annulated with spines and with annulations alternating brown and black. Leg measurements: I 15.06 (4.25, 1.45, 3.78, 3.81, 1.77), II 13.68 (4.04, 1.34, 3.27, 3.45, 1.58), III 10.68 (3.20, 1.19, 2.47, 2.56, 1.26), IV 13.16 (3.83, 1.20, 3.18, 3.55, 1.40). All metatarsal trichobothria 0.33 approximately (TmI–IV), located on the dorsum near the retrolateral surface. Abdomen with white spots in anterior part and irregular patches dorsally. Palp: tibia short, with one dorsal and two retrolateral trichobothria and one distal seta prolaterally (Figs 4A, C, D, 6C, D). Cymbium with a conical ectal process in medial retrolateral margin (Figs 4B, 6C), apically pointed. Paracymbium glabrous, concave and bowl-like, with apical end broad and nearly square in ventral view, connected to cymbial base by means of a membrane, ventral to a sclerotized cymbial apophysis (Figs 4B–D, 6C). Tegulum large and pointed apically, with tegular base connecting to suprategulum which is divided into three processes, embolus, embolic process and distal suprategular apophysis (Figs 4C, E, 6D). The embolus “U” shaped (especially in prolateral view), developed and sclerotized, flattening in distal half part (Figs 4A, C, E, 6D). Embolic process thin and small, close to embolus apex (Figs 4C, E, 6D). Distal suprategular apophysis short and spiculate, curved in retrolateral view, extending towards end of tegulum across the gap between bulb and cymbium (Figs 4C, D, 6C, D). Conductor triangular, membranous, pointing apically end of tegulum (Figs 4D, 6D). Median apophysis absent. Female (paratype, HNU287). Total length 9.85 (Fig. 3B). Cephalothorax 4.06 long, 3.01wide. Sternum 2.27 long, 2.08 wide (Fig. 4D). Abdomen 6.86 long, 5.13 wide. Anterior median eye diameter 0.18. Clypeus height 1.4 times one anterior median eye diameter. Chelicerae with five prolateral and five retrolateral teeth (Fig. 5E); stridulatory striae absent. Legs annulated with spines and with annulations alternating brown and black. Leg measurements: I 15.68 (4.44, 1.50, 3.89, 3.97, 1.88), II 14.62 (4.18, 1.54, 3.60, 3.60, 1.70), III 11.57 (3.38, 1.35, 2.68, 2.73, 1.40), IV 13.96 (4.10, 1.34, 3.46, 3.55, 1.51). All metatarsal trichobothria 0.33 approximately (TmI–IV), located on the dorsum near the retrolateral surface. Similar to male in somatic features. Epigynum: basal plate slightly forming an obtuse triangle, and with the median area of posterior margin protruding caudally (Figs 3E, F, 5F). Atrium divided in two by septum. Copulatory openings visible ventrally (Fig. 3E). Copulatory ducts short and thick. Spermathecae spherical, almost in contact with each other. Fertilization ducts medially oriented, arising from the middle of spermathecae (Figs 3F, 5F). Distribution. China (Hunan) (Fig. 8).

Putaoa annulata new species (Ịƥgḓ) Figures 1, 2, 5A–C, 6A, B, 7B Holotype ♁ (HNU250): CHINA, Hunan Province, Hengyang City, Hengshan Mountain, Banshanting, 26°16.73'N, 112°42.49'E, 620m, 17 X 2015, Ailan He, Bing Zhou, Wang Liu, Chen Zeng, Zhuo’er Chen. Paratypes: 1♁ (HNU392) 1♀ (HNU251), same data as the holotype; 1♁ (HNU252) 1♀ (HNU253), CHINA, Hunan Province, Shaoyang City, Xinning County, Tianyixiang, 26°21.22'N, 110°48.25'E, 586m, 21 XI 2014, Haiqiang Yin, Cheng Wang, Jiahui Gan, Bing Zhou, Yuhui Gong; 2♁ (HNU255, HNU256) 2♀ (HNU257, HNU258), CHINA, Hunan Province, Hengyang City, Hengshan Mountain, Wanshou village, 27°31.51'N, 112°10.63'E, 487m, 20 X 2015, Ailan He, Bing Zhou, Wang Liu, Chen Zeng, Zhuo’er Chen; 2♀ (HNU430, HNU431), CHINA, Hunan Province, Zhangjiajie City, Wulingyuan District, Suoxiyu Town, portal of the Laoxiyu Tunnel, 29°17.33'N, 110°29.08'E, 456m, 23 IIII 2016, Wang Liu, Chen Zeng, Tian Tian; 1♁ (HNU479) 3♀ (HNU712–714), CHINA, Guangxi Zhuang Autonomous Region, Laibin City, Jinxiu Yao Autonomous County, Dayaoshan Mountain, Shengtangshan scenic area, 23°59.12'N, 110°8.23'E, 12 X 2021, Ailan He, Jinxin Liu, Zongguang Huang, Rongrong Liao, Qing Li, Xuemei Yang, Yingli Wen, Zhao Ye, Yang Liu. Other specimens examined.: 2♁ (HNU 489, HNU 490), CHINA, Hunan Province, Hengshan City, Hengshan Mountain, 14 XI 2007, Xiaoqi Mi [no more information in the label]; 1♁ (HNU691) 1♀ (HNU692), CHINA, Hunan Province, Hengshan City, Hengshan Mountain, 28 VII 2007, Guo Tang, Xiaoqi Mi [no more information in the label]; 3♁ (HNU264–266) 13♀ (HNU267–279), CHINA, Hunan Province, Chenzhou City, Mangshan Mountain, Hejiawan, 24°55.58'N, 112°50.35'E, 640m, 11 XII 2017, Haiqiang Yin, Ailan He, Jinxin Liu; 1♁ (HNU280) 2♀ (HNU281, HNU282), CHINA, Hunan Province, Chenzhou City, Mangshan Mountain, Yanziyan, 25°2.31'N, 112°47.12'E, 390m, 10 XII 2017, Haiqiang Yin, Ailan He, Jinxin Liu; 1♀ (HNU283), CHINA, Hunan Province, Chenzhou City, Mangshan Mountain, Xiangsikeng, 24°56.98'N, 112°59.33'E, 1300m, 8 XII 2017, Haiqiang Yin, Ailan He, Jinxin Liu; aftermentioned specimens are collected in Langshan Mountain, Xinning County, Shaoyang City, Hunan Province, China, by Haiqiang Yin, Cheng Wang, Jiahui Gan, Bing Zhou and Yuhui Gong: 5♁ (HNU403–407) 6♀ (HNU254, HNU408–412), Tianyixiang, 26°21.22'N, 110°48.25'E, 586m, 21 XI 2014; 1♁ (HNU422), Feiliandong, 26°21.48'N, 110°47.92'E, 403m, 23 XI 2014; 1♁ (HNU425) 3♀ (HNU426–428), Tianyixiang, 26°21.45'N, 110°48.19'E, 556m, 25 XI 2014; 1♀ (HNU423), Zixiadong, 26°23.13'N, 110°48.46'E, 465m, 26 XI 2014; 1♀ (HNU424), Lajiaofeng, 26°23.14'N, 110°48.46'E, 653m, 27 XI 2014; 1♀ (HNU393), Luotuofeng, 26°20.47'E, 110°46.06'E, 482m, 27 XI 2014. The following specimens were collected in Hengshan Mountain, Hengyang City, Hunan Province, China, by Ailan He, Bing Zhou, Wang Liu, Chen Zeng, Zhuo’er Chen: 1♁ (HNU260) 2♀ (HNU259, HNU261), Wanshou village, 27°31.51'N, 112°10.63'E, 487m, 20 X 2015; 3♁ (HNU394–396) 7♀ (HNU262, HNU397–402), Wanshou Village, 27°31.51'N, 112°10.63'E, 487m, 19 X 2015; 1♀ (HNU413), Martyry (Zhonglieci), 27°15.96'N, 112°42.67'E, 427m, 19 X 2015; 1♁ (HNU420) 1♀ (HNU421), Depository of Buddhist texts (Cangjingge), 27°16.06'N, 112°41.53'E, 999m, 21 X 2015; 4♁ (HNU414– 417) 1♀ (HNU418), Martyry (Zhonglieci), 27°31.58'N, 112°21.63'E, 487m, 21 X 2015. Etymology. The species epithet is derived from the Latin word for ring (annulus), in reference to the proximal half of the embolus which in prolateral view resembles a ring. Diagnosis. Males of this new species can be distinguished from those of Putaoa seediq Hormiga and Dimitrov, 2017 by having the distal suprategular apophysis more developed and lobate, and the embolic process robust and sclerotized, while the distal suprategular apophysis is small and spiculate and the embolic process thin and membranous in P. seediq (compare Figs 2A–D, 6A, B with figs 2C, D, 3B, D in Hormiga and Dimitrov (2017)). Females of this new species resemble those of Putaoa titanoverpa n. sp. in having a flat basal epigynal plate, but can be distinguished by having the posterior margin of the basal plate nearly straight, and the copulatory ducts long and curved, while the medial posterior margin of the basal plate protruded towards apically to form an obtuse triangle, and the copulatory ducts are short and thick in P. titanoverpa n. sp. (compare Figs 1E–G, 5C with Figs 3E, F, 5F). Description. Male (holotype; HNU250, Figs 1A, 2). Total length 9.12. Cephalothorax 4.94 long, 3.82 wide. Sternum 2.16 long, 1.92 wide. Abdomen 4.40 long, 3.38 wide. Anterior median eye diameter 0.18. Clypeus height 1.5 times one anterior median eye diameter. Carapace with shallow and longitudinal fovea (Fig. 1A). Chelicerae with four prolateral and four retrolateral teeth (Fig. 5A); stridulatory striae absent. Legs annulated with spines and with annulations alternating brown and black. Leg measurements: I 15.26 (4.33, 1.46, 3.71, 3.89, 1.87), II 14.01 (4.12, 1.39, 3.38, 3.51, 1.61), III 1.16 (3.33, 1.25, 2.58, 2.69, 1.31), IV 13.60 (3.94, 1.26, 3.28, 3.66, 1.46). Metatarsus I and II trichobothria (TmI and TmII) 0.25 approximately; Metatarsus III and IV trichobothria (TmIII and TmIV) 0.33 approximately.All metatarsal trichobothria located on the dorsum, near the retrolateral surface. Abdomen with some large brown patches and a few of white spots dorsally. Palp: tibia short, with three retrolateral trichobothria, one distal prolateral seta and one thick and short dorsal macroseta retrolaterally (Figs 2C, D, 6A, B). Cymbium with a conical ectal process (Figs 2B, 6A), apically pointed, and two basal cymbial apophyses near the edge of paracymbium (Figs 2E, 6A). Paracymbium glabrous, concave and bowl-like, apically hooked, with edge of median margin curved inward (Figs 2E, 6A), connected to cymbial base by means of a membrane, ventral to a bipartite basal cymbial apophysis. Tegulum large and pointed apically, and with base connecting to suprategulum which is prolonged and divided into three processes, embolus, embolic process and distal suprategular apophysis (Figs 2C, 6B). Embolus sinuous, well developed, with proximal half ring-like, flat and wide, and distal half gradually becoming narrow and somewhat sharp (Figs 2A, 6A). Embolic process sclerotized, pointed apically, running parallel to embolus end (Figs 2A, C, 6B). Distal suprategular apophysis sclerotized and lobate, almost in contact with apicoventral area of cymbium, extending towards tegulum apex across gap between bulb and cymbium (Figs 2B, C, 6A, B). Conductor membranous, on apical end on tegulum (Figs 2A, C, 6A, B). Median apophysis absent. Female (paratype, HNU251). Total length 9.95 (Fig. 1B). Cephalothorax 4.56 long, 3.47 wide. Sternum 2.33 long, 2.06 wide (Fig. 1D). Abdomen 5.68 long, 4.83 wide. Anterior median eye diameter 0.19. Clypeus height 1.4 times one anterior median eye diameter. Chelicerae with five prolateral and five retrolateral teeth (Fig. 5B); stridulatory striae absent. Legs annulated with sporadic spines. Leg measurements: I 15.73 (4.48, 1.51, 3.83, 4.01, 1.90), II 14.48 (4.22, 1.53, 3.51, 3.60, 1.62), III 11.46 (3.40, 1.31, 2.70, 2.70, 1.35), IV 13.92 (4.05, 1.31, 3.48, 3.61, 1.47). Metatarsus I trichobothrium (TmI) 0.25 approximately; Metatarsus II, III and IV trichobothria (TmII–IV) 0.33 approximately. All metatarsal trichobothria located on the dorsum, near the retrolateral surface. Similar to males in somatic features. Epigynum: basal plate trapeziform, anterior margin 1.5 times longer than posterior margin, and the posterior margin nearly straight but with slightly protruded in the middle.Atrium separated into two parts by septum. Copulatory openings visible (Fig. 1E). Copulatory ducts elongated, flat and looping. Spermathecae small and spherical, approximately one spermatheca diameter from each other (Figs 1F, G, 5C). Fertilization ducts medially oriented (Figs 1G, 5C). Distribution. China (Hunan, Guangxi) (Fig. 8)., Published as part of Liu, Jinxin, Xu, Xiang, Hormiga, Gustavo, Yin, Haiqiang & Li, Hao, 2023, Two new species of the spider genus Putaoa (Araneae, Linyphiidae) from southern China, pp. 553-564 in Zootaxa 5277 (3) on pages 554-556, DOI: 10.11646/zootaxa.5277.3.7, http://zenodo.org/record/7890227, {"references":["Hormiga, G. & Dimitrov, D. (2017) The discovery of the spider genus Putaoa (Araneae, Pimoidae) in Taiwan with the description of a new species, including its web architecture. Zootaxa, 4341 (1), 97 - 104. https: // doi. org / 10.11646 / zootaxa. 4341.1.8"]}

Sathytes magnus Yin & Li, 2012 (Figs 84A, 98C, D) Chinese common name: 巨型ḛà甲 Sathytes magnus Yin & Li, 2012a: 845. Type locality: Xizang A. R., Motuo County, Hanmi, alt. 2200 m. Distribution. Mêdog County, Tibet, SW China (Figs 84A, 98C, D). Remarks. Sathytes magnus was described based on one male and one female collected at Hanmi of Mêdog County (Yin & Li 2012). This species morphologically resembles a number of congeners whose males share a single basal protuberance of antennomere 11, and markedly oblique mesal margins of antennomeres 9. It differs in the large body size (3.36 mm), the relatively broad antennomeres 11 (Yin & Li 2012: fig. 4F), the unique forms of antennomeres 9 and their apophyses (Yin & Li 2012: fig. 7F), an apically expanded aedeagus (Yin & Li 2012: fig. 8F), as well as its distribution.

Sathytes linzhiensis Yin & Li, 2020 (Figs 83C, 99A, F) Chinese common name: ffżḛà甲 Sathytes linzhiensis Yin & Shen, 2020: 47. Type locality: China, Xizang, Motuo County, path from Bari village to Renqingbeng Temple, 29°18’44.82”N, 95°21’27.18”E, alt. 1833 m. Distribution. Mêdog County, Tibet, SW China (Figs 83C, 99A, F). Remarks. Sathytes linzhiensis was described from four males found at Bari Village and Beibeng Township of Mêdog. This species is distinct in possessing a small body size (1.7–1.9 mm), the markedly elongate and straight hyaline apophyses on antennomeres 9, and there are two small basal projections of antennomeres 11., Published as part of Yin, Zi-Wei, 2022, The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earth's " Third Pole " (Coleoptera, Staphylinidae, Pselaphinae), pp. 1-211 in Zootaxa 5111 (1) on page 127, DOI: 10.11646/zootaxa.5111.1.1, http://zenodo.org/record/6340729, {"references":["Shen, Q. & Yin, Z. - W. (2020) Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4751 (2), 386 - 394. https: // doi. org / 10.11646 / zootaxa. 4751.2.13"]}

Labomimus shii (Yin & Li, 2012), comb. nov. Pselaphodes shii Yin & Li, 2012 in Y I N et al. (2012: 79) Type locality. China: Hainan Prov., Ledong County, Jianfengling N. R., Yulingu, 18.74686°N, 108.92988°E, ca. 635 m. Additional material examined (2 specimens). VIETNAM: HÒA BÌNH: 2 ♂♂, ‘Vietnam, Hoa bing, 4.- 7.6.1986, Ha son binh prov., Jan Horák lgt.’ (MHNG). Comments. This species was originally described based on three males from Hainan, as a member of the genus Pselaphodes. It is characterized primarily by the male antennomere 9 with an acute anteromedial triangular process, the long horn-like metaventral processes, and a broad median lobe of aedeagus in lateral view. Due to the presence of a distinct metaventral fovea between the metaventral processes, this species is here transferred from Pselaphodes to Labomimus. No distinct intraspecific variation has been detected between the populations of China and Vietnam. Distribution. China: Hainan; Vietnam: Hòa Bình (Y I N et al. 2012; this paper). First country record for Vietnam., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on pages 39-40, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2012: Two new species of Pselaphodes Westwood and new record of Taiwanophodes minor Hlavac from South China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 175: 75 - 86.","YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2012: Two new species of Pselaphodes Westwood and new record of Taiwanophodes minor Hlavac from South China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 1 ID: 75 - 86."]}

Pselaphodes hainanensis Yin & Li, 2013 Pselaphodes hainanensis Yin & Li, 2013 in Y I N et al. (2013b: 43) Type locality. China, Hainan, Baisha County,Yuanmeng, near Yinggezui Station, N19°03′10″, E109°33’ 55, 660 m. Additional material examined (1 specimen). LAOS: HOUAPHANH: 1 ♂, ‘LAOS-NE, Hua Phan prov., Ban Saluei, Phu Phan Mt., 20°15′N, 104°02′E, 1500-2000 m, 26. iv.-11.v. 2001, J. Bezděk leg’ (NMPC). Comments. This species belongs to the P. tianmuensis species group and differs from related species primarily in the relatively elongate median lobe of aedeagus. The single male from Laos is morphologically largely identical to those of Hainan, even the positions of the two endophalic sclerites are almost the same. Distribution. China: Hainan; Laos: Houaphanh (Y I N et al. 2013b; this paper). First country record for Laos.

Linan fortunatus Yin & Li, 2013 Linan fortunatus Yin & Li, 2013a: 147. Type locality. China, Guangxi Province, Jinxiu County, Lianhuashan Mountain, 1100 m, 24°09′21″N, 110°06′48″E. Additional material examined (35 specimens). VIETNAMN: VĩNH PHÚC: 4 ♂♂ 6 ♀♀, ‘ VIETNAM, Tam Dao, N21°27′48″, E 105°38′46″, 16.V. 2012, 1229 m, sift 06, V. Grebennikov lgt’ (MHNG, SNUC); 5 ♂♂ 5 ♀♀, ‘ VIETNAM, Tam Dao, N21°29′27″, E105°37′49″, 13.V. 2012, 1128 m, sift 03, V.Grebennikov lgt’ (MHNG, SNUC); 1 ♂ 4♀♀, ‘ VIETNAM, Tam Dao, N21°27′44″, E105°38′55″, 17.V. 2012, 1185 m, sift 07, V. Grebennikov lgt’ (MHNG); 2 ♂♂ 2♀♀, ‘ VIETNAM, Tam Dao, N21°27′54″, E105°38′44″, 12.V. 2012, 1218 m, sift 02, V. Grebennikov’ (MHNG); 3 ♂♂ 1♀♀, ‘ VIETNAM, Tam Dao, N 21°27′54″, E 105°38′56″, 14.V. 2012, 1237 m, sift 04, V. Grebennikov lgt’ (MHNG); 1 ♂, ‘ Mt. Tam Dao (950), Vinh Phu Prov C N-VIETNAM], 13.v.2003. S Nomura leg’ (MHNG); 1 ♀, ‘same locality and collector, except ‘ 21. v. 2003 ’ (MHNG). Comments. The male of L. fortunatus can be readily separated from all congeners by denser setae at the posterolateral margins of pronotum and the anterolateral margins of elytra, as well as by apically broadened antennomere 9, which is massively impressed on the lateral surface. The female, which was previously unknown, lacks these dense setae on pronotum and elytra. This species was originally described from Guangxi, southern China, and is here reported from northern Vietnam. The populations from China and Vietnam differ only slightly in the shape of the apex of the median lobe and the position of the endophallic sclerites of aedeagus. Distribution. China: Guangxi; Vietnam: Vĩnh Phúc (Y I N & L I 2013 a; this paper). First country record for Vietnam., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on pages 40-41, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464, {"references":["YIN Z. - W. & LI L. - Z. 2013 a: Five new species of the genera Labomimus and Linan from Guangxi, South China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae 53 (1): 141 - 153."]}

Pselaphodes linae Yin & Li, 2012 Pselaphodes linae Yin & Li, 2012 in Y I N et al. (2012: 76) Type locality. China: Hainan Prov., Wuzhishan Mt., Guzhandao, 18.86657°N, 109.68285°E, ca. 640 m. Additional material examined (4 specimens). VIETNAM: HÒABÌNH: 3 ♂♂, ‘ Vietnam, Hoa binh, 4.- 7.6.1986, Ha son binh prov. Jan Horák lgt’ (NMPC); 1 ♂, ‘TOEIN, RÈGION DE HOA-BINH / MUÉSUM PARIS, 1933, A. de COOMAN’ (MHNG). Comments. Pselaphodes linae was originally described from Hainan and Fujian, southern China and later reported from Taiwan. Here it is recorded from northern Vietnam for the first time. This species is readily recognizable by the deeply emarginate ventral and lateral surfaces of antennomeres 11 and 10, respectively, presence of a highly raised disc-like projection near the apex of antennomere 9, and metatibia with an acute tooth at apical 1/3. The male from Vietnam shows little morphological variation in both external features and aedeagus. Distribution. China: Fujian, Hainan, Taiwan; Vietnam: Hòa Bình (Y I N et al. 2012, 2013a; this paper). First country record for Vietnam., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on pages 45-46, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2012: Two new species of Pselaphodes Westwood and new record of Taiwanophodes minor Hlavac from South China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 175: 75 - 86.","YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2012: Two new species of Pselaphodes Westwood and new record of Taiwanophodes minor Hlavac from South China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 1 ID: 75 - 86."]}

Sathytes tangliangi Yin & Li, 2012 Sathytes yunnanicus Yin & Li in Yin et al. 2012: 850, figs 2b, 3k, 5e, 7k, 8k. Material examined (1 specimen) CHINA • 1 ♂; Yunnan, Baoshan City, Tengchong County, Mingguang Town, Baihua Ling; 25°18′10″ N, 98°47′31″ E; alt. 1870 m; 21 Apr. 2013; Dai, Song and Peng leg.; SNUC. Remarks This species was originally described based on a male and a female from Qiqi, Gongshan County. An additional specimen shows it has a wider distribution in Yunnan. Distribution China: Yunnan., Published as part of Yin, Zi-Wei & Shen, Qi, 2020, Fifteen new species of Sathytes Westwood from China (Coleoptera, Staphylinidae, Pselaphinae), with an updated checklist of world species, pp. 37-74 in European Journal of Taxonomy 722 on pages 56-57, DOI: 10.5852/ejt.2020.722.1133, http://zenodo.org/record/4134339, {"references":["Yin Z. - W., Li L. - Z. & Zhao M. - J. 2012. Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History 46: 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651"]}

Sathytes yunnanicus Yin & Li, 2012 Sathytes yunnanicus Yin & Li in Yin et al. 2012: 853, figs 3o, 6c, j, 7o, 8o. Material examined (3 specimens) CHINA • 1 ♂; Nujiang Pref., Lushui Co., Pianma To., Gangfang Vill., 26°06′42.64″ N, 98°35′47.54″ E; alt. 2100 m; 7 Jun. 2015; Wen-Xuan Bi leg. [ẔwaeứṄoeLj]; SNUC • 2 ♂♂; Yunnan, Baoshan City, Tengchong County, Mingguang Town, Baihua Ling; 25°18′10″ N, 98°47′31″ E; alt. 1870 m; 21 Apr. 2013; Dai, Song and Peng leg.; SNUC. Remarks This species was originally described based on a single male from Qiqi, Gongshan County. Additional specimens show that it has a wider distribution in Yunnan. Distribution China: Yunnan., Published as part of Yin, Zi-Wei & Shen, Qi, 2020, Fifteen new species of Sathytes Westwood from China (Coleoptera, Staphylinidae, Pselaphinae), with an updated checklist of world species, pp. 37-74 in European Journal of Taxonomy 722 on page 65, DOI: 10.5852/ejt.2020.722.1133, http://zenodo.org/record/4134339, {"references":["Yin Z. - W., Li L. - Z. & Zhao M. - J. 2012. Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History 46: 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651"]}

Sathytes longitrabis Yin & Li, 2012 Fig. 7 Sathytes longitrabis Yin & Li in Yin et al. 2012: 843, figs 1a, 3d, 4d, 7d, 8d. Material examined (8 specimens) CHINA • 2 ♂♂, 4 ♀♀; Shaanxi Prov., Ankang City (Ȑaem), Ningshaan County (ṪŘn), Huoditang Forestry Centre (KṈṚWĸ); 33°26′ N, 108°27′ E; alt. 1500–1700 m; 12 Jul. 2012; Chen and W.R. Li leg.; SNUC • 1 ♂, 1 ♀, same collection data as for previous; 33°26′02″ N, 108°27′07″ E; alt. 1500–1700 m; 12 Jul. 2012; Y.H. Pan leg.; mixed leaf litter, sifted; SNUC. Remarks This species was originally described based on a single male specimen from Houzhenzi, Qinling (Ṁẃ), and is here recorded from Huoditang in the same mountain range. The habitus and diagnostic characters are shown in Fig. 7. Distribution China: Shaanxi., Published as part of Yin, Zi-Wei & Shen, Qi, 2020, Fifteen new species of Sathytes Westwood from China (Coleoptera, Staphylinidae, Pselaphinae), with an updated checklist of world species, pp. 37-74 in European Journal of Taxonomy 722 on pages 49-50, DOI: 10.5852/ejt.2020.722.1133, http://zenodo.org/record/4134339, {"references":["Yin Z. - W., Li L. - Z. & Zhao M. - J. 2012. Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History 46: 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651"]}

Pselaphodes parapectinatus sp. nov. (Figs 10 B, 12) Type material (11 specimens). HOLOTYPE: VIETNAM: LÀO CAI: ♂, ‘VIETNAM N, 22′20′N, 103′50′E, SAPA (Lao Cai) 25.5- 10.6.1991, leg. E. Jendek’ (MHNG). PARATYPES: VIETNAM: LÀO CAI: 9 ♂♂, same label data as for holotype (MHNG, SNUC); 1 ♂, ‘C N-Vietnam], 23.v.1998 / Mt. Phang Si Pang (1,840, beating), Lao Cai Prov.’ (MHNG). Diagnosis. Male length 3.23–3.28 mm. Antennomeres 9–11 greatly elongate and forming distinct club, antennomere 9 with large disc-like process near apex. Horn-like metaventral processes short and apically blunt. Profemur with ventral spine before middle expansion; protibia strongly curved, with small middle spine and large apical projection. Median lobe of aedeagus almost symmetric dorso-ventrally; parameres long and broad; endophallus composed of broad, plate-like structure and elongate sclerite. Female unknow. Description. Male (Fig. 10 B). Body length 3.23–3.28 mm. Head slightly longer than wide, HL 0.65–0.70 mm, HW 0.63 mm; eyes prominent, each composed of about 42 facets. Antennomeres 2–8 of similar form, each slightly longer than wide, antennomeres 9–11 (Fig. 12 A) each much longer than wide, antennomere 9 with distinct disclike process near apex, antennomere 10 subcylindrical, slightly broadened from base toward apex, antennomere 11 sub-oval, narrowing apically. Pronotum (Fig. 12 B) approximately as long as wide, PL 0.63–0.65 mm, PW 0.60–0.63 mm, sides roundly narrowing apically at apical 2/5. Elytra slightly longer than wide, EL 0.93–0.95 mm, EW 0.90 mm. Horn-like metaventral processes (Fig. 12 C) in lateral view roundly triangular, short and blunt at apices. Profemur (Fig. 12 D) with triangular spine at basal 1/3, angularly expanded at middle; protibia (Fig. 12 E) strongly curved in apical half, with small spine near middle and large apical projection; mesotrochanter, mesofemur (Fig. 12 F), metatrochanter and metafemur (Fig. 12 G) simple. Abdomen broad at base and narrowed posteriorly, AL 0.90–0.95 mm, AW 1.08–1.15 mm; tergite 1 (IV) longest, more than twice as long as tergite V; sternite 7 (IX) (Fig. 12 H) hemi-membranous, suboval.Aedeagus (Figs 12 I – E) 0.76 mm long; median lobe nearly symmetric dorso-ventrally; parameres each elongate and broad, with enlarged, round apices; endophallus comprising one plate-like sclerite with many small spines at apex, and one elongate, curved sclerite. Female. Unknown. Comparative notes. The new species is most similar to P. pectinatus Yin, Li & Zhao, 2011 from Hainan, southern China, by sharing similar position and structure of the leg modifications, and short horn-like metaventral processes of the male. These two species differ in the presence of a large disc-like process on antennomere 9 (process lacking in P. pectinatus), a much shorter and differently shaped apical projection of protibia, each paramere of aedeagus lacking the apical setae and roundly broadened at the apex (each paramere with many thick apical and preapical setae, and rounded but not broadened at the apex in P. pectinatus), and different structure of the endophallic sclerites of P. parapectinatus sp. nov. Etymology. The new specific name reflects the morphological similarity between the new species and P. pectinatus. The name consists of the Greek prefix pará - (beside; next to, near, from) and the Latin adjective pectinatus (- a, - um; meaning combed). Distribution. Vietnam: Lào Cai., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on page 50, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464

Pselaphodes olexai sp. nov. (Figs 10 A, 11) Type material (1 specimen). HOLOTYPE: VIETNAM: ♂, ‘ Tam Dao, 20-27.6, N. VIETNAM, A. Olexa, 1990’ (MHNG). Diagnosis. Male length 3.08 mm. Antennomere 9–11 elongate and forming distinct club, antennomere 9 with large disc-like protuberance near apex. Protrochanter with hook-like ventral spine, profemur strongly expanded at middle, with long, thin ventral spine, protibia with broad and apically truncate projection; mesotrochanter with acute ventral spine. Horn-like metaventral processes short and apically blunt. Median lobe of aedeagus asymmetric, narrowing and curved rightwards at apex in dorsal view; parameres broad; endophallus composed of two elongate sclerites. Female unknown. Description. Male (Fig. 10 A). Body length 3.08 mm. Head slightly wider than long, HL 0.56 mm, HW 0.59 mm; eyes prominent, each composed of about 42 facets. Antennomeres 2–8 similarly moniliform, antennomeres 9–11 (Fig. 11 A) forming distinct club, antennomere 9 with large disc-like protuberance near apex, antennomere 10 subcylindrical, antennomere 11 sub-oval, narrowing from middle toward apex. Pronotum (Fig. 11 B) slight longer than wide, PL 0.61mm, PW 0.57 mm, with lateral margins rounded and narrowing apically at apical 1/3. Elytra much broader than long, EL 0.81 mm, EW 1.10 mm. Horn-like metaventral processes (Fig. 11 C) short and straight, blunt at apices. Protrochanter (Fig. 11 D) with hook-like ventral spine, profemur strongly expanded and with acute and long ventral spine near middle, protibia (Fig. 11 E) with large, apically truncate projection before apex; mesotrochanter (Fig. 11 F) with short ventral spine; mesofemur (Fig. 11 F), metatrochanter and metafemur (Fig. 11 G) simple. Abdomen broad at base and narrowed apically, AL and AW 1.0 mm. Sternite 7 (IX) (Fig. 11 H) hemi-membranous, roundly elongate. Aedeagus (Figs 11 I – E) 0.55 mm long, median lobe asymmetric dorso-ventrally, narrowing and curved rightwards at apex in dorsal view; parameres each with two long and two short apical setae; endophallus with one elongate sclerites at middle and another elongate one at base. Female. Unknown. Comparative notes. Antennomere 9 with a distinct disclike protuberance, thickened protibia with a large, apically truncate projection, and broad profemur of the male collectively provide a ready separation of P. olexai sp. nov. from all known congeners. Etymology. The new species is named after the Czech entomologist Aldo Olexa, collector of the holotype. Distribution. Northern Vietnam.

Pselaphodes jendeki sp. nov. (Figs 7 A, 8) Type material (8 specimens). HOLOTYPE: VIETNAM: LÀO CAI: ♂, ‘VIETNAM N 22°20′N, 103°50′E, SAPA (Lao Cai) 25.5- 10.6.1991, leg. E. Jendek’ (MHNG). PARATYPES: VIETNAM: LÀO CAI: 7 ♂♂, same label data as for holotype (MHNG, SNUC). Diagnosis. Male length 3.80–3.85 mm. antennomeres 9–11 each elongate, almost simple, antennomere 9 with small disc-like process near apex. Protrochanter with acute ventral spine, profemur with sharp spine at middle; mesotrochanter with acute ventral spine, mesotibia with small, blunt apical tubercle. Horn-like metaventral processes considerably short, apices truncate and bent downwards. Median lobe of aedeagus asymmetric; parameres short; endophallus composed of three sclerites. Female unknow. Description. Male (Fig. 7 A). Body length 3.80–3.85 mm. Head slightly longer than wide, HL 0.68–0.73 mm, HW 0.68–0.70 mm; eyes prominent, each composed of about 40 facets. Antennomeres 2–8 each moniliform, antennomeres 9–11 (Fig. 8 A) each strongly elongate and enlarged, antennomere 9 with small disc-like structure near apex. Pronotum (Fig. 8 B) approximately as long as wide, PL 0.65–0.68 mm, PW 0.62–0.67 mm, sides narrowing anteriorly at apical two-fifths. Elytra broader than long, EL 1.10–1.20 mm, EW 1.30–1.40 mm. Horn-like metaventral processes (Fig. 8 C) short and broad in lateral view, apically bifid, upper apex acute, apex truncate.Protrochanter (Fig. 8 D) with thin, acute ventral spine; profemur with sharp ventral spine at middle; mesotrochanter (Fig. 8 E) with acute ventral spine, mesofemur simple, mesotibia (Fig. 8 F) with small apical tubercle; metatrochanters and metafemur (Fig. 8 G) simple. Abdomen broad at base and narrowed apically, AL 1.15–1.20 mm, AW 1.30 mm. Sternite 7 (IX) (Fig. 8 H) hemi-membranous. Aedeagus (Figs 8 I – E) 0.75 mm long, median lobe asymmetric dorso-ventrally, narrowing apically and curved rightwards at apex in dorsal view; parameres much shorter than median lobe, each with three apical setae, endophallus composed of one triangle and two slightly curved, elongate sclerites. Female. Unknown. Comparative notes. This species belongs to the P. tianmuensis species group whose males share almost simple antennal clubs except for antennomere 9 which has a disc-like process near the apex. The new species may be readily separated from allied species by short and truncate horn-like metaventral processes, unique configuration of the aedeagus, as well as by its distribution. Etymology. The new species is named after the Slovak entomologist Eduard Jendek, collector of the holotype. Distribution. Vietnam: Lào Cai., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on pages 44-45, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464

Pselaphodes banphabat sp. nov. (Figs 4 A, 5) Type material (1 specimen). HOLOTYPE: LAOS: BOLIKHAMSAI: ♂, ‘LAOS: 27.iv-1.v.97, 70 km NE Vientiane BAN PHABAT, 150 m, N18°16′, E103°11′, Sausa & Jendek’ (MHNG). Diagnosis. Male body length 3.05 mm. Antennomeres 9–11 enlarged and forming club, antennomeres 9 and 10 each with small tubercle at apex and base, respectively. Horn-like metaventral processes long, in lateral view narrowing apically. Protrochanter with tiny ventral spine, mesotrochanter with acute ventral spine; mesofemur with small tooth at middle. Median lobe of aedeagus slightly asymmetrical dorso-ventrally; parameres elongate; endophallus composed of spine-like structures and a single sclerite. Female unknown. Description. Male (Fig. 4 A). Body length 3.05 mm, Head approximately as long as wide, HL 0.59 mm, HW 0.57 mm; eyes prominent, each composed of about 44 facets. Antennomeres 9–11 (Fig. 5 A) enlarged and forming distinct club, antennomere 9 with a round tubercle at apex, antennomere 10 with similar but larger tubercle at base. Pronotum (Fig. 5 B) as long as wide, PL and PW 0.59 mm, roundly narrowing apically at apical 2/5. Elytra much broader than long, EL 0.83 mm, EW 1.05 mm. Horn-like metaventral processes (Fig. 5 C) long, in lateral view narrowing apically. Protrochanter (Fig. 5 D) with tiny spine at ventral margin, profemur simple; mesotrochanter (Fig. 5 E) with acute ventral spine, mesofemur with small tooth at middle; metatrochanter and metafemur (Fig. 5 F) simple. Abdomen broad at base and narrowed posteriorly, AL 0.98 mm, AW 1.05 mm; tergite 1 (IV) longest, about 2.5 times as long as tergite V; sternite 7 (IX) (Fig. 5 G) hemi-membranous, semi-ellipse. Aedeagus (Fig. 5 H – J) 0.59 mm long; median lobe dorso-ventrally slightly asymmetric; parameres elongate, broadening to near apices, then greatly narrowed at apices, each paramere with one long apical seta; endophallus composed of numerous spine-like structures, one membranous part, and one forked sclerite. Female. Unknown. Comparative notes. The new species is quite distinctive in the form of the male antennal clubs, which leads to a ready separation of P. banphabat from all other congeners. Etymology. The new species is named after the type locality, Ban Phabat. The name is a noun in apposition. Distribution. Laos: Bolikhamsai., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on page 42, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464

Labomimus cucphuong sp. nov. (Figs 1 B, 3) Type material (1 specimen). HOLOTYPE: VIETNAM: NINHBÌNH: ♂, ‘VIETNAM N, 100 EM s from Hanoi CUC-PHUONG nat. Park, 02.- 12.5.1991, leg. E. Jendek’ (MHNG). Diagnosis. Male body length 2.84 mm. Antennomere 9 with round, deep ventral concavity in apical half; antennomere 10 strongly constricted at base and forming nodule-like stem, broadly impressed for most area of ventral side. Horn-like metaventral processes in lateral view slightly broaden at apices. Protrochanter with blunt ventral projection, protibia with small tubercle at apex; mesotrochanter with small triangular protuberance. Median lobe of aedeagus almost symmetric dorsoventrally; parameres thin and elongate; endophallus with seven sclerites and one structure. Female unknown. Description. Male (Fig. 1 B). Body length 2.84 mm. Head winder than long, HL 0.48 mm, HW 0.56 mm; eyes prominent, each composed of about 43 facets. Antennomeres 3–8 gradually becoming shorter, antennomeres 9–11 (Fig. 3 A) forming distinct club; antennomere 9 with round, deep ventral concavity in apical half, antennomere 10 strongly constricted near base and forming nodule-like stem, most part of ventral surface broadly impressed, antennomere 11 much shorter than 10, truncate at base and with round apex. Pronotum (Fig. 3 B) slightly longer than wide, PL 0.56 mm, PW 0.50 mm, narrowed anteriorly at apical 2/5. Elytra broader than long, EL 0.74 mm, EW 1.05 mm. Horn-like metaventral processes (Fig. 3 C) in lateral view slightly expand at apices. Protrochanter (Fig. 3 D) with blunt projection; profemur angularly expanded at middle; protibia (Fig. 3 E) with small tubercle at apex; mesotrochanter (Fig. 3 F) with short triangle protuberance; mesofemur (Fig. 3 F), metatrochanter and metafemur (Fig. 3 G) simple. Abdomen broad at base and narrowed apically, AL 0.96 mm, AW 1.07 mm. Aedeagus (Figs 3 H – J) 0.51 mm long; median lobe almost symmetric dorsoventrally, in lateral view curved and narrowed at apex; each paramere with two preapical setae; endophallus composed of seven sclerites and one membranous structure. Female. Unknown. Comparative notes. This species can be readily recognized and distinguished from all other members of the genus by uniquely modified male antennal club. Etymology. The new species is named after the type locality, Cúc PhƯƠng National Park. The species name is a noun in apposition. Distribution. Vietnam: Ninh Bình., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on pages 37-38, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464

Pselaphodes nongfa sp. nov. (Figs 7 B, 9) Type material (1 specimen). HOLOTYPE: LAOS: ATTAPEU: ♂: ‘ LAOS, ATTAPEU prov.Annam Highlands Mts. Dong Amphan NBCA, ca. 1160 m, NONG FA C crater lake], env. 15°05.9′N, 107°25.6′E, Jiří Hájek leg. 30.iv.-6.v.2010 ’ (NMPC). Diagnosis. Male length 3.05 mm. Antennomeres 9–11 distinctly elongate and enlarged, lacking modification. Horn-like metaventral processes long and broad at apices. Protrochanter with blunt ventral projection, profemur roundly angulate at middle, protibia with distinct apical protuberance; mesotrochanter bluntly protuberant. Median lobe of aedeagus almost symmetric dorso-ventrally; parameres elongate and slender; endophallus composed of 13 sclerites. Female unknown. Description. Male (Fig. 7 B). Body length 3.05 mm. Head slightly longer than wide, HL 0.61 mm, HW 0.58 mm; eyes prominent, each composed of about 38 facets. Antennomeres 2–8 similarly moniliform; antennomeres 9–10 each elongate, antennomere10 truncate at base and with round apex, together they forming distinct club (Fig. 9 A), which lacks modification. Pronotum (Fig. 9 B) slightly longer than wide, PL 0.62 mm, PW 0.58 mm, with lateral margins slightly expanding and then narrowing anteriorly at apical third. Elytra much broader than long, EL 0.80 mm, EW 1.10 mm; disc with linear microsculpture. Horn-like metaventral processes (Fig. 9 C) elongate, in lateral view broadened at apex. Protrochanter (Fig. 9 D) with blunt ventral protuberance; profemur roundly protuberant at middle; protibia (Fig. 9 E) with large apical tubercle; mesotrochanter (Fig. 9 F) with roundly triangular ventral projection; mesofemur (Fig. 9 F), metatrochanter and metafemur (Fig. 9 G) simple. Abdomen broad at base and narrowed apically, AL 0.92 mm, AW 1.12 mm. Sternite 7 (IX) (Fig. 9 H) elongate, hemi-membranous. Aedeagus (Figs 9 I – E) 0.60 mm long, almost symmetric dorso-ventrally; median lobe broad at base and narrowing apicad; parameres each elongate and slender (right paramere); with two short preapical setae; endophallus composed of six short sclerites at base, five similar ones at middle, and two elongate sclerites on the left side. Female. Unknown. Comparative notes. The new species is most similar to P. maolanensis Huang, Li & Yin, 2018 from Guizhou, southwestern China by sharing many external features. The male of these two species can be readily separated by relatively much more elongate antennomere 10 in relation to antennomeres 9 and 11, the narrow apical half of the aedeagus, as well as the more complex structure of endophallus of P. nongfa sp. nov. Etymology. The new species is named after its type locality, Nong Fa Lake (Blue Lake). The name is a noun in apposition. Distribution. Laos: Attapeu.

Pselaphodes dalatensis sp. nov. (Figs 4 B, 6) Type material (1 specimen). HOLOTYPE: VIETNAM: LÂM Đồ NG: ♂, ‘VIETNAM.Da Lat, Institute of Biology, at light, No. 765., 10.XII.1994, S. Mahunka, Gy. Sziráki & L. Zombori’ (MHNG). Diagnosis. Male body length 3.85 mm.Antennomeres 9–11 greatly modified and forming distinct club, antennomere 9 broadly impressed on ventral surface, with short basal protuberance and setose tuft above protuberance, apical margin strongly elevated and expanded to form irregular lamina-like projection, antennomere 10 with mesal margin bluntly projected at base, antennomere 11 strongly constricted at base. Protrochanter with sharp ventral spine at base and tiny ventral tooth near apex, profemur with large triangular ventral projection at middle, protibia with two roundly-triangular projections along mesal margin in apical half; mesotrochanter with acute ventral spine. Hornlike metaventral processes short, in lateral view triangular, narrowing apically. Aedeagus with moderately long parameres; endophallus composed of two elongate sclerites. Female unknown. Description. Male (Fig. 4 B). Body length 3.85 mm. Head approximately as long as wide, HL 0.68 mm, HW 0.70 mm; eyes prominent, each composed of about 45 facets. Antennomeres 2–7 each moniliform, except for antennomere 3 distinctly longer than 2 and 4; antennomeres 9–11 (Fig. 6 A) greatly modified, antennomere 9 broadly excavated on ventral surface, with short basal protuberance, apical part strongly elevated and expanded to form irregular lamina-like projection, with setose tuft below projection, antennomere 10 moderately curved in lateral view, its mesal margin bluntly projected at base, antennomere 11 strongly constricted at base. Pronotum (Fig. 6 B) slightly longer than wide, PL 0.70 mm, PW 0.67 mm, narrowing apically at apical 2/5. Elytra much broader than long, EL 1.09 mm, EW 1.35 mm. Hornlike metaventral processes (Fig. 6 C) short, in lateral view sub-triangular and narrowing apically. Protrochanter (Fig. 6 D) with sharp ventral spine at base, and tiny ventral tooth near apex; profemur with large triangular ventral projection at middle; protibia (Fig. 6 E) with two roundly-triangular projections along mesal margin in apical half; mesotrochanter (Fig. 6 F) with acute ventral spine; mesofemur (Fig. 6 F), metatrochanter and metafemur (Fig. 6 G) simple. Abdomen broad at base and narrowed posteriorly, AL 1.28 mm, AW 1.28 mm. Aedeagus (Fig. 6 H – J) with elongate parameres, each (Fig. 6 E) having four thick apical setae; endophallus composed of two elongate sclerites (note: the median lobe of the aedeagus is broken in holotype). Female. Unknown. Comparative notes. This is another distinctive species, the male of which can be rather easily distinguished from all congeners by unusual forms of the antennal club and protibia. Etymology. The new species name, dalatensis (- is, - e), is a latinized adjective given after the type locality, Da Lat city. Distribution. Vietnam: Lâm Đồng., Published as part of Yin, Zi-Wei & Li, Ning, 2021, Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae), pp. 35-53 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 61 (1) on page 42, DOI: 10.37520/aemnp.2021.002, http://zenodo.org/record/5037464

Sathytes larinus Yin & Li, 2012 (Fig. 4) Sathytes larinus Yin & Li (in: Yin et al. 2012, p842). Material examined. 1 ♂, ��� China: Zhejiang, Longquan City, Fengyangshan N. R., Huang-mao-jian (���Ŧ��), 27��53���23.75���N, 119��10���32.88���E, mixed leaf litter, sifted, 1580 m, 01.v.2016, Jiang, Liu & Zhou leg.��� (SNUC). Comments. Sathytes larinus was described from one male and one female at Guiheshang Village (OiAE��LJ) in southern Fujian, and is here newly recorded from the main mountain peak of Fengyangshan Nature Reserve. The new record extends the known distributional range of this species for some 400 km in the northeast direction. The male antennal club (Fig. 4 B���D) and the aedeagus (Fig. 4E) of the Zhejiang population (Fig. 4A) are both inconspicuously relatively longer than those of the Fujian population, which is regarded as intraspecific variation. Distribution. China: Zhejiang (new provincial record), Fujian., Published as part of Shen, Qi & Yin, Zi-Wei, 2020, Five new species and a new record of Sathytes Westwood from eastern China (Coleoptera: Staphylinidae: Pselaphinae), pp. 386-394 in Zootaxa 4751 (2) on page 391, DOI: 10.11646/zootaxa.4751.2.13, http://zenodo.org/record/3713034, {"references":["Yin, Z. - W., Li, L. - Z. & Zhao, M. - J. (2012) Taxonomic study on Sathytes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Journal of Natural History, 46, 831 - 857. https: // doi. org / 10.1080 / 00222933.2011.651651"]}

Genus Zopherobatrus Yin & Li, 2015 This monotypic genus is represented by a single eyeless species, Zopherobatrus tianmingyii Yin & Li, 2015 from Guizhou, southwestern China (YIN & LI 2015). The complete loss of compound eyes, coupled with the extremely elongate appendages, and its occurrence in the dark zone of the cave suggest Z. tianmingyii is an obligate troglobiont. Here, a second species with strongly reduced (but still present) eyes is described from Chongqing. In light of the discovery of a second species, the generic diagnosis of Zopherobatrus needs to be modified as follows. Revised diagnosis. General body form slender, with extremely elongate appendages. Head narrowed apically; median longitudinal carina on vertex faintly present; lacking frontal rostrum; with indistinct antennal tubercles; eyes strongly reduced to lacking. Pronotal disc moderately convex; median longitudinal sulcus present or absent, lateral longitudinal sulci usually present; lacking antebasal spines; lacking spines on lateral margins. Elytra strongly constricted at base, each elytron with two basal foveae; discal stria lacking to distinct; disc moderately to strongly convex. Tergite IV (first visible tergite) longest, with single marginal carina., Published as part of Yin, Zi-Wei, 2020, New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae), pp. 163-168 in Acta Entomologica Musei Nationalis Pragae 60 (1) on pages 166-167, DOI: 10.37520/aemnp.2020.009, http://zenodo.org/record/3879323, {"references":["YIN Z. - W. & LI L. - Z. 2015: Zopherobatrus gen. n. (Coleoptera: Staphylinidae: Pselaphinae), a new troglobitic batrisine from southwestern China. Zootaxa 3985 (2): 291 - 295."]}

Pselaphodes cuonaus Yin, Li & Zhao, 2011 Pselaphodes cuonaus Yin, Li & Zhao, 2011: 465. Material examined: 9 ♂♂, 1 ♀; CHINA, ���MDOi���B �����, ���LJĀ������ď 13 km ��, 2016. VI.4 D2, 3356 m, N�� 27.8050, E�� 91.7620, ������^: ��������, K��� (China: Xizang, Cuona County, Le Township, 13 km of Bian-Fang-Liu-Lian Highway, Hong-Bin Liang leg., beating)��� (IZCAS, SNUC). Comments: The male of this species can be readily recognized based on the unique modification of the antennal clubs, which was illustrated in Yin et al. (2011: 468, fig. 16). Distribution: This species was known from a single male collected at Le Township. The present new record confirms the presence of this species in Cuona., Published as part of Yin, Zi-Wei, 2019, Two new species and new records of the Pselaphodes - complex of genera from China (Coleoptera: Staphylinidae: Pselaphinae), pp. 197-202 in Revue suisse de Zoologie 126 (2) on page 200, DOI: 10.5281/zenodo.3463447, {"references":["Yin Z. - W., Li L. - Z., Zhao M. - J. 2011. Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part II. Annales Zoologici 61: 463 - 481."]}

Parabolitobius sinensis Yin & Li, sp. nov. (Figs 1���2) Type material. Holotype: CHINA: ♂, ���China: Henan, Nanyang, Funiu Mountain, 1400���1700 m, 2.viii.2004, L. Tang, L. Zhu & J. Hu leg. (��DZ�� ������Dz���+��)��� (SNUC). Paratype: CHINA: 1 ♀, same label data as for the holotype (SNUC). Diagnosis. Body medium-sized, total body length (TBL) 6.0��� 6.7 mm; fore body length (FBL) 2.72���2.85 mm. Elytral length/width 0.77 (male) and 0.68 (female); disc evenly pubescent, lacking distinct sutural, discal or lateral rows of coarse punctures. Male sternite VIII broadly emarginate at posterior margin. Aedeagus with apices of parameres greatly exceeding that of median lobe, each paramere roundly expanded on mesal margin near apex, bearing one apical and two preapical setae; endophallus with unique, complex structure comprised of multiple sclerites. Description. Male (Fig. 1A). Body medium-sized, total body length (from anterior margin of clypeus to apex of abdomen) 6.7 mm, fore body length (from anterior margin of clypeus to apex of elytra along midline) 2.85 mm. Color in general reddish-brown; head, antennae, mouthparts, pronotum, and legs paler. Head small, length from anterior margin of clypeus to base 0.69 mm, width across eyes (also maximum width) 0.72 mm; surface vaguely punctate, lacking microsculpture. Eyes moderately large, elliptical, slightly longer than tempora, convex, slightly prominent laterally; with long ocular seta located near inner posterior margin of each eye, separated from margin by distance of twice diameter of one puncture.Antennae elongate, length 2.23 mm, with apex of antennomere 8 reaching posterior margin of pronotum; antennomeres 1���3 more sparsely setose; antennomeres 4���11 densely and finely pubescent and slightly compressed; relative lengths of antennomeres 1���11: 2.3: 1.1: 1.7: 1.1: 1.4: 1.3: 1.1: 1.2: 1.1: 1.0: 1.6. Maxillary palpi with palpomeres 1 small, palpomeres 2���3 much larger, rather widened apically and sparsely pubescent, palpomeres 4 conical, 1.2 times as long as, and 0.8 times as wide as palpomeres 3. Labial palpi with palpomeres 3 largest, 3.7 times as long as, and 2.2 times as wide as palpomeres 2. Pronotum broadly transverse, length along midline 1.12 mm, maximum width 1.40 mm, widest at basal 1/4, disc moderately convex, less so at basal portion; with arcuate lateral margins; posterolateral corners roundly angulate; disc with punctures as those of head, without microsculpture; anterior, lateral, and basal margins each with four, three, and four coarse punctures; preapical punctures separated from anterior margin, located at apical 1/4; each puncture bearing one long, suberect seta. Scutellum small, subtriangular and impunctate, with transverse microsculpture. Elytra (2A) moderately transverse, length along suture 1.08 mm, maximum width 1.40 mm, sides slightly divergent posteriorly; conjoint apical margin broadly emarginate at middle, anterolateral corners broadly rounded; surface clothed with fine punctures and evenly pubescent; posterior margin of each elytron with row of 14 punctures, each puncture bearing one seta similar to those of disc. Metathoracic wings strongly reduced. Legs elongate and slender; apical spines of meso- and metatibiae sub-equal in length; metatibiae 0.55 times as long as metatarsi; relative lengths of metatarsomeres 1���5: 3.7: 1.7: 1.4: 1.0: 1.1. Abdomen elongate, gradually tapering from base to apex; surface densely punctate and pubescent; punctures moderately coarse; pubescence long and gradually becoming longer posteriorly; microsculpture absent; apical margin of tergites III���VII bearing 4���5 pairs of long bristles. Sternite VII (Fig. 2B) with posterior margin truncate and indistinctly emarginate, bearing 12 long bristles; sternite VIII (Fig. 2C) with posterior margin broadly emarginate, bearing six long bristles and four much shorter setae. Aedeagus (Fig. 2D) elongate; median lobe evenly tapering apically; parameres rather narrow from base to apical 1/3, then gradually broadened apicad and roundly narrowing at apices, each paramere (Fig. 3F) bearing one apical and two preapical short setae; endophallus (Fig. 2E) with complex structure comprised of multiple sclerites. Female (Fig. 1B). General appearance similar to male, with slightly shorter antennae and elytra; sternite VIII with posterior margin broadly rounded. Measurements (as for male): TBL 6.0 mm; FBL 2.72 mm; length/width of head, pronotum, and elytra 0.72/0.70, 1.04/1.29, 0.92/ 1.35 mm; antennae 2.12 mm, relative lengths of antennomeres 1���11: 2.9: 1.3: 1.8: 1.5: 1.4: 1.2: 1.2: 1.2: 1.1: 1.0: 1.6; metatibiae 0.57 times as long as metatarsi; relative lengths of metatarsomeres 1���5: 2.6: 1.7: 1.5: 1.0: 1.2. Comparison. The new species is similar to P. prolongatus and P. yezoensis in general appearance. It shares with P. prolongatus the strongly reduced metathoracic wings, the presence of many long bristles on tergite and sternite VIII, and the aedeagus with parameres extending far beyond the apex of the median lobe; it shares with P. yezoensis the relatively more elongate elytra of the male, and each paramere with one apical and two preapical short setae. Parabolitobius sinensis is distinctly smaller (TBL 6.0��� 6.7 mm; FBL 2.72���2.85 mm) than both P. prolongatus (TBL 6.9���8.5 mm; FBL 2.8���3.0 mm) and P. yezoensis (TBL 6.9���7.3 mm; FBL 3.3���3.6 mm). It can be further separated from P. prolongatus by the relatively more elongate elytra of the male, the broader and shallower posterior emargination of male sternite VIII, and the aedeagal parameres each being much broader at their apices, and bearing one apical and two preapical setae (parameres much thinner at the apices and bearing two apical and one preapical setae in P. prolongatus); P. sinensis further differs from P. yezoensis in the lack of palisade fringe along the posterior margin of tergite VII, the denser and longer bristles of male tergite and sternite VIII, the broader and shallower posterior emargination of male sternite VIII, and the aedeagus with relatively much longer parameres that are broader at the apices. Distribution. China: Henan Province. Etymology. The new specific epithet refers to the country where the type locality is located, i.e., China., Published as part of Yin, Zi-Wei & Li, Li-Zhen, 2019, A new species of Parabolitobius Li, Zhao & Sakai, 2000 from central China (Coleoptera: Staphylinidae: Tachyporinae), pp. 492-496 in Zootaxa 4695 (5) on pages 493-495, DOI: 10.11646/zootaxa.4695.5.6, http://zenodo.org/record/3542643

Olophrinus parastriatus Chang, Yin & Li, sp. nov. Type material. HOLOTYPE: J, CHINA: YUNNAN: ‘China: Yunnan Prov., Nabanhe N. R., Xiao-nuo-you-xia-zhai, N22°14.121’, E100°37.09 ’’, alt. 950 m, 20.xi.2008, J.-Y. Hu & L. Tang leg.’ (deposited in the Insect Collection of Shanghai Normal University, Shanghai, China; abbreviation: SNUC). PARATYPE: CHINA: YUNNAN: 1 J, ‘ China: Yunnan Prov., Nabanhe N. R., Manfei, alt. 700 m, 05.v.2009, J.-Y. Hu & Z.-W. Yin leg.’ (SNUC). Diagnosis. Olophrinus parastriatus is most similar to O. striatus in sharing the head, pronotum, and elytra with microsculpture consisting of transverse waves, and the right paramere of the aedeagus much broader than the left one. The new species may be separated by the much deeper emargination of male sternite VI, and the relatively much shorter parameres of the aedeagus without a preapical denticle on the ventral surface. For species description and illustrations see CHANG et al. (2019)., Published as part of Chang, Yuan, Yin, Zi-Wei & Li, Li-Zhen, 2019, Validation of Olophrinus parastriatus (Coleoptera: Staphylinidae: Tachyporinae), pp. 490 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 59 (2) on page 490, DOI: 10.2478/aemnp-2019-0038, http://zenodo.org/record/5340337, {"references":["CHANG Y., YIN Z. - W. & LI L. - Z. & SCHULKE M. 2019: A review of the genus Olophrinus Fauvel (Coleoptera: Staphylinidae: Tachyporinae) from China. Acta Entomologica Musei Nationalis Pragae 59 (1): 307 - 324."]}

Olophrinus qian Chang, Yin, Li & Schülke, sp. nov. (Figs 15–17, 18) Type material. HOLOTYPE: ♁, CHINA: GUIZHOU: ‘China: Guizhou, Zunyi City, (Suiyang County Ō ṚH), Kuankuoshui N. R. (ĒāķssṂüae ffl), Gongtonggou (AEÑḿ), nr. Dadong (†ḄDzã), 18.viii.2010, 1550 m, T. Feng & Z.-W. Yin leg.’ (SNUC). PARATYPES: CHINA: GUIZHOU: 9 ♁♁ 7 ♀♀, same collecting data as holotype; 7 ♁♁ 4 ♀♀, also from Kuankuoshui, ‘Houshan (ÜƜ), 1550 m, 15.viii.2010, Z.-W. Yin leg.’ (SNUC). YUNNAN: 1 ♁ 3 ♀♀, ‘ CHINA: Yunnan, mountains WNW Wuding, 25°38’45’’N, 102°06’55’’E, 2390 m, mixed forest margin with alder and pine, litter sifted, 1.IX.2014, Schülke [CH 14-12a]’ (cSch); 2 ♁♁ 1 ♀, ‘ CHINA [12a] – Yunnan, mt. WNW Wuding, mix. forest, 25°38’45’’N, 102°06’55’’E, 2390 m, 1.IX.2014, V. Assing’ (cAss, cSch); 1 ♀, ‘ CHINA [12] – Yunnan, mt. WNW Wuding, mix. forest, 25°38’45’’N, 102°06’55’’E, 2390 m, 18.VIII.2014, V.Assing’ (cAss); 1 ♀, ‘ CHINA: Yunnan [CH 07-35], Dali Bai Auton. Pref., Wuliang Shan, 9 km SW Weishan, 2450–2500 m, 25°10′14″N, 100°14′22″E, W slope, oaks and pines, sifted, 13.VI.2007, M. Schülke’ (cSch); 1 ♀, ‘ CHINA: Yunnan, Pu’er Pref., Ailao Shan, 37 km NW Jingdong, 24°45′12″N, 100°41′24.5″E, devastated forest remnant, litter & dead wood sifted, 13.IX.2009, leg. M. Schülke [CH 09-48]’ (cSch). Diagnosis description. Male. Habitus as in Fig. 15A; body slightly convex, TBL 7.07–7.90 mm, FBL 4.90–5.32 mm; head, elytra and abdomen black, antennae and tarsi yellowish-brown, pronotal disc, and most parts of legs reddish-brown. Head (Figs 16A, 16B) sub-pentagon, HL 0.84–1.11 mm, HW 1.29–1.32 mm, with microsculpture consisting of small isodiametric or nearly isodiametric meshes; eyes prominent, EyL 0.34–0.37 mm; antennae elongate, AnL 2.48–2.51 mm, relative lengths of antennomeres 1–11: 1.62: 1.00: 1.56: 1.25: 1.68: 1.56: 1.31: 1.25: 1.50: 1.18: 1.56. Pronotum (Fig. 16C) strongly transverse, PL 1.52–1.71 mm, PW 3.20–3.22 mm, PL/PW 0.47–0.53, with microsculpture as of head. Elytra (Figs 16D, E) transverse, EL 2.19–2.36 mm, EW 3.23–3.34 mm, EL/EW 0.65–0.73, with microsculpture consisting of transverse waves, each elytron with five complete and one additional shorter rows of punctures. Abdominal surface (Fig. 16F) with coarse punctures, and with microsculpture as elytra; tergite VIII (Fig. 17A) with six relatively short lobes, deeply and narrowly incised between inner lobes and median ones; tergite IX (Fig. 17B) divided laterally to four lobes; sternite VI (Figs 17 C–D) with posterior margin shallowly emarginate, thickened setae arranged along emargination, short bristles arranged in sub-triangular area; sternite VII (Figs 17 E–F) with posterior margin deeply and broadly emarginate, peg setae continuously arranged along emargination; sternite VIII (Fig. 17G) with four lobes, deeply incised between inner lobes, mid-length 1.06 times as long as depth of incision; sternite IX elongate and asymmetric (Fig. 17H). Length of aedeagus (Figs 17 I–J) 2.07 mm, parameres short, and asymmetric, left paramere slightly broader than right one in ventral view, each with preapical denticle on ventral surface; median lobe broad at base and narrowing toward apex. Female. General habitus (Fig. 15B) similar to male. Measurements: TBL 7.85–8.27 mm, FBL 4.80–5.26 mm, HL 1.07–1.08 mm, HW 1.28–1.31 mm, EyL 0.35–0.36 mm, AnL 2.42–2.48 mm, PL 1.58–1.63 mm, PW 3.09–3.16 mm, PL/PW 0.50–0.52, EL 2.24–2.28 mm, EW 3.12–3.19 mm, EL/EW 0.71–0.72. Tergite VIII (Fig. 17K) with five lobes, inner lobe longer than others, outer lobes short; sternite VIII (Fig. 17L) with six lobes, fimbriate lobes longer than others; styli as in Fig. 17M. Comparative notes. Olophrinus qian is similar to O. nepalensis in sharing the head and pronotum with meshlike microsculpture, each elytron with five complete and one additional shorter row of punctures, and presence of a preapical denticle of the parameres. The new species can be separated by the much shallower incision between the inner and median lobes of male tergite VIII, the parameres of the aedeagus each with a preapical denticle, and the much broader base of the inner lobe of female tergite VIII. Etymology. ‘Qian (Ŗ)’ is an abbreviation for Guizhou Province, where the type locality is located; noun in apposition. Distribution. China: Guizhou, Yunnan (Fig. 18)., Published as part of Chang, Yuan, Yin, Zi-Wei, Li, Li-Zhen & Schülke, Michael, 2019, A review of the genus Olophrinus from China (Coleoptera: Staphylinidae: Tachyporinae), pp. 307-324 in Acta Entomologica Musei Nationalis Pragae 59 (1) on pages 317-319, DOI: 10.2478/aemnp-2019-0024, http://zenodo.org/record/4489004, {"references":["SHIBATA Y. 1992: A new species of the genus Olophrinus Fauvel (Coleoptera: Staphylinidae: Tachyporinae) first recorded from Taiwan. Elytra 20 (1): 41 - 46."]}

Olophrinus setiventris Chang, Yin & Li, sp. nov. (Figs 8B, 9–10, 18) Type material. HOLOTYPE: ♁, CHINA: YUNNAN: ‘ China: Yunnan Prov., Nabanhe N. R., Hui-lao-xin-zhai (OiZǛḃ), Lao-yin-shui-chi (Zửķ ±), alt. 1250 m, 04.v.2009, J.-Y. Hu & Z.-W. Yin leg.’ (SNUC). Diagnostic description. Male. Habitus as in Fig. 8B. Body slightly convex, TBL 7.01 mm, FBL 4.21 mm; head, pronotal disc and abdomen black, elytra and most parts of legs reddish-brown. Head (Figs 9A, 9B) sub-triangular, HL 0.92 mm, HW 1.05 mm, with fine punctures, posterolateral and posterior margins with microsculpture consisting of transverse waves; eyes prominent, EyL 0.31 mm; antennae broken. Pronotum (Fig. 9C) strongly transverse, PL 1.37 mm, PW 2.59 mm, PL/PW 0.53, with fine punctures, lacking microsculpture. Elytra (Figs 9D, 9E) transverse, EL 1.74 mm, EW 2.59 mm, EL/EW 0.67, each elytron with seven complete rows of punctures. Abdominal (Fig. 9F) surface with coarse punctures, and microsculpture consisting of transverse waves; tergite VIII (Fig. 10A) with six relatively short lobes, narrowly and shallowly incised between inner lobes and outer ones; tergite IX (Fig. 10B) divided laterally to four lobes; sternite VI (Figs 10 C–E) with posterior margin narrowly and shallowly emarginate, dense and elongate pubescence arranged along emargination; sternite VII (Figs 10 F–G) with posterior margin deeply and broadly emarginate, peg setae evenly arranged along posterior margin; sternite VIII (Fig. 10H) with four lobes, shallowly incised between inner and outer lobes, mid-length 1.52 times as long as depth of incision; sternite IX elongate and asymmetric (Fig. 10I). Length of aedeagus (Figs 10 J–K) 2.04 mm, parameres short, slightly asymmetric, right paramere longer than left one in ventral view, median lobe broad at base and narrowing toward apex. Female. Unknown. Comparative notes. Olophrinus setiventris is similar to O. lantschangensis in sharing the head with fine punctures, the posterolateral and posterior margins of the head with microsculpture consisting of transverse waves, and each elytron with seven complete rows of punctures. The new species can be readily separated by the densely setose emargination of the posterior margin of male sternite VI. Distribution. China: Yunnan (Fig. 18). Etymology. The specific epithet is a noun combined from Latin ‘ seti- ’ (seta, bristle) and ‘ ventris ’ (venter), referring to the setose posterior margin of male sternite VI., Published as part of Chang, Yuan, Yin, Zi-Wei, Li, Li-Zhen & Schülke, Michael, 2019, A review of the genus Olophrinus from China (Coleoptera: Staphylinidae: Tachyporinae), pp. 307-324 in Acta Entomologica Musei Nationalis Pragae 59 (1) on page 314, DOI: 10.2478/aemnp-2019-0024, http://zenodo.org/record/4489004

A new species of the genus Confusacris Yin & Li, 1987, namely i.e. Confusacris shanxiensis sp. nov. is described in this paper from Shanxi province of China. The new species is similar to Confusacris brachypterus Yin & Li, 1987 but differs from the latter by tegmen of male extending over middle of hind femur distinctly, maximum width of cubital vein area 1.5 times median vein area, furculae large, epiphallus without acute projection on both sides and length of interspace of mesosternum1.6 times minimum width in female. A key to known species of the genus Confusacris Yin & Li, 1987 is given in this paper. The type specimens are deposited in the College of Life Sciences, Hebei University, Baoding, China.

Smetanabatrus ghecu Yin & Li, 2015 Smetanabatrus ghecu Yin & Li, 2015: 377. Type locality: Carin Asciuii Ghec��, southern Shan State, central Myanmar. Material examined: MHNG-ENTO-00013302; ♀; Thailand, Chiang Mai, Doi Suthep; 3. 87; coll. de Rougemont; ��� Smetanabatrus ghecu Yin & Li 2015, Det. Yin 2018���. Distribution: Myanmar (southern Shan State); Thailand (Chiang Mai), new country record (Fig. 2). Comments: Given the similar body size (3.38 mm) and proportions, roundly broadened inner and outer margins of the maxillary palpomere IV, presence of a reversed ���U���-shaped sulcus on the vertex, and coarsely punctate head and pronotum, the present specimen can be readily identified as conspecific with the population from central Myanmar. The present record expands the distributional range of this species some 220 km to the southeast from the type locality. Key to males 1 Vertex lacking reversed ���U���-shaped sulcus connecting foveae (Fig. 1B); maxillary palpomere IV distinctly lobed on inner margin (Fig. 1C; Yin & Li, 2013: fig. 2A); protibia slightly broadened on outer margin (Fig. 1A; Yin & Li, 2013: fig. 1A), metatibia strongly dilated laterally on apical portion (Fig. 1A, 1E; Yin & Li, 2013: figs 1A, 2C).....................................................................................................................................................................................2 - Vertex with distinct reversed ���U���-shaped sulcus connecting foveae (Yin & Li, 2015: fig. 1); maxillary palpomere IV with both outer and inner margins roundly expanded (Yin & Li, 2015: fig. 2E); protibia slightly expanded on inner margin (Yin & Li, 2015: figs 1A, 2D), metatibia unmodified. Distribution: Myanmar, Thailand (Fig. 2)............................................................................................................................................................................ S. ghecu 2 Protrochanter simple, profemur with large ventral spine (Yin & Li, 2013: fig. 2B); tergite 1 (IV) with large lateral expansions and broad, deep median cavity, tergites 2-3 (V-VI) strongly raised at middle (Yin & Li, 2013: figs 1A, 2D-E). Distribution: East Malaysia (Fig. 2)............................................................................................ S. kinabalu - Protrochanter with large, flat protuberance on ventral margin, profemur simple (Fig. 1D); tergite 1 (IV) with median patch at basal half, submedial area moderately raised, tergites 2-3 (V-VI) unmodified (Fig. 1 F-G). Distribution: West Malaysia (Fig. 2).................................................................................................................... S. loebli sp. nov., Published as part of Yin, Zi-Wei & Cuccodoro, Giulio, 2018, Notes on Smetanabatrus from Southeast Asia (Coleoptera: Staphylinidae: Pselaphinae), pp. 245-248 in Revue suisse de Zoologie 125 (2) on pages 247-248, DOI: 10.5281/zenodo.1414213, {"references":["Yin Z. - W., Li L. - Z. 2015. A second species of Smetanabatrus (Coleoptera: Staphylinidae: Pselaphinae). Revue suisse de Zoologie 122 (2): 377 - 380.","Yin Z. - W., Li L. - Z. 2013. Smetanabatrus kinabalu (Staphylinidae: Pselaphinae: Batrisitae), a new genus and new species from Sabah, Borneo. Zootaxa 3718 (5): 477 - 482."]}

Ji, Li-Li, Zhi, Yong-Chao, Li, Xin-Jiang (2019): A new species and key to all known species of Confusacris Yin & Li, 1987 (Orthoptera: Acrididae) from China. Zootaxa 4555 (4): 595-599, DOI: https://doi.org/10.11646/zootaxa.4555.4.11

29. Labomimus schuelkei Yin & Li, 2012: 96. Distribution. China: Shaanxi., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 529, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2012) Notes on Michael Schulke's pselaphine collections from China. - Tyrini. I. genera Labomimus Sharp, Linan Hlavac and Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae). Zookeys, 251, 83 - 118. https: // doi. org / 10.3897 / zookeys. 251.4099"]}

37. Labomimus vespertilio Yin & Li, 2012: 98. Distribution. China: Yunnan., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 530, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2012) Notes on Michael Schulke's pselaphine collections from China. - Tyrini. I. genera Labomimus Sharp, Linan Hlavac and Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae). Zookeys, 251, 83 - 118. https: // doi. org / 10.3897 / zookeys. 251.4099"]}

18. Labomimus maoershanus Yin & Li, 2013: 142. Distribution. China: Guangxi., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 528, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2013) Five new species of the genera Labomimus and Linan from Guangxi, South China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae, 53, 141 - 153."]}

24. Labomimus paratorus Yin & Li, 2012: 91. Distribution. China: Shaanxi., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 529, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2012) Notes on Michael Schulke's pselaphine collections from China. - Tyrini. I. genera Labomimus Sharp, Linan Hlavac and Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae). Zookeys, 251, 83 - 118. https: // doi. org / 10.3897 / zookeys. 251.4099"]}

3. Labomimus bannaus Yin & Li, 2013 in Yin et al. (2013b: 344). Distribution. China: Yunnan., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 527, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2013) Five new species of the genera Labomimus and Linan from Guangxi, South China (Coleoptera: Staphylinidae: Pselaphinae). Acta Entomologica Musei Nationalis Pragae, 53, 141 - 153.","Yin, Z. - W., Nomura, S. & Li, L. - Z. (2013 b) New species and new records of the Pselaphodes complex of genera (Staphylinidae: Pselaphinae: Tyrini) from China. Annales Zoologici, 63, 343 - 356. https: // doi. org / 10.3161 / 000345413 X 669595"]}

5. Labomimus cognatus Yin & Li, 2012: 85. Distribution. China: Yunnan., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 528, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2012) Notes on Michael Schulke's pselaphine collections from China. - Tyrini. I. genera Labomimus Sharp, Linan Hlavac and Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae). Zookeys, 251, 83 - 118. https: // doi. org / 10.3897 / zookeys. 251.4099"]}

26. Labomimus quadratithorax Yin & Li, 2013: 143. Distribution. China: Guangxi.

7. Labomimus dabashanus Yin & Li, 2012: 88. Distribution. China: Hubei, Yunnan., Published as part of Zhang, Yu-Qing, Li, Li-Zhen & Yin, Zi-Wei, 2019, Fifteen new species and a new country record of Labomimus Sharp from China, with a checklist of world species (Coleoptera: Staphylinidae: Pselaphinae), pp. 497-531 in Zootaxa 4554 (2) on page 528, DOI: 10.11646/zootaxa.4554.2.7, http://zenodo.org/record/2623895, {"references":["Yin, Z. - W. & Li, L. - Z. (2012) Notes on Michael Schulke's pselaphine collections from China. - Tyrini. I. genera Labomimus Sharp, Linan Hlavac and Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae). Zookeys, 251, 83 - 118. https: // doi. org / 10.3897 / zookeys. 251.4099"]}

48. Pselaphodes pseudowalkeri Yin & Li, 2013: 330. China: Zhejiang, Fujian, Jiangxi., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 477, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W. & LI L. - Z. 2013: On the identity of Pselaphodes walkeri (Sharp, 1892) (Coleoptera: Staphylinidae: Pselaphinae), with description of a new related species. Zootaxa 3609 (3): 327 - 334."]}

25. Pselaphodes hainanensis Yin & Li, 2013 in YIN et al. (2013a: 43). China: Hainan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 476, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W. & LI L. - Z. 2013: On the identity of Pselaphodes walkeri (Sharp, 1892) (Coleoptera: Staphylinidae: Pselaphinae), with description of a new related species. Zootaxa 3609 (3): 327 - 334.","YIN Z. - W., HLAVAC P. & LI L. - Z. 2013 a: Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 275: 23 - 65."]}

38. Pselaphodes meniscus Yin, Li & Zhao, 2011: 472. China: Sichuan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 477, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2011: Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part II. Annales Zoologici 61: 463 - 481."]}

Pselaphodes condylus Yin, Li & Zhao 2011 Material examined (2 3, 7 ♀♀). 1 3, labelled ‘ CHINA: Guangxi Province / Xing’an County / Maoershan Mountain / 10.vii. 2011, 2,000–2,140 m / Chen, Ma, Peng & Zhu leg.’; 2 ♀♀, same label data; 2 ♀♀, same label data, except ‘Zhong Peng leg’; 1 3, same label data, except ‘ 11.vii. 2011, 2,000 m / W. J. He & L. Tang leg’; 1 ♀, same label data; 2 ♀♀, same label data, except ‘ 10.vii. 2011, 2,100 m’ Diagnosis. Pselaphodes condylus can be separated from the other Pselaphodes species by the large body size (ca. 3.5 mm); robust male antennomere IX, with a distinct disc-shaped hump at the anteromesal margin; distinct spine pattern on legs, and broad and dorsally rightward-curved median lobe of the aedeagus. Habitat. All specimens were sifted from a mixed leaf litter of a broad-leaved forest. Distribution. Southwest China: Guizhou; South China: Guangxi

15. Pselaphodes dayaoensis Yin & Li, 2012a: 30. China: Guangxi., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 476, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & GU F. - K. 2012 a: Taxonomic study on the genus Pselaphodes Westwood (Coleoptera, Staphylinidae, Pselaphinae) from China. Part III. Zootaxa 3189: 29 - 38."]}

12. Pselaphodes cuonaus Yin, Li & Zhao, 2011: 465. China: Xizang.

3. Pselaphodes anhuianus Yin & Li, 2013 in YIN et al. (2013a: 38). China: Anhui, Hubei.

43. Pselaphodes parvus Yin, Li & Zhao, 2011: 473. China: Guizhou, Hubei, Sichuan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 477, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2011: Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part II. Annales Zoologici 61: 463 - 481."]}

27. Pselaphodes hlavaci Yin, Li & Zhao, 2010: 14. China: Sichuan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 476, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2010: Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Zootaxa 2512: 1 - 25."]}

32. Pselaphodes kuankuoshuiensis Yin & Li, 2013 in YIN et al. (2013a: 45). China: Guizhou., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 477, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W. & LI L. - Z. 2013: On the identity of Pselaphodes walkeri (Sharp, 1892) (Coleoptera: Staphylinidae: Pselaphinae), with description of a new related species. Zootaxa 3609 (3): 327 - 334.","YIN Z. - W., HLAVAC P. & LI L. - Z. 2013 a: Further studies on the Pselaphodes complex of genera from China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 275: 23 - 65."]}

1. Pselaphodes aculeus Yin, Li & Zhao, 2010: 8. China: Anhui, Fujian, Hainan, Guangxi, Yunnan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 475, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2010: Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part I. Zootaxa 2512: 1 - 25."]}

22. Pselaphodes flexus Yin & Li, 2012: 106. China: Yunnan., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 476, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739

9. Pselaphodes bomiensis Yin, Li & Zhao, 2011: 465. China: Xizang., Published as part of Huang, Meng-Chi, Li, Li-Zhen & Yin, Zi-Wei, 2018, Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species, pp. 457-478 in Acta Entomologica Musei Nationalis Pragae 58 (2) on page 476, DOI: 10.2478/aemnp-2018-0035, http://zenodo.org/record/4488739, {"references":["YIN Z. - W., LI L. - Z. & ZHAO M. - J. 2011: Taxonomical study on the genus Pselaphodes Westwood (Coleoptera: Staphylinidae: Pselaphinae) from China. Part II. Annales Zoologici 61: 463 - 481."]}