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Haploops similis Stephensen 1925 Type species collected by the R. V “ Ingolf”, one specimen, (sex?) length: 4.5 mm, Station 36: 61°50’N – 56°21’W, depth 2702 m, bottom: sediment type unknown. BIOICE material examined. Station 2314: nine specimens, 11September 1992, depth 156 m, 63° 42.16’N – 23°03.50’W; bottom:muddy sand.Station 2328: four specimens, 3 May 1993, depth 429m, 63°20.00’N – 10°57.00’W; bottom: sand. Station 2616: four specimens, 11 July 1994, depth 535 m, 67°11.38’N – 16°50.40’W; bottom: sandy silt. Station 2873: one specimen, 22 August 1996, depth 555 m, 64°37.41’N – 27°14.28’W; bottom: gravely sand. Station 2946: two specimens, 28 August 1996, depth 229 m, 65°47.90’N – 25°38.70’W; bottom: sandy silt. Station 3026: five specimens, 8 July 1997, depth 564 m, 63°53.96’N – 12°44.18’W; sediment type unknown. Station 3099: two specimens, 21August 1999, depth 229 m, 67°11.02’N – 21°45.68’W; bottom: gravely sandy silt. Station 3282: four specimens, 16 September 2001, depth 1808 m, 62°48.00’N – 16°14.80’W; bottom: sandy silt. Station 3531: two specimens, 9 September 2002, depth 1712 m, 62°43.26’N – 14°34.70’W; bottom: sandy silt. Station 3544: 16 specimens, 11September 2002, depth 1632 m, 61°33.005’N – 13°40.14’W; bottom: gravely sand. Station 3598: one specimen, 10 September 2003, depth 768 m, 62°17.37’N – 26°37.58’W; bottom: silty sand. All specimens of these stations are females. Diagnosis. Blind species; body without long dorsal setae on the pereon, pleon and urosome; A1 = 8–9/10 A2, A2 = 2/3 body; coxa 4 heart-shaped, Pereopod 7: basis narrow, anterior distal lobe of the carpus developed. Description. Head: Square shaped, without corneal lenses, anterior margin oblique and straight. Antenna 1 length is 8–9/10 of Antenna 2 length; Antenna 2 length is 2/3 body length. Pereon: without long dorsal setae. Gnathopod 1: coxa 1 roughly oval, distal margin rounded and fringed with long setae; basis long, slightly curved, approximately rectangular with many long setae, as long as ischium + merus + carpus, the merus, carpus and propodus bearing long setae; propodus oval. Gnathopod 2: coxa 2 approximately rectangular, distal margin straight with few setae; basis long, slightly curved, with many long setae, as long as ischium + merus + carpus; the merus, carpus and propodus bearing long setae; propodus oval. Pereopod 3: coxa 3 approximately rectangular, distal margin straight with few setae; basis, merus and carpus with long setae on the posterior margin; dactylus slender, curved and longer than propodus. Pereopod 4: coxa 4 heart-shaped, basis, merus and carpus with long setae on the posterior margin, and tuft of long setae on distal anterior margin of merus and propodus; dactylus slender, curved and shorter than propodus. Pereopod 5: coxa 5 rectangular, basis roughly rectangular with few setae on the anterior margin; carpus rectangular, anterior margin with few setae, posterior margin with rows of little spines, postero-distal lobe with short spines and a long one; propodus rectangular, longer than carpus, with few short setae on anterior margin, and long distal setae; dactylus short and curved. Pereopod 6: basis rounded with few short setae on the anterior margin; carpus rectangular, posterior margin with rows of little spines, postero-distal lobe ornamented with short spines and a very long one; propodus a little longer than carpus, with few short setae on anterior margin, and long distal setae; dactylus short and curved. Pereopod 7: basis narrow, anterior and posterior margin slightly concave, lobe not deflected, rounded with few long setae, not reaching the ischium-merus joint; ischium quadrangular; merus rectangular with an antero-inferior lobe, anterior and posterior margin ornamented with few little spines and long postero-distal setae; carpus oval, with spines on the anterior and posterior margin, anterior distal lobe developed; propodus and dactylus short and narrow. Pleon: without long dorsal setae. Epimeral plate 1: anterior margin oblique and straight, ventral margin and posterior margin straight, corners rounded. Epimeral plate 2: inferior and posterior margins slightly convex, postero-inferior corner rounded. Epimeral plate 3: anterior margin straight, posterior margin oblique and straight, inferior one slightly convex, anterior corner rounded, posterior corner slightly acute. Urosome: carina small. Uropod 1: the rami are long and slender, of unequal length (2/3). Uropod 2: rami short, of equal size, and armed with a row of spines on each ramus. Uropod 3: peduncle short and strong; rami of unequal length, ornamented with numerous setae. Telson: triangular, apically rounded, cleft on 2/3 of the length; two setae at the apex of each lobe. Distribution: North Atlantic Ocean; wide bathymetric range species: this species was collected at 2702m depth (Stephensen,1925), between 100–2900 m (Mills,1971), between 156–1808 m, on sand, sandy silt, silty sand, muddy sand, gravely sand, gravely sandy silt (BIOICE samples), one specimen at 1024 m on bathyal mud in the south of the Bay of Biscay (Dauvin & Bellan-Santini,1996). Taxonomic remarks. H. similis belongs to the sub-group of blind species with a narrow Pereopod 7 basis and without dorsal tuft setae, which includes six other species: H. abyssorum, H. vallifera, H. lodo, H. dauvini, H. bjarnii, and H. faroensis. It differs from these species by having: — Antennae: Antenna 1= 8–9/10 Antenna 2, Antenna 2 = 2/3 body length; the antennae are longer than the body length for H. bjarnii; they are half body length for H. vallifera, H. dauvini, and H. faroensis; for H. lodo, A1= ped A2, A2 shorter than body (in Barnard, 1961). — Coxa 4 heart-shaped (also the case for H. bjarnii and H. vallifera); but square shaped for H. abyssorum, H. lodo, H. dauvini and H. faroensis. — Epimeral plates 3 with posterior margin oblique and straight (also for H. faroensis); slightly convex for H. bjarnii and H. dauvini; rounded for H. vallifera and H. lodo (not indicated by Chevreux 1908 for H. abyssorum). — Uropod 1: the rami are long and slender, of unequal length (2/3); the rami are subequal for H. bjarnii, H. vallifera; H. lodo and H. dauvini; they have the same size for H. abyssorum and H. faroensis., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 166-168, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Mills, E. L. (1971) Deep-Sea Amphipoda from the Western North Atlantic Ocean, the family Ampeliscidae. Limnology and Oceanography, 16, 357 - 386. https: // doi. org / 10.4319 / lo. 1971.16.2.0357","Dauvin, J. C. & Bellan-Santini, D. (1996) Ampeliscidae (Amphipoda) from the Bay of Biscay. Journal of Crustacean Biology, 16, 149 - 168. https: // doi. org / 10.2307 / 1548938","Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de l a Princesse Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13."]}

Haploops vallifera Stephensen 1925. (Figures 9–12) BIOFAR material examined. Station 124, one specimen, 26 July 1987, depth 600 m, 62°16.94’N – 09°38.93’W; sediment type unknown. Station 261, nine specimens, 14 May 1988, depth 1003 m, 61°35.57’N – 09°35.47’W; sediment type unknown. Station 731, 182 specimens, 29 September 1990, depth 1042 m, 60°29.70’N – 07°14.10’W; bottom: sand, gravel, cobbles and stones. Station 738, one specimen, 1 October 1990, depth 749 m, 62°19.30’N – 10°13.30’W; bottom: gravel, cobbles and stones. Male unknown. Diagnosis. Body with few small dorsal setae on the pereon and the pleon. Corneal lenses absent. Segments of dorsal side of mesosome, marked. Urosome segment 1 with a small rounded carena. Epimeral plate 3 rounded. The two pairs of antennae of equal length not longer than half of body. Description. (Fig. 9–12). Adult female (Fig. 9) length 6 mm Biofar 731. Head: nearly square, with a small rostrum, without corneal lenses, blind species, straight lateral lobe. Antenna 1 (Fig. 9, 10 A): nearly equal to half of the body length. Peduncle with article 3 Antenna 2 (Fig. 9, 10 B): Equal to antenna 1 in length. Peduncle segment 5> 4. Flagellum with 12 articles. Mandible (Fig. 10 C): molar strong, palp very long, article 2> article 3, artcles 2, 3 bear long setae, article 3 with a long apical seta. Lower lip bilobate (Fig. 10 F). Maxilla 1 (Fig. 10 E): inner plate conical, outer plate with spine teeth distally, last article of palp ended with some setae and 5 strong teeth. Maxilla 2 (Fig. 10 G): the two plates are subequal in length and apically densely setose. Maxilliped (Fig. 10 D): inner plate sub-rectangular with short setae on the lateral and distal part. Internal margin of the outer plate with numerous long setae and strong teeth. Palp elongate, longer than outer plate, articles 2 and 3 with long setae, article 3 distally enlarged, dactylus elongate slender with 4 setae at the inferior margin. Pereon: small setae on the posterodorsal part of segment 6 and 7 (Fig. 9). Gnathopod 1 (Fig. 11A): longer than gnathopod 2. Coxa 1 as long as basis, distal margin rounded and fringed with long setae. Basis a little longer than carpus+propodus, setose. Merus, carpus and propodus bears long setae, dactylus curved with 5 short setae. Gnathopod 2 (Fig. 11B): Coxa 2 shorter than basis, triangular, distal margin pointed. Basis long with long setae, egal to merus+carpus+propodus. Carpus longer than propodus, roughtly rectangular with long setae mainly on the inferior margin. Propodus roughtly rectangular. Dactylus curved and ornamed with 5 setae. Pereopod 3 (Fig. 11C): Coxa 3 triangular, distal margin pointed. Basis rectangular, basis = ischium + merus + carpus, the margins of the different articles except dactylus, bearing long setae. Dactylus strong, curved, very long, longer than propodus. Pereopod 4 (Fig. 11D): Coxa 4 pointed, shorter and wider than coxa 3, posteriorly excavate, ventral margin straight with one seta on the distal part. Basis rectangular, longer than ischium + merus + carpus. Dactylus curved, as long as propodus. Long setae are present on the margins of all articles except dactylus. Pereopod 5 (Fig. 11E): coxa 5 bilobate, posterior margin rounded. Basis oval anterior margin fringed with short setae. Carpus sub-rectangular with a small postero-distal lobe, posterior and distal margins ornamented with 4 rows of spines. Propodus rectangular, longer than carpus. Dactylus very strong and curved. Pereopod 6 (Fig. 11F): Coxa 6 triangular, distal edge rounded. Basis pyriform with a small indentation at the posterodistal corner. Carpus rectangular with a small distal lobe, posterior and distal margins ornamented with 3 rows of spines. Pereopod 7 (Fig. 12A, B): coxa rectangular, distally rounded. Basis broad, width / length = 1/2, anterior and posterior margins with numerous long setae. Ischium quadrangular with 2 small strong spines on the posterior margin. Merus quadrangular but longer than width. Carpus shorter and narrower of merus with inferior corner elongate. Propodus and dactylus short. Pleon: Segment 1 and 2 with short setae (Fig. 9). Epimeral plate 1: short, ventral margin rounded. Epimeral plate 2 (Fig. 12 F): Rounded distal margin with 2 setae. Epimeral plate 3 (Fig. 12 F): anterior margin straight, anteroventral corner round, vental margin slightly convex, postero-ventral corner rounded, posterior margin convex with 4 setae. Urosome (Fig. 9): segment 1 with a small hump anteriorly at a high rounded dorsal carina. Uropod 1 (Fig. 12C): long, the rami unequal and slightly curved, rami inermous, peduncle longer than rami (peduncle/outer ramus/inner ramous = 40/36/30), ornamented with only one seta at the inner distal corner. Uropod 2 (Fig. 12D): rami subequal, equal to peduncle, one spine on the distal inner corner of the peduncle and 3 on the edge of the inner ramus. Uropod 3 (Fig. 12E): peduncle short and strong, rami foliaceous, longer than peduncle, inner ramus with 2 strong spines, outer ramus with long apical setae and some ones on the margin. Telson (Fig. 12 G): Triangular, rounded, slightly wide than long, cleft on 3/4 of the length, 2 or 3 long setae on each lobe. Known distribution: North Atlantic Ocean; wide bathymetric range species. BIOFAR material, this study, bathymetric range: 600–1042 m; the nature of the bottom is indicated for only two samples: sand, gravel, cobbles and stones. Offshore Iceland, 913–1960 m (Dauvin and Bellan-Santini,1990). Faroe Islands: 600-1098 m (Dauvin 1996), Iceland one station 1392 m (Bellan-Santini and Dauvin, 1997), south and west of Iceland, 13 stations from 285 to 1963 m (Dauvin et al., 2012). Taxonomic remarks. H. vallifera belongs to the sub-group of blind species with a narrow Pereopod 7 basis and without dorsal tuft setae which includes six other species: H. abyssorum, H. similis, H. lodo, H. dauvini, H. bjarnii, and H. faroensis. It differs from these species by having: — Dorsal side of pereon and urosome carinate. — Coxa 1 rectangular with the distal margin rounded. — Coxa 2 triangular, with the distal margin pointed., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 162-166, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Dauvin, J. C. & Bellan-Santini, D. (1990) An overview of the amphipod genus Haploops (Ampeliscidae). Journal of the Biological Association of the United Kingdom 70, 887 - 903. https: // doi. org / 10.1017 / S 0025315400059129","Bellan-Santini, D. & Dauvin, J. C. (1997) Ampeliscidae (Amphipoda) from Iceland with a description of a new species: Ampelisca islandica (Contribution to the BIOICE programme research). Journal of Natural History, 31, 1157 - 1173. https: // doi. org / 10.1080 / 00222939700770621","Dauvin, J. C., Alizier, S., Weppe, A. & Gudmundsson, G. (2012) Diversity and zoogeography of Icelandic deep-sea Ampeliscidae (Crustacea: amphipoda). Deep Sea Research I, 68, 12 - 23. https: // doi. org / 10.1016 / j. dsr. 2012.04.013"]}

Haploops truncata spec. nov. (Figures 5–8) Type material. HOLOTYPE. One female without oostegite. Length: 5.57 mm (Fig. 5). BIOICE: Station 3140; one specimen, 25 August 1999, depth: 768 m, 67°51.90’N – 22°14.89’W, bottom: sediment type unknown. Holotype IINH 42245 + slides IINH 42247. The specimen is deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2011: one specimen, 20 July 1991, depth: 768 m, 65°41.43’N – 11°16.77’W; sediment type unknown (IINH 42251). Station 2414: three specimens, 20 July 1991, depth 978 m, 65°35.01’N – 10°59.98’W; bottom: brown sandy, silt mixed with foraminifera (IINH 42250). Station 2897: eight specimens, 24 August 1996, depth 672 m, 65°29.44’N – 27°32.55’W; sediment type unknown (IINH 42249). Station 2898: one specimen, 24 August 1996, depth 672 m, 65°29.30’N – 27°32.70’W; sediment type unknown (IINH 42248). Station 3140: one specimen, 25 August 1999, depth 768 m, 67°51.90’N – 22°14.89’W; sediment type unknown (IINH 42245). Station 3501: two specimens, 31 August 2002, depth 829 m, 62°59.84’N – 20°30.25’W; sediment type unknown (IINH 42246). Male unknown. Ethymology. The species name refers to the shape of the head (square) with the lateral lobe truncate. Diagnosis. Pereon, Pleon and Urosome without long dorsal setae, Head square with anterior margin straight (lateral lobe truncate). One pair of superior corneal lenses. Antenna 1 length = 0.8 Antenna 2 length; Antenna 2 as long as body length (0.876). Coxa 4 external side bearing numerous short setae. Description. Holotype. Female without oostegites. Length: 5.57 mm (Fig. 5). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 6A): square with anterior margin straight, one pair of superior corneal lenses. Antenna 1 (Fig. 6B): shorter than antenna 2, peduncle of A1 with article 3 Antenna 2 (Fig. 6C): longer than antenna 1 (A1/A2 = 0.81). Peduncle segment 5 longer than 4 (23/19). Flagellum with 23 articles. Antenna 2 near the body length. The two antennae bearing long numerous setae (Fig. 6B, C). Upper lip and Lower lip bilobate (Fig. 6D, G). Mandible (Fig. 6H): molar strong; palp long, article 2 and 3 of same length; setae are present on the margin of each article, article 3 with very long apical setae. Maxilla 1 (Fig. 6E). the inner plate is conical shaped with two apical setae and some short hairs; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 6F): dense apically setae on the two plates. Maxilliped (Fig. 6J): inner plate sub-rectangular, elongated, with few distal setae; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, article 2 and 3 with long setae, article 3 triangular shaped, dactylus elongate and slender. Pereon: without long dorsal setae (Fig. 5). Gnathopod 1 (Fig. 7A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis long, slightly curved, approximately rectangular with some setae on the two margins, length longer than ischium + merus + carpus, the merus, carpus and propodus bearing long setae; propodus oval; dactylus slender and curved with small setae, dactylus length = 0.51 propodus length. Gnathopod 2 (Fig. 7B): coxa 2 approximately rectangular and short (coxa length / basis length= 65/85), distal margin slightly rounded with few short setae; basis long with some setae, basis longer than ischium + merus +carpus; carpus longer than propodus (carpus length / propodus length = 53/30), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 7C), dactylus length = 0.46 (17/37) propodus length. Pereopod 3 (Fig. 7D): coxa 3 triangular, distal margin rounded; basis rectangular, basis> ischium + merus + carpus. The margins of the different articles of pereopod 3, except dactylus, bearing few long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 1.35) (27/20). Pereopod 4 (Fig. 7E): coxa 4 square-shaped, external side bearing numerous short setae; anterior margin straight, length / width = 43/40, antero-ventral corner quadrate but not sharp, ventral margin straight with short setae, posterior concavity = 0.46 (17/37) length of the posterior length of the coxa 4, posterior hook broad, length = 17/42 width of coxa 4, posterior corner blunted; basis length> ischium + merus; the margins of the different articles of pereopod 4, except dactylus, bearing few long setae; dactylus slender, slightly curved and longer than propodus (dactylus length / propodus length = 1.125 (27/24). Pereopod 5 (Fig. 7F): coxa 5 rectangular, basis roughly rectangular with few setae on the anterior margin; carpus rectangular, anterior margin with few setae, posterior margin with two rows of little spines, distal margin with short spines and a long one; propodus rectangular, longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 0.23 (12/51). Pereopod 6 (Fig. 8A): coxa 6 roughly trapezoidal shaped, posterior margin rounded; basis rounded with few short setae and spines on the anterior margin; carpus rectangular, anterior margin with few short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a very long one; propodus a little longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 11/40. Pereopod 7 (Fig. 8B): coxa 7 roughly rectangular; basis narrow (length without lobe / width = 80/35 = 2.28), anterior and posterior margin slightly concave, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with an antero-inferior lobe ending with a spine, posterior margin ornamented with few little spines; carpus pyriform (width / length = 17/20), with strong spines on the anterior and posterior margin (propodus length / carpus length = 15/20; propodus + dactylus / carpus = 23/20); propodus narrow, width / length = 5/15 (propodus width / carpus width = 5/17), with short apical setae; dactylus rectangular, short and narrow (length / width = 7/1) with two very small apical setae (dactylus length / propodus length = 8/15). Pleon (Fig. 5): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 8C): anterior margin oblique and straight, ventral margin rounded, posterior margin slightly convex. Epimeral plate 2 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, postero-inferior corner rounded. Epimeral plate 3 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, corners rounded. Urosome (Fig. 5): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 8D): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the ramus, with one spine on the distal margin. length rami / length peduncle = 54–56 / 64. Uropod 2 (Fig. 8E): shorter than Uropod 1; rami triangular and short, inner ramus shorter than the outer one, with a row of spines on each ramus, outer ramus length / peduncle length = 78 /102; peduncle rectangular, robust with a strong spines on the distal margin. Uropod 3 (Fig. 8F): peduncle short and strong (length / width = 58/35); rami of unequal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 90–100/58); inner ramus with three spines and some apical setae, outer ramus with apical long setae and also some long setae on the outer margin. Telson (Fig. 8G): triangular, apically rounded, cleft on 25/35 of the length; one seta present on the apical part of each lobe. Distribution: North Atlantic Ocean; this species was collected between 672 m and 978 m. The nature of the bottom is indicated for only one sample: brown sandy silt mixed with foraminifera. Taxonomic remarks. H. truncata is probably a non mature female, but the original characters in the sub-group with superior corneal lenses justify its description as a new species. H. truncata belongs to the sub-group with the superior pair of corneal lenses clearly visible; the inferior pair is absent; a narrow Pereopod 7 basis, and the absence of dorsal tuft, this sub-group includes six other species: H. tubicola, H. spinosa, H. descansa, H. fundiensis, H. oonah, and H. antennata. H. truncata differs from the other neighbouring species by the following characters: — Head square with anterior margin straight (oblique in H. tubicola; H. spinosa; H. fundiensis; H. antennata). — Antennae subequal, Antenna 2 near the body length; in H. tubicola, the antennae are subequal and reach 2/3 of the body length; they reach 1/3 of the body length for H. fundiensis; in H. spinosa the Antenna 2 length is include between 1/3 and 1/2 of body length. For the two other species, H. descansa and H. oonah, the Antenna 1 is half the length of Antenna 2; whereas in H. antennata, the Antenna 1 is longer than Antenna 2. — Coxa 4 square-shaped, external side bearing numerous short setae. — Epimeral plate 3 square shaped; the posterior margin is oblique or slightly curved in the other species (except for H. descansa). — Uropod 1 long, with rami slender, curved, and of equal length; they are of unequal length for H. tubicola, H. descansa, H. fundiensis, H. oonah and H. antennata (except H. spinosa with rami of equal length, and especially as Uropod 2 rami are strongly armed). In the introduction we indicate that it is necessary to supplement the description of two species described by Stephensen in 1925: Haploops vallifera and Haploops similis. This is the object of the following section.

Haploops truncata spec. nov. (Figures 5–8) Type material. HOLOTYPE. One female without oostegite. Length: 5.57 mm (Fig. 5). BIOICE: Station 3140; one specimen, 25 August 1999, depth: 768 m, 67°51.90’N – 22°14.89’W, bottom: sediment type unknown. Holotype IINH 42245 + slides IINH 42247. The specimen is deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2011: one specimen, 20 July 1991, depth: 768 m, 65°41.43’N – 11°16.77’W; sediment type unknown (IINH 42251). Station 2414: three specimens, 20 July 1991, depth 978 m, 65°35.01’N – 10°59.98’W; bottom: brown sandy, silt mixed with foraminifera (IINH 42250). Station 2897: eight specimens, 24 August 1996, depth 672 m, 65°29.44’N – 27°32.55’W; sediment type unknown (IINH 42249). Station 2898: one specimen, 24 August 1996, depth 672 m, 65°29.30’N – 27°32.70’W; sediment type unknown (IINH 42248). Station 3140: one specimen, 25 August 1999, depth 768 m, 67°51.90’N – 22°14.89’W; sediment type unknown (IINH 42245). Station 3501: two specimens, 31 August 2002, depth 829 m, 62°59.84’N – 20°30.25’W; sediment type unknown (IINH 42246). Male unknown. Ethymology. The species name refers to the shape of the head (square) with the lateral lobe truncate. Diagnosis. Pereon, Pleon and Urosome without long dorsal setae, Head square with anterior margin straight (lateral lobe truncate). One pair of superior corneal lenses. Antenna 1 length = 0.8 Antenna 2 length; Antenna 2 as long as body length (0.876). Coxa 4 external side bearing numerous short setae. Description. Holotype. Female without oostegites. Length: 5.57 mm (Fig. 5). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 6A): square with anterior margin straight, one pair of superior corneal lenses. Antenna 1 (Fig. 6B): shorter than antenna 2, peduncle of A1 with article 3 Antenna 2 (Fig. 6C): longer than antenna 1 (A1/A2 = 0.81). Peduncle segment 5 longer than 4 (23/19). Flagellum with 23 articles. Antenna 2 near the body length. The two antennae bearing long numerous setae (Fig. 6B, C). Upper lip and Lower lip bilobate (Fig. 6D, G). Mandible (Fig. 6H): molar strong; palp long, article 2 and 3 of same length; setae are present on the margin of each article, article 3 with very long apical setae. Maxilla 1 (Fig. 6E). the inner plate is conical shaped with two apical setae and some short hairs; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 6F): dense apically setae on the two plates. Maxilliped (Fig. 6J): inner plate sub-rectangular, elongated, with few distal setae; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, article 2 and 3 with long setae, article 3 triangular shaped, dactylus elongate and slender. Pereon: without long dorsal setae (Fig. 5). Gnathopod 1 (Fig. 7A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis long, slightly curved, approximately rectangular with some setae on the two margins, length longer than ischium + merus + carpus, the merus, carpus and propodus bearing long setae; propodus oval; dactylus slender and curved with small setae, dactylus length = 0.51 propodus length. Gnathopod 2 (Fig. 7B): coxa 2 approximately rectangular and short (coxa length / basis length= 65/85), distal margin slightly rounded with few short setae; basis long with some setae, basis longer than ischium + merus +carpus; carpus longer than propodus (carpus length / propodus length = 53/30), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 7C), dactylus length = 0.46 (17/37) propodus length. Pereopod 3 (Fig. 7D): coxa 3 triangular, distal margin rounded; basis rectangular, basis> ischium + merus + carpus. The margins of the different articles of pereopod 3, except dactylus, bearing few long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 1.35) (27/20). Pereopod 4 (Fig. 7E): coxa 4 square-shaped, external side bearing numerous short setae; anterior margin straight, length / width = 43/40, antero-ventral corner quadrate but not sharp, ventral margin straight with short setae, posterior concavity = 0.46 (17/37) length of the posterior length of the coxa 4, posterior hook broad, length = 17/42 width of coxa 4, posterior corner blunted; basis length> ischium + merus; the margins of the different articles of pereopod 4, except dactylus, bearing few long setae; dactylus slender, slightly curved and longer than propodus (dactylus length / propodus length = 1.125 (27/24). Pereopod 5 (Fig. 7F): coxa 5 rectangular, basis roughly rectangular with few setae on the anterior margin; carpus rectangular, anterior margin with few setae, posterior margin with two rows of little spines, distal margin with short spines and a long one; propodus rectangular, longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 0.23 (12/51). Pereopod 6 (Fig. 8A): coxa 6 roughly trapezoidal shaped, posterior margin rounded; basis rounded with few short setae and spines on the anterior margin; carpus rectangular, anterior margin with few short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a very long one; propodus a little longer than carpus, with few short setae, and a long distal seta; dactylus short and curved, dactylus length / propodus length = 11/40. Pereopod 7 (Fig. 8B): coxa 7 roughly rectangular; basis narrow (length without lobe / width = 80/35 = 2.28), anterior and posterior margin slightly concave, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with an antero-inferior lobe ending with a spine, posterior margin ornamented with few little spines; carpus pyriform (width / length = 17/20), with strong spines on the anterior and posterior margin (propodus length / carpus length = 15/20; propodus + dactylus / carpus = 23/20); propodus narrow, width / length = 5/15 (propodus width / carpus width = 5/17), with short apical setae; dactylus rectangular, short and narrow (length / width = 7/1) with two very small apical setae (dactylus length / propodus length = 8/15). Pleon (Fig. 5): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 8C): anterior margin oblique and straight, ventral margin rounded, posterior margin slightly convex. Epimeral plate 2 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, postero-inferior corner rounded. Epimeral plate 3 (Fig. 8C): square shaped, anterior and posterior margin straight, inferior one slightly convex, corners rounded. Urosome (Fig. 5): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 8D): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the ramus, with one spine on the distal margin. length rami / length peduncle = 54–56 / 64. Uropod 2 (Fig. 8E): shorter than Uropod 1; rami triangular and short, inner ramus shorter than the outer one, with a row of spines on each ramus, outer ramus length / peduncle length = 78 /102; peduncle rectangular, robust with a strong spines on the distal margin. Uropod 3 (Fig. 8F): peduncle short and strong (length / width = 58/35); rami of unequal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 90–100/58); inner ramus with three spines and some apical setae, outer ramus with apical long setae and also some long setae on the outer margin. Telson (Fig. 8G): triangular, apically rounded, cleft on 25/35 of the length; one seta present on the apical part of each lobe. Distribution: North Atlantic Ocean; this species was collected between 672 m and 978 m. The nature of the bottom is indicated for only one sample: brown sandy silt mixed with foraminifera. Taxonomic remarks. H. truncata is probably a non mature female, but the original characters in the sub-group with superior corneal lenses justify its description as a new species. H. truncata belongs to the sub-group with the superior pair of corneal lenses clearly visible; the inferior pair is absent; a narrow Pereopod 7 basis, and the absence of dorsal tuft, this sub-group includes six other species: H. tubicola, H. spinosa, H. descansa, H. fundiensis, H. oonah, and H. antennata. H. truncata differs from the other neighbouring species by the following characters: — Head square with anterior margin straight (oblique in H. tubicola; H. spinosa; H. fundiensis; H. antennata). — Antennae subequal, Antenna 2 near the body length; in H. tubicola, the antennae are subequal and reach 2/3 of the body length; they reach 1/3 of the body length for H. fundiensis; in H. spinosa the Antenna 2 length is include between 1/3 and 1/2 of body length. For the two other species, H. descansa and H. oonah, the Antenna 1 is half the length of Antenna 2; whereas in H. antennata, the Antenna 1 is longer than Antenna 2. — Coxa 4 square-shaped, external side bearing numerous short setae. — Epimeral plate 3 square shaped; the posterior margin is oblique or slightly curved in the other species (except for H. descansa). — Uropod 1 long, with rami slender, curved, and of equal length; they are of unequal length for H. tubicola, H. descansa, H. fundiensis, H. oonah and H. antennata (except H. spinosa with rami of equal length, and especially as Uropod 2 rami are strongly armed). In the introduction we indicate that it is necessary to supplement the description of two species described by Stephensen in 1925: Haploops vallifera and Haploops similis. This is the object of the following section., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 157-161, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919

Key to species of Haploops 1 Corneal lenses present................................................................................. 2 - Corneal lenses absent................................................................................. 17 2 Two pairs of corneal lenses............................................................................. 3 - Only one pair of corneal lenses.......................................................................... 8 3 Pereopod 7 basis broad................................................................................. 4 - Pereopod 7 basis narrow................................................................................ 5 4 P7 carpus with anterior distal lobe; coxa1 inferior margin straight.......................... Haploops laevis Hoek, 1882 - P7 carpus without anterior distal lobe; coxa1 inferior margin rounded............................................................................................ Haploops quebecoisis Bellan-Santini Kaïm-Malka & Dauvin, 2018 5 Inferior lenses in the middle of the lobe head................................... Haploops sibirica Gurjanova, 1929 - Inferior lenses near or on the margin head.................................................................. 6 6 Inferior lenses near the margin head.......................................... Haploops dellavallei Chevreux, 1900 - Inferior lenses on the margin head........................................................................ 7 7 Inferior lenses on the margin, antennae = body length............................. Haploops nirae Kaïm-Malka, 1976 - Inferior lenses on the internal side of the margin, antennae = 1/2 body length........... Haploops tenuis Kanneworff, 1966 8 Only inferior lenses present (superior absent)............................................................... 9 - Only superior lenses present (inferior absent).............................................................. 10 9 Long setae on Md, back, telson, A1 Haploops longiseta Kaïm-Malka, 2010 - Few long dorsal setae, A1> A 2.............................................. Haploops proxima Chevreux, 1919 10 Pereopod 7 basis broad, A 2> body................................ Haploops gascogni Dauvin & Bellan-Santini, 1996 - Pereopod 7 basis narrow............................................................................... 11 11 Head truncated straight................................................................................ 12 - Head transversely truncated............................................................................ 15 12 A1> A 2.............................................................. Haploops antennata Kaïm-Malka, 2012 - A1 Haploops descansa Barnard, 1961 - A1 length> A2 peduncle.............................................................................. 14 14 A1 length a little more A 2 peduncle; coxa 1 oval; U1 rami unequal length.......... Haploops oonah Lowry & Poore, 1985 - A1= 2/3 A2, A2 = body; coxa 1 triangular; outer side coxa 4 setose; U1 rami equal length.... Haploops truncata spec. nov. 15 A 2 = 1/2 body; corneal lenses great size......................................... Haploops tubicola Liljborg, 1856 - A 2 = 1/3 body; corneal lenses small size.................................................................. 16 16. U1–2 strongly armed....................................................... Haploops spinosa Shoemaker 1931 - U1–2 weakly armed............................................. Haploops fundiensis Wildish & Dickinson, 1982 17 Pereopod 7 basis narrow............................................................................... 18 - Pereopod 7 basis broad................................................................................ 24 18 P7 basis, merus and carpus, rectangular and slender (male)....................... Haploops abyssorum Chevreux, 1908 - P7 basis, merus and carpus, not rectangular and slender...................................................... 19 19 Coxa 4 heart-shaped.................................................................................. 20 - Coxa 4 not heart-shaped............................................................................... 22 20 Dorsal side of the body carinate, coxa 2 acute................................... Haploops vallifera Stephensen,1925 - Dorsal side of the body not carinate...................................................................... 21 21 A1= 8–9/10 A 2, A2= 2/3 body............................................... Haploops similis Stephensen, 1925 - Antennae longer than the body length..................... Haploops bjarnii Bellan-Santini Kaïm-Malka & Dauvin, 2018 22 A1= A 2 peduncle, coxa1 oval.................................................... Haploops lodo Barnard, 1961 - Antennae Haploops dauvini Peart, 2018 - A1=A2=1/2 body length......................................................... Haploops faroensis spec. nov. 24 Long dorsal tuft of setae absent......................................................................... 25 - Long dorsal tuft of setae present........................................................................ 26 25 P7 basis lobe exceeding ischium distal edge; A1Haploops antarctica Bellan-Santini & Dauvin, 2008 - P7 basis lobe not exceeding ischium distal edge; A1=A2; coxa 4 heart-shaped............ Haploops kaimmalkai Peart, 2018 26 Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe triangular................... Haploops setosa Boeck, 1870 - Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe rounded............................................27 27 Urosome 1 high upturned carina.............................................. Haploops carinata Liljeborg, 1856 - Urosome 1 carina straight.............................................................................. 28 28 Antenna length> body length, antennae very strong...... Haploops islandica Kaïm-Malka,Bellan-Santini & Dauvin, 2016 - Antenna length =1/2 body length............................................... Haploops robusta G.O. Sars, 1891, Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on page 173, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Hoek, P. P. C. (1882) Die Crustaceen, gesammelt waehrend der Fahrten des \" Willem Barents \" in den Jahren 1878 und 1879, Niederlandisches Archiv fur Zoologie, Supplement band I, 7, 61.","Gurjanova, E. F. (1929) Neue Forrnen arktischer Isopoden und Amphipoden. Zoologischer Anzeiger, 81, 309 - 317.","Chevreux, E. (1900) Amphipodes provenant des campagnes de l'Hirondelle (1885 - 1888). Resultats des Campagnes Scientifiques Accomplies par le Prince Albert I. Monaco. 16, 195 pp.","Kaim-Malka, R. A. (1976) Revision des Haploops (Crustacea, Amphipoda) de Mediterranee. Bolletino del Museo Civico di Storia Naturale, Verona, 3, 269 - 308.","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207.","Kaim-Malka, R. A. (2010) Haploops longiseta, a new species from the Atlantic Ocean (Crustacea, Gammaridea, Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 6]. Zootaxa, 2356 (1), 57 - 68. https: // doi. org / 10.11646 / zootaxa. 2356.1.3","Chevreux, E. (1919) Note preliminaire sur les Amphipodes recueillis par les expeditions du \" Travailleur \" et du \" Talisman \" (1880 - 1883). Bulletin du Museum d'Histoire Naturelle, 25, 574 - 580. https: // doi. org / 10.5962 / bhl. part. 7932","Dauvin, J. C. & Bellan-Santini, D. (1996) Ampeliscidae (Amphipoda) from the Bay of Biscay. Journal of Crustacean Biology, 16, 149 - 168. https: // doi. org / 10.2307 / 1548938","Kaim-Malka, R. A. (2012) Haploops antennata, a new species from the North Atlantic Ocean (Crustacea: Gammaridea: Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 7]. Zootaxa, 3320 (1), 36 - 46. https: // doi. org / 10.11646 / zootaxa. 3320.1.2","Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Lowry, J. K. & Poore, G. C. B. (1985) The ampeliscid amphipods of south-eastern Australia (Crustacea). Records of the Australian Museum, 36, 259 - 298. https: // doi. org / 10.3853 / j. 0067 - 1975.36.1985.348","Shoemaker, C. R. (1931) The Stegocephalid and Ampeliscid Amphipod Crustaceans of Newfoundland, Nova Scotia, and New Brunswick in t he United States National Museum. Proceedings of the United States National Museum, 79 (2888), 1 - 18, 6 figs. https: // doi. org / 10.5479 / si. 00963801.79 - 2888.1","Wildish, D. J. & Dickinson, J. J. (1982) A new species of Haploops (Amphipoda, Ampeliscidae) from the Bay of Fundy. Canadian Journal of Zoology, 60, 962 - 967. https: // doi. org / 10.1139 / z 82 - 133","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de l a Princesse Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13.","Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Peart, R. A. (2018) Ampeliscidae (Crustacea, Amphipoda) from the IceAGE expeditions. Zookeys, 731, 145 - 173, 2 tabs. https: // doi. org / 10.3897 / zookeys. 731.19948","Boeck, A. (1870) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Videnskabs-Selskabet i Christiania 1870, pp. 83 - 280. https: // doi. org / 10.5962 / bhl. title. 2056","Liljeborg, W. (1856) Om Hafs-Crustaceer vid Kullaberg i Skane. Ofversigt af Kongliga Vetenskaps - Akademien Forhandlingar, Stockholm, 12, 117 - 138.","Kaim-Malka, R., Bellan-Santini, D. & Dauvin, J. C. (2016) On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8]. Zootaxa, 4179 (1), 42 - 76. https: // doi. org / 10.11646 / zootaxa. 4179.1.2","Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway with short descriptions and figures of all the species, 1, 191 - 196, pls. 67 - 68."]}

Haploops faroensis spec. nov. (Figures 1–4) Type material. HOLOTYPE. One adult female with oostegites not completely developed. Length: 9.18 mm (Fig.1). BIOFAR: Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’ W; bottom: clay and silt, sand, gravel. Holotype + slides (NHMD – 873179). The specimen is deposited in the Natural History Museum of Denmark (Danish Statens Naturhistoriske Museum). BIOFAR material. The specimens are deposited in the Natural History Museum of Denmark. Station 269, five specimens, 15 May 1988, depth: 510 m, 62° 49.84’N – 08° 15.55’ W; bottom: mud, sand, cobbles and stones (NHMD-873180). Station 271, four specimens, 16 May 1988, depth: 559 m, 62° 52.30’N – 08° 09.24’W; bottom: soft bottom, foraminifers (NHMD-873181). Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’W; bottom: clay and silt, sand, gravel (NHMD-873179). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2673, nine specimens, 15 July 1994, depth: 227– 222 m, 66° 50.20’N – 16°15.74’W; bottom: sponge spicules. (IINH 42254). Station 3522, five specimens, 7 September 2002, depth: 1940 m, 62°31.14’N – 17° 09.87’W, bottom: sediment type unknown (IINH 42253). Station 3544, one specimen, 11 September 2002, depth: 1635– 1632 m, 61°33.00’N – 13°40.14’W, bottom: gravely sand. (IINH 42252). Male unknown. Etymology. The species name refers to the area of the Faroe Islands where the species was collected. Diagnosis. Blind species. Pereon, Pleon and Urosome without long dorsal setae. Antenna 1 and Antenna 2 have the same length. Antennae half length of the body. Pereopod 7 basis narrow. Description. Holotype. Adult female, 9.18 mm (Fig.1). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 2A): rectangular, with a short rostrum pointed, without corneal lenses, blind species. Anterior margin oblique and straight. Antenna 1 (Fig. 2B): as long as antenna 2, half length of the body, peduncle of A1 with article 3 Antenna 2 (Fig. 2C): as long as antenna 1. Peduncle segment 5 longer than 4 (24/20). Flagellum with 17 articles. The two antennae bearing long setae. Upper lip, and Lower lip bilobate. Mandible (Fig. 2D): molar strong; palp long, article 3 longer than article 2 (3/2 = 40/35), setae are present on the margin of each article, article 3 with 3 long apical setae. Maxilla 1 (Fig. 2G): the inner plate is conical shaped with one long apical seta; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 2H): dense apically setae on the two plates. Maxilliped (Fig. 2E): inner plate sub-rectangular, elongated; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, articles 2 and 3 with long setae, article 3 oval shape, dactylus elongate and slender (Fig. 2F). Pereon: without long dorsal setae (Fig. 1). Gnathopod 1 (Fig. 3A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis as long as ischium + merus + carpus, more broad on the distal part than proximal one, with some setae on the two margins, the merus, carpus and propodus bearing long setae; propodus oval; dactylus strongly curved, (Fig. 3B), dactylus length = 0.6 propodus length. Gnathopod 2 (Fig. 3C): coxa 2 approximately rectangular and short (coxa length / basis length = 30/33), distal margin straight with few short setae; basis long, rectangular, with some long setae, basis = ischium + merus + carpus; carpus longer than propodus (carpus length / propodus length = 26/15), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 3D), dactylus length = 1/3 propodus length. Pereopod 3 (Fig. 3E): coxa 3 rectangular, distal margin rounded; basis rectangular, basis longer than ischium + merus + carpus. The margins of the different articles of pereopod 3, except ischium and dactylus, bearing few long setae; dactylus slender, curved and as long as propodus (dactylus length / propodus length = 1). Pereopod 4 (Fig. 3F): coxa 4 square-shaped, anterior margin straight, length / width = 40/34, antero-ventral corner rounded, ventral margin slightly curved with short setae, posterior concavity 1/2 length of the posterior length of the coxa 4, posterior hook broad, length = 12/34 width of coxa 4, posterior corner rounded; basis longer than ischium + merus + carpus; the margins of the different articles of pereopod 4, except dactylus, bearing many long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 21/17). Pereopod 5 (Fig. 3G): coxa 5 roughly rectangular, bi-lobated; basis oval with few short setae on the anterior margin; carpus rectangular, anterior margin with two short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long seta; propodus rectangular a little longer than carpus, with few short setae on the anterior margin, and long distal setae; dactylus slender and curved, dactylus length / propodus length = 12/40 (0.3). Pereopod 6 (Fig. 4A): coxa 6 roughly rectangular, posterior margin rounded; basis rounded with few short setae on the anterior margin; carpus rectangular, anterior margin with few short spines, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long one; propodus rectangular a little shorter than carpus, with few short setae, and a long distal seta; dactylus slender and curved, dactylus length / propodus length = 8/21. Pereopod 7 (Fig. 4B): coxa 7 roughly rectangular, posterior margin rounded; basis narrow (length without lobe / width = 40/20), anterior margin slightly concave, numerous long setae on posterior margin, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with spines on the margins and long setae on the posterior one; carpus rectangular (width / length = 8/12), with small spines on the anterior and posterior margins and one spine on the posterodistal corners (propodus length / carpus length = 4/12; propodus + dactylus / carpus = 8/12); propodus narrow, width / length = 2/4 (propodus width / carpus width = 2/8), with a short apical seta; dactylus rectangular, short and narrow (length / width = 3/1) with two very small apical setae (dactylus length / propodus length = 3/4). Pleon (Fig. 1): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 4G): anterior margin oblique and straight, ventral margin and posterior one straight, inferior corners rounded. Epimeral plate 2 (Fig. 4G): rounded. Epimeral plate 3 (Fig. 4G): anterior margin straight, antero-ventral corner rounded, ventral margin straight, postero-ventral corner acute, posterior margin oblique and straight. Urosome (Fig. 4H): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 4C): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the rami, with 1 spine on the distal margin. length rami / length peduncle = 38 / 52. Uropod 2 (Fig. 4D): shorter than uropod 1; rami triangular, inner ramus longer than the outer one, with a row of five little spines on inner ramus and only two spines on outer, outer ramus length / peduncle length = 36 /41; peduncle rectangular, robust with spines on the inner margin. Uropod 3 (Fig. 4E): peduncle short and strong (length / width = 25/15); rami of equal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 35/25); inner ramus with three spines and some apical setae, outer ramus with an apical tuft of long setae and also some long setae on the outer margin. Telson (Fig. 4F): rectangular, apically rounded, cleft on 33/45 of the length; one spine and one seta are present on the apical part of each lobe. Distribution: North Atlantic Ocean; wide bathymetric range species: 222–1940 m, present mainly in soft sediments: mud, sand, gravely sand, clay and silt, gravel, cobbles and stones, foraminifers, sponge spicules. Taxonomic remarks. H. faroensis spec. nov. belongs to the sub-group of blind species with a narrow pereopod 7 basis and without dorsal tuft seta e. This sub-group includes six other species: H. abyssorum, H. similis, H. vallifera, H. lodo, H. dauvini and H. bjarnii. It differs from these species in having: — Antennae of the same length and as long as half body length; Antenna 2 = 2/3 body length for H. similis; the antennae are longer than the body length for H. bjarnii; for H. lodo, A1= ped A2, A2 shorter than body (in Barnard 1961) (they are half body length for H. vallifera, H. dauvini and H. faroensis), — Coxa 4 square-shaped, heart-shaped for H. bjarnii, H. vallifera, H. similis, but also square- shaped for H. abyssorum, H. lodo and H. dauvini. — Epimeral plate 3 with posterior margin oblique and straight (also for H. similis); slightly convex for H. bjarnii, H. dauvini, rounded for H. vallifera, H. lodo (not indicated by Chevreux 1908 for H. abyssorum). — Uropod 1: rami of same size (also for H. abyssorum), they are of unequal length (2/3 for H. similis), the rami are subequal for H. bjarnii, H. vallifera, H. lodo and H. dauvini.

Haploops spinosa Shoemaker 1931. Shoemaker (1931) described Haploops spinosa in the amphipods collections of the United States National Museum, indicating that this species is morphologically close to H. tubicola, being often confused with it, but differs by several characters. He indicated that all the specimens studied were females, and the illustration of antennae, confirm that the specimens were females. Later Kanneworff (1966) stated that H. spinosa Shoemaker 1931 was partly established because Shoemaker’s specimens differed from Sars’ illustrations of H. tubicola, and considered that the two species were synonymous. In his opinion Shoemaker was describing males of H. tubicola (as H. spinosa). However, this assertion is not correct, if we consider the previous indications of Shoemaker 1931. Kanneworff (1966), Dickinson (1983), Barnard & Karaman (1991) had supported this synonymy between H. spinosa and H. tubicola. We have examined in detail some specimens from the BIOICE collections which show morphological characters close to the species described as H. spinosa by Shoemaker (1931). BIOICE material examined. Station 2075: one adult female, great size, oostegits presents; 3 July 1992; depth 563–572 m; 67° 11.65’N – 17° 32. 04’W; bottom: silty sand, gravel, stone. Station 2903: three specimens; 24 August 1996; depth: 1066– 1057 m; 65° 22.80’N – 26° 20.63’W; bottom: sandy silt. Station 2950: one adult female, great size; 29 August 1996; depth: 153 m; 65°42.10’N – 25°16.21’W; bottom: sand. After detailed examination, we consider that Haploops spinosa should be re-established as a valid species. The main characters of the large-sized female collected at the BIOICE 2950 station are as follows: — Only superior corneal lenses present (inferior absent) and corneal lenses being of small size. — Antennae 1 a little shorter than the Antennae 2. — Antennae 2 length (L), 1/3 Distribution: Bay of Fundy, Nova Scotia Coast; depth 12–1255 fathoms (around 20–2300 m) (Shoemaker,1931); Iceland area, 153–1066 m (this study). Taxonomic remarks. Three Haploops species with only superior corneal lenses are morphologically close: H. tubicola, H. spinosa, H. fundiensis. These species can be separated mainly by the following characters: H. tubicola: corneal lenses of great size; A2 = 2/3 body; Uropods 1 and 2 unequal sized rami, moderatly armed. H. fundiensis: small corneal lenses; antennae short, A2 =1/3 body; Uropods 1 and 2 equal sized rami, weakly armed. H. spinosa: small corneal lenses; A2 length include between 1/3 and 1/2 body length; Uropods 1 and 2, rami of same size, strongly armed., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on page 168, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Shoemaker, C. R. (1931) The Stegocephalid and Ampeliscid Amphipod Crustaceans of Newfoundland, Nova Scotia, and New Brunswick in t he United States National Museum. Proceedings of the United States National Museum, 79 (2888), 1 - 18, 6 figs. https: // doi. org / 10.5479 / si. 00963801.79 - 2888.1","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207.","Dickinson, J. J. (1983) The systematics and distributional ecology of the superfamily Ampeliscoidea (Amphipoda: Gammaridea) in the northeastern Pacific region. 11. The genera Byblis and Haploops. Publications in Natural scicnces, National Museum of Natural Sciences. Vol. 1. National Museums of Canada, Ottawa, 38 pp., 17 figs, 6 tabs.","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, Supplement 13, Part 1, 84 - 90. https: // doi. org / 10.3853 / j. 0812 - 7387.13.1991.91"]}

Haploops faroensis spec. nov. (Figures 1–4) Type material. HOLOTYPE. One adult female with oostegites not completely developed. Length: 9.18 mm (Fig.1). BIOFAR: Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’ W; bottom: clay and silt, sand, gravel. Holotype + slides (NHMD – 873179). The specimen is deposited in the Natural History Museum of Denmark (Danish Statens Naturhistoriske Museum). BIOFAR material. The specimens are deposited in the Natural History Museum of Denmark. Station 269, five specimens, 15 May 1988, depth: 510 m, 62° 49.84’N – 08° 15.55’ W; bottom: mud, sand, cobbles and stones (NHMD-873180). Station 271, four specimens, 16 May 1988, depth: 559 m, 62° 52.30’N – 08° 09.24’W; bottom: soft bottom, foraminifers (NHMD-873181). Station 299, one specimen, 17 July 1988, depth: 923 m, 60° 10.80’N – 08° 17.30’W; bottom: clay and silt, sand, gravel (NHMD-873179). BIOICE material. The specimens are deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Station 2673, nine specimens, 15 July 1994, depth: 227– 222 m, 66° 50.20’N – 16°15.74’W; bottom: sponge spicules. (IINH 42254). Station 3522, five specimens, 7 September 2002, depth: 1940 m, 62°31.14’N – 17° 09.87’W, bottom: sediment type unknown (IINH 42253). Station 3544, one specimen, 11 September 2002, depth: 1635– 1632 m, 61°33.00’N – 13°40.14’W, bottom: gravely sand. (IINH 42252). Male unknown. Etymology. The species name refers to the area of the Faroe Islands where the species was collected. Diagnosis. Blind species. Pereon, Pleon and Urosome without long dorsal setae. Antenna 1 and Antenna 2 have the same length. Antennae half length of the body. Pereopod 7 basis narrow. Description. Holotype. Adult female, 9.18 mm (Fig.1). Body without long dorsal setae on the pereon, pleon and urosome. Head (Fig. 2A): rectangular, with a short rostrum pointed, without corneal lenses, blind species. Anterior margin oblique and straight. Antenna 1 (Fig. 2B): as long as antenna 2, half length of the body, peduncle of A1 with article 3 Antenna 2 (Fig. 2C): as long as antenna 1. Peduncle segment 5 longer than 4 (24/20). Flagellum with 17 articles. The two antennae bearing long setae. Upper lip, and Lower lip bilobate. Mandible (Fig. 2D): molar strong; palp long, article 3 longer than article 2 (3/2 = 40/35), setae are present on the margin of each article, article 3 with 3 long apical setae. Maxilla 1 (Fig. 2G): the inner plate is conical shaped with one long apical seta; outer plate with spine teeth distally; palp ended with strong teeth and spines. Maxilla 2 (Fig. 2H): dense apically setae on the two plates. Maxilliped (Fig. 2E): inner plate sub-rectangular, elongated; internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, articles 2 and 3 with long setae, article 3 oval shape, dactylus elongate and slender (Fig. 2F). Pereon: without long dorsal setae (Fig. 1). Gnathopod 1 (Fig. 3A): coxa 1 roughly triangular, distal margin rounded and fringed with long setae; basis as long as ischium + merus + carpus, more broad on the distal part than proximal one, with some setae on the two margins, the merus, carpus and propodus bearing long setae; propodus oval; dactylus strongly curved, (Fig. 3B), dactylus length = 0.6 propodus length. Gnathopod 2 (Fig. 3C): coxa 2 approximately rectangular and short (coxa length / basis length = 30/33), distal margin straight with few short setae; basis long, rectangular, with some long setae, basis = ischium + merus + carpus; carpus longer than propodus (carpus length / propodus length = 26/15), approximately rectangular with long setae on the margins; propodus oval, long setae on the margins; dactylus slightly curved with short setae (Fig. 3D), dactylus length = 1/3 propodus length. Pereopod 3 (Fig. 3E): coxa 3 rectangular, distal margin rounded; basis rectangular, basis longer than ischium + merus + carpus. The margins of the different articles of pereopod 3, except ischium and dactylus, bearing few long setae; dactylus slender, curved and as long as propodus (dactylus length / propodus length = 1). Pereopod 4 (Fig. 3F): coxa 4 square-shaped, anterior margin straight, length / width = 40/34, antero-ventral corner rounded, ventral margin slightly curved with short setae, posterior concavity 1/2 length of the posterior length of the coxa 4, posterior hook broad, length = 12/34 width of coxa 4, posterior corner rounded; basis longer than ischium + merus + carpus; the margins of the different articles of pereopod 4, except dactylus, bearing many long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 21/17). Pereopod 5 (Fig. 3G): coxa 5 roughly rectangular, bi-lobated; basis oval with few short setae on the anterior margin; carpus rectangular, anterior margin with two short setae, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long seta; propodus rectangular a little longer than carpus, with few short setae on the anterior margin, and long distal setae; dactylus slender and curved, dactylus length / propodus length = 12/40 (0.3). Pereopod 6 (Fig. 4A): coxa 6 roughly rectangular, posterior margin rounded; basis rounded with few short setae on the anterior margin; carpus rectangular, anterior margin with few short spines, posterior margin with two rows of little spines, postero-distal lobe ornamented with short spines and a long one; propodus rectangular a little shorter than carpus, with few short setae, and a long distal seta; dactylus slender and curved, dactylus length / propodus length = 8/21. Pereopod 7 (Fig. 4B): coxa 7 roughly rectangular, posterior margin rounded; basis narrow (length without lobe / width = 40/20), anterior margin slightly concave, numerous long setae on posterior margin, lobe slightly deflected, rounded with few long setae, reaching the merus; ischium quadrangular; merus rectangular with spines on the margins and long setae on the posterior one; carpus rectangular (width / length = 8/12), with small spines on the anterior and posterior margins and one spine on the posterodistal corners (propodus length / carpus length = 4/12; propodus + dactylus / carpus = 8/12); propodus narrow, width / length = 2/4 (propodus width / carpus width = 2/8), with a short apical seta; dactylus rectangular, short and narrow (length / width = 3/1) with two very small apical setae (dactylus length / propodus length = 3/4). Pleon (Fig. 1): the postero-dorsal segments of the pleon without setae. Epimeral plate 1 (Fig. 4G): anterior margin oblique and straight, ventral margin and posterior one straight, inferior corners rounded. Epimeral plate 2 (Fig. 4G): rounded. Epimeral plate 3 (Fig. 4G): anterior margin straight, antero-ventral corner rounded, ventral margin straight, postero-ventral corner acute, posterior margin oblique and straight. Urosome (Fig. 4H): the urosome segment 1 has a dorsal carina straight, moderately hight, with the apex rounded. Uropod 1 (Fig. 4C): long, rami slender, curved, and equal length; inner ramus with two little spines; peduncle longer than the rami, with 1 spine on the distal margin. length rami / length peduncle = 38 / 52. Uropod 2 (Fig. 4D): shorter than uropod 1; rami triangular, inner ramus longer than the outer one, with a row of five little spines on inner ramus and only two spines on outer, outer ramus length / peduncle length = 36 /41; peduncle rectangular, robust with spines on the inner margin. Uropod 3 (Fig. 4E): peduncle short and strong (length / width = 25/15); rami of equal length, roughly rectangular, longer than peduncle (rami length / peduncle length = 35/25); inner ramus with three spines and some apical setae, outer ramus with an apical tuft of long setae and also some long setae on the outer margin. Telson (Fig. 4F): rectangular, apically rounded, cleft on 33/45 of the length; one spine and one seta are present on the apical part of each lobe. Distribution: North Atlantic Ocean; wide bathymetric range species: 222–1940 m, present mainly in soft sediments: mud, sand, gravely sand, clay and silt, gravel, cobbles and stones, foraminifers, sponge spicules. Taxonomic remarks. H. faroensis spec. nov. belongs to the sub-group of blind species with a narrow pereopod 7 basis and without dorsal tuft seta e. This sub-group includes six other species: H. abyssorum, H. similis, H. vallifera, H. lodo, H. dauvini and H. bjarnii. It differs from these species in having: — Antennae of the same length and as long as half body length; Antenna 2 = 2/3 body length for H. similis; the antennae are longer than the body length for H. bjarnii; for H. lodo, A1= ped A2, A2 shorter than body (in Barnard 1961) (they are half body length for H. vallifera, H. dauvini and H. faroensis), — Coxa 4 square-shaped, heart-shaped for H. bjarnii, H. vallifera, H. similis, but also square- shaped for H. abyssorum, H. lodo and H. dauvini. — Epimeral plate 3 with posterior margin oblique and straight (also for H. similis); slightly convex for H. bjarnii, H. dauvini, rounded for H. vallifera, H. lodo (not indicated by Chevreux 1908 for H. abyssorum). — Uropod 1: rami of same size (also for H. abyssorum), they are of unequal length (2/3 for H. similis), the rami are subequal for H. bjarnii, H. vallifera, H. lodo and H. dauvini., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, J. C., 2021, Complement to the knowledge of the Haploops species (Crustacea, Gammaridea Ampeliscidae), with the description of two new species from North Atlantic Ocean [Contribution to the knowledge of the Haploops genus. 10.], pp. 151-175 in Zootaxa 5048 (2) on pages 152-157, DOI: 10.11646/zootaxa.5048.2.1, http://zenodo.org/record/5551919, {"references":["Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de l a Princesse Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13."]}

Genus Haploops Lilljeborg, 1855 Haploops Lilljeborg, 1855:135; Barnard, 1969: 132; Karaman, 1975: 57; Lincoln, 1979: 124; Bellan-Santini, 1982: 64; Barnard & Karaman, 1991: 90. Type species. Haploops tubicola Lilljeborg, 1855, original designation. Diagnosis. Flagella of antennae 1���2 with 5 or more articles. Article 3 of maxillipedal palp produced or inflated. Article 2 of pereopod 7 variable, usually narrow and with anterior and posterior margins parallel, or posterior oblique and article 2 expanding ventrally, or article 2 tapering, anterior margin of posteroventral lobe near junction with article 2 (?) setose or face of lobe with large setae. Telson varying from scarcely longer to as long as, or shorter than broad, cleft much more than half its length (Barnard & Karaman, 1991). Distribution. Marine, cosmopolitan, cold water, submergent, 6���3570 m (Barnard & Karaman, 1991)., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Lilljeborg, W. (1855) Om Hafs-Crustaceer vid Kullaberg i Skane. Ofversigt af Kongliga Vetenskaps - Akademien Forhandlingar (Stockholm 1856), 12, 117 - 138.","Barnard, J. L. (1969) The families and genera of marine gammaridean Amphipoda. Bulletin of the United States national Museum, 271, 1 - 535, 173 figs.","Karaman, G. S. (1975) The family Ampeliscidae of the Adriatic Sea. Acta Adriatica, Institut Za Oceanografiju I Ribarstvo, Split, 17, (3), 67 pp, 32 figs.","Lincoln, R. J. (1979) British Marine Amphipoda: Gammaridea. London: British Museum (Natural History), 124 - 126, fig. 53 - 54.","Bellan-Santini, D. (1982). Family Ampeliscidae. In: Ruffo, S. (Ed.), The Amphipoda of the Mediterranean, Part 1. Memoires de l'Institut oceanographique, Monaco, 13, 19 - 69.","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, supplement 13, part 1, 84 - 90."]}

Haploops carinata Lilljeborg, 1855 Haploops carinata Lilljeborg, 1855 Haploops carinata Lilljeborg (in Boeck, 1870) Haploops setosa Boeck (in G.O.Sars, 1891) Haploops setosa Boeck (in part in Gurjanova, 1951) Haploops setosa Boeck (in Mills 1971) Haploops setosa Boeck (in Lincoln 1979) nec Haploops tubicola Lilljeborg 1855 (in Stebbing 1906; in Barnard & Karaman 1991). nec Ampelica eschrichti Kr��yer 1842 (in Lilljeborg 1855), Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Lilljeborg, W. (1855) Om Hafs-Crustaceer vid Kullaberg i Skane. Ofversigt af Kongliga Vetenskaps - Akademien Forhandlingar (Stockholm 1856), 12, 117 - 138.","Boeck, A. (1870) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Videnskabs - Selskabet i Christiania 1870, pp. 83 - 280. http: // dx. doi. org / 10.5962 / bhl. title. 2056","Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway With Short Descriptions and Figures of All the Species, 1, 191 - 196, pls. 67 - 68.","Gurjanova, E. F (1951) Bokoplavy Morei SSSR e sopredelny vod (Amphipoda- Gammaridea). Akademiia Nauk SSSR, Opedeliteli po Faune SSSR, 41, 1029 pp, 705 figs.","Mills, E. L. (1971) Deep-Sea Amphipoda from the Western North Atlantic Ocean, the family Ampeliscidae. Limnology and Oceanography, 16, 3377 - 386. http: // dx. doi. org / 10.4319 / lo. 1971.16.2.0357","Lincoln, R. J. (1979) British Marine Amphipoda: Gammaridea. London: British Museum (Natural History), 124 - 126, fig. 53 - 54.","Stebbing, TRR. (1906) Amphipoda 1. Gammaridea. Das Tierreich, 21, 806 pp., 127 figs.","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, supplement 13, part 1, 84 - 90."]}

Haploops islandica nov. sp. Type material. HOLOTYPE. One adult female with oostegites not completely developped. Length: 16.24 mm (Fig. 15). Bioice 3625, 15 July 2004, depth: 1630��� 1627 m, 66��59' 03'' N���8��48' 06'' W.���Holotype IMNHR-33741 + slides IMNHR-33742. This specimen is deposited in the Icelandic Museum of Natural History in Reykjavik (IMNHR). Biofar material. Station 343, 5 specimens, 22 July 1988, depth: 710 m, 62��14.6���N ��� 09��32.6 W, bottom: coarse sand, cobbles and stones. Bioice material Station 2937, 1 specimen, 28 August 1996, depth: 285��� 283 m, 65�� 55' 90'' N���26�� 41' 70'' W.(IMNHR-33743) Station 3198, 1 specimen, 0 8 July 2001, depth: 935 m, 65�� 30' 50'' N��� 08�� 31' 30'' W. (IMNHR-33744) Station 3199, 1 specimen, 0 8 July 2001, depth: 934���938 m, 65�� 30' 08'' N���08�� 31' 59'' W, sandy silt. (IMNHR- 33745) Station 3515, 3 specimens, 0 4 September 2002, depth 1331���1334 m, 62�� 22' 20'' N���18�� 23' 35'' W. (IMNHR- 33746) Station 3595, 1 specimen, 10 September 2003, depth 916���917 m, 61�� 49' 58'' N���26�� 09' 97'' W, sponges spicules. (IMNHR-33747) Station 3596, 1 specimen, 10 September 2003, depth 895���914 m, 61�� 49' 73'' N���26�� 10' 63'' W, with sponges, corals and echinoderms. (IMNHR-33748) Station 3597, 2 specimens, 10 September 2003, depth: 774��� 765 m, 62�� 17' 43'' N���25�� 37' 52'' W, silty sand.(IMNHR-33749) Station 3599, 1 specimen, 10 September 2003, depth: 765���784 m, 62�� 17' 38'' N���25�� 37' 68'' W.(IMNHR- 3350) Station 3625, 1 specimen, 15 July 2004, depth: 1630��� 1627 m, 66��59' 03'' N���08��48' 06'' W. (IMNHR-33741; 33742) Station 3632, 4 specimens, 16 July 2004, depth: 1727��� 1726 m, 68��00' 92'' N���09��14' 78'' W, sponges (IMNHR- 33751) Station 3657, 18 specimens, 23 July 2004, depth: 1499���1501 m, 68��45' 37'' N���15��22' 80'' W.(IMNHR-33752) Station 3657, 2 specimens, 23 July 2004, depth: 1499���1501 m, 68��45' 37'' N��� 15��22' 80'' W (MSNVR Cr 586)+ slides (MSNVR Cr 587). These specimens and slides are deposited in the Museo Civico di Storia Naturale di Verona (Italia). Etymology. The name species refers to the Iceland where the species was collected. Diagnosis. Blind species. Antennae as long as the body length. Peduncles of Antenna 1 and Antenna 2 long and strong. Pereon and Pleon with long setae (broken). Coxa 2 very short, Basis Gnathopod 2 long. Dactylus Pereopod 3 and 4 very long. Pereopod 7 basis moderatly broad, lobe slightly deflected, rounded. Carina straight, moderately hight. Telson with long apical setae. Description. Holotype. Adult female, 16.24 mm (Fig. 15). Body with long dorsal setae (broken) on the Pereon segment 5���7 and Pleon segment 1���3. Head: nearly square, with a rostrum pointed, without corneal lenses, blind species. Anterior margin slightly convex (Fig. 17 A). Antenna 1: as long as antenna 2, and as long as the body length. Peduncle of A1 with article 3 Antenna 2: as long as antenna 1, and as long as the body. Peduncle segment 5 longer than 4 (23 / 19). Flagellum with 37 articles (Fig. 17 C). The peduncles of Antenna 1 and Antenna 2 are long and strong. Peduncles and flagellum are densely setose with very long setae (Figs. 15 & 16). Upper lip: as in the other Haploops species. Mandible (Fig. 17 D): molar strong; palp very long, article 2 longer than article 3 (art.2/art.3 = 3/2,4), each article bears long setae, article 3 with 3 long apical setae. Lower lip bilobed (Fig. 17 E). Maxilla 1 (Fig. 17 F): the inner plate is conical shaped with two apical setae; outer plate with spine teeth distally; palp ended with strong teeth. Maxilla 2 (Fig. 17 G): the two plates are densely setosed apically. Maxilliped (Fig. 18 A): inner plate subrectangular, elongated, characterized by the presence of two strong teeth and long setae on the distal part (Fig. 18 B); internal margin of the outer plate with long setae and strong teeth; palp elongate, longer than outer plate, article 2 and 3 with long setae, dactylus elongate, slender and bearing short setae (Fig. 18 C). Pereon. Long setae (broken) are present on the posterodorsal part of the pereon segment 5���7; they are frequently broken, particularly on the holotype (Fig. 15). Gnathopod 1 (Fig. 18 D): coxa 1 roughly rectangular, distal margin rounded and fringed with long setae; basis long, approximatively rectangular with some setae, length a little less than merus + carpus + propodus, the merus, carpus and propodus bearing long setae; propodus oval; dactylus curved, with short setae (Fig.18 E), length of dactylus = �� length of propodus. Gnathopod 2 (Fig. 18 F): coxa 2 rectangular and very short (coxa length / basis length = 34/51), distal margin rounded with long setae; basis very long with small setae, basis = carpus + propodus; carpus longer than propodus (carpus length / propodus length = 32/18), roughtly rectangular with long setae on the margins; propodus roughly rectangular, long setae on the margins and also covered by many short setae; dactylus long and curved, dactylus length> �� propodus length. Pereopod 3 (Fig. 18 G): coxa 3 roughly rectangular, distal margin rounded with some setae; basis rectangular, basis = ischium + merus. The margins of the different articles of pereopod 3, except dactylus, bearing few long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 18/ 14). Pereopod 4 (Fig. 18H): coxa 4 anterior margin slightly concave, length / width = 26 / 37, anteroventral corner rounded, ventral margin convex with long setae, posterior concavity 1/2 length of the coxa 4, posterior hook length = 1/5 width of coxa 4, posterior corner very short and straight (Fig. 18 J); basis length = merus + carpus; The margins of the different articles of pereopod 4, except dactylus, bearing many long setae; dactylus slender, curved and longer than propodus (dactylus length / propodus length = 18/ 16). Pereopod 5 (Fig. 19 A): coxa 5 roughly oval, posterior margin rounded; basis pyriform, anterior margin with few short setae; merus anterior margin with short spines; carpus rectangular, anterior margin with strong spines, posterodistal lobe with a long spine; propodus rectangular, shorter than carpus, with strong spines on the anterior margin and short spines on the posterior one; dactylus slender and curved, dactylus length / propodus length =7/16. Pereopod 6 (Fig. 19 B): coxa 6 roughly rectangular, posterior margin rounded; basis oval with few short setae on the anterior margin; carpus rectangular, anterior margin and posterodistal lobe with distal strong spines; propodus rectangular and slightly curved, a little shorter than carpus, with strong spines and setae on the anterior margin; dactylus slender and curved, dactylus length / propodus length =7/17. Pereopod 7 (Fig. 19 C): coxa 7 roughly rectangular, posterior margin rounded; basis moderatly broad (length without lobe / width = 37 /20), anterior and posterior margin sligthly concave, numerous long setae on inner side, lobe slightly deflected, rounded with few setae, reaching the middle of merus; merus quadrangular with robust spines on the posterodistal margin; carpus subrectangular (width / length = 8/15), with 5 strong spines on the posterior margin (propodus length / carpus length = 11 /15); propodus + dactylus / carpus = 18/15); propodus narrow, width / length = 2/11 (propodus width / carpus width = 2 /8), with 2 spines on the posterior margin; dactylus triangular, long and narrow (length /width = 6/ 1) with 2 very little apical setae (dactylus length / propodus length = 6 / 11). Pleon. Long setae (broken) are present on the posterodorsal part of the pleon (segment 1���3), they are frequently broken (Figs. 15 & 16). Epimeral plate 1 (Fig. 19 D): anterior margin oblique and straight, anteroventral corner rounded, ventral and posterior margin straight. Epimeral plate 2 (Fig. 19 E): anterior margin scarcely concave, anteroventral corner rounded with two short setae, ventral margin convex, posteroventral corner rounded, posterior margin convex. Epimeral plate 3 (Fig. 19 F); anterior margin straight, anteroventral corner roughly rounded with 3 short setae, ventral margin straight, posteroventral corner ended with a little acute tooth, posterior margin slightly convex. Urosome (Figs. 15, 16): the urosome segment 1 has a dorsal carina straight, moderately hight with very small setae; a small dorsolateral process is present on each side of urosome segment 2. Uropod 1 (Fig. 19 G): long, rami slender of inequal length, inner ramus slightly shorter than outer ramus (39 / 41); inner ramus with 5 spines; outer ramus with 2 spines; peduncle longer than rami (rami length / peduncle length = 39���41 / 47), with 4 strong spines and 2 long setose setae. Uropod 2 (Fig. 19H): shorter than uropod 1; rami short and subequal, shorter than peduncle (rami length / peduncle length = 30���32/ 35); strong spines on rami, strong spines and setae on peduncle. Uropod 3 (Fig. 19 J): peduncle short and strong (length / width = 22 / 15); rami of equal length, foliaceous, longer than peduncle (rami length/ peduncle length = 31 / 22); inner ramus with long apical setose setae and few short spines; outer ramus with long setose setae on the margins and the apical part. Telson (Fig. 19 K): triangular rounded, slightly longer than wide (length / width = 35 / 30), cleft on 26/35 of the length, two short setae and one longer are present on the apical part of each lobe. In summary, the new species Haploops islandica, is morphologically near to the three other species studied in this work, but can be distinguished by its long and strong setose antennae, the very small rectangular coxa 2, very long dactylus of the Pereopods 3 and 4, the coxa 4 anterior margin slightly concave, and a triangular rounded, slightly longer than wide telson with two short setae and one longer present on the apical part of each lobe. Male unknown. Distribution. Wide bathymetric range species: 283���1727 m. Present in a wide variety of sediment: sandy silt, silty sand, coarse sand, cobbles and stones, with sediment including pieces of corals, sponges, and echinoderms., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566

Key to species of Haploops 1 Corneal lenses present................................................................................. 2 - Corneal lenses absent................................................................................ 14 2 Two pairs of corneal lenses............................................................................ 3 - Only one pair of corneal lenses...........................................................................7 3 Inferior lenses in the middle of the lobe head..................................... Haploops sibirica Gurjanova, 1929 - Inferior lenses near or on the margin head..................................................................4 4 Inferior lenses near the margin head.......................................... Haploops dellavallei Chevreux, 1900 - Inferior lenses on the margin............................................................................. 5 5 Short down-like setae on the back, coxa 1 strongly truncated............................. Haploops laevis Hoek, 1882 - These characters absent................................................................................ 6 6 Inferior lenses on the margin, antennae =body length............................. Haploops nirae Kaïm-Malka, 1976 - Inferior lenses on the internal side of the margin, antennae = 1/2 body lenth........... Haploops tenuis Kanneworff, 1966 7 Only superior lenses present (inferior absent).............................................................. 8 - Only inferior lenses present (superior absent).............................................................. 13 8 A1 A 2........................................................................................... 12 9 A1 length A 2 peduncle............................................................................. 11 10 A1 length Haploops descansa Barnard, 1961 - A1 length = A 2 peduncle, head transversely truncated.......................... Haploops oonah Lowry & Poore, 1985 11 A 2 = 1 / 3 body................................................. Haploops fundiensis Wildish & Dickinson, 1982 - A 2 = 1 / 2 body............................................................ Haploops tubicola Lilljborg, 1856 12 A 2> body, head truncated straight................................ Haploops gascogni Dauvin & Bellan-Santini, 1996 - A 2 =1 / 2 body, head transversely truncated................................. Haploops antennata Kaïm-Malka, 2012 13 Long setae on Md, back, telson, A1 Haploops longiseta Kaïm-Malka, 2010 - Few long dorsal setae, A1> A 2.............................................. Haploops proxima Chevreux, 1919 14 Pereopod 7 basis narrow.............................................................................. 15 - Pereopod 7 basis broad............................................................................... 19 15 Uropod 1, rami equal................................................................................. 16 - Uropod 1, rami unequal.............................................................................. 18 16 Dorsal side of pleon and urosome carinate..................................... Haploops vallifera Stephensen, 1925 - Dorsal side of pleon and urosome not carinate............................................................. 17 17 P7 not elongate, basis lobe reaching longer than ischium............. Haploops antarctica Bellan-Santini & Dauvin, 2008 - P7 elongate, basis lobe reaching only the end of ischium (male only)............... Haploops abyssorum Chevreux, 1908 18 A1 = A 2, Coxa 4 heart-shaped............................................... Haploops similis Stephensen, 1925 - A1 = A 2 peduncle, coxa 4 not heart shaped.......................................... Haploops lodo Barnard, 1961 19 Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe triangular.................... Haploops setosa Boeck, 1870 - Long dorsal tuft of setae (pereon 5–7 & pleon), P7 basis lobe rounded.......................................... 20 20 Urosome 1 high upturned carina.............................................. Haploops carinata Lilljeborg, 1855 - Urosome 1 carina straight............................................................................. 21 21 Antenna length> body length, antennae very strong...................................... Haploops islandica n. sp. - Antenna length =1/2 body length............................................... Haploops robusta G.O.Sars, 1891 19' Short posterodorsal setae (Pl 2–3 & U 1).............................. H. meloi Valerio-Berardo, 2008 (dubious species)

Kaim-Malka, R. A., Bellan-Santini, D., Dauvin, C. (2016): On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.]. Zootaxa 4179 (1), DOI: http://doi.org/10.11646/zootaxa.4179.1.2

Key to species of Haploops 1 Corneal lenses present................................................................................. 2 - Corneal lenses absent................................................................................ 14 2 Two pairs of corneal lenses............................................................................ 3 - Only one pair of corneal lenses...........................................................................7 3 Inferior lenses in the middle of the lobe head..................................... Haploops sibirica Gurjanova, 1929 - Inferior lenses near or on the margin head..................................................................4 4 Inferior lenses near the margin head.......................................... Haploops dellavallei Chevreux, 1900 - Inferior lenses on the margin............................................................................. 5 5 Short down-like setae on the back, coxa 1 strongly truncated............................. Haploops laevis Hoek, 1882 - These characters absent................................................................................ 6 6 Inferior lenses on the margin, antennae =body length............................. Haploops nirae Ka��m-Malka, 1976 - Inferior lenses on the internal side of the margin, antennae = 1/2 body lenth........... Haploops tenuis Kanneworff, 1966 7 Only superior lenses present (inferior absent).............................................................. 8 - Only inferior lenses present (superior absent).............................................................. 13 8 A1 A 2........................................................................................... 12 9 A1 length A 2 peduncle............................................................................. 11 10 A1 length Haploops descansa Barnard, 1961 - A1 length = A 2 peduncle, head transversely truncated.......................... Haploops oonah Lowry & Poore, 1985 11 A 2 = 1 / 3 body................................................. Haploops fundiensis Wildish & Dickinson, 1982 - A 2 = 1 / 2 body............................................................ Haploops tubicola Lilljborg, 1856 12 A 2> body, head truncated straight................................ Haploops gascogni Dauvin & Bellan-Santini, 1996 - A 2 =1 / 2 body, head transversely truncated................................. Haploops antennata Ka��m-Malka, 2012 13 Long setae on Md, back, telson, A1 Haploops longiseta Ka��m-Malka, 2010 - Few long dorsal setae, A1> A 2.............................................. Haploops proxima Chevreux, 1919 14 Pereopod 7 basis narrow.............................................................................. 15 - Pereopod 7 basis broad............................................................................... 19 15 Uropod 1, rami equal................................................................................. 16 - Uropod 1, rami unequal.............................................................................. 18 16 Dorsal side of pleon and urosome carinate..................................... Haploops vallifera Stephensen, 1925 - Dorsal side of pleon and urosome not carinate............................................................. 17 17 P7 not elongate, basis lobe reaching longer than ischium............. Haploops antarctica Bellan-Santini & Dauvin, 2008 - P7 elongate, basis lobe reaching only the end of ischium (male only)............... Haploops abyssorum Chevreux, 1908 18 A1 = A 2, Coxa 4 heart-shaped............................................... Haploops similis Stephensen, 1925 - A1 = A 2 peduncle, coxa 4 not heart shaped.......................................... Haploops lodo Barnard, 1961 19 Long dorsal tuft of setae (pereon 5���7 & pleon), P7 basis lobe triangular.................... Haploops setosa Boeck, 1870 - Long dorsal tuft of setae (pereon 5���7 & pleon), P7 basis lobe rounded.......................................... 20 20 Urosome 1 high upturned carina.............................................. Haploops carinata Lilljeborg, 1855 - Urosome 1 carina straight............................................................................. 21 21 Antenna length> body length, antennae very strong...................................... Haploops islandica n. sp. - Antenna length =1/2 body length............................................... Haploops robusta G.O.Sars, 1891 19' Short posterodorsal setae (Pl 2���3 & U 1).............................. H. meloi Valerio-Berardo, 2008 (dubious species), Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Gurjanova, E. F. (1929) Neue Forrnen arktischer Isopoden und Amphipoden. Zoologischer Anzeiger, 81, 309 - 317.","Chevreux, E. (1900) Amphipodes provenant des campagnes de l' Hirondelle (1885 - 1888). Resultats des Campagnes Scientifiques Accomplies par le Prince Albert I. Monaco. 16, 195 pp.","Hoek, P. P. C. (1882) Die Crustaceen, gesammelt waehrend der Fahrten des \" Willem Barents \" in den Jahren 1878 und 1879, Niederlandisches Archiv fur Zoologie, Supplementband I, 7, p 61.","Kaim-Malka, R. A. (1976) Revision des Haploops (Crustacea, Amphipoda) de Mediterranee. Bolletino del Museo Civico di Storia Naturale, Verona, 3, 269 - 308.","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207. http: // dx. doi. org / 10.1080 / 00785326.1966.10409642","Barnard, J. L. (1961) Gammaridean Amphipoda from depths of 400 to 6000 meters. Galathea Report, 5, 23 - 128.","Lowry, J. K. & Poore, G. C. B. (1985) The ampeliscid amphipods of south-eastern Australia (Crustacea). Records of the Australian Museum, 36, 259 - 298. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.36.1985.348","Wildish, D. J. & Dickinson, J. J. (1982) A new species of Haploops (Amphipoda, Ampeliscidae) from the Bay of Fundy. Canadian Journal of Zoology, 60, 962 - 967.","Dauvin, J. C. & Bellan-Santini, D. (1996) Ampeliscidae (Amphipoda) from the Bay of Biscay. Journal of Crustacean Biology, 16, 149 - 168. http: // dx. doi. org / 10.1163 / 193724096 X 00351","Kaim-Malka, R. A. (2012) Haploops antennata, a new species from the North Atlantic Ocean (Crustacea: Gammaridea: Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 7.]. Zootaxa, 3320, 36 - 46.","Kaim-Malka, R. A. (2010) Haploops longiseta, a new species from the Atlantic Ocean (Crustacea, Gammaridea, Ampeliscidae). [Contribution to the knowledge of the Haploops genus. 6.]. Zootaxa, 2356, 57 - 68.","Chevreux, E. (1919) Note preliminaire sur les Amphipodes recueillis par les expeditions du \" Travailleur \" et du \" Talisman \" (1880 - 1883). Bulletin du Museum d'Histoire Naturelle, 25, 574 - 580. http: // dx. doi. org / 10.5962 / bhl. part. 7932","Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Chevreux, E. (1908) Diagnoses d'amphipodes nouveaux provenant des campagnes de la Princesse-Alice dans l'Atlantique nord. Bulletin de l'Institut Oceanographique, Monaco, l 17, 1 - 13.","Boeck, A. (1870) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Videnskabs - Selskabet i Christiania 1870, pp. 83 - 280. http: // dx. doi. org / 10.5962 / bhl. title. 2056","Lilljeborg, W. (1855) Om Hafs-Crustaceer vid Kullaberg i Skane. Ofversigt af Kongliga Vetenskaps - Akademien Forhandlingar (Stockholm 1856), 12, 117 - 138.","Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway With Short Descriptions and Figures of All the Species, 1, 191 - 196, pls. 67 - 68.","Valerio-Berardo, M. T. (2008) Six new species of deep water Ampeliscidae (Crustacea: Amphipoda) from Brazilian Coast. Zootaxa, 1844, 1 - 24."]}

Family Ampeliscidae Costa, 1857 Diagnosis. Head very large. Eyes when present composed of internal pigment masses served by 2���4 external cuticular lenses. Accessory flagellum absent. Gnathopods feeble. Article 4 of pereopods 3���4 elongate, article 6 much shorter than 4 and article 5 much shorter than 6, these pereopods glandular. Pereopods 5���6 alike but pereopod 7 of distinct structure, article 2 with distinct, usually broad posteroventral lobe, article 2 of pereopods 5��� 6 rhomboid or diamond shaped and poorly lobed. Urosomites 2���3 coalesced. Uropod 3 biramous. Telson laminar (Barnard & Karaman, 1991)., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Costa, A. (1857) Ricerche sui crostacei amfipodi del regno di Napoli. Memorie della Reale Accademia de Scienze di Napoli, 1, 165 - 235, 4 pls.","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, supplement 13, part 1, 84 - 90."]}

Haploops robusta G.O. Sars, 1891 Haploops robusta G.O.Sars, 1891. Haploops robusta G.O.Sars: in Stebbing, 1906; in Stephensen 1925; in Gurjanova 1951. nec Haploops setosa Boeck: in Dunbar 1954; in Kanneworff 1966; in Mills 1971; in Barnard& Karaman 1991. nec Haploops setosa f. sarsi Schellenberg 1925: in Barnard & Karaman 1991. nec Haploops sibirica Gurjanova, 1929; in Barnard & Karaman, 1991. Bioice material (Fig. 1). Station 3208, 1 specimen, 9 July 2001, depth: 2002���2004 m, 65�� 43' 93'' N���7�� 00' 95'' W. Station 3539, 1 specimen, 10 September 2002, depth: 1377��� 1370 m, 61�� 59' 63'' N���13�� 33' 09'' W. Station 3656, 1 specimen, 23 July 2004, depth: 1492��� 1490 m, 68�� 47' 11'' N���15�� 18' 67'' W. Description. Adult female with oostegites not completely developped. Length: 12.85 mm (Fig. 11). Bioice 3656, 23 July 2004, depth: 1490���1492 m, 68�� 47' 11'' N ��� 15�� 18' 67'' W. Body with long dorsal setae (broken) on the pereon, and pleon (see discussion)(Fig. 11) Head: nearly square, with a rostrum but without corneal lenses, blind species. The anterior margin is convex (Fig. 12 A). Antenna 1: near as long as antenna 2, �� body length. Peduncle of A1 with article 3 Antenna 2: a little longer than antenna 1, �� body length. Two teeth are present on the peduncle segment 2 (Fig. 12 C). Peduncle segment 5 is slightly longer than 4 (51 / 47). Flagellum with 38 articles. Antenna 1 and Antenna 2 are densely setose with very long setae. Their peduncles are long and strong. Upper lip (Fig. 12 J): without hair. Mandible (Fig. 12 D): molar strong; palp very long, article 2 slightly longer than article 3, each article bears long setae. Article 3 with 4 long apical setae. Lower lip bilobed. Maxilla 1 (Fig. 12 G): the inner plate is conical shaped with two sub apical setae; outer plate with spine teeth distally; palp ended with some setae and strong teeth. Maxilla 2 (Fig. 12 H); the two plates are subequal in length and apically densely setose. Maxilliped (Fig. 12 E): inner plate subrectangular, elongate, characterized by the presence of two strong teeth and setae on the distal part (Fig. 12 F); internal margin of the outer plate with numerous long setae; palp elongate, longer than outer plate, article 2 and 3 with long setae, dactylus elongate and slender. Pereon. Long setae (broken) are present on the posterodorsal part of the pereon segment 6 and 7 (Fig. 11). Gnathopod 1 (Fig. 13 A): coxa 1 shorter than basis, roughly rectangular, distal margin rounded and fringed with long setae; basis very long and slightly curved, basis = merus + carpus + propodus, merus, carpus and propodus bear very long setae; propodus roughly rectangular and cover by numerous short setae; dactylus curved, with short spines, in the specimen studied a very strong subapical tooth was present, the size of this tooth was very different between the right dactylus (Fig. 13 B) and the left one (Fig. 13 C), the origin of this dimorphism cannot be given. Gnathopod 2 (Fig. 13 D): coxa 2 is roughly rectangular, the distal margin is rounded with some short setae. Basis long with long setae, basis = ischium + merus + carpus; carpus longer than propodus (36/18), roughly rectangular with long setae on the margins; propodus subrectangular, covered by numerous short setae; dactylus curved and ornamented with setae and spines (Fig. 13 E). Pereopod 3 (Fig. 13 F): coxa 3 is roughly rectangular, distal margin rounded with few setae; basis rectangular, basis = ischium + merus; the margins of the different articles of pereopod 3 bear few long setae, except dactylus; dactylus slender, curved and slightly longer than propodus. Pereopod 4 (Fig. 13 G): coxa 4 wide, length / width = 25 / 36, the anterior margin of coxa 4 is divided in two part, the 1/3 superior part is straight, the 2/3 inferior part is directed obliquely towards the head, anteroventral corner very slightly rounded, ventral margin convex with long setae, posterior concavity 2/3 length of the coxa 4, posterior hook length = 1/7 width of coxa 4, corner rounded; basis roughly rectangular, a little longer than ischium + merus; dactylus slender and slightly curved, a little longer than the propodus; the margins of the different articles of pereopod 4, except dactylus, bear long setae. Pereopod 5 (Fig. 13H): coxa 5 roughly triangular; basis oval, anterior margin fringed with short setae; carpus rectangular with a posterodistal lobe, anterior margin with long setae, posterior margin and distal margin ornamented with rows of spines; propodus rectangular, as long as carpus, with long strong spines on the anterior margin and a very long spine on the distal part; dactylus strong and curved, dactylus length = �� length of propodus. Pereopod 6 (Fig.14 A): the structure is the same that pereopod 5, long strong spines on the distal part of the carpus and the propodus. Pereopod 7 (Fig.14 B): coxa 7 bilobed; basis moderatly broad (length without lobe / width = 32 / 18), anterior and posterior margins straight, numerous long setae on inner side, external side glabrous; lobe not deflected, rounded and setose, reaching the middle of merus; ischium and merus quadrangular with strong spines on the posterior margin; carpus suboval (width / length = 8 / 13), with 5 strong spines on the posterior margin (propodus / carpus = 10 /13; propodus + dactylus / carpus = 17 / 13); propodus narrow: width / length = 3 / 10 (propodus width / carpus width = 3 / 8), with 2 strong spines on the posterior margin and on the distal margin; dactylus very long and narrow (length /width = 32 / 4) with 2 little apical setae (dactylus length / propodus length = 6 /10) (Fig. 14 C). Pleon. Long setae (broken) are present on the posterodorsal part of the pleon segment 1 and 2 (Fig. 11). Epimeral plate 1 (Fig. 14 D): ventral margin rounded, with a little seta. Epimeral plate 2 (Fig. 14 E): ventral margin straight, with a little spine. Epimeral plate 3 (Fig. 14 F): anterior margin straight, anteroventral corner rounded with 4 setae, ventral margin very slightly convex, posteroventral corner ended with a little acute tooth, posterior margin convex. Urosome (Fig. 14 L): the urosome segment 1 has a small dorsal convex carina; a small dorsolateral process is present on each side of urosome segment 2. Uropod 1 (Fig. 14 G) moderatly long, the rami are subequal (inner ramus length /outer ramus length = 5 / 5.5); outer ramus slightly curved. Each ramus bears strong spines; peduncle longer than the rami (rami length / peduncle length = 5���5.5 / 6.5), ornamented with strong spines on each margin and also with a row of four spines in its middle part. Uropod 2 (Fig. 14H): shorter than uropod 1; rami subequal and shorter than peduncle (rami length / peduncle length = 58 / 75); long setae on peduncle, very long spines on peduncle and rami (peduncle partly broken). Uropod 3 (Fig. 14 J): rami subequal, more or less triangular, longer than peduncle (rami length / peduncle length = 5.5 / 4); inner ramus with long setae on each margin; outer ramus with short spines and setae on each margin and some long apical setae; peduncle short and strong (length / width = 4 / 2.7) Telson (Fig. 14 K): triangular rounded, longer than wide (length / width =110 / 85), cleft on 7/11 of the length, three apical setae of different size are present on each lobe, the longuest is 8/11 the length of the telson. Male unknown. Distribution in the examined collection. Deep species: 1370���2004 m. Present on deep sandy-silt substrates, in the same habitats than the decapod Bythocaris spp., the mysid Boreomysis spp. and the fish of the genera Lycodes and Paraliparis., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway With Short Descriptions and Figures of All the Species, 1, 191 - 196, pls. 67 - 68.","Stebbing, TRR. (1906) Amphipoda 1. Gammaridea. Das Tierreich, 21, 806 pp., 127 figs.","Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Gurjanova, E. F (1951) Bokoplavy Morei SSSR e sopredelny vod (Amphipoda- Gammaridea). Akademiia Nauk SSSR, Opedeliteli po Faune SSSR, 41, 1029 pp, 705 figs.","Dunbar, M. J. (1954) The amphipod Crustacea of Ungava Bay, Canadian Eastern Arctic. Journal of Fisheries Research Board of Canada, 11, 721 - 723. http: // dx. doi. org / 10.1139 / f 54 - 044","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207. http: // dx. doi. org / 10.1080 / 00785326.1966.10409642","Mills, E. L. (1971) Deep-Sea Amphipoda from the Western North Atlantic Ocean, the family Ampeliscidae. Limnology and Oceanography, 16, 3377 - 386. http: // dx. doi. org / 10.4319 / lo. 1971.16.2.0357","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, supplement 13, part 1, 84 - 90.","Schellenberg, A. (1925) Die Gammariden Spitzbergens nebst einer Uebersicht der von Romer & Schaudinn 1898 im nordlichen Eismeer gesammelten Arten. Mittheilungen ausdem Zoologischen Zoologischen Museum en Berlin, 11, 195 - 231, 10 figs.","Gurjanova, E. F. (1929) Neue Forrnen arktischer Isopoden und Amphipoden. Zoologischer Anzeiger, 81, 309 - 317."]}

Haploops setosa Boeck, 1870 Haploops setosa Boeck 1870 Haploops setosa Boeck in Stebbing 1906 (in part); in Stephensen 1925 (in part); in Kanneworff 1966 (in part). Haploops setosa f. sarsi Schellenberg 1925, Gurjanova 1951 in part. nec Haploops setosa Boeck 1870: in G.O.Sars, 1891; in Kanneworff 1966 (as H.setosa Sars); in Mills 1971. nec Haploops robusta G.O.Sars, 1891: in Dunbar 1954; in Mills 1971; in Dickinson 1983; in Barnard & Karaman, 1991. nec Haploops sibirica Gurjanova, 1929; in Barnard & Karaman, 1991., Published as part of Kaim-Malka, R. A., Bellan-Santini, D. & Dauvin, C., 2016, On some Haploops species collected in the North Atlantic Ocean with the description of Haploops islandica n. sp. (Crustacea: Gammaridea: Ampeliscidae) [Contribution to the knowledge of the Haploops genus. 8.] in Zootaxa 4179 (1), DOI: 10.11646/zootaxa.4179.1.2, http://zenodo.org/record/260566, {"references":["Boeck, A. (1870) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Videnskabs - Selskabet i Christiania 1870, pp. 83 - 280. http: // dx. doi. org / 10.5962 / bhl. title. 2056","Stebbing, TRR. (1906) Amphipoda 1. Gammaridea. Das Tierreich, 21, 806 pp., 127 figs.","Stephensen, K. (1925) Crustacea Malacostraca. VI. (Amphipoda. Il). Danish Ingolf-Expedition, 3, 101 - 178.","Kanneworff, E. (1966) On some amphipod species of the genus Haploops, with special reference to H. tubicola Liljeborg and H. tenuis sp. nov. from the Oresund. Ophelia, 3, 183 - 207. http: // dx. doi. org / 10.1080 / 00785326.1966.10409642","Schellenberg, A. (1925) Die Gammariden Spitzbergens nebst einer Uebersicht der von Romer & Schaudinn 1898 im nordlichen Eismeer gesammelten Arten. Mittheilungen ausdem Zoologischen Zoologischen Museum en Berlin, 11, 195 - 231, 10 figs.","Gurjanova, E. F (1951) Bokoplavy Morei SSSR e sopredelny vod (Amphipoda- Gammaridea). Akademiia Nauk SSSR, Opedeliteli po Faune SSSR, 41, 1029 pp, 705 figs.","Sars, G. O. (1891) Amphipoda. An Account of the Crustacea of Norway With Short Descriptions and Figures of All the Species, 1, 191 - 196, pls. 67 - 68.","Mills, E. L. (1971) Deep-Sea Amphipoda from the Western North Atlantic Ocean, the family Ampeliscidae. Limnology and Oceanography, 16, 3377 - 386. http: // dx. doi. org / 10.4319 / lo. 1971.16.2.0357","Dunbar, M. J. (1954) The amphipod Crustacea of Ungava Bay, Canadian Eastern Arctic. Journal of Fisheries Research Board of Canada, 11, 721 - 723. http: // dx. doi. org / 10.1139 / f 54 - 044","Dickinson, J. J. (1983) The systematics and distributional ecology of the superfamily Ampeliscoidea (Amphipoda: Gammaridea) in the northeastern Pacific region. II. The genera Byblis and Haploops. Publications in Natural Sciences, National Museum of Natural Sciences, 1, 1 - 138, 17 figs, 6 tables.","Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine Gammaridea Amphipod (except marine Gammaroid). Records of the Australian Museum, supplement 13, part 1, 84 - 90.","Gurjanova, E. F. (1929) Neue Forrnen arktischer Isopoden und Amphipoden. Zoologischer Anzeiger, 81, 309 - 317."]}

Comment on: "Pediatric-type" gastrointestinal stromal tumors in adults: distinctive histology predicts genotype and clinical behavior. Rege TA, Wagner AJ, Corless CL, Heinrich MC, Hornick JL. Am J Surg Pathol. 2011 Apr; 35(4):495-504.

Nanotechnology-based drug delivery was born as a chance for pharmaceutical weapons to be delivered in the body sites where drug action is required. Specifically, the incorporation of anti-cancer agents in nanodevices of 100-300 nm allows their delivery in tissues that have a fenestrated vasculature and a reduced lymphatic drainage. These two features are typical of neoplastic tissues and, therefore, allow the accumulation of nanostructured devices in tumours. An important issue of anti-cancer pharmacological strategies is the overcoming of anatomical barriers such as the blood-brain-barrier (BBB) that protects brain from toxicological injuries but, at the same time, makes impossible for most of the pharmacological agents with anti-cancer activity to reach tumour cells placed in the brain and derived from either primary tumours or metastases. In fact, only highly lipophilic molecules can passively diffuse through BBB to reach central nervous system(CNS). Another possibility is to use nanotechnological approaches as powerful tools to across BBB, by both prolonging the plasma half-life of the drugs and crossing fenestrations of BBB damaged by brain metastases. Moreover, modifications of nanocarrier surface with specific endogenous or exogenous ligands can promote the crossing of intact BBB as in the case of primary brain tumours. This aim can be achieved through the binding of the nanocarriers to carriers or receptors expressed by the endothelial cells of BBB and that can favour the internalization of the nanostructured devices delivering anti-cancer drugs. This review summarizes the most meaningful advances in the field of nanotechnologies for brain delivery of drugs.

Renal cell carcinoma is the most common type of kidney cancer in adults. It accounts for approximately 3% of adult malignancies and 90–95% of neoplasms arising from the kidney. At the moment several biological agents are used for the treatment of metastatic renal cell carcinoma. We describe the case of a man who has been treated with pazopanib (Votrient) for metastatic renal cell carcinoma since July 2011. At the time of writing, the patient is still receiving treatment (29 months) and is showing a long-lasting response with a favorable safety profile. This is an excellent example of chronic neoplastic disease in a patient who can be defined as long-surviving.

INTRODUCTION Angiogenesis has been correlated with increased invasion and metastases in a variety of human neoplasms. Inadequate inhibition of the growth of tumor microvessels by anticancer agents may result in treatment failure, rated clinically as progressive or stable disease. We designed this trial to investigate the modification of the vascular endothelial growth factor (VEGF) and interferon-gamma (IFN-gamma) in advanced colorectal cancer patients during treatment with a weekly combination of cetuximab plus irinotecan. MATERIALS AND METHODS Forty-five metastatic colorectal cancer patients were prospectively evaluated for circulating levels of VEGF and IFN-gamma during the treatment with cetuximab (initial dose of 400 mg/m(2), followed by weekly infusions of 250 mg/m(2)) plus weekly irinotecan (90 mg/m(2)). The circulating levels of the cytokines were assessed at the following time points: just before and at 1, 21, 50 and 92 days after the start of cetuximab plus irinotecan treatment. RESULTS Basal serum VEGF median levels were significantly decreased just at the first day (after the first treatment infusion (P = 0.016). The VEGF persisted at the following time points reaching the highest statistical significance 92 days after the first infusion (P < 0.0001). On the contrary, IFN-gamma values showed a statistical significant increase one day after the first infusion (P < 0.0001). This effect persisted 21 days after the treatment start (P = 0.001), but was no more evident at the following time points. Moreover, a linear regression model with variance analysis showed a significant negative correlation between VEGF and IFN-gamma values 1, 21 and 50 days after the treatment beginning (P = 0.002, 0.001 and 0.047, respectively). CONCLUSIONS This study suggests that a cetuximab may induce a modulation of VEGF circulating levels. The reduction of VEGF serum levels is a sudden and long lasting phenomenon. Moreover, in our study we identified a IFN-gamma increase, even if the specific role of this behavior remains to be investigated.

To assess sonographically brain anatomy in fetuses with severe anemia due to red blood cell alloimmunization undergoing intrauterine intravascular transfusions.Multiplanar neurosonography was performed in seven consecutive hydropic fetuses undergoing intrauterine transfusions (mean gestational age 22 +/- 2.5 weeks; mean hemoglobin concentration at the first transfusion 2.3 +/- 1.0 g/dL).Abnormal cerebral findings were identified in four out of seven fetuses. An intracerebellar hemorrhage developed in two fetuses after the first transfusion and one fetus that had severe brain edema before the first transfusion was later found to have cystic periventricular leukomalacia. In one fetus unilateral ventriculomegaly was noted after the first transfusion. Two fetuses were terminated. The remaining pregnancies had an uneventful course, the infants were delivered between 34 and 36 gestational weeks and were alive and well at the time of writing. Prenatal diagnosis of brain injury was always confirmed except for the case with ventriculomegaly that underwent spontaneous intrauterine resolution.Fetuses with extreme anemia due to red blood cell alloimmunization can be salvaged by intrauterine transfusion. In some of these cases brain injury may occur prenatally, and the risk seems to be particularly high when the hemoglobin concentration at the time of the first transfusion isor= 2 g/dL. We suggest that in these pregnancies detailed fetal neuroimaging by either multiplanar sonography and/or magnetic resonance imaging is indicated.

In order to identify genes relevant for melanoma development, we carried out cDNA array experiments employing an in vitro model of human melanoma progression, consisting of two cell lines: one, LP, derived from a primary melanoma and the other, LM, from its metastatic supraclavicular lymph node. Basic cDNA array data identified 26 genes as down-regulated in the LM cell line. Northern blot analysis confirmed an effective transcriptional down-regulation for five out of 13 genes analyzed. The products of these five genes belong to different functional protein types, such as transcription and translation regulators (Edg-2, eIF-3 p110, and RNPL/RBM3), extracellular communicators (PRSS11) and members of the major histocompatibility complex (β2-microglobulin). Some previously described differences in expression patterns, such as loss of HLA I, were confirmed by our array data. In addition, we identified and validated for the first time the reduced expression level of several genes during melanoma progression. In particular, reduced Edg-2 gene product expression was also confirmed in a group of 50 primary melanomas and unrelated metastases. In conclusion, comparative hybridization by means of cDNA arrays assisted in identifying a series of novel progression-associated changes in gene expression, confirming, at the same time, a number of previously described results. © Blackwell Munksgaard, 2003.

Cell adhesion is a basic count in inter- and intracellular communication and plays an important role in tumor progression. In this study, the expression of E-selectin, ICAM-1 and VCAM-1 were evaluated by means of immunohistochemistry in a group of 153 lung cancer specimens. E-selectin immunoreactivity was localized mostly on endothelial cell venules and capillaries with an average staining intensity of 75% of cells in the NSCLC, while in SCLC the intensity of the staining was 69%. The staining pattern for ICAM-1 reached an average intensity of 57%, in both NSCLC and SCLC. Finally, VCAM-1 immunoreactivity was detected only in NSCLC with an average intensity of 12% on endothelial cell venules and capillaries. This study provides a contribution towards the understanding of the basic mechanisms of cell adhesion in lung cancer progression.

The first cyclin dependent kinase inhibitor to be discovered was the p21 cdk interacting protein (a.k.a., WAF1, Cip1, CAP20, Sdi1, mda6). p21 expression may or may not be dependent on p53. This pathway also inhibits DNA replication by merit of p21's interaction with PCNA, but it has also been shown that this same inhibitory interaction with p21 does not affect PCNA DNA repair abilities. We assessed the immunohistochemical expression of p21 protein in 60 curative surgical resected non small cell lung cancers relating it to the expression of PCNA to clarify the contribution of the p21/PCNA pathway to the development of NSCLC. We did not find any relationship between PCNA and p21 expression. This last result may indicate that the mechanism by which PCNA controls the DNA repair is the most important activity of this protein during lung cancer progression and development, compared to its contribution to cell proliferation. In fact, this last event is strongly counteracted by p21 expression, which in this last case works as an inhibitor of PCNA expression. In conclusion this study highlighted the important role of the p21/PCNA pathway in lung carcinogenesis, pointing out the contribution of PCNA to the response to lung aggression and not only it's role as a proliferation index. Therefore, these results offer a background to further study to evaluate potential novel therapeutic approaches to lung cancer treatment.

Recently, reverse transcriptase-polymerase chain reaction (RT-PCR) for the detection of circulating tumor cells has been suggested as a potential technique for staging cancer. In this report, 43 melanoma patients (including 4 in situ melanoma patients) were tested for tyrosinase mRNA in blood by RT-PCR. All patients had melanoma thinner than 1.5 mm (stage I). Circulating melanoma cells were detected in 8 (18.6%) out of 43 MM patients tested: 5 (16.1%) of 31 patients with melanoma thinner than 0.76 mm and 3 (42.8%) out of 7 patients with melanoma thicker than 0. 76 mm. Moreover, in the tyrosinase-negative group we found only 4/31 patients (13%) with histologic signs of regression, but in the tyrosinase-positive group, 3 out of 8 patients (37.5%) showed, at histologic examination, signs of regression. At the time of this analysis all the patients enrolled (tyrosinase-negative and tyrosinase-positive ones) were free of disease, probably due to the short median time of follow-up after the inclusion in the study. The presence of regression is an important cause of melanoma understaging and the tyrosinase test could represent an effective tool in order to achieve a realistic staging in this subgroup of melanoma patients. Probably, maximum sensitivity of the diagnostic RT-PCR approach to monitor MM patients with either localized or advanced disease could be achieved by using additional markers expressed with high frequencies in melanoma. We propose that one such marker could be the sign of regression.

Viral diseases are reported to cause several detrimental effects on grapevine. Leafroll, due to single or mixed infection of GLRaV-1 and GLRaV-3, and Rugose wood associated to GVA, are considered the most widespread and dangerous. The virus influence on grape qualitative (i.e enological) aspects is still controversial and the grapevine response may vary depending on the specific virus involved or the environmental conditions. The aim of the present study was to deeply investigate the effect due to specific viruses. For this purpose the healthy and the infected descendants of two clones of Nebbiolo (Vitis vinifera L.) were compared in a vineyard. The viruses present in the infected lines were GLRaV1+GVA in the clone 'A' and GLRaV3+GVA in the clone 'B'. During 2008 and 2009, canopy physiological efficiency was investigated as well as field performances, juice composition, berry phenols amount and berry anthocyanin profile. The results indicated the infected plants showed alternate field performances, lower canopy efficiency and an anthocyanin profile poorer of malvidine-3-gucoside.

Twenty Grapevine leafroll-associated virus 1 (GLRaV-1) and Grapevine virus A (GVA) doubly infected plants were identified among progeny of one Vitis vinifera cv. 'Nebbiolo' clone, originally infected with these two viruses. In the same vineyard, 20 healthy plants, derived from thermotherapy sanitation of the doubly-infected original clone, were also chosen as controls. The vineyard was located in Neive (north-western Italy). The sanitary status of each vine was confirmed by ELISA and the virus load was measured by quantitative real time RT-PCR (qRT-PCR). Vegetative behaviour, yield, and grape and wine quality of the vines were compared. The protein pattern of the ripe berries was analysed using 2D electrophoresis and the proteins differentially expressed in relation to the sanitary status were identified by MALDI mass spectrometry.