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Picobia magellani sp. nov. (Figs. 30 and 31) Description. FEMALE NPF (holotype). Gnathosoma. Hypostomal apex tapering. Infracapitulum and stylophore punctate. Movable cheliceral digit edentate on proximal end. Each medial branch of peritremes with 5���7 chambers, each lateral branch with ill-visible chambers. Idiosoma. Propodonotal shield absent. Setae vi situated distinctly posterior to level of setae ve. Setae se and c1 situated at same transverse level. Setae vi, ve and si lightly beaded. Pygidial shield well developed, with vertical furrow, double-leaf, apunctate. Setae ag2 situated posterolateral to ag1. Large, punctate genital plate bearing setae ag1���3. Setae g1 filiform, situated on sharp-ended genital lobes. All coxal fields punctate. Legs. Paraxial claws of legs III and IV thiner than antaxial claws III���IV. Solenidia �� on genua I setiform. Mesurements. Total body length 495. Length of stylophore 125. Lengths of setae: vi 115, ve 115, si 170, se 180, c1 170, c2 180, d1 170, d2 170, e2 100, f1 55, f2 60, h1 55, h2 265, ag1 60, ag2 35, ag3 80, g1 50, ps1 25, ps2 30, tc���III���IV and tc���III���IV 55. Length ratios of setae: vi: ve: si 1:1:1.5, d2: e2 1.7:1, f1: f2 1:1, f1: h1 1:1, f2: h2 1:4.4, h1: h2 1:4.8, ag1:ag2:ag3 1.7:1:2.3, g1: ps2: ps1 2:1.2:1, tc���III���IV: tc���III���IV 1:1. FEMALE PF (7 paratypes). Total body length 745���795. MALE. Unknown. Type material. Female holotype (NPF) and 7 female (PF) paratypes from Scytalopus magellanicus (Gmelin) (Passeriformes: Rhinocryptidae), COLOMBIA: Bogota, no other data. Type deposition. All type material is deposited in the AMU, except 1 female (PF) paratype in ZSM and 1 female (PF) paratype in ZISP. Differential diagnosis. Picobia magellani sp. nov. is morphologically most simillar to P. schmidti Glowska and Milensky, 2014. In females of both species, the hypostomal apex is tapering; the lateral branches of the peritremes are with ill-visible chambers; setae vi are situated distinctly posterior to the level of setae ve; the pygidial shield is double-leaf, each with 1 spin-like outhgrowth; setae ag2 are situated posterolateral to ag1. This new species differs from P. schmidti as follow: in females of P. m a g e l l a n i, the propodonotal shield is absent; the length ratios of setae ve: si and f1: f2 are 1:1.5 and 1:1. In females of P. schmidti, the lateral propodonotal shields are present, each bearing bases of setae ve and si; the length ratios of setae ve: si and f1: f2 are 1:4 and 1:1.8���2.3. Etymology. The name of this species is derived from the specific name of the host��� Scytalopus magellanicus, and is a noun in apposition., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 43, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Milensky C. M. (2014) New species of the genus Picobia (Cheyletoidea: Syringophilidae) parasitizing tyrannid birds (Passeriformes: Tyrannidae). Zootaxa, 3821 (3), 373 - 383. http: // dx. doi. org / 10.11646 / zootaxa. 3821.3.6"]}

Picobia caudati Skoracki and Hebda, 2004 (Figs. 39 G���H) Picobia caudati Skoracki and Hebda, 2004: 2, figs. 1���9; Skoracki 2011: 352, figs. 261���262. Types deposited in AMU and ZISP, examined. Type host: Aegithalos caudatus (Linnaeus) (Passeriformes: Aegithalidae). Type locality: Poland. Host range and distribution. Monoxenous species inhabiting Aegithalos caudatus (Linnaeus) (Passeriformes: Aegithalidae) in Poland (Skoracki & Hebda 2004)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 52-53, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Hebda, G. (2004) Quill mites (Acari: Syringophilidae) from Aegithalos caudatus (Passeriformes: Aegithalidae). Zootaxa, 691, 1 - 6.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Picobia ploceus Klimovicova, Skoracki, Wamiti and Hromada, 2014 (Figs. 38 D���F) Picobia ploceus Klimovicova et al., 2014: 395, figs. 1���2. Types deposited in AMU, examined. Type host: Ploceus ocularis Smith (Passeriformes: Ploceidae). Type locality: Kenya. Host range and distribution. Monoxenous species inhabiting Ploceus ocularis Smith (Passeriformes: Ploceidae) in Kenya (Klimovicova et al. 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 51, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Klimovicova, M., Skoracki, M., Wamiti, W. & Hromada, M. (2014) New data on the systematics of quill mites of the subfamily Picobiinae (Acari: Syringophilidae) parasitising African birds with description of two new species. Folia Parasitologica, 61, 394 - 400."]}

Picobia magellani sp. nov. (Figs. 30 and 31) Description. FEMALE NPF (holotype). Gnathosoma. Hypostomal apex tapering. Infracapitulum and stylophore punctate. Movable cheliceral digit edentate on proximal end. Each medial branch of peritremes with 5–7 chambers, each lateral branch with ill-visible chambers. Idiosoma. Propodonotal shield absent. Setae vi situated distinctly posterior to level of setae ve. Setae se and c1 situated at same transverse level. Setae vi, ve and si lightly beaded. Pygidial shield well developed, with vertical furrow, double-leaf, apunctate. Setae ag2 situated posterolateral to ag1. Large, punctate genital plate bearing setae ag1–3. Setae g1 filiform, situated on sharp-ended genital lobes. All coxal fields punctate. Legs. Paraxial claws of legs III and IV thiner than antaxial claws III–IV. Solenidia σ on genua I setiform. Mesurements. Total body length 495. Length of stylophore 125. Lengths of setae: vi 115, ve 115, si 170, se 180, c1 170, c2 180, d1 170, d2 170, e2 100, f1 55, f2 60, h1 55, h2 265, ag1 60, ag2 35, ag3 80, g1 50, ps1 25, ps2 30, tc’III–IV and tc”III–IV 55. Length ratios of setae: vi: ve: si 1:1:1.5, d2: e2 1.7:1, f1: f2 1:1, f1: h1 1:1, f2: h2 1:4.4, h1: h2 1:4.8, ag1:ag2:ag3 1.7:1:2.3, g1: ps2: ps1 2:1.2:1, tc’III–IV: tc”III–IV 1:1. FEMALE PF (7 paratypes). Total body length 745–795. MALE. Unknown. Type material. Female holotype (NPF) and 7 female (PF) paratypes from Scytalopus magellanicus (Gmelin) (Passeriformes: Rhinocryptidae), COLOMBIA: Bogota, no other data. Type deposition. All type material is deposited in the AMU, except 1 female (PF) paratype in ZSM and 1 female (PF) paratype in ZISP. Differential diagnosis. Picobia magellani sp. nov. is morphologically most simillar to P. schmidti Glowska and Milensky, 2014. In females of both species, the hypostomal apex is tapering; the lateral branches of the peritremes are with ill-visible chambers; setae vi are situated distinctly posterior to the level of setae ve; the pygidial shield is double-leaf, each with 1 spin-like outhgrowth; setae ag2 are situated posterolateral to ag1. This new species differs from P. schmidti as follow: in females of P. m a g e l l a n i, the propodonotal shield is absent; the length ratios of setae ve: si and f1: f2 are 1:1.5 and 1:1. In females of P. schmidti, the lateral propodonotal shields are present, each bearing bases of setae ve and si; the length ratios of setae ve: si and f1: f2 are 1:4 and 1:1.8–2.3. Etymology. The name of this species is derived from the specific name of the host— Scytalopus magellanicus, and is a noun in apposition.

Neopicobia freya Skoracki and Unsoeld, 2014 (Figs. 27 K, L) Neopicobia freya Skoracki et al., 2014b: 74, figs. 4–6. Types deposited in AMU, ZISP and ZSM, examined. Type host: Dryocopus galeatus (Temminck) (Piciformes: Picidae). Type locality: Paraguay. Host range and distribution. Mesostenoxenous species associated with the woodpeckers (Piciformes: Picidae): Dryocopus galeatus (Temminck) from Paraguay, Piculus rubiginosus (Swainson) from Panama (Skoracki et al. 2014b), Dendropicos griseocephalus (Boddaert) from Tanzania and D. goertae (Muller) from Kenya (Klimovicova et al. 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 37, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Unsoeld, M., Kavetska, K. & Kaszewska, K. (2014 b) Quill mites of the subfamily Picobiinae (Acari: Syringophilidae) associated with woodpeckers (Aves: Piciformes: Picidae). Acta Parasitologica, 59, 68 - 79. http: // dx. doi. org / 10.2478 / s 11686 - 014 - 0210 - x","Klimovicova, M., Skoracki, M., Wamiti, W. & Hromada, M. (2014) New data on the systematics of quill mites of the subfamily Picobiinae (Acari: Syringophilidae) parasitising African birds with description of two new species. Folia Parasitologica, 61, 394 - 400."]}

Genus Gunabopicobia Skoracki and Hromada, 2013 Gunabopicobia Skoracki and Hromada, 2013: 196. Type species: Picobia zumpti Lawrence, 1959, by original designation. Diagnosis. FEMALE. Hypostomal apex bumpy. Peritremes V-shaped. Setae vi situated anterior to level of setae ve. Propodonotal shield divided into 2 wide lateral shields bearing bases of setae vi, ve, si and se, and large unpaired medial shield bearing bases of setae c1. Opisthonotal and genital lobes absent. Bases of setae 1a coalesced. Pseudanal setal series represented by 2 pairs. Genital setae absent. Apodemes I without thorn-like protuberances. Legs with full complement of solenidia. Physogastric form with distinctly enlarged idiosoma, worm-shaped in outline. MALE. Features as in females except: peritremes M-shaped; propodonotal shield entire, bearing bases of all propodonotal setae except c2; genital setal series represented by 2 pairs. Species included. Two species: G. masalaje Kaszewska et al., 2014 and G. zumpti (Lawrence, 1959). Host range. Columbiformes: Columbidae. Distribution. Afrotropical (South Africa), Nearctic (USA), Oceanian (Papua New Guinea), Oriental (Indonesia), and Palaearctic (Poland) regions., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 26, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Hromada, M. (2013) A review of picobiine mites (Acari: Syringophilidae: Picobiinae) parasitising African birds. Folia Parasitologica, 60, 192 - 212. http: // dx. doi. org / 10.14411 / fp. 2013.022","Lawrence, R. F. (1959) New mite parasites of African birds. Parasitology, 49, 425 - 427. http: // dx. doi. org / 10.1017 / S 0031182000026950","Kaszewska, K., Kavetska, K. & Skoracki, M. (2014) Two new species of quill mites of the family Syringophilidae (Acariformes: Cheyletoidea) associated with treronine doves (Columbiformes: Columbidae: Treroninae). Zootaxa, 3846 (2), 293 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 3846.2.9"]}

Neopicobia cardinalis (Skoracki, Hendricks and Spicer, 2010) (Figs. 26 H–K) Picobia cardinalis Skoracki et al., 2010b: 733, figs. 25–36. Types deposited in AMU and USNM, examined. Neopicobia cardinalis, Skoracki 2011: 384. Type host: Cardinalis cardinalis (Linnaeus) (Passeriformes: Cardinalidae). Type locality: USA. Host range and distribution. Monoxenous species parasitising Cardinalis cardinalis (Linnaeus) (Passeriformes: Cardinalidae) in USA (Skoracki et al. 2010b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 35-36, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Hendricks, S. & Spicer, G. S. (2010 b) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727 - 742. http: // dx. doi. org / 10.1603 / ME 09265","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Picobia cetti Skoracki, 2011 (Figs. 39 D���F) Picobia cetti Skoracki, 2011: 357, figs. 266, 267. Types deposited in AMU, examined. Type host: Cettia cetti (Temminck) (Passeriformes: Cettiidae). Type locality: Turkey. Host range. Monoxenous species inhabiting Cettia cetti (Temminck) (Passeriformes: Cettiidae) in Turkey (Skoracki 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 51, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Picobia galerida Skoracki, 2011 (Figs. 44 D���F) Picobia galerida Skoracki, 2011: 367, figs. 275 and 276. Types deposited in AMU, examined. Type host: Galerida cristata (Linnaeus) (Passeriformes: Alaudidae). Type locality: Poland. Host range and distribution. Monoxenous species inhabiting Galerida cristata (Linnaeus) (Passeriformes: Alaudidae) in Poland (Skoracki 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 58-59, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Pseudopicobia hapaloptila Skoracki, Scibek and Sikora, 2012 (Figs. 56 C, D) Pseudopicobia hapaloptila Skoracki et al., 2012a: 234, figs. 24���29. Types deposited in AMU, examined. Type host: Hapaloptila castanea (Verreaux) (Galbuliformes: Bucconidae). Type locality: Ecuador. Host range and distribution. Monoxenous species parasitising Hapaloptila castanea (Verreaux) (Galbuliformes: Bucconidae) in Ecuador (Skoracki et al. 2012)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 72, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Scibek, K. & Sikora, B. (2012 a) New genus and three new species of quill mites (Acari: Syringophilidae: Picobiinae) parasitising puffbirds (Aves: Piciformes). Folia Parasitologica, 59, 229 - 236. http: // dx. doi. org / 10.14411 / fp. 2012.031"]}

Lawrencipicobia poicephali (Skoracki and Dabert, 2002) (Figs. 21–23) Picobia poicephali Skoracki and Dabert, 2002: 143, figs. 16–24; Skoracki et al. 2004: 17; Skoracki 2011: 329, as incertae sedis. Types deposited in RMCA, examined. Lawrencipicobia poicephali, Skoracki & Hromada 2013: 199, figs. 4–6. Type host: Poicephalus senegalus versteri Finsch (Psittaciformes: Psittacidae). Type locality: Cameroon. Host range. Monoxenous species associated with Poicephalus senegalus versteri Finsch (Psittaciformes: Psittacidae) in Cameroon (Skoracki & Dabert 2002)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 29, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Dabert, J. (2002) A review of parasitic mites of the family Syringophilidae (Acari, Prostigmata) from African birds, with descriptions of four new species. Acta Parasitologica, 47, 137 - 146.","Skoracki, M., Bochkov, A. V. & Wauthy, G. (2004) Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes on their host-parasite relationships. Insect Systematics & Evolution, 35, 155 - 176. http: // dx. doi. org / 10.1163 / 187631204788912409","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Skoracki, M. & Hromada, M. (2013) A review of picobiine mites (Acari: Syringophilidae: Picobiinae) parasitising African birds. Folia Parasitologica, 60, 192 - 212. http: // dx. doi. org / 10.14411 / fp. 2013.022"]}

Charadriineopicobia apricaria sp. nov. (Figs. 11 and 12) Description. FEMALE (holotype). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 3 chambers, and lateral branch with 4 chambers. Stylophore densely punctate. Idiosoma. Propodonotal shield densely punctate in anterior half; 4 apunctate patches present. Setae vi, ve and si beaded. Setae c1 situated slightly anterior to level of setae se. Hysteronotal shields absent. Pygidial shield apunctate, pear-like in shape. Agenital and genital plates absent. Setae ag2 situated slightly posteromedial to setae ag3. Pseudanal setae ps1 short but not as microsetae. Coxal fields I punctate, II–IV apunctate. Legs. Claws of legs III and IV equal in size. Apodemes I without thorn-like protuberances. Measurements. Total body length 560. Length of stylophore 120. Lengths of setae: vi 25, ve 70, si 100, se 170, c1 195, c2 175, d1 140, d2 165, e2 140, f1 15, f2 30, h1 15, h2 >255, ag1 100, ag2 70, ag3 135, ps1 10, 3b and 4b 25, 3c and 4c 70, tc’III–IV 40, tc”III–IV 55. Length ratios of setae: vi: ve: si 1:2.8:4, f1: f2 1:2, h1: f1 1:1, ag1: ag2: ag3 1.4:1:1.9, tc’III–IV: tc”III–IV 1:1.4, 3b: 3c 1:2.8. MALE. Unknown. Type material. Female holotype from Pluvialis apricaria (Linnaeus) (Charadriiformes: Charadriidae), FRANCE: St. Malo, 1970, no other data. Type deposition. Holotype in AMU. Differential diagnosis. Female of Charadriineopicobia apricaria sp. nov. differs from two other described species of this genus by the absence of the hysteronotal shields; presence of short pseudanal setae but not as miocrosetae, and by the position of agenital setae ag2 which are situated posteromedial to setae ag3. In females of Ch. calidris and Ch. leucophaeus, the hysteronotal shield is present; pseudanal setae are as microsetae and setae ag2 are situated anteromedial to setae ag3. Etymology. The name of this species refers to specific name of the host— Pluvialis apricaria.

Picobia hylocichlae Skoracki, Spicer and OConnor, 2014 (Figs. 42 C, D) Picobia hylocichlae Skoracki et al., 2014a: 105, figs. 2���3. Types deposited in AMU, UMMZ and ZISP, examined. Type host: Hylocichla mustelina (Gmelin) (Passeriformes: Turdidae). Type locality: USA. Host range and distribution. Monoxenous species inhabiting Hylocichla mustelina (Gmelin) (Passeriformes: Turdidae) in USA (Skoracki et al. 2014a)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 56, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Spicer, G. S. & OConnor, B. M. (2014 a) A review of mites of the subfamily Picobiinae Johnston & Kethley, 1973 (Prostigmata: Syringophilidae) from North American birds. Systematic Parasitology, 87, 99 - 110. http: // dx. doi. org / 10.1007 / s 11230 - 013 - 9460 - 5"]}

Picobia sayornis Glowska, 2014 (Figs. 35 C) Picobia sayornis Glowska, 2014: 140, figs. 1���5. Types deposited in AMU and USNM, examined. Type host: Sayornis nigricans (Swainson) (Passeriformes: Tyrannidae). Type locality: Peru. Host range and distribution. Monoxenous species associated with Sayornis nigricans (Swainson) (Passeriformes: Tyrannidae) in Peru (Glowska 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 47, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. (2014) New quill mites (Cheyletoidea: Syringophilidae) parasitizing tyrannid birds (Passeriformes: Tyrannidae) in Peru. Zootaxa, 3814 (1), 139 - 145. http: // dx. doi. org / 10.11646 / zootaxa. 3814.1.9"]}

Rafapicobia brotogeris (Fain, Bochkov and Mironov, 2000) comb. nov. (Figs. 57, 58) Picobia brotogeris Fain et al., 2000: 62, figs. 90–94. Types deposited in RBINS, examined. Type host: Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae). Type locality: Brasil. Redescription. FEMALE (holotype and 2 paratypes). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 4–6 chambers, each lateral branch with ill-visible borders between chambers. Movable cheliceral digit edentate on proximal end. Stylophore apunctate. Idiosoma. Propodonotal shield entire, punctate, slightly concave on anterior margin. Setae vi, ve and si strongly beaded. Setae c1 and se situated at same transverse level. Two hysteronotal shields apunctate. Pygidial shield apunctate. Agenital and genital plates absent. Setae ag2 short, situated posterior to setae ag1. Pseudanal setae thin and minute. Coxal fields I–IV apunctate. Legs. Claws of legs III–IV equal in size. Measurements. Total body length 455–595. Length of stylophore 140–150. Lengths of setae: vi 30, ve 85, si 100, se 160–175, c2 150–190; d1 165, d2 135–140, e2 160–180, f1 15–25, f2 30–35, h1 15, ps1, ps2 and g1 5, ag1 110, ag2 5, ag3 130–140, tc’III–IV 35–40, tc”III–IV 75, l’RIII 25–35, 3b 35, 3c 75. Length ratios of setae: vi: ve: si 1:2.8:3.3, d1: d2: e2 1.2:1:1.3, f1: f2 1:1.2–1.4, f1: h1 1.7:1, ag1: ag2: ag3 >20:1:>20, 3b: 3c 1:2, tc’III–IV: tc”III–IV 1:1.5–2. MALE (1 paratype). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 5 chambers, each lateral branch with ill-visible borders between chambers. Stylophore apunctate. Idiosoma. Propodonotal shield entire, punctate. Setae vi, ve and si lightly beaded. Setae c1 and se situated at same transverse level. Hysteronotal shield entire, punctate bearing setae d1 and e2. Pygidial shield apunctate. Two agenital plates apunctate, bearing bases of setae ag1. All coxal fields apunctate. Measurements. Total body length 335. Lengths of setae: se 50, c2 45, c1 55, h2 15, ag1 20, other setae vi, ve, si, d1, d2, e2, f2, ag2 about 10. Type material examined. Female holotype, 2 female and 1 male paratypes from Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae), BRAZIL: Upper Amazonia, this bird died in the Zoo of Antwerp, 24 April 1966, coll. A. Fain, all material in RBINS. Host range. Monoxenous species associated with Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae) in Brazil (Fain et al. 2000).

Picobia phoeniculi Fain, Bochkov and Mironov, 2000 (Figs. 48 A–C) Picobia phoeniculi Fain et al., 2000: 69, figs. 110–117; Skoracki et al. 2004: 168; Skoracki 2011: 328; Skoracki & Hromada 2013: 206, figs. 10–12. Types deposited in RBINS, examined. Type host: Phoeniculus purpureus (Coraciiformes: Phoeniculidae). Type locality: Rwanda. Additional material. One female (NPF) and 1 female (PF) from Upupa epops epops Linnaeus (Coraciiformes: Upupidae), NEPAL: Khumbu, Chukhung (Himalayas, S. Mount Everest), elev. 4,800 m. a.s.l., 29 August 1962, all material in AMU; 2 females (PF) from same host species, MACEDONIA: Uskub, Ljubliana, 15 April 1918, coll. F. Doflein, all material in AMU; 1 female (PF) from same host species, KAZAKHSTAN: Semipalatinsk, 14 July 1922, coll. P. P. Schestowsky, specimen in AMU; 1 female (NPF) and 2 females (PF) from same host species, PAKISTAN: N.W. Frontier Prov., Malakand Div., Swat Distr., Yakh Tangai, 2,076 m. a.s.l., 23 April 1990, coll. S. M. Goodman, all material in FMNH. Host range and distribution. Metastenoxenous species associated with the coraciiform birds (Coraciiformes): Phoeniculus purpureus (Muller) (Phoeniculidae) from Rwanda (Fain et al. 2000), Upupa epops africana Bechstein (Upupidae) from Tanzania (Skoracki & Hromada 2013) and U. e. epops Linnaeus from Kazakhstan, Macedonia, Nepal, and Pakistan (present paper)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 62, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Fain, A., Bochkov, A. V. & Mironov, S. V. (2000) New genera and species of quill mites of the family Syringophilidae (Acari: Prostigmata). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 70, 33 - 70.","Skoracki, M., Bochkov, A. V. & Wauthy, G. (2004) Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes on their host-parasite relationships. Insect Systematics & Evolution, 35, 155 - 176. http: // dx. doi. org / 10.1163 / 187631204788912409","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Skoracki, M. & Hromada, M. (2013) A review of picobiine mites (Acari: Syringophilidae: Picobiinae) parasitising African birds. Folia Parasitologica, 60, 192 - 212. http: // dx. doi. org / 10.14411 / fp. 2013.022"]}

Picobia chloris Bochkov, Mironov and Kravtsova, 2000 (Figs. 44 A–C) Picobia chloris Bochkov et al., 2000: 352, figs. 1–10; Skoracki 2011: 369, figs. 277–279. Types deposited in ZISP, examined. Type host: Chloris chloris (Linnaeus) (Passeriformes: Fringillidae). Type locality: Kirghizia. Host range and distribution. Monoxenous species inhabiting Chloris chloris (Linnaeus) (Passeriformes: Fringillidae) in Kirghizia (Bochkov et al. 2000)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 58, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Bochkov, A. V., Mironov, S. V. & Kravtsova, N. T. (2000) Two new syringophilid mites from the Greenfinch Carduelis chloris (Passeriformes: Fringillidae) from Kirghizia (Acari: Syringophilidae). Genus, 11, 351 - 358.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Picobia currucae Skoracki and Magowski, 2001 (Figs. 39 I–K) Picobia currucae Skoracki and Magowski, 2001: 116, figs. 9–15; Skoracki 2011: 348, figs. 258–260. Types deposited in AMU, examined. Type host: Sylvia curruca (Linaneus) (Passeriformes: Sylviidae). Type locality: Poland. Host range and distribution. Oligoxenous species associated with the sylviid warblers of the genus Sylvia (Passeriformes: Sylviidae): S. curruca (Linnaeus) from Poland (Skoracki & Magowski 2001), Jordan (Skoracki 2011), S. hortensis (Gmelin) from Jordan (Glowska et al. 2007), and S. nisoria (Bechstein) from Turkey (Skoracki 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 53, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Magowski, W. (2001) Two new species of quill mites of the genus Picobia (Acari: Prostigmata: Syringophilidae) from passerine birds of Poland. Acarina, 9, 113 - 119.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Glowska, E., Skoracki, M. & Khourly, F. (2007) A new species and new records of syringophilid mites (Acari: Prostigmata: Syringophilidae) from birds of Jordan. Zootaxa, 1635, 63 - 68."]}

Pipicobia terpsiphoni Glowska and Schmidt, 2014 (Figs. 52 F, G) Pipicobia terpsiphoni Glowska and Schmidt, 2014a: 59, figs. 1���5. Types deposited in AMU and USNM, examined. Type host: Terpsiphone rufiventer (Swainson) (Passeriformes: Monarchidae): Type locality: Gabon. Host range and distribution. Monoxenous species parasitising Terpsiphone rufiventer (Swainson) (Passeriformes: Monarchidae) in Gabon (Glowska & Schmidt 2014a)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 68, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Schmidt, B. K. (2014 a) New quill mites (Cheyletoidea: Syringophilidae) parasitizing the black-headed paradiseflycatcher Terpsiphone rufiventer (Passeriformes: Monarchidae) in Gabon. Zootaxa, 3786 (1), 57 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3786.1.3"]}

Picobia phoenicuri Skoracki and Hromada, 2013 (Figs. 42 E, F) Picobia phoenicuri Skoracki and Hromada, 2013: 208, figs. 13���15. Types deposited in AMU, examined. Type host: Phoenicurus moussieri (Olphe-Galliard) (Passeriformes: Muscicapidae). Type locality: Tunisia. Host range and distribution. Monoxenous species inhabiting Phoenicurus moussieri (Olphe-Galliard) (Passeriformes: Muscicapidae) in Tunisia (Skoracki & Hromada 2013)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 56, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Hromada, M. (2013) A review of picobiine mites (Acari: Syringophilidae: Picobiinae) parasitising African birds. Folia Parasitologica, 60, 192 - 212. http: // dx. doi. org / 10.14411 / fp. 2013.022"]}

Genus Lawrencipicobia Skoracki and Hromada, 2013 Lawrencipicobia Skoracki and Hromada, 2013: 199. Type species: Picobia poicephali Skoracki and Dabert, 2002, by original designation. Diagnosis. FEMALE. Hypostomal apex flat. Peritremes M-shaped. Setae vi situated anterior to level of setae ve. Propodonotal shield entire, bearing all propodonotal setae except c2. Opisthonotal and genital lobes absent. Bases of setae 1a separated. Pseudanal setal series with 2 pairs. Genital series with 1 pair of setae. Apodemes I with small thorn-like protuberances. Legs with full complement of solenidia. Physogastric form unknown. MALE. Features as in females except: genital setal series represented by 2 pairs. Species included. Monotypic genus, L. poicephali (Skoracki and Dabert, 2002) only. Host range. Psittaciformes: Psittacidae. Distribution. Afrotropical region (Cameroon)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 29, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Hromada, M. (2013) A review of picobiine mites (Acari: Syringophilidae: Picobiinae) parasitising African birds. Folia Parasitologica, 60, 192 - 212. http: // dx. doi. org / 10.14411 / fp. 2013.022","Skoracki, M. & Dabert, J. (2002) A review of parasitic mites of the family Syringophilidae (Acari, Prostigmata) from African birds, with descriptions of four new species. Acta Parasitologica, 47, 137 - 146."]}

Picobia myrmecocichla Skoracki, Solarczyk and Sikora, 2012 (Figs. 43 A���C) Picobia myrmecocichla Skoracki et al., 2012b: 132, figs. 22���32. Types deposited in AMU and ZSM, examined. Type host: Myrmecocichla arnotti (Tristram) (Passeriformes: Muscicapidae). Type locality: Tanzania. Host range and distribution. Monoxenous species inhabiting Myrmecocichla arnotti (Tristram) (Passeriformes: Muscicapidae) in Tanzania (Skoracki et al. 2012b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 57, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Solarczyk, P. & Sikora, B. (2012 b) Three new species of picobiine mites (Acari: Syringophilidae) parasitizing African flycatchers (Aves: Muscicapidae). Systematic Parasitology, 83, 123 - 135. http: // dx. doi. org / 10.1007 / s 11230 - 012 - 9376 - 5"]}

Picobia schmidti Glowska and Milensky, 2014 (Figs. 35 E) Picobia schmidti Glowska and Milensky, 2014: 378, figs. 9–12, 19, 23. Types deposited in AMU and USNM, examined. Type host: Lophotriccus pileatus (Tschudi) (Passeriformes: Tyrannidae). Type locality: Peru. Host range and distribution. Mesotenoxenous species associated with the tyrant flycatchers (Passeriformes: Tyrannidae): Hemitriccus margaritaceiventer (Orbigny and Lafresnaye) from Guyana and Lophotriccus pileatus (Tschudi) from Peru (Glowska & Milensky 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 48, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Milensky C. M. (2014) New species of the genus Picobia (Cheyletoidea: Syringophilidae) parasitizing tyrannid birds (Passeriformes: Tyrannidae). Zootaxa, 3821 (3), 373 - 383. http: // dx. doi. org / 10.11646 / zootaxa. 3821.3.6"]}

Columbiphilus odontophoridus Skoracki and Sikora, 2011 (Figs. 17 C, D) Columbiphilus odontophoridus Skoracki and Sikora, 2011: 27, figs. 31���40. Types deposited in AMU and ZSM, examined. Type host: Colinus cristatus (Linnaeus) (Galliformes: Odontophoridae). Type locality: Colombia. Host range. Monoxenous species associated with Colinus cristatus (Linnaeus) (Galliformes: Odontophoridae) in Colombia and Surinam (Skoracki & Sikora 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 25, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Sikora, B. (2011) Quill mites (Acari: Syringophilidae) associated with galliform birds (Aves: Galliformes). Zootaxa, 2966, 13 - 30."]}

Phipicobia pygiptilae Glowska and Schmidt, 2014 (Figs. 28 and 29) Phipicobia pygiptilae Glowska and Schmidt, 2014b: 194, figs. 1���5. Types deposited in AMU and USNM, examined. Type host: Pygiptila stellaris (Spix) (Passeriformes: Thamnophilidae). Type locality: Guyana. Host range and distribution. Monoxenous species associated with Pygiptila stellaris (Spix) (Passeriformes: Thamnophilidae) in Guyana (Glowska & Schmidt 2014b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 39, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Schmidt, B. K. (2014 a) New quill mites (Cheyletoidea: Syringophilidae) parasitizing the black-headed paradiseflycatcher Terpsiphone rufiventer (Passeriformes: Monarchidae) in Gabon. Zootaxa, 3786 (1), 57 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3786.1.3","Glowska, E. & Schmidt, B. K. (2014 b) New taxa of the subfamily Picobiinae (Cheyletoidea: Syringophilidae) paraziting antbirds and gnateaters (Passeriformes: Thamnophilidae: Conopophagidae) in Guyana. Zootaxa, 3861 (2), 193 - 200. http: // dx. doi. org / 10.11646 / zootaxa. 3861.2.7"]}

Genus Pipicobia Glowska and Schmidt, 2014 Pipicobia Glowska and Schmidt, 2014a: 58. Type species: Picobia locustella Skoracki, Bochkov and Wauthy, 2004, by original designation. Diagnosis. FEMALE. Hypostomal apex tapering. Peritremes mouth-shaped. Setae vi situated anterior to level of setae ve. Propodonotal shield entire, bearing setae vi, ve, si, c1 and se or divided into 2 narrow lateral sclerites with setae si, se, and unpaired medial large sclerite with setae vi, ve, c1. Opisthonotal and genital lobes absent. Bases of setae 1a separated. Each pseudanal and genital setal series represented by 1 pair. Apodemes I with small thorn-like protuberances. Solenidia phi (φ) on tibiae I absent. Physogastric form with weakly enlarged idiosoma, bulb-shaped in outline. MALE. Unknown. Species included. Four species: Pipicobia glossopsitta (Skoracki et al., 2008), P. locustella (Skoracki et al., 2004), P. pyrrholaemus (Skoracki and Glowska, 2008) and P. terpsiphoni Glowska and Schmidt, 2014. Host range. Passeriformes (Acanthizidae, Locustellidae, Monarchidae) and Psittaciformes (Psittaculidae). Distribution. Afrotropical (Gabon), Palaearctic (Poland), and Australian (Australia) regions., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 65-66, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Schmidt, B. K. (2014 a) New quill mites (Cheyletoidea: Syringophilidae) parasitizing the black-headed paradiseflycatcher Terpsiphone rufiventer (Passeriformes: Monarchidae) in Gabon. Zootaxa, 3786 (1), 57 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3786.1.3","Skoracki, M., Bochkov, A. V. & Wauthy, G. (2004) Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes on their host-parasite relationships. Insect Systematics & Evolution, 35, 155 - 176. http: // dx. doi. org / 10.1163 / 187631204788912409","Skoracki, M. & Glowska, E. (2008) Two new species of the genus Picobia Haller (Acari: Syringophilidae) from Australian and Indonesian passeriform birds. New Zealand Journal of Zoology, 35, 281 - 286. http: // dx. doi. org / 10.1080 / 03014220809510125"]}

Charadriineopicobia apricaria sp. nov. (Figs. 11 and 12) Description. FEMALE (holotype). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 3 chambers, and lateral branch with 4 chambers. Stylophore densely punctate. Idiosoma. Propodonotal shield densely punctate in anterior half; 4 apunctate patches present. Setae vi, ve and si beaded. Setae c1 situated slightly anterior to level of setae se. Hysteronotal shields absent. Pygidial shield apunctate, pear-like in shape. Agenital and genital plates absent. Setae ag2 situated slightly posteromedial to setae ag3. Pseudanal setae ps1 short but not as microsetae. Coxal fields I punctate, II���IV apunctate. Legs. Claws of legs III and IV equal in size. Apodemes I without thorn-like protuberances. Measurements. Total body length 560. Length of stylophore 120. Lengths of setae: vi 25, ve 70, si 100, se 170, c1 195, c2 175, d1 140, d2 165, e2 140, f1 15, f2 30, h1 15, h2 >255, ag1 100, ag2 70, ag3 135, ps1 10, 3b and 4b 25, 3c and 4c 70, tc���III���IV 40, tc���III���IV 55. Length ratios of setae: vi: ve: si 1:2.8:4, f1: f2 1:2, h1: f1 1:1, ag1: ag2: ag3 1.4:1:1.9, tc���III���IV: tc���III���IV 1:1.4, 3b: 3c 1:2.8. MALE. Unknown. Type material. Female holotype from Pluvialis apricaria (Linnaeus) (Charadriiformes: Charadriidae), FRANCE: St. Malo, 1970, no other data. Type deposition. Holotype in AMU. Differential diagnosis. Female of Charadriineopicobia apricaria sp. nov. differs from two other described species of this genus by the absence of the hysteronotal shields; presence of short pseudanal setae but not as miocrosetae, and by the position of agenital setae ag2 which are situated posteromedial to setae ag3. In females of Ch. calidris and Ch. leucophaeus, the hysteronotal shield is present; pseudanal setae are as microsetae and setae ag2 are situated anteromedial to setae ag3. Etymology. The name of this species refers to specific name of the host��� Pluvialis apricaria., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 12-17, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Casto, S. D. (1977) Cuculiphilus lobatus gen. n., sp. n. representing a new subfamily of quill mites (Acarina: Syringophilidae) from the groove-billed ani, Crotophaga sulcirostris (Cuculiformes: Cuculidae). Southwestern Naturalist, 22, 169 - 176. http: // dx. doi. org / 10.2307 / 3669807"]}

Columbiphilus pteroclesi (Skoracki and OConnor, 2010) (Figs. 17 E, F) Picobia pteroclesi Skoracki and OConnor, 2010: 27, figs. 78–88. Types deposited in UMMZ, ZISP and AMU, examined. Columbiphilus pteroclesi, Skoracki 2011: 379. Type host: Pterocles senegallus (Linnaeus) (Pteroclidiformes: Pteroclididae). Type locality: Egypt. Host range. Oligoxenous species associated with pteroclidiform birds of the genus Pterocles (Pteroclidiformes: Pteroclididae): P. coronatus (Linnaeus) and P. senegallus Lichtenstein both from Egypt (Skoracki & OConnor, 2010)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 26, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & OConnor, B. (2010) New taxa of quill mites (Acari: Cheyletoidea: Syringophilidae). Zootaxa, 2341, 1 - 32.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Neopicobia troglodytes (Skoracki, Hendricks and Spicer, 2010) (Figs. 26 L) Picobia troglodytes Skoracki et al., 2010b: 731, figs. 13–24. Types deposited in AMU and USNM, examined. Neopicobia troglodytes, Skoracki 2011: 384; Skoracki et al. 2014a: 104. Type host: Troglodytes aedon Vieillot (Passeriformes: Troglodytidae). Type locality: USA. Host range and distribution. Metastenoxenous species associated with Thryomanes bewickii (Audubon) (Passeriformes: Troglodytidae) in USA (Skoracki et al. 2014a), Troglodytes aedon Vieillot (Troglodytidae) in USA (Skoracki et al. 2010b), and Cardellina pusilla (Wilson) (Passeriformes: Parulidae) in USA (Skoracki et al. 2014a)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 36, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Hendricks, S. & Spicer, G. S. (2010 b) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727 - 742. http: // dx. doi. org / 10.1603 / ME 09265","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Skoracki, M., Spicer, G. S. & OConnor, B. M. (2014 a) A review of mites of the subfamily Picobiinae Johnston & Kethley, 1973 (Prostigmata: Syringophilidae) from North American birds. Systematic Parasitology, 87, 99 - 110. http: // dx. doi. org / 10.1007 / s 11230 - 013 - 9460 - 5"]}

Neopicobia hepburni Glowska and Laniecka, 2014 (Figs. 27 A���E) Neopicobia hepburni Glowska and Laniecka, 2014: 636, figs. 1���5. Types depsoited in AMU and USNM, examined. Type host: Picumnus sclateri Taczanowski (Piciformes: Picidae). Type locality: Peru. Additional material. 14 females and 3 males from Picumnus temminckii Lafresnaye (Piciformes: Picidae), PARAGUAY: 3 November 1938, coll. A. Neunteufel, all material in AMU. Host range and distribution. Oligoxenous species associated with the piculets of the genus Picumnus (Piciformes: Picidae): P. o l i v a c e u s Lafresnaye and P. sclateri Taczanowski, both from Peru (Glowska & Laniecka 2014), and P. temminckii Lafresnaye [new host] from Paraguay (present paper)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 37, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Laniecka, I. (2014) A new quill mite species Neopicobia hepburni sp. nov. (Cheyletoidea: Syringophilidae) parasitizing picid birds (Piciformes: Picidae) in Peru. Acta Parasitologica, 59, 635 - 637. http: // dx. doi. org / 10.2478 / s 11686 - 014 - 0284 - 5"]}

Picobia riparius Skoracki, 2011 (Figs. 45 C, D) Picobia riparius Skoracki, 2011: 360, figs. 268–271. Types deposited in AMU, examined. Type host: Riparia riparia (Linnaeus) (Passeriformes: Hirundinidae). Type locality: Poland. Host range and distribution. Monoxenous species parasitising Riparia riparia (Linnaeus) (Passeriformes: Hirundinidae) in Poland (Skoracki 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 59, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Rafapicobia brotogeris (Fain, Bochkov and Mironov, 2000) comb. nov. (Figs. 57, 58) Picobia brotogeris Fain et al., 2000: 62, figs. 90���94. Types deposited in RBINS, examined. Type host: Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae). Type locality: Brasil. Redescription. FEMALE (holotype and 2 paratypes). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 4���6 chambers, each lateral branch with ill-visible borders between chambers. Movable cheliceral digit edentate on proximal end. Stylophore apunctate. Idiosoma. Propodonotal shield entire, punctate, slightly concave on anterior margin. Setae vi, ve and si strongly beaded. Setae c1 and se situated at same transverse level. Two hysteronotal shields apunctate. Pygidial shield apunctate. Agenital and genital plates absent. Setae ag2 short, situated posterior to setae ag1. Pseudanal setae thin and minute. Coxal fields I���IV apunctate. Legs. Claws of legs III���IV equal in size. Measurements. Total body length 455���595. Length of stylophore 140���150. Lengths of setae: vi 30, ve 85, si 100, se 160���175, c2 150���190; d1 165, d2 135���140, e2 160���180, f1 15���25, f2 30���35, h1 15, ps1, ps2 and g1 5, ag1 110, ag2 5, ag3 130���140, tc���III���IV 35���40, tc���III���IV 75, l���RIII 25���35, 3b 35, 3c 75. Length ratios of setae: vi: ve: si 1:2.8:3.3, d1: d2: e2 1.2:1:1.3, f1: f2 1:1.2���1.4, f1: h1 1.7:1, ag1: ag2: ag3 >20:1:>20, 3b: 3c 1:2, tc���III���IV: tc���III���IV 1:1.5���2. MALE (1 paratype). Gnathosoma. Infracapitulum apunctate. Each medial branch of peritremes with 5 chambers, each lateral branch with ill-visible borders between chambers. Stylophore apunctate. Idiosoma. Propodonotal shield entire, punctate. Setae vi, ve and si lightly beaded. Setae c1 and se situated at same transverse level. Hysteronotal shield entire, punctate bearing setae d1 and e2. Pygidial shield apunctate. Two agenital plates apunctate, bearing bases of setae ag1. All coxal fields apunctate. Measurements. Total body length 335. Lengths of setae: se 50, c2 45, c1 55, h2 15, ag1 20, other setae vi, ve, si, d1, d2, e2, f2, ag2 about 10. Type material examined. Female holotype, 2 female and 1 male paratypes from Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae), BRAZIL: Upper Amazonia, this bird died in the Zoo of Antwerp, 24 April 1966, coll. A. Fain, all material in RBINS. Host range. Monoxenous species associated with Brotogeris cyanoptera (Pelzeln) (Psittaciformes: Psittacidae) in Brazil (Fain et al. 2000)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 73, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Fain, A., Bochkov, A. V. & Mironov, S. V. (2000) New genera and species of quill mites of the family Syringophilidae (Acari: Prostigmata). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 70, 33 - 70."]}

Genus Calamincola Casto, 1978 Calamincola Casto, 1978: 158, non Cuculiphilus Casto, 1977: 170 (name preoc.); Fain et al. 2000: 34; Skoracki 2011: 326. Type species: Calamincola lobatus Casto, 1978, by original designation. Diagnosis. FEMALE. Hypostomal apex rounded. Peritremes U-shaped. Setae vi situated anterior to level of setae ve. Propodonotal shield divided into 2 wide lateral shields bearing bases of setae vi, ve, si and se and unpaired posteromedial shield bearing bases of setae c1. Opisthonotal lobes present. Bases of setae 1a coalesced. Pseudanal series represented by 2 pairs of setae. Genital series with 1 pair of setae. Genital lobes absent. Apodemes I with small thorn-like protuberances. Legs with full complement of solenidia. Physogastric form with extremely enlarged idiosoma, worm-shaped in outline. MALE. Features as in females except: peritremes M-shaped; propodonotal shield entire; opisthonotal lobes absent; genital setal series with 2 pairs. Species included. Monotypic genus, C. lobatus Casto, 1977 only. Host range. Cuculiformes: Cuculidae. Distribution. Nearctic region (USA, Mexico)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 8, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Casto, S. D. (1978) A new name for Cuculiphilus (Acarina: Syringophilidae). Southwestern Naturalist, 23, 158. http: // dx. doi. org / 10.2307 / 3669996","Casto, S. D. (1977) Cuculiphilus lobatus gen. n., sp. n. representing a new subfamily of quill mites (Acarina: Syringophilidae) from the groove-billed ani, Crotophaga sulcirostris (Cuculiformes: Cuculidae). Southwestern Naturalist, 22, 169 - 176. http: // dx. doi. org / 10.2307 / 3669807","Fain, A., Bochkov, A. V. & Mironov, S. V. (2000) New genera and species of quill mites of the family Syringophilidae (Acari: Prostigmata). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 70, 33 - 70.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Picobia lamprotornis Klimovicova, Skoracki, Wamiti and Hromada, 2014 (Figs. 47 C���E) Picobia lamprotornis Klimovicova et al., 2014: 395, figs. 3���5. Types deposited in AMU, examined. Type host: Lamprotornis superbus (R��ppell) (Passeriformes: Sturnidae). Type locality: Kenya. Host range and distribution. Monoxenous species inhabiting Lamprotornis superbus (Ruppell) (Passeriformes: Sturnidae) in Kenya (Klimovicova et al. 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 61, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Klimovicova, M., Skoracki, M., Wamiti, W. & Hromada, M. (2014) New data on the systematics of quill mites of the subfamily Picobiinae (Acari: Syringophilidae) parasitising African birds with description of two new species. Folia Parasitologica, 61, 394 - 400."]}

Charadriineopicobia leucophaeus (Skoracki, Hendricks and Spicer, 2010) (Figs. 13 A–D) Picobia leucophaeus Skoracki et al., 2010b: 730, figs. 1–10. Types deposited in AMU, examined. Charadriineopicobia leucophaeus, Skoracki et al. 2014a: 104. Type host: Leucophaeus atricilla (Linnaeus) (Charadriiformes: Laridae). Type locality: USA (Texas). Host range and distribution. Monoxenous species associated with Leucophaeus atricilla (Linnaeus) (Charadriiformes: Laridae) in USA (Texas) (Skoracki et al. 2010b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 17-18, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Hendricks, S. & Spicer, G. S. (2010 b) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727 - 742. http: // dx. doi. org / 10.1603 / ME 09265","Skoracki, M., Spicer, G. S. & OConnor, B. M. (2014 a) A review of mites of the subfamily Picobiinae Johnston & Kethley, 1973 (Prostigmata: Syringophilidae) from North American birds. Systematic Parasitology, 87, 99 - 110. http: // dx. doi. org / 10.1007 / s 11230 - 013 - 9460 - 5","Skoracki, M., Glowska, E., Lontkowski J. & Stawarczyk, T. (2010 a) Picobia ictericus sp. n., (Acari: Prostigmata: Syringophilidae) an ectoparasite of two icterid bird species from Brazil. Genus, 21, 143 - 148.","Skoracki, M. & Sikora, M. (2014) Two new quill mite species of the family Syringophilidae (Acari: Prostigmata) parasitising the house sparrow Passer domesticus (L.) (Aves: Passeriformes). Zootaxa, 3765 (2), 194 - 200. http: // dx. doi. org / 10.11646 / zootaxa. 3765.2.7"]}

Key to species of the genus Neopicobia (Females) 1. Hysteronotal shields well developed, bearing bases of setae d1 and e2. Agenital and genital plates present … / freya species group /............................................................................................. 2 - Hysteronotal shields reduced to small rings surrounded bases of setae d1 or absent. Agenital and genital plates absent … / anthi species group /........................................................................................ 4 2. Pygidial shield with well developed wing-like lobes. Setae 4c situated out of coxal fields IV.............................................................................................. N. hepburni Glowska and Laniecka, 2014 - Pygidial shield without wing-like lobes. Setae 4c situated on coxal fields IV...................................... 3 3. Length ratios of setae vi: ve: si 1:1.8:2–2.2, f1: f2 1:4.5–5.5 times. Agenital plates fused to genital plate. In males, setae d2 are about 3.5–4 times longer than e2.............................................. N. ea Skoracki and Unsoeld, 2014 - Length ratios of setae vi: ve: si 1:1.3–1.5:1.5–1.7, f1: f2 1:11. Agenital plates not fused to genital plate. In males, setae d2 are 9 times longer than e2...................................................... N. freya Skoracki and Unsoeld, 2014 4. Setae f1 situated out of pygidial shield.......................... N. troglodytes (Skoracki, Hendricks and Spicer, 2010) - Setae f1 situated on pygidial shield....................................................................... 5 5. Hysteronotal shield as small punctate rings surrounded bases of setae d1..................................................................................................... N. cardinalis (Skoracki, Hendricks and Spicer, 2010) - Hysteronotal shields absent............................................................................. 6 6. Setae ag1 and ag3 no more than twice longer than ag2....................................................... 7 - Setae ag1 and ag3 4–5 times longer than ag2............................ N. modularis (Skoracki and Magowski, 2001) 7. Setae ag1 and ag3 twice longer than ag2.................................................................. 8 - Setae ag1 and ag3 1.4–1.6 times longer than ag2............................................................. 9 8. Coxal fields III and IV densely punctate. Pygidial shield punctate... N. carpodacus (Skoracki, Hendricks and Spicer, 2010) - Coxal fields III and IV apunctate. Pygidial shield apunctate.................................. N. anthi (Fritsch, 1958) 9. Coxal fields I and II apunctate. Length of setae si, c1, d1 and e2 70 –75, 105, 80–95 and 85–100, respectively...................................................................................................... N. pari sp. nov. - Coxal fields I and II punctate. Length of setae si, c1, d1 and e2 165–175, 180–190, 165–170 and 155–170, respectively.............................................................. N. ictericus (Skoracki and Glowska, 2010) comb. nov., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 38-39, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Laniecka, I. (2014) A new quill mite species Neopicobia hepburni sp. nov. (Cheyletoidea: Syringophilidae) parasitizing picid birds (Piciformes: Picidae) in Peru. Acta Parasitologica, 59, 635 - 637. http: // dx. doi. org / 10.2478 / s 11686 - 014 - 0284 - 5","Skoracki, M., Unsoeld, M., Kavetska, K. & Kaszewska, K. (2014 b) Quill mites of the subfamily Picobiinae (Acari: Syringophilidae) associated with woodpeckers (Aves: Piciformes: Picidae). Acta Parasitologica, 59, 68 - 79. http: // dx. doi. org / 10.2478 / s 11686 - 014 - 0210 - x","Skoracki, M., Hendricks, S. & Spicer, G. S. (2010 b) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727 - 742. http: // dx. doi. org / 10.1603 / ME 09265","Skoracki, M. & Magowski, W. (2001) Two new species of quill mites of the genus Picobia (Acari: Prostigmata: Syringophilidae) from passerine birds of Poland. Acarina, 9, 113 - 119.","Fritsch, W. (1958) Die milbengattung Syringophilus Heller, 1880 (subordo Trombidiformes, Fam. Myobiidae Megnin, 1877). Zoologische Jahrbucher Systematik, 86, 227 - 234.","Skoracki, M., Glowska, E., Lontkowski J. & Stawarczyk, T. (2010 a) Picobia ictericus sp. n., (Acari: Prostigmata: Syringophilidae) an ectoparasite of two icterid bird species from Brazil. Genus, 21, 143 - 148."]}

Pseudopicobia malacoptila Skoracki, Scibek and Sikora, 2012 (Figs. 56 A, B) Pseudopicobia malacoptila Skoracki et al., 2012a: 233, figs. 14���23. Types deposited in AMU, ZSM and IPCAS, examined. Type host: Malacoptila panamensis Lafresnaye (Galbuliformes: Bucconidae). Type locality: Colombia. Host range and distribution. Monoxenous species associated with Malacoptila panamensis Lafresnaye (Galbuliformes: Bucconidae) in Colombia (Skoracki et al. 2012a)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 72, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Scibek, K. & Sikora, B. (2012 a) New genus and three new species of quill mites (Acari: Syringophilidae: Picobiinae) parasitising puffbirds (Aves: Piciformes). Folia Parasitologica, 59, 229 - 236. http: // dx. doi. org / 10.14411 / fp. 2012.031"]}

Picobia biarmicus Skoracki, Bochkov and Wauthy, 2004 (Figs. 36 A, B) Picobia biarmicus Skoracki et al., 2004: 165, figs. 28���36; Skoracki 2011: 341, figs. 252���255. Types deposited in AMU, examined. Type host: Panurus biarmicus (Linnaeus) (Passeriformes: Panuridae). Type locality: Poland. Host range and distribution. Monoxenous species inhabiting Panurus biarmicus (Linnaeus) (Passeriformes: Panuridae) in Poland (Skoracki et al. 2004)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 48, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Bochkov, A. V. & Wauthy, G. (2004) Revision of the quill mites of the genus Picobia Haller, 1878 (Acari: Syringophilidae) with notes on their host-parasite relationships. Insect Systematics & Evolution, 35, 155 - 176. http: // dx. doi. org / 10.1163 / 187631204788912409","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Rafapicobia zirnitra Skoracki, 2011 (Figs. 60 A���C) Rafapicobia zirnitra Skoracki, 2011: 392, figs. 293���296. Types deposited in AMU, examined. Type host: Saxicola rubetra (Linnaeus) (Passeriformes: Muscicapidae). Type locality: Poland. Additional material. Two females from Luscinia cyanura (Pallas) (Passeriformes: Muscicapidae), JAPAN: Shinano Prov., 3 October 1911, coll. R. Frenk, all material in AMU; 1 female and 2 males from same host species, CHINA: 21 March 1912, coll. unknown, all material in AMU; 3 females and 1 male from Luscinia akahige (Temminck) (Muscicapidae), JAPAN: 3 May 1912, coll. J. Gengler, all material in AMU; 1 female from Luscinia sibilans (Swinhoe) (Muscicapidae), CHINA: Weihai, 12 May 1911, coll. H. Lynes, specimen in AMU; 1 female from Phoenicurus auroreus (Pallas) (Muscicapidae), CHINA: Jangtsze, Schaschi, 19 December 1899, coll. Haberer, specimen in AMU. Host range and distribution. Mesostenoxenous species associated with the Old World flycatchers (Passeriformes: Muscicapidae): Ficedula hypoleuca (Pallas) from Poland (Skoracki 2011), Luscinia cyanura (Pallas) [new host] from Japan and China, Luscinia akahige (Temminck) [new host] from Japan, Luscinia sibilans (Swinhoe) [new host] from China, Phoenicurus auroreus (Pallas) [new host] from China (present paper), and Saxicola rubetra (Linnaeus) from Poland (Skoracki 2011)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 77, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Rafapicobia melzeri Skoracki, Unsoeld, Skorupski and Kavetska, 2014 (Fig. 61 E) Rafapicobia melzeri Skoracki et al., 2014: 229, figs. 1, 2. Types deposited in ZSM, AMU and ZISP, examined. Type host: Rallus aquaticus Linnaeus (Gruiformes: Rallidae). Type locality: Germany. Additional material. 16 females from Porzana fusca (Linnaeus) (Gruiformes: Ralidae), JAPAN: Yokohama area, coll. Haberer, all material in AMU and ZSM. Host range and distribution. Mesostenoxenous species associated with the rallid birds (Gruiformes: Rallidae): Pardirallus sanguinolentus (Swainson) from Chile, P. parva (Scopoli) from Kirghizia, P. porzana (Linnaeus) from France, Rallus aquaticus Linnaeus from Germany (Skoracki et al. 2014), and Porzana fusca (Linnaeus) [new host] from Japan (present paper)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 79, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Unsoeld, M, Skorupski, M. & Kavetska, K. (2014 c) Syringophilid mites associated with the rails (Aves: Rallidae) and a key to the species of the genus Rafapicobia Skoracki, 2011. Systematic Parasitology, 88, 227 - 232. http: // dx. doi. org / 10.1007 / s 11230 - 014 - 9502 - 7","Skoracki, M. & Sikora, M. (2014) Two new quill mite species of the family Syringophilidae (Acari: Prostigmata) parasitising the house sparrow Passer domesticus (L.) (Aves: Passeriformes). Zootaxa, 3765 (2), 194 - 200. http: // dx. doi. org / 10.11646 / zootaxa. 3765.2.7"]}

Picobia lemi Skoracki, Glowska and Sikora, 2008 Picobia lemi Skoracki et al., 2008: 60, figs. 1–12. Types deposited in AMU, examined. Type host: Manucodia chalybatus (Forster) (Passeriformes: Paradisaeidae). Type locality: New Guinea. Host range and distribution. Oligoxenous species inhabiting the birds of paradise (Passeriformes: Paradisaeidae) of the genus Manucodia: M. ater (Lesson) and M. chalybatus (Forster), both from New Guinea (Skoracki et al. 2008)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 56, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Glowska, E. & Sikora, B. (2008) Four new species of the quill mite genus Picobia Haller (Acari: Syringophilidae) parasitizing birds in the Australian Region. Zootaxa, 1961, 58 - 68.","Skoracki, M. & Glowska, E. (2008) Two new species of the genus Picobia Haller (Acari: Syringophilidae) from Australian and Indonesian passeriform birds. New Zealand Journal of Zoology, 35, 281 - 286. http: // dx. doi. org / 10.1080 / 03014220809510125"]}

Pipicobia pyrrholaemus (Skoracki and Glowska, 2008) (Fig. 51) Picobia pyrrholaemus Skoracki and Glowska, 2008: 284, figs. 14–19. Types deposited in AMU, examined. Neopicobia pyrrholaemus, Skoracki 2011: 384. Pipicobia pyrrholaemus, Glowska & Schmidt 2014a: 59. Type host: Pyrrholaemus sagittatus (Latham) (Passeriformes: Acanthizidae). Type locality: Australia. Host range and distribution. Monoxenous species parasitising Pyrrholaemus sagittatus (Latham) (Passeriformes: Acanthizidae) in Australia (Skoracki & Glowska 2008)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 67, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M. & Glowska, E. (2008) Two new species of the genus Picobia Haller (Acari: Syringophilidae) from Australian and Indonesian passeriform birds. New Zealand Journal of Zoology, 35, 281 - 286. http: // dx. doi. org / 10.1080 / 03014220809510125","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Glowska, E. & Schmidt, B. K. (2014 a) New quill mites (Cheyletoidea: Syringophilidae) parasitizing the black-headed paradiseflycatcher Terpsiphone rufiventer (Passeriformes: Monarchidae) in Gabon. Zootaxa, 3786 (1), 57 - 64. http: // dx. doi. org / 10.11646 / zootaxa. 3786.1.3"]}

Picobia dryobatis (Fritsch, 1958) (Figs. 48 G–I) Syringophilus dryobatis Fritsch, 1958: 242, fig. 10. Type material is lost (in Kethley: 65). Material from type host species from Poland, examined. Picobia dryobatis, Kethley 1970: 65; Skoracki 2011: 334, figs. 245–247; Skoracki et al. 2014a: 72; 2014b: 108. Type host: Dendrocopos major Linnaeus (Piciformes: Picidae). Type locality: Germany. Additional material. Three females (PF) and 2 males from Dendrocopos macei (Vieillot) (Piciformes: Picidae), INDIA: Uttarakhand, Naghar, 26 March 1966, coll. unknown, all material in AMU; 2 females (PF) and 1 male from Dendrocopos syriacus (Hemprich and Ehrenberg) (Picidae), IRAN: near Teheran, 7 March 1958, coll. S. H. J. Read, all material in AMU; 5 females (NPF), 2 females (PF) and 4 males from Dendrocopos borealis (Vieillot) (Picidae), USA: no other data, all material in AMU; 1 female (NPF) and 2 females (PF) from Jynx torquilla Linnaeus (Picidae), JAPAN: 7 September 1912, coll. J. Gengler, all material in AMU; 1 female (NPF), 5 females (PF) and 2 males from Picoides mixtus (Boddaert) (Picidae), PARAGUAY: San Luis, 16 September 1931, coll. E. Schuhmacher, all material in AMU; 2 females (NPF) and 3 females (PF) from Picoides tridactylus (Linnaeus) (Picidae), SLOVAKIA: Rejdova, 27 September 1975, coll. T. Wiesz, all material in AMU; 2 females (NFP), 2 females (PF) and 8 males from same host species, BOSNIA AND HERZEGOVINA: Herzegovina region, Mostar, coll. Sandre, all material in AMU; 2 females (NPF), 7 females (PF) and 6 males from same host species, POLAND: Bukowina, February 1999, coll. unknown, all material in AMU; 2 females (PF) from same host species, KIRGHIZIA: Tien-Schan, Naryn Distr., 7 February 1910, coll. Merzbacher, all material in AMU; 2 females (PF) and 2 males from same host species, GERMANY: Bayern, Graseck, Garmisch-Partenkirchen, 13 March 1925, coll. Pinzinger, all material in AMU and ZSM. Host range and distribution. Mesostenoxenous species associated with the woodpeckers (Piciformes: Picidae): Campethera nubica (Boddaert) from Kenya (Klimovicova et al. 2014), Dendrocopos borealis (Vieillot) [new host] from USA (present paper), D. darjellensis Blyth from Nepal (Skoracki et al. 2014b), D. leucotos (Bechstein) from Slovakia (Skoracki 2011), D. macei (Vieillot) [new host] from India (present paper), D. major (Linnaeus) from Germany (Fritsch 1958), Poland, Slovakia and England (Skoracki 2011; Nattress & Skoracki 2009), D. minor hortorum (Brehm) from Slovakia (Skoracki 2011), D. syriacus (Hemprich and Ehrenberg) [new host] from Iran (present paper), Dendropicos goertae St. Muller from Ethiopia (Skoracki et al. 2014b), D. fuscescens (Vieillot) from Zambia (Klimovicova et al. 2014), D. namaquus Lichteinstein from Tanzania, D. stierlingi Reichenov from Tanzania (Skoracki et al. 2014b), Dryocopus pileatus (Linnaeus) from USA (Skoracki et al. 2014a), Jynx torquilla Linnaeus [new host] from Japan (present paper), Melanerpes erythrocephalus (Linnaeus) from USA (Michigan), M. carolinus (Linnaeus) from USA (Michigan) (Skoracki et al. 2014a), Picoides mixtus (Boddaert) [new host] from Paraguay (present paper), P. pubescens (Linnaeus) from USA (Texas, Virginia), P. scalaris (Wagler) from USA (California) (Skoracki et al. 2010), P. tridactylus (Linnaeus) from Slovakia (Skoracki 2011), Bosnia and Herzegovina, Poland, Kirghizia and Germany (present paper), Veniliornis cassini (Malherbe) from Suriname, V. dingus (Sclater and Salvin) from Ecuador, V. fumigatus (Orbigny) from Venezuela, V. kirkii (Malherbe) from Trinidad and Tobago, V. maculifrons (Spix) from Brazil, V. passerinus (Linnaeus) from Brazil, V. sanguineus (Lichtenstein) from Suriname, and V. spilogaster (Wagler) from Paraguay (Skoracki et al. 2014b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on pages 62-63, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Fritsch, W. (1958) Die milbengattung Syringophilus Heller, 1880 (subordo Trombidiformes, Fam. Myobiidae Megnin, 1877). Zoologische Jahrbucher Systematik, 86, 227 - 234.","Kethley, J. B. (1970) A revision of the family Syringophilidae (Prostigmata: Acarina). Contributions of the American Entomological Institute, 5, 1 - 76.","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415.","Skoracki, M., Spicer, G. S. & OConnor, B. M. (2014 a) A review of mites of the subfamily Picobiinae Johnston & Kethley, 1973 (Prostigmata: Syringophilidae) from North American birds. Systematic Parasitology, 87, 99 - 110. http: // dx. doi. org / 10.1007 / s 11230 - 013 - 9460 - 5","Skoracki, M., Unsoeld, M., Kavetska, K. & Kaszewska, K. (2014 b) Quill mites of the subfamily Picobiinae (Acari: Syringophilidae) associated with woodpeckers (Aves: Piciformes: Picidae). Acta Parasitologica, 59, 68 - 79. http: // dx. doi. org / 10.2478 / s 11686 - 014 - 0210 - x","Fain, A., Bochkov, A. V. & Mironov, S. V. (2000) New genera and species of quill mites of the family Syringophilidae (Acari: Prostigmata). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 70, 33 - 70.","Haller, G. (1878) Freyana und Picobia. Zeitschrift fur Wissenschafliche Zoologie, 30, 81 - 98.","Klimovicova, M., Skoracki, M., Wamiti, W. & Hromada, M. (2014) New data on the systematics of quill mites of the subfamily Picobiinae (Acari: Syringophilidae) parasitising African birds with description of two new species. Folia Parasitologica, 61, 394 - 400.","Nattress, B. & Skoracki, M. (2009) A new species and further records of quill mites (Acari: Cheyletoidea: Syringophilidae) parasitic on birds (Aves) in England. Zootaxa, 2133, 49 - 54.","Skoracki, M., Glowska, E., Lontkowski J. & Stawarczyk, T. (2010 a) Picobia ictericus sp. n., (Acari: Prostigmata: Syringophilidae) an ectoparasite of two icterid bird species from Brazil. Genus, 21, 143 - 148."]}

Picobia troglodytidus Sikora and Skoracki, 2012 (Figs. 37 D) Picobia troglodytidus Sikora et al., 2012: 62, figs. 11���19. Types deposited in AMU, ZISP and ZSM, examined. Type host: Henicorhina leucophrys (Tschudi) (Passeriformes: Troglodytidae). Type locality: Costa Rica. Host range. Monoxenous species inhabiting Henicorhina leucophrys (Tschudi) (Passeriformes: Troglodytidae) in Costa Rica (Sikora & Skoracki 2012)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 49, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Scibek, K. & Sikora, B. (2012 a) New genus and three new species of quill mites (Acari: Syringophilidae: Picobiinae) parasitising puffbirds (Aves: Piciformes). Folia Parasitologica, 59, 229 - 236. http: // dx. doi. org / 10.14411 / fp. 2012.031","Sikora, B., Fajfer, M., Kavetska, K. & Skoracki, M. (2012) Three new species of quill mites (Acari: Syringophilidae) parasitizing the wrens (Aves: Troglodytidae). Zootaxa, 3167, 57 - 65."]}

Picobia ochoi Glowska and Milensky, 2014 (Figs. 35 D) Picobia ochoi Glowska and Milensky, 2014: 374, figs. 1���8, 18, 22. Types deposited in AMU and USNM, examined. Type host: Tolmomyias sulphurescens (Spix) (Passeriformes: Tyrannidae). Type locality: Guyana. Host range and distribution. Monoxenous species parasitising Tolmomyias sulphurescens (Spix) (Passeriformes: Tyrannidae) in Guyana (Glowska & Milensky 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 48, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Glowska, E. & Milensky C. M. (2014) New species of the genus Picobia (Cheyletoidea: Syringophilidae) parasitizing tyrannid birds (Passeriformes: Tyrannidae). Zootaxa, 3821 (3), 373 - 383. http: // dx. doi. org / 10.11646 / zootaxa. 3821.3.6"]}

Neopicobia carpodacus (Skoracki, Hendricks and Spicer, 2010) (Figs. 26 E, F) Picobia carpodacus Skoracki et al., 2010b: 737, figs. 37–43. Type deposited in AMU, examined. Neopicobia carpodacus, Skoracki 2011: 384. Type host: Haemorhous purpureus (Gmelin) (Passeriformes: Fringillidae). Type locality: USA. Host range and distribution. Monoxenous species parasitising Haemorhous purpureus (Gmelin) (Passeriformes: Fringillidae) in USA (Skoracki et al. 2010b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 35, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Hendricks, S. & Spicer, G. S. (2010 b) New species of parasitic quill mites of the genus Picobia (Acari: Syringophilidae: Picobiinae) from North American birds. Journal of Medical Entomology, 47, 727 - 742. http: // dx. doi. org / 10.1603 / ME 09265","Skoracki, M. (2011) Quill mites (Acari: syringophilidae) of the Palaearctic region. Zootaxa, 2840, 1 - 415."]}

Gunabopicobia masalaje Kaszewska, Kavetska and Skoracki, 2014 (Figs. 18, 19 and 20 A–D) Gunabopicobia masalaje Kaszewska et al., 2014: 294, figs. 1–3. Types deposited in AMU, ZISP and ZSM, examined. Type host: Ptilinopus iozonus Gray (Columbiformes: Columbidae). Type locality: Papua New Guinea. Host range and distribution. Mesostenoxenous species associated with columbiform birds of the family Columbidae (Columbiformes): Ducula luctuosa (Temminick) and D. rosacea (Temminick) both from from Indonesia, D. pistrinaria Bonaparte, D. rufigaster (Quoy and Gaimard), D. spilorrhoa (Gray), and Ptilinopus iozonus Gray all from Papua New Guinea (Kaszewska et al. 2014)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 26, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Kaszewska, K., Kavetska, K. & Skoracki, M. (2014) Two new species of quill mites of the family Syringophilidae (Acariformes: Cheyletoidea) associated with treronine doves (Columbiformes: Columbidae: Treroninae). Zootaxa, 3846 (2), 293 - 300. http: // dx. doi. org / 10.11646 / zootaxa. 3846.2.9"]}

Picobia oritis Skoracki, Antczak and Riegert, 2009 (Figs. 38 G–I) Picobia oritis Skoracki et al., 2009: 133, figs. 15–25; Klimovicova et al. 2014: 258. Types deposited in AMU, RMCA and ZISP, examined. Type host: Cyanomitra oritis (Reichenow) (Passeriformes: Nectariniidae). Type locality: Cameroon. Additional material. One female (PF) from Cinnyris chalybeus (Linnaeus) (Passeriformes: Nectariniidae), SOUTH AFRICA: 20 August 1955, coll. F. J. R. Junor, specimen in FMNH. Host range and distribution. Stenoxenous species associated with the sunbirds (Passeriformes: Nectariniidae): Cinnyris chalybeus (Linnaeus) [new host] from South Africa (present paper), C. erythrocercus (Hartlaub) from Uganda, C. talatala Smith from Botswana and Zambia, C. oustaleti (Bocage) from Angola, C. venustus (Shaw) from Somalia (Klimovicova et al. 2014), Cyanomitra olivacea (Smith) from Kenya (Skoracki et al. 2011), and C. oritis (Reichenow) from Cameroon (Skoracki et al. 2009)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 51, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Antczak, M., Riegert, J., Fainova, D. & Mikes, V. (2009) New species and new records of quill mites (Acari: Syringophilidae) inhabiting african passerines (Aves: Passeriformes). Acta Zoologica Academiae Scientiarum Hungaricae, 55, 123 - 137.","Klimovicova, M., Skoracki, M., Wamiti, W. & Hromada, M. (2014) New data on the systematics of quill mites of the subfamily Picobiinae (Acari: Syringophilidae) parasitising African birds with description of two new species. Folia Parasitologica, 61, 394 - 400.","Skoracki, M., Hromada, M. & Wamiti, W. (2011) A New species and New host records of syringophilid mites (Acari: Syringophilidae) from passerines from Kenya. Zootaxa, 2922, 34 - 40."]}

Genus Pseudopicobia Skoracki, Scibek and Sikora, 2012 Pseudopicobia Skoracki et al., 2012a: 230. Type species: Pseudopicobia nonnula Skoracki, Scibek and Sikora, 2012, by original designation. Diagnosis. FEMALE. Hypostomal apex rounded. Peritremes M-shaped. Setae vi situated posterior to level of setae ve. Propodonotal shield divided into 2 longitudinal sclerites, bearing bases of setae ve, si and se. Opisthonotal and genital lobes absent. Bases of setae 1a coalesced. Pseudanal series represented by 2 pairs of setae. Genital setae absent. Apodemes I with thorn-like protuberances. Legs with full complement of solenidia. Antaxial and paraxial claws of legs III and IV unequal in size and shape. Physogastric form with distinctly enlarged idiosoma, wormshaped in outline. MALE. Features as in females except: each pseudanal and genital setal series represented by 2 pairs. Species included. Three species: Psudopicobia nonnula Skoracki et al., 2012; P. malacoptila Skoracki et al., 2012, and P. hapaloptila Skoracki et al., 2012. Host range. Galbuliformes (Bucconidae). Distribution. Neotropical region (Colombia, Ecuador)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 69, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Scibek, K. & Sikora, B. (2012 a) New genus and three new species of quill mites (Acari: Syringophilidae: Picobiinae) parasitising puffbirds (Aves: Piciformes). Folia Parasitologica, 59, 229 - 236. http: // dx. doi. org / 10.14411 / fp. 2012.031"]}

Picobia cichladusa Skoracki, Solarczyk and Sikora, 2012 (Figs. 43 D���F) Picobia cichladusa Skoracki et al., 2012b: 124, figs. 1���9. Types deposited in AMU and ZISP, examined. Type host: Cichladusa arquata Peters (Passeriformes: Muscicapidae). Type locality: Tanzania. Host range and distribution. Monoxenous species inhabiting Cichladusa arquata Peters (Passeriformes: Muscicapidae) in Tanzania (Skoracki et al. 2012b)., Published as part of Skoracki, Maciej, Sikora, Bozena & Spicer, Greg S., 2016, A review of the subfamily Picobiinae Johnston and Kethley, 1973 (Acariformes: Prostigmata: Syringophilidae), pp. 1-95 in Zootaxa 4113 (1) on page 58, DOI: 10.11646/zootaxa.4113.1.1, http://zenodo.org/record/271508, {"references":["Skoracki, M., Solarczyk, P. & Sikora, B. (2012 b) Three new species of picobiine mites (Acari: Syringophilidae) parasitizing African flycatchers (Aves: Muscicapidae). Systematic Parasitology, 83, 123 - 135. http: // dx. doi. org / 10.1007 / s 11230 - 012 - 9376 - 5"]}